818 lines
81 KiB
XML
818 lines
81 KiB
XML
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<mods:title id="69BEAB339A8826E795CC4BF276804B77">Unravelling the taxonomy of an interstitial fish radiation: Three new species of Gouania (Teleostei: Gobiesocidae) from the Mediterranean Sea and redescriptions of G. willdenowi and G. pigra</mods:title>
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<mods:namePart id="56779D5A0BE93EBFCA7634FFBDCEE05C">Wagner, Maximilian</mods:namePart>
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<mods:affiliation id="883C0F7BE8121A442FEB16EF7F80A121">Institute of Biology, University of Graz, Graz, Austria & Department of Biology, University of Antwerp, Antwerp, Belgium</mods:affiliation>
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<mods:namePart id="A050660CBFD541D5D03D3B10E59398DC">Kovac, Marcelo</mods:namePart>
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<mods:affiliation id="937E3A679ED77F8212F5F2A0FE2B06E1">Natural History Museum Rijeka, Rijeka, Croatia</mods:affiliation>
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<mods:namePart id="530CAA9A9CC2312AB94848136DE4604F">Koblmüller, Stephan</mods:namePart>
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<mods:affiliation id="C8476706870A6755A62531171C02A234">Institute of Biology, University of Graz, Graz, Austria</mods:affiliation>
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<mods:title id="5DBD1B8FC644E06B78417E97D92BDE91">Journal of Fish Biology</mods:title>
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<mods:part id="AE9E5698B8B090F802D2CABB33645541">
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<mods:date id="91C002FC246A22BC6070CEC750E47D33">2021</mods:date>
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<mods:number id="8369144D10E8C3F9CF2215AB25194537">2020-11-03</mods:number>
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<mods:number id="ECE8E1FBFD0E74349DB5D2F0E71B4540">98</mods:number>
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<treatment id="195D87A2FFB85B21515FF9DC0FF85C11" LSID="urn:lsid:plazi:treatment:195D87A2FFB85B21515FF9DC0FF85C11" httpUri="http://treatment.plazi.org/id/195D87A2FFB85B21515FF9DC0FF85C11" lastPageId="11" lastPageNumber="75" pageId="8" pageNumber="72">
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<subSubSection id="D9EE653FFFB85B22515FF9DC08895975" box="[821,1251,1628,1654]" pageId="8" pageNumber="72" type="nomenclature">
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<paragraph id="914B36B4FFB85B22515FF9DC08895975" blockId="8.[821,1251,1628,1654]" box="[821,1251,1628,1654]" pageId="8" pageNumber="72">
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<heading id="CA0381D8FFB85B22515FF9DC08895975" box="[821,1251,1628,1654]" fontSize="10" level="3" pageId="8" pageNumber="72" reason="9">
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<emphasis id="A380EAA6FFB85B22515FF9DC0F055975" bold="true" box="[821,879,1629,1654]" pageId="8" pageNumber="72">3.1.3</emphasis>
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|
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<emphasis id="A380EAA6FFB85B2251DFF9DD08895975" bold="true" box="[949,1251,1628,1654]" pageId="8" pageNumber="72">
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<taxonomicName id="56F44D37FFB85B2251DFF9DD08E85975" authority="Wagner & Kovac & Koblmüller, 2021" authorityName="Wagner & Kovac & Koblmüller" authorityYear="2021" box="[949,1154,1628,1654]" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="8" pageNumber="72" phylum="Chordata" rank="species" species="orientalis" status="sp. nov.">
|
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<emphasis id="A380EAA6FFB85B2251DFF9DD08E85975" bold="true" box="[949,1154,1628,1654]" italics="true" pageId="8" pageNumber="72">Gouania orientalis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="B8B357DDFFB85B2256E3F9DC08895975" box="[1161,1251,1629,1654]" pageId="8" pageNumber="72" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<subSubSection id="D9EE653FFFB85B22515FF927086259FC" pageId="8" pageNumber="72" type="description">
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<emphasis id="A380EAA6FFB85B22515FF927088C59B9" bold="true" box="[821,1254,1702,1722]" pageId="8" pageNumber="72">
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English name:
|
||
<collectingRegion id="5330F856FFB85B2251D1F927086259B9" box="[955,1032,1702,1722]" country="Oman" name="Shamal ash Sharqiyah" pageId="8" pageNumber="72">Oriental</collectingRegion>
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blunt-snouted clingfish
|
||
</emphasis>
|
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</paragraph>
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<paragraph id="914B36B4FFB85B225137F948086259FC" blockId="8.[821,1470,1702,1965]" pageId="8" pageNumber="72">
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<emphasis id="A380EAA6FFB85B225137F9480F8D59DE" box="[861,999,1737,1757]" italics="true" pageId="8" pageNumber="72">
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ZooBank
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<collectionCode id="F7E5AE71FFB85B2251D4F9480F8D59DE" box="[958,999,1737,1757]" pageId="8" pageNumber="72">LSID</collectionCode>
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</emphasis>
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:
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<uri id="E5653AB6FFB85B225195F948086259FC" pageId="8" pageNumber="72">
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urn:lsid:zoobank.org:act:
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</uri>
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</paragraph>
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<paragraph id="914B36B4FFB85B225137F88F0FA25864" blockId="8.[821,1470,1702,1965]" pageId="8" pageNumber="72">
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<materialsCitation id="219C3CE9FFB85B225137F88F0FA25864" collectingDate="2016-08-09" collectionCode="PMR" collectorName="M. Wagner" country="Greece" location="Crete" pageId="8" pageNumber="72" specimenCode="VP4585" specimenCount="1" stateProvince="Kriti" typeStatus="holotype">
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<emphasis id="A380EAA6FFB85B225137F88F0FD05821" bold="true" box="[861,954,1806,1826]" pageId="8" pageNumber="72">
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<typeStatus id="4E4F8816FFB85B225137F88F0FDC5821" box="[861,950,1806,1826]" pageId="8" pageNumber="72" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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<collectionCode id="F7E5AE71FFB85B2251AEF88F0F985821" box="[964,1010,1806,1826]" country="0" httpUri="http://grbio.org/cool/xq4z-q5gq" name="Prirodoslovni muzej Rijeka" pageId="8" pageNumber="72">PMR</collectionCode>
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<specimenCode id="C1529ECFFFB85B225196F88F08215821" box="[1020,1099,1806,1826]" collectionCode="VP" pageId="8" pageNumber="72">VP4585</specimenCode>
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, male, 32.8 +
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<quantity id="560C9B51FFB85B2256BEF88F09425821" box="[1236,1320,1806,1826]" metricMagnitude="-3" metricUnit="m" metricValue="4.93" pageId="8" pageNumber="72" unit="mm" value="4.93">4.93 mm</quantity>
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, Plakias,
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<collectingRegion id="5330F856FFB85B2257EEF88F09D05821" box="[1412,1466,1806,1826]" country="Greece" name="Kriti" pageId="8" pageNumber="72">Crete</collectingRegion>
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,
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<collectingCountry id="E9E37624FFB85B22515FF8B00F105846" box="[821,890,1841,1861]" name="Greece" pageId="8" pageNumber="72">Greece</collectingCountry>
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, 35 11
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<emphasis id="A380EAA6FFB85B2251A8F8AF0FAC5838" bold="true" box="[962,966,1838,1851]" italics="true" pageId="8" pageNumber="72">
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<superScript id="66819BFCFFB85B2251A8F8AF0FAC5838" attach="none" box="[962,966,1838,1851]" fontSize="6" pageId="8" pageNumber="72">0</superScript>
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</emphasis>
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40.8
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||
<emphasis id="A380EAA6FFB85B22519BF8AF0F935838" bold="true" box="[1009,1017,1838,1851]" italics="true" pageId="8" pageNumber="72">
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</emphasis>
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N, 24 22
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||
<emphasis id="A380EAA6FFB85B22563FF8AF08335838" bold="true" box="[1109,1113,1838,1851]" italics="true" pageId="8" pageNumber="72">
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<superScript id="66819BFCFFB85B22563FF8AF08335838" attach="none" box="[1109,1113,1838,1851]" fontSize="6" pageId="8" pageNumber="72">0</superScript>
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</emphasis>
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||
50.9
|
||
<emphasis id="A380EAA6FFB85B2256EEF8AF08E65838" bold="true" box="[1156,1164,1838,1851]" italics="true" pageId="8" pageNumber="72">
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<superScript id="66819BFCFFB85B2256EEF8AF08E65838" attach="right" box="[1156,1164,1838,1851]" fontSize="6" pageId="8" pageNumber="72">00</superScript>
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</emphasis>
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E, coll.
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||
<collectorName id="3C015362FFB85B2256B7F8B009245846" box="[1245,1358,1841,1861]" pageId="8" pageNumber="72">M. Wagner</collectorName>
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,
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||
<date id="E54A1074FFB85B225737F8B00F0C5864" pageId="8" pageNumber="72" value="2016-08-09">
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||
<collectingDate id="F50EE99CFFB85B225737F8B00F0C5864" pageId="8" pageNumber="72" value="2016-08-09">August 9, 2016</collectingDate>
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||
</date>
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||
(Figure 6).
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||
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</paragraph>
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||
<paragraph id="914B36B4FFB85B225137F8F709D758AE" blockId="8.[821,1470,1702,1965]" pageId="8" pageNumber="72">
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<materialsCitation id="219C3CE9FFB85B225137F8F709FE58AE" collectionCode="PMR" collectorName="M. Wagner" country="Greece" location="Gulf of Corinth" pageId="8" pageNumber="72" specimenCode="VP4719" specimenCount="1" typeStatus="paratype">
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||
<emphasis id="A380EAA6FFB85B225137F8F70FAA5889" bold="true" box="[861,960,1910,1930]" pageId="8" pageNumber="72">
|
||
<typeStatus id="4E4F8816FFB85B225137F8F70FD65889" box="[861,956,1910,1930]" pageId="8" pageNumber="72" type="paratype">Paratypes</typeStatus>
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.
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||
</emphasis>
|
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<collectionCode id="F7E5AE71FFB85B2251AEF8F70F985889" box="[964,1010,1910,1930]" country="0" httpUri="http://grbio.org/cool/xq4z-q5gq" name="Prirodoslovni muzej Rijeka" pageId="8" pageNumber="72">PMR</collectionCode>
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<specimenCode id="C1529ECFFFB85B22519DF8F7082F5889" box="[1015,1093,1910,1930]" collectionCode="VP" pageId="8" pageNumber="72">VP4719</specimenCode>
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, male, 37.53 +
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<quantity id="560C9B51FFB85B2256BAF8F7094E5889" box="[1232,1316,1910,1930]" metricMagnitude="-3" metricUnit="m" metricValue="5.59" pageId="8" pageNumber="72" unit="mm" value="5.59">5.59 mm</quantity>
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,
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<location id="942B606FFFB85B225746F8F709D05889" LSID="urn:lsid:plazi:treatment:195D87A2FFB85B21515FF9DC0FF85C11:942B606FFFB85B225746F8F709D05889" box="[1324,1466,1910,1930]" country="Greece" name="Gulf of Corinth" pageId="8" pageNumber="72">Gulf of Corinth</location>
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||
,
|
||
<collectingCountry id="E9E37624FFB85B22515FF8180F1058AE" box="[821,890,1945,1965]" name="Greece" pageId="8" pageNumber="72">Greece</collectingCountry>
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, 38 10
|
||
<emphasis id="A380EAA6FFB85B2251AAF8170FAE58A0" bold="true" box="[960,964,1942,1955]" italics="true" pageId="8" pageNumber="72">
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</emphasis>
|
||
17.0
|
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<emphasis id="A380EAA6FFB85B225185F8170F9D58A0" bold="true" box="[1007,1015,1942,1955]" italics="true" pageId="8" pageNumber="72">
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<superScript id="66819BFCFFB85B225185F8170F9D58A0" attach="right" box="[1007,1015,1942,1955]" fontSize="6" pageId="8" pageNumber="72">00</superScript>
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</emphasis>
|
||
N, 22 16
|
||
<emphasis id="A380EAA6FFB85B225638F817083C58A0" bold="true" box="[1106,1110,1942,1955]" italics="true" pageId="8" pageNumber="72">
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<superScript id="66819BFCFFB85B225638F817083C58A0" attach="none" box="[1106,1110,1942,1955]" fontSize="6" pageId="8" pageNumber="72">0</superScript>
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</emphasis>
|
||
26.7
|
||
<emphasis id="A380EAA6FFB85B2256EAF81708E258A0" bold="true" box="[1152,1160,1942,1955]" italics="true" pageId="8" pageNumber="72">
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<superScript id="66819BFCFFB85B2256EAF81708E258A0" attach="right" box="[1152,1160,1942,1955]" fontSize="6" pageId="8" pageNumber="72">00</superScript>
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</emphasis>
|
||
E, coll.
