treatments-xml/data/19/5D/87/195D87A2FFBB5B27515AFCE3080A5A5C.xml
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<document id="9AF2CFD74D2D454E6919016D4C07DC83" ID-DOI="10.1111/jfb.14558" ID-ISSN="1095-8649" ID-PMC="PMC7821206" ID-PubMed="32985685" ID-Zenodo-Dep="10928926" IM.bibliography_approvedBy="julia" IM.illustrations_approvedBy="julia" IM.materialsCitations_approvedBy="julia" IM.tables_approvedBy="julia" IM.taxonomicNames_approvedBy="julia" IM.treatmentCitations_approvedBy="julia" IM.treatments_approvedBy="julia" checkinTime="1712261492542" checkinUser="felipe" docAuthor="Wagner, Maximilian, Kovac, Marcelo &amp; Koblmüller, Stephan" docDate="2021" docId="195D87A2FFBB5B27515AFCE3080A5A5C" docLanguage="en" docName="JFishBiol.98.1.64-88.pdf" docOrigin="Journal of Fish Biology 98 (1)" docSource="http://dx.doi.org/10.1111/jfb.14558" docStyle="DocumentStyle:E312FF628AF650F200AD49ACE691BDEB.1:JFishBiol.2018-.journal_article.open" docStyleId="E312FF628AF650F200AD49ACE691BDEB" docStyleName="JFishBiol.2018-.journal_article.open" docStyleVersion="1" docTitle="Gouania hofrichteri Wagner &amp; Kovac &amp; Koblmüller 2021, sp. nov." docType="treatment" docVersion="3" lastPageNumber="77" masterDocId="E564FFDAFFB05B2A526AFF810C6A5F03" masterDocTitle="Unravelling the taxonomy of an interstitial fish radiation: Three new species of Gouania (Teleostei: Gobiesocidae) from the Mediterranean Sea and redescriptions of G. willdenowi and G. pigra" masterLastPageNumber="88" masterPageNumber="64" pageNumber="75" updateTime="1712674921547" updateUser="julia" zenodo-license-document="CC-BY-NC-ND-4.0">
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<mods:title id="2F54A2764206A7F15B11F29DD02FB9AF">Unravelling the taxonomy of an interstitial fish radiation: Three new species of Gouania (Teleostei: Gobiesocidae) from the Mediterranean Sea and redescriptions of G. willdenowi and G. pigra</mods:title>
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<mods:namePart id="F0DA6804EA09D1FF7ABF8227516FB11E">Wagner, Maximilian</mods:namePart>
<mods:affiliation id="E031F7B55A21C1C67CE2D48EFFA7E504">Institute of Biology, University of Graz, Graz, Austria &amp; Department of Biology, University of Antwerp, Antwerp, Belgium</mods:affiliation>
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<mods:namePart id="4F83D5089EC5794F19B47DF167768626">Kovac, Marcelo</mods:namePart>
<mods:affiliation id="BD5CB26B6E69C70F2545CBF3DCBED7AE">Natural History Museum Rijeka, Rijeka, Croatia</mods:affiliation>
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<mods:namePart id="B85EEFC5E81188B6CF0B030B41D313D1">Koblmüller, Stephan</mods:namePart>
<mods:affiliation id="A8AACAAA5EEB8189B0441EDE9B7CACFD">Institute of Biology, University of Graz, Graz, Austria</mods:affiliation>
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<mods:title id="531E8B784CBA184F832DAF5214CC21D8">Journal of Fish Biology</mods:title>
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<mods:date id="E3EFC62B97C0DB2192C1706DFECFBA86">2021</mods:date>
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<mods:number id="E6CCDF6E012E379E2C8378136A68E879">2020-11-03</mods:number>
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<mods:number id="88B537251267A2CC23DB4B4FB82F439A">98</mods:number>
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<mods:identifier id="A3AED08C4A1001152A584564B31401C1" type="ISSN">1095-8649</mods:identifier>
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<treatment id="195D87A2FFBB5B27515AFCE3080A5A5C" LSID="urn:lsid:plazi:treatment:195D87A2FFBB5B27515AFCE3080A5A5C" httpUri="http://treatment.plazi.org/id/195D87A2FFBB5B27515AFCE3080A5A5C" lastPageId="13" lastPageNumber="77" pageId="11" pageNumber="75">
<subSubSection id="D9EE653FFFBB5B21515AFCE308865C78" box="[816,1260,865,891]" pageId="11" pageNumber="75" type="nomenclature">
<paragraph id="914B36B4FFBB5B21515AFCE308865C78" blockId="11.[816,1260,865,891]" box="[816,1260,865,891]" pageId="11" pageNumber="75">
<heading id="CA0381D8FFBB5B21515AFCE308865C78" box="[816,1260,865,891]" fontSize="10" level="3" pageId="11" pageNumber="75" reason="9">
<emphasis id="A380EAA6FFBB5B21515AFCE30F005C78" bold="true" box="[816,874,866,891]" pageId="11" pageNumber="75">3.1.4</emphasis>
|
<emphasis id="A380EAA6FFBB5B2151DBFCE008865C78" bold="true" box="[945,1260,865,891]" pageId="11" pageNumber="75">
<taxonomicName id="56F44D37FFBB5B2151DBFCE008E15C78" authority="Wagner &amp; Kovac &amp; Koblmüller, 2021" authorityName="Wagner &amp; Kovac &amp; Koblmüller" authorityYear="2021" box="[945,1163,865,891]" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="11" pageNumber="75" phylum="Chordata" rank="species" species="hofrichteri" status="sp. nov.">
<emphasis id="A380EAA6FFBB5B2151DBFCE008E15C78" bold="true" box="[945,1163,865,891]" italics="true" pageId="11" pageNumber="75">Gouania hofrichteri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B8B357DDFFBB5B2156F8FCE308865C78" box="[1170,1260,866,891]" pageId="11" pageNumber="75" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="D9EE653FFFBB5B21515AFC2A086E5B06" pageId="11" pageNumber="75" type="description">
<paragraph id="914B36B4FFBB5B21515AFC2A08105CBC" blockId="11.[816,1465,939,1965]" box="[816,1146,939,959]" pageId="11" pageNumber="75">
<emphasis id="A380EAA6FFBB5B21515AFC2A08105CBC" bold="true" box="[816,1146,939,959]" pageId="11" pageNumber="75">English name: Hofrichter's clingfish</emphasis>
</paragraph>
<paragraph id="914B36B4FFBB5B215132FC4F086E5B06" blockId="11.[816,1465,939,1965]" pageId="11" pageNumber="75">
<emphasis id="A380EAA6FFBB5B215132FC4F0F8B5CE1" box="[856,993,974,994]" italics="true" pageId="11" pageNumber="75">
ZooBank
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</emphasis>
:
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urn:lsid:zoobank.org:act:
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</uri>
</paragraph>
</subSubSection>
<subSubSection id="D9EE653FFFBB5B215132FB920FBB5951" pageId="11" pageNumber="75" type="materials_examined">
<paragraph id="914B36B4FFBB5B215132FB9208475B6E" blockId="11.[816,1465,939,1965]" pageId="11" pageNumber="75">
<materialsCitation id="219C3CE9FFBB5B215132FB9208475B6E" collectingDate="2016-08-17" collectionCode="PMR" collectorName="M. Wagner" country="Greece" location="Crete" pageId="11" pageNumber="75" specimenCode="VP4595" specimenCount="1" stateProvince="Kriti" typeStatus="holotype">
<emphasis id="A380EAA6FFBB5B215132FB920FDF5B24" bold="true" box="[856,949,1043,1063]" pageId="11" pageNumber="75">
<typeStatus id="4E4F8816FFBB5B215132FB920FDB5B24" box="[856,945,1043,1063]" pageId="11" pageNumber="75" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectionCode id="F7E5AE71FFBB5B2151D4FB920F865B24" box="[958,1004,1043,1063]" country="0" httpUri="http://grbio.org/cool/xq4z-q5gq" name="Prirodoslovni muzej Rijeka" pageId="11" pageNumber="75">PMR</collectionCode>
<specimenCode id="C1529ECFFFBB5B21519FFB92082E5B24" box="[1013,1092,1043,1063]" collectionCode="VP" pageId="11" pageNumber="75">VP4595</specimenCode>
, male, 30.35 +
<quantity id="560C9B51FFBB5B2156BDFB9209415B24" box="[1239,1323,1043,1063]" metricMagnitude="-3" metricUnit="m" metricValue="3.67" pageId="11" pageNumber="75" unit="mm" value="3.67">3.67 mm</quantity>
, Souda Beach, Plakias,
<collectingRegion id="5330F856FFBB5B215111FBB70FDB5B49" box="[891,945,1078,1098]" country="Greece" name="Kriti" pageId="11" pageNumber="75">Crete</collectingRegion>
,
<collectingCountry id="E9E37624FFBB5B2151D0FBB70F955B49" box="[954,1023,1078,1098]" name="Greece" pageId="11" pageNumber="75">Greece</collectingCountry>
, 35 11
<emphasis id="A380EAA6FFBB5B21562BFBB2082F5B43" bold="true" box="[1089,1093,1075,1088]" italics="true" pageId="11" pageNumber="75">
<superScript id="66819BFCFFBB5B21562BFBB2082F5B43" attach="none" box="[1089,1093,1075,1088]" fontSize="6" pageId="11" pageNumber="75">0</superScript>
</emphasis>
32.1
<emphasis id="A380EAA6FFBB5B21561AFBB208125B43" bold="true" box="[1136,1144,1075,1088]" italics="true" pageId="11" pageNumber="75">
<superScript id="66819BFCFFBB5B21561AFBB208125B43" attach="right" box="[1136,1144,1075,1088]" fontSize="6" pageId="11" pageNumber="75">00</superScript>
</emphasis>
N, 24 22
<emphasis id="A380EAA6FFBB5B2156A4FBB208B85B43" bold="true" box="[1230,1234,1075,1088]" italics="true" pageId="11" pageNumber="75">
<superScript id="66819BFCFFBB5B2156A4FBB208B85B43" attach="none" box="[1230,1234,1075,1088]" fontSize="6" pageId="11" pageNumber="75">0</superScript>
</emphasis>
04.9
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<superScript id="66819BFCFFBB5B215697FBB2096F5B43" attach="right" box="[1277,1285,1075,1088]" fontSize="6" pageId="11" pageNumber="75">00</superScript>
</emphasis>
E, coll.
<collectorName id="3C015362FFBB5B215720FBB709DF5B49" box="[1354,1461,1078,1098]" pageId="11" pageNumber="75">M. Wagner</collectorName>
,
<date id="E54A1074FFBB5B21515AFBD80FA05B6E" box="[816,970,1113,1133]" pageId="11" pageNumber="75" value="2016-08-17">
<collectingDate id="F50EE99CFFBB5B21515AFBD80FA05B6E" box="[816,970,1113,1133]" pageId="11" pageNumber="75" value="2016-08-17">August 17, 2016</collectingDate>
</date>
(Figure 7).
