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<document id="9BBE2BD8442D9B26A4EAD852EEF4AE6D" ID-CLB-Dataset="55598" ID-DOI="10.5281/zenodo.5307016" ID-GBIF-Dataset="ec834b82-4c86-4d14-9a55-44b6a00f9d0e" ID-ISSN="0253-116X" ID-Zenodo-Dep="5307016" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630114470423" checkinUser="marcus" docAuthor="Assing, V. &amp; I, North" docDate="2014" docId="443BC10CA5296A02B8CDFDEEFE18FAB5" docLanguage="en" docName="LinzerbiolBeitr.2014.46.2.1087-1096.pdf" docOrigin="Linzer biologische Beiträge 46 (2)" docStyle="DocumentStyle:24E5EB772D355DE2EE23B1E8D07853E0.9:LinzerbiolBeitr.2006-.journal_article" docStyleId="24E5EB772D355DE2EE23B1E8D07853E0" docStyleName="LinzerbiolBeitr.2006-.journal_article" docStyleVersion="9" docTitle="Echiaster discrepans Assing &amp; I 2014, nov.sp." docType="treatment" docVersion="11" lastPageNumber="1094" masterDocId="B802B974A52F6A05B87FFFDCFFEBFF91" masterDocTitle="Three new species and additional records of Echiaster from Asia (Coleoptera: Staphylinidae: Paederinae)" masterLastPageNumber="1096" masterPageNumber="1087" pageNumber="1093" updateTime="1699156644472" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="2C8C9FEE27B6099EBC300B94E84752AF">Three new species and additional records of Echiaster from Asia (Coleoptera: Staphylinidae: Paederinae)</mods:title>
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<mods:namePart id="963351A4CDC520BF5F381321FA6E350E">Assing, V.</mods:namePart>
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<mods:namePart id="4BD4DBDF9DE84FB135D080125C7DD096">I, North</mods:namePart>
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<mods:title id="6279ED56768F4A064CC3448F0BA81E29">Linzer biologische Beiträge</mods:title>
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<mods:part id="19F52A5CB8246971665FD065475A41E8">
<mods:date id="FA01370741A7F00CB367FCA07CE796FF">2014</mods:date>
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<mods:number id="C08B0C9950D8689BF440A17C03A75542">2014-12-19</mods:number>
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<mods:number id="A02BE8BAF25F3AD285DC0D22D2C953CB">46</mods:number>
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<mods:classification id="65314B27E24BBA64C7F7FBC36970D3A0">journal article</mods:classification>
<mods:identifier id="72D9209AA162AB066B6FD1CB7BEA90C0" type="CLB-Dataset">55598</mods:identifier>
<mods:identifier id="E9285FCFF97DDD296F333917CA8B6CC3" type="DOI">10.5281/zenodo.5307016</mods:identifier>
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<treatment id="443BC10CA5296A02B8CDFDEEFE18FAB5" ID-DOI="http://doi.org/10.5281/zenodo.6536708" ID-GBIF-Taxon="195298159" ID-Zenodo-Dep="6536708" LSID="urn:lsid:plazi:treatment:443BC10CA5296A02B8CDFDEEFE18FAB5" httpUri="http://treatment.plazi.org/id/443BC10CA5296A02B8CDFDEEFE18FAB5" lastPageId="7" lastPageNumber="1094" pageId="6" pageNumber="1093">
<subSubSection id="84882391A5296A03B8CDFDEEFE36FDD9" box="[178,477,562,584]" pageId="6" pageNumber="1093" type="nomenclature">
<paragraph id="CC2D701AA5296A03B8CDFDEEFE36FDD9" blockId="6.[178,687,562,585]" box="[178,477,562,584]" pageId="6" pageNumber="1093">
<heading id="9765C776A5296A03B8CDFDEEFE36FDD9" box="[178,477,562,584]" fontSize="8" level="2" pageId="6" pageNumber="1093" reason="2">
<emphasis id="FEE6AC08A5296A03B8CDFDEEFE36FDD9" bold="true" box="[178,477,562,584]" pageId="6" pageNumber="1093">
<taxonomicName id="0B920B99A5296A03B8CDFDEEFE60FDD9" ID-CoL="38FCH" authority="Assing &amp; I, 2014" authorityName="Assing &amp; I" authorityYear="2014" box="[178,395,562,584]" class="Insecta" family="Staphylinidae" genus="Echiaster" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="1093" phylum="Arthropoda" rank="species" species="discrepans" status="sp. nov.">