|
||
<collectorName id="3C015362FFB85B2256BCF818092F58AE" box="[1238,1349,1945,1965]" pageId="8" pageNumber="72">M. Wagner</collectorName>
|
||
, August
|
||
</materialsCitation>
|
||
23,
|
||
</paragraph>
|
||
<caption id="C58B663CFFB95B23521DFB1409ED5BE1" ID-DOI="http://doi.org/10.5281/zenodo.10928934" ID-Zenodo-Dep="10928934" httpUri="https://zenodo.org/record/10928934/files/figure.png" pageId="9" pageNumber="73" startId="9.[119,131,1173,1192]" targetBox="[264,1318,138,1132]" targetPageId="9" targetType="figure">
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<paragraph id="914B36B4FFB95B23521DFB1409ED5BE1" blockId="9.[119,1458,1172,1250]" pageId="9" pageNumber="73">
|
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<emphasis id="A380EAA6FFB95B23521DFB140C8D5BAB" bold="true" box="[119,231,1173,1192]" pageId="9" pageNumber="73">F I G U R E 5</emphasis>
|
||
Head osteology of
|
||
<taxonomicName id="56F44D37FFB95B2353DEFB150E245BAB" box="[436,590,1172,1192]" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="9" pageNumber="73" phylum="Chordata" rank="species" species="undetermined">
|
||
<emphasis id="A380EAA6FFB95B2353DEFB150D945BAB" box="[436,510,1172,1192]" italics="true" pageId="9" pageNumber="73">Gouania</emphasis>
|
||
species.
|
||
</taxonomicName>
|
||
(a)
|
||
<taxonomicName id="56F44D37FFB95B235007FB150EB95BAB" authority="Wagner & Kovac & Koblmüller, 2021" authorityName="Wagner & Kovac & Koblmüller" authorityYear="2021" box="[621,723,1172,1192]" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="9" pageNumber="73" phylum="Chordata" rank="species" species="adriatica" status="sp. nov.">
|
||
<emphasis id="A380EAA6FFB95B235007FB150EB95BAB" box="[621,723,1172,1192]" italics="true" pageId="9" pageNumber="73">G. adriatica</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A380EAA6FFB95B2350B3FB150F4B5BAB" bold="true" box="[729,801,1172,1192]" pageId="9" pageNumber="73">
|
||
<taxonomicNameLabel id="B8B357DDFFB95B2350B3FB150F4B5BAB" box="[729,801,1172,1192]" pageId="9" pageNumber="73" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(PMR VP4618
|
||
<emphasis id="A380EAA6FFB95B2351DAFB140FD65BAB" bold="true" box="[944,956,1173,1192]" pageId="9" pageNumber="73">–</emphasis>
|
||
Holotype), (b)
|
||
<taxonomicName id="56F44D37FFB95B23562EFB1508C65BAB" authority="Wagner & Kovac & Koblmüller, 2021" authorityName="Wagner & Kovac & Koblmüller" authorityYear="2021" box="[1092,1196,1172,1192]" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="9" pageNumber="73" phylum="Chordata" rank="species" species="orientalis" status="sp. nov.">
|
||
<emphasis id="A380EAA6FFB95B23562EFB1508C65BAB" box="[1092,1196,1172,1192]" italics="true" pageId="9" pageNumber="73">G. orientalis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A380EAA6FFB95B2356D8FB1508915BAB" bold="true" box="[1202,1275,1172,1192]" pageId="9" pageNumber="73">
|
||
<taxonomicNameLabel id="B8B357DDFFB95B2356D8FB1508915BAB" box="[1202,1275,1172,1192]" pageId="9" pageNumber="73" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(PMR VP4585
|
||
<emphasis id="A380EAA6FFB95B2357E3FB1409FF5BAB" bold="true" box="[1417,1429,1173,1192]" pageId="9" pageNumber="73">–</emphasis>
|
||
Holotype), (c)
|
||
<taxonomicName id="56F44D37FFB95B235292FB300D065BC6" authority="Wagner & Kovac & Koblmüller, 2021" authorityName="Wagner & Kovac & Koblmüller" authorityYear="2021" box="[248,364,1201,1221]" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="9" pageNumber="73" phylum="Chordata" rank="species" species="hofrichteri" status="sp. nov.">
|
||
<emphasis id="A380EAA6FFB95B235292FB300D065BC6" box="[248,364,1201,1221]" italics="true" pageId="9" pageNumber="73">G. hofrichteri</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A380EAA6FFB95B23531BFB300DD05BC6" bold="true" box="[369,442,1201,1221]" pageId="9" pageNumber="73">
|
||
<taxonomicNameLabel id="B8B357DDFFB95B23531BFB300DD05BC6" box="[369,442,1201,1221]" pageId="9" pageNumber="73" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(ZSM-PIS- 047656
|
||
<emphasis id="A380EAA6FFB95B23501BFB330E175BC6" bold="true" box="[625,637,1202,1221]" pageId="9" pageNumber="73">–</emphasis>
|
||
Paratype), (d)
|
||
<taxonomicName id="56F44D37FFB95B23516BFB300FAD5BC6" authority="(Nardo 1827)" baseAuthorityName="Nardo" baseAuthorityYear="1827" box="[769,967,1201,1221]" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="9" pageNumber="73" phylum="Chordata" rank="species" species="pigra">
|
||
<emphasis id="A380EAA6FFB95B23516BFB300F2C5BC6" box="[769,838,1201,1221]" italics="true" pageId="9" pageNumber="73">G. pigra</emphasis>
|
||
(Nardo 1827)
|
||
</taxonomicName>
|
||
(PMR VP3531
|
||
<emphasis id="A380EAA6FFB95B23563CFB3308085BC6" bold="true" box="[1110,1122,1202,1221]" pageId="9" pageNumber="73">–</emphasis>
|
||
Other material) and (e)
|
||
<taxonomicName id="56F44D37FFB95B235757FB300C835BE1" authority="(Risso 1810)" baseAuthorityName="Risso" baseAuthorityYear="1810" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="9" pageNumber="73" phylum="Chordata" rank="species" species="willdenowi">
|
||
<emphasis id="A380EAA6FFB95B235757FB3009D85BC6" box="[1341,1458,1201,1221]" italics="true" pageId="9" pageNumber="73">G. willdenowi</emphasis>
|
||
(
|
||
<bibRefCitation id="F5654B45FFB95B235216FB4F0C8E5BE1" author="Risso, A." box="[124,228,1230,1250]" pageId="9" pageNumber="73" refId="ref28299" refString="Risso, A. (1810). Ichthyologie de Nice, ou histoire naturelle des poissons du Departement des Alpes Maritimes. Paris, France: F. Schoell." type="book" year="1810">Risso 1810</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
(ZSM-PIS-0476654
|
||
<emphasis id="A380EAA6FFB95B2353C3FB4E0DDF5BE1" bold="true" box="[425,437,1231,1250]" pageId="9" pageNumber="73">–</emphasis>
|
||
Other material). Red, ceratobranchial 5; orange, premaxillary bone; blue, maxillary bone; green, nasal bone
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="914B36B4FFB95B23521DFAA80ED159DE" blockId="9.[119,769,1321,1965]" pageId="9" pageNumber="73">
|
||
2018;
|
||
<materialsCitation id="219C3CE9FFB95B2352DFFAA80EAB5A5C" collectingDate="2016-08-09" collectionCode="PMR" collectorName="M. Wagner" country="Greece" location="Plakias" pageId="9" pageNumber="73" specimenCode="VP4584" specimenCount="1" stateProvince="Kriti">
|
||
<collectionCode id="F7E5AE71FFB95B2352DFFAA80C895A3E" box="[181,227,1321,1341]" country="0" httpUri="http://grbio.org/cool/xq4z-q5gq" name="Prirodoslovni muzej Rijeka" pageId="9" pageNumber="73">PMR</collectionCode>
|
||
<specimenCode id="C1529ECFFFB95B235281FAA80D525A3E" box="[235,312,1321,1341]" collectionCode="VP" pageId="9" pageNumber="73">VP4584</specimenCode>
|
||
, female, 26.0 +
|
||
<quantity id="560C9B51FFB95B2353A4FAA80E495A3E" box="[462,547,1321,1341]" metricMagnitude="-3" metricUnit="m" metricValue="4.56" pageId="9" pageNumber="73" unit="mm" value="4.56">4.56 mm</quantity>
|
||
,
|
||
<location id="942B606FFFB95B235047FAA80E055A3E" LSID="urn:lsid:plazi:treatment:195D87A2FFB85B21515FF9DC0FF85C11:942B606FFFB95B235047FAA80E055A3E" box="[557,623,1321,1341]" country="Greece" name="Plakias" pageId="9" pageNumber="73" stateProvince="Kriti">Plakias</location>
|
||
,
|
||
<collectingRegion id="5330F856FFB95B235012FAA80EC45A3E" box="[632,686,1321,1341]" country="Greece" name="Kriti" pageId="9" pageNumber="73">Crete</collectingRegion>
|
||
,
|
||
<collectingCountry id="E9E37624FFB95B2350D2FAA80E975A3E" box="[696,765,1321,1341]" name="Greece" pageId="9" pageNumber="73">Greece</collectingCountry>
|
||
, 35 11
|
||
<emphasis id="A380EAA6FFB95B2352DAFAC80CDE5A55" bold="true" box="[176,180,1353,1366]" italics="true" pageId="9" pageNumber="73">
|
||
<superScript id="66819BFCFFB95B2352DAFAC80CDE5A55" attach="none" box="[176,180,1353,1366]" fontSize="6" pageId="9" pageNumber="73">0</superScript>
|
||
</emphasis>
|
||
40.8
|
||
<emphasis id="A380EAA6FFB95B2352B4FAC80C8C5A55" bold="true" box="[222,230,1353,1366]" italics="true" pageId="9" pageNumber="73">
|
||
<superScript id="66819BFCFFB95B2352B4FAC80C8C5A55" attach="right" box="[222,230,1353,1366]" fontSize="6" pageId="9" pageNumber="73">00</superScript>
|
||
</emphasis>
|
||
N, 24 22
|
||
<emphasis id="A380EAA6FFB95B235356FAC80D2A5A55" bold="true" box="[316,320,1353,1366]" italics="true" pageId="9" pageNumber="73">
|
||
<superScript id="66819BFCFFB95B235356FAC80D2A5A55" attach="none" box="[316,320,1353,1366]" fontSize="6" pageId="9" pageNumber="73">0</superScript>
|
||
</emphasis>
|
||
50.9
|
||
<emphasis id="A380EAA6FFB95B235300FAC80D185A55" bold="true" box="[362,370,1353,1366]" italics="true" pageId="9" pageNumber="73">
|
||
<superScript id="66819BFCFFB95B235300FAC80D185A55" attach="right" box="[362,370,1353,1366]" fontSize="6" pageId="9" pageNumber="73">00</superScript>
|
||
</emphasis>
|
||
E, coll.