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</paragraph>
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<materialsCitation id="219C3CE9FFBB5B215132FBFA0FE35BD6" collectingDate="2016-08-15" collectionCode="PMR" collectorName="M. Wagner" country="Greece" location="Mades" pageId="11" pageNumber="75" specimenCode="VP4591" specimenCount="1" stateProvince="Kriti" typeStatus="paratype">
<emphasis id="A380EAA6FFBB5B215132FBFA0FDF5B8C" bold="true" box="[856,949,1147,1167]" pageId="11" pageNumber="75">
<typeStatus id="4E4F8816FFBB5B215132FBFA0FDF5B8C" box="[856,949,1147,1167]" pageId="11" pageNumber="75" type="paratype">Paratypes</typeStatus>
</emphasis>
.
<collectionCode id="F7E5AE71FFBB5B2151A1FBFA0F935B8C" box="[971,1017,1147,1167]" country="0" httpUri="http://grbio.org/cool/xq4z-q5gq" name="Prirodoslovni muzej Rijeka" pageId="11" pageNumber="75">PMR</collectionCode>
<specimenCode id="C1529ECFFFBB5B215660FBFA08325B8C" box="[1034,1112,1147,1167]" collectionCode="VP" pageId="11" pageNumber="75">VP4591</specimenCode>
, female, 21.42 +
<quantity id="560C9B51FFBB5B215767FBFA090B5B8C" box="[1293,1377,1147,1167]" metricMagnitude="-3" metricUnit="m" metricValue="2.62" pageId="11" pageNumber="75" unit="mm" value="2.62">2.62 mm</quantity>
,
<location id="942B606FFFBB5B21571FFBFA09DC5B8C" LSID="urn:lsid:plazi:treatment:195D87A2FFBB5B27515AFCE3080A5A5C:942B606FFFBB5B21571FFBFA09DC5B8C" box="[1397,1462,1147,1167]" country="Greece" name="Mades" pageId="11" pageNumber="75" stateProvince="Kriti">Mades</location>
,
<collectingRegion id="5330F856FFBB5B21515AFB1F0F0C5BB1" box="[816,870,1182,1202]" country="Greece" name="Kriti" pageId="11" pageNumber="75">Crete</collectingRegion>
,
<collectingCountry id="E9E37624FFBB5B215105FB1F0FDE5BB1" box="[879,948,1182,1202]" name="Greece" pageId="11" pageNumber="75">Greece</collectingCountry>
, 35 24
<emphasis id="A380EAA6FFBB5B21519DFB1A0F915BAB" bold="true" box="[1015,1019,1179,1192]" italics="true" pageId="11" pageNumber="75">
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</emphasis>
01.1
<emphasis id="A380EAA6FFBB5B21564CFB1A08445BAB" bold="true" box="[1062,1070,1179,1192]" italics="true" pageId="11" pageNumber="75">
<superScript id="66819BFCFFBB5B21564CFB1A08445BAB" attach="right" box="[1062,1070,1179,1192]" fontSize="6" pageId="11" pageNumber="75">00</superScript>
</emphasis>
N, 25 02
<emphasis id="A380EAA6FFBB5B2156EFFB1A08E35BAB" bold="true" box="[1157,1161,1179,1192]" italics="true" pageId="11" pageNumber="75">
<superScript id="66819BFCFFBB5B2156EFFB1A08E35BAB" attach="none" box="[1157,1161,1179,1192]" fontSize="6" pageId="11" pageNumber="75">0</superScript>
</emphasis>
01.6
<emphasis id="A380EAA6FFBB5B2156DEFB1A08D65BAB" bold="true" box="[1204,1212,1179,1192]" italics="true" pageId="11" pageNumber="75">
<superScript id="66819BFCFFBB5B2156DEFB1A08D65BAB" attach="right" box="[1204,1212,1179,1192]" fontSize="6" pageId="11" pageNumber="75">00</superScript>
</emphasis>
E, coll.
<collectorName id="3C015362FFBB5B21576BFB1F09075BB1" box="[1281,1389,1182,1202]" pageId="11" pageNumber="75">M. Wagner</collectorName>
,
<date id="E54A1074FFBB5B21571CFB1F0FE35BD6" pageId="11" pageNumber="75" value="2016-08-15">
<collectingDate id="F50EE99CFFBB5B21571CFB1F0FE35BD6" pageId="11" pageNumber="75" value="2016-08-15">August 15, 2016</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="219C3CE9FFBB5B2151F9FB4009405BD6" box="[915,1322,1217,1237]" collectionCode="PMR" country="Greece" pageId="11" pageNumber="75" specimenCode="VP4599" specimenCount="1" stateProvince="Kriti" typeStatus="paratype">
<collectionCode id="F7E5AE71FFBB5B2151F9FB400FAB5BD6" box="[915,961,1217,1237]" country="0" httpUri="http://grbio.org/cool/xq4z-q5gq" name="Prirodoslovni muzej Rijeka" pageId="11" pageNumber="75">PMR</collectionCode>
<specimenCode id="C1529ECFFFBB5B2151A3FB4008725BD6" box="[969,1048,1217,1237]" collectionCode="VP" pageId="11" pageNumber="75">VP4599</specimenCode>
, female, 26.8 +
<quantity id="560C9B51FFBB5B2156C4FB40096A5BD6" box="[1198,1280,1217,1237]" metricMagnitude="-3" metricUnit="m" metricValue="3.23" pageId="11" pageNumber="75" unit="mm" value="3.23">3.23 mm</quantity>
and
</materialsCitation>
<materialsCitation id="219C3CE9FFBB5B21575BFB400F0E5A3E" collectingDate="2016-08-17" collectionCode="PMR" collectorName="M. Wagner" country="Greece" location="Souda Beach" municipality="Plakias" pageId="11" pageNumber="75" specimenCode="VP4600" specimenCount="1" stateProvince="Kriti" typeStatus="paratype">
<collectionCode id="F7E5AE71FFBB5B21575BFB4009355BD6" box="[1329,1375,1217,1237]" country="0" httpUri="http://grbio.org/cool/xq4z-q5gq" name="Prirodoslovni muzej Rijeka" pageId="11" pageNumber="75">PMR</collectionCode>
<specimenCode id="C1529ECFFFBB5B21570DFB4009DF5BD6" box="[1383,1461,1217,1237]" collectionCode="VP" pageId="11" pageNumber="75">VP4600</specimenCode>
, male, 29.75 +
<quantity id="560C9B51FFBB5B2151D7FB62087B5BF4" box="[957,1041,1251,1271]" metricMagnitude="-3" metricUnit="m" metricValue="3.49" pageId="11" pageNumber="75" unit="mm" value="3.49">3.49 mm</quantity>
, both from
<location id="942B606FFFBB5B2156F2FB6209715BF4" LSID="urn:lsid:plazi:treatment:195D87A2FFBB5B27515AFCE3080A5A5C:942B606FFFBB5B2156F2FB6209715BF4" box="[1176,1307,1251,1271]" country="Greece" municipality="Plakias" name="Souda Beach" pageId="11" pageNumber="75" stateProvince="Kriti">Souda Beach</location>
,
<collectingMunicipality id="712FACCEFFBB5B215747FB6209055BF4" box="[1325,1391,1251,1271]" pageId="11" pageNumber="75">Plakias</collectingMunicipality>
,
<collectingRegion id="5330F856FFBB5B215715FB6209DF5BF4" box="[1407,1461,1251,1271]" country="Greece" name="Kriti" pageId="11" pageNumber="75">Crete</collectingRegion>
,
<collectingCountry id="E9E37624FFBB5B21515AFA870F1F5A19" box="[816,885,1286,1306]" name="Greece" pageId="11" pageNumber="75">Greece</collectingCountry>
, 35 11
<emphasis id="A380EAA6FFBB5B2151D1FA820FD55A13" bold="true" box="[955,959,1283,1296]" italics="true" pageId="11" pageNumber="75">
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32.1
<emphasis id="A380EAA6FFBB5B215180FA820F985A13" bold="true" box="[1002,1010,1283,1296]" italics="true" pageId="11" pageNumber="75">
<superScript id="66819BFCFFBB5B215180FA820F985A13" attach="right" box="[1002,1010,1283,1296]" fontSize="6" pageId="11" pageNumber="75">00</superScript>
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N, 24 22
<emphasis id="A380EAA6FFBB5B215627FA82083B5A13" bold="true" box="[1101,1105,1283,1296]" italics="true" pageId="11" pageNumber="75">
<superScript id="66819BFCFFBB5B215627FA82083B5A13" attach="none" box="[1101,1105,1283,1296]" fontSize="6" pageId="11" pageNumber="75">0</superScript>
</emphasis>
04.9
<emphasis id="A380EAA6FFBB5B215616FA8208EE5A13" bold="true" box="[1148,1156,1283,1296]" italics="true" pageId="11" pageNumber="75">
<superScript id="66819BFCFFBB5B215616FA8208EE5A13" attach="right" box="[1148,1156,1283,1296]" fontSize="6" pageId="11" pageNumber="75">00</superScript>
</emphasis>
E, coll.
<collectorName id="3C015362FFBB5B2156BBFA87092B5A19" box="[1233,1345,1286,1306]" pageId="11" pageNumber="75">M. Wagner</collectorName>
,
<date id="E54A1074FFBB5B215724FA870F0E5A3E" pageId="11" pageNumber="75" value="2016-08-17">
<collectingDate id="F50EE99CFFBB5B215724FA870F0E5A3E" pageId="11" pageNumber="75" value="2016-08-17">August 17, 2016</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="219C3CE9FFBB5B21511AFAA80FA35A81" collectingDate="2016-08-02" collectionCode="PMR" collectorName="M. Wagner" country="Greece" location="Saronida" pageId="11" pageNumber="75" specimenCode="VP4605" specimenCount="1" typeStatus="paratype">
<collectionCode id="F7E5AE71FFBB5B21511AFAA80FF45A3E" box="[880,926,1321,1341]" country="0" httpUri="http://grbio.org/cool/xq4z-q5gq" name="Prirodoslovni muzej Rijeka" pageId="11" pageNumber="75">PMR</collectionCode>
<specimenCode id="C1529ECFFFBB5B2151CDFAA80F9F5A3E" box="[935,1013,1321,1341]" collectionCode="VP" pageId="11" pageNumber="75">VP4605</specimenCode>
, juvenile of unidentified sex, 20.62 +
<quantity id="560C9B51FFBB5B21570BFAA809DF5A3E" box="[1377,1461,1321,1341]" metricMagnitude="-3" metricUnit="m" metricValue="2.63" pageId="11" pageNumber="75" unit="mm" value="2.63">2.63 mm</quantity>
,
<location id="942B606FFFBB5B21515AFACA0FE95A5C" LSID="urn:lsid:plazi:treatment:195D87A2FFBB5B27515AFCE3080A5A5C:942B606FFFBB5B21515AFACA0FE95A5C" box="[816,899,1355,1375]" country="Greece" name="Saronida" pageId="11" pageNumber="75">Saronida</location>
,
<collectingCountry id="E9E37624FFBB5B2151FEFACA0FB35A5C" box="[916,985,1355,1375]" name="Greece" pageId="11" pageNumber="75">Greece</collectingCountry>
, 37 43
<emphasis id="A380EAA6FFBB5B215648FAC8084C5A55" bold="true" box="[1058,1062,1353,1366]" italics="true" pageId="11" pageNumber="75">
<superScript id="66819BFCFFBB5B215648FAC8084C5A55" attach="none" box="[1058,1062,1353,1366]" fontSize="6" pageId="11" pageNumber="75">0</superScript>
</emphasis>
12.2
<emphasis id="A380EAA6FFBB5B21563BFAC808335A55" bold="true" box="[1105,1113,1353,1366]" italics="true" pageId="11" pageNumber="75">
<superScript id="66819BFCFFBB5B21563BFAC808335A55" attach="right" box="[1105,1113,1353,1366]" fontSize="6" pageId="11" pageNumber="75">00</superScript>
</emphasis>
N, 23 55
<emphasis id="A380EAA6FFBB5B2156DDFAC808D15A55" bold="true" box="[1207,1211,1353,1366]" italics="true" pageId="11" pageNumber="75">
<superScript id="66819BFCFFBB5B2156DDFAC808D15A55" attach="none" box="[1207,1211,1353,1366]" fontSize="6" pageId="11" pageNumber="75">0</superScript>
</emphasis>
41.0
<emphasis id="A380EAA6FFBB5B21568CFAC808845A55" bold="true" box="[1254,1262,1353,1366]" italics="true" pageId="11" pageNumber="75">
<superScript id="66819BFCFFBB5B21568CFAC808845A55" attach="right" box="[1254,1262,1353,1366]" fontSize="6" pageId="11" pageNumber="75">00</superScript>
</emphasis>
E, coll.