<emphasis id="FEE6AC08A5296A03B8CDFDEEFE60FDD9" bold="true" box="[178,395,562,584]" italics="true" pageId="6" pageNumber="1093">Echiaster discrepans</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E5D51173A5296A03B9EEFDEFFE36FDD9" box="[401,477,563,584]" pageId="6" pageNumber="1093" rank="species">nov.sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="84882391A5296A03B99CFDEFFD44FDD8" box="[483,687,563,585]" pageId="6" pageNumber="1093" type="description">
<paragraph id="CC2D701AA5296A03B99CFDEFFD44FDD8" blockId="6.[178,687,562,585]" box="[483,687,563,585]" pageId="6" pageNumber="1093">
<heading id="9765C776A5296A03B99CFDEFFD44FDD8" box="[483,687,563,585]" fontSize="8" level="2" pageId="6" pageNumber="1093" reason="2">
(
<figureCitation id="54A96C9FA5296A03B994FDEFFDB1FDD9" box="[491,602,563,585]" captionStart="Figs 14-19" captionStartId="7.[178,218,935,957]" captionTargetBox="[193,1062,212,917]" captionTargetId="figure-1@7.[178,1082,212,926]" captionTargetPageId="7" captionText="Figs 14-19: Echiaster discrepans nov.sp.: (14) habitus; (15) forebody; (16) male sternite VIII; (17- 19) aedeagus in lateral and in ventral view. Scale bars: 14: 1.0 mm; 15: 0.5 mm; 16-19: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5361375" httpUri="https://zenodo.org/record/5361375/files/figure.png" pageId="6" pageNumber="1093">Figs 14-19</figureCitation>
,
<figureCitation id="54A96C9FA5296A03BA19FDEFFD4EFDD8" box="[614,677,563,585]" captionStart="Map 1" captionStartId="1.[178,223,1240,1261]" captionTargetBox="[178,1085,558,1214]" captionTargetId="figure-142@1.[178,1085,558,1214]" captionTargetPageId="1" captionText="Map 1: Distribution of Echiaster species in Asia. Echiaster discrepans (white circles); E. bivirgatus (white diamonds); E. maior (white triangles); E. paulexcisus (black diamond); E. unicolor (black circles)." figureDoi="http://doi.org/10.5281/zenodo.5361369" httpUri="https://zenodo.org/record/5361369/files/figure.png" pageId="6" pageNumber="1093">Map 1</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="84882391A5296A03B8CDFD84FD4EFD67" pageId="6" pageNumber="1093" type="materials_examined">
<paragraph id="CC2D701AA5296A03B8CDFD84FD4EFD67" blockId="6.[178,1086,600,758]" pageId="6" pageNumber="1093">
Type material:
<materialsCitation id="7CFA7A47A5296A03B9CFFD84FEE2FD0A" ID-GBIF-Occurrence="3765333323" collectionCode="MHNG" collectorName="Besuchet Lobl Mussard" country="India" elevation="2100" location="Kodai Kanal" municipality="Palni H." pageId="6" pageNumber="1093" specimenCount="1" stateProvince="22 Madras" typeStatus="holotype">
<typeStatus id="1329CEB8A5296A03B9CFFD84FE17FDFF" box="[432,508,600,622]" pageId="6" pageNumber="1093" type="holotype">Holotype</typeStatus>
: &quot;
<collectingCountry id="B485308AA5296A03BA4AFD84FD9CFDFC" box="[565,631,600,621]" name="India" pageId="6" pageNumber="1093">INDIA</collectingCountry>
<collectingRegion id="0E56BEF8A5296A03BA00FD84FD08FDFF" box="[639,739,600,622]" pageId="6" pageNumber="1093">22 Madras</collectingRegion>
,
<collectingMunicipality id="2C49EA60A5296A03BA90FD84FCD6FDFC" box="[751,829,600,622]" pageId="6" pageNumber="1093">Palni H.</collectingMunicipality>
,
<location id="C94D26C1A5296A03BB37FD84FC2BFDFF" LSID="urn:lsid:plazi:treatment:443BC10CA5296A02B8CDFDEEFE18FAB5:C94D26C1A5296A03BB37FD84FC2BFDFF" box="[840,960,600,622]" country="India" municipality="Palni H." name="Kodai Kanal" pageId="6" pageNumber="1093" stateProvince="22 Madras">Kodai Kanal</location>
,
<quantity id="0B6ADDFFA5296A03BBB4FD84FBFFFDFC" box="[971,1044,600,621]" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="6" pageNumber="1093" unit="m" value="2100.0">
<elevation id="47BF9729A5296A03BBB4FD84FBFFFDFC" box="[971,1044,600,621]" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="6" pageNumber="1093" unit="m" value="2100.0">2100 m</elevation>
