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<document id="85C5709F7A5E169321045CC3826C51A2" ID-DOI="10.1111/jfb.14558" ID-ISSN="1095-8649" ID-PMC="PMC7821206" ID-PubMed="32985685" ID-Zenodo-Dep="10928926" IM.bibliography_approvedBy="julia" IM.illustrations_approvedBy="julia" IM.materialsCitations_approvedBy="julia" IM.tables_approvedBy="julia" IM.taxonomicNames_approvedBy="julia" IM.treatmentCitations_approvedBy="julia" IM.treatments_approvedBy="julia" checkinTime="1712261492542" checkinUser="felipe" docAuthor="Wagner, Maximilian, Kovac, Marcelo &amp; Koblmüller, Stephan" docDate="2021" docId="195D87A2FFBD5B3A515AFA2E0DA95AC4" docLanguage="en" docName="JFishBiol.98.1.64-88.pdf" docOrigin="Journal of Fish Biology 98 (1)" docSource="http://dx.doi.org/10.1111/jfb.14558" docStyle="DocumentStyle:E312FF628AF650F200AD49ACE691BDEB.1:JFishBiol.2018-.journal_article.open" docStyleId="E312FF628AF650F200AD49ACE691BDEB" docStyleName="JFishBiol.2018-.journal_article.open" docStyleVersion="1" docTitle="Gouania pigra" docType="treatment" docVersion="3" lastPageNumber="80" masterDocId="E564FFDAFFB05B2A526AFF810C6A5F03" masterDocTitle="Unravelling the taxonomy of an interstitial fish radiation: Three new species of Gouania (Teleostei: Gobiesocidae) from the Mediterranean Sea and redescriptions of G. willdenowi and G. pigra" masterLastPageNumber="88" masterPageNumber="64" pageNumber="77" updateTime="1712674921547" updateUser="julia" zenodo-license-document="CC-BY-NC-ND-4.0">
<mods:mods id="A1C6829D8573B5CF551E76A43B2643D9" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="54A39A90CB6D49BC2B642992AF898EAD">
<mods:title id="6F0964EDD1E1E509057336B842C52724">Unravelling the taxonomy of an interstitial fish radiation: Three new species of Gouania (Teleostei: Gobiesocidae) from the Mediterranean Sea and redescriptions of G. willdenowi and G. pigra</mods:title>
</mods:titleInfo>
<mods:name id="80FE897646F437696BCE241E79D8A74B" type="personal">
<mods:role id="762F4AB9AC2CBD1A0ACA311077622560">
<mods:roleTerm id="A5FB7632CA2F4B1AA1A32A2109DA190F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B55A12258A226870C95C0908C9742349">Wagner, Maximilian</mods:namePart>
<mods:affiliation id="91E20675FC8FCF29B686E0BE6231DB80">Institute of Biology, University of Graz, Graz, Austria &amp; Department of Biology, University of Antwerp, Antwerp, Belgium</mods:affiliation>
</mods:name>
<mods:name id="9EA99617A8FC61E6BE0172986DF69B37" type="personal">
<mods:role id="E4BEDEF1BCA1DD29725AED4B31AA9F3B">
<mods:roleTerm id="BA61BE2B51DD8DB2A45AC30627A6BBF7">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="DD576755D5665FA46EDA3823810250B4">Kovac, Marcelo</mods:namePart>
<mods:affiliation id="6DECB4EA8146AC39BB0D4812AE861BB8">Natural History Museum Rijeka, Rijeka, Croatia</mods:affiliation>
</mods:name>
<mods:name id="917EB9020126464257B9C27891530F98" type="personal">
<mods:role id="C4B25D4A968167C3D3190EB22A835028">
<mods:roleTerm id="0EA6F18D7EB14D52C466D82C80460BDF">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="57BE606BEFFD7E1C173C0246F65FFCAC">Koblmüller, Stephan</mods:namePart>
<mods:affiliation id="1FDF6200BE17F9B63858A6AB86E6F973">Institute of Biology, University of Graz, Graz, Austria</mods:affiliation>
</mods:name>
<mods:typeOfResource id="B13E6C9E99A4C70FB86A8D5A1FC127C4">text</mods:typeOfResource>
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<mods:titleInfo id="C8D17EAD721818C8EC5E24E35985E88C">
<mods:title id="2FAD5CFA516FDF3197F75FAFE475A881">Journal of Fish Biology</mods:title>
</mods:titleInfo>
<mods:part id="75AE4E656BF34B1EEB3967095535BD67">
<mods:date id="FC7B7AEFFA8B19BE09A254B1461047AF">2021</mods:date>
<mods:detail id="F39C376DCFF1DABED93BB66EB8BE1CC7" type="pubDate">
<mods:number id="5AF96F87DD73F31EED38184A5D5A2966">2020-11-03</mods:number>
</mods:detail>
<mods:detail id="D7BD3751A516ED38C98A7AFC25561120" type="volume">
<mods:number id="2BE616CA92104DA8BB42DC067FE0DAF7">98</mods:number>
</mods:detail>
<mods:detail id="1232D9E5A3BAB6F2471171360497F0FC" type="issue">
<mods:number id="192EAFD8725EC0055C54FD726087B730">1</mods:number>
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<mods:extent id="D98F2CD9589E84011071B10F10CB8434" unit="page">
<mods:start id="8D2A5187774A9BE1A36C5C19D54CD485">64</mods:start>
<mods:end id="9D23139F5CDFE4E5F4095544CD722746">88</mods:end>
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<mods:location id="373103C9550B30F056DC5C7BDFC911DE">
<mods:url id="5B12948EC7EDC023F90EEC3664F4BDDF">http://dx.doi.org/10.1111/jfb.14558</mods:url>
</mods:location>
<mods:classification id="1A8036CE9BF02D6794D1E1100D5E330E">journal article</mods:classification>
<mods:identifier id="F74DF847407149CC42B71B50A08F8576" type="DOI">10.1111/jfb.14558</mods:identifier>
<mods:identifier id="647590F3E29D3CCCA06CC24B84193DE0" type="ISSN">1095-8649</mods:identifier>
<mods:identifier id="704FD468512321E65A841DE9EF741919" type="PMC">PMC7821206</mods:identifier>
<mods:identifier id="871F4C6E3EBC462F646A8E9F514D88DC" type="PubMed">32985685</mods:identifier>
<mods:identifier id="0EBD5A78E6971AB2CE8F08B0B2183758" type="Zenodo-Dep">10928926</mods:identifier>
</mods:mods>
<treatment id="195D87A2FFBD5B3A515AFA2E0DA95AC4" LSID="urn:lsid:plazi:treatment:195D87A2FFBD5B3A515AFA2E0DA95AC4" httpUri="http://treatment.plazi.org/id/195D87A2FFBD5B3A515AFA2E0DA95AC4" lastPageId="16" lastPageNumber="80" pageId="13" pageNumber="77">
<subSubSection id="D9EE653FFFBD5B27515AFA2E08985ACB" box="[816,1266,1455,1481]" pageId="13" pageNumber="77" type="nomenclature">
<paragraph id="914B36B4FFBD5B27515AFA2E08985ACB" blockId="13.[816,1266,1455,1481]" box="[816,1266,1455,1481]" pageId="13" pageNumber="77">
<heading id="CA0381D8FFBD5B27515AFA2E08985ACB" box="[816,1266,1455,1481]" fontSize="10" level="3" pageId="13" pageNumber="77" reason="9">
<emphasis id="A380EAA6FFBD5B27515AFA2E0F005ACB" bold="true" box="[816,874,1455,1480]" pageId="13" pageNumber="77">3.1.5</emphasis>
|
<taxonomicName id="56F44D37FFBD5B2751DBFA2E08985ACB" ID-CoL="3H4M9" authority="(Nardo 1827)" baseAuthorityName="Nardo" baseAuthorityYear="1827" box="[945,1266,1455,1481]" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="13" pageNumber="77" phylum="Chordata" rank="species" species="pigra">
<emphasis id="A380EAA6FFBD5B2751DBFA2E08985ACB" bold="true" box="[945,1266,1455,1481]" pageId="13" pageNumber="77">
<emphasis id="A380EAA6FFBD5B2751DBFA2E08255ACA" bold="true" box="[945,1103,1455,1481]" italics="true" pageId="13" pageNumber="77">Gouania pigra</emphasis>
(Nardo 1827)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="D9EE653FFFBD5B27515AFA790FE75976" pageId="13" pageNumber="77" type="description">
<paragraph id="914B36B4FFBD5B27515AFA79085A590F" blockId="13.[816,1465,1528,1965]" box="[816,1072,1528,1548]" pageId="13" pageNumber="77">
<emphasis id="A380EAA6FFBD5B27515AFA79085A590F" bold="true" box="[816,1072,1528,1548]" pageId="13" pageNumber="77">English name: Piglet sucker</emphasis>
</paragraph>
<paragraph id="914B36B4FFBD5B275132F99A0FE75976" blockId="13.[816,1465,1528,1965]" pageId="13" pageNumber="77">
<materialsCitation id="219C3CE9FFBD5B275132F99A0FE75976" collectingDate="2015-05-16" collectionCode="PMR" collectorName="M. Wagner" country="Croatia" location="Croatia" pageId="13" pageNumber="77" specimenCode="VP3529" specimenCount="1" typeStatus="neotype">
<emphasis id="A380EAA6FFBD5B275132F99A0FDA592C" bold="true" box="[856,944,1563,1583]" pageId="13" pageNumber="77">
<typeStatus id="4E4F8816FFBD5B275132F99A0FC6592C" box="[856,940,1563,1583]" pageId="13" pageNumber="77" type="neotype">Neotype</typeStatus>
.
</emphasis>
<collectionCode id="F7E5AE71FFBD5B2751DFF99A0F89592C" box="[949,995,1563,1583]" country="0" httpUri="http://grbio.org/cool/xq4z-q5gq" name="Prirodoslovni muzej Rijeka" pageId="13" pageNumber="77">PMR</collectionCode>
<specimenCode id="C1529ECFFFBD5B275183F99A085D592C" box="[1001,1079,1563,1583]" collectionCode="VP" pageId="13" pageNumber="77">VP3529</specimenCode>
, female, 43.12 +
<quantity id="560C9B51FFBD5B2756BDF99A0941592C" box="[1239,1323,1563,1583]" metricMagnitude="-3" metricUnit="m" metricValue="5.03" pageId="13" pageNumber="77" unit="mm" value="5.03">5.03 mm</quantity>
, Glavotok, Krk,
<collectingCountry id="E9E37624FFBD5B27515AF9BF0F1F5951" box="[816,885,1598,1618]" name="Croatia" pageId="13" pageNumber="77">Croatia</collectingCountry>
, 45 05
<emphasis id="A380EAA6FFBD5B2751D2F9BA0FD6594B" bold="true" box="[952,956,1595,1608]" italics="true" pageId="13" pageNumber="77">
<superScript id="66819BFCFFBD5B2751D2F9BA0FD6594B" attach="none" box="[952,956,1595,1608]" fontSize="6" pageId="13" pageNumber="77">0</superScript>
</emphasis>
44.9
<emphasis id="A380EAA6FFBD5B27518DF9BA0F85594B" bold="true" box="[999,1007,1595,1608]" italics="true" pageId="13" pageNumber="77">
<superScript id="66819BFCFFBD5B27518DF9BA0F85594B" attach="right" box="[999,1007,1595,1608]" fontSize="6" pageId="13" pageNumber="77">00</superScript>
</emphasis>
N, 14 26
<emphasis id="A380EAA6FFBD5B27562FF9BA0823594B" bold="true" box="[1093,1097,1595,1608]" italics="true" pageId="13" pageNumber="77">
<superScript id="66819BFCFFBD5B27562FF9BA0823594B" attach="none" box="[1093,1097,1595,1608]" fontSize="6" pageId="13" pageNumber="77">0</superScript>
</emphasis>
32.4
<emphasis id="A380EAA6FFBD5B27561EF9BA0816594B" bold="true" box="[1140,1148,1595,1608]" italics="true" pageId="13" pageNumber="77">
<superScript id="66819BFCFFBD5B27561EF9BA0816594B" attach="right" box="[1140,1148,1595,1608]" fontSize="6" pageId="13" pageNumber="77">00</superScript>
</emphasis>
E, coll.