<collectorName id="3C015362FFBB5B215728FACA09DF5A5C" box="[1346,1461,1355,1375]" pageId="11" pageNumber="75">M. Wagner</collectorName>
,
<date id="E54A1074FFBB5B21515AFAEF0FA35A81" box="[816,969,1390,1410]" pageId="11" pageNumber="75" value="2016-08-02">
<collectingDate id="F50EE99CFFBB5B21515AFAEF0FA35A81" box="[816,969,1390,1410]" pageId="11" pageNumber="75" value="2016-08-02">August 2, 2016</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="219C3CE9FFBB5B2151BEFAEF08F15AA6" collectionCode="PMR" country="Greece" pageId="11" pageNumber="75" specimenCode="VP4606" specimenCount="1" typeStatus="paratype">
<collectionCode id="F7E5AE71FFBB5B2151BEFAEF08685A81" box="[980,1026,1390,1410]" country="0" httpUri="http://grbio.org/cool/xq4z-q5gq" name="Prirodoslovni muzej Rijeka" pageId="11" pageNumber="75">PMR</collectionCode>
<specimenCode id="C1529ECFFFBB5B215661FAEF08305A81" box="[1035,1114,1390,1410]" collectionCode="VP" pageId="11" pageNumber="75">VP4606</specimenCode>
, female, 29.47 +
<quantity id="560C9B51FFBB5B215694FAEF09395A81" box="[1278,1363,1390,1410]" metricMagnitude="-3" metricUnit="m" metricValue="3.41" pageId="11" pageNumber="75" unit="mm" value="3.41">3.41 mm</quantity>
, ZSM-PIS-047656, female, 35.86 +
<quantity id="560C9B51FFBB5B215674FA10081B5AA6" box="[1054,1137,1425,1445]" metricMagnitude="-3" metricUnit="m" metricValue="4.49" pageId="11" pageNumber="75" unit="mm" value="4.49">4.49 mm</quantity>
and
</materialsCitation>
<materialsCitation id="219C3CE9FFBB5B2156C9FA1008145AE9" collectingDate="2016-08-04" collectionCode="PMR" collectorName="M. Wagner" country="Greece" location="Chamolia" pageId="11" pageNumber="75" specimenCode="VP4608" specimenCount="1" typeStatus="paratype">
<collectionCode id="F7E5AE71FFBB5B2156C9FA1008BB5AA6" box="[1187,1233,1425,1445]" country="0" httpUri="http://grbio.org/cool/xq4z-q5gq" name="Prirodoslovni muzej Rijeka" pageId="11" pageNumber="75">PMR</collectionCode>
<specimenCode id="C1529ECFFFBB5B2156B3FA10094D5AA6" box="[1241,1319,1425,1445]" collectionCode="VP" pageId="11" pageNumber="75">VP4608</specimenCode>
, female, 26.69 +
<quantity id="560C9B51FFBB5B215128FA320FFC5AC4" box="[834,918,1459,1479]" metricMagnitude="-3" metricUnit="m" metricValue="3.32" pageId="11" pageNumber="75" unit="mm" value="3.32">3.32 mm</quantity>
, all from
<location id="942B606FFFBB5B215199FA3208215AC4" LSID="urn:lsid:plazi:treatment:195D87A2FFBB5B27515AFCE3080A5A5C:942B606FFFBB5B215199FA3208215AC4" box="[1011,1099,1459,1479]" country="Greece" name="Chamolia" pageId="11" pageNumber="75">Chamolia</location>
,
<collectingCountry id="E9E37624FFBB5B21563CFA3208F15AC4" box="[1110,1179,1459,1479]" name="Greece" pageId="11" pageNumber="75">Greece</collectingCountry>
, 37 54
<emphasis id="A380EAA6FFBB5B2156B5FA3008895ABD" bold="true" box="[1247,1251,1457,1470]" italics="true" pageId="11" pageNumber="75">
<superScript id="66819BFCFFBB5B2156B5FA3008895ABD" attach="none" box="[1247,1251,1457,1470]" fontSize="6" pageId="11" pageNumber="75">0</superScript>
</emphasis>
58.5
<emphasis id="A380EAA6FFBB5B215764FA30097C5ABD" bold="true" box="[1294,1302,1457,1470]" italics="true" pageId="11" pageNumber="75">
<superScript id="66819BFCFFBB5B215764FA30097C5ABD" attach="right" box="[1294,1302,1457,1470]" fontSize="6" pageId="11" pageNumber="75">00</superScript>
</emphasis>
N, 24 02
<emphasis id="A380EAA6FFBB5B215704FA3009185ABD" bold="true" box="[1390,1394,1457,1470]" italics="true" pageId="11" pageNumber="75">
<superScript id="66819BFCFFBB5B215704FA3009185ABD" attach="none" box="[1390,1394,1457,1470]" fontSize="6" pageId="11" pageNumber="75">0</superScript>
</emphasis>
08.7
<emphasis id="A380EAA6FFBB5B2157F7FA3009CF5ABD" bold="true" box="[1437,1445,1457,1470]" italics="true" pageId="11" pageNumber="75">
<superScript id="66819BFCFFBB5B2157F7FA3009CF5ABD" attach="right" box="[1437,1445,1457,1470]" fontSize="6" pageId="11" pageNumber="75">00</superScript>
</emphasis>
E, coll.
<collectorName id="3C015362FFBB5B21510BFA570FBB5AE9" box="[865,977,1494,1514]" pageId="11" pageNumber="75">M. Wagner</collectorName>
,
<date id="E54A1074FFBB5B21518AFA5708145AE9" box="[992,1150,1494,1514]" pageId="11" pageNumber="75" value="2016-08-04">
<collectingDate id="F50EE99CFFBB5B21518AFA5708145AE9" box="[992,1150,1494,1514]" pageId="11" pageNumber="75" value="2016-08-04">August 4, 2016</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="219C3CE9FFBB5B2156E6FA570FBB5951" collectingDate="2018-08-16" collectorName="M. Wagner" country="Greece" location="Chamolia" pageId="11" pageNumber="75" specimenCount="1" typeStatus="paratype">
ZSM-PIS-047657, male, 36.91 +
<quantity id="560C9B51FFBB5B215128FA780FFE590E" box="[834,916,1529,1549]" metricMagnitude="-3" metricUnit="m" metricValue="4.24" pageId="11" pageNumber="75" unit="mm" value="4.24">4.24 mm</quantity>
and ZSM-PIS-047657, male, 33.27 + 4.31, mm, both from
<location id="942B606FFFBB5B21515AF99A0FE2592C" LSID="urn:lsid:plazi:treatment:195D87A2FFBB5B27515AFCE3080A5A5C:942B606FFFBB5B21515AF99A0FE2592C" box="[816,904,1563,1583]" country="Greece" name="Chamolia" pageId="11" pageNumber="75">Chamolia</location>
,
<collectingCountry id="E9E37624FFBB5B2151F2F99A0FB7592C" box="[920,989,1563,1583]" name="Greece" pageId="11" pageNumber="75">Greece</collectingCountry>
, 37 54
<emphasis id="A380EAA6FFBB5B21564CF99808405925" bold="true" box="[1062,1066,1561,1574]" italics="true" pageId="11" pageNumber="75">
<superScript id="66819BFCFFBB5B21564CF99808405925" attach="none" box="[1062,1066,1561,1574]" fontSize="6" pageId="11" pageNumber="75">0</superScript>
</emphasis>
58.5
<emphasis id="A380EAA6FFBB5B21563FF99808375925" bold="true" box="[1109,1117,1561,1574]" italics="true" pageId="11" pageNumber="75">
<superScript id="66819BFCFFBB5B21563FF99808375925" attach="right" box="[1109,1117,1561,1574]" fontSize="6" pageId="11" pageNumber="75">00</superScript>
</emphasis>
N, 24 02
<emphasis id="A380EAA6FFBB5B2156D0F99808D45925" bold="true" box="[1210,1214,1561,1574]" italics="true" pageId="11" pageNumber="75">
<superScript id="66819BFCFFBB5B2156D0F99808D45925" attach="none" box="[1210,1214,1561,1574]" fontSize="6" pageId="11" pageNumber="75">0</superScript>
</emphasis>
08.7
<emphasis id="A380EAA6FFBB5B215683F998089B5925" bold="true" box="[1257,1265,1561,1574]" italics="true" pageId="11" pageNumber="75">
<superScript id="66819BFCFFBB5B215683F998089B5925" attach="right" box="[1257,1265,1561,1574]" fontSize="6" pageId="11" pageNumber="75">00</superScript>
</emphasis>
E, coll.