</quantity>
, 11- XI-72,
<collectorName id="616715CCA5296A03B888FDB2FE3DFD15" box="[247,470,622,644]" pageId="6" pageNumber="1093">Besuchet Löbl Mussard</collectorName>
/ Holotypus
<originalDetermination id="EF8CC8E6A5296A03BA0BFDB2FC80FD15" box="[628,875,622,644]" pageId="6" pageNumber="1093">
<taxonomicName id="0B920B99A5296A03BA0BFDB2FCDDFD12" authority="Assing &amp; I, 2014" authorityName="Assing &amp; I" authorityYear="2014" box="[628,822,622,643]" class="Insecta" family="Staphylinidae" genus="Echiaster" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="1093" phylum="Arthropoda" rank="species" species="discrepans" status="sp. nov.">
<emphasis id="FEE6AC08A5296A03BA0BFDB2FCDDFD12" box="[628,822,622,643]" italics="true" pageId="6" pageNumber="1093">Echiaster discrepans</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E5D51173A5296A03BB40FDB3FC80FD15" box="[831,875,623,644]" pageId="6" pageNumber="1093" rank="species">sp.n.</taxonomicNameLabel>
</originalDetermination>
det.
<originalDetermination id="EF8CC8E6A5296A03BBE1FDB3FC16FD15" box="[926,1021,622,644]" pageId="6" pageNumber="1093">V. Assing</originalDetermination>
2014&quot; (
<collectionCode id="AA83E8DFA5296A03B8C7FD59FEE8FD0A" box="[184,259,645,667]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="6" pageNumber="1093" type="Museum">MHNG</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="7CFA7A47A5296A03B96BFD59FC8DFD0B" ID-GBIF-Occurrence="3765333313" box="[276,870,645,667]" collectionCode="MHNG, cAss, cRou" collectorName="Besuchet Lobl Mussard" country="India" elevation="2100" location="Kodai Kanal" municipality="Palni H." pageId="6" pageNumber="1093" specimenCount="5" stateProvince="22 Madras" typeStatus="paratype">
<typeStatus id="1329CEB8A5296A03B96BFD59FE87FD0B" box="[276,364,645,666]" pageId="6" pageNumber="1093" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="DA94BB93A5296A03B904FD59FE5DFD0B" box="[379,438,645,667]" pageId="6" pageNumber="1093" type="generic">5 exs.</specimenCount>
: same data as holotype (
<collectionCode id="AA83E8DFA5296A03BAE1FD59FD0CFD0B" box="[670,743,645,666]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="6" pageNumber="1093" type="Museum">MHNG</collectionCode>
,
<collectionCode id="AA83E8DFA5296A03BA8DFD59FCCBFD0B" box="[754,800,645,666]" pageId="6" pageNumber="1093">cAss</collectionCode>
,
<collectionCode id="AA83E8DFA5296A03BB53FD59FC8BFD0B" box="[812,864,645,666]" pageId="6" pageNumber="1093">cRou</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="7CFA7A47A5296A03BB0EFD59FEA8FD58" ID-GBIF-Occurrence="3765333317" collectingDate="1972-11-15" collectionCode="MHNG, cAss" collectorName="Besuchet Lobl Mussard" country="India" elevation="2150" location="Kodai Kanal" municipality="Palni H." pageId="6" pageNumber="1093" specimenCount="2" stateProvince="27 Madras">
<specimenCount id="DA94BB93A5296A03BB0EFD59FC47FD0B" box="[881,940,645,666]" pageId="6" pageNumber="1093" type="generic">2 exs.</specimenCount>
: &quot;
<collectingCountry id="B485308AA5296A03BBC1FD59FBEBFD0B" box="[958,1024,645,666]" name="India" pageId="6" pageNumber="1093">INDIA</collectingCountry>
N.
<collectingRegion id="0E56BEF8A5296A03BC59FD59FF12FD23" pageId="6" pageNumber="1093">27 Madras</collectingRegion>
,
<collectingMunicipality id="2C49EA60A5296A03B978FD40FEB2FD20" box="[263,345,668,690]" pageId="6" pageNumber="1093">Palni H.</collectingMunicipality>
,
<locationDeviation id="DD9F6938A5296A03B917FD40FE35FD20" box="[360,478,668,690]" pageId="6" pageNumber="1093">
<quantity id="0B6ADDFFA5296A03B917FD40FE4DFD23" box="[360,422,668,690]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="1093" unit="km" value="10.0">10 km</quantity>
N.O.