|
||
<collectorName id="3C015362FFB95B2353D2FACA0E495A5C" box="[440,547,1355,1375]" pageId="9" pageNumber="73">M. Wagner</collectorName>
|
||
,
|
||
<date id="E54A1074FFB95B235047FACA0EAB5A5C" box="[557,705,1355,1375]" pageId="9" pageNumber="73" value="2016-08-09">
|
||
<collectingDate id="F50EE99CFFB95B235047FACA0EAB5A5C" box="[557,705,1355,1375]" pageId="9" pageNumber="73" value="2016-08-09">August 9, 2016</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
; ZSM- PIS-047658, female, 30.55 +
|
||
<quantity id="560C9B51FFB95B2353E2FAEF0DB05A81" box="[392,474,1390,1410]" metricMagnitude="-3" metricUnit="m" metricValue="5.25" pageId="9" pageNumber="73" unit="mm" value="5.25">5.25 mm</quantity>
|
||
and
|
||
<materialsCitation id="219C3CE9FFB95B235064FAEF0EA95AC4" collectingDate="2016-08-13" collectionCode="PMR" collectorName="M. Wagner" country="Greece" location="Vatos" pageId="9" pageNumber="73" specimenCode="VP4588" specimenCount="1" stateProvince="Kriti">
|
||
<collectionCode id="F7E5AE71FFB95B235064FAEF0E515A81" box="[526,571,1390,1410]" country="0" httpUri="http://grbio.org/cool/xq4z-q5gq" name="Prirodoslovni muzej Rijeka" pageId="9" pageNumber="73">PMR</collectionCode>
|
||
<specimenCode id="C1529ECFFFB95B23502EFAEF0EFB5A81" box="[580,657,1390,1410]" collectionCode="VP" pageId="9" pageNumber="73">VP4588</specimenCode>
|
||
, juvenile of unidentified sex, 17.92 +
|
||
<quantity id="560C9B51FFB95B235302FA100DD15AA6" box="[360,443,1425,1445]" metricMagnitude="-3" metricUnit="m" metricValue="2.98" pageId="9" pageNumber="73" unit="mm" value="2.98">2.98 mm</quantity>
|
||
, both from
|
||
<location id="942B606FFFB95B23505EFA100E005AA6" LSID="urn:lsid:plazi:treatment:195D87A2FFB85B21515FF9DC0FF85C11:942B606FFFB95B23505EFA100E005AA6" box="[564,618,1425,1445]" country="Greece" name="Vatos" pageId="9" pageNumber="73" stateProvince="Kriti">Vatos</location>
|
||
,
|
||
<collectingRegion id="5330F856FFB95B235012FA100EC65AA6" box="[632,684,1425,1445]" country="Greece" name="Kriti" pageId="9" pageNumber="73">Crete</collectingRegion>
|
||
,
|
||
<collectingCountry id="E9E37624FFB95B2350D0FA100E975AA6" box="[698,765,1425,1445]" name="Greece" pageId="9" pageNumber="73">Greece</collectingCountry>
|
||
, 34 59
|
||
<emphasis id="A380EAA6FFB95B2352DAFA300CDE5ABD" bold="true" box="[176,180,1457,1470]" italics="true" pageId="9" pageNumber="73">
|
||
<superScript id="66819BFCFFB95B2352DAFA300CDE5ABD" attach="none" box="[176,180,1457,1470]" fontSize="6" pageId="9" pageNumber="73">0</superScript>
|
||
</emphasis>
|
||
41.0
|
||
<emphasis id="A380EAA6FFB95B2352B4FA300C8C5ABD" bold="true" box="[222,230,1457,1470]" italics="true" pageId="9" pageNumber="73">
|
||
<superScript id="66819BFCFFB95B2352B4FA300C8C5ABD" attach="right" box="[222,230,1457,1470]" fontSize="6" pageId="9" pageNumber="73">00</superScript>
|
||
</emphasis>
|
||
N, 25 33
|
||
<emphasis id="A380EAA6FFB95B235351FA300D555ABD" bold="true" box="[315,319,1457,1470]" italics="true" pageId="9" pageNumber="73">
|
||
<superScript id="66819BFCFFB95B235351FA300D555ABD" attach="none" box="[315,319,1457,1470]" fontSize="6" pageId="9" pageNumber="73">0</superScript>
|
||
</emphasis>
|
||
17.3
|
||
<emphasis id="A380EAA6FFB95B235302FA300D1A5ABD" bold="true" box="[360,368,1457,1470]" italics="true" pageId="9" pageNumber="73">
|
||
<superScript id="66819BFCFFB95B235302FA300D1A5ABD" attach="right" box="[360,368,1457,1470]" fontSize="6" pageId="9" pageNumber="73">00</superScript>
|
||
</emphasis>
|
||
E, coll.
|
||
<collectorName id="3C015362FFB95B2353D9FA320E775AC4" box="[435,541,1459,1479]" pageId="9" pageNumber="73">M. Wagner</collectorName>
|
||
,
|
||
<date id="E54A1074FFB95B23504CFA320EA95AC4" box="[550,707,1459,1479]" pageId="9" pageNumber="73" value="2016-08-13">
|
||
<collectingDate id="F50EE99CFFB95B23504CFA320EA95AC4" box="[550,707,1459,1479]" pageId="9" pageNumber="73" value="2016-08-13">August 13, 2016</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
; ZSM- PIS-047659, female, 17.88 +
|
||
<quantity id="560C9B51FFB95B2353EAFA570DBB5AE9" box="[384,465,1494,1514]" metricMagnitude="-3" metricUnit="m" metricValue="2.79" pageId="9" pageNumber="73" unit="mm" value="2.79">2.79 mm</quantity>
|
||
and
|
||
<materialsCitation id="219C3CE9FFB95B235397FA570EA9592C" collectingDate="2016-08-17" collectionCode="PMR" collectorName="M. Wagner" country="Greece" location="Souda Beach" municipality="Plakias" pageId="9" pageNumber="73" specimenCode="VP4596" specimenCount="1" stateProvince="Kriti">
|
||
<collectionCode id="F7E5AE71FFB95B235397FA570E405AE9" box="[509,554,1494,1514]" country="0" httpUri="http://grbio.org/cool/xq4z-q5gq" name="Prirodoslovni muzej Rijeka" pageId="9" pageNumber="73">PMR</collectionCode>
|
||
<specimenCode id="C1529ECFFFB95B23505AFA570E115AE9" box="[560,635,1494,1514]" collectionCode="VP" pageId="9" pageNumber="73">VP4596</specimenCode>
|
||
, female, 17.09 +
|
||
<quantity id="560C9B51FFB95B2352E3FA780CB6590E" box="[137,220,1529,1549]" metricMagnitude="-3" metricUnit="m" metricValue="2.69" pageId="9" pageNumber="73" unit="mm" value="2.69">2.69 mm</quantity>
|
||
, both from
|
||
<location id="942B606FFFB95B23531FFA780D97590E" LSID="urn:lsid:plazi:treatment:195D87A2FFB85B21515FF9DC0FF85C11:942B606FFFB95B23531FFA780D97590E" box="[373,509,1529,1549]" country="Greece" municipality="Plakias" name="Souda Beach" pageId="9" pageNumber="73" stateProvince="Kriti">Souda Beach</location>
|
||
,
|
||
<collectingMunicipality id="712FACCEFFB95B23507FFA780E3F590E" box="[533,597,1529,1549]" pageId="9" pageNumber="73">Plakias</collectingMunicipality>
|
||
,
|
||
<collectingRegion id="5330F856FFB95B235007FA780ECB590E" box="[621,673,1529,1549]" country="Greece" name="Kriti" pageId="9" pageNumber="73">Crete</collectingRegion>
|
||
,
|
||
<collectingCountry id="E9E37624FFB95B2350D0FA780E97590E" box="[698,765,1529,1549]" name="Greece" pageId="9" pageNumber="73">Greece</collectingCountry>
|
||
, 35 11
|
||
<emphasis id="A380EAA6FFB95B2352DAF9980CDE5925" bold="true" box="[176,180,1561,1574]" italics="true" pageId="9" pageNumber="73">
|
||
<superScript id="66819BFCFFB95B2352DAF9980CDE5925" attach="none" box="[176,180,1561,1574]" fontSize="6" pageId="9" pageNumber="73">0</superScript>
|
||
</emphasis>
|
||
32.1
|
||
<emphasis id="A380EAA6FFB95B2352B4F9980C8C5925" bold="true" box="[222,230,1561,1574]" italics="true" pageId="9" pageNumber="73">
|
||
<superScript id="66819BFCFFB95B2352B4F9980C8C5925" attach="right" box="[222,230,1561,1574]" fontSize="6" pageId="9" pageNumber="73">00</superScript>
|
||
</emphasis>
|
||
N, 24 22
|
||
<emphasis id="A380EAA6FFB95B235351F9980D555925" bold="true" box="[315,319,1561,1574]" italics="true" pageId="9" pageNumber="73">
|
||
<superScript id="66819BFCFFB95B235351F9980D555925" attach="none" box="[315,319,1561,1574]" fontSize="6" pageId="9" pageNumber="73">0</superScript>
|
||
</emphasis>
|
||
04.9
|
||
<emphasis id="A380EAA6FFB95B235302F9980D1A5925" bold="true" box="[360,368,1561,1574]" italics="true" pageId="9" pageNumber="73">
|
||
<superScript id="66819BFCFFB95B235302F9980D1A5925" attach="right" box="[360,368,1561,1574]" fontSize="6" pageId="9" pageNumber="73">00</superScript>
|
||
</emphasis>
|
||
E, coll.
|
||
<collectorName id="3C015362FFB95B2353D9F99A0E77592C" box="[435,541,1563,1583]" pageId="9" pageNumber="73">M. Wagner</collectorName>
|
||
,
|
||
<date id="E54A1074FFB95B23504CF99A0EA9592C" box="[550,707,1563,1583]" pageId="9" pageNumber="73" value="2016-08-17">
|
||
<collectingDate id="F50EE99CFFB95B23504CF99A0EA9592C" box="[550,707,1563,1583]" pageId="9" pageNumber="73" value="2016-08-17">August 17, 2016</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="219C3CE9FFB95B2350A6F99A0E255994" collectingDate="2018-08-16" collectorName="M. Wagner" country="Greece" location="Chamolia" pageId="9" pageNumber="73" specimenCount="1">
|
||
ZSM- PIS-047660, female, 24.87 +
|
||
<quantity id="560C9B51FFB95B2353E0F9BF0DB05951" box="[394,474,1598,1618]" metricMagnitude="-3" metricUnit="m" metricValue="3.56" pageId="9" pageNumber="73" unit="mm" value="3.56">3.56 mm</quantity>
|
||
and ZSM-PIS-047660, female, 23.68 + 3.39, mm, both from
|
||
<location id="942B606FFFB95B2353D2F9E00E645976" LSID="urn:lsid:plazi:treatment:195D87A2FFB85B21515FF9DC0FF85C11:942B606FFFB95B2353D2F9E00E645976" box="[440,526,1633,1653]" country="Greece" name="Chamolia" pageId="9" pageNumber="73">Chamolia</location>
|
||
,
|
||
<collectingCountry id="E9E37624FFB95B235049F9E00E0C5976" box="[547,614,1633,1653]" name="Greece" pageId="9" pageNumber="73">Greece</collectingCountry>
|
||
, 37 54
|
||
<emphasis id="A380EAA6FFB95B2350DEF9DF0ED25968" bold="true" box="[692,696,1630,1643]" italics="true" pageId="9" pageNumber="73">
|
||
<superScript id="66819BFCFFB95B2350DEF9DF0ED25968" attach="none" box="[692,696,1630,1643]" fontSize="6" pageId="9" pageNumber="73">0</superScript>
|
||
</emphasis>
|
||
58.5
|
||
<emphasis id="A380EAA6FFB95B23508BF9DF0E835968" bold="true" box="[737,745,1630,1643]" italics="true" pageId="9" pageNumber="73">
|
||
<superScript id="66819BFCFFB95B23508BF9DF0E835968" attach="right" box="[737,745,1630,1643]" fontSize="6" pageId="9" pageNumber="73">00</superScript>
|
||
</emphasis>
|
||
N, 24 02
|
||
<emphasis id="A380EAA6FFB95B2352DAF9000CDE598D" bold="true" box="[176,180,1665,1678]" italics="true" pageId="9" pageNumber="73">
|
||
<superScript id="66819BFCFFB95B2352DAF9000CDE598D" attach="none" box="[176,180,1665,1678]" fontSize="6" pageId="9" pageNumber="73">0</superScript>
|
||
</emphasis>
|
||
08.7
|
||
<emphasis id="A380EAA6FFB95B2352B4F9000C8C598D" bold="true" box="[222,230,1665,1678]" italics="true" pageId="9" pageNumber="73">
|
||
<superScript id="66819BFCFFB95B2352B4F9000C8C598D" attach="right" box="[222,230,1665,1678]" fontSize="6" pageId="9" pageNumber="73">00</superScript>
|
||
</emphasis>
|
||
E, coll.