<collectorName id="3C015362FFBB5B215729F99A09DF592C" box="[1347,1461,1563,1583]" pageId="11" pageNumber="75">M. Wagner</collectorName>
,
<date id="E54A1074FFBB5B21515AF9BF0FA65951" box="[816,972,1598,1618]" pageId="11" pageNumber="75" value="2018-08-16">
<collectingDate id="F50EE99CFFBB5B21515AF9BF0FA65951" box="[816,972,1598,1618]" pageId="11" pageNumber="75" value="2018-08-16">August 16, 2018</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="D9EE653FFFBB5B265132F9E10E815A3E" lastPageId="12" lastPageNumber="76" pageId="11" pageNumber="75" type="diagnosis">
<paragraph id="914B36B4FFBB5B265132F9E10E815A3E" blockId="11.[816,1465,939,1965]" lastBlockId="12.[124,773,905,1965]" lastPageId="12" lastPageNumber="76" pageId="11" pageNumber="75">
<emphasis id="A380EAA6FFBB5B215132F9E10FD35977" bold="true" box="[856,953,1632,1652]" pageId="11" pageNumber="75">Diagnosis.</emphasis>
<taxonomicName id="56F44D37FFBB5B2151A9F9E008195976" authority="Wagner &amp; Kovac &amp; Koblmüller, 2021" authorityName="Wagner &amp; Kovac &amp; Koblmüller" authorityYear="2021" box="[963,1139,1633,1653]" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="11" pageNumber="75" phylum="Chordata" rank="species" species="hofrichteri" status="sp. nov.">
<emphasis id="A380EAA6FFBB5B2151A9F9E008195976" box="[963,1139,1633,1653]" italics="true" pageId="11" pageNumber="75">Gouania hofrichteri</emphasis>
</taxonomicName>
<emphasis id="A380EAA6FFBB5B215617F9E108A15977" bold="true" box="[1149,1227,1632,1652]" pageId="11" pageNumber="75">
<taxonomicNameLabel id="B8B357DDFFBB5B215617F9E108A15977" box="[1149,1227,1632,1652]" pageId="11" pageNumber="75" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from congeneric species by the combination of the following characters: (1) dorsal head profile in lateral view
<emphasis id="A380EAA6FFBB5B215669F926086159B9" bold="true" box="[1027,1035,1703,1722]" pageId="11" pageNumber="75"></emphasis>
S
<emphasis id="A380EAA6FFBB5B21567CF926087459B9" bold="true" box="[1046,1054,1703,1722]" pageId="11" pageNumber="75"></emphasis>
curved, concave above eye and convex at nape; (2) posterior angle of jaws extends to between vertical line drawn through posterior edge of anterior nostril and vertical line drawn through anterior edge of eye; (3) infraorbital invagination below anterior half of eye or below mideye; (4) posterior opercular edge with pointed upper tip and rounded lower edge; (5) longitudinal infralateral and suborbital transversal rows of superficial neuromasts placed in shallow groove disappearing in posterior part of longitudinal infralateral row; (6) body cross-section behind pectoral fin base triangular with ventral flat and dorsal pointed; (7) granules on body, at least on posterior part and nape, large and prominent; (8) upper attachment of gill membrane opposite to 3rd
<emphasis id="A380EAA6FFBC5B26531EFC700DEA5B07" bold="true" box="[372,384,1009,1028]" pageId="12" pageNumber="76"></emphasis>
4th pectoral ray; (9) vertical eye diameter 2.0
<emphasis id="A380EAA6FFBC5B2652F2FB950CCE5B24" bold="true" box="[152,164,1044,1063]" pageId="12" pageNumber="76"></emphasis>
2.7% of standard length; (10) head length 18.9
<emphasis id="A380EAA6FFBC5B26503AFB950E365B24" bold="true" box="[592,604,1044,1063]" pageId="12" pageNumber="76"></emphasis>
23.4% of standard length; (11) pectoral-fin length 5.4
<emphasis id="A380EAA6FFBC5B2653D3FBB60DAF5B49" bold="true" box="[441,453,1079,1098]" pageId="12" pageNumber="76"></emphasis>
6.4% of standard length; (12) prepectoral distance 19.5
<emphasis id="A380EAA6FFBC5B26532FFBD80D3B5B6F" bold="true" box="[325,337,1113,1132]" pageId="12" pageNumber="76"></emphasis>
23.7% of standard length; (13) ventral adhesive disc length 10.2
<emphasis id="A380EAA6FFBC5B265371FBFD0D4D5B8C" bold="true" box="[283,295,1148,1167]" pageId="12" pageNumber="76"></emphasis>
13.4% of standard length; (14) caudal-fin length 11.5
<emphasis id="A380EAA6FFBC5B2652C9FB1E0CC55BB1" bold="true" box="[163,175,1183,1202]" pageId="12" pageNumber="76"></emphasis>
13.4% of standard length; (15) high total number of vertebrae (Supporting Information Table S2; 38
<emphasis id="A380EAA6FFBC5B2653B3FB400D8F5BD7" bold="true" box="[473,485,1217,1236]" pageId="12" pageNumber="76"></emphasis>
40); (16) pharyngeal jaws with ceratobranchial 5 elongated, having several larger elongated conical teeth; (17) nasal bones hook-shaped; (18) star-like pigmentation around eyes absent, body pigmentation striped or marbled, small iridophores visible.
</paragraph>
</subSubSection>
<caption id="C58B663CFFBC5B265216FD300CAE5C39" ID-DOI="http://doi.org/10.5281/zenodo.10928938" ID-Zenodo-Dep="10928938" httpUri="https://zenodo.org/record/10928938/files/figure.png" pageId="12" pageNumber="76" startId="12.[124,136,689,708]" targetBox="[130,769,142,646]" targetPageId="12" targetType="figure">
<paragraph id="914B36B4FFBC5B265216FD300CAE5C39" blockId="12.[124,760,688,826]" pageId="12" pageNumber="76">
<emphasis id="A380EAA6FFBC5B265216FD300C865DC7" bold="true" box="[124,236,689,708]" pageId="12" pageNumber="76">F I G U R E 7</emphasis>
<taxonomicName id="56F44D37FFBC5B265360FD300DDE5DC6" authority="Wagner &amp; Kovac &amp; Koblmüller, 2021" authorityName="Wagner &amp; Kovac &amp; Koblmüller" authorityYear="2021" box="[266,436,689,709]" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="12" pageNumber="76" phylum="Chordata" rank="species" species="hofrichteri" status="sp. nov.">
<emphasis id="A380EAA6FFBC5B265360FD300DDE5DC6" box="[266,436,689,709]" italics="true" pageId="12" pageNumber="76">Gouania hofrichteri</emphasis>
</taxonomicName>
<emphasis id="A380EAA6FFBC5B2653D3FD310E685DC7" bold="true" box="[441,514,688,708]" pageId="12" pageNumber="76">
<taxonomicNameLabel id="B8B357DDFFBC5B2653D3FD310E685DC7" box="[441,514,688,708]" pageId="12" pageNumber="76" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, PMR VP4595, holotype, male, 30.35+3.67 mm, Souda Beach, Plakias, Crete, Greece. Lateral view of specimen preserved in 4% formaldehyde (top). Lateral, dorsal and ventral view, alive (below). Photographs by M. Wagner and M. Kovacˇic
</paragraph>
</caption>
<subSubSection id="D9EE653FFFBC5B2752CEFACA0DE45B8C" lastPageId="13" lastPageNumber="77" pageId="12" pageNumber="76" type="description">
<paragraph id="914B36B4FFBC5B2652CEFACA097F5A81" blockId="12.[124,773,905,1965]" lastBlockId="12.[821,1470,142,1965]" pageId="12" pageNumber="76">
<emphasis id="A380EAA6FFBC5B2652CEFACA0D7D5A5C" bold="true" box="[164,279,1355,1375]" pageId="12" pageNumber="76">Description.</emphasis>
<emphasis id="A380EAA6FFBC5B265375FACA0DBE5A5C" box="[287,468,1355,1375]" italics="true" pageId="12" pageNumber="76">General morphology</emphasis>
: Body proportions are given in Table 1. Body very slender and elongated, posteriorly laterally compressed, body depth at pectoral fins
<quantity id="560C9B51FFBC5B26538BFA100E245AA6" box="[481,590,1425,1445]" metricMagnitude="-1" metricUnit="m" metricValue="2.5019" metricValueMax="2.667" metricValueMin="2.3367999999999998" pageId="12" pageNumber="76" unit="in" value="9.85" valueMax="10.5" valueMin="9.2">
9.2
<emphasis id="A380EAA6FFBC5B265394FA100E605AA7" bold="true" box="[510,522,1425,1444]" pageId="12" pageNumber="76"></emphasis>
10.5 in
</quantity>
SL, body depth at anus 11.0
<emphasis id="A380EAA6FFBC5B2652B3FA350C8F5AC4" bold="true" box="[217,229,1460,1479]" pageId="12" pageNumber="76"></emphasis>
<quantity id="560C9B51FFBC5B26528FFA320D425AC4" box="[229,296,1459,1479]" metricMagnitude="-1" metricUnit="m" metricValue="3.429" pageId="12" pageNumber="76" unit="in" value="13.5">13.5 in</quantity>
SL, body depth in width at pectoral fins 1.0
<emphasis id="A380EAA6FFBC5B2650BFFA350E8B5AC4" bold="true" box="[725,737,1460,1479]" pageId="12" pageNumber="76"></emphasis>
1.0, body depth in width at anus 0.7
<emphasis id="A380EAA6FFBC5B2653C2FA560DDE5AE9" bold="true" box="[424,436,1495,1514]" pageId="12" pageNumber="76"></emphasis>
0.9. Body cross-section behind pectoral fin base triangular with ventral flat and dorsal pointed. Granules on body, at least on posterior part and nape, large and prominent. Head dorsoventrally compressed, head depth in width at orbit 1.5
<emphasis id="A380EAA6FFBC5B2652F3F9E00CCF5977" bold="true" box="[153,165,1633,1652]" pageId="12" pageNumber="76"></emphasis>
1.8, and moderately small, head length
<quantity id="560C9B51FFBC5B26507EF9E00E195976" box="[532,627,1633,1653]" metricMagnitude="-1" metricUnit="m" metricValue="1.2192" metricValueMax="1.3462" metricValueMin="1.0921999999999998" pageId="12" pageNumber="76" unit="in" value="4.8" valueMax="5.3" valueMin="4.3">
4.3
<emphasis id="A380EAA6FFBC5B265058F9E00E545977" bold="true" box="[562,574,1633,1652]" pageId="12" pageNumber="76"></emphasis>
5.3 in
</quantity>
SL, head wider than body width maximum, head width at anterior sucking disc edge
<quantity id="560C9B51FFBC5B265216F9270CB059B9" box="[124,218,1702,1722]" metricMagnitude="-2" metricUnit="m" metricValue="1.905" metricValueMax="2.032" metricValueMin="1.7779999999999998" pageId="12" pageNumber="76" unit="in" value="0.75" valueMax="0.8" valueMin="0.7">
0.7
<emphasis id="A380EAA6FFBC5B2652F3F9260CCF59B9" bold="true" box="[153,165,1703,1722]" pageId="12" pageNumber="76"></emphasis>
0.8 in
</quantity>
body width at pectoral fins. Dorsal head profile
<emphasis id="A380EAA6FFBC5B2650F4F9260ECC59B9" bold="true" box="[670,678,1703,1722]" pageId="12" pageNumber="76"></emphasis>
S
<emphasis id="A380EAA6FFBC5B2650D8F9260ED059B9" bold="true" box="[690,698,1703,1722]" pageId="12" pageNumber="76"></emphasis>
curved, concave above eye and convex at nape. Head rounded in dorsal view. Snout large compared to eyes, preorbital distance
<quantity id="560C9B51FFBC5B265000F96A0EA659FC" box="[618,716,1771,1791]" metricMagnitude="-2" metricUnit="m" metricValue="8.255" metricValueMax="9.398" metricValueMin="7.111999999999999" pageId="12" pageNumber="76" unit="in" value="3.25" valueMax="3.7" valueMin="2.8">
2.8
<emphasis id="A380EAA6FFBC5B2650E2F96D0EFE59FC" bold="true" box="[648,660,1772,1791]" pageId="12" pageNumber="76"></emphasis>
3.7 in
</quantity>
head length,
<quantity id="560C9B51FFBC5B2652ABF88F0D755821" box="[193,287,1806,1826]" metricMagnitude="-2" metricUnit="m" metricValue="1.016" metricValueMax="1.27" metricValueMin="0.762" pageId="12" pageNumber="76" unit="in" value="0.4" valueMax="0.5" valueMin="0.3">
0.3
<emphasis id="A380EAA6FFBC5B2652B4F88E0C805821" bold="true" box="[222,234,1807,1826]" pageId="12" pageNumber="76"></emphasis>
0.5 in
</quantity>
horizontal eye diameter. Snout wide, not produced, blunt. Internostril space almost triangular in cross-section, with ridge top, conspicuously convex. Eyes dorsolateral, with lower eye edge rounded. Eyes small,
<quantity id="560C9B51FFBC5B26532AF8F70DC15889" box="[320,427,1910,1930]" metricMagnitude="-1" metricUnit="m" metricValue="2.3367999999999998" metricValueMax="2.7177999999999995" metricValueMin="1.9558" pageId="12" pageNumber="76" unit="in" value="9.2" valueMax="10.7" valueMin="7.7">
7.7
<emphasis id="A380EAA6FFBC5B265337F8F60D035889" bold="true" box="[349,361,1911,1930]" pageId="12" pageNumber="76"></emphasis>
10.7 in
</quantity>
head length, vertical diameter of the eye
<quantity id="560C9B51FFBC5B2652C9F8180D6A58AE" box="[163,256,1945,1965]" metricMagnitude="-2" metricUnit="m" metricValue="2.413" metricValueMax="2.794" metricValueMin="2.032" pageId="12" pageNumber="76" unit="in" value="0.95" valueMax="1.1" valueMin="0.8">
0.8
<emphasis id="A380EAA6FFBC5B2652AAF8180CA658AF" bold="true" box="[192,204,1945,1964]" pageId="12" pageNumber="76"></emphasis>
1.1 in
</quantity>
horizontal eye diameter. Infraorbital invagination below anterior half of eye or below mideye. Interorbital distance wide,
<quantity id="560C9B51FFBC5B26515FFF300FFE5FC6" box="[821,916,177,197]" metricMagnitude="-2" metricUnit="m" metricValue="1.016" metricValueMax="1.27" metricValueMin="0.762" pageId="12" pageNumber="76" unit="in" value="0.4" valueMax="0.5" valueMin="0.3">
0.3
<emphasis id="A380EAA6FFBC5B265138FF300F345FC7" bold="true" box="[850,862,177,196]" pageId="12" pageNumber="76"></emphasis>
0.5 in
</quantity>
horizontal eye diameter.