</locationDeviation>
de
<location id="C94D26C1A5296A03BA76FD40FD6FFD23" LSID="urn:lsid:plazi:treatment:443BC10CA5296A02B8CDFDEEFE18FAB5:C94D26C1A5296A03BA76FD40FD6FFD23" box="[521,644,668,690]" country="India" municipality="Palni H." name="Kodai Kanal" pageId="6" pageNumber="1093" stateProvince="27 Madras">Kodai Kanal</location>
,
<quantity id="0B6ADDFFA5296A03BAEDFD40FD35FD20" box="[658,734,668,690]" metricMagnitude="3" metricUnit="m" metricValue="2.15" pageId="6" pageNumber="1093" unit="m" value="2150.0">
<elevation id="47BF9729A5296A03BAEDFD40FD35FD20" box="[658,734,668,690]" metricMagnitude="3" metricUnit="m" metricValue="2.15" pageId="6" pageNumber="1093" unit="m" value="2150.0">2150 m</elevation>
</quantity>
,
<collectingDate id="A868AF32A5296A03BA92FD40FCA8FD23" box="[749,835,668,690]" pageId="6" pageNumber="1093">
<date id="B82C56DAA5296A03BA92FD40FCA8FD23" box="[749,835,668,690]" pageId="6" pageNumber="1093" value="1972-11-15">15-XI-72</date>
</collectingDate>
,
<collectorName id="616715CCA5296A03BB2DFD40FBDEFD23" box="[850,1077,668,690]" pageId="6" pageNumber="1093">Besuchet Löbl Mussard</collectorName>
&quot; (
<collectionCode id="AA83E8DFA5296A03B8C7FD6FFEEAFD59" box="[184,257,691,712]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="6" pageNumber="1093" type="Museum">MHNG</collectionCode>
,
<collectionCode id="AA83E8DFA5296A03B973FD6FFED7FD58" box="[268,316,691,713]" pageId="6" pageNumber="1093">cAss</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="7CFA7A47A5296A03B931FD6FFDD2FD4E" ID-GBIF-Occurrence="3765333309" collectingDate="1972-11-12" collectionCode="MHNG" collectorName="Besuchet Lobl Mussard" country="India" elevation="2200" location="Kodai Kanal" municipality="Palni H." pageId="6" pageNumber="1093" specimenCount="2" stateProvince="23 Madras">
<specimenCount id="DA94BB93A5296A03B931FD6FFE62FD59" box="[334,393,691,712]" pageId="6" pageNumber="1093" type="generic">2 exs.</specimenCount>
: &quot;
<collectingCountry id="B485308AA5296A03B9E5FD6FFE37FD59" box="[410,476,691,712]" name="India" pageId="6" pageNumber="1093">INDIA</collectingCountry>
N.
<collectingRegion id="0E56BEF8A5296A03BA7DFD6FFD8DFD59" box="[514,614,690,712]" pageId="6" pageNumber="1093">23 Madras</collectingRegion>
,
<collectingMunicipality id="2C49EA60A5296A03BA0DFD6EFD2BFD59" box="[626,704,690,712]" pageId="6" pageNumber="1093">Palni H.</collectingMunicipality>
, au-dessus de
<location id="C94D26C1A5296A03BB37FD6EFC2BFD59" LSID="urn:lsid:plazi:treatment:443BC10CA5296A02B8CDFDEEFE18FAB5:C94D26C1A5296A03BB37FD6EFC2BFD59" box="[840,960,690,712]" country="India" municipality="Palni H." name="Kodai Kanal" pageId="6" pageNumber="1093" stateProvince="23 Madras">Kodai Kanal</location>
,
<quantity id="0B6ADDFFA5296A03BBB4FD6FFBFFFD59" box="[971,1044,691,712]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="6" pageNumber="1093" unit="m" value="2200.0">
<elevation id="47BF9729A5296A03BBB4FD6FFBFFFD59" box="[971,1044,691,712]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="6" pageNumber="1093" unit="m" value="2200.0">2200 m</elevation>
</quantity>
,
<collectingDate id="A868AF32A5296A03BC60FD6FFF00FD4F" pageId="6" pageNumber="1093">
<date id="B82C56DAA5296A03BC60FD6FFF00FD4F" pageId="6" pageNumber="1093" value="1972-11-12">12- XI-72</date>
</collectingDate>
,
<collectorName id="616715CCA5296A03B88AFD15FE38FD4E" box="[245,467,713,735]" pageId="6" pageNumber="1093">Besuchet Löbl Mussard</collectorName>
&quot; (
<collectionCode id="AA83E8DFA5296A03B997FD15FDD8FD4E" box="[488,563,713,735]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="6" pageNumber="1093" type="Museum">MHNG</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="7CFA7A47A5296A03BA3BFD15FD4AFD67" ID-GBIF-Occurrence="3765333310" collectingDate="1972-11-14" collectionCode="MHNG" collectorName="Besuchet Lobl Mussard" country="India" elevation="2150" location="Berijan Lake" municipality="Palni H." pageId="6" pageNumber="1093" specimenCount="2" stateProvince="26 Madras">
<specimenCount id="DA94BB93A5296A03BA3BFD15FD94FD4E" box="[580,639,713,735]" pageId="6" pageNumber="1093" type="generic">2 exs.</specimenCount>
: &quot;
<collectingCountry id="B485308AA5296A03BAEFFD15FD39FD4F" box="[656,722,713,734]" name="India" pageId="6" pageNumber="1093">INDIA</collectingCountry>
N.