|
||
<collectorName id="3C015362FFB95B23535AF9020DF45994" box="[304,414,1667,1687]" pageId="9" pageNumber="73">M. Wagner</collectorName>
|
||
,
|
||
<date id="E54A1074FFB95B2353C0F9020E255994" box="[426,591,1667,1687]" pageId="9" pageNumber="73" value="2018-08-16">
|
||
<collectingDate id="F50EE99CFFB95B2353C0F9020E255994" box="[426,591,1667,1687]" pageId="9" pageNumber="73" value="2018-08-16">August 16, 2018</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="219C3CE9FFB95B235036F9020EDD59DE" collectingDate="2018-08-21" collectorName="M. Wagner" country="Greece" county="Kythira" location="Feloti Beach" municipality="Kapsali" pageId="9" pageNumber="73" specimenCount="1">
|
||
ZSM-PIS-047661, male, 28.52 +
|
||
<quantity id="560C9B51FFB95B23536DF9270D3359B9" box="[263,345,1702,1722]" metricMagnitude="-3" metricUnit="m" metricValue="4.04" pageId="9" pageNumber="73" unit="mm" value="4.04">4.04 mm</quantity>
|
||
,
|
||
<location id="942B606FFFB95B23531AF9270D9B59B9" LSID="urn:lsid:plazi:treatment:195D87A2FFB85B21515FF9DC0FF85C11:942B606FFFB95B23531AF9270D9B59B9" box="[368,497,1702,1722]" country="Greece" county="Kythira" municipality="Kapsali" name="Feloti Beach" pageId="9" pageNumber="73">Feloti Beach</location>
|
||
,
|
||
<collectingMunicipality id="712FACCEFFB95B23506DF9270E2359B9" box="[519,585,1702,1722]" pageId="9" pageNumber="73">Kapsáli</collectingMunicipality>
|
||
,
|
||
<collectingCounty id="782A4E38FFB95B23500AF9270ECE59B9" box="[608,676,1702,1722]" pageId="9" pageNumber="73">Kythira</collectingCounty>
|
||
,
|
||
<collectingCountry id="E9E37624FFB95B2350D0F9270E9759B9" box="[698,765,1702,1722]" name="Greece" pageId="9" pageNumber="73">Greece</collectingCountry>
|
||
, 36 09
|
||
<emphasis id="A380EAA6FFB95B2352DAF9470CDE59D0" bold="true" box="[176,180,1734,1747]" italics="true" pageId="9" pageNumber="73">
|
||
<superScript id="66819BFCFFB95B2352DAF9470CDE59D0" attach="none" box="[176,180,1734,1747]" fontSize="6" pageId="9" pageNumber="73">0</superScript>
|
||
</emphasis>
|
||
16.2
|
||
<emphasis id="A380EAA6FFB95B2352B4F9470C8C59D0" bold="true" box="[222,230,1734,1747]" italics="true" pageId="9" pageNumber="73">
|
||
<superScript id="66819BFCFFB95B2352B4F9470C8C59D0" attach="right" box="[222,230,1734,1747]" fontSize="6" pageId="9" pageNumber="73">00</superScript>
|
||
</emphasis>
|
||
N, 22 57
|
||
<emphasis id="A380EAA6FFB95B235353F9470D5759D0" bold="true" box="[313,317,1734,1747]" italics="true" pageId="9" pageNumber="73">
|
||
<superScript id="66819BFCFFB95B235353F9470D5759D0" attach="none" box="[313,317,1734,1747]" fontSize="6" pageId="9" pageNumber="73">0</superScript>
|
||
</emphasis>
|
||
52.0
|
||
<emphasis id="A380EAA6FFB95B23530DF9470D0559D0" bold="true" box="[359,367,1734,1747]" italics="true" pageId="9" pageNumber="73">
|
||
<superScript id="66819BFCFFB95B23530DF9470D0559D0" attach="right" box="[359,367,1734,1747]" fontSize="6" pageId="9" pageNumber="73">00</superScript>
|
||
</emphasis>
|
||
E, coll.
|
||
<collectorName id="3C015362FFB95B2353C4F9480E7D59DE" box="[430,535,1737,1757]" pageId="9" pageNumber="73">M. Wagner</collectorName>
|
||
,
|
||
<date id="E54A1074FFB95B235074F9480EDD59DE" box="[542,695,1737,1757]" pageId="9" pageNumber="73" value="2018-08-21">
|
||
<collectingDate id="F50EE99CFFB95B235074F9480EDD59DE" box="[542,695,1737,1757]" pageId="9" pageNumber="73" value="2018-08-21">August 21, 2018</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D9EE653FFFB95B2352F5F96A09145864" pageId="9" pageNumber="73" type="diagnosis">
|
||
<paragraph id="914B36B4FFB95B2352F5F96A09145864" blockId="9.[119,769,1321,1965]" lastBlockId="9.[816,1465,1321,1965]" pageId="9" pageNumber="73">
|
||
<emphasis id="A380EAA6FFB95B2352F5F96A0D6B59FC" bold="true" box="[159,257,1771,1791]" pageId="9" pageNumber="73">Diagnosis.</emphasis>
|
||
<taxonomicName id="56F44D37FFB95B235362F96A0DC159FC" authority="Wagner & Kovac & Koblmüller, 2021" authorityName="Wagner & Kovac & Koblmüller" authorityYear="2021" box="[264,427,1771,1791]" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="9" pageNumber="73" phylum="Chordata" rank="species" species="orientalis" status="sp. nov.">
|
||
<emphasis id="A380EAA6FFB95B235362F96A0DC159FC" box="[264,427,1771,1791]" italics="true" pageId="9" pageNumber="73">Gouania orientalis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A380EAA6FFB95B2353DEF96A0E6A59FC" bold="true" box="[436,512,1771,1791]" pageId="9" pageNumber="73">
|
||
<taxonomicNameLabel id="B8B357DDFFB95B2353DEF96A0E6A59FC" box="[436,512,1771,1791]" pageId="9" pageNumber="73" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs from its congeners by the combination of the following characters: (1) dorsal head profile a straight line from nape above eye to upper lip tip; (2) posterior angle of jaws extends to between a vertical line drawn through anterior edge of eye and a vertical line drawn through anterior part of eye; (3) infraorbital invagination vertical to posterior part of eye; (4) posterior opercular edge w-shaped with two equally long tips; (5) longitudinal infralateral and suborbital transversal rows of superficial neuromasts placed in the well-defined deep groove; (6) trunk cross-section behind pectoral fin base half oval with straight ventral side; (7) granules on body shallow and inconspicuous; (8) upper attachment of gill membrane opposite to 5th to 6th pectoral ray; (9) pectoral rays 17
|
||
<emphasis id="A380EAA6FFB95B2357E7FA5609F35AE9" bold="true" box="[1421,1433,1495,1514]" pageId="9" pageNumber="73">–</emphasis>
|
||
19; (10) upper attachment of disc membrane attaching to base of pectoral fin at 16th
|
||
<emphasis id="A380EAA6FFB95B2351FDF99D0FC9592C" bold="true" box="[919,931,1564,1583]" pageId="9" pageNumber="73">–</emphasis>
|
||
18th pectoral ray; (11) principal caudal rays 10
|
||
<emphasis id="A380EAA6FFB95B235708F99D0904592C" bold="true" box="[1378,1390,1564,1583]" pageId="9" pageNumber="73">–</emphasis>
|
||
11; (12) head length 25.0
|
||
<emphasis id="A380EAA6FFB95B2351BBF9BE0FB75951" bold="true" box="[977,989,1599,1618]" pageId="9" pageNumber="73">–</emphasis>
|
||
28.9% of standard length; (13) pectoral fin length 8.2
|
||
<emphasis id="A380EAA6FFB95B235127F9E00F335977" bold="true" box="[845,857,1633,1652]" pageId="9" pageNumber="73">–</emphasis>
|
||
10.3% of standard length; (14) prepectoral distance 24.7
|
||
<emphasis id="A380EAA6FFB95B23571BF9E009175977" bold="true" box="[1393,1405,1633,1652]" pageId="9" pageNumber="73">–</emphasis>
|
||
28.2% of standard length; (15) ventral adhesive disc length 14.9
|
||
<emphasis id="A380EAA6FFB95B23573FF905090B5994" bold="true" box="[1365,1377,1668,1687]" pageId="9" pageNumber="73">–</emphasis>
|
||
19.0% of standard length; (16) caudal-fin length 14.2
|
||
<emphasis id="A380EAA6FFB95B2356D7F92608A359B9" bold="true" box="[1213,1225,1703,1722]" pageId="9" pageNumber="73">–</emphasis>
|
||
17.5% of standard length; (17) low number of vertebrae (Supporting Information Table S2; 35
|
||
<emphasis id="A380EAA6FFB95B235122F96D0F3E59FC" bold="true" box="[840,852,1772,1791]" pageId="9" pageNumber="73">–</emphasis>
|
||
36) (18) pharyngeal jaws with small ceratobranchial 5, having a few hardly recognizable small conical teeth; (19) nasal bones with inconspicuous frontal end; (20) in life star-like pigmentation around eyes, body colouration dark, sometimes marbled or with stripes.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D9EE653FFFB95B215132F8F70E815EFE" lastPageId="11" lastPageNumber="75" pageId="9" pageNumber="73" type="description">
|
||
<paragraph id="914B36B4FFB95B205132F8F708385C54" blockId="9.[816,1465,1321,1965]" lastBlockId="10.[821,1470,142,1965]" lastPageId="10" lastPageNumber="74" pageId="9" pageNumber="73">
|
||
<emphasis id="A380EAA6FFB95B235132F8F70FA15889" bold="true" box="[856,971,1910,1930]" pageId="9" pageNumber="73">Description.</emphasis>
|
||
<emphasis id="A380EAA6FFB95B2351B9F8F708E35889" box="[979,1161,1910,1930]" italics="true" pageId="9" pageNumber="73">General morphology</emphasis>
|
||
: Body proportions are given in Table 1. Body slender and elongated, posteriorly laterally compressed, body depth at pectoral fins
|
||
<quantity id="560C9B51FFBA5B205314FC2A0DB15CBC" box="[382,475,939,959]" metricMagnitude="-1" metricUnit="m" metricValue="1.9685" metricValueMax="2.1843999999999997" metricValueMin="1.7526" pageId="10" pageNumber="74" unit="in" value="7.75" valueMax="8.6" valueMin="6.9">
|
||
6.9
|
||
<emphasis id="A380EAA6FFBA5B2053F1FC2D0DCD5CBC" bold="true" box="[411,423,940,959]" pageId="10" pageNumber="74">–</emphasis>