<collectorName id="3C015362FFBD5B2756AAF9BF09415951" box="[1216,1323,1598,1618]" pageId="13" pageNumber="77">M. Wagner</collectorName>
,
<date id="E54A1074FFBD5B275759F9BF09D25951" box="[1331,1464,1598,1618]" pageId="13" pageNumber="77" value="2015-05-16">
<collectingDate id="F50EE99CFFBD5B275759F9BF09D25951" box="[1331,1464,1598,1618]" pageId="13" pageNumber="77" value="2015-05-16">May 16, 2015</collectingDate>
</date>
(Figure 8).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="D9EE653FFFBD5B245132F9020CB45D1C" lastPageId="14" lastPageNumber="78" pageId="13" pageNumber="77" type="materials_examined">
<paragraph id="914B36B4FFBD5B245132F9020D6A5E2E" blockId="13.[816,1465,1528,1965]" lastBlockId="14.[124,773,139,1167]" lastPageId="14" lastPageNumber="78" pageId="13" pageNumber="77">
<emphasis id="A380EAA6FFBD5B275132F90208EA5994" bold="true" box="[856,1152,1667,1687]" pageId="13" pageNumber="77">Additional material examined.</emphasis>
<materialsCitation id="219C3CE9FFBD5B2756E1F9020FAA5821" collectingDate="2019-09-16" collectionCode="PMR" collectorName="M. Wagner" country="Croatia" location="Stara Baska" municipality="Krk" pageId="13" pageNumber="77" specimenCode="VP3531" specimenCount="1">
ZSM-PIS-047648, male, 37.71 +
<quantity id="560C9B51FFBD5B275128F9270FFC59B9" box="[834,918,1702,1722]" metricMagnitude="-3" metricUnit="m" metricValue="4.36" pageId="13" pageNumber="77" unit="mm" value="4.36">4.36 mm</quantity>
, ZSM-PIS-047648, male, 37.2 +
<quantity id="560C9B51FFBD5B2756BEF927094C59B9" box="[1236,1318,1702,1722]" metricMagnitude="-3" metricUnit="m" metricValue="4.31" pageId="13" pageNumber="77" unit="mm" value="4.31">4.31 mm</quantity>
and ZSM-PIS-047648, female, 32.58 +
<quantity id="560C9B51FFBD5B275677F948081B59DE" box="[1053,1137,1737,1757]" metricMagnitude="-3" metricUnit="m" metricValue="3.94" pageId="13" pageNumber="77" unit="mm" value="3.94">3.94 mm</quantity>
, all from
<location id="942B606FFFBD5B2756A0F948095359DE" LSID="urn:lsid:plazi:treatment:195D87A2FFBD5B3A515AFA2E0DA95AC4:942B606FFFBD5B2756A0F948095359DE" box="[1226,1337,1737,1757]" country="Croatia" municipality="Krk" name="Stara Baska" pageId="13" pageNumber="77">Stara Baška</location>
,
<collectingMunicipality id="712FACCEFFBD5B275729F948090F59DE" box="[1347,1381,1737,1757]" pageId="13" pageNumber="77">Krk</collectingMunicipality>
,
<collectingCountry id="E9E37624FFBD5B27571AF94809DC59DE" box="[1392,1462,1737,1757]" name="Croatia" pageId="13" pageNumber="77">Croatia</collectingCountry>
, 44 56
<emphasis id="A380EAA6FFBD5B275100F9680F0459F5" bold="true" box="[874,878,1769,1782]" italics="true" pageId="13" pageNumber="77">
<superScript id="66819BFCFFBD5B275100F9680F0459F5" attach="none" box="[874,878,1769,1782]" fontSize="6" pageId="13" pageNumber="77">0</superScript>
</emphasis>
45.3
<emphasis id="A380EAA6FFBD5B2751F2F9680FCA59F5" bold="true" box="[920,928,1769,1782]" italics="true" pageId="13" pageNumber="77">
<superScript id="66819BFCFFBD5B2751F2F9680FCA59F5" attach="right" box="[920,928,1769,1782]" fontSize="6" pageId="13" pageNumber="77">00</superScript>
</emphasis>
N, 14 42
<emphasis id="A380EAA6FFBD5B275192F9680F9659F5" bold="true" box="[1016,1020,1769,1782]" italics="true" pageId="13" pageNumber="77">
<superScript id="66819BFCFFBD5B275192F9680F9659F5" attach="none" box="[1016,1020,1769,1782]" fontSize="6" pageId="13" pageNumber="77">0</superScript>
</emphasis>
22.2
<emphasis id="A380EAA6FFBD5B27564DF968084559F5" bold="true" box="[1063,1071,1769,1782]" italics="true" pageId="13" pageNumber="77">
<superScript id="66819BFCFFBD5B27564DF968084559F5" attach="right" box="[1063,1071,1769,1782]" fontSize="6" pageId="13" pageNumber="77">00</superScript>
</emphasis>
E, coll.
<collectorName id="3C015362FFBD5B27561DF96A088E59FC" box="[1143,1252,1771,1791]" pageId="13" pageNumber="77">M. Wagner</collectorName>
,
<date id="E54A1074FFBD5B275685F96A09DF59FC" box="[1263,1461,1771,1791]" pageId="13" pageNumber="77" value="2019-09-16">
<collectingDate id="F50EE99CFFBD5B275685F96A09DF59FC" box="[1263,1461,1771,1791]" pageId="13" pageNumber="77" value="2019-09-16">September 16, 2019</collectingDate>
</date>
;
<collectionCode id="F7E5AE71FFBD5B27515AF88F0F345821" box="[816,862,1806,1826]" country="0" httpUri="http://grbio.org/cool/xq4z-q5gq" name="Prirodoslovni muzej Rijeka" pageId="13" pageNumber="77">PMR</collectionCode>
<specimenCode id="C1529ECFFFBD5B275118F88F0FAA5821" box="[882,960,1806,1826]" collectionCode="VP" pageId="13" pageNumber="77">VP3531</specimenCode>
</materialsCitation>
,
<materialsCitation id="219C3CE9FFBD5B2751BDF88F0F175864" collectingDate="2015-05-16" collectionCode="PMR" collectorName="M. Wagner" country="Croatia" location="Glavotok" municipality="Krk." pageId="13" pageNumber="77" specimenCode="VP4619" specimenCount="1">
male, 39.48 +
<quantity id="560C9B51FFBD5B275600F88F08D85821" box="[1130,1202,1806,1826]" metricMagnitude="-3" metricUnit="m" metricValue="4.4" pageId="13" pageNumber="77" unit="mm" value="4.4">4.4 mm</quantity>
,
<location id="942B606FFFBD5B2756A3F88F09755821" LSID="urn:lsid:plazi:treatment:195D87A2FFBD5B3A515AFA2E0DA95AC4:942B606FFFBD5B2756A3F88F09755821" box="[1225,1311,1806,1826]" country="Croatia" municipality="Krk." name="Glavotok" pageId="13" pageNumber="77">Glavotok</location>
,
<collectingMunicipality id="712FACCEFFBD5B27575CF88F09375821" box="[1334,1373,1806,1826]" pageId="13" pageNumber="77">Krk.</collectingMunicipality>
<collectingCountry id="E9E37624FFBD5B27571AF88F09DC5821" box="[1392,1462,1806,1826]" name="Croatia" pageId="13" pageNumber="77">Croatia</collectingCountry>
, 45 05
<emphasis id="A380EAA6FFBD5B275100F8AF0F045838" bold="true" box="[874,878,1838,1851]" italics="true" pageId="13" pageNumber="77">
<superScript id="66819BFCFFBD5B275100F8AF0F045838" attach="none" box="[874,878,1838,1851]" fontSize="6" pageId="13" pageNumber="77">0</superScript>
</emphasis>
44.9
<emphasis id="A380EAA6FFBD5B2751F2F8AF0FCA5838" bold="true" box="[920,928,1838,1851]" italics="true" pageId="13" pageNumber="77">
<superScript id="66819BFCFFBD5B2751F2F8AF0FCA5838" attach="right" box="[920,928,1838,1851]" fontSize="6" pageId="13" pageNumber="77">00</superScript>
</emphasis>
N, 14 26
<emphasis id="A380EAA6FFBD5B275193F8AF0F975838" bold="true" box="[1017,1021,1838,1851]" italics="true" pageId="13" pageNumber="77">
<superScript id="66819BFCFFBD5B275193F8AF0F975838" attach="none" box="[1017,1021,1838,1851]" fontSize="6" pageId="13" pageNumber="77">0</superScript>
</emphasis>
32.4
<emphasis id="A380EAA6FFBD5B275642F8AF085A5838" bold="true" box="[1064,1072,1838,1851]" italics="true" pageId="13" pageNumber="77">
<superScript id="66819BFCFFBD5B275642F8AF085A5838" attach="right" box="[1064,1072,1838,1851]" fontSize="6" pageId="13" pageNumber="77">00</superScript>
</emphasis>
E, coll.
<collectorName id="3C015362FFBD5B275613F8B0088D5846" box="[1145,1255,1841,1861]" pageId="13" pageNumber="77">M. Wagner</collectorName>
,
<date id="E54A1074FFBD5B275698F8B009145846" box="[1266,1406,1841,1861]" pageId="13" pageNumber="77" value="2015-05-16">
<collectingDate id="F50EE99CFFBD5B275698F8B009145846" box="[1266,1406,1841,1861]" pageId="13" pageNumber="77" value="2015-05-16">May 16, 2015</collectingDate>
</date>
;
<collectionCode id="F7E5AE71FFBD5B2757E0F8B009D25846" box="[1418,1464,1841,1861]" country="0" httpUri="http://grbio.org/cool/xq4z-q5gq" name="Prirodoslovni muzej Rijeka" pageId="13" pageNumber="77">PMR</collectionCode>
<specimenCode id="C1529ECFFFBD5B27515AF8D20F175864" box="[816,893,1875,1895]" collectionCode="VP" pageId="13" pageNumber="77">VP4619</specimenCode>
</materialsCitation>
,
<materialsCitation id="219C3CE9FFBD5B2751ECF8D2096B5889" collectingDate="2016-07-17" collectorName="M. Wagner" country="Croatia" location="Stoja" municipality="Pula" pageId="13" pageNumber="77" specimenCount="1">
female, 35.86 +
<quantity id="560C9B51FFBD5B275676F8D2081A5864" box="[1052,1136,1875,1895]" metricMagnitude="-3" metricUnit="m" metricValue="4.14" pageId="13" pageNumber="77" unit="mm" value="4.14">4.14 mm</quantity>
,
<location id="942B606FFFBD5B275612F8D208C35864" LSID="urn:lsid:plazi:treatment:195D87A2FFBD5B3A515AFA2E0DA95AC4:942B606FFFBD5B275612F8D208C35864" box="[1144,1193,1875,1895]" country="Croatia" municipality="Pula" name="Stoja" pageId="13" pageNumber="77">Stoja</location>
,
<collectingMunicipality id="712FACCEFFBD5B2756DBF8D208B15864" box="[1201,1243,1875,1895]" pageId="13" pageNumber="77">Pula</collectingMunicipality>
,
<collectingCountry id="E9E37624FFBD5B275689F8D209435864" box="[1251,1321,1875,1895]" name="Croatia" pageId="13" pageNumber="77">Croatia</collectingCountry>
, 44 51
<emphasis id="A380EAA6FFBD5B275700F8D00904585D" bold="true" box="[1386,1390,1873,1886]" italics="true" pageId="13" pageNumber="77">
<superScript id="66819BFCFFBD5B275700F8D00904585D" attach="none" box="[1386,1390,1873,1886]" fontSize="6" pageId="13" pageNumber="77">0</superScript>
</emphasis>
38.4
<emphasis id="A380EAA6FFBD5B2757F3F8D009CB585D" bold="true" box="[1433,1441,1873,1886]" italics="true" pageId="13" pageNumber="77">
<superScript id="66819BFCFFBD5B2757F3F8D009CB585D" attach="right" box="[1433,1441,1873,1886]" fontSize="6" pageId="13" pageNumber="77">00</superScript>
</emphasis>
N, 13 49
<emphasis id="A380EAA6FFBD5B275100F8F20F045883" bold="true" box="[874,878,1907,1920]" italics="true" pageId="13" pageNumber="77">
<superScript id="66819BFCFFBD5B275100F8F20F045883" attach="none" box="[874,878,1907,1920]" fontSize="6" pageId="13" pageNumber="77">0</superScript>
</emphasis>
05.0
<emphasis id="A380EAA6FFBD5B2751F2F8F20FCA5883" bold="true" box="[920,928,1907,1920]" italics="true" pageId="13" pageNumber="77">
<superScript id="66819BFCFFBD5B2751F2F8F20FCA5883" attach="right" box="[920,928,1907,1920]" fontSize="6" pageId="13" pageNumber="77">00</superScript>
</emphasis>
E, coll.