<collectingRegion id="0E56BEF8A5296A03BA88FD15FCB7FD4E" box="[759,860,713,735]" pageId="6" pageNumber="1093">26 Madras</collectingRegion>
,
<collectingMunicipality id="2C49EA60A5296A03BB19FD15FC5EFD4E" box="[870,949,713,735]" pageId="6" pageNumber="1093">Palni H.</collectingMunicipality>
,
<location id="C94D26C1A5296A03BBBFFD15FBD1FD4E" LSID="urn:lsid:plazi:treatment:443BC10CA5296A02B8CDFDEEFE18FAB5:C94D26C1A5296A03BBBFFD15FBD1FD4E" box="[960,1082,713,735]" country="India" municipality="Palni H." name="Berijan Lake" pageId="6" pageNumber="1093" stateProvince="26 Madras">Berijan Lake</location>
,
<quantity id="0B6ADDFFA5296A03B8CDFD3CFF12FD64" box="[178,249,736,758]" metricMagnitude="3" metricUnit="m" metricValue="2.15" pageId="6" pageNumber="1093" unit="m" value="2150.0">
<elevation id="47BF9729A5296A03B8CDFD3CFF12FD64" box="[178,249,736,758]" metricMagnitude="3" metricUnit="m" metricValue="2.15" pageId="6" pageNumber="1093" unit="m" value="2150.0">2150 m</elevation>
</quantity>
,
<collectingDate id="A868AF32A5296A03B97DFD3CFEB1FD64" box="[258,346,736,757]" pageId="6" pageNumber="1093">
<date id="B82C56DAA5296A03B97DFD3CFEB1FD64" box="[258,346,736,757]" pageId="6" pageNumber="1093" value="1972-11-14">14-XI-72</date>
</collectingDate>
,
<collectorName id="616715CCA5296A03B91DFD3CFDD7FD67" box="[354,572,736,758]" pageId="6" pageNumber="1093">Besuchet Löbl Mussard</collectorName>
&quot; (
<collectionCode id="AA83E8DFA5296A03BA2FFD3CFD70FD67" box="[592,667,736,758]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="6" pageNumber="1093" type="Museum">MHNG</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="84882391A5296A03B8CDFCD8FDB6FCC5" pageId="6" pageNumber="1093" type="etymology">
<paragraph id="CC2D701AA5296A03B8CDFCD8FDB6FCC5" blockId="6.[178,1085,772,852]" pageId="6" pageNumber="1093">E t y m o l o g y: The specific epithet is the present participle of the Latin verb discrepare (to differ) and alludes to the numerous characters distinguishing this species from other congeners recorded from Asia.</paragraph>
</subSubSection>
<subSubSection id="84882391A5296A03B8CDFCBFFCDAF9D0" pageId="6" pageNumber="1093" type="description">
<paragraph id="CC2D701AA5296A03B8CDFCBFFC9BFBB0" blockId="6.[178,1085,866,1057]" pageId="6" pageNumber="1093">
Description: Conspicuously small species; body length
<quantity id="0B6ADDFFA5296A03BB2BFCBEFC3BFCE9" box="[852,976,866,888]" metricMagnitude="-3" metricUnit="m" metricValue="2.85" metricValueMax="3.1" metricValueMin="2.6" pageId="6" pageNumber="1093" unit="mm" value="2.85" valueMax="3.1" valueMin="2.6">2.6-3.1 mm</quantity>
; length of forebody
<quantity id="0B6ADDFFA5296A03B968FCA2FE79FC05" box="[279,402,894,916]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="1.6" metricValueMin="1.4" pageId="6" pageNumber="1093" unit="mm" value="1.5" valueMax="1.6" valueMin="1.4">1.4-1.6 mm</quantity>
. Habitus as in