<collectingRegion id="5330F856FFBC5B2656ECFF3008AF5FC6" box="[1158,1221,177,197]" country="France" name="Centre" pageId="12" pageNumber="76">Centre</collectingRegion>
of eye much closer to tip of snout than to posterior margin of operculum, preorbital distance in postorbital distance 1.7
<emphasis id="A380EAA6FFBC5B265667FF7608735E09" bold="true" box="[1037,1049,247,266]" pageId="12" pageNumber="76"></emphasis>
2.3. Anterior and posterior nostrils long tubes of about equal length. Nostrils well separated and posterior nostril located behind and dorsally to the anterior edge of eyes. Single large dermal flap at the posterior margin of anterior nostril leaf shaped, longer than nostril. Posterior nostril rim crenate with no extension. Head lateral line system with canals with pores and with superficial neuromasts arranged in rows. Head canals reduced and pores small. Single pore in nasal canal near posterior nostril. Single pore in postorbital canal close to posterior eye edge. Two pores in mandibular canal, anterior one close to anteriolateral angle of mouth, posterior pore slightly in front of vertical of posterior angle of jaws, posterior pore usually more prominent. Lachrymal as well as preopercular canals and pores absent. Rows of superficial neuromasts as follows: SR 2, NR 3, LIR 22
<emphasis id="A380EAA6FFBC5B265118FD380F145DCF" bold="true" box="[882,894,697,716]" pageId="12" pageNumber="76"></emphasis>
29, STR 2
<emphasis id="A380EAA6FFBC5B2651B0FD380F8C5DCF" bold="true" box="[986,998,697,716]" pageId="12" pageNumber="76"></emphasis>
4, POR 3, PTR 2, SLR 4
<emphasis id="A380EAA6FFBC5B2656A9FD3808A55DCF" bold="true" box="[1219,1231,697,716]" pageId="12" pageNumber="76"></emphasis>
5, MR 9
<emphasis id="A380EAA6FFBC5B265771FD38094D5DCF" bold="true" box="[1307,1319,697,716]" pageId="12" pageNumber="76"></emphasis>
11, AVR 2, PVR 1, ADR 2
<emphasis id="A380EAA6FFBC5B2651F9FD5D0FF55DEC" bold="true" box="[915,927,732,751]" pageId="12" pageNumber="76"></emphasis>
3, PDR 1, HR 3
<emphasis id="A380EAA6FFBC5B265655FD5D08215DEC" bold="true" box="[1087,1099,732,751]" pageId="12" pageNumber="76"></emphasis>
4, SR1 3
<emphasis id="A380EAA6FFBC5B2656C9FD5D08C55DEC" bold="true" box="[1187,1199,732,751]" pageId="12" pageNumber="76"></emphasis>
4, SR2 1
<emphasis id="A380EAA6FFBC5B26576DFD5D09795DEC" bold="true" box="[1287,1299,732,751]" pageId="12" pageNumber="76"></emphasis>
2, DLR 5
<emphasis id="A380EAA6FFBC5B265704FD5D09105DEC" bold="true" box="[1390,1402,732,751]" pageId="12" pageNumber="76"></emphasis>
8, VLR 10
<emphasis id="A380EAA6FFBC5B265127FD7E0F335C11" bold="true" box="[845,857,767,786]" pageId="12" pageNumber="76"></emphasis>
13. STR and LIR rows of superficial neuromasts placed in shallow groove disappearing in posterior part of LIR. DLR row of superficial neuromasts anteriorly starts above pectoral fin, continuously dorsolateral and ends posteriorly downwards at or above midlateral level vertical to anus. VLR anteriorly starts behind pectoral fin base, continuous ventrolaterally and ends posteriorly upwards with last papilla near or at midlateral level at or close to caudal-fin base. Mouth terminal, upper and lower lips ends about equally, lips fleshy, upper lip larger than the lower lip. Posterior angle of jaws extends to between vertical line drawn through posterior edge of anterior nostril and vertical line drawn through anterior edge of eye. Chin with bilobed or single lobe fold at anterior edge not covering MR row of superficial neuromasts. The gill membrane is attached to isthmus, gill opening starting at the base of pectoral fin, with the upper attachment of the gill membrane is opposite to 3rd
<emphasis id="A380EAA6FFBC5B26560AFB6508065BF4" bold="true" box="[1120,1132,1252,1271]" pageId="12" pageNumber="76"></emphasis>
4th pectoral ray. Posterior opercular edge with pointed upper tip and rounded lower edge. No subopercular spine. No fleshy pad present on lower pectoral base. Urogenital papilla present. Preanus length in postanus length 0.7
<emphasis id="A380EAA6FFBC5B2657E4FACD09F05A5C" bold="true" box="[1422,1434,1356,1375]" pageId="12" pageNumber="76"></emphasis>
1.0. Anal papillae absent, the area around anus wrinkled.
</paragraph>
<paragraph id="914B36B4FFBC5B265137FA10086158AE" blockId="12.[821,1470,142,1965]" pageId="12" pageNumber="76">
<emphasis id="A380EAA6FFBC5B265137FA100FEA5AA6" box="[861,896,1425,1445]" italics="true" pageId="12" pageNumber="76">Fins</emphasis>
. Rudimentary dorsal and anal fins located well posteriorly and short, reduced to low ridges with the very weak rays, connected to the caudal fin. Pectoral rays 14
<emphasis id="A380EAA6FFBC5B26564FFA56085B5AE9" bold="true" box="[1061,1073,1495,1514]" pageId="12" pageNumber="76"></emphasis>
16. Caudal fin rounded, principal caudal rays 10
<emphasis id="A380EAA6FFBC5B265126FA780F32590F" bold="true" box="[844,856,1529,1548]" pageId="12" pageNumber="76"></emphasis>
12. Ventral adhesive disc (Figure 4c) of
<emphasis id="A380EAA6FFBC5B2656DEFA7808D1590F" bold="true" box="[1204,1211,1529,1548]" pageId="12" pageNumber="76"></emphasis>
double
<emphasis id="A380EAA6FFBC5B26569DFA780895590F" bold="true" box="[1271,1279,1529,1548]" pageId="12" pageNumber="76"></emphasis>
<typeStatus id="4E4F8816FFBC5B26576EFA780947590E" box="[1284,1325,1529,1549]" pageId="12" pageNumber="76">type</typeStatus>
, anterior margin crenate with large invagination on each lateral side and central invagination at midventral; posterior margin crenate. Disc very small, disc length
<quantity id="560C9B51FFBC5B26515FF9E00FFB5976" box="[821,913,1633,1653]" metricMagnitude="-1" metricUnit="m" metricValue="2.1971" metricValueMax="2.4892" metricValueMin="1.905" pageId="12" pageNumber="76" unit="in" value="8.65" valueMax="9.8" valueMin="7.5">
7.5
<emphasis id="A380EAA6FFBC5B26513AF9E00F365977" bold="true" box="[848,860,1633,1652]" pageId="12" pageNumber="76"></emphasis>
9.8 in
</quantity>
SL, its width slightly larger than its length, width in length 0.8
<emphasis id="A380EAA6FFBC5B26513AF9050F365994" bold="true" box="[848,860,1668,1687]" pageId="12" pageNumber="76"></emphasis>
0.9. No papillae in region A and flattened papillae in regions B and C. In region B one or two rows of papillae with total papillae count 12
<emphasis id="A380EAA6FFBC5B2657F2F92609CE59B9" bold="true" box="[1432,1444,1703,1722]" pageId="12" pageNumber="76"></emphasis>
30 and in region C one or two rows of papillae with total papillae count 4
<emphasis id="A380EAA6FFBC5B26512AF96D0F2659FC" bold="true" box="[832,844,1772,1791]" pageId="12" pageNumber="76"></emphasis>
14. No inner row of papillae on lateral sides of the central part of the anterior disc. Upper attachment of disc membrane attaching to base of pectoral fin at 14th
<emphasis id="A380EAA6FFBC5B2651B4F8B00F805847" bold="true" box="[990,1002,1841,1860]" pageId="12" pageNumber="76"></emphasis>
16th pectoral ray (
<emphasis id="A380EAA6FFBC5B2656E6F8B008F45846" box="[1164,1182,1841,1861]" italics="true" pageId="12" pageNumber="76">i.e</emphasis>
., at ultimate or penultimate ray). Males can have two prominent seemingly perfused finger-like extensions on each site of sucking disc that are of equal size or exceeding length of disc region A (Figure 4f).