<collectorName id="3C015362FFBD5B27519BF8F708095889" box="[1009,1123,1910,1930]" pageId="13" pageNumber="77">M. Wagner</collectorName>
,
<date id="E54A1074FFBD5B275618F8F7096B5889" box="[1138,1281,1910,1930]" pageId="13" pageNumber="77" value="2016-07-17">
<collectingDate id="F50EE99CFFBD5B275618F8F7096B5889" box="[1138,1281,1910,1930]" pageId="13" pageNumber="77" value="2016-07-17">July 17, 2016</collectingDate>
</date>
</materialsCitation>
; ZSM-PIS-047649, female, 35.14 +
<quantity id="560C9B51FFBD5B2751BDF818084358AE" box="[983,1065,1945,1965]" metricMagnitude="-3" metricUnit="m" metricValue="4.03" pageId="13" pageNumber="77" unit="mm" value="4.03">4.03 mm</quantity>
and ZSM-PIS-047649, male, 37.07 +
<materialsCitation id="219C3CE9FFBE5B2452E4FF0F0E5D5FC6" collectingDate="2019-07-12" collectorName="M. Wagner" country="Croatia" location="Croatia" pageId="14" pageNumber="78" specimenCount="1">
<quantity id="560C9B51FFBE5B2452E4FF0F0C885FA1" box="[142,226,142,162]" metricMagnitude="-3" metricUnit="m" metricValue="4.51" pageId="14" pageNumber="78" unit="mm" value="4.51">4.51 mm</quantity>
, both from Pecine, Rijeka,
<collectingCountry id="E9E37624FFBE5B24504BFF0F0E0D5FA1" box="[545,615,142,162]" name="Croatia" pageId="14" pageNumber="78">Croatia</collectingCountry>
, 45 18
<emphasis id="A380EAA6FFBE5B2450DCFF0A0ED05F9B" bold="true" box="[694,698,139,152]" italics="true" pageId="14" pageNumber="78">
<superScript id="66819BFCFFBE5B2450DCFF0A0ED05F9B" attach="none" box="[694,698,139,152]" fontSize="6" pageId="14" pageNumber="78">0</superScript>
</emphasis>
52.4
<emphasis id="A380EAA6FFBE5B24508FFF0A0E875F9B" bold="true" box="[741,749,139,152]" italics="true" pageId="14" pageNumber="78">
<superScript id="66819BFCFFBE5B24508FFF0A0E875F9B" attach="right" box="[741,749,139,152]" fontSize="6" pageId="14" pageNumber="78">00</superScript>
</emphasis>
N, 14 28
<emphasis id="A380EAA6FFBE5B2452DFFF2F0CD35FB8" bold="true" box="[181,185,174,187]" italics="true" pageId="14" pageNumber="78">
<superScript id="66819BFCFFBE5B2452DFFF2F0CD35FB8" attach="none" box="[181,185,174,187]" fontSize="6" pageId="14" pageNumber="78">0</superScript>
</emphasis>
11.7
<emphasis id="A380EAA6FFBE5B24528EFF2F0C865FB8" bold="true" box="[228,236,174,187]" italics="true" pageId="14" pageNumber="78">
<superScript id="66819BFCFFBE5B24528EFF2F0C865FB8" attach="right" box="[228,236,174,187]" fontSize="6" pageId="14" pageNumber="78">00</superScript>
</emphasis>
E, coll.
<collectorName id="3C015362FFBE5B24535CFF300DCE5FC6" box="[310,420,177,197]" pageId="14" pageNumber="78">M. Wagner</collectorName>
,
<date id="E54A1074FFBE5B2453C5FF300E5D5FC6" box="[431,567,177,197]" pageId="14" pageNumber="78" value="2019-07-12">
<collectingDate id="F50EE99CFFBE5B2453C5FF300E5D5FC6" box="[431,567,177,197]" pageId="14" pageNumber="78" value="2019-07-12">July 12, 2019</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="219C3CE9FFBE5B245029FF300C915E2E" collectingDate="2015-07-18" collectionCode="PMR" collectorName="M. Wagner" country="Croatia" location="Istria" pageId="14" pageNumber="78" specimenCode="VP4581, VP4583" specimenCount="1" stateProvince="Istarska">
<collectionCode id="F7E5AE71FFBE5B245029FF300E1B5FC6" box="[579,625,177,197]" country="0" httpUri="http://grbio.org/cool/xq4z-q5gq" name="Prirodoslovni muzej Rijeka" pageId="14" pageNumber="78">PMR</collectionCode>
<specimenCode id="C1529ECFFFBE5B245010FF300EAD5FC6" box="[634,711,177,197]" collectionCode="VP" pageId="14" pageNumber="78">VP4581</specimenCode>
, male, 38.67 +
<quantity id="560C9B51FFBE5B2452A0FF520D765FE4" box="[202,284,211,231]" metricMagnitude="-3" metricUnit="m" metricValue="4.91" pageId="14" pageNumber="78" unit="mm" value="4.91">4.91 mm</quantity>
and
<collectionCode id="F7E5AE71FFBE5B245331FF520DE35FE4" box="[347,393,211,231]" country="0" httpUri="http://grbio.org/cool/xq4z-q5gq" name="Prirodoslovni muzej Rijeka" pageId="14" pageNumber="78">PMR</collectionCode>
<specimenCode id="C1529ECFFFBE5B2453FDFF520D8F5FE4" box="[407,485,211,231]" collectionCode="VP" pageId="14" pageNumber="78">VP4583</specimenCode>
, male, 36.87 +
<quantity id="560C9B51FFBE5B2450E9FF520EBD5FE4" box="[643,727,211,231]" metricMagnitude="-3" metricUnit="m" metricValue="4.08" pageId="14" pageNumber="78" unit="mm" value="4.08">4.08 mm</quantity>
, Sv. Marina,
<collectingRegion id="5330F856FFBE5B2452A3FF770C935E09" box="[201,249,246,266]" country="Croatia" name="Istarska" pageId="14" pageNumber="78">Istria</collectingRegion>
,
<collectingCountry id="E9E37624FFBE5B245368FF770D225E09" box="[258,328,246,266]" name="Croatia" pageId="14" pageNumber="78">Croatia</collectingCountry>
, 45 01
<emphasis id="A380EAA6FFBE5B2453E1FF720DE55E03" bold="true" box="[395,399,243,256]" italics="true" pageId="14" pageNumber="78">
<superScript id="66819BFCFFBE5B2453E1FF720DE55E03" attach="none" box="[395,399,243,256]" fontSize="6" pageId="14" pageNumber="78">0</superScript>
</emphasis>
42.1
<emphasis id="A380EAA6FFBE5B2453D0FF720DA85E03" bold="true" box="[442,450,243,256]" italics="true" pageId="14" pageNumber="78">
<superScript id="66819BFCFFBE5B2453D0FF720DA85E03" attach="right" box="[442,450,243,256]" fontSize="6" pageId="14" pageNumber="78">00</superScript>
</emphasis>
N, 14 09
<emphasis id="A380EAA6FFBE5B245073FF720E775E03" bold="true" box="[537,541,243,256]" italics="true" pageId="14" pageNumber="78">
<superScript id="66819BFCFFBE5B245073FF720E775E03" attach="none" box="[537,541,243,256]" fontSize="6" pageId="14" pageNumber="78">0</superScript>
</emphasis>
17.4
<emphasis id="A380EAA6FFBE5B245022FF720E3A5E03" bold="true" box="[584,592,243,256]" italics="true" pageId="14" pageNumber="78">
<superScript id="66819BFCFFBE5B245022FF720E3A5E03" attach="right" box="[584,592,243,256]" fontSize="6" pageId="14" pageNumber="78">00</superScript>
</emphasis>
E, coll.
<collectorName id="3C015362FFBE5B2450FFFF770F6B5E09" box="[661,769,246,266]" pageId="14" pageNumber="78">M. Wagner</collectorName>
,
<date id="E54A1074FFBE5B245216FE980C915E2E" box="[124,251,281,301]" pageId="14" pageNumber="78" value="2015-07-18">
<collectingDate id="F50EE99CFFBE5B245216FE980C915E2E" box="[124,251,281,301]" pageId="14" pageNumber="78" value="2015-07-18">July 18, 2015</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
<paragraph id="914B36B4FFBE5B2452CEFEBA0CB45D1C" blockId="14.[124,773,139,1167]" pageId="14" pageNumber="78">
<emphasis id="A380EAA6FFBE5B2452CEFEBA0D615E4C" bold="true" box="[164,267,315,335]" pageId="14" pageNumber="78">Synonyms.</emphasis>
<taxonomicName id="56F44D37FFBE5B245365FEBA0E295E4C" authority="Nardo 1827: 9" authorityName="Nardo" authorityPageNumber="9" authorityYear="1827" box="[271,579,315,335]" family="Gobiesocidae" genus="Lepadogaster" kingdom="Animalia" order="Gobiesociformes" pageId="14" pageNumber="78" phylum="Chordata" rank="species" species="piger">
<emphasis id="A380EAA6FFBE5B245365FEBA0DDC5E4C" box="[271,438,315,335]" italics="true" pageId="14" pageNumber="78">Lepadogaster piger</emphasis>
Nardo 1827: 9
</taxonomicName>
(original description; type locality: Rovinj;
<typeStatus id="4E4F8816FFBE5B24532FFEDF0DF05E71" box="[325,410,350,370]" pageId="14" pageNumber="78" type="holotype">holotype</typeStatus>
: unknown);
<taxonomicName id="56F44D37FFBE5B245065FEDF0C8C5E96" authority="Nardo 1833: 548" authorityName="Nardo" authorityPageNumber="548" authorityYear="1833" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="14" pageNumber="78" phylum="Chordata" rank="species" species="prototypus">
<emphasis id="A380EAA6FFBE5B245065FEDF0EAB5E71" box="[527,705,350,370]" italics="true" pageId="14" pageNumber="78">Gouania prototypus</emphasis>
<bibRefCitation id="F5654B45FFBE5B2450A0FEDF0C8C5E96" author="Nardo, G. D." pageId="14" pageNumber="78" pagination="548 - 549" refId="ref28068" refString="Nardo, G. D. (1833). Eine neue Art von Lepadogaster (L. piger). Isis (Oken), 26, 548 - 549." type="journal article" year="1833">Nardo 1833: 548</bibRefCitation>
</taxonomicName>
(original description; type locality: Rovinj?;
<typeStatus id="4E4F8816FFBE5B2450C6FE000F6B5E96" box="[684,769,385,405]" pageId="14" pageNumber="78" type="holotype">holotype</typeStatus>
: unknown),
<taxonomicName id="56F44D37FFBE5B24528DFE220E525EB4" authority="Bonaparte 1846: 64" authorityName="Bonaparte" authorityPageNumber="64" authorityYear="1846" box="[231,568,419,439]" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="14" pageNumber="78" phylum="Chordata" rank="species" species="piger">
<emphasis id="A380EAA6FFBE5B24528DFE220D025EB4" box="[231,360,419,439]" italics="true" pageId="14" pageNumber="78">Gouania piger</emphasis>
Bonaparte 1846: 64
</taxonomicName>
(original description; type locality: unknown;
<typeStatus id="4E4F8816FFBE5B245333FE470DC45ED9" box="[345,430,454,474]" pageId="14" pageNumber="78" type="holotype">holotype</typeStatus>
: unknown);
<taxonomicName id="56F44D37FFBE5B245076FE470D635EFE" authority="Gunther 1861: 515" authorityName="Gunther" authorityPageNumber="515" authorityYear="1861" family="Gobiesocidae" genus="Leptopterygius" kingdom="Animalia" order="Gobiesociformes" pageId="14" pageNumber="78" phylum="Chordata" rank="species" species="piger">
<emphasis id="A380EAA6FFBE5B245076FE470EA55ED9" box="[540,719,454,474]" italics="true" pageId="14" pageNumber="78">Leptopterygius piger</emphasis>
Günther 1861: 515
</taxonomicName>
(original description; type locality: unknown;
<typeStatus id="4E4F8816FFBE5B2450C6FE680F6B5EFE" box="[684,769,489,509]" pageId="14" pageNumber="78" type="holotype">holotype</typeStatus>
: unknown).