<figureCitation id="54A96C9FA5296A03BA4CFCA3FD6AFC05" box="[563,641,895,917]" captionStart="Figs 14-19" captionStartId="7.[178,218,935,957]" captionTargetBox="[193,1062,212,917]" captionTargetId="figure-1@7.[178,1082,212,926]" captionTargetPageId="7" captionText="Figs 14-19: Echiaster discrepans nov.sp.: (14) habitus; (15) forebody; (16) male sternite VIII; (17- 19) aedeagus in lateral and in ventral view. Scale bars: 14: 1.0 mm; 15: 0.5 mm; 16-19: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5361375" httpUri="https://zenodo.org/record/5361375/files/figure.png" pageId="6" pageNumber="1093">Fig. 14</figureCitation>
. Coloration: head dark-brown to blackishbrown, with the anterior portion reddish to reddish-yellow; pronotum brown; elytra brown, with the postero-lateral portions usually more or less extensively infuscate, sometimes leaving only the humeral angles, the anterior margins, and the anterior sutural portion paler; abdomen brown to dark-brown, with the apex, as well as the posterior and lateral margins of the segments reddish; legs and antennae yellowish.
</paragraph>
<paragraph id="CC2D701AA5296A03B8CDFBF3FC11FB24" blockId="6.[178,1086,1070,1205]" pageId="6" pageNumber="1093">
Head (
<figureCitation id="54A96C9FA5296A03B887FBF3FEAFFBD5" box="[248,324,1070,1093]" captionStart="Figs 14-19" captionStartId="7.[178,218,935,957]" captionTargetBox="[193,1062,212,917]" captionTargetId="figure-1@7.[178,1082,212,926]" captionTargetPageId="7" captionText="Figs 14-19: Echiaster discrepans nov.sp.: (14) habitus; (15) forebody; (16) male sternite VIII; (17- 19) aedeagus in lateral and in ventral view. Scale bars: 14: 1.0 mm; 15: 0.5 mm; 16-19: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5361375" httpUri="https://zenodo.org/record/5361375/files/figure.png" pageId="6" pageNumber="1093">Fig. 15</figureCitation>
) oblong and of subquadrangular shape, 1.05-1.10 times as long as broad and with rounded, but noticeable posterior angles; punctation coarse, very dense, and umbilicate; interstices reduced to narrow ridges, but glossy, surface not completely matt; eyes slightly more than half as long as distance between posterior margin of eye to posterior constriction of head in dorsal view. Antenna approximately
<quantity id="0B6ADDFFA5296A03BB0DFB43FC2BFB25" box="[882,960,1183,1205]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="6" pageNumber="1093" unit="mm" value="0.6">0.6 mm</quantity>
long.
</paragraph>
<paragraph id="CC2D701AA5296A03B8CDFB1FFC7FFA80" blockId="6.[178,1085,1219,1297]" pageId="6" pageNumber="1093">
Pronotum (
<figureCitation id="54A96C9FA5296A03B95AFB1FFE9BFB48" box="[293,368,1219,1241]" captionStart="Figs 14-19" captionStartId="7.[178,218,935,957]" captionTargetBox="[193,1062,212,917]" captionTargetId="figure-1@7.[178,1082,212,926]" captionTargetPageId="7" captionText="Figs 14-19: Echiaster discrepans nov.sp.: (14) habitus; (15) forebody; (16) male sternite VIII; (17- 19) aedeagus in lateral and in ventral view. Scale bars: 14: 1.0 mm; 15: 0.5 mm; 16-19: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5361375" httpUri="https://zenodo.org/record/5361375/files/figure.png" pageId="6" pageNumber="1093">Fig. 15</figureCitation>
) conspicuously small (in relation to head and elytra) and not particularly slender, approximately 1.15 times as long as broad and 0.75 times as broad as head; middle elevated or flat, not impressed; punctation similar to that of head.