|
||
8.6 in
|
||
</quantity>
|
||
SL, body depth at anus
|
||
<quantity id="560C9B51FFBA5B2050D7FC2A0CE75CE1" metricMagnitude="-1" metricUnit="m" metricValue="2.1717" metricValueMax="2.3622" metricValueMin="1.9812" pageId="10" pageNumber="74" unit="in" value="8.55" valueMax="9.3" valueMin="7.8">
|
||
7.8
|
||
<emphasis id="A380EAA6FFBA5B2050B0FC2D0E8C5CBC" bold="true" box="[730,742,940,959]" pageId="10" pageNumber="74">–</emphasis>
|
||
9.3 in
|
||
</quantity>
|
||
SL, body depth in width at pectoral fins 1.1
|
||
<emphasis id="A380EAA6FFBA5B205029FC4E0E255CE1" bold="true" box="[579,591,975,994]" pageId="10" pageNumber="74">–</emphasis>
|
||
1.2, body depth in width at anus 0.8
|
||
<emphasis id="A380EAA6FFBA5B20534FFC700D5B5B07" bold="true" box="[293,305,1009,1028]" pageId="10" pageNumber="74">–</emphasis>
|
||
0.9. Trunk cross-section behind pectoral fin base half oval with straight ventral side. Granules on body shallow and inconspicuous. Head dorsoventrally compressed, head depth in width at orbit 1.5
|
||
<emphasis id="A380EAA6FFBA5B205289FBD80C855B6F" bold="true" box="[227,239,1113,1132]" pageId="10" pageNumber="74">–</emphasis>
|
||
1.9 and moderately large, head length 3.5
|
||
<emphasis id="A380EAA6FFBA5B205005FBD80E115B6F" bold="true" box="[623,635,1113,1132]" pageId="10" pageNumber="74">–</emphasis>
|
||
4.0 in SL, head wider than body width maximum, head width at anterior sucking disc edge
|
||
<quantity id="560C9B51FFBA5B2052DFFB1F0D735BB1" box="[181,281,1182,1202]" metricMagnitude="-2" metricUnit="m" metricValue="1.905" metricValueMax="2.032" metricValueMin="1.7779999999999998" pageId="10" pageNumber="74" unit="in" value="0.75" valueMax="0.8" valueMin="0.7">
|
||
0.7
|
||
<emphasis id="A380EAA6FFBA5B2052B8FB1E0CB45BB1" bold="true" box="[210,222,1183,1202]" pageId="10" pageNumber="74">–</emphasis>
|
||
0.8 in
|
||
</quantity>
|
||
body width at pectoral fins. Dorsal head profile straight between nape above eye and upper lip tip. Head rounded in dorsal view. Snout large compared to eyes, preorbital distance
|
||
<quantity id="560C9B51FFBA5B205216FA870CB55A19" box="[124,223,1286,1306]" metricMagnitude="-2" metricUnit="m" metricValue="8.763" metricValueMax="9.652" metricValueMin="7.874" pageId="10" pageNumber="74" unit="in" value="3.45" valueMax="3.8" valueMin="3.1">
|
||
3.1
|
||
<emphasis id="A380EAA6FFBA5B2052F3FA860CCF5A19" bold="true" box="[153,165,1287,1306]" pageId="10" pageNumber="74">–</emphasis>
|
||
3.8 in
|
||
</quantity>
|
||
head length,
|
||
<quantity id="560C9B51FFBA5B205301FA870DA75A19" box="[363,461,1286,1306]" metricMagnitude="-2" metricUnit="m" metricValue="1.016" metricValueMax="1.27" metricValueMin="0.762" pageId="10" pageNumber="74" unit="in" value="0.4" valueMax="0.5" valueMin="0.3">
|
||
0.3
|
||
<emphasis id="A380EAA6FFBA5B2053E2FA860DFE5A19" bold="true" box="[392,404,1287,1306]" pageId="10" pageNumber="74">–</emphasis>
|
||
0.5 in
|
||
</quantity>
|
||
horizontal eye diameter. Snout wide, not produced, blunt. Internostril space gently convex. Eyes dorsolateral, with lower eye edge rounded. Eyes small, 7.0
|
||
<emphasis id="A380EAA6FFBA5B2050EBFACD0EE75A5C" bold="true" box="[641,653,1356,1375]" pageId="10" pageNumber="74">–</emphasis>
|
||
<quantity id="560C9B51FFBA5B2050E4FACA0EA55A5C" box="[654,719,1355,1375]" metricMagnitude="-1" metricUnit="m" metricValue="2.5654" pageId="10" pageNumber="74" unit="in" value="10.1">10.1 in</quantity>
|
||
head length, vertical diameter of the eye
|
||
<quantity id="560C9B51FFBA5B2053A8FAEF0E755A81" box="[450,543,1390,1410]" metricMagnitude="-2" metricUnit="m" metricValue="2.413" metricValueMax="2.794" metricValueMin="2.032" pageId="10" pageNumber="74" unit="in" value="0.95" valueMax="1.1" valueMin="0.8">
|
||
0.8
|
||
<emphasis id="A380EAA6FFBA5B20538AFAEE0D865A81" bold="true" box="[480,492,1391,1410]" pageId="10" pageNumber="74">–</emphasis>
|
||
1.1 in
|
||
</quantity>
|
||
horizontal eye diameter. Infraorbital invagination vertical to posterior part of eye. Interorbital distance wide,
|
||
<quantity id="560C9B51FFBA5B20536FFA320D085AC4" box="[261,354,1459,1479]" metricMagnitude="-2" metricUnit="m" metricValue="1.016" metricValueMax="1.27" metricValueMin="0.762" pageId="10" pageNumber="74" unit="in" value="0.4" valueMax="0.5" valueMin="0.3">
|
||
0.3
|
||
<emphasis id="A380EAA6FFBA5B205348FA350D445AC4" bold="true" box="[290,302,1460,1479]" pageId="10" pageNumber="74">–</emphasis>
|
||
0.5 in
|
||
</quantity>
|
||
horizontal eye diameter.
|
||
<collectingRegion id="5330F856FFBA5B205026FA320EE15AC4" box="[588,651,1459,1479]" country="France" name="Centre" pageId="10" pageNumber="74">Centre</collectingRegion>
|
||
of eye much closer to tip of snout than to posterior margin of operculum, preorbital distance in postorbital distance 1.9
|
||
<emphasis id="A380EAA6FFBA5B20538FFA780D9B590F" bold="true" box="[485,497,1529,1548]" pageId="10" pageNumber="74">–</emphasis>
|
||
2.3. Anterior and posterior nostrils long tubes of about equal length. Nostrils well separated and posterior nostril located behind and dorsally to anterior edge of eyes. Single large dermal flap at the posterior margin of anterior nostril leaf shaped, longer than nostril. Posterior nostril rim slightly crenate with no extension. Head lateral line system with canals with pores and with superficial neuromasts arranged in rows. Head canals reduced and pores small. Single pore in nasal canal near posterior nostril. Single pore in postorbital canal close to posterior eye edge. Two pores in mandibular canal, anterior one close to anteriolateral angle of mouth, posterior pore slightly in front of vertical of posterior angle of jaws, posterior pore usually more prominent. Lachrymal as well as preopercular canals and pores absent. Rows of superficial neuromasts as follows: SR 2, NR 3
|
||
<emphasis id="A380EAA6FFBA5B20566FFF0E087B5FA1" bold="true" box="[1029,1041,143,162]" pageId="10" pageNumber="74">–</emphasis>
|
||
5, LIR 23
|
||
<emphasis id="A380EAA6FFBA5B20560CFF0E08185FA1" bold="true" box="[1126,1138,143,162]" pageId="10" pageNumber="74">–</emphasis>
|
||
28, STR 3
|
||
<emphasis id="A380EAA6FFBA5B2056A4FF0E08B05FA1" bold="true" box="[1230,1242,143,162]" pageId="10" pageNumber="74">–</emphasis>
|
||
4, POR 3
|
||
<emphasis id="A380EAA6FFBA5B20575AFF0E09565FA1" bold="true" box="[1328,1340,143,162]" pageId="10" pageNumber="74">–</emphasis>
|
||
4, PTR 2, SLR 5, MR 9
|
||
<emphasis id="A380EAA6FFBA5B2051EAFF300FE65FC7" bold="true" box="[896,908,177,196]" pageId="10" pageNumber="74">–</emphasis>
|
||
13, AVR 2, PVR 1, ADR 1
|
||
<emphasis id="A380EAA6FFBA5B205616FF3008E25FC7" bold="true" box="[1148,1160,177,196]" pageId="10" pageNumber="74">–</emphasis>
|
||
3, PDR 1, HR 3, SR1 3, SR2 1
|
||
<emphasis id="A380EAA6FFBA5B2057F5FF3009C15FC7" bold="true" box="[1439,1451,177,196]" pageId="10" pageNumber="74">–</emphasis>
|
||
2, DLR 6
|
||
<emphasis id="A380EAA6FFBA5B20511EFF550FEA5FE4" bold="true" box="[884,896,212,231]" pageId="10" pageNumber="74">–</emphasis>
|
||
8, VLR 9
|
||
<emphasis id="A380EAA6FFBA5B2051B3FF550F8F5FE4" bold="true" box="[985,997,212,231]" pageId="10" pageNumber="74">–</emphasis>
|
||
13. STR and LIR rows of superficial neuromasts placed in the well-defined deep groove. DLR row of superficial neuromasts anteriorly starts above pectoral fin, continuously dorsolateral and ends posteriorly downwards at midlateral level and variably, vertical to anus or in front of it or behind it. VLR anteriorly starts behind pectoral fin base, continuous ventrolaterally and ends posteriorly upwards with last papilla nearly at midlateral level at caudal fin base or close to it. Mouth terminal, upper and lower lips end about equally, lips fleshy, upper lip larger than the lower lip. Posterior angle of jaws extends to between vertical line drawn through anterior edge of eyes and vertical line drawn through anterior part of eye. Chin with bilobed or slightly bilobed fold at anterior edge covering MR row of superficial neuromasts. The gill membrane is attached to isthmus, gill opening starting at the base of pectoral fin, with the upper attachment of the gill membrane is opposite to 5th to 6th pectoral ray. Posterior opercular edge w-shaped with two equally long tips. No subopercular spine. No fleshy pad present on lower pectoral base. Urogenital papilla present. Preanus length in postanus length 0.6
|
||
<emphasis id="A380EAA6FFBA5B2056A2FCA008BE5C37" bold="true" box="[1224,1236,801,820]" pageId="10" pageNumber="74">–</emphasis>