</paragraph>
<paragraph id="914B36B4FFBD5B2752F5FF0F0E005EB4" blockId="13.[119,769,142,1965]" pageId="13" pageNumber="77">
<emphasis id="A380EAA6FFBD5B2752F5FF0F0D6C5FA1" box="[159,262,142,162]" italics="true" pageId="13" pageNumber="77">Colouration</emphasis>
. Background colour of live specimens bright to skin-coloured (Figure 7). Pigmentation behind head region sometimes with a clearly visible striped or marbled (
<emphasis id="A380EAA6FFBD5B2753AFFF520DB25FE4" box="[453,472,211,231]" italics="true" pageId="13" pageNumber="77">i.e</emphasis>
., irregularly distributed patches of pigments) pattern. Sometimes small evenly distributed iridophores visible in lateral view (Figure 7) in life. No star-like pigmentation around the eyes visible. Formaldehyde fixed specimens white-yellow and without pigments. Ethanol fixed specimens white to skin coloured, striped or marbled pigmentation visible. For more pictures of live colouration see Supporting Information File S1.
</paragraph>
<paragraph id="914B36B4FFBD5B2752F5FE470DE45B8C" blockId="13.[119,769,142,1965]" pageId="13" pageNumber="77">
<emphasis id="A380EAA6FFBD5B2752F5FE470D1F5ED9" box="[159,373,454,474]" italics="true" pageId="13" pageNumber="77">Dentition and osteology</emphasis>
. Upper jaw with outer row of about eight (one side) medium-sized caniniforms frontally. Behind them inner small conical teeth irregularly scattered in two separate (left and right) drop-like patches medially wide about 5
<emphasis id="A380EAA6FFBD5B275381FDAE0D9D5D41" bold="true" box="[491,503,559,578]" pageId="13" pageNumber="77"></emphasis>
6 teeth, becoming narrowed to a single row of teeth laterally. Outer row continues laterally laterally as four large caniniforms of variable size, followed behind by about six medium-sized caniniforms. Lower jaw with outer row of eight to 10 (one side) medium-sized caniniforms frontally. Behind them single broad patch of small conical inner teeth medially wide about 5
<emphasis id="A380EAA6FFBD5B2752D5FD7E0CA15C11" bold="true" box="[191,203,767,786]" pageId="13" pageNumber="77"></emphasis>
6 teeth, becoming narrowed to a single row of teeth laterally. The single row of about five larger caniniforms continuous laterally. Pharyngeal jaws with elongated ceratobranchial 5, having several (about 5) larger elongated conical teeth (Figure 5c), pharyngobranchial 3 toothplate not visible on 3D models from micro-computed tomography (microCT) images. Number of vertebrae 38
<emphasis id="A380EAA6FFBD5B275041FC2D0E5D5CBC" bold="true" box="[555,567,940,959]" pageId="13" pageNumber="77"></emphasis>
40, abdominal 17 and caudal 21
<emphasis id="A380EAA6FFBD5B2752BFFC4E0C8B5CE1" bold="true" box="[213,225,975,994]" pageId="13" pageNumber="77"></emphasis>
22 (Supporting Information Table S2). The first gill arch with hemibranch, the 2nd to 4th gill arches with holobranchs. Subopercle indistinguishable from opercle, shaped as its posterior elongated extension, not forming or having subopercular spine. Six branchiostegals. Hook-shaped nasal bones. Maxillary and premaxillary bones shaped as on Figure 5c.
</paragraph>
</subSubSection>
<subSubSection id="D9EE653FFFBD5B2752F5FB1F0DB65A3E" pageId="13" pageNumber="77" type="etymology">
<paragraph id="914B36B4FFBD5B2752F5FB1F0DB65A3E" blockId="13.[119,769,142,1965]" pageId="13" pageNumber="77">
<emphasis id="A380EAA6FFBD5B2752F5FB1F0D605BB1" bold="true" box="[159,266,1182,1202]" pageId="13" pageNumber="77">Etymology.</emphasis>
Named
<taxonomicName id="56F44D37FFBD5B27530AFB1F0DD15BB1" authorityName="Wagner &amp; Kovac &amp; Koblmüller" authorityYear="2021" box="[352,443,1182,1202]" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="13" pageNumber="77" phylum="Chordata" rank="species" species="hofrichteri">
<emphasis id="A380EAA6FFBD5B27530AFB1F0DD15BB1" box="[352,443,1182,1202]" italics="true" pageId="13" pageNumber="77">hofrichteri</emphasis>
</taxonomicName>
, in honour of Robert Hofrichter, whose work on European clingfishes sparked our interest in these enigmatic fishes. The species epithet was formed from the personal name, as the noun in the genitive case, with
<emphasis id="A380EAA6FFBD5B275077FA860E4F5A19" bold="true" box="[541,549,1287,1306]" pageId="13" pageNumber="77"></emphasis>
i
<emphasis id="A380EAA6FFBD5B275040FA860E585A19" bold="true" box="[554,562,1287,1306]" pageId="13" pageNumber="77"></emphasis>
added to the stem of the name (Article 31.1.2.,
<collectionCode id="F7E5AE71FFBD5B27530EFAA80DFD5A3E" box="[356,407,1321,1341]" pageId="13" pageNumber="77">ICZN</collectionCode>
, 1999).
</paragraph>
</subSubSection>
<subSubSection id="D9EE653FFFBD5B2752F5FACA0D2259DE" pageId="13" pageNumber="77" type="biology_ecology">
<paragraph id="914B36B4FFBD5B2752F5FACA0D2259DE" blockId="13.[119,769,142,1965]" pageId="13" pageNumber="77">
<emphasis id="A380EAA6FFBD5B2752F5FACA0E115A5C" bold="true" box="[159,635,1355,1375]" pageId="13" pageNumber="77">Ecology and geographical distribution (Figure 1a).</emphasis>
Species widespread in the eastern Mediterranean Sea with a single record from the Adriatic Sea (Pelješac; Figure 1c).
<taxonomicName id="56F44D37FFBD5B2753F6FA100E625AA6" authority="Wagner &amp; Kovac &amp; Koblmüller, 2021" authorityName="Wagner &amp; Kovac &amp; Koblmüller" authorityYear="2021" box="[412,520,1425,1445]" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="13" pageNumber="77" phylum="Chordata" rank="species" species="hofrichteri" status="sp. nov.">
<emphasis id="A380EAA6FFBD5B2753F6FA100E625AA6" box="[412,520,1425,1445]" italics="true" pageId="13" pageNumber="77">G. hofrichteri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B8B357DDFFBD5B275064FA110E395AA6" box="[526,595,1424,1445]" pageId="13" pageNumber="77" rank="species">
<emphasis id="A380EAA6FFBD5B275064FA110E245AA7" bold="true" box="[526,590,1424,1444]" pageId="13" pageNumber="77">sp. nov</emphasis>
.
</taxonomicNameLabel>
is very abundant in the northern and southern Ionian Sea (Corfu, Stomio), the
<collectingRegion id="5330F856FFBD5B27501FFA320EF15AC4" box="[629,667,1459,1479]" country="Papua New Guinea" name="Gulf" pageId="13" pageNumber="77">Gulf</collectingRegion>
of Corinth, the islands of Kythira and
<collectingRegion id="5330F856FFBD5B275337FA570DE45AE9" box="[349,398,1494,1514]" country="Greece" name="Kriti" pageId="13" pageNumber="77">Crete</collectingRegion>
and the
<collectingRegion id="5330F856FFBD5B2753B6FA570E755AE9" box="[476,543,1494,1514]" country="Greece" name="Voreio Aigaio" pageId="13" pageNumber="77">Aegean</collectingRegion>
Sea (
<collectingRegion id="5330F856FFBD5B275024FA570EEE5AE9" box="[590,644,1494,1514]" country="Greece" name="Attiki" pageId="13" pageNumber="77">Attica</collectingRegion>
). Quantitative data on ecology is largely lacking. The species inhabits intertidal pebble beaches and probably shows passive emergence behaviour (compare with
<bibRefCitation id="F5654B45FFBD5B27521DF9BF0D705951" author="Bilecenoglu, M." box="[119,282,1598,1618]" pageId="13" pageNumber="77" pagination="270" refId="ref24740" refString="Bilecenoglu, M. (2015). First observation of the amphibious behaviour of Gouania willdenowi (Gobiesocidae) in the eastern Mediterranean Sea. New Mediterranean Biodiversity Records, 16, 270." type="journal article" year="2015">Bilecenoglu, 2015</bibRefCitation>
). Throughout its distribution range this species occurs in sympatry with
<taxonomicName id="56F44D37FFBD5B275370F9E00DEA5976" authority="Wagner &amp; Kovac &amp; Koblmüller, 2021" authorityName="Wagner &amp; Kovac &amp; Koblmüller" authorityYear="2021" box="[282,384,1633,1653]" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="13" pageNumber="77" phylum="Chordata" rank="species" species="orientalis" status="sp. nov.">
<emphasis id="A380EAA6FFBD5B275370F9E00DEA5976" box="[282,384,1633,1653]" italics="true" pageId="13" pageNumber="77">G. orientalis</emphasis>
</taxonomicName>
<emphasis id="A380EAA6FFBD5B2753E2F9E10DBB5977" bold="true" box="[392,465,1632,1652]" pageId="13" pageNumber="77">
<taxonomicNameLabel id="B8B357DDFFBD5B2753E2F9E10DBB5977" box="[392,465,1632,1652]" pageId="13" pageNumber="77" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and on the island Corfu with
<taxonomicName id="56F44D37FFBD5B275086F9E00CAA5994" authority="Wagner &amp; Kovac &amp; Koblmüller, 2021" authorityName="Wagner &amp; Kovac &amp; Koblmüller" authorityYear="2021" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="13" pageNumber="77" phylum="Chordata" rank="species" species="adriatica" status="sp. nov.">
<emphasis id="A380EAA6FFBD5B275086F9E00CAA5994" italics="true" pageId="13" pageNumber="77">G. adriatica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B8B357DDFFBD5B2752ACF9020D615994" box="[198,267,1667,1687]" pageId="13" pageNumber="77" rank="species">
<emphasis id="A380EAA6FFBD5B2752ACF9020D6C5994" bold="true" box="[198,262,1667,1687]" pageId="13" pageNumber="77">sp. nov</emphasis>
.