</paragraph>
<paragraph id="CC2D701AA5296A03B8CDFAC3FD5CFAFC" blockId="6.[178,1085,1311,1389]" pageId="6" pageNumber="1093">
Elytra (
<figureCitation id="54A96C9FA5296A03B97EFAC3FEA6FAA4" box="[257,333,1311,1333]" captionStart="Figs 14-19" captionStartId="7.[178,218,935,957]" captionTargetBox="[193,1062,212,917]" captionTargetId="figure-1@7.[178,1082,212,926]" captionTargetPageId="7" captionText="Figs 14-19: Echiaster discrepans nov.sp.: (14) habitus; (15) forebody; (16) male sternite VIII; (17- 19) aedeagus in lateral and in ventral view. Scale bars: 14: 1.0 mm; 15: 0.5 mm; 16-19: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5361375" httpUri="https://zenodo.org/record/5361375/files/figure.png" pageId="6" pageNumber="1093">Fig. 15</figureCitation>
) relatively long and large, approximately 1.2 times as long and 1.5-1.6 times as large as pronotum; humeral angles pronounced; punctation very dense and coarse, not granulose. Hind wings fully developed.
</paragraph>
<paragraph id="CC2D701AA5296A03B8CDFAA7FCA9FA74" blockId="6.[178,1086,1403,1509]" pageId="6" pageNumber="1093">Abdomen distinctly narrower than elytra; tergites III-V with moderately coarse and moderately dense, tergites VI-VIII with fine and moderately dense punctation; interstices glossy; posterior margin of tergite VII very weakly convex, with palisade fringe; tergite VIII distinctly oblong and with weakly convex posterior margin.</paragraph>
<paragraph id="CC2D701AA5296A03B8B8FA2FFCDAF9D0" blockId="6.[178,1085,1523,1601]" pageId="6" pageNumber="1093">
: sternite VIII (
<figureCitation id="54A96C9FA5296A03B91CFA2FFE46F998" box="[355,429,1523,1545]" captionStart="Figs 14-19" captionStartId="7.[178,218,935,957]" captionTargetBox="[193,1062,212,917]" captionTargetId="figure-1@7.[178,1082,212,926]" captionTargetPageId="7" captionText="Figs 14-19: Echiaster discrepans nov.sp.: (14) habitus; (15) forebody; (16) male sternite VIII; (17- 19) aedeagus in lateral and in ventral view. Scale bars: 14: 1.0 mm; 15: 0.5 mm; 16-19: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5361375" httpUri="https://zenodo.org/record/5361375/files/figure.png" pageId="6" pageNumber="1093">Fig. 16</figureCitation>
) oblong, with moderately dense pubescence composed of stout, long, and dark setae, posterior excision shallow; aedeagus (
<figureCitation id="54A96C9FA5296A03BB6BF9D3FC6FF9B4" box="[788,900,1551,1573]" captionStart="Figs 14-19" captionStartId="7.[178,218,935,957]" captionTargetBox="[193,1062,212,917]" captionTargetId="figure-1@7.[178,1082,212,926]" captionTargetPageId="7" captionText="Figs 14-19: Echiaster discrepans nov.sp.: (14) habitus; (15) forebody; (16) male sternite VIII; (17- 19) aedeagus in lateral and in ventral view. Scale bars: 14: 1.0 mm; 15: 0.5 mm; 16-19: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5361375" httpUri="https://zenodo.org/record/5361375/files/figure.png" pageId="6" pageNumber="1093">Figs 17-19</figureCitation>
) minute, approximately
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long, weakly sclerotized, and of simple shape.
</paragraph>
</subSubSection>
<caption id="98ED2092A5286A02B8CDFC7BFBF6FC45" ID-DOI="http://doi.org/10.5281/zenodo.5361375" ID-Zenodo-Dep="5361375" httpUri="https://zenodo.org/record/5361375/files/figure.png" pageId="7" pageNumber="1094" startId="7.[178,218,935,957]" targetBox="[193,1062,212,917]" targetPageId="7">
<paragraph id="CC2D701AA5286A02B8CDFC7BFBF6FC45" blockId="7.[178,1085,935,980]" pageId="7" pageNumber="1094">
<emphasis id="FEE6AC08A5286A02B8CDFC7BFEFDFC2D" bold="true" box="[178,278,935,957]" pageId="7" pageNumber="1094">Figs 14-19</emphasis>
:
<taxonomicName id="0B920B99A5286A02B95CFC7BFE09FC2D" authority="Assing &amp; I, 2014" authorityName="Assing &amp; I" authorityYear="2014" box="[291,482,935,956]" class="Insecta" family="Staphylinidae" genus="Echiaster" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="1094" phylum="Arthropoda" rank="species" species="discrepans" status="sp. nov.">
<emphasis id="FEE6AC08A5286A02B95CFC7BFE09FC2D" box="[291,482,935,956]" italics="true" pageId="7" pageNumber="1094">Echiaster discrepans</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E5D51173A5286A02B996FC74FDC5FC2C" box="[489,558,936,957]" pageId="7" pageNumber="1094" rank="species">nov.sp.</taxonomicNameLabel>
: (
<emphasis id="FEE6AC08A5286A02BA41FC7BFDBEFC2D" bold="true" box="[574,597,935,956]" pageId="7" pageNumber="1094">14</emphasis>
) habitus; (
<emphasis id="FEE6AC08A5286A02BAC6FC7BFD3CFC2C" bold="true" box="[697,727,935,957]" pageId="7" pageNumber="1094">15)</emphasis>
forebody; (
<emphasis id="FEE6AC08A5286A02BB3CFC7BFC89FC2C" bold="true" box="[835,866,935,957]" pageId="7" pageNumber="1094">16)</emphasis>
male sternite VIII; (
<emphasis id="FEE6AC08A5286A02BC60FC7BFF22FC42" bold="true" pageId="7" pageNumber="1094">17- 19</emphasis>
) aedeagus in lateral and in ventral view. Scale bars: 14: 1.0 mm; 15: 0.5 mm; 16-19: 0.1 mm.