|
||
0.8. Anal papillae absent, the area around anus wrinkled.
|
||
</paragraph>
|
||
<caption id="C58B663CFFBA5B205216FD680EE15C57" ID-DOI="http://doi.org/10.5281/zenodo.10928936" ID-Zenodo-Dep="10928936" httpUri="https://zenodo.org/record/10928936/files/figure.png" pageId="10" pageNumber="74" startId="10.[124,136,745,764]" targetBox="[129,769,143,698]" targetPageId="10" targetType="figure">
|
||
<paragraph id="914B36B4FFBA5B205216FD680EE15C57" blockId="10.[124,764,744,852]" pageId="10" pageNumber="74">
|
||
<emphasis id="A380EAA6FFBA5B205216FD680C865DFF" bold="true" box="[124,236,745,764]" pageId="10" pageNumber="74">F I G U R E 6</emphasis>
|
||
<taxonomicName id="56F44D37FFBA5B205360FD690DC35DFF" authority="Wagner & Kovac & Koblmüller, 2021" authorityName="Wagner & Kovac & Koblmüller" authorityYear="2021" box="[266,425,744,764]" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="10" pageNumber="74" phylum="Chordata" rank="species" species="orientalis" status="sp. nov.">
|
||
<emphasis id="A380EAA6FFBA5B205360FD690DC35DFF" box="[266,425,744,764]" italics="true" pageId="10" pageNumber="74">Gouania orientalis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A380EAA6FFBA5B2053C5FD690D9D5DFF" bold="true" box="[431,503,744,764]" pageId="10" pageNumber="74">
|
||
<taxonomicNameLabel id="B8B357DDFFBA5B2053C5FD690D9D5DFF" box="[431,503,744,764]" pageId="10" pageNumber="74" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, PMR VP4585, holotype, male, 32.8+4.93 mm, Plakias, Crete, Greece. Lateral view of specimen preserved in 4% formaldehyde (top). Lateral, dorsal and ventral view, alive (below). Photographs by M. Wagner and M. Kovacˇic
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="914B36B4FFBA5B205136FCE708B65A19" blockId="10.[821,1470,142,1965]" pageId="10" pageNumber="74">
|
||
<emphasis id="A380EAA6FFBA5B205136FCE70F155C79" box="[860,895,870,890]" italics="true" pageId="10" pageNumber="74">Fins</emphasis>
|
||
. Rudimentary dorsal and anal fins located well posteriorly and short, reduced to low ridges with very weak rays, connected to the caudal fin. Pectoral rays 17
|
||
<emphasis id="A380EAA6FFBA5B20518FFC2D0F9B5CBC" bold="true" box="[997,1009,940,959]" pageId="10" pageNumber="74">–</emphasis>
|
||
19. Caudal fin rounded, principal caudal rays 10
|
||
<emphasis id="A380EAA6FFBA5B2057FEFC2D09CA5CBC" bold="true" box="[1428,1440,940,959]" pageId="10" pageNumber="74">–</emphasis>
|
||
11. Ventral adhesive disc (Figure 4b) of
|
||
<emphasis id="A380EAA6FFBA5B205604FC4E081F5CE1" bold="true" box="[1134,1141,975,994]" pageId="10" pageNumber="74">“</emphasis>
|
||
double
|
||
<emphasis id="A380EAA6FFBA5B2056DBFC4E08D35CE1" bold="true" box="[1201,1209,975,994]" pageId="10" pageNumber="74">”</emphasis>
|
||
<typeStatus id="4E4F8816FFBA5B2056D4FC4F088D5CE1" box="[1214,1255,974,994]" pageId="10" pageNumber="74">type</typeStatus>
|
||
, anterior margin crenate with large invagination on each lateral side and central invagination at midventral; posterior margin slightly crenate. Disc small, disc length
|
||
<quantity id="560C9B51FFBA5B2057FEFB920F0D5B49" metricMagnitude="-1" metricUnit="m" metricValue="1.524" metricValueMax="1.7018" metricValueMin="1.3462" pageId="10" pageNumber="74" unit="in" value="6.0" valueMax="6.7" valueMin="5.3">
|
||
5.3
|
||
<emphasis id="A380EAA6FFBA5B2057DAFB9509D65B24" bold="true" box="[1456,1468,1044,1063]" pageId="10" pageNumber="74">–</emphasis>
|
||
6.7 in
|
||
</quantity>
|
||
SL, its width slightly larger than its length, width in length 0.9
|
||
<emphasis id="A380EAA6FFBA5B2057FAFBB609F65B49" bold="true" box="[1424,1436,1079,1098]" pageId="10" pageNumber="74">–</emphasis>
|
||
1.1. No papillae in region A and flattened papillae in regions B and C. In region B two to three rows of papillae with total papillae count 21
|
||
<emphasis id="A380EAA6FFBA5B205732FBFD090E5B8C" bold="true" box="[1368,1380,1148,1167]" pageId="10" pageNumber="74">–</emphasis>
|
||
37 and in region C two rows of papillae with total papillae count 9
|
||
<emphasis id="A380EAA6FFBA5B205756FB1E09225BB1" bold="true" box="[1340,1352,1183,1202]" pageId="10" pageNumber="74">–</emphasis>
|
||
15. No inner row of papillae on lateral sides of the central part of the anterior disc. Upper attachment of disc membrane attaching to base of pectoral fin at 16th
|
||
<emphasis id="A380EAA6FFBA5B205134FA860F005A19" bold="true" box="[862,874,1287,1306]" pageId="10" pageNumber="74">–</emphasis>
|
||
18th pectoral ray (
|
||
<emphasis id="A380EAA6FFBA5B205661FA8708775A19" box="[1035,1053,1286,1306]" italics="true" pageId="10" pageNumber="74">i.e</emphasis>
|
||
., on penultimate ray).
|
||
</paragraph>
|
||
<paragraph id="914B36B4FFBA5B205137FAA8083F5994" blockId="10.[821,1470,142,1965]" pageId="10" pageNumber="74">
|
||
<emphasis id="A380EAA6FFBA5B205137FAA80FAE5A3E" box="[861,964,1321,1341]" italics="true" pageId="10" pageNumber="74">Colouration</emphasis>
|
||
. Background colour of live specimens bright yellow to brownish red (Figure 6) and prominent star-like pigmentation around eyes present. Body pigments reduced (juveniles) or, behind head, with clearly visible regular stripes (juveniles,
|
||
<emphasis id="A380EAA6FFBA5B2056C0FA1008A95AA6" box="[1194,1219,1425,1445]" italics="true" pageId="10" pageNumber="74">e.g</emphasis>
|
||
.,
|
||
<collectingRegion id="5330F856FFBA5B2056BEFA1009675AA6" box="[1236,1293,1425,1445]" country="Greece" name="Attiki" pageId="10" pageNumber="74">Attica</collectingRegion>
|
||
,
|
||
<collectingRegion id="5330F856FFBA5B205772FA1009245AA6" box="[1304,1358,1425,1445]" country="Greece" name="Kriti" pageId="10" pageNumber="74">Crete</collectingRegion>
|
||
, Kythira) or marbled (
|
||
<emphasis id="A380EAA6FFBA5B2051E7FA320FCA5AC4" box="[909,928,1459,1479]" italics="true" pageId="10" pageNumber="74">i.e</emphasis>
|
||
., irregular pattern;
|
||
<emphasis id="A380EAA6FFBA5B205632FA32081B5AC4" box="[1112,1137,1459,1479]" italics="true" pageId="10" pageNumber="74">e.g</emphasis>
|
||
.,
|
||
<collectingRegion id="5330F856FFBA5B2056E8FA3208C05AC4" box="[1154,1194,1459,1479]" country="Papua New Guinea" name="Gulf" pageId="10" pageNumber="74">Gulf</collectingRegion>
|
||
of Corinth). Specimens from
|
||
<collectingRegion id="5330F856FFBA5B20515FFA570F015AE9" box="[821,875,1494,1514]" country="Greece" name="Kriti" pageId="10" pageNumber="74">Crete</collectingRegion>
|
||
, Kythira and the
|
||
<collectingRegion id="5330F856FFBA5B20567DFA5708555AE9" box="[1047,1087,1494,1514]" country="Papua New Guinea" name="Gulf" pageId="10" pageNumber="74">Gulf</collectingRegion>
|
||
of Corinth have stronger pigmentation (see Figure 6), hence, the pigmentation pattern can be less clearly visible. Formaldehyde fixed specimens white-yellow and without pigments. Ethanol fixed specimens white to skin-coloured, pigmentation present (also stripes). For more pictures of life colouration see Supporting Information File S1.
|
||
</paragraph>
|
||
<paragraph id="914B36B4FFBA5B215137F9270E815EFE" blockId="10.[821,1470,142,1965]" lastBlockId="11.[119,769,142,1965]" lastPageId="11" lastPageNumber="75" pageId="10" pageNumber="74">
|
||
<emphasis id="A380EAA6FFBA5B205137F927084159B9" box="[861,1067,1702,1722]" italics="true" pageId="10" pageNumber="74">Dentition and osteology</emphasis>
|
||
. Upper jaw with outer row of about 10 (one side) medium-sized caniniforms frontally. Behind them inner small conical teeth irregularly scattered in two separate (left and right) drop-like patches medially wide about 5
|
||
<emphasis id="A380EAA6FFBA5B205642F88E085E5821" bold="true" box="[1064,1076,1807,1826]" pageId="10" pageNumber="74">–</emphasis>
|
||
6 teeth, becoming narrowed to a single row of teeth laterally. Outer row continues laterally as two large caniniforms, followed behind by about eight medium-sized caniniforms. Lower jaw with outer row of about 10 (one side) medium-sized caniniforms frontally. Behind them single broad patch of small conical inner teeth wide medially about 5
|
||
<emphasis id="A380EAA6FFBB5B2152D4FF0E0CA05FA1" bold="true" box="[190,202,143,162]" pageId="11" pageNumber="75">–</emphasis>
|
||
6 teeth, becoming narrowed to a single row of teeth laterally. The single row of about six larger caniniforms continuous laterally. Pharyngeal jaws with small ceratobranchial 5, having a few (1
|
||
<emphasis id="A380EAA6FFBB5B215010FF550EEC5FE4" bold="true" box="[634,646,212,231]" pageId="11" pageNumber="75">–</emphasis>
|
||
2) poorly recognizable small conical teeth (Figure 5b), pharyngobranchial 3 toothplate not visible on 3D models from microCT images. Number of vertebrae 35
|
||
<emphasis id="A380EAA6FFBB5B2152E5FEBD0CF15E4C" bold="true" box="[143,155,316,335]" pageId="11" pageNumber="75">–</emphasis>
|
||
36, abdominal 16 and caudal 21 (Supporting Information Table S2). The first gill arch with hemibranch, the 2nd to 4th gill arches with holobranchs. Subopercle indistinguishable from opercle, shaped as its posterior elongated extension, not forming or having subopercular spine. Six branchiostegals. Nasal bones with inconspicuous frontal end. Maxillary, premaxillary, nasal and ceratobranchial 5 bones shaped as on Figure 5b.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D9EE653FFFBB5B2152F5FD8A0EC75D84" pageId="11" pageNumber="75" type="etymology">
|
||
<paragraph id="914B36B4FFBB5B2152F5FD8A0EC75D84" blockId="11.[119,769,142,1965]" pageId="11" pageNumber="75">
|
||
<emphasis id="A380EAA6FFBB5B2152F5FD8A0D605D1C" bold="true" box="[159,266,523,543]" pageId="11" pageNumber="75">Etymology.</emphasis>
|
||
Named
|
||
<taxonomicName id="56F44D37FFBB5B21530AFD8A0DDA5D1C" authorityName="Wagner & Kovac & Koblmüller" authorityYear="2021" box="[352,432,523,543]" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="11" pageNumber="75" phylum="Chordata" rank="species" species="orientalis">
|
||
<emphasis id="A380EAA6FFBB5B21530AFD8A0DDA5D1C" box="[352,432,523,543]" italics="true" pageId="11" pageNumber="75">orientalis</emphasis>
|
||
</taxonomicName>
|
||
, from the Latin word
|
||
<emphasis id="A380EAA6FFBB5B2150FBFD8D0EF35D1C" bold="true" box="[657,665,524,543]" pageId="11" pageNumber="75">“</emphasis>
|
||
oriens
|
||
<emphasis id="A380EAA6FFBB5B2150B8FD8D0EB05D1C" bold="true" box="[722,730,524,543]" pageId="11" pageNumber="75">”</emphasis>
|
||
for
|
||
<emphasis id="A380EAA6FFBB5B21521DFDAE0C155D41" bold="true" box="[119,127,559,578]" pageId="11" pageNumber="75">“</emphasis>
|
||
east
|
||
<emphasis id="A380EAA6FFBB5B2152CFFDAE0CC75D41" bold="true" box="[165,173,559,578]" pageId="11" pageNumber="75">”</emphasis>
|
||
, which describes the distribution range of the species that is restricted to the oriental Mediterranean basin. The name is an adjective in the nominative singular (Article 11.9.1.1.,
|
||
<collectionCode id="F7E5AE71FFBB5B21505FFDF20E025D84" box="[565,616,627,647]" pageId="11" pageNumber="75">ICZN</collectionCode>
|
||
, 1999).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D9EE653FFFBB5B2152F5FD170D575C79" pageId="11" pageNumber="75" type="biology_ecology">
|
||
<paragraph id="914B36B4FFBB5B2152F5FD170D575C79" blockId="11.[119,769,142,1965]" pageId="11" pageNumber="75">
|
||
<emphasis id="A380EAA6FFBB5B2152F5FD170EE25DA9" bold="true" box="[159,648,662,682]" pageId="11" pageNumber="75">Ecology and geographical distribution (Figure 1a).</emphasis>
|
||
The known species distribution range encompasses the
|
||
<collectingRegion id="5330F856FFBB5B215059FD380E315DCE" box="[563,603,697,717]" country="Papua New Guinea" name="Gulf" pageId="11" pageNumber="75">Gulf</collectingRegion>
|
||
of Corinth, the
|
||
<collectingRegion id="5330F856FFBB5B21521DFD5A0CD65DEC" box="[119,188,731,751]" country="Greece" name="Voreio Aigaio" pageId="11" pageNumber="75">Aegean</collectingRegion>
|
||
Sea (
|
||
<collectingRegion id="5330F856FFBB5B215284FD5A0D4C5DEC" box="[238,294,731,751]" country="Greece" name="Attiki" pageId="11" pageNumber="75">Attica</collectingRegion>
|
||
) and on the islands
|
||
<collectingRegion id="5330F856FFBB5B2153B7FD5A0E7A5DEC" box="[477,528,731,751]" country="Greece" name="Kriti" pageId="11" pageNumber="75">Crete</collectingRegion>
|
||
and Kythira. Quantitative data on ecology is largely lacking.