</paragraph>
</subSubSection>
<subSubSection id="D9EE653FFFBE5B2452CEFDAF0FEC5E09" pageId="14" pageNumber="78" type="diagnosis">
<paragraph id="914B36B4FFBE5B2452CEFDAF0FEC5E09" blockId="14.[124,773,139,1167]" lastBlockId="14.[821,1470,142,1965]" pageId="14" pageNumber="78">
<emphasis id="A380EAA6FFBE5B2452CEFDAF0D6F5D41" bold="true" box="[164,261,558,578]" pageId="14" pageNumber="78">Diagnosis.</emphasis>
<taxonomicName id="56F44D37FFBE5B245367FDAF0DE15D41" baseAuthorityName="Nardo" baseAuthorityYear="1827" box="[269,395,558,578]" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="14" pageNumber="78" phylum="Chordata" rank="species" species="pigra">
<emphasis id="A380EAA6FFBE5B245367FDAF0DE15D41" box="[269,395,558,578]" italics="true" pageId="14" pageNumber="78">Gouania pigra</emphasis>
</taxonomicName>
differs from the congeneric species by the combination of the following characters: (1) dorsal head profile in lateral view
<emphasis id="A380EAA6FFBE5B245298FDF50C905D84" bold="true" box="[242,250,628,647]" pageId="14" pageNumber="78">“</emphasis>
S
<emphasis id="A380EAA6FFBE5B24536FFDF50D675D84" bold="true" box="[261,269,628,647]" pageId="14" pageNumber="78">”</emphasis>
curved, concave above eye and convex at nape; (2) posterior angle of jaws extends to, or close to, vertical line drawn through anterior edge of anterior nostril; (3) infraorbital invagination below anterior half of eye or below mideye; (4) posterior opercular edge with two tips, upper longer or equal to lower; (5) longitudinal infralateral and suborbital transversal rows of superficial neuromasts placed in the well-defined deep groove; (6) body cross-section behind pectoral fin base half oval to pentagonal with straight ventral side; (7) granules on body shallow and inconspicuous; (8) pectoral rays 13
<emphasis id="A380EAA6FFBE5B2450B3FC080E8F5C9F" bold="true" box="[729,741,905,924]" pageId="14" pageNumber="78">–</emphasis>
16; (9) upper attachment of disc membrane attaching to base of pectoral fin at 12th
<emphasis id="A380EAA6FFBE5B245288FC4E0C845CE1" bold="true" box="[226,238,975,994]" pageId="14" pageNumber="78">–</emphasis>
15th pectoral ray; (10) principal caudal rays 10
<emphasis id="A380EAA6FFBE5B2450C7FC4E0ED35CE1" bold="true" box="[685,697,975,994]" pageId="14" pageNumber="78">–</emphasis>
11; (11) head length 19.5
<emphasis id="A380EAA6FFBE5B245374FC700D405B07" bold="true" box="[286,298,1009,1028]" pageId="14" pageNumber="78">–</emphasis>
22.9% of standard length; (12) pectoral-fin length 5.6
<emphasis id="A380EAA6FFBE5B2452F3FB950CCF5B24" bold="true" box="[153,165,1044,1063]" pageId="14" pageNumber="78">–</emphasis>
7.5% of standard length; (13) prepectoral distance 19.5
<emphasis id="A380EAA6FFBE5B2450CEFB950EDA5B24" bold="true" box="[676,688,1044,1063]" pageId="14" pageNumber="78">–</emphasis>
23.4% of standard length; (14) ventral adhesive disc length, 12.1
<emphasis id="A380EAA6FFBE5B2450F4FBB60EC05B49" bold="true" box="[670,682,1079,1098]" pageId="14" pageNumber="78">–</emphasis>
14.6% of standard length; (15) caudal-fin length 11.1
<emphasis id="A380EAA6FFBE5B245063FBD80E7F5B6F" bold="true" box="[521,533,1113,1132]" pageId="14" pageNumber="78">–</emphasis>
12.7% of standard length; (16) large total number of vertebrae (Supporting Information Table S2; 39
<emphasis id="A380EAA6FFBE5B245127FF0E0F335FA1" bold="true" box="[845,857,143,162]" pageId="14" pageNumber="78">–</emphasis>
40); (17) pharyngeal jaws with elongated ceratobranchial 5, having several larger elongated conical teeth; (18) nasal bones hook-shaped; (19) no star-like pigmentation around eyes, pigmentation generally reduced.
</paragraph>
</subSubSection>
<subSubSection id="D9EE653FFFBE5B255137FE990E3858AE" lastPageId="15" lastPageNumber="79" pageId="14" pageNumber="78" type="description">
<paragraph id="914B36B4FFBE5B255137FE990D2F5E2E" blockId="14.[821,1470,142,1965]" lastBlockId="15.[119,769,142,1965]" lastPageId="15" lastPageNumber="79" pageId="14" pageNumber="78">
<emphasis id="A380EAA6FFBE5B245137FE990FBA5E2F" bold="true" box="[861,976,280,300]" pageId="14" pageNumber="78">Description.</emphasis>
<emphasis id="A380EAA6FFBE5B2451B2FE9808E75E2E" box="[984,1165,281,301]" italics="true" pageId="14" pageNumber="78">General morphology</emphasis>
: Body proportions are given in Table 1. Body very slender and elongated, posteriorly laterally compressed, body depth at pectoral fins
<quantity id="560C9B51FFBE5B2456ECFEDF08895E71" box="[1158,1251,350,370]" metricMagnitude="-1" metricUnit="m" metricValue="2.2733" metricValueMax="2.3876" metricValueMin="2.159" pageId="14" pageNumber="78" unit="in" value="8.95" valueMax="9.4" valueMin="8.5">
8.5
<emphasis id="A380EAA6FFBE5B2456C9FEDE08C55E71" bold="true" box="[1187,1199,351,370]" pageId="14" pageNumber="78">–</emphasis>
9.4 in
</quantity>
SL, body depth at anus
<quantity id="560C9B51FFBE5B24515FFE000FC85E96" box="[821,930,385,405]" metricMagnitude="-1" metricUnit="m" metricValue="2.6162" metricValueMax="2.9972" metricValueMin="2.2352" pageId="14" pageNumber="78" unit="in" value="10.3" valueMax="11.8" valueMin="8.8">
8.8
<emphasis id="A380EAA6FFBE5B245138FE000F345E97" bold="true" box="[850,862,385,404]" pageId="14" pageNumber="78">–</emphasis>
11.8 in
</quantity>
SL, body depth in width at pectoral fins 1.0
<emphasis id="A380EAA6FFBE5B24573CFE0009085E97" bold="true" box="[1366,1378,385,404]" pageId="14" pageNumber="78">–</emphasis>
1.2, body depth in width at anus 0.7
<emphasis id="A380EAA6FFBE5B24565CFE2508285EB4" bold="true" box="[1078,1090,420,439]" pageId="14" pageNumber="78">–</emphasis>
0.9. Body cross-section behind pectoral fin base half oval to pentagonal with straight ventral side. Granules on body shallow and inconspicuous. Head dorsoventrally compressed, head depth in width at orbit 1.5
<emphasis id="A380EAA6FFBE5B245612FD8D08EE5D1C" bold="true" box="[1144,1156,524,543]" pageId="14" pageNumber="78">–</emphasis>
1.9, and moderately small, head length
<quantity id="560C9B51FFBE5B245112FDAF0FB05D41" box="[888,986,558,578]" metricMagnitude="-1" metricUnit="m" metricValue="1.2065" metricValueMax="1.2953999999999999" metricValueMin="1.1176" pageId="14" pageNumber="78" unit="in" value="4.75" valueMax="5.1" valueMin="4.4">
4.4
<emphasis id="A380EAA6FFBE5B2451FFFDAE0FCB5D41" bold="true" box="[917,929,559,578]" pageId="14" pageNumber="78">–</emphasis>
5.1 in
</quantity>
SL, head wider than body width maximum, head width at anterior sucking disc edge
<quantity id="560C9B51FFBE5B2456E9FDD0088B5D66" box="[1155,1249,593,613]" metricMagnitude="-2" metricUnit="m" metricValue="1.905" metricValueMax="2.032" metricValueMin="1.7779999999999998" pageId="14" pageNumber="78" unit="in" value="0.75" valueMax="0.8" valueMin="0.7">
0.7
<emphasis id="A380EAA6FFBE5B2456CAFDD008C65D67" bold="true" box="[1184,1196,593,612]" pageId="14" pageNumber="78">–</emphasis>
0.8 in
</quantity>
body width at pectoral fins. Dorsal head profile
<emphasis id="A380EAA6FFBE5B245672FDF5084A5D84" bold="true" box="[1048,1056,628,647]" pageId="14" pageNumber="78">“</emphasis>
S
<emphasis id="A380EAA6FFBE5B245641FDF508595D84" bold="true" box="[1067,1075,628,647]" pageId="14" pageNumber="78">”</emphasis>
curved, concave above eye and convex at nape. Head rounded in dorsal view. Snout large compared to eyes, preorbital distance
<quantity id="560C9B51FFBE5B245181FD3808215DCE" box="[1003,1099,697,717]" metricMagnitude="-2" metricUnit="m" metricValue="8.001" metricValueMax="9.144" metricValueMin="6.858" pageId="14" pageNumber="78" unit="in" value="3.15" valueMax="3.6" valueMin="2.7">
2.7
<emphasis id="A380EAA6FFBE5B245662FD38087E5DCF" bold="true" box="[1032,1044,697,716]" pageId="14" pageNumber="78">–</emphasis>
3.6 in
</quantity>
head length,
<quantity id="560C9B51FFBE5B2456A5FD3809445DCE" box="[1231,1326,697,717]" metricMagnitude="-3" metricUnit="m" metricValue="8.889999999999999" metricValueMax="10.16" metricValueMin="7.62" pageId="14" pageNumber="78" unit="in" value="0.35" valueMax="0.4" valueMin="0.3">
0.3
<emphasis id="A380EAA6FFBE5B245686FD3808925DCF" bold="true" box="[1260,1272,697,716]" pageId="14" pageNumber="78">–</emphasis>
0.4 in
</quantity>
horizontal eye diameter. Snout wide, not produced, blunt. Internostril space convex to gently convex. Eyes dorsolateral, rounded or drop-like with slightly pointed lower eye edge. Eyes small,
<quantity id="560C9B51FFBE5B2456E6FCA0089D5C36" box="[1164,1271,801,821]" metricMagnitude="-1" metricUnit="m" metricValue="2.5780999999999996" metricValueMax="2.8447999999999998" metricValueMin="2.3114" pageId="14" pageNumber="78" unit="in" value="10.149999999999999" valueMax="11.2" valueMin="9.1">
9.1
<emphasis id="A380EAA6FFBE5B2456C0FCA008DC5C37" bold="true" box="[1194,1206,801,820]" pageId="14" pageNumber="78">–</emphasis>
11.2 in
</quantity>
head length, vertical diameter of the eye
<quantity id="560C9B51FFBE5B245197FCC208345C54" box="[1021,1118,835,855]" metricMagnitude="-2" metricUnit="m" metricValue="2.032" metricValueMax="2.286" metricValueMin="1.7779999999999998" pageId="14" pageNumber="78" unit="in" value="0.8" valueMax="0.9" valueMin="0.7">
0.7
<emphasis id="A380EAA6FFBE5B245670FCC5084C5C54" bold="true" box="[1050,1062,836,855]" pageId="14" pageNumber="78">–</emphasis>
0.9 in
</quantity>
horizontal eye diameter. Infraorbital invagination below anterior half of eye or below mideye. Interorbital distance wide,
<quantity id="560C9B51FFBE5B2451D4FC0808715C9E" box="[958,1051,905,925]" metricMagnitude="-3" metricUnit="m" metricValue="8.889999999999999" metricValueMax="10.16" metricValueMin="7.62" pageId="14" pageNumber="78" unit="in" value="0.35" valueMax="0.4" valueMin="0.3">
0.3
<emphasis id="A380EAA6FFBE5B2451B1FC080F8D5C9F" bold="true" box="[987,999,905,924]" pageId="14" pageNumber="78">–</emphasis>
0.4 in
</quantity>
horizontal eye diameter.
<collectingRegion id="5330F856FFBE5B24576FFC08092E5C9E" box="[1285,1348,905,925]" country="France" name="Centre" pageId="14" pageNumber="78">Centre</collectingRegion>
of eye much closer to tip of snout than to posterior margin of operculum, preorbital distance in postorbital distance 1.8
<emphasis id="A380EAA6FFBE5B2456F7FC4E08C35CE1" bold="true" box="[1181,1193,975,994]" pageId="14" pageNumber="78">–</emphasis>
2.4. Anterior and posterior nostrils long tubes of about equal length. Nostrils well separated and posterior nostril located behind and dorsally to the anterior edge of eyes. Single large dermal flap of leaf shape at the posterior margin of anterior nostril, longer than nostril. Posterior nostril rim crenate or villose with no extension. Head lateral line system with canals with pores and with superficial neuromasts arranged in rows. Head canals reduced and pores small. Single pore in nasal canal near posterior nostril. Single pore in postorbital canal close to posterior eye edge. Two pores in mandibular canal, anterior one close to anteriolateral angle of mouth, posterior pore slightly in front of vertical of posterior angle of jaws, posterior pore usually more prominent. Lachrymal as well as preopercular canals and pores absent. Rows of superficial neuromasts as follows: SR 2, NR 3, LIR 21
<emphasis id="A380EAA6FFBE5B245624FA1008305AA7" bold="true" box="[1102,1114,1425,1444]" pageId="14" pageNumber="78">–</emphasis>
26, STR 2
<emphasis id="A380EAA6FFBE5B2456DCFA1008A85AA7" bold="true" box="[1206,1218,1425,1444]" pageId="14" pageNumber="78">–</emphasis>
3, POR 2
<emphasis id="A380EAA6FFBE5B245772FA10094E5AA7" bold="true" box="[1304,1316,1425,1444]" pageId="14" pageNumber="78">–</emphasis>
3, PTR 2, SLR 5, MR 8
<emphasis id="A380EAA6FFBE5B245100FA350F1C5AC4" bold="true" box="[874,886,1460,1479]" pageId="14" pageNumber="78">–</emphasis>
12, AVR 2, PVR 1, ADR 3, PDR 1
<emphasis id="A380EAA6FFBE5B2456D1FA3508AD5AC4" bold="true" box="[1211,1223,1460,1479]" pageId="14" pageNumber="78">–</emphasis>
2, HR 3, SR1 3, SR2 1
<emphasis id="A380EAA6FFBE5B2457F5FA3509C15AC4" bold="true" box="[1439,1451,1460,1479]" pageId="14" pageNumber="78">–</emphasis>
2, DLR 6
<emphasis id="A380EAA6FFBE5B24511BFA560F175AE9" bold="true" box="[881,893,1495,1514]" pageId="14" pageNumber="78">–</emphasis>
10, VLR 12
<emphasis id="A380EAA6FFBE5B245180FA560F9C5AE9" bold="true" box="[1002,1014,1495,1514]" pageId="14" pageNumber="78">–</emphasis>
16. STR and LIR rows of superficial neuromasts placed in the well-defined deep groove. DLR row of superficial neuromasts anteriorly starts above pectoral fin, continuously dorsolateral and ends posteriorly variably: downwards at midlateral level or above it, at vertical from anus, or in front or behind it. VLR anteriorly starts behind pectoral fin base, continuously ventrolateral and ends posteriorly upwards with last papilla nearly at midlateral level at caudal fin base or more distant from it. Mouth terminal, upper and lower lips end about equally, lips fleshy, upper lip larger than the lower lip. Posterior angle of jaws extends to, or close to, a vertical line drawn through the anterior edge of the anterior nostril. Chin with bilobed or single lobed fold at anterior edge covering MR row of superficial neuromasts. Gill membrane is attached to isthmus, gill opening starting at base of pectoral fin, with upper attachment of gill membrane opposite to 3rd to 5th pectoral ray. Posterior opercular edge with two tips, upper longer or equal to lower. No subopercular spine. No fleshy pad present on lower pectoral base. Urogenital papilla present. Preanus length in postanus length 0.7
<emphasis id="A380EAA6FFBF5B2553F7FF760DC35E09" bold="true" box="[413,425,247,266]" pageId="15" pageNumber="79">–</emphasis>
0.9. Anal papillae absent, the area around anus wrinkled.
</paragraph>
<caption id="C58B663CFFBE5B245216F8A50CAE58AE" ID-DOI="http://doi.org/10.5281/zenodo.10928940" ID-Zenodo-Dep="10928940" httpUri="https://zenodo.org/record/10928940/files/figure.png" pageId="14" pageNumber="78" startId="14.[124,136,1828,1847]" targetBox="[131,768,1235,1786]" targetPageId="14" targetType="figure">
<paragraph id="914B36B4FFBE5B245216F8A50CAE58AE" blockId="14.[124,752,1827,1965]" pageId="14" pageNumber="78">
<emphasis id="A380EAA6FFBE5B245216F8A50C865834" bold="true" box="[124,236,1828,1847]" pageId="14" pageNumber="78">F I G U R E 8</emphasis>
<taxonomicName id="56F44D37FFBE5B245360F8A20E665834" authority="(Nardo, 1827)" baseAuthorityName="Nardo" baseAuthorityYear="1827" box="[266,524,1827,1847]" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="14" pageNumber="78" phylum="Chordata" rank="species" species="pigra">
<emphasis id="A380EAA6FFBE5B245360F8A20DEF5834" box="[266,389,1827,1847]" italics="true" pageId="14" pageNumber="78">Gouania pigra</emphasis>
(Nardo, 1827)
</taxonomicName>
, PMR VP3529, neotype, female, 43.12+5.03 mm, Glavotok, Krk, Croatia. Lateral view of specimen preserved in 4% formaldehyde (top). Lateral, dorsal and ventral view, alive (below). Photographs by M. Wagner and M. Kovacˇic
</paragraph>
</caption>
<paragraph id="914B36B4FFBF5B2552F5FEBA0D335C79" blockId="15.[119,769,142,1965]" pageId="15" pageNumber="79">
<emphasis id="A380EAA6FFBF5B2552F5FEBA0CA85E4C" box="[159,194,315,335]" italics="true" pageId="15" pageNumber="79">Fins</emphasis>
. Rudimentary dorsal and anal fins located well posteriorly and short, reduced to low ridges with very weak rays, connected to caudal fin. Pectoral rays 13
<emphasis id="A380EAA6FFBF5B255354FE000D205E97" bold="true" box="[318,330,385,404]" pageId="15" pageNumber="79">–</emphasis>
16. Caudal fin rounded, principal caudal rays 10
<emphasis id="A380EAA6FFBF5B2552FAFE250CF65EB4" bold="true" box="[144,156,420,439]" pageId="15" pageNumber="79">–</emphasis>
11. Ventral adhesive disc (Figure 4d) of
<emphasis id="A380EAA6FFBF5B25504CFE250E445EB4" bold="true" box="[550,558,420,439]" pageId="15" pageNumber="79">“</emphasis>
double
<emphasis id="A380EAA6FFBF5B255007FE250E1F5EB4" bold="true" box="[621,629,420,439]" pageId="15" pageNumber="79">”</emphasis>
<typeStatus id="4E4F8816FFBF5B255014FE220EC35EB4" box="[638,681,419,439]" pageId="15" pageNumber="79">type</typeStatus>
, anterior margin crenate or straight with large invagination on each lateral side and central invagination at midventral; posterior margin crenate or straight. Disc very small, disc length
<quantity id="560C9B51FFBF5B2553BCFD8A0E5D5D1C" box="[470,567,523,543]" metricMagnitude="-1" metricUnit="m" metricValue="1.9304" metricValueMax="2.1082" metricValueMin="1.7526" pageId="15" pageNumber="79" unit="in" value="7.6" valueMax="8.3" valueMin="6.9">
6.9
<emphasis id="A380EAA6FFBF5B25539EFD8D0E6A5D1C" bold="true" box="[500,512,524,543]" pageId="15" pageNumber="79">–</emphasis>
8.3 in
</quantity>
SL, its width slightly larger than its length, width in length 0.8
<emphasis id="A380EAA6FFBF5B25539EFDAE0E6A5D41" bold="true" box="[500,512,559,578]" pageId="15" pageNumber="79">–</emphasis>
1.0. No papillae in region A and flattened papillae in regions B and C. In region B one or two rows of papillae with total papillae count 11
<emphasis id="A380EAA6FFBF5B2553B1FDF50D8D5D84" bold="true" box="[475,487,628,647]" pageId="15" pageNumber="79">–</emphasis>
23 and in region C one or two rows of papillae with total papillae count 4
<emphasis id="A380EAA6FFBF5B255068FD160E645DA9" bold="true" box="[514,526,663,682]" pageId="15" pageNumber="79">–</emphasis>
11. No inner row of papillae on lateral sides of the central part of the anterior disc. Upper attachment of disc membrane attaching to base of pectoral fin at 12th
<emphasis id="A380EAA6FFBF5B2552C9FD7E0CC55C11" bold="true" box="[163,175,767,786]" pageId="15" pageNumber="79">–</emphasis>
15th pectoral ray,
<emphasis id="A380EAA6FFBF5B255331FD7F0D055C11" box="[347,367,766,786]" italics="true" pageId="15" pageNumber="79">i.e</emphasis>
., at ultimate or penultimate ray. Males can have two prominent seemingly perfused finger-like extensions on each site of sucking disc that sometimes equal or exceed length of disc region A (Figure 4f).
</paragraph>
<paragraph id="914B36B4FFBF5B2552F5FC080C8F5BB1" blockId="15.[119,769,142,1965]" pageId="15" pageNumber="79">
<emphasis id="A380EAA6FFBF5B2552F5FC080D6C5C9E" box="[159,262,905,925]" italics="true" pageId="15" pageNumber="79">Colouration</emphasis>
. Background colouration of live specimens white to flesh-coloured, slightly transparent (Figures 1b and 8) and no starshaped pigmentation around eyes. In life body almost pigmentless or with very small pigments, leading to an irregular marbled pattern, but never as strong as in other
<taxonomicName id="56F44D37FFBF5B2553E8FB920E7D5B24" box="[386,535,1043,1063]" pageId="15" pageNumber="79">
<emphasis id="A380EAA6FFBF5B2553E8FB920DA65B24" box="[386,460,1043,1063]" italics="true" pageId="15" pageNumber="79">Gouania</emphasis>
species
</taxonomicName>
(Supporting Information File S1). Formaldehyde fixed specimens white-yellow and without pigments. Ethanol fixed specimens white to skin-coloured pigmentation reduced. For more pictures of life colouration see Supporting Information File S1.
</paragraph>
<paragraph id="914B36B4FFBF5B2552F5FB400E3858AE" blockId="15.[119,769,142,1965]" pageId="15" pageNumber="79">
<emphasis id="A380EAA6FFBF5B2552F5FB400D1F5BD6" box="[159,373,1217,1237]" italics="true" pageId="15" pageNumber="79">Dentition and osteology</emphasis>
. Upper jaw with outer row of about eight (one side) medium-sized caniniforms frontally. Behind them inner small conical teeth irregularly scattered in two separate (left and right) drop-like patches medially wide about 5
<emphasis id="A380EAA6FFBF5B255381FAA80D9D5A3F" bold="true" box="[491,503,1321,1340]" pageId="15" pageNumber="79">–</emphasis>
6 teeth, becoming narrowed to a single row of teeth laterally. Outer row continues laterally as four large caniniforms of variable size, followed behind by four to five medium-sized caniniforms. Lower jaw with outer row of 10 to 12 (one side) medium-sized caniniforms frontally. Behind them single broad patch of small conical inner teeth medially wide about 5
<emphasis id="A380EAA6FFBF5B2550CDFA560ED95AE9" bold="true" box="[679,691,1495,1514]" pageId="15" pageNumber="79">–</emphasis>
6 teeth, becoming narrowed to a single row of teeth laterally. The single row of about six larger caniniforms continuous laterally. Pharyngeal jaws with elongated ceratobranchial 5, having several (about 4
<emphasis id="A380EAA6FFBF5B255021F9BE0E3D5951" bold="true" box="[587,599,1599,1618]" pageId="15" pageNumber="79">–</emphasis>
5) larger elongated conical teeth (Figure 5d), pharyngobranchial 3 toothplate not visible on 3D models from microcomputed tomography (microCT) images. Number of vertebrae 39
<emphasis id="A380EAA6FFBF5B25535AF9260D5659B9" bold="true" box="[304,316,1703,1722]" pageId="15" pageNumber="79">–</emphasis>
40, abdominal 17 and caudal 22
<emphasis id="A380EAA6FFBF5B255000F9260E1C59B9" bold="true" box="[618,630,1703,1722]" pageId="15" pageNumber="79">–</emphasis>
23 (Supporting Information Table S2). The first gill arch with hemibranch, the 2nd
<emphasis id="A380EAA6FFBF5B2550BEF9480E8A59DF" bold="true" box="[724,736,1737,1756]" pageId="15" pageNumber="79">–</emphasis>
4th gill arches with holobranchs. Six small, pointed rakers on third gill arch. The subopercular element is present/absent as the terminal bone posteriorly. Subopercle indistinguishable from opercle, shaped as its posterior elongated extension, not forming or having subopercular spine. Six branchiostegals. Nasal bones hook-shaped. Maxillary, premaxillary, nasal and ceratobranchial 5 bones shaped as on Figure 5d.
</paragraph>
</subSubSection>
<subSubSection id="D9EE653FFFBF5B255132FF0F08495E2E" pageId="15" pageNumber="79" type="etymology">
<paragraph id="914B36B4FFBF5B255132FF0F08495E2E" blockId="15.[816,1465,142,1965]" pageId="15" pageNumber="79">
<emphasis id="A380EAA6FFBF5B255132FF0F0FA85FA1" bold="true" box="[856,962,142,162]" pageId="15" pageNumber="79">Etymology.</emphasis>
The Latin adjective masculine singular nominative
<taxonomicName id="56F44D37FFBF5B25515AFF300F0C5FC6" authorityName="Gunther" authorityYear="1861" box="[816,870,177,197]" family="Gobiesocidae" genus="Leptopterygius" kingdom="Animalia" order="Gobiesociformes" pageId="15" pageNumber="79" phylum="Chordata" rank="species" species="piger">
<emphasis id="A380EAA6FFBF5B25515AFF300F525FC7" bold="true" box="[816,824,177,196]" pageId="15" pageNumber="79">“</emphasis>
piger
</taxonomicName>
<emphasis id="A380EAA6FFBF5B25510CFF300F045FC7" bold="true" box="[870,878,177,196]" pageId="15" pageNumber="79">”</emphasis>
in
<taxonomicName id="56F44D37FFBF5B2551E7FF3008D85FC6" authority="Nardo 1827" authorityName="Nardo" authorityYear="1827" box="[909,1202,177,197]" family="Gobiesocidae" genus="Lepadogaster" kingdom="Animalia" order="Gobiesociformes" pageId="15" pageNumber="79" phylum="Chordata" rank="species" species="piger">
<emphasis id="A380EAA6FFBF5B2551E7FF30085C5FC6" box="[909,1078,177,197]" italics="true" pageId="15" pageNumber="79">Lepadogaster piger</emphasis>
Nardo 1827
</taxonomicName>
, meaning slow-moving, was changed to
<taxonomicName id="56F44D37FFBF5B2551F7FF550FB95FE4" baseAuthorityName="Nardo" baseAuthorityYear="1827" box="[925,979,211,231]" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="15" pageNumber="79" phylum="Chordata" rank="species" species="pigra">
<emphasis id="A380EAA6FFBF5B2551F7FF550FCF5FE4" bold="true" box="[925,933,212,231]" pageId="15" pageNumber="79">“</emphasis>
pigra
</taxonomicName>
<emphasis id="A380EAA6FFBF5B2551B8FF550FB05FE4" bold="true" box="[978,986,212,231]" pageId="15" pageNumber="79">”</emphasis>
in
<taxonomicName id="56F44D37FFBF5B25519DFF5208195FE4" baseAuthorityName="Nardo" baseAuthorityYear="1827" box="[1015,1139,211,231]" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="15" pageNumber="79" phylum="Chordata" rank="species" species="pigra">
<emphasis id="A380EAA6FFBF5B25519DFF5208195FE4" box="[1015,1139,211,231]" italics="true" pageId="15" pageNumber="79">Gouania pigra</emphasis>
</taxonomicName>
which is an adjective feminine singular nominative, following the necessity of agreement in gender (Article 31.2,
<collectionCode id="F7E5AE71FFBF5B2551C1FE980FB45E2E" box="[939,990,281,301]" pageId="15" pageNumber="79">ICZN</collectionCode>
, 1999).
</paragraph>
</subSubSection>
<subSubSection id="D9EE653FFFBF5B255132FEBA08015DA9" pageId="15" pageNumber="79" type="biology_ecology">
<paragraph id="914B36B4FFBF5B255132FEBA08015DA9" blockId="15.[816,1465,142,1965]" pageId="15" pageNumber="79">
<emphasis id="A380EAA6FFBF5B255132FEBA09585E4C" bold="true" box="[856,1330,315,335]" pageId="15" pageNumber="79">Ecology and geographical distribution (Figure 1a).</emphasis>
<taxonomicName id="56F44D37FFBF5B255753FEBA09DD5E4C" baseAuthorityName="Nardo" baseAuthorityYear="1827" box="[1337,1463,315,335]" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="15" pageNumber="79" phylum="Chordata" rank="species" species="pigra">
<emphasis id="A380EAA6FFBF5B255753FEBA09DD5E4C" box="[1337,1463,315,335]" italics="true" pageId="15" pageNumber="79">Gouania pigra</emphasis>
</taxonomicName>
is endemic to the Adriatic Sea and hence the only purely marine endemic fish known for this basin. The southernmost record is from
<collectingRegion id="5330F856FFBF5B255751FE0009015E96" box="[1339,1387,385,405]" country="Albania" name="Vlore" pageId="15" pageNumber="79">Vlorë</collectingRegion>
(Albany) and it was (so far) not found on Corfu. Quantitative data on ecology is largely lacking. Throughout its distribution range the species occurs in sympatry with
<taxonomicName id="56F44D37FFBF5B2551D0FE6808755EFE" authority="Wagner &amp; Kovac &amp; Koblmüller, 2021" authorityName="Wagner &amp; Kovac &amp; Koblmüller" authorityYear="2021" box="[954,1055,489,509]" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="15" pageNumber="79" phylum="Chordata" rank="species" species="adriatica" status="sp. nov.">
<emphasis id="A380EAA6FFBF5B2551D0FE6808755EFE" box="[954,1055,489,509]" italics="true" pageId="15" pageNumber="79">G. adriatica</emphasis>
</taxonomicName>
<emphasis id="A380EAA6FFBF5B255642FE69081B5EFF" bold="true" box="[1064,1137,488,508]" pageId="15" pageNumber="79">
<taxonomicNameLabel id="B8B357DDFFBF5B255642FE69081B5EFF" box="[1064,1137,488,508]" pageId="15" pageNumber="79" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Sympatric and syntopic occurrence with both
<taxonomicName id="56F44D37FFBF5B2551FAFD8A0F995D1C" authority="Wagner &amp; Kovac &amp; Koblmüller, 2021" authorityName="Wagner &amp; Kovac &amp; Koblmüller" authorityYear="2021" box="[912,1011,523,543]" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="15" pageNumber="79" phylum="Chordata" rank="species" species="adriatica" status="sp. nov.">
<emphasis id="A380EAA6FFBF5B2551FAFD8A0F995D1C" box="[912,1011,523,543]" italics="true" pageId="15" pageNumber="79">G. adriatica</emphasis>
</taxonomicName>
<emphasis id="A380EAA6FFBF5B255191FD8A08295D1C" bold="true" box="[1019,1091,523,543]" pageId="15" pageNumber="79">
<taxonomicNameLabel id="B8B357DDFFBF5B255191FD8A08295D1C" box="[1019,1091,523,543]" pageId="15" pageNumber="79" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="56F44D37FFBF5B255618FD8A08885D1C" authority="Wagner &amp; Kovac &amp; Koblmüller, 2021" authorityName="Wagner &amp; Kovac &amp; Koblmüller" authorityYear="2021" box="[1138,1250,523,543]" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="15" pageNumber="79" phylum="Chordata" rank="species" species="hofrichteri" status="sp. nov.">
<emphasis id="A380EAA6FFBF5B255618FD8A08885D1C" box="[1138,1250,523,543]" italics="true" pageId="15" pageNumber="79">G. hofrichteri</emphasis>
</taxonomicName>
<emphasis id="A380EAA6FFBF5B255683FD8A09585D1C" bold="true" box="[1257,1330,523,543]" pageId="15" pageNumber="79">
<taxonomicNameLabel id="B8B357DDFFBF5B255683FD8A09585D1C" box="[1257,1330,523,543]" pageId="15" pageNumber="79" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is only known from Pelješac (Figure 1c). The species inhabits intertidal pebble beaches, where it might be found even above the waterline during low tide, and only rarely occurs in fields of larger boulders. See Supporting Information Video S1 for locomotion behaviour.
</paragraph>
</subSubSection>
<subSubSection id="D9EE653FFFBF5B3A5132FD390DA95AC4" lastPageId="16" lastPageNumber="80" pageId="15" pageNumber="79" type="discussion">
<paragraph id="914B36B4FFBF5B3A5132FD390E645DEC" blockId="15.[816,1465,142,1965]" lastBlockId="16.[124,773,142,1479]" lastPageId="16" lastPageNumber="80" pageId="15" pageNumber="79">
<emphasis id="A380EAA6FFBF5B255132FD390FDA5DCF" bold="true" box="[856,944,696,716]" pageId="15" pageNumber="79">Remarks.</emphasis>
<taxonomicName id="56F44D37FFBF5B2551D3FD3808E75DCE" authority="(Nardo 1827)" baseAuthorityName="Nardo" baseAuthorityYear="1827" box="[953,1165,697,717]" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="15" pageNumber="79" phylum="Chordata" rank="species" species="pigra">
<emphasis id="A380EAA6FFBF5B2551D3FD3808685DCE" box="[953,1026,697,717]" italics="true" pageId="15" pageNumber="79">G. pigra</emphasis>
(Nardo 1827)
</taxonomicName>
differs from all other
<taxonomicName id="56F44D37FFBF5B255707FD380F195DEC" pageId="15" pageNumber="79">
<emphasis id="A380EAA6FFBF5B255707FD3809DD5DCE" box="[1389,1463,697,717]" italics="true" pageId="15" pageNumber="79">Gouania</emphasis>
species
</taxonomicName>
by the posterior angle of jaws extending to, or close to, a vertical line drawn through the anterior edge of the anterior nostril (
<emphasis id="A380EAA6FFBF5B2557CAFD7F09D85C11" box="[1440,1458,766,786]" italics="true" pageId="15" pageNumber="79">vs</emphasis>
. posterior angle of jaws extending to between a vertical line drawn through the posterior edge of the anterior nostril and a vertical line drawn through the anterior edge of the eye in three other
<taxonomicName id="56F44D37FFBF5B255707FCE70F195C9E" pageId="15" pageNumber="79">
<emphasis id="A380EAA6FFBF5B255707FCE709DD5C79" box="[1389,1463,870,890]" italics="true" pageId="15" pageNumber="79">Gouania</emphasis>
species
</taxonomicName>
or even to below the anterior edge of the eyes to the anterior part of the eye in
<taxonomicName id="56F44D37FFBF5B2551B6FC2A082C5CBC" authority="Wagner &amp; Kovac &amp; Koblmüller, 2021" authorityName="Wagner &amp; Kovac &amp; Koblmüller" authorityYear="2021" box="[988,1094,939,959]" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="15" pageNumber="79" phylum="Chordata" rank="species" species="orientalis" status="sp. nov.">
<emphasis id="A380EAA6FFBF5B2551B6FC2A082C5CBC" box="[988,1094,939,959]" italics="true" pageId="15" pageNumber="79">G. orientalis</emphasis>
</taxonomicName>
<emphasis id="A380EAA6FFBF5B255627FC2A08FD5CBC" bold="true" box="[1101,1175,939,959]" pageId="15" pageNumber="79">
<taxonomicNameLabel id="B8B357DDFFBF5B255627FC2A08FD5CBC" box="[1101,1175,939,959]" pageId="15" pageNumber="79" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
).
<taxonomicName id="56F44D37FFBF5B2556C2FC2A08845CBC" baseAuthorityName="Nardo" baseAuthorityYear="1827" box="[1192,1262,939,959]" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="15" pageNumber="79" phylum="Chordata" rank="species" species="pigra">
<emphasis id="A380EAA6FFBF5B2556C2FC2A08845CBC" box="[1192,1262,939,959]" italics="true" pageId="15" pageNumber="79">G. pigra</emphasis>
</taxonomicName>
differs from all other congeneric species also by its overall reduced pigmentation.