</paragraph>
</caption>
<subSubSection id="84882391A5286A02B8CDFC3EFCE5FB54" pageId="7" pageNumber="1094" type="discussion">
<paragraph id="CC2D701AA5286A02B8CDFC3EFCE5FB54" blockId="7.[178,1086,994,1221]" pageId="7" pageNumber="1094">
Comparative notes
<taxonomicName id="0B920B99A5286A02BA70FC3EFD0CFC66" authorityName="Assing &amp; I" authorityYear="2014" box="[527,743,994,1015]" class="Insecta" family="Staphylinidae" genus="Echiaster" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="1094" phylum="Arthropoda" rank="species" species="discrepans">
<emphasis id="FEE6AC08A5286A02BA70FC3EFD0CFC66" box="[527,743,994,1015]" italics="true" pageId="7" pageNumber="1094">Echiaster discrepans</emphasis>
</taxonomicName>
differs from other Asian representatives of the genus by numerous significant external and sexual characters, particularly much smaller body size, completely different habitus (much smaller and shorter pronotum, different head shape, much larger elytra), different punctation of the forebody and of the abdomen, posteriorly weakly convex posterior margins of the abdominal tergites VII and VIII, different chaetotaxy of the male sternite VII and VIII, and by the different general morphology of the aedeagus. The number and quality of these differences suggest that it probably belongs to a separate lineage.
</paragraph>
</subSubSection>
<subSubSection id="84882391A5286A02B8CDFB08FE18FAB5" pageId="7" pageNumber="1094" type="distribution">
<paragraph id="CC2D701AA5286A02B8CDFB08FE18FAB5" blockId="7.[178,1085,1236,1317]" pageId="7" pageNumber="1094">
Distribution and natural history The species was collected in several localities near Kodaikanal [
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,
<geoCoordinate id="A9A616DDA5286A02BAFFFB2DFD38FA96" box="[640,723,1265,1287]" degrees="77" direction="east" minutes="29" orientation="longitude" pageId="7" pageNumber="1094" precision="925" value="77.48333">77°29'E</geoCoordinate>
] in
<collectingRegion id="0E56BEF8A5286A02BA82FB2EFC90FA99" box="[765,891,1266,1288]" country="India" name="Tamil Nadu" pageId="7" pageNumber="1094">Tamil Nadu</collectingRegion>
, South
<collectingCountry id="B485308AA5286A02BBB5FB2EFC15FA99" box="[970,1022,1266,1288]" name="India" pageId="7" pageNumber="1094">India</collectingCountry>
(
<figureCitation id="54A96C9FA5286A02BC72FB2EFF56FAB4" captionStart="Map 1" captionStartId="1.[178,223,1240,1261]" captionTargetBox="[178,1085,558,1214]" captionTargetId="figure-142@1.[178,1085,558,1214]" captionTargetPageId="1" captionText="Map 1: Distribution of Echiaster species in Asia. Echiaster discrepans (white circles); E. bivirgatus (white diamonds); E. maior (white triangles); E. paulexcisus (black diamond); E. unicolor (black circles)." figureDoi="http://doi.org/10.5281/zenodo.5361369" httpUri="https://zenodo.org/record/5361369/files/figure.png" pageId="7" pageNumber="1094">Map 1</figureCitation>
), at altitudes of
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.
</paragraph>
</subSubSection>
</treatment>
</document>
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