|
||
<taxonomicName id="56F44D37FFBB5B2153DFFD7F0E3F5C11" authority="Wagner & Kovac & Koblmüller, 2021" authorityName="Wagner & Kovac & Koblmüller" authorityYear="2021" box="[437,597,766,786]" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="11" pageNumber="75" phylum="Chordata" rank="species" species="orientalis" status="sp. nov.">
|
||
<emphasis id="A380EAA6FFBB5B2153DFFD7F0E3F5C11" box="[437,597,766,786]" italics="true" pageId="11" pageNumber="75">Gouania orientalis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A380EAA6FFBB5B215036FD7F0ECC5C11" bold="true" box="[604,678,766,786]" pageId="11" pageNumber="75">
|
||
<taxonomicNameLabel id="B8B357DDFFBB5B215036FD7F0ECC5C11" box="[604,678,766,786]" pageId="11" pageNumber="75" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
occurs in sympatry or even syntopy with
|
||
<taxonomicName id="56F44D37FFBB5B2153C2FCA00E755C36" authority="Wagner & Kovac & Koblmüller, 2021" authorityName="Wagner & Kovac & Koblmüller" authorityYear="2021" box="[424,543,801,821]" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="11" pageNumber="75" phylum="Chordata" rank="species" species="hofrichteri" status="sp. nov.">
|
||
<emphasis id="A380EAA6FFBB5B2153C2FCA00E755C36" box="[424,543,801,821]" italics="true" pageId="11" pageNumber="75">G. hofrichteri</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A380EAA6FFBB5B21504CFCA10E1B5C37" bold="true" box="[550,625,800,820]" pageId="11" pageNumber="75">
|
||
<taxonomicNameLabel id="B8B357DDFFBB5B21504CFCA10E1B5C37" box="[550,625,800,820]" pageId="11" pageNumber="75" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
throughout its distribution range. The species inhabits intertidal and subtidal pebble and boulder beaches.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D9EE653FFFBB5B2152F5FC090FF85C11" pageId="11" pageNumber="75" type="discussion">
|
||
<paragraph id="914B36B4FFBB5B2152F5FC090FF85C11" blockId="11.[119,769,142,1965]" lastBlockId="11.[816,1465,142,786]" pageId="11" pageNumber="75">
|
||
<emphasis id="A380EAA6FFBB5B2152F5FC090C9D5C9F" bold="true" box="[159,247,904,924]" pageId="11" pageNumber="75">Remarks.</emphasis>
|
||
<taxonomicName id="56F44D37FFBB5B215294FC080DCB5C9E" authority="Wagner & Kovac & Koblmüller, 2021" authorityName="Wagner & Kovac & Koblmüller" authorityYear="2021" box="[254,417,905,925]" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="11" pageNumber="75" phylum="Chordata" rank="species" species="orientalis" status="sp. nov.">
|
||
<emphasis id="A380EAA6FFBB5B215294FC080DCB5C9E" box="[254,417,905,925]" italics="true" pageId="11" pageNumber="75">Gouania orientalis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A380EAA6FFBB5B2153C3FC090D9F5C9F" bold="true" box="[425,501,904,924]" pageId="11" pageNumber="75">
|
||
<taxonomicNameLabel id="B8B357DDFFBB5B2153C3FC090D9F5C9F" box="[425,501,904,924]" pageId="11" pageNumber="75" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs from slender-bodied
|
||
<taxonomicName id="56F44D37FFBB5B21521DFC2A0D665CBC" box="[119,268,939,959]" pageId="11" pageNumber="75">
|
||
<emphasis id="A380EAA6FFBB5B21521DFC2A0CAB5CBC" box="[119,193,939,959]" italics="true" pageId="11" pageNumber="75">Gouania</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName id="56F44D37FFBB5B215373FC2A0D355CBC" baseAuthorityName="Nardo" baseAuthorityYear="1827" box="[281,351,939,959]" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="11" pageNumber="75" phylum="Chordata" rank="species" species="pigra">
|
||
<emphasis id="A380EAA6FFBB5B215373FC2A0D355CBC" box="[281,351,939,959]" italics="true" pageId="11" pageNumber="75">G. pigra</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="56F44D37FFBB5B2153FAFC2A0E6C5CBC" authority="Wagner & Kovac & Koblmüller, 2021" authorityName="Wagner & Kovac & Koblmüller" authorityYear="2021" box="[400,518,939,959]" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="11" pageNumber="75" phylum="Chordata" rank="species" species="hofrichteri" status="sp. nov.">
|
||
<emphasis id="A380EAA6FFBB5B2153FAFC2A0E6C5CBC" box="[400,518,939,959]" italics="true" pageId="11" pageNumber="75">G. hofrichteri</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A380EAA6FFBB5B215067FC2A0E3D5CBC" bold="true" box="[525,599,939,959]" pageId="11" pageNumber="75">
|
||
<taxonomicNameLabel id="B8B357DDFFBB5B215067FC2A0E3D5CBC" box="[525,599,939,959]" pageId="11" pageNumber="75" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
) by a dorsal head profile that forms a straight line between nape above eye and upper lip tip (
|
||
<emphasis id="A380EAA6FFBB5B2152DCFC700CA25B06" box="[182,200,1009,1029]" italics="true" pageId="11" pageNumber="75">vs</emphasis>
|
||
. dorsal head profile in lateral view
|
||
<emphasis id="A380EAA6FFBB5B215061FC700E795B07" bold="true" box="[523,531,1009,1028]" pageId="11" pageNumber="75">“</emphasis>
|
||
S
|
||
<emphasis id="A380EAA6FFBB5B215074FC700E4C5B07" bold="true" box="[542,550,1009,1028]" pageId="11" pageNumber="75">”</emphasis>
|
||
curved, concave above eye and convex at nape), an infraorbital invagination vertical to posterior part of eye (
|
||
<emphasis id="A380EAA6FFBB5B215365FBB70D4B5B49" box="[271,289,1078,1098]" italics="true" pageId="11" pageNumber="75">vs</emphasis>
|
||
. infraorbital invagination below anterior half of eye or below mideye), lower number of vertebrae (Supporting Information Table S2; 35
|
||
<emphasis id="A380EAA6FFBB5B215284FBFD0C905B8C" bold="true" box="[238,250,1148,1167]" pageId="11" pageNumber="75">–</emphasis>
|
||
36
|
||
<emphasis id="A380EAA6FFBB5B215373FBFA0D415B8C" box="[281,299,1147,1167]" italics="true" pageId="11" pageNumber="75">vs</emphasis>
|
||
. 38
|
||
<emphasis id="A380EAA6FFBB5B215325FBFD0D315B8C" bold="true" box="[335,347,1148,1167]" pageId="11" pageNumber="75">–</emphasis>
|
||
40) and a star-like pigmentation around eyes (
|
||
<emphasis id="A380EAA6FFBB5B215217FB1F0CE55BB1" box="[125,143,1182,1202]" italics="true" pageId="11" pageNumber="75">vs</emphasis>
|
||
. no star-like pigmentation around eyes). Nine morphometric characters as percentages of standard length of
|
||
<taxonomicName id="56F44D37FFBB5B215076FB400EED5BD6" authority="Wagner & Kovac & Koblmüller, 2021" authorityName="Wagner & Kovac & Koblmüller" authorityYear="2021" box="[540,647,1217,1237]" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="11" pageNumber="75" phylum="Chordata" rank="species" species="orientalis" status="sp. nov.">
|
||
<emphasis id="A380EAA6FFBB5B215076FB400EED5BD6" box="[540,647,1217,1237]" italics="true" pageId="11" pageNumber="75">G. orientalis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A380EAA6FFBB5B2150E5FB410EB15BD7" bold="true" box="[655,731,1216,1236]" pageId="11" pageNumber="75">
|
||
<taxonomicNameLabel id="B8B357DDFFBB5B2150E5FB410EB15BD7" box="[655,731,1216,1236]" pageId="11" pageNumber="75" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
are nonoverlapping in range with both slender-bodied
|
||
<taxonomicName id="56F44D37FFBB5B21501CFB620EAA5BF4" authorityName="Nardo" authorityYear="1833" box="[630,704,1251,1271]" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="11" pageNumber="75" phylum="Chordata" rank="genus">
|
||
<emphasis id="A380EAA6FFBB5B21501CFB620EAA5BF4" box="[630,704,1251,1271]" italics="true" pageId="11" pageNumber="75">Gouania</emphasis>
|
||
</taxonomicName>
|
||
: head length, horizontal eye diameter, body width at pectoral fins, pectoral-fin length, prepectoral distance, ventral adhesive disc length, distance between the posterior margin of sucking disc and anus, caudal base depth and caudal-fin length (values in Table 1). There are also morphometric characters nonoverlapping in range with only one of the two slender-bodied
|
||
<taxonomicName id="56F44D37FFBB5B21535BFA320D115AC4" authorityName="Nardo" authorityYear="1833" box="[305,379,1459,1479]" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="11" pageNumber="75" phylum="Chordata" rank="genus">
|
||
<emphasis id="A380EAA6FFBB5B21535BFA320D115AC4" box="[305,379,1459,1479]" italics="true" pageId="11" pageNumber="75">Gouania</emphasis>
|
||
</taxonomicName>
|
||
(Table 1). In addition,
|
||
<taxonomicName id="56F44D37FFBB5B215022FA320EDB5AC4" authority="Wagner & Kovac & Koblmüller, 2021" authorityName="Wagner & Kovac & Koblmüller" authorityYear="2021" box="[584,689,1459,1479]" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="11" pageNumber="75" phylum="Chordata" rank="species" species="orientalis" status="sp. nov.">
|
||
<emphasis id="A380EAA6FFBB5B215022FA320EDB5AC4" box="[584,689,1459,1479]" italics="true" pageId="11" pageNumber="75">G. orientalis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A380EAA6FFBB5B2150DDFA320F6A5AC4" bold="true" box="[695,768,1459,1479]" pageId="11" pageNumber="75">
|
||
<taxonomicNameLabel id="B8B357DDFFBB5B2150DDFA320F6A5AC4" box="[695,768,1459,1479]" pageId="11" pageNumber="75" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs from
|
||
<taxonomicName id="56F44D37FFBB5B215293FA570D2F5AE9" baseAuthorityName="Nardo" baseAuthorityYear="1827" box="[249,325,1494,1514]" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="11" pageNumber="75" phylum="Chordata" rank="species" species="pigra">
|
||
<emphasis id="A380EAA6FFBB5B215293FA570D2F5AE9" box="[249,325,1494,1514]" italics="true" pageId="11" pageNumber="75">G. pigra</emphasis>
|
||
</taxonomicName>
|
||
by a posterior angle of jaws extending to between vertical line drawn through anterior edge of eye and vertical line drawn through anterior part of eye (
|
||
<emphasis id="A380EAA6FFBB5B215068F99A0E7E592C" box="[514,532,1563,1583]" italics="true" pageId="11" pageNumber="75">vs</emphasis>
|
||
. posterior angle of jaws extending to, or close to, a vertical line drawn through the anterior edge of the anterior nostril), pectoral rays 17
|
||
<emphasis id="A380EAA6FFBB5B21505AF9E00E565977" bold="true" box="[560,572,1633,1652]" pageId="11" pageNumber="75">–</emphasis>
|
||
19 (
|
||
<emphasis id="A380EAA6FFBB5B215009F9E00E1F5976" box="[611,629,1633,1653]" italics="true" pageId="11" pageNumber="75">vs</emphasis>
|
||
. pectoral rays 13
|
||
<emphasis id="A380EAA6FFBB5B2152FAF9050CF65994" bold="true" box="[144,156,1668,1687]" pageId="11" pageNumber="75">–</emphasis>
|
||
16) and upper attachment of disc membrane attaching to base of pectoral fin at 16th
|
||
<emphasis id="A380EAA6FFBB5B21535BF9260D5759B9" bold="true" box="[305,317,1703,1722]" pageId="11" pageNumber="75">–</emphasis>
|
||
18th pectoral ray (
|
||
<emphasis id="A380EAA6FFBB5B21539AF9270E6859B9" box="[496,514,1702,1722]" italics="true" pageId="11" pageNumber="75">vs</emphasis>
|
||
. upper attachment of disc membrane attaching to base of pectoral fin at 12th
|
||
<emphasis id="A380EAA6FFBB5B215018F9480E1459DF" bold="true" box="[626,638,1737,1756]" pageId="11" pageNumber="75">–</emphasis>
|
||
15th pectoral ray).