<taxonomicName id="56F44D37FFBF5B25571AFC4F09DD5CE1" baseAuthorityName="Nardo" baseAuthorityYear="1827" box="[1392,1463,974,994]" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="15" pageNumber="79" phylum="Chordata" rank="species" species="pigra">
<emphasis id="A380EAA6FFBF5B25571AFC4F09DD5CE1" box="[1392,1463,974,994]" italics="true" pageId="15" pageNumber="79">G. pigra</emphasis>
</taxonomicName>
differs from the three stout-bodied
<taxonomicName id="56F44D37FFBF5B2556E2FC7009775B06" box="[1160,1309,1009,1029]" pageId="15" pageNumber="79">
<emphasis id="A380EAA6FFBF5B2556E2FC7008B85B06" box="[1160,1234,1009,1029]" italics="true" pageId="15" pageNumber="79">Gouania</emphasis>
species
</taxonomicName>
(
<taxonomicName id="56F44D37FFBF5B255741FC7009FF5B06" authority="Wagner &amp; Kovac &amp; Koblmüller, 2021" authorityName="Wagner &amp; Kovac &amp; Koblmüller" authorityYear="2021" box="[1323,1429,1009,1029]" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="15" pageNumber="79" phylum="Chordata" rank="species" species="adriatica" status="sp. nov.">
<emphasis id="A380EAA6FFBF5B255741FC7009FF5B06" box="[1323,1429,1009,1029]" italics="true" pageId="15" pageNumber="79">G. adriatica</emphasis>
</taxonomicName>
<emphasis id="A380EAA6FFBF5B2557F7FC710F335B24" bold="true" pageId="15" pageNumber="79">
<taxonomicNameLabel id="B8B357DDFFBF5B2557F7FC710F335B24" pageId="15" pageNumber="79" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="56F44D37FFBF5B255109FB920FA75B24" authority="Wagner &amp; Kovac &amp; Koblmüller, 2021" authorityName="Wagner &amp; Kovac &amp; Koblmüller" authorityYear="2021" box="[867,973,1043,1063]" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="15" pageNumber="79" phylum="Chordata" rank="species" species="orientalis" status="sp. nov.">
<emphasis id="A380EAA6FFBF5B255109FB920FA75B24" box="[867,973,1043,1063]" italics="true" pageId="15" pageNumber="79">G. orientalis</emphasis>
</taxonomicName>
<emphasis id="A380EAA6FFBF5B2551B9FB9208775B24" bold="true" box="[979,1053,1043,1063]" pageId="15" pageNumber="79">
<taxonomicNameLabel id="B8B357DDFFBF5B2551B9FB9208775B24" box="[979,1053,1043,1063]" pageId="15" pageNumber="79" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="56F44D37FFBF5B255620FB9208AB5B24" baseAuthorityName="Risso" baseAuthorityYear="1810" box="[1098,1217,1043,1063]" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="15" pageNumber="79" phylum="Chordata" rank="species" species="willdenowi">
<emphasis id="A380EAA6FFBF5B255620FB9208AB5B24" box="[1098,1217,1043,1063]" italics="true" pageId="15" pageNumber="79">G. willdenowi</emphasis>
</taxonomicName>
) by dorsal head profile
<emphasis id="A380EAA6FFBF5B2557F6FB9509CE5B24" bold="true" box="[1436,1444,1044,1063]" pageId="15" pageNumber="79">“</emphasis>
S
<emphasis id="A380EAA6FFBF5B2557DAFB9509D25B24" bold="true" box="[1456,1464,1044,1063]" pageId="15" pageNumber="79">”</emphasis>
curved, concave above eye and convex at nape (
<emphasis id="A380EAA6FFBF5B255685FBB7096B5B49" box="[1263,1281,1078,1098]" italics="true" pageId="15" pageNumber="79">vs</emphasis>
. dorsal head profile straight between nape above eye and upper lip tip), the absence of star-like pigmentation around eyes (
<emphasis id="A380EAA6FFBF5B2556EBFBFA08F95B8C" box="[1153,1171,1147,1167]" italics="true" pageId="15" pageNumber="79">vs</emphasis>
. no pigmentation around eyes) and a large number of vertebrae (Supporting Information Table S2; 39
<emphasis id="A380EAA6FFBF5B255122FB400F3E5BD7" bold="true" box="[840,852,1217,1236]" pageId="15" pageNumber="79">–</emphasis>
40
<emphasis id="A380EAA6FFBF5B25511CFB400FE25BD6" box="[886,904,1217,1237]" italics="true" pageId="15" pageNumber="79">vs</emphasis>
. 35
<emphasis id="A380EAA6FFBF5B2551DAFB400FD65BD7" bold="true" box="[944,956,1217,1236]" pageId="15" pageNumber="79">–</emphasis>
38). Five morphometric characters as percentages of standard length of
<taxonomicName id="56F44D37FFBF5B255188FB6208425BF4" baseAuthorityName="Nardo" baseAuthorityYear="1827" box="[994,1064,1251,1271]" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="15" pageNumber="79" phylum="Chordata" rank="species" species="pigra">
<emphasis id="A380EAA6FFBF5B255188FB6208425BF4" box="[994,1064,1251,1271]" italics="true" pageId="15" pageNumber="79">G. pigra</emphasis>
</taxonomicName>
are nonoverlapping in range with all three stout-bodied
<taxonomicName id="56F44D37FFBF5B2551D8FA870F965A19" authorityName="Nardo" authorityYear="1833" box="[946,1020,1286,1306]" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="15" pageNumber="79" phylum="Chordata" rank="genus">
<emphasis id="A380EAA6FFBF5B2551D8FA870F965A19" box="[946,1020,1286,1306]" italics="true" pageId="15" pageNumber="79">Gouania</emphasis>
</taxonomicName>
: head length, pectoral-fin length, prepectoral distance, ventral adhesive disc length and caudal fin-length (values in the Table 1). There are also morphometric characters nonoverlapping in range with one or two out of three stout-bodied
<taxonomicName id="56F44D37FFBF5B255748FAEF09DD5A81" box="[1314,1463,1390,1410]" pageId="15" pageNumber="79">
<emphasis id="A380EAA6FFBF5B255748FAEF09065A81" box="[1314,1388,1390,1410]" italics="true" pageId="15" pageNumber="79">Gouania</emphasis>
species
</taxonomicName>
(Table 1). In addition, it differs from
<taxonomicName id="56F44D37FFBF5B255616FA1008985AA6" baseAuthorityName="Risso" baseAuthorityYear="1810" box="[1148,1266,1425,1445]" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="15" pageNumber="79" phylum="Chordata" rank="species" species="willdenowi">
<emphasis id="A380EAA6FFBF5B255616FA1008985AA6" box="[1148,1266,1425,1445]" italics="true" pageId="15" pageNumber="79">G. willdenowi</emphasis>
</taxonomicName>
by infraorbital invagination below anterior half of eye or below mideye (
<emphasis id="A380EAA6FFBF5B25575BFA3209295AC4" box="[1329,1347,1459,1479]" italics="true" pageId="15" pageNumber="79">vs</emphasis>
. infraorbital invagination vertical to posterior part of eye); from
<taxonomicName id="56F44D37FFBF5B25574DFA5709FE5AE9" authority="Wagner &amp; Kovac &amp; Koblmüller, 2021" authorityName="Wagner &amp; Kovac &amp; Koblmüller" authorityYear="2021" box="[1319,1428,1494,1514]" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="15" pageNumber="79" phylum="Chordata" rank="species" species="orientalis" status="sp. nov.">
<emphasis id="A380EAA6FFBF5B25574DFA5709FE5AE9" box="[1319,1428,1494,1514]" italics="true" pageId="15" pageNumber="79">G. orientalis</emphasis>
</taxonomicName>
<emphasis id="A380EAA6FFBF5B2557F7FA570F33590F" bold="true" pageId="15" pageNumber="79">
<taxonomicNameLabel id="B8B357DDFFBF5B2557F7FA570F33590F" pageId="15" pageNumber="79" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
by pectoral rays 13
<emphasis id="A380EAA6FFBF5B255672FA78084E590F" bold="true" box="[1048,1060,1529,1548]" pageId="15" pageNumber="79">–</emphasis>
16 (
<emphasis id="A380EAA6FFBF5B255620FA780836590E" box="[1098,1116,1529,1549]" italics="true" pageId="15" pageNumber="79">vs</emphasis>
. pectoral rays 17
<emphasis id="A380EAA6FFBF5B255768FA780964590F" bold="true" box="[1282,1294,1529,1548]" pageId="15" pageNumber="79">–</emphasis>
19), upper attachment of disc membrane attaching to base of pectoral fin at 12th
<emphasis id="A380EAA6FFBF5B2557EAF99D09E6592C" bold="true" box="[1408,1420,1564,1583]" pageId="15" pageNumber="79">–</emphasis>
15th pectoral ray (
<emphasis id="A380EAA6FFBF5B2551C3F9BF0FD15951" box="[937,955,1598,1618]" italics="true" pageId="15" pageNumber="79">vs</emphasis>
. upper attachment of disc membrane attaching to base of pectoral fin at 16th
<emphasis id="A380EAA6FFBF5B255664F9E008705977" bold="true" box="[1038,1050,1633,1652]" pageId="15" pageNumber="79">–</emphasis>
18th pectoral ray), infraorbital invagination below anterior half of eye or below mideye (
<emphasis id="A380EAA6FFBF5B2556B2F90208805994" box="[1240,1258,1667,1687]" italics="true" pageId="15" pageNumber="79">vs</emphasis>
. infraorbital invagination vertical to posterior part of eye); and from
<taxonomicName id="56F44D37FFBF5B25568CF927092759B9" authority="Wagner &amp; Kovac &amp; Koblmüller, 2021" authorityName="Wagner &amp; Kovac &amp; Koblmüller" authorityYear="2021" box="[1254,1357,1702,1722]" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="15" pageNumber="79" phylum="Chordata" rank="species" species="adriatica" status="sp. nov.">
<emphasis id="A380EAA6FFBF5B25568CF927092759B9" box="[1254,1357,1702,1722]" italics="true" pageId="15" pageNumber="79">G. adriatica</emphasis>
</taxonomicName>
<emphasis id="A380EAA6FFBF5B255739F92709F659B9" bold="true" box="[1363,1436,1702,1722]" pageId="15" pageNumber="79">
<taxonomicNameLabel id="B8B357DDFFBF5B255739F92709F659B9" box="[1363,1436,1702,1722]" pageId="15" pageNumber="79" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
by posterior opercular edge with two tips, upper longer or equal to lower (
<emphasis id="A380EAA6FFBF5B25515CF96A0F2259FC" box="[822,840,1771,1791]" italics="true" pageId="15" pageNumber="79">vs</emphasis>
. posterior opercular edge with pointed upper tip and rounded lower edge) and principal caudal rays 10
<emphasis id="A380EAA6FFBF5B2556DCF88E08A85821" bold="true" box="[1206,1218,1807,1826]" pageId="15" pageNumber="79">–</emphasis>
11 (
<emphasis id="A380EAA6FFBF5B255683F88F08915821" box="[1257,1275,1806,1826]" italics="true" pageId="15" pageNumber="79">vs</emphasis>
. principal caudal-fin rays 12
<emphasis id="A380EAA6FFBF5B255119F8B00F155847" bold="true" box="[883,895,1841,1860]" pageId="15" pageNumber="79">–</emphasis>
13).
<taxonomicName id="56F44D37FFBF5B2551CDF8B0084B5846" baseAuthorityName="Nardo" baseAuthorityYear="1827" box="[935,1057,1841,1861]" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="15" pageNumber="79" phylum="Chordata" rank="species" species="pigra">
<emphasis id="A380EAA6FFBF5B2551CDF8B0084B5846" box="[935,1057,1841,1861]" italics="true" pageId="15" pageNumber="79">Gouania pigra</emphasis>
</taxonomicName>
differs from another slender-bodied species,
<taxonomicName id="56F44D37FFBF5B25515AF8D20FC85864" authority="Wagner &amp; Kovac &amp; Koblmüller, 2021" authorityName="Wagner &amp; Kovac &amp; Koblmüller" authorityYear="2021" box="[816,930,1875,1895]" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="15" pageNumber="79" phylum="Chordata" rank="species" species="hofrichteri" status="sp. nov.">
<emphasis id="A380EAA6FFBF5B25515AF8D20FC85864" box="[816,930,1875,1895]" italics="true" pageId="15" pageNumber="79">G. hofrichteri</emphasis>
</taxonomicName>
<emphasis id="A380EAA6FFBF5B2551C2F8D20F9A5864" bold="true" box="[936,1008,1875,1895]" pageId="15" pageNumber="79">
<taxonomicNameLabel id="B8B357DDFFBF5B2551C2F8D20F9A5864" box="[936,1008,1875,1895]" pageId="15" pageNumber="79" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, by posterior opercular edge with two tips, upper longer or equal to lower (
<emphasis id="A380EAA6FFBF5B255647F8F708555889" box="[1069,1087,1910,1930]" italics="true" pageId="15" pageNumber="79">vs</emphasis>
. posterior opercular edge with pointed upper tip and rounded lower edge), longitudinal infralateral and suborbital transversal rows of superficial neuromasts placed in the well-defined deep groove (
<emphasis id="A380EAA6FFA05B3A5320FF300D365FC6" box="[330,348,177,197]" italics="true" pageId="16" pageNumber="80">vs</emphasis>
. longitudinal infralateral and suborbital transversal rows of superficial neuromasts placed in shallow groove disappearing in posterior part of longitudinal infralateral row), body cross-section behind pectoral fin base half oval to pentagonal with straight ventral side (
<emphasis id="A380EAA6FFA05B3A529EFEBA0D6C5E4C" box="[244,262,315,335]" italics="true" pageId="16" pageNumber="80">vs</emphasis>
. body cross-section behind pectoral fin base triangular with ventral flat and dorsal pointed), and the granules on body shallow and inconspicuous (
<emphasis id="A380EAA6FFA05B3A535AFE000D285E96" box="[304,322,385,405]" italics="true" pageId="16" pageNumber="80">vs</emphasis>
. granules on body, at least on posterior part and nape, large and prominent). It is known from the Adriatic Sea and has a nonoverlapping geographic distribution range with
<taxonomicName id="56F44D37FFA05B3A5021FE470ED85ED9" authority="Wagner &amp; Kovac &amp; Koblmüller, 2021" authorityName="Wagner &amp; Kovac &amp; Koblmüller" authorityYear="2021" box="[587,690,454,474]" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="16" pageNumber="80" phylum="Chordata" rank="species" species="orientalis" status="sp. nov.">
<emphasis id="A380EAA6FFA05B3A5021FE470ED85ED9" box="[587,690,454,474]" italics="true" pageId="16" pageNumber="80">G. orientalis</emphasis>
</taxonomicName>
<emphasis id="A380EAA6FFA05B3A50D2FE470F6A5ED9" bold="true" box="[696,768,454,474]" pageId="16" pageNumber="80">
<taxonomicNameLabel id="B8B357DDFFA05B3A50D2FE470F6A5ED9" box="[696,768,454,474]" pageId="16" pageNumber="80" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, and
<taxonomicName id="56F44D37FFA05B3A52CEFE680D725EFE" baseAuthorityName="Risso" baseAuthorityYear="1810" box="[164,280,489,509]" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="16" pageNumber="80" phylum="Chordata" rank="species" species="willdenowi">
<emphasis id="A380EAA6FFA05B3A52CEFE680D725EFE" box="[164,280,489,509]" italics="true" pageId="16" pageNumber="80">G. willdenowi</emphasis>
</taxonomicName>
. Based on the distributional data presented here, differential morphological and genetic characters as well as the taxonomic positioning of the species
<taxonomicName id="56F44D37FFA05B3A5300FDAF0DC45D41" baseAuthorityName="Nardo" baseAuthorityYear="1827" box="[362,430,558,578]" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="16" pageNumber="80" phylum="Chordata" rank="species" species="pigra">
<emphasis id="A380EAA6FFA05B3A5300FDAF0DC45D41" box="[362,430,558,578]" italics="true" pageId="16" pageNumber="80">G. pigra</emphasis>
</taxonomicName>
(see Discussion,
<emphasis id="A380EAA6FFA05B3A5023FDAF0D555D66" italics="true" pageId="16" pageNumber="80">Taxonomical and systematic considerations</emphasis>
below) and the fact that no original types (
<typeStatus id="4E4F8816FFA05B3A50BAFDD00F6B5D66" box="[720,769,593,613]" pageId="16" pageNumber="80" type="holotype">holo-</typeStatus>
,
<typeStatus id="4E4F8816FFA05B3A5216FDF20CB05D84" box="[124,218,627,647]" pageId="16" pageNumber="80" type="lectotype">lectotypes</typeStatus>
) exist we designate a
<typeStatus id="4E4F8816FFA05B3A53D8FDF20D975D84" box="[434,509,627,647]" pageId="16" pageNumber="80" type="neotype">neotype</typeStatus>
for this species. The designated
<typeStatus id="4E4F8816FFA05B3A52DCFD170D6B5DA9" box="[182,257,662,682]" pageId="16" pageNumber="80" type="neotype">neotype</typeStatus>
was collected on the island of Krk, close to the original locus typicus (Rovinj, northern Adriatic Sea) and is accessible at the voucher numbers
<collectionCode id="F7E5AE71FFA05B3A5374FD5A0D215DEC" box="[286,331,731,751]" country="0" httpUri="http://grbio.org/cool/xq4z-q5gq" name="Prirodoslovni muzej Rijeka" pageId="16" pageNumber="80">PMR</collectionCode>
<collectionCode id="F7E5AE71FFA05B3A533BFD5A0D065DEC" box="[337,364,731,751]" pageId="16" pageNumber="80">VP</collectionCode>
3529 at the
<collectionCode id="F7E5AE71FFA05B3A53B0FD5A0E635DEC" box="[474,521,731,751]" country="0" httpUri="http://grbio.org/cool/xq4z-q5gq" name="Prirodoslovni muzej Rijeka" pageId="16" pageNumber="80">PMR</collectionCode>
.
</paragraph>
<paragraph id="914B36B4FFA05B3A52CEFD7F0DA95AC4" blockId="16.[124,773,142,1479]" pageId="16" pageNumber="80">
<emphasis id="A380EAA6FFA05B3A52CEFD7F0D1A5C11" bold="true" box="[164,368,766,786]" pageId="16" pageNumber="80">
<typeStatus id="4E4F8816FFA05B3A52CEFD7F0C9C5C11" box="[164,246,766,786]" pageId="16" pageNumber="80" type="neotype">Neotype</typeStatus>
designation.
</emphasis>
We designated a
<typeStatus id="4E4F8816FFA05B3A5076FD7F0E025C11" box="[540,616,766,786]" pageId="16" pageNumber="80" type="neotype">neotype</typeStatus>
for
<taxonomicName id="56F44D37FFA05B3A50FBFD7F0EB25C11" baseAuthorityName="Nardo" baseAuthorityYear="1827" box="[657,728,766,786]" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="16" pageNumber="80" phylum="Chordata" rank="species" species="pigra">
<emphasis id="A380EAA6FFA05B3A50FBFD7F0EB25C11" box="[657,728,766,786]" italics="true" pageId="16" pageNumber="80">G. pigra</emphasis>
</taxonomicName>
, fulfilling the qualifying conditions (Article 75.3,
<collectionCode id="F7E5AE71FFA05B3A507CFCA00E235C36" box="[534,585,801,821]" pageId="16" pageNumber="80">ICZN</collectionCode>
, 1999). We are positive that no name-bearing type specimens exist for
<taxonomicName id="56F44D37FFA05B3A501FFCC20ED75C54" baseAuthorityName="Nardo" baseAuthorityYear="1827" box="[629,701,835,855]" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="16" pageNumber="80" phylum="Chordata" rank="species" species="pigra">
<emphasis id="A380EAA6FFA05B3A501FFCC20ED75C54" box="[629,701,835,855]" italics="true" pageId="16" pageNumber="80">G. pigra</emphasis>
</taxonomicName>
(
<bibRefCitation id="F5654B45FFA05B3A50A0FCC20C9C5C79" author="Fricke, R. &amp; Eschmeyer, W. N. &amp; Van der Laan, R." pageId="16" pageNumber="80" refId="ref25987" refString="Fricke, R., Eschmeyer, W. N. &amp; Van der Laan, R. (eds) (2020 b). Eschmeyer's catalog of fishes: Genera, species, references. Retrieved from http: // researcharchive. calacademy. org / research / ichthyology / catalog / fishcatmain. asp" type="url" year="2020">
Fricke
<emphasis id="A380EAA6FFA05B3A5216FCE70CCC5C79" box="[124,166,870,890]" italics="true" pageId="16" pageNumber="80">et al</emphasis>
., 2020b
</bibRefCitation>
; Article 75.3.4,
<collectionCode id="F7E5AE71FFA05B3A53E7FCE70DAA5C79" box="[397,448,870,890]" pageId="16" pageNumber="80">ICZN</collectionCode>
, 1999). The genus
<taxonomicName id="56F44D37FFA05B3A5012FCE70EA85C79" authorityName="Nardo" authorityYear="1833" box="[632,706,870,890]" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="16" pageNumber="80" phylum="Chordata" rank="genus">
<emphasis id="A380EAA6FFA05B3A5012FCE70EA85C79" box="[632,706,870,890]" italics="true" pageId="16" pageNumber="80">Gouania</emphasis>
</taxonomicName>
represents a complex zoological problem (Article 75.2,
<collectionCode id="F7E5AE71FFA05B3A502CFC080E135C9E" box="[582,633,905,925]" pageId="16" pageNumber="80">ICZN</collectionCode>
, 1999) of morphologically very similar congenerics. The redescription of
<taxonomicName id="56F44D37FFA05B3A509AFC2A0CB35CE1" baseAuthorityName="Risso" baseAuthorityYear="1810" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="16" pageNumber="80" phylum="Chordata" rank="species" species="willdenowi">
<emphasis id="A380EAA6FFA05B3A509AFC2A0CB35CE1" italics="true" pageId="16" pageNumber="80">G. willdenowi</emphasis>
</taxonomicName>
, resurrection of
<taxonomicName id="56F44D37FFA05B3A531CFC4F0DD65CE1" baseAuthorityName="Nardo" baseAuthorityYear="1827" box="[374,444,974,994]" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="16" pageNumber="80" phylum="Chordata" rank="species" species="pigra">
<emphasis id="A380EAA6FFA05B3A531CFC4F0DD65CE1" box="[374,444,974,994]" italics="true" pageId="16" pageNumber="80">G. pigra</emphasis>
</taxonomicName>
and description of three new species of
<taxonomicName id="56F44D37FFA05B3A52ABFC710D615B07" authorityName="Nardo" authorityYear="1833" box="[193,267,1008,1028]" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="16" pageNumber="80" phylum="Chordata" rank="genus">
<emphasis id="A380EAA6FFA05B3A52ABFC710D615B07" box="[193,267,1008,1028]" italics="true" pageId="16" pageNumber="80">Gouania</emphasis>
</taxonomicName>
in the hitherto monotypic genus thus represented an exceptional need for designation of a
<typeStatus id="4E4F8816FFA05B3A5399FB920E555B24" box="[499,575,1043,1063]" pageId="16" pageNumber="80" type="neotype">neotype</typeStatus>
for
<taxonomicName id="56F44D37FFA05B3A5004FB920EDD5B24" baseAuthorityName="Nardo" baseAuthorityYear="1827" box="[622,695,1043,1063]" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="16" pageNumber="80" phylum="Chordata" rank="species" species="pigra">
<emphasis id="A380EAA6FFA05B3A5004FB920EDD5B24" box="[622,695,1043,1063]" italics="true" pageId="16" pageNumber="80">G. pigra</emphasis>
</taxonomicName>
(Article 75.3,
<collectionCode id="F7E5AE71FFA05B3A52DEFBB70C8D5B49" box="[180,231,1078,1098]" pageId="16" pageNumber="80">ICZN</collectionCode>
, 1999). The situation could become even more complicated if more
<taxonomicName id="56F44D37FFA05B3A5296FBD80D2C5B6E" authorityName="Nardo" authorityYear="1833" box="[252,326,1113,1133]" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="16" pageNumber="80" phylum="Chordata" rank="genus">
<emphasis id="A380EAA6FFA05B3A5296FBD80D2C5B6E" box="[252,326,1113,1133]" italics="true" pageId="16" pageNumber="80">Gouania</emphasis>
</taxonomicName>
linages were to be found around the Mediterranean. In that case, the name-bearing material of the present species,
<typeStatus id="4E4F8816FFA05B3A5216FB1F0CBD5BB1" box="[124,215,1182,1202]" pageId="16" pageNumber="80" type="holotype">holotypes</typeStatus>
and
<typeStatus id="4E4F8816FFA05B3A537AFB1F0D0D5BB1" box="[272,359,1182,1202]" pageId="16" pageNumber="80" type="neotype">neotypes</typeStatus>
, should be available for comparison with potential new material. The diagnostic characters are stated in the species redescription (Article 75.3.1,
<collectionCode id="F7E5AE71FFA05B3A53B2FB620E615BF4" box="[472,523,1251,1271]" pageId="16" pageNumber="80">ICZN</collectionCode>
, 1999), which is sufficient to ensure the recognition of the species (Article 75.3.3,
<collectionCode id="F7E5AE71FFA05B3A50E6FA870ED55A19" box="[652,703,1286,1306]" pageId="16" pageNumber="80">ICZN</collectionCode>
, 1999). The
<typeStatus id="4E4F8816FFA05B3A52DAFAA80C965A3E" box="[176,252,1321,1341]" pageId="16" pageNumber="80" type="neotype">neotype</typeStatus>
fits the original species descriptions of
<taxonomicName id="56F44D37FFA05B3A50DEFAA80F695A3E" baseAuthorityName="Nardo" baseAuthorityYear="1827" box="[692,771,1321,1341]" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="16" pageNumber="80" phylum="Chordata" rank="species" species="pigra">
<emphasis id="A380EAA6FFA05B3A50DEFAA80F695A3E" box="[692,771,1321,1341]" italics="true" pageId="16" pageNumber="80">G. pigra</emphasis>
</taxonomicName>
(
<bibRefCitation id="F5654B45FFA05B3A52EBFACA0D6B5A5C" author="Nardo, G. D." box="[129,257,1355,1375]" pageId="16" pageNumber="80" pagination="102 - 105" refId="ref27986" refString="Nardo, G. D. (1827 a). Estratto da una memoria ittiologica inedita. Giornale di fisica, chimica, storia naturale, medicina ed arti (series 2), 10, 102 - 105." type="journal article" year="1827">Nardo, 1827a</bibRefCitation>
and
<bibRefCitation id="F5654B45FFA05B3A535AFACA0DDE5A5C" author="Nardo, G. D." box="[304,436,1355,1375]" pageId="16" pageNumber="80" pagination="22 - 40" refId="ref28027" refString="Nardo, G. D. (1827 b). Prodromus observationum et disquisitionum Adriaticae ichthyologiae. Giornale di fisica, chimica e storia naturale, medicina ed arti (series 2), 10, 22 - 40." type="journal article" year="1827">Nardo, 1827b</bibRefCitation>
; Article 75.3.5,
<collectionCode id="F7E5AE71FFA05B3A502FFACA0E125A5C" box="[581,632,1355,1375]" pageId="16" pageNumber="80">ICZN</collectionCode>
, 1999) and the
<typeStatus id="4E4F8816FFA05B3A5216FAEF0CA25A81" box="[124,200,1390,1410]" pageId="16" pageNumber="80" type="neotype">neotype</typeStatus>
were collected close to the original type localities (Article 75.3.6,
<collectionCode id="F7E5AE71FFA05B3A52ABFA100C9E5AA6" box="[193,244,1425,1445]" pageId="16" pageNumber="80">ICZN</collectionCode>
, 1999). The
<typeStatus id="4E4F8816FFA05B3A530DFA100DD95AA6" box="[359,435,1425,1445]" pageId="16" pageNumber="80" type="neotype">neotype</typeStatus>
is stored in a scientific museum collection (Article 75.3.7,
<collectionCode id="F7E5AE71FFA05B3A5326FA320D155AC4" box="[332,383,1459,1479]" pageId="16" pageNumber="80">ICZN</collectionCode>
, 1999).
</paragraph>
</subSubSection>
</treatment>
</document>
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