</taxonomicNameLabel>
There is a single record of the species inside the Adriatic basin from Pelješac, where it occurs in low densities together with
<taxonomicName id="56F44D37FFBD5B2750D7F9270E9559B9" baseAuthorityName="Nardo" baseAuthorityYear="1827" box="[701,767,1702,1722]" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="13" pageNumber="77" phylum="Chordata" rank="species" species="pigra">
<emphasis id="A380EAA6FFBD5B2750D7F9270E9559B9" box="[701,767,1702,1722]" italics="true" pageId="13" pageNumber="77">G. pigra</emphasis>
</taxonomicName>
and
<taxonomicName id="56F44D37FFBD5B2752F7F9480C9459DE" authority="Wagner &amp; Kovac &amp; Koblmüller, 2021" authorityName="Wagner &amp; Kovac &amp; Koblmüller" authorityYear="2021" box="[157,254,1737,1757]" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="13" pageNumber="77" phylum="Chordata" rank="species" species="adriatica" status="sp. nov.">
<emphasis id="A380EAA6FFBD5B2752F7F9480C9459DE" box="[157,254,1737,1757]" italics="true" pageId="13" pageNumber="77">G. adriatica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B8B357DDFFBD5B275369F9490D2259DE" box="[259,328,1736,1757]" pageId="13" pageNumber="77" rank="species">
<emphasis id="A380EAA6FFBD5B275369F9490D2959DF" bold="true" box="[259,323,1736,1756]" pageId="13" pageNumber="77">sp. nov</emphasis>
.
</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="D9EE653FFFBD5B2752F5F96A080A5A5C" pageId="13" pageNumber="77" type="discussion">
<paragraph id="914B36B4FFBD5B2752F5F96A080A5A5C" blockId="13.[119,769,142,1965]" lastBlockId="13.[816,1465,142,1375]" pageId="13" pageNumber="77">
<emphasis id="A380EAA6FFBD5B2752F5F96A0C9D59FC" bold="true" box="[159,247,1771,1791]" pageId="13" pageNumber="77">Remarks.</emphasis>
<taxonomicName id="56F44D37FFBD5B275296F96A0DC259FC" authority="Wagner &amp; Kovac &amp; Koblmüller, 2021" authorityName="Wagner &amp; Kovac &amp; Koblmüller" authorityYear="2021" box="[252,424,1771,1791]" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="13" pageNumber="77" phylum="Chordata" rank="species" species="hofrichteri" status="sp. nov.">
<emphasis id="A380EAA6FFBD5B275296F96A0DC259FC" box="[252,424,1771,1791]" italics="true" pageId="13" pageNumber="77">Gouania hofrichteri</emphasis>
</taxonomicName>
<emphasis id="A380EAA6FFBD5B2753C4F96A0D9259FC" bold="true" box="[430,504,1771,1791]" pageId="13" pageNumber="77">
<taxonomicNameLabel id="B8B357DDFFBD5B2753C4F96A0D9259FC" box="[430,504,1771,1791]" pageId="13" pageNumber="77" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from all other
<taxonomicName id="56F44D37FFBD5B2750D3F96A0CDC5821" pageId="13" pageNumber="77">
<emphasis id="A380EAA6FFBD5B2750D3F96A0E9559FC" box="[697,767,1771,1791]" italics="true" pageId="13" pageNumber="77">Gouania</emphasis>
species
</taxonomicName>
by longitudinal infralateral and suborbital transversal rows of superficial neuromasts placed in shallow groove disappearing in posterior part of longitudinal infralateral row (
<emphasis id="A380EAA6FFBD5B2753E9F8D20DFE5864" box="[387,404,1875,1895]" italics="true" pageId="13" pageNumber="77">vs</emphasis>
. longitudinal infralateral and suborbital transversal rows of superficial neuromasts placed in the well-defined deep groove in all other
<taxonomicName id="56F44D37FFBD5B275371F8180DC258AE" box="[283,424,1945,1965]" pageId="13" pageNumber="77">
<emphasis id="A380EAA6FFBD5B275371F8180D0B58AE" box="[283,353,1945,1965]" italics="true" pageId="13" pageNumber="77">Gouania</emphasis>
species
</taxonomicName>
); body cross-section behind pectoral fin base triangular with ventral flat and dorsal pointed (
<emphasis id="A380EAA6FFBD5B27569DFF0F09625FA1" box="[1271,1288,142,162]" italics="true" pageId="13" pageNumber="77">vs</emphasis>
. body cross-section behind pectoral fin base half oval to pentagonal (pentagonal only in
<taxonomicName id="56F44D37FFBD5B2757CFFF300F335FE4" baseAuthorityName="Nardo" baseAuthorityYear="1827" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="13" pageNumber="77" phylum="Chordata" rank="species" species="pigra">
<emphasis id="A380EAA6FFBD5B2757CFFF300F335FE4" italics="true" pageId="13" pageNumber="77">G. pigra</emphasis>
</taxonomicName>
) with straight ventral side in all other
<taxonomicName id="56F44D37FFBD5B2756D9FF5209285FE4" box="[1203,1346,211,231]" pageId="13" pageNumber="77">
<emphasis id="A380EAA6FFBD5B2756D9FF5208935FE4" box="[1203,1273,211,231]" italics="true" pageId="13" pageNumber="77">Gouania</emphasis>
species
</taxonomicName>
); granules on body, at least on posterior part and nape, large and prominent (
<emphasis id="A380EAA6FFBD5B275716FF7709E75E09" box="[1404,1421,246,266]" italics="true" pageId="13" pageNumber="77">vs</emphasis>
. the granules on body shallow and inconspicuous in all other
<taxonomicName id="56F44D37FFBD5B27574DFE9809D95E2E" box="[1319,1459,281,301]" pageId="13" pageNumber="77">
<emphasis id="A380EAA6FFBD5B27574DFE9809075E2E" box="[1319,1389,281,301]" italics="true" pageId="13" pageNumber="77">Gouania</emphasis>
species
</taxonomicName>
) and hook-like nasal bones.
<taxonomicName id="56F44D37FFBD5B27564AFEBA08E75E4C" authority="Wagner &amp; Kovac &amp; Koblmüller, 2021" authorityName="Wagner &amp; Kovac &amp; Koblmüller" authorityYear="2021" box="[1056,1165,315,335]" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="13" pageNumber="77" phylum="Chordata" rank="species" species="hofrichteri" status="sp. nov.">
<emphasis id="A380EAA6FFBD5B27564AFEBA08E75E4C" box="[1056,1165,315,335]" italics="true" pageId="13" pageNumber="77">G. hofrichteri</emphasis>
</taxonomicName>
<emphasis id="A380EAA6FFBD5B2756FEFEBA08B05E4C" bold="true" box="[1172,1242,315,335]" pageId="13" pageNumber="77">
<taxonomicNameLabel id="B8B357DDFFBD5B2756FEFEBA08B05E4C" box="[1172,1242,315,335]" pageId="13" pageNumber="77" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from three stout-bodied
<taxonomicName id="56F44D37FFBD5B275119FEDF0F955E71" box="[883,1023,350,370]" pageId="13" pageNumber="77">
<emphasis id="A380EAA6FFBD5B275119FEDF0FD35E71" box="[883,953,350,370]" italics="true" pageId="13" pageNumber="77">Gouania</emphasis>
species
</taxonomicName>
(
<taxonomicName id="56F44D37FFBD5B275661FEDF08075E71" authority="Wagner &amp; Kovac &amp; Koblmüller, 2021" authorityName="Wagner &amp; Kovac &amp; Koblmüller" authorityYear="2021" box="[1035,1133,350,370]" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="13" pageNumber="77" phylum="Chordata" rank="species" species="adriatica" status="sp. nov.">
<emphasis id="A380EAA6FFBD5B275661FEDF08075E71" box="[1035,1133,350,370]" italics="true" pageId="13" pageNumber="77">G. adriatica</emphasis>
</taxonomicName>
<emphasis id="A380EAA6FFBD5B27561EFEDF08D05E71" bold="true" box="[1140,1210,350,370]" pageId="13" pageNumber="77">
<taxonomicNameLabel id="B8B357DDFFBD5B27561EFEDF08D05E71" box="[1140,1210,350,370]" pageId="13" pageNumber="77" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="56F44D37FFBD5B2756AFFEDF09435E71" authority="Wagner &amp; Kovac &amp; Koblmüller, 2021" authorityName="Wagner &amp; Kovac &amp; Koblmüller" authorityYear="2021" box="[1221,1321,350,370]" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="13" pageNumber="77" phylum="Chordata" rank="species" species="orientalis" status="sp. nov.">
<emphasis id="A380EAA6FFBD5B2756AFFEDF09435E71" box="[1221,1321,350,370]" italics="true" pageId="13" pageNumber="77">G. orientalis</emphasis>
</taxonomicName>
<emphasis id="A380EAA6FFBD5B27575AFEDF091D5E71" bold="true" box="[1328,1399,350,370]" pageId="13" pageNumber="77">
<taxonomicNameLabel id="B8B357DDFFBD5B27575AFEDF091D5E71" box="[1328,1399,350,370]" pageId="13" pageNumber="77" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="56F44D37FFBD5B2757CFFEDF0FEC5E96" baseAuthorityName="Risso" baseAuthorityYear="1810" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="13" pageNumber="77" phylum="Chordata" rank="species" species="willdenowi">
<emphasis id="A380EAA6FFBD5B2757CFFEDF0FEC5E96" italics="true" pageId="13" pageNumber="77">G. willdenowi</emphasis>
</taxonomicName>
) by dorsal head profile in lateral view
<emphasis id="A380EAA6FFBD5B2756BBFE0008B25E97" bold="true" box="[1233,1240,385,404]" pageId="13" pageNumber="77"></emphasis>
S
<emphasis id="A380EAA6FFBD5B275689FE0008815E97" bold="true" box="[1251,1259,385,404]" pageId="13" pageNumber="77"></emphasis>
curved, concave above eye and convex at nape (
<emphasis id="A380EAA6FFBD5B275676FE2208475EB4" box="[1052,1069,419,439]" italics="true" pageId="13" pageNumber="77">vs</emphasis>
. dorsal head profile straight between nape above eye and upper lip tip), absence of star-like pigmentation around eyes and large number of vertebrae (Supporting Information Table S2; 38
<emphasis id="A380EAA6FFBD5B2757FEFE6809F55EFF" bold="true" box="[1428,1439,489,508]" pageId="13" pageNumber="77"></emphasis>
40
<emphasis id="A380EAA6FFBD5B27515AFD8A0F2B5D1C" box="[816,833,523,543]" italics="true" pageId="13" pageNumber="77">vs</emphasis>
. 35
<emphasis id="A380EAA6FFBD5B27510CFD8D0F1B5D1C" bold="true" box="[870,881,524,543]" pageId="13" pageNumber="77"></emphasis>
38). Eleven morphometric characters as percentages of standard length of
<taxonomicName id="56F44D37FFBD5B2751ECFDAF0F995D41" authority="Wagner &amp; Kovac &amp; Koblmüller, 2021" authorityName="Wagner &amp; Kovac &amp; Koblmüller" authorityYear="2021" box="[902,1011,558,578]" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="13" pageNumber="77" phylum="Chordata" rank="species" species="hofrichteri" status="sp. nov.">
<emphasis id="A380EAA6FFBD5B2751ECFDAF0F995D41" box="[902,1011,558,578]" italics="true" pageId="13" pageNumber="77">G. hofrichteri</emphasis>
</taxonomicName>
<emphasis id="A380EAA6FFBD5B275193FDAF082A5D41" bold="true" box="[1017,1088,558,578]" pageId="13" pageNumber="77">
<taxonomicNameLabel id="B8B357DDFFBD5B275193FDAF082A5D41" box="[1017,1088,558,578]" pageId="13" pageNumber="77" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
are nonoverlapping in range with all three stout-bodied
<taxonomicName id="56F44D37FFBD5B2751CDFDD00F865D66" authorityName="Nardo" authorityYear="1833" box="[935,1004,593,613]" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="13" pageNumber="77" phylum="Chordata" rank="genus">
<emphasis id="A380EAA6FFBD5B2751CDFDD00F865D66" box="[935,1004,593,613]" italics="true" pageId="13" pageNumber="77">Gouania</emphasis>
</taxonomicName>
: vertical eye diameter, head length, postorbital distance, all three head widths, body width at pectoral fins, pectoral-fin length, prepectoral distance, ventral adhesive disc length and caudal-fin length (values in the Table 1). In addition, there are numerous morphometric characters nonoverlapping in range with one or two out of three stout-bodied
<taxonomicName id="56F44D37FFBD5B27511FFD7F08695C11" box="[885,1027,766,786]" pageId="13" pageNumber="77">
<emphasis id="A380EAA6FFBD5B27511FFD7F0FD15C11" box="[885,955,766,786]" italics="true" pageId="13" pageNumber="77">Gouania</emphasis>
species
</taxonomicName>
(Table 1).
<taxonomicName id="56F44D37FFBD5B275602FD7F08B25C11" authority="Wagner &amp; Kovac &amp; Koblmüller, 2021" authorityName="Wagner &amp; Kovac &amp; Koblmüller" authorityYear="2021" box="[1128,1240,766,786]" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="13" pageNumber="77" phylum="Chordata" rank="species" species="hofrichteri" status="sp. nov.">
<emphasis id="A380EAA6FFBD5B275602FD7F08B25C11" box="[1128,1240,766,786]" italics="true" pageId="13" pageNumber="77">G. hofrichteri</emphasis>
</taxonomicName>
<emphasis id="A380EAA6FFBD5B27568BFD7F09405C11" bold="true" box="[1249,1322,766,786]" pageId="13" pageNumber="77">
<taxonomicNameLabel id="B8B357DDFFBD5B27568BFD7F09405C11" box="[1249,1322,766,786]" pageId="13" pageNumber="77" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from
<taxonomicName id="56F44D37FFBD5B2757CFFD7F0F125C36" authority="Wagner &amp; Kovac &amp; Koblmüller, 2021" authorityName="Wagner &amp; Kovac &amp; Koblmüller" authorityYear="2021" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="13" pageNumber="77" phylum="Chordata" rank="species" species="adriatica" status="sp. nov.">
<emphasis id="A380EAA6FFBD5B2757CFFD7F0F125C36" italics="true" pageId="13" pageNumber="77">G. adriatica</emphasis>
</taxonomicName>
<emphasis id="A380EAA6FFBD5B275114FCA10FAE5C37" bold="true" box="[894,964,800,820]" pageId="13" pageNumber="77">
<taxonomicNameLabel id="B8B357DDFFBD5B275114FCA10FAE5C37" box="[894,964,800,820]" pageId="13" pageNumber="77" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="56F44D37FFBD5B275185FCA0083B5C36" authority="Wagner &amp; Kovac &amp; Koblmüller, 2021" authorityName="Wagner &amp; Kovac &amp; Koblmüller" authorityYear="2021" box="[1007,1105,801,821]" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="13" pageNumber="77" phylum="Chordata" rank="species" species="orientalis" status="sp. nov.">
<emphasis id="A380EAA6FFBD5B275185FCA0083B5C36" box="[1007,1105,801,821]" italics="true" pageId="13" pageNumber="77">G. orientalis</emphasis>
</taxonomicName>
<emphasis id="A380EAA6FFBD5B27563DFCA108F75C37" bold="true" box="[1111,1181,800,820]" pageId="13" pageNumber="77">
<taxonomicNameLabel id="B8B357DDFFBD5B27563DFCA108F75C37" box="[1111,1181,800,820]" pageId="13" pageNumber="77" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
by the upper attachment of the gill membrane opposite to 3rd
<emphasis id="A380EAA6FFBD5B275657FCC508225C54" bold="true" box="[1085,1096,836,855]" pageId="13" pageNumber="77"></emphasis>
4th pectoral ray (
<emphasis id="A380EAA6FFBD5B275689FCC2089E5C54" box="[1251,1268,835,855]" italics="true" pageId="13" pageNumber="77">vs</emphasis>
. upper attachment of the gill membrane opposite to 5th
<emphasis id="A380EAA6FFBD5B275636FCE6080D5C79" bold="true" box="[1116,1127,871,890]" pageId="13" pageNumber="77"></emphasis>
6th pectoral ray) and from
<taxonomicName id="56F44D37FFBD5B27573FFCE709DD5C79" authority="Wagner &amp; Kovac &amp; Koblmüller, 2021" authorityName="Wagner &amp; Kovac &amp; Koblmüller" authorityYear="2021" box="[1365,1463,870,890]" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="13" pageNumber="77" phylum="Chordata" rank="species" species="orientalis" status="sp. nov.">
<emphasis id="A380EAA6FFBD5B27573FFCE709DD5C79" box="[1365,1463,870,890]" italics="true" pageId="13" pageNumber="77">G. orientalis</emphasis>
</taxonomicName>
<emphasis id="A380EAA6FFBD5B27515AFC090F1D5C9F" bold="true" box="[816,887,904,924]" pageId="13" pageNumber="77">
<taxonomicNameLabel id="B8B357DDFFBD5B27515AFC090F1D5C9F" box="[816,887,904,924]" pageId="13" pageNumber="77" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="56F44D37FFBD5B2751CCFC08087D5C9E" baseAuthorityName="Risso" baseAuthorityYear="1810" box="[934,1047,905,925]" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="13" pageNumber="77" phylum="Chordata" rank="species" species="willdenowi">
<emphasis id="A380EAA6FFBD5B2751CCFC08087D5C9E" box="[934,1047,905,925]" italics="true" pageId="13" pageNumber="77">G. willdenowi</emphasis>
</taxonomicName>
by the infraorbital invagination below anterior half of eye or below mideye (
<emphasis id="A380EAA6FFBD5B275659FC2A082E5CBC" box="[1075,1092,939,959]" italics="true" pageId="13" pageNumber="77">vs</emphasis>
. infraorbital invagination vertical to posterior part of eye) and by posterior opercular edge with pointed upper tip and rounded lower edge (
<emphasis id="A380EAA6FFBD5B275679FC70084E5B06" box="[1043,1060,1009,1029]" italics="true" pageId="13" pageNumber="77">vs</emphasis>
. posterior opercular edge w-shaped with two equally long tips).
<taxonomicName id="56F44D37FFBD5B2751BAFB9208545B24" authority="Wagner &amp; Kovac &amp; Koblmüller, 2021" authorityName="Wagner &amp; Kovac &amp; Koblmüller" authorityYear="2021" box="[976,1086,1043,1063]" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="13" pageNumber="77" phylum="Chordata" rank="species" species="hofrichteri" status="sp. nov.">
<emphasis id="A380EAA6FFBD5B2751BAFB9208545B24" box="[976,1086,1043,1063]" italics="true" pageId="13" pageNumber="77">G. hofrichteri</emphasis>
</taxonomicName>
<emphasis id="A380EAA6FFBD5B27562EFB9208E15B24" bold="true" box="[1092,1163,1043,1063]" pageId="13" pageNumber="77">
<taxonomicNameLabel id="B8B357DDFFBD5B27562EFB9208E15B24" box="[1092,1163,1043,1063]" pageId="13" pageNumber="77" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from other slender-bodied species,
<taxonomicName id="56F44D37FFBD5B275113FBB70FD05B49" baseAuthorityName="Nardo" baseAuthorityYear="1827" box="[889,954,1078,1098]" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="13" pageNumber="77" phylum="Chordata" rank="species" species="pigra">
<emphasis id="A380EAA6FFBD5B275113FBB70FD05B49" box="[889,954,1078,1098]" italics="true" pageId="13" pageNumber="77">G. pigra</emphasis>
</taxonomicName>
, by the posterior angle of jaws extending to between vertical line drawn through posterior edge of anterior nostril and vertical line drawn through anterior edge of eye (
<emphasis id="A380EAA6FFBD5B27561DFBFA08E25B8C" box="[1143,1160,1147,1167]" italics="true" pageId="13" pageNumber="77">vs</emphasis>
. posterior angle of jaws extending to, or close to, vertical line drawn through anterior edge of anterior nostril) and by posterior opercular edge with pointed upper tip and rounded lower edge (
<emphasis id="A380EAA6FFBD5B27510CFB620F1D5BF4" box="[870,887,1251,1271]" italics="true" pageId="13" pageNumber="77">vs</emphasis>
. posterior opercular edge with two tips, upper longer or equal to lower).
<taxonomicName id="56F44D37FFBD5B275119FA870F8B5A19" authority="Wagner &amp; Kovac &amp; Koblmüller, 2021" authorityName="Wagner &amp; Kovac &amp; Koblmüller" authorityYear="2021" box="[883,993,1286,1306]" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="13" pageNumber="77" phylum="Chordata" rank="species" species="hofrichteri" status="sp. nov.">
<emphasis id="A380EAA6FFBD5B275119FA870F8B5A19" box="[883,993,1286,1306]" italics="true" pageId="13" pageNumber="77">G. hofrichteri</emphasis>
</taxonomicName>
<emphasis id="A380EAA6FFBD5B275183FA87085B5A19" bold="true" box="[1001,1073,1286,1306]" pageId="13" pageNumber="77">
<taxonomicNameLabel id="B8B357DDFFBD5B275183FA87085B5A19" box="[1001,1073,1286,1306]" pageId="13" pageNumber="77" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is known from the Adriatic (single findings from Pelješac),
<collectingRegion id="5330F856FFBD5B2751DEFAA80F9C5A3E" box="[948,1014,1321,1341]" country="Greece" name="Voreio Aigaio" pageId="13" pageNumber="77">Aegean</collectingRegion>
and Ionian Sea and has nonoverlapping geographic distribution only with
<taxonomicName id="56F44D37FFBD5B275184FACA08315A5C" baseAuthorityName="Risso" baseAuthorityYear="1810" box="[1006,1115,1355,1375]" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="13" pageNumber="77" phylum="Chordata" rank="species" species="willdenowi">
<emphasis id="A380EAA6FFBD5B275184FACA08315A5C" box="[1006,1115,1355,1375]" italics="true" pageId="13" pageNumber="77">G. willdenowi</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>