|
||
<taxonomicName id="56F44D37FFBB5B2152CCF96A0D7B59FC" authority="Wagner & Kovac & Koblmüller, 2021" authorityName="Wagner & Kovac & Koblmüller" authorityYear="2021" box="[166,273,1771,1791]" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="11" pageNumber="75" phylum="Chordata" rank="species" species="orientalis" status="sp. nov.">
|
||
<emphasis id="A380EAA6FFBB5B2152CCF96A0D7B59FC" box="[166,273,1771,1791]" italics="true" pageId="11" pageNumber="75">G. orientalis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A380EAA6FFBB5B215373F96A0D0E59FC" bold="true" box="[281,356,1771,1791]" pageId="11" pageNumber="75">
|
||
<taxonomicNameLabel id="B8B357DDFFBB5B215373F96A0D0E59FC" box="[281,356,1771,1791]" pageId="11" pageNumber="75" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is also different from
|
||
<taxonomicName id="56F44D37FFBB5B215052F96A0EC459FC" authority="Wagner & Kovac & Koblmüller, 2021" authorityName="Wagner & Kovac & Koblmüller" authorityYear="2021" box="[568,686,1771,1791]" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="11" pageNumber="75" phylum="Chordata" rank="species" species="hofrichteri" status="sp. nov.">
|
||
<emphasis id="A380EAA6FFBB5B215052F96A0EC459FC" box="[568,686,1771,1791]" italics="true" pageId="11" pageNumber="75">G. hofrichteri</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A380EAA6FFBB5B2150DFF96A0F6A59FC" bold="true" box="[693,768,1771,1791]" pageId="11" pageNumber="75">
|
||
<taxonomicNameLabel id="B8B357DDFFBB5B2150DFF96A0F6A59FC" box="[693,768,1771,1791]" pageId="11" pageNumber="75" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
by posterior opercular edge w-shaped with two equally long tips (
|
||
<emphasis id="A380EAA6FFBB5B215082F88F0E905821" box="[744,762,1806,1826]" italics="true" pageId="11" pageNumber="75">vs</emphasis>
|
||
. pointed upper tip and rounded lower posterior opercular edge), longitudinal infralateral and suborbital transversal rows of superficial neuromasts placed in the well-defined deep groove (
|
||
<emphasis id="A380EAA6FFBB5B215005F8F70EEB5889" box="[623,641,1910,1930]" italics="true" pageId="11" pageNumber="75">vs</emphasis>
|
||
. longitudinal infralateral and suborbital transversal rows of superficial neuromasts placed in shallow groove disappearing in posterior part of longitudinal infralateral row), body cross-section behind pectoral fin base half oval with straight ventral side (
|
||
<emphasis id="A380EAA6FFBB5B215640FF5208565FE4" box="[1066,1084,211,231]" italics="true" pageId="11" pageNumber="75">vs</emphasis>
|
||
. trunk cross-section behind pectoral fin base triangular with ventral flat and dorsal pointed), granules on body shallow and inconspicuous (
|
||
<emphasis id="A380EAA6FFBB5B215659FE98082F5E2E" box="[1075,1093,281,301]" italics="true" pageId="11" pageNumber="75">vs</emphasis>
|
||
. granules on body, at least on posterior part and nape, large and prominent) and upper attachment of gill membrane opposite to 5th
|
||
<emphasis id="A380EAA6FFBB5B21564DFEDE08595E71" bold="true" box="[1063,1075,351,370]" pageId="11" pageNumber="75">–</emphasis>
|
||
6th pectoral ray (
|
||
<emphasis id="A380EAA6FFBB5B2156BBFEDF08895E71" box="[1233,1251,350,370]" italics="true" pageId="11" pageNumber="75">vs</emphasis>
|
||
. the upper attachment of the gill membrane opposite to 3rd
|
||
<emphasis id="A380EAA6FFBB5B2156FEFE0008CA5E97" bold="true" box="[1172,1184,385,404]" pageId="11" pageNumber="75">–</emphasis>
|
||
4th pectoral ray).
|
||
<taxonomicName id="56F44D37FFBB5B215721FE0009DD5E96" authority="Wagner & Kovac & Koblmüller, 2021" authorityName="Wagner & Kovac & Koblmüller" authorityYear="2021" box="[1355,1463,385,405]" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="11" pageNumber="75" phylum="Chordata" rank="species" species="orientalis" status="sp. nov.">
|
||
<emphasis id="A380EAA6FFBB5B215721FE0009DD5E96" box="[1355,1463,385,405]" italics="true" pageId="11" pageNumber="75">G. orientalis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A380EAA6FFBB5B21515AFE220F105EB4" bold="true" box="[816,890,419,439]" pageId="11" pageNumber="75">
|
||
<taxonomicNameLabel id="B8B357DDFFBB5B21515AFE220F105EB4" box="[816,890,419,439]" pageId="11" pageNumber="75" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs from the stout-bodied species
|
||
<taxonomicName id="56F44D37FFBB5B21568CFE2209255EB4" authority="Wagner & Kovac & Koblmüller, 2021" authorityName="Wagner & Kovac & Koblmüller" authorityYear="2021" box="[1254,1359,419,439]" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="11" pageNumber="75" phylum="Chordata" rank="species" species="adriatica" status="sp. nov.">
|
||
<emphasis id="A380EAA6FFBB5B21568CFE2209255EB4" box="[1254,1359,419,439]" italics="true" pageId="11" pageNumber="75">G. adriatica</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A380EAA6FFBB5B21573FFE2209CA5EB4" bold="true" box="[1365,1440,419,439]" pageId="11" pageNumber="75">
|
||
<taxonomicNameLabel id="B8B357DDFFBB5B21573FFE2209CA5EB4" box="[1365,1440,419,439]" pageId="11" pageNumber="75" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in posterior opercular edge w-shaped with two equally long tips (
|
||
<emphasis id="A380EAA6FFBB5B215718FE4709EE5ED9" box="[1394,1412,454,474]" italics="true" pageId="11" pageNumber="75">vs</emphasis>
|
||
. posterior opercular edge with pointed upper tip and rounded lower edge), principal caudal rays 10
|
||
<emphasis id="A380EAA6FFBB5B21567BFD8D08775D1C" bold="true" box="[1041,1053,524,543]" pageId="11" pageNumber="75">–</emphasis>
|
||
11 (
|
||
<emphasis id="A380EAA6FFBB5B215629FD8A083F5D1C" box="[1091,1109,523,543]" italics="true" pageId="11" pageNumber="75">vs</emphasis>
|
||
. principal caudal-fin rays 12
|
||
<emphasis id="A380EAA6FFBB5B215709FD8D09055D1C" bold="true" box="[1379,1391,524,543]" pageId="11" pageNumber="75">–</emphasis>
|
||
13) and pattern of pigmentation (stripes, marbles
|
||
<emphasis id="A380EAA6FFBB5B2156C7FDAF08D55D41" box="[1197,1215,558,578]" italics="true" pageId="11" pageNumber="75">vs</emphasis>
|
||
. reduced pigmentation).
|
||
<taxonomicName id="56F44D37FFBB5B2157CEFDAF0FEA5D66" authority="Wagner & Kovac & Koblmüller, 2021" authorityName="Wagner & Kovac & Koblmüller" authorityYear="2021" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="11" pageNumber="75" phylum="Chordata" rank="species" species="orientalis" status="sp. nov.">
|
||
<emphasis id="A380EAA6FFBB5B2157CEFDAF0FEA5D66" italics="true" pageId="11" pageNumber="75">G. orientalis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A380EAA6FFBB5B2151E2FDD10FBE5D67" bold="true" box="[904,980,592,612]" pageId="11" pageNumber="75">
|
||
<taxonomicNameLabel id="B8B357DDFFBB5B2151E2FDD10FBE5D67" box="[904,980,592,612]" pageId="11" pageNumber="75" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
has no nonoverlapping external morphological differences to
|
||
<taxonomicName id="56F44D37FFBB5B2151C9FDF208715D84" baseAuthorityName="Risso" baseAuthorityYear="1810" box="[931,1051,627,647]" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="11" pageNumber="75" phylum="Chordata" rank="species" species="willdenowi">
|
||
<emphasis id="A380EAA6FFBB5B2151C9FDF208715D84" box="[931,1051,627,647]" italics="true" pageId="11" pageNumber="75">G. willdenowi</emphasis>
|
||
</taxonomicName>
|
||
but differs by its low number of vertebrae (Supporting Information Table S2; 35
|
||
<emphasis id="A380EAA6FFBB5B2156E5FD1608F15DA9" bold="true" box="[1167,1179,663,682]" pageId="11" pageNumber="75">–</emphasis>
|
||
36
|
||
<emphasis id="A380EAA6FFBB5B2156D6FD1708A45DA9" box="[1212,1230,662,682]" italics="true" pageId="11" pageNumber="75">vs</emphasis>
|
||
. 37
|
||
<emphasis id="A380EAA6FFBB5B215699FD1608955DA9" bold="true" box="[1267,1279,663,682]" pageId="11" pageNumber="75">–</emphasis>
|
||
38).
|
||
<taxonomicName id="56F44D37FFBB5B215740FD1709FF5DA9" authority="Wagner & Kovac & Koblmüller, 2021" authorityName="Wagner & Kovac & Koblmüller" authorityYear="2021" box="[1322,1429,662,682]" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="11" pageNumber="75" phylum="Chordata" rank="species" species="orientalis" status="sp. nov.">
|
||
<emphasis id="A380EAA6FFBB5B215740FD1709FF5DA9" box="[1322,1429,662,682]" italics="true" pageId="11" pageNumber="75">G. orientalis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A380EAA6FFBB5B2157F7FD170F335DCF" bold="true" pageId="11" pageNumber="75">
|
||
<taxonomicNameLabel id="B8B357DDFFBB5B2157F7FD170F335DCF" pageId="11" pageNumber="75" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is known from
|
||
<collectingRegion id="5330F856FFBB5B215199FD3808525DCE" box="[1011,1080,697,717]" country="Greece" name="Voreio Aigaio" pageId="11" pageNumber="75">Aegean</collectingRegion>
|
||
and Ionian Sea and has nonoverlapping geographic distribution with
|
||
<taxonomicName id="56F44D37FFBB5B215621FD5A08DD5DEC" authority="Wagner & Kovac & Koblmüller, 2021" authorityName="Wagner & Kovac & Koblmüller" authorityYear="2021" box="[1099,1207,731,751]" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="11" pageNumber="75" phylum="Chordata" rank="species" species="adriatica" status="sp. nov.">
|
||
<emphasis id="A380EAA6FFBB5B215621FD5A08DD5DEC" box="[1099,1207,731,751]" italics="true" pageId="11" pageNumber="75">G. adriatica</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A380EAA6FFBB5B2156A9FD5A097B5DEC" bold="true" box="[1219,1297,731,751]" pageId="11" pageNumber="75">
|
||
<taxonomicNameLabel id="B8B357DDFFBB5B2156A9FD5A097B5DEC" box="[1219,1297,731,751]" pageId="11" pageNumber="75" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
,
|
||
<taxonomicName id="56F44D37FFBB5B21574AFD5A09015DEC" baseAuthorityName="Nardo" baseAuthorityYear="1827" box="[1312,1387,731,751]" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="11" pageNumber="75" phylum="Chordata" rank="species" species="pigra">
|
||
<emphasis id="A380EAA6FFBB5B21574AFD5A09015DEC" box="[1312,1387,731,751]" italics="true" pageId="11" pageNumber="75">G. pigra</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="56F44D37FFBB5B2157CEFD5A0FE75C11" baseAuthorityName="Risso" baseAuthorityYear="1810" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="11" pageNumber="75" phylum="Chordata" rank="species" species="willdenowi">
|
||
<emphasis id="A380EAA6FFBB5B2157CEFD5A0FE75C11" italics="true" pageId="11" pageNumber="75">G. willdenowi</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |