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<document id="B2F84E124C8E3DC421BAB033492F915D" ID-DOI="10.1111/jfb.14558" ID-ISSN="1095-8649" ID-PMC="PMC7821206" ID-PubMed="32985685" ID-Zenodo-Dep="10928926" checkinTime="1712261492542" checkinUser="felipe" docAuthor="Wagner, Maximilian, Kovac, Marcelo &amp; Koblmüller, Stephan" docDate="2021" docId="195D87A2FFB65B225216FAA408EA590E" docLanguage="en" docName="JFishBiol.98.1.64-88.pdf" docOrigin="Journal of Fish Biology 98 (1)" docSource="http://dx.doi.org/10.1111/jfb.14558" docStyle="DocumentStyle:E312FF628AF650F200AD49ACE691BDEB:JFishBiol.2018-.journal_article.open" docStyleId="E312FF628AF650F200AD49ACE691BDEB" docStyleName="JFishBiol.2018-.journal_article.open" docTitle="Gouania adriatica Wagner &amp; Kovac &amp; Koblmüller 2021, sp. nov." docType="treatment" docVersion="2" lastPageNumber="72" masterDocId="E564FFDAFFB05B2A526AFF810C6A5F03" masterDocTitle="Unravelling the taxonomy of an interstitial fish radiation: Three new species of Gouania (Teleostei: Gobiesocidae) from the Mediterranean Sea and redescriptions of G. willdenowi and G. pigra" masterLastPageNumber="88" masterPageNumber="64" pageNumber="70" updateTime="1712261854377" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-ND-4.0">
<mods:mods id="BAD7004658652E95FD530C6DE89F9234" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="A3218DD133F3D37CFE5E848F1F4EC8B7">
<mods:title id="7B5F562726D5005240B4455B8481254C">Unravelling the taxonomy of an interstitial fish radiation: Three new species of Gouania (Teleostei: Gobiesocidae) from the Mediterranean Sea and redescriptions of G. willdenowi and G. pigra</mods:title>
</mods:titleInfo>
<mods:name id="E93B267B9FECA5021524CC3C1CC7AAFE" type="personal">
<mods:role id="4C289120910DC8C7472A6D0D58F2DAE1">
<mods:roleTerm id="F8341F65F4CA1CC977ED591471748287">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="ACC8508F25E0AB32FD62694F6CB55B3B">Wagner, Maximilian</mods:namePart>
<mods:affiliation id="825151C25020E339EEB03BA0836D171E">Institute of Biology, University of Graz, Graz, Austria &amp; Department of Biology, University of Antwerp, Antwerp, Belgium</mods:affiliation>
</mods:name>
<mods:name id="28263BF8AADF25BC4948E78A4225505E" type="personal">
<mods:role id="720E6E675D4B015C6443AC7ABB5C4E1E">
<mods:roleTerm id="E34F4CD8E522804239EA80D459F6B9D6">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="1B2342ED98BC44FC6995564667EC2F32">Kovac, Marcelo</mods:namePart>
<mods:affiliation id="C4EC6E496555B70E363A3D0F297B07B9">Natural History Museum Rijeka, Rijeka, Croatia</mods:affiliation>
</mods:name>
<mods:name id="F4A9BB1A3B092B3830474D3F8128E2CE" type="personal">
<mods:role id="A6588F7BB005C682ABC062226D7031D0">
<mods:roleTerm id="6D6A5BA744BCD68A9DA7415554EC7F5C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="3F6386005B3B7CB9FE26BC3F02474007">Koblmüller, Stephan</mods:namePart>
<mods:affiliation id="AB4F335563EBD5425712B7E26550BA19">Institute of Biology, University of Graz, Graz, Austria</mods:affiliation>
</mods:name>
<mods:typeOfResource id="BDB05CEFFA4E3620B8AACC44AAF9F903">text</mods:typeOfResource>
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<mods:titleInfo id="38BFEDD6F10529FD4275892D6F5FABED">
<mods:title id="546680165CB7E36767C9A41566514079">Journal of Fish Biology</mods:title>
</mods:titleInfo>
<mods:part id="7394E08586F32BEA96DAA1957B054A7D">
<mods:date id="49EE3B3D2542AF5430FD84E9627E2081">2021</mods:date>
<mods:detail id="9D066EA8D41F074DC2706A56A58D4827" type="pubDate">
<mods:number id="96262823B0AC57C9A487B700DD63332E">2020-11-03</mods:number>
</mods:detail>
<mods:detail id="FFB0883E5FEBD9B52619E7340D91F38C" type="volume">
<mods:number id="F61103866066055E006BE388B36A04AC">98</mods:number>
</mods:detail>
<mods:detail id="D28C35D68B3F1139C0E49A9E6F023F14" type="issue">
<mods:number id="DC65479B7A0AA4E9BA37DA73BE28AB02">1</mods:number>
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<mods:start id="7AF9056E3D4BED26F91E6D380CEFFCDF">64</mods:start>
<mods:end id="D27B109597B5D455E2C32046CFC65942">88</mods:end>
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<mods:url id="BD84A3C2079A5B374C82E5A79894460D">http://dx.doi.org/10.1111/jfb.14558</mods:url>
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<mods:classification id="D35D0E558B01363622BBEE49F2ED30E5">journal article</mods:classification>
<mods:identifier id="AA515C51A7C2862606C728EACB2C7AF3" type="DOI">10.1111/jfb.14558</mods:identifier>
<mods:identifier id="F33EE8CC2B9EBD383895DFECD49052B9" type="ISSN">1095-8649</mods:identifier>
<mods:identifier id="02D9E1B871A64571B34B17EF9F4D90A9" type="PMC">PMC7821206</mods:identifier>
<mods:identifier id="A95307D93B693CC390CCEBA4674A1BEF" type="PubMed">32985685</mods:identifier>
<mods:identifier id="E6662C0359E5DB099615EA75CA43AEAB" type="Zenodo-Dep">10928926</mods:identifier>
</mods:mods>
<treatment id="195D87A2FFB65B225216FAA408EA590E" LSID="urn:lsid:plazi:treatment:195D87A2FFB65B225216FAA408EA590E" httpUri="http://treatment.plazi.org/id/195D87A2FFB65B225216FAA408EA590E" lastPageId="8" lastPageNumber="72" pageId="6" pageNumber="70">
<subSubSection id="D9EE653FFFB65B2C5216FAA40E4D5A3D" box="[124,551,1316,1342]" pageId="6" pageNumber="70" type="nomenclature">
<paragraph id="914B36B4FFB65B2C5216FAA40E4D5A3D" blockId="6.[124,551,1316,1342]" box="[124,551,1316,1342]" pageId="6" pageNumber="70">
<heading id="CA0381D8FFB65B2C5216FAA40E4D5A3D" box="[124,551,1316,1342]" fontSize="10" level="3" pageId="6" pageNumber="70" reason="9">
<emphasis id="A380EAA6FFB65B2C5216FAA40CDC5A3D" bold="true" box="[124,182,1317,1342]" pageId="6" pageNumber="70">3.1.2</emphasis>
|
<emphasis id="A380EAA6FFB65B2C5297FAA50E4D5A3D" bold="true" box="[253,551,1316,1342]" pageId="6" pageNumber="70">
<taxonomicName id="56F44D37FFB65B2C5297FAA50DAF5A3D" authority="Wagner &amp; Kovac &amp; Koblmüller, 2021" authorityName="Wagner &amp; Kovac &amp; Koblmüller" authorityYear="2021" box="[253,453,1316,1342]" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="6" pageNumber="70" phylum="Chordata" rank="species" species="adriatica" status="sp. nov.">
<emphasis id="A380EAA6FFB65B2C5297FAA50DAF5A3D" bold="true" box="[253,453,1316,1342]" italics="true" pageId="6" pageNumber="70">Gouania adriatica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B8B357DDFFB65B2C53A7FAA40E4D5A3D" box="[461,551,1317,1342]" pageId="6" pageNumber="70" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="D9EE653FFFB65B2C5216FAEF0D3C5AC4" pageId="6" pageNumber="70" type="description">
<paragraph id="914B36B4FFB65B2C5216FAEF0E405A81" blockId="6.[124,773,1390,1965]" box="[124,554,1390,1410]" pageId="6" pageNumber="70">
<emphasis id="A380EAA6FFB65B2C5216FAEF0E405A81" bold="true" box="[124,554,1390,1410]" pageId="6" pageNumber="70">English name: Adriatic blunt-snouted clingfish</emphasis>
</paragraph>
<paragraph id="914B36B4FFB65B2C52CEFA100D3C5AC4" blockId="6.[124,773,1390,1965]" pageId="6" pageNumber="70">
<emphasis id="A380EAA6FFB65B2C52CEFA100D465AA6" box="[164,300,1425,1445]" italics="true" pageId="6" pageNumber="70">
ZooBank
<collectionCode id="F7E5AE71FFB65B2C5369FA100D465AA6" box="[259,300,1425,1445]" pageId="6" pageNumber="70">LSID</collectionCode>
</emphasis>
:
<uri id="E5653AB6FFB65B2C532BFA100D3C5AC4" pageId="6" pageNumber="70">
urn:lsid:zoobank.org:act:
<uuid id="E5520C61FFB65B2C504FFA100D3C5AC4" pageId="6" pageNumber="70">F054E7C9-604F-41DA-8068-145DDCCC1FBE</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="D9EE653FFFB65B2C52CEFA5709025B49" pageId="6" pageNumber="70" type="materials_examined">
<paragraph id="914B36B4FFB65B2C52CEFA570CB3592C" blockId="6.[124,773,1390,1965]" pageId="6" pageNumber="70">
<materialsCitation id="219C3CE9FFB65B2C52CEFA570CB3592C" collectingDate="2016-07-17" collectionCode="PMR" collectorName="M. Wagner" country="Croatia" location="Croatia" pageId="6" pageNumber="70" specimenCode="VP4618" specimenCount="1" typeStatus="holotype">
<emphasis id="A380EAA6FFB65B2C52CEFA570C965AE9" bold="true" box="[164,252,1494,1514]" pageId="6" pageNumber="70">
<typeStatus id="4E4F8816FFB65B2C52CEFA570C965AE9" box="[164,252,1494,1514]" pageId="6" pageNumber="70" type="holotype">Holotype</typeStatus>
</emphasis>
.
<collectionCode id="F7E5AE71FFB65B2C536CFA570D5E5AE9" box="[262,308,1494,1514]" country="0" httpUri="http://grbio.org/cool/xq4z-q5gq" name="Prirodoslovni muzej Rijeka" pageId="6" pageNumber="70">PMR</collectionCode>
<specimenCode id="C1529ECFFFB65B2C5350FA570DED5AE9" box="[314,391,1494,1514]" collectionCode="VP" pageId="6" pageNumber="70">VP4618</specimenCode>
, male, 41.41 +
<quantity id="560C9B51FFB65B2C5079FA570E0D5AE9" box="[531,615,1494,1514]" metricMagnitude="-3" metricUnit="m" metricValue="6.77" pageId="6" pageNumber="70" unit="mm" value="6.77">6.77 mm</quantity>
, Stoja, Pula,
<collectingCountry id="E9E37624FFB65B2C50B0FA570CCA590E" name="Croatia" pageId="6" pageNumber="70">Croatia</collectingCountry>
, 44 51
<emphasis id="A380EAA6FFB65B2C528DFA770C815900" bold="true" box="[231,235,1526,1539]" italics="true" pageId="6" pageNumber="70">
<superScript id="66819BFCFFB65B2C528DFA770C815900" attach="none" box="[231,235,1526,1539]" fontSize="6" pageId="6" pageNumber="70">0</superScript>
</emphasis>
38.4
<emphasis id="A380EAA6FFB65B2C537CFA770D745900" bold="true" box="[278,286,1526,1539]" italics="true" pageId="6" pageNumber="70">
<superScript id="66819BFCFFB65B2C537CFA770D745900" attach="right" box="[278,286,1526,1539]" fontSize="6" pageId="6" pageNumber="70">00</superScript>
</emphasis>
N, 13 49
<emphasis id="A380EAA6FFB65B2C5313FA770D175900" bold="true" box="[377,381,1526,1539]" italics="true" pageId="6" pageNumber="70">
<superScript id="66819BFCFFB65B2C5313FA770D175900" attach="none" box="[377,381,1526,1539]" fontSize="6" pageId="6" pageNumber="70">0</superScript>
</emphasis>
05.0
<emphasis id="A380EAA6FFB65B2C53C2FA770DDA5900" bold="true" box="[424,432,1526,1539]" italics="true" pageId="6" pageNumber="70">
<superScript id="66819BFCFFB65B2C53C2FA770DDA5900" attach="right" box="[424,432,1526,1539]" fontSize="6" pageId="6" pageNumber="70">00</superScript>
</emphasis>
E, coll.
<collectorName id="3C015362FFB65B2C5397FA780E07590E" box="[509,621,1529,1549]" pageId="6" pageNumber="70">M. Wagner</collectorName>
,
<date id="E54A1074FFB65B2C5011FA780F6E590E" box="[635,772,1529,1549]" pageId="6" pageNumber="70" value="2016-07-17">
<collectingDate id="F50EE99CFFB65B2C5011FA780F6E590E" box="[635,772,1529,1549]" pageId="6" pageNumber="70" value="2016-07-17">July 17, 2016</collectingDate>
</date>
(Figure 3).
</materialsCitation>
</paragraph>
<paragraph id="914B36B4FFB65B2C52CEF9BF09025B49" blockId="6.[124,773,1390,1965]" lastBlockId="6.[821,1470,939,1965]" pageId="6" pageNumber="70">
<materialsCitation id="219C3CE9FFB65B2C52CEF9BF0CBE5994" collectingDate="2019-08-14" collectorName="M. Wagner" country="Albania" location="5.89 mm Envi" pageId="6" pageNumber="70" specimenCount="1" stateProvince="Vlore" typeStatus="paratype">
<emphasis id="A380EAA6FFB65B2C52CEF9BF0D6B5951" bold="true" box="[164,257,1598,1618]" pageId="6" pageNumber="70">
<typeStatus id="4E4F8816FFB65B2C52CEF9BF0D6B5951" box="[164,257,1598,1618]" pageId="6" pageNumber="70" type="paratype">Paratypes</typeStatus>
</emphasis>
. ZSM-PIS-047650, male, 41.03 +
<location id="942B606FFFB65B2C502AF9BF0EAB5951" LSID="urn:lsid:plazi:treatment:195D87A2FFB65B225216FAA408EA590E:942B606FFFB65B2C502AF9BF0EAB5951" box="[576,705,1598,1618]" country="Albania" name="5.89 mm Envi" pageId="6" pageNumber="70" stateProvince="Vlore">
<quantity id="560C9B51FFB65B2C502AF9BF0EF85951" box="[576,658,1598,1618]" metricMagnitude="-3" metricUnit="m" metricValue="5.89" pageId="6" pageNumber="70" unit="mm" value="5.89">5.89 mm</quantity>
Envi
</location>
beach,
<collectingRegion id="5330F856FFB65B2C5216F9E00CDA5976" box="[124,176,1633,1653]" country="Albania" name="Vlore" pageId="6" pageNumber="70">Vlorë</collectingRegion>
,
<collectingCountry id="E9E37624FFB65B2C52D0F9E00D6B5976" box="[186,257,1633,1653]" name="Albania" pageId="6" pageNumber="70">Albania</collectingCountry>
, 40 23
<emphasis id="A380EAA6FFB65B2C5329F9DF0D2D5968" bold="true" box="[323,327,1630,1643]" italics="true" pageId="6" pageNumber="70">
<superScript id="66819BFCFFB65B2C5329F9DF0D2D5968" attach="none" box="[323,327,1630,1643]" fontSize="6" pageId="6" pageNumber="70">0</superScript>
</emphasis>
16.3
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<superScript id="66819BFCFFB65B2C5318F9DF0D105968" attach="right" box="[370,378,1630,1643]" fontSize="6" pageId="6" pageNumber="70">00</superScript>
</emphasis>
N, 19 28
<emphasis id="A380EAA6FFB65B2C53BBF9DF0DBF5968" bold="true" box="[465,469,1630,1643]" italics="true" pageId="6" pageNumber="70">
<superScript id="66819BFCFFB65B2C53BBF9DF0DBF5968" attach="none" box="[465,469,1630,1643]" fontSize="6" pageId="6" pageNumber="70">0</superScript>
</emphasis>
58.2
<emphasis id="A380EAA6FFB65B2C506AF9DF0E625968" bold="true" box="[512,520,1630,1643]" italics="true" pageId="6" pageNumber="70">
<superScript id="66819BFCFFB65B2C506AF9DF0E625968" attach="right" box="[512,520,1630,1643]" fontSize="6" pageId="6" pageNumber="70">00</superScript>
</emphasis>
E, coll.
<collectorName id="3C015362FFB65B2C5027F9E00ED35976" box="[589,697,1633,1653]" pageId="6" pageNumber="70">M. Wagner</collectorName>
,
<date id="E54A1074FFB65B2C50A8F9E00CBE5994" pageId="6" pageNumber="70" value="2019-08-14">
<collectingDate id="F50EE99CFFB65B2C50A8F9E00CBE5994" pageId="6" pageNumber="70" value="2019-08-14">August 14, 2019</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="219C3CE9FFB65B2C52B7F9020CDA59DE" collectingDate="2019-08-18" collectorName="M. Wagner" country="Croatia" location="Trstenik Peljesac" pageId="6" pageNumber="70" specimenCount="1" typeStatus="paratype">
ZSM-PIS-047652, male, 27.3 +
<quantity id="560C9B51FFB65B2C506FF9020E335994" box="[517,601,1667,1687]" metricMagnitude="-3" metricUnit="m" metricValue="4.27" pageId="6" pageNumber="70" unit="mm" value="4.27">4.27 mm</quantity>
,
<location id="942B606FFFB65B2C5008F9020F6B5994" LSID="urn:lsid:plazi:treatment:195D87A2FFB65B225216FAA408EA590E:942B606FFFB65B2C5008F9020F6B5994" box="[610,769,1667,1687]" country="Croatia" name="Trstenik Peljesac" pageId="6" pageNumber="70">Trstenik Pelješac</location>
,
<collectingCountry id="E9E37624FFB65B2C5216F9270CA859B9" box="[124,194,1702,1722]" name="Croatia" pageId="6" pageNumber="70">Croatia</collectingCountry>
, 42 54
<emphasis id="A380EAA6FFB65B2C5362F9220D6659B3" bold="true" box="[264,268,1699,1712]" italics="true" pageId="6" pageNumber="70">
<superScript id="66819BFCFFB65B2C5362F9220D6659B3" attach="none" box="[264,268,1699,1712]" fontSize="6" pageId="6" pageNumber="70">0</superScript>
</emphasis>
07.7
<emphasis id="A380EAA6FFB65B2C535DF9220D5559B3" bold="true" box="[311,319,1699,1712]" italics="true" pageId="6" pageNumber="70">
<superScript id="66819BFCFFB65B2C535DF9220D5559B3" attach="right" box="[311,319,1699,1712]" fontSize="6" pageId="6" pageNumber="70">00</superScript>
</emphasis>
N, 17 25
<emphasis id="A380EAA6FFB65B2C53F0F9220DF459B3" bold="true" box="[410,414,1699,1712]" italics="true" pageId="6" pageNumber="70">
<superScript id="66819BFCFFB65B2C53F0F9220DF459B3" attach="none" box="[410,414,1699,1712]" fontSize="6" pageId="6" pageNumber="70">0</superScript>
</emphasis>
47.0
<emphasis id="A380EAA6FFB65B2C53A3F9220DBB59B3" bold="true" box="[457,465,1699,1712]" italics="true" pageId="6" pageNumber="70">
<superScript id="66819BFCFFB65B2C53A3F9220DBB59B3" attach="right" box="[457,465,1699,1712]" fontSize="6" pageId="6" pageNumber="70">00</superScript>
</emphasis>
E, coll.
<collectorName id="3C015362FFB65B2C5077F9270EE759B9" box="[541,653,1702,1722]" pageId="6" pageNumber="70">M. Wagner</collectorName>
,
<date id="E54A1074FFB65B2C50F0F9270CDA59DE" pageId="6" pageNumber="70" value="2019-08-18">
<collectingDate id="F50EE99CFFB65B2C50F0F9270CDA59DE" pageId="6" pageNumber="70" value="2019-08-18">August 18, 2019</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="219C3CE9FFB65B2C52D4F9480CDA5821" collectingDate="2019-09-16" collectorName="M. Wagner" country="Croatia" location="Stara Baska" municipality="Krk" pageId="6" pageNumber="70" specimenCount="1" typeStatus="paratype">
ZSM-PIS-047651, male, 34.53 +
<quantity id="560C9B51FFB65B2C5390F9480E2559DE" box="[506,591,1737,1757]" metricMagnitude="-3" metricUnit="m" metricValue="5.65" pageId="6" pageNumber="70" unit="mm" value="5.65">5.65 mm</quantity>
,
<location id="942B606FFFB65B2C5037F9480EBA59DE" LSID="urn:lsid:plazi:treatment:195D87A2FFB65B225216FAA408EA590E:942B606FFFB65B2C5037F9480EBA59DE" box="[605,720,1737,1757]" country="Croatia" municipality="Krk" name="Stara Baska" pageId="6" pageNumber="70">Stara Baška</location>
,
<collectingMunicipality id="712FACCEFFB65B2C50B5F9480F6B59DE" box="[735,769,1737,1757]" pageId="6" pageNumber="70">Krk</collectingMunicipality>
,
<collectingCountry id="E9E37624FFB65B2C5216F96A0CA859FC" box="[124,194,1771,1791]" name="Croatia" pageId="6" pageNumber="70">Croatia</collectingCountry>
, 44 56
<emphasis id="A380EAA6FFB65B2C5369F9680D6D59F5" bold="true" box="[259,263,1769,1782]" italics="true" pageId="6" pageNumber="70">
<superScript id="66819BFCFFB65B2C5369F9680D6D59F5" attach="none" box="[259,263,1769,1782]" fontSize="6" pageId="6" pageNumber="70">0</superScript>
</emphasis>
45.3
<emphasis id="A380EAA6FFB65B2C5358F9680D5059F5" bold="true" box="[306,314,1769,1782]" italics="true" pageId="6" pageNumber="70">
<superScript id="66819BFCFFB65B2C5358F9680D5059F5" attach="right" box="[306,314,1769,1782]" fontSize="6" pageId="6" pageNumber="70">00</superScript>
</emphasis>
N, 14 42
<emphasis id="A380EAA6FFB65B2C53FAF9680DFE59F5" bold="true" box="[400,404,1769,1782]" italics="true" pageId="6" pageNumber="70">
<superScript id="66819BFCFFB65B2C53FAF9680DFE59F5" attach="none" box="[400,404,1769,1782]" fontSize="6" pageId="6" pageNumber="70">0</superScript>
</emphasis>
22.2
<emphasis id="A380EAA6FFB65B2C53D4F9680DAC59F5" bold="true" box="[446,454,1769,1782]" italics="true" pageId="6" pageNumber="70">
<superScript id="66819BFCFFB65B2C53D4F9680DAC59F5" attach="right" box="[446,454,1769,1782]" fontSize="6" pageId="6" pageNumber="70">00</superScript>
</emphasis>
E, coll.
<collectorName id="3C015362FFB65B2C5063F96A0E1E59FC" box="[521,628,1771,1791]" pageId="6" pageNumber="70">M. Wagner</collectorName>
,
<date id="E54A1074FFB65B2C5016F96A0CDA5821" pageId="6" pageNumber="70" value="2019-09-16">
<collectingDate id="F50EE99CFFB65B2C5016F96A0CDA5821" pageId="6" pageNumber="70" value="2019-09-16">September 16, 2019</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="219C3CE9FFB65B2C52D7F88F0D625864" collectingDate="2015-05-13" collectionCode="PMR" collectorName="M. Wagner" country="Croatia" location="Glavotok" municipality="Krk" pageId="6" pageNumber="70" specimenCode="VP3523" specimenCount="1" typeStatus="paratype">
<collectionCode id="F7E5AE71FFB65B2C52D7F88F0C815821" box="[189,235,1806,1826]" country="0" httpUri="http://grbio.org/cool/xq4z-q5gq" name="Prirodoslovni muzej Rijeka" pageId="6" pageNumber="70">PMR</collectionCode>
<specimenCode id="C1529ECFFFB65B2C529CF88F0D2E5821" box="[246,324,1806,1826]" collectionCode="VP" pageId="6" pageNumber="70">VP3523</specimenCode>
, juvenile of unidentified sex, 22.95 +
<quantity id="560C9B51FFB65B2C50D3F88F0F6B5821" box="[697,769,1806,1826]" metricMagnitude="-3" metricUnit="m" metricValue="3.6" pageId="6" pageNumber="70" unit="mm" value="3.6">3.6 mm</quantity>
,
<location id="942B606FFFB65B2C5216F8B00CB85846" LSID="urn:lsid:plazi:treatment:195D87A2FFB65B225216FAA408EA590E:942B606FFFB65B2C5216F8B00CB85846" box="[124,210,1841,1861]" country="Croatia" municipality="Krk" name="Glavotok" pageId="6" pageNumber="70">Glavotok</location>
,
<collectingMunicipality id="712FACCEFFB65B2C52B1F8B00C975846" box="[219,253,1841,1861]" pageId="6" pageNumber="70">Krk</collectingMunicipality>
,
<collectingCountry id="E9E37624FFB65B2C536CF8B00D265846" box="[262,332,1841,1861]" name="Croatia" pageId="6" pageNumber="70">Croatia</collectingCountry>
, 45 05
<emphasis id="A380EAA6FFB65B2C53E4F8AF0DF85838" bold="true" box="[398,402,1838,1851]" italics="true" pageId="6" pageNumber="70">
<superScript id="66819BFCFFB65B2C53E4F8AF0DF85838" attach="none" box="[398,402,1838,1851]" fontSize="6" pageId="6" pageNumber="70">0</superScript>
</emphasis>
44.9
<emphasis id="A380EAA6FFB65B2C53D7F8AF0DAF5838" bold="true" box="[445,453,1838,1851]" italics="true" pageId="6" pageNumber="70">
<superScript id="66819BFCFFB65B2C53D7F8AF0DAF5838" attach="right" box="[445,453,1838,1851]" fontSize="6" pageId="6" pageNumber="70">00</superScript>
</emphasis>
N, 14 26
<emphasis id="A380EAA6FFB65B2C5071F8AF0E755838" bold="true" box="[539,543,1838,1851]" italics="true" pageId="6" pageNumber="70">
<superScript id="66819BFCFFB65B2C5071F8AF0E755838" attach="none" box="[539,543,1838,1851]" fontSize="6" pageId="6" pageNumber="70">0</superScript>
</emphasis>
32.4
<emphasis id="A380EAA6FFB65B2C5020F8AF0E385838" bold="true" box="[586,594,1838,1851]" italics="true" pageId="6" pageNumber="70">
<superScript id="66819BFCFFB65B2C5020F8AF0E385838" attach="right" box="[586,594,1838,1851]" fontSize="6" pageId="6" pageNumber="70">00</superScript>
</emphasis>
E, coll.
<collectorName id="3C015362FFB65B2C50FCF8B00F6B5846" box="[662,769,1841,1861]" pageId="6" pageNumber="70">M. Wagner</collectorName>
,
<date id="E54A1074FFB65B2C5216F8D20D625864" box="[124,264,1875,1895]" pageId="6" pageNumber="70" value="2015-05-13">
<collectingDate id="F50EE99CFFB65B2C5216F8D20D625864" box="[124,264,1875,1895]" pageId="6" pageNumber="70" value="2015-05-13">May 13, 2015</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="219C3CE9FFB65B2C537EF8D20F6B58AE" collectingDate="2015-05-15" collectionCode="PMR" collectorName="M. Wagner" country="Croatia" location="Glavotok" municipality="Krk" pageId="6" pageNumber="70" specimenCode="VP3524, VP3525" specimenCount="1" typeStatus="paratype">
<collectionCode id="F7E5AE71FFB65B2C537EF8D20D285864" box="[276,322,1875,1895]" country="0" httpUri="http://grbio.org/cool/xq4z-q5gq" name="Prirodoslovni muzej Rijeka" pageId="6" pageNumber="70">PMR</collectionCode>
<specimenCode id="C1529ECFFFB65B2C5321F8D20DF25864" box="[331,408,1875,1895]" collectionCode="VP" pageId="6" pageNumber="70">VP3524</specimenCode>
, male.
<quantity id="560C9B51FFB65B2C53B4F8D20E555864" box="[478,575,1875,1895]" metricMagnitude="-2" metricUnit="m" metricValue="2.619" pageId="6" pageNumber="70" unit="mm" value="26.19">26.19 mm</quantity>
, caudal fin damaged and
<collectionCode id="F7E5AE71FFB65B2C52CFF8F70CB95889" box="[165,211,1910,1930]" country="0" httpUri="http://grbio.org/cool/xq4z-q5gq" name="Prirodoslovni muzej Rijeka" pageId="6" pageNumber="70">PMR</collectionCode>
<specimenCode id="C1529ECFFFB65B2C52B0F8F70D425889" box="[218,296,1910,1930]" collectionCode="VP" pageId="6" pageNumber="70">VP3525</specimenCode>
, female, 28.4 +
<quantity id="560C9B51FFB65B2C53D6F8F70E7B5889" box="[444,529,1910,1930]" metricMagnitude="-3" metricUnit="m" metricValue="4.23" pageId="6" pageNumber="70" unit="mm" value="4.23">4.23 mm</quantity>
, both from
<location id="942B606FFFB65B2C5015F8F70EBF5889" LSID="urn:lsid:plazi:treatment:195D87A2FFB65B225216FAA408EA590E:942B606FFFB65B2C5015F8F70EBF5889" box="[639,725,1910,1930]" country="Croatia" municipality="Krk" name="Glavotok" pageId="6" pageNumber="70">Glavotok</location>
,
<collectingMunicipality id="712FACCEFFB65B2C50B5F8F70F6B5889" box="[735,769,1910,1930]" pageId="6" pageNumber="70">Krk</collectingMunicipality>
,
<collectingCountry id="E9E37624FFB65B2C5216F8180CA858AE" box="[124,194,1945,1965]" name="Croatia" pageId="6" pageNumber="70">Croatia</collectingCountry>
, 45 05
<emphasis id="A380EAA6FFB65B2C5369F8170D6D58A0" bold="true" box="[259,263,1942,1955]" italics="true" pageId="6" pageNumber="70">
<superScript id="66819BFCFFB65B2C5369F8170D6D58A0" attach="none" box="[259,263,1942,1955]" fontSize="6" pageId="6" pageNumber="70">0</superScript>
</emphasis>
44.9
<emphasis id="A380EAA6FFB65B2C5358F8170D5058A0" bold="true" box="[306,314,1942,1955]" italics="true" pageId="6" pageNumber="70">
<superScript id="66819BFCFFB65B2C5358F8170D5058A0" attach="right" box="[306,314,1942,1955]" fontSize="6" pageId="6" pageNumber="70">00</superScript>
</emphasis>
N, 14 26
<emphasis id="A380EAA6FFB65B2C53FAF8170DFE58A0" bold="true" box="[400,404,1942,1955]" italics="true" pageId="6" pageNumber="70">
<superScript id="66819BFCFFB65B2C53FAF8170DFE58A0" attach="none" box="[400,404,1942,1955]" fontSize="6" pageId="6" pageNumber="70">0</superScript>
</emphasis>
32.4
<emphasis id="A380EAA6FFB65B2C53D4F8170DAC58A0" bold="true" box="[446,454,1942,1955]" italics="true" pageId="6" pageNumber="70">
<superScript id="66819BFCFFB65B2C53D4F8170DAC58A0" attach="right" box="[446,454,1942,1955]" fontSize="6" pageId="6" pageNumber="70">00</superScript>
</emphasis>
E, coll.
<collectorName id="3C015362FFB65B2C5063F8180E1958AE" box="[521,627,1945,1965]" pageId="6" pageNumber="70">M. Wagner</collectorName>
,
<date id="E54A1074FFB65B2C5011F8180F6B58AE" box="[635,769,1945,1965]" pageId="6" pageNumber="70" value="2015-05-15">
<collectingDate id="F50EE99CFFB65B2C5011F8180F6B58AE" box="[635,769,1945,1965]" pageId="6" pageNumber="70" value="2015-05-15">May 15, 2015</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="219C3CE9FFB65B2C515FFC2A09E85B06" collectingDate="2019-07-12" collectorName="M. Wagner" country="Croatia" location="Pecine" municipality="Rijeka" pageId="6" pageNumber="70" specimenCount="1" typeStatus="paratype">
ZSM-PIS-047653, female, 34.42 +
<quantity id="560C9B51FFB65B2C56EDFC2A08B35CBC" box="[1159,1241,939,959]" metricMagnitude="-3" metricUnit="m" metricValue="4.85" pageId="6" pageNumber="70" unit="mm" value="4.85">4.85 mm</quantity>
and ZSM-PIS-047653, female, 35.23 +
<quantity id="560C9B51FFB65B2C51B7FC4F084F5CE1" box="[989,1061,974,994]" metricMagnitude="-3" metricUnit="m" metricValue="4.9" pageId="6" pageNumber="70" unit="mm" value="4.9">4.9 mm</quantity>
, both from
<location id="942B606FFFB65B2C56ACFC4F096C5CE1" LSID="urn:lsid:plazi:treatment:195D87A2FFB65B225216FAA408EA590E:942B606FFFB65B2C56ACFC4F096C5CE1" box="[1222,1286,974,994]" country="Croatia" municipality="Rijeka" name="Pecine" pageId="6" pageNumber="70">Pecine</location>
,
<collectingMunicipality id="712FACCEFFB65B2C574AFC4F09305CE1" box="[1312,1370,974,994]" pageId="6" pageNumber="70">Rijeka</collectingMunicipality>
,
<collectingCountry id="E9E37624FFB65B2C571EFC4F09D05CE1" box="[1396,1466,974,994]" name="Croatia" pageId="6" pageNumber="70">Croatia</collectingCountry>
, 45 18
<emphasis id="A380EAA6FFB65B2C5104FC6F0F185CF8" bold="true" box="[878,882,1006,1019]" italics="true" pageId="6" pageNumber="70">
<superScript id="66819BFCFFB65B2C5104FC6F0F185CF8" attach="none" box="[878,882,1006,1019]" fontSize="6" pageId="6" pageNumber="70">0</superScript>
</emphasis>
52.4
<emphasis id="A380EAA6FFB65B2C51F7FC6F0FCF5CF8" bold="true" box="[925,933,1006,1019]" italics="true" pageId="6" pageNumber="70">
<superScript id="66819BFCFFB65B2C51F7FC6F0FCF5CF8" attach="right" box="[925,933,1006,1019]" fontSize="6" pageId="6" pageNumber="70">00</superScript>
</emphasis>
N, 14 28
<emphasis id="A380EAA6FFB65B2C5194FC6F08685CF8" bold="true" box="[1022,1026,1006,1019]" italics="true" pageId="6" pageNumber="70">
<superScript id="66819BFCFFB65B2C5194FC6F08685CF8" attach="none" box="[1022,1026,1006,1019]" fontSize="6" pageId="6" pageNumber="70">0</superScript>
</emphasis>
11.7
<emphasis id="A380EAA6FFB65B2C5647FC6F085F5CF8" bold="true" box="[1069,1077,1006,1019]" italics="true" pageId="6" pageNumber="70">
<superScript id="66819BFCFFB65B2C5647FC6F085F5CF8" attach="right" box="[1069,1077,1006,1019]" fontSize="6" pageId="6" pageNumber="70">00</superScript>
</emphasis>
E, coll.
<collectorName id="3C015362FFB65B2C5615FC7008845B06" box="[1151,1262,1009,1029]" pageId="6" pageNumber="70">M. Wagner</collectorName>
,
<date id="E54A1074FFB65B2C5690FC7009E85B06" box="[1274,1410,1009,1029]" pageId="6" pageNumber="70" value="2019-07-12">
<collectingDate id="F50EE99CFFB65B2C5690FC7009E85B06" box="[1274,1410,1009,1029]" pageId="6" pageNumber="70" value="2019-07-12">July 12, 2019</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="219C3CE9FFB65B2C57E4FC70090E5B49" collectingDate="2015-07-18" collectionCode="PMR" collectorName="M. Wagner" country="Croatia" location="Sv." pageId="6" pageNumber="70" specimenCode="VP4580" specimenCount="1" stateProvince="Istarska" typeStatus="paratype">
<collectionCode id="F7E5AE71FFB65B2C57E4FC7009D65B06" box="[1422,1468,1009,1029]" country="0" httpUri="http://grbio.org/cool/xq4z-q5gq" name="Prirodoslovni muzej Rijeka" pageId="6" pageNumber="70">PMR</collectionCode>
<specimenCode id="C1529ECFFFB65B2C515FFB920FE85B24" box="[821,898,1043,1063]" collectionCode="VP" pageId="6" pageNumber="70">VP4580</specimenCode>
, female, 34.7 +
<quantity id="560C9B51FFB65B2C565EFB9208E35B24" box="[1076,1161,1043,1063]" metricMagnitude="-3" metricUnit="m" metricValue="4.99" pageId="6" pageNumber="70" unit="mm" value="4.99">4.99 mm</quantity>
,
<location id="942B606FFFB65B2C56CAFB9208D75B24" LSID="urn:lsid:plazi:treatment:195D87A2FFB65B225216FAA408EA590E:942B606FFFB65B2C56CAFB9208D75B24" box="[1184,1213,1043,1063]" country="Croatia" name="Sv." pageId="6" pageNumber="70" stateProvince="Istarska">Sv.</location>
<taxonomicName id="56F44D37FFB65B2C56BBFB92097E5B24" box="[1233,1300,1043,1063]" class="Magnoliopsida" family="Fabaceae" genus="Marina" kingdom="Plantae" order="Fabales" pageId="6" pageNumber="87" phylum="Tracheophyta" rank="genus">Marina</taxonomicName>
,
<collectingRegion id="5330F856FFB65B2C5746FB9209365B24" box="[1324,1372,1043,1063]" country="Croatia" name="Istarska" pageId="6" pageNumber="70">Istria</collectingRegion>
,
<collectingCountry id="E9E37624FFB65B2C571EFB9209D05B24" box="[1396,1466,1043,1063]" name="Croatia" pageId="6" pageNumber="70">Croatia</collectingCountry>
, 45 01
<emphasis id="A380EAA6FFB65B2C5104FBB20F185B43" bold="true" box="[878,882,1075,1088]" italics="true" pageId="6" pageNumber="70">
<superScript id="66819BFCFFB65B2C5104FBB20F185B43" attach="none" box="[878,882,1075,1088]" fontSize="6" pageId="6" pageNumber="70">0</superScript>
</emphasis>
42.1
<emphasis id="A380EAA6FFB65B2C51F7FBB20FCF5B43" bold="true" box="[925,933,1075,1088]" italics="true" pageId="6" pageNumber="70">
<superScript id="66819BFCFFB65B2C51F7FBB20FCF5B43" attach="right" box="[925,933,1075,1088]" fontSize="6" pageId="6" pageNumber="70">00</superScript>
</emphasis>
N, 14 09
<emphasis id="A380EAA6FFB65B2C5190FBB20F945B43" bold="true" box="[1018,1022,1075,1088]" italics="true" pageId="6" pageNumber="70">
<superScript id="66819BFCFFB65B2C5190FBB20F945B43" attach="none" box="[1018,1022,1075,1088]" fontSize="6" pageId="6" pageNumber="70">0</superScript>
</emphasis>
17.4
<emphasis id="A380EAA6FFB65B2C5643FBB2085B5B43" bold="true" box="[1065,1073,1075,1088]" italics="true" pageId="6" pageNumber="70">
<superScript id="66819BFCFFB65B2C5643FBB2085B5B43" attach="right" box="[1065,1073,1075,1088]" fontSize="6" pageId="6" pageNumber="70">00</superScript>
</emphasis>
E, coll.
<collectorName id="3C015362FFB65B2C5619FBB708B75B49" box="[1139,1245,1078,1098]" pageId="6" pageNumber="70">M. Wagner</collectorName>
,
<date id="E54A1074FFB65B2C568FFBB7090E5B49" box="[1253,1380,1078,1098]" pageId="6" pageNumber="70" value="2015-07-18">
<collectingDate id="F50EE99CFFB65B2C568FFBB7090E5B49" box="[1253,1380,1078,1098]" pageId="6" pageNumber="70" value="2015-07-18">July 18, 2015</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="D9EE653FFFB65B2C5137FBD909E85821" pageId="6" pageNumber="70" type="diagnosis">
<paragraph id="914B36B4FFB65B2C5137FBD909E85821" blockId="6.[821,1470,939,1965]" pageId="6" pageNumber="70">
<emphasis id="A380EAA6FFB65B2C5137FBD90FD25B6F" bold="true" box="[861,952,1112,1132]" pageId="6" pageNumber="70">Diagnosis</emphasis>
.
<taxonomicName id="56F44D37FFB65B2C51A8FBD8080A5B6E" authority="Wagner &amp; Kovac &amp; Koblmüller, 2021" authorityName="Wagner &amp; Kovac &amp; Koblmüller" authorityYear="2021" box="[962,1120,1113,1133]" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="6" pageNumber="70" phylum="Chordata" rank="species" species="adriatica" status="sp. nov.">
<emphasis id="A380EAA6FFB65B2C51A8FBD8080A5B6E" box="[962,1120,1113,1133]" italics="true" pageId="6" pageNumber="70">Gouania adriatica</emphasis>
</taxonomicName>
<emphasis id="A380EAA6FFB65B2C560FFBD908C45B6F" bold="true" box="[1125,1198,1112,1132]" pageId="6" pageNumber="70">
<taxonomicNameLabel id="B8B357DDFFB65B2C560FFBD908C45B6F" box="[1125,1198,1112,1132]" pageId="6" pageNumber="70" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from its congeners by the combination of the following characters: (1) dorsal head profile a straight line from nape above eye to upper lip tip; (2) posterior angle of jaws extends to between a vertical line drawn through posterior edge of anterior nostril and a vertical line drawn through anterior edge of eye; (3) pointed upper and rounded lower posterior opercular edge; (4) longitudinal infralateral and suborbital transversal rows of superficial neuromasts placed in the well-defined deep groove; (5) body cross-section behind pectoral fin base half oval with straight ventral side; (6) the granules on body shallow and inconspicuous; (7) upper attachment of gill membrane opposite to 5th to 6th pectoral ray; (8) principal caudal-fin rays 12
<emphasis id="A380EAA6FFB65B2C565BFA5608575AE9" bold="true" box="[1073,1085,1495,1514]" pageId="6" pageNumber="70">–</emphasis>
13; (9) vertical eye diameter 3.4
<emphasis id="A380EAA6FFB65B2C5702FA56091E5AE9" bold="true" box="[1384,1396,1495,1514]" pageId="6" pageNumber="70">–</emphasis>
4.3% of standard length; (10) horizontal eye diameter 3.0
<emphasis id="A380EAA6FFB65B2C5766FA780972590F" bold="true" box="[1292,1304,1529,1548]" pageId="6" pageNumber="70">–</emphasis>
3.7% of standard length; (11) head length 26.2
<emphasis id="A380EAA6FFB65B2C5655F99D0821592C" bold="true" box="[1087,1099,1564,1583]" pageId="6" pageNumber="70">–</emphasis>
30.0% of standard length; (12) pectoral-fin length 8.7
<emphasis id="A380EAA6FFB65B2C51D3F9BE0FAF5951" bold="true" box="[953,965,1599,1618]" pageId="6" pageNumber="70">–</emphasis>
11.3% of standard length; (13) prepectoral distance 25.1
<emphasis id="A380EAA6FFB65B2C5134F9E00F005977" bold="true" box="[862,874,1633,1652]" pageId="6" pageNumber="70">–</emphasis>
28.5% of standard length; (14) ventral adhesive disc length 15.9
<emphasis id="A380EAA6FFB65B2C5134F9050F005994" bold="true" box="[862,874,1668,1687]" pageId="6" pageNumber="70">–</emphasis>
20.1% of standard length; (15) caudal-fin length 13.9%
<emphasis id="A380EAA6FFB65B2C5706F90509125994" bold="true" box="[1388,1400,1668,1687]" pageId="6" pageNumber="70">–</emphasis>
16.4 of standard length; (16) low number of vertebrae (= 35); (17) pharyngeal jaws with ceratobranchial 5 small, having several (about 5) small, conical teeth; (18) nasal bones club-shaped; (19) star-like pigmentation around eyes, reduced body pigmentation with no visible stripes.
</paragraph>
</subSubSection>
<subSubSection id="D9EE653FFFB65B2D5137F8B1087B5AE9" lastPageId="7" lastPageNumber="71" pageId="6" pageNumber="70" type="description">
<paragraph id="914B36B4FFB65B2D5137F8B10ED85821" blockId="6.[821,1470,939,1965]" lastBlockId="7.[119,769,142,1965]" lastPageId="7" lastPageNumber="71" pageId="6" pageNumber="70">
<emphasis id="A380EAA6FFB65B2C5137F8B10FA05847" bold="true" box="[861,970,1840,1860]" pageId="6" pageNumber="70">Description</emphasis>
.
<emphasis id="A380EAA6FFB65B2C51B2F8B008E75846" box="[984,1165,1841,1861]" italics="true" pageId="6" pageNumber="70">General morphology</emphasis>
: Body proportions are given in Table 1. Body slender and elongated, posteriorly laterally compressed, body depth at pectoral fins
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6.4
<emphasis id="A380EAA6FFB65B2C561CF8F608E85889" bold="true" box="[1142,1154,1911,1930]" pageId="6" pageNumber="70">–</emphasis>
8.9 in
</quantity>
SL, body depth at anus
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7.7
<emphasis id="A380EAA6FFB65B2C5138F8180F3458AF" bold="true" box="[850,862,1945,1964]" pageId="6" pageNumber="70">–</emphasis>
10.2 in
</quantity>
SL, body depth in width at pectoral fins 1.1
<emphasis id="A380EAA6FFB65B2C573CF818090858AF" bold="true" box="[1366,1378,1945,1964]" pageId="6" pageNumber="70">–</emphasis>
1.4, body depth in width at anus 0.7
<emphasis id="A380EAA6FFB75B2D5313FF0E0DEF5FA1" bold="true" box="[377,389,143,162]" pageId="7" pageNumber="71">–</emphasis>
0.8. Body cross-section behind pectoral fin base half oval with straight ventral side. Granules on body shallow and inconspicuous, making skin surface more dotted than granulose. Head dorsoventrally compressed, head depth in width at orbit 1.5
<emphasis id="A380EAA6FFB75B2D52FFFE980CCB5E2F" bold="true" box="[149,161,281,300]" pageId="7" pageNumber="71">–</emphasis>
1.8, and moderately large, head length
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3.3
<emphasis id="A380EAA6FFB75B2D5046FE980E525E2F" bold="true" box="[556,568,281,300]" pageId="7" pageNumber="71">–</emphasis>
3.8 in
</quantity>
SL, head wider than body width maximum, head width at anterior sucking disc edge
<quantity id="560C9B51FFB75B2D521DFEDF0CBD5E71" box="[119,215,350,370]" metricMagnitude="-2" metricUnit="m" metricValue="2.032" metricValueMax="2.286" metricValueMin="1.7779999999999998" pageId="7" pageNumber="71" unit="in" value="0.8" valueMax="0.9" valueMin="0.7">
0.7
<emphasis id="A380EAA6FFB75B2D52FFFEDE0CCB5E71" bold="true" box="[149,161,351,370]" pageId="7" pageNumber="71">–</emphasis>
0.9 in
</quantity>
body width at pectoral fins. Dorsal head profile a straight line from nape above eye to upper lip tip. Head rounded in dorsal view. Snout large compared to eyes, preorbital distance
<quantity id="560C9B51FFB75B2D50F4FE220E955EB4" box="[670,767,419,439]" metricMagnitude="-2" metricUnit="m" metricValue="8.382" metricValueMax="9.398" metricValueMin="7.366" pageId="7" pageNumber="71" unit="in" value="3.3" valueMax="3.7" valueMin="2.9">
2.9
<emphasis id="A380EAA6FFB75B2D50D1FE250EAD5EB4" bold="true" box="[699,711,420,439]" pageId="7" pageNumber="71">–</emphasis>
3.7 in
</quantity>
head length,
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0.3
<emphasis id="A380EAA6FFB75B2D5367FE460D735ED9" bold="true" box="[269,281,455,474]" pageId="7" pageNumber="71">–</emphasis>
0.5 in
</quantity>
horizontal eye diameter. Snout wide, not produced, blunt. Internostril space gently convex. Eyes dorsolateral, with lower eye edge rounded. Eyes small,
<quantity id="560C9B51FFB75B2D53B3FD8A0E535D1C" box="[473,569,523,543]" metricMagnitude="-1" metricUnit="m" metricValue="2.1336" metricValueMax="2.413" metricValueMin="1.8541999999999998" pageId="7" pageNumber="71" unit="in" value="8.4" valueMax="9.5" valueMin="7.3">
7.3
<emphasis id="A380EAA6FFB75B2D539CFD8D0E685D1C" bold="true" box="[502,514,524,543]" pageId="7" pageNumber="71">–</emphasis>
9.5 in
</quantity>
head length, vertical diameter of the eye
<quantity id="560C9B51FFB75B2D532AFDAF0DCB5D41" box="[320,417,558,578]" metricMagnitude="-2" metricUnit="m" metricValue="2.413" metricValueMax="2.794" metricValueMin="2.032" pageId="7" pageNumber="71" unit="in" value="0.95" valueMax="1.1" valueMin="0.8">
0.8
<emphasis id="A380EAA6FFB75B2D5337FDAE0D035D41" bold="true" box="[349,361,559,578]" pageId="7" pageNumber="71">–</emphasis>
1.1 in
</quantity>
horizontal eye diameter. Infraorbital invagination vertical to posterior part of eye or to mideye. Interorbital distance wide,
<quantity id="560C9B51FFB75B2D536AFDF20D345D84" box="[256,350,627,647]" metricMagnitude="-2" metricUnit="m" metricValue="1.1429999999999998" metricValueMax="1.524" metricValueMin="0.762" pageId="7" pageNumber="71" unit="in" value="0.44999999999999996" valueMax="0.6" valueMin="0.3">
0.3
<emphasis id="A380EAA6FFB75B2D5374FDF50D405D84" bold="true" box="[286,298,628,647]" pageId="7" pageNumber="71">–</emphasis>
0.6 in
</quantity>
horizontal eye diameter.
<collectingRegion id="5330F856FFB75B2D5022FDF20EED5D84" box="[584,647,627,647]" country="France" name="Centre" pageId="7" pageNumber="71">Centre</collectingRegion>
of eye much closer to tip of snout than to posterior margin of operculum, preorbital distance in postorbital distance 1.8
<emphasis id="A380EAA6FFB75B2D538AFD380D865DCF" bold="true" box="[480,492,697,716]" pageId="7" pageNumber="71">–</emphasis>
2.2. Anterior and posterior nostrils long tubes of about equal length. Nostrils well separated and posterior nostril located behind and dorsally to the anterior edge of eyes. Single large lobe at the posterior margin of anterior nostril or bilobed, longer than nostril. Posterior nostril rim crenate with no extension. Head lateral line system with canals with pores and with superficial neuromasts arranged in rows. Head canals reduced and pores small. Single pore in nasal canal near posterior nostril. Single pore in postorbital canal close to posterior eye edge. Two pores in mandibular canal, anterior one close to anterolateral angle of mouth, posterior pore slightly in front of vertical of posterior angle of jaws, posterior pore usually more prominent. Lachrymal as well as preopercular canals and pores absent. Rows of superficial neuromasts as follows: SR 2, NR 3, LIR 24
<emphasis id="A380EAA6FFB75B2D53F7FBFD0DC35B8C" bold="true" box="[413,425,1148,1167]" pageId="7" pageNumber="71">–</emphasis>
27, STR 1
<emphasis id="A380EAA6FFB75B2D5062FBFD0E7E5B8C" bold="true" box="[520,532,1148,1167]" pageId="7" pageNumber="71">–</emphasis>
3, POR 3
<emphasis id="A380EAA6FFB75B2D5007FBFD0E135B8C" bold="true" box="[621,633,1148,1167]" pageId="7" pageNumber="71">–</emphasis>
4, PTR 2, SLR 4
<emphasis id="A380EAA6FFB75B2D52EEFB1E0CFA5BB1" bold="true" box="[132,144,1183,1202]" pageId="7" pageNumber="71">–</emphasis>
5, MR 9
<emphasis id="A380EAA6FFB75B2D52B4FB1E0C805BB1" bold="true" box="[222,234,1183,1202]" pageId="7" pageNumber="71">–</emphasis>
11, AVR 2, PVR 1, ADR 2, PDR 1, HR 3, SR1 3, SR2 2, DLR 6
<emphasis id="A380EAA6FFB75B2D52DFFB400CAB5BD7" bold="true" box="[181,193,1217,1236]" pageId="7" pageNumber="71">–</emphasis>
8, VLR 10
<emphasis id="A380EAA6FFB75B2D534EFB400D5A5BD7" bold="true" box="[292,304,1217,1236]" pageId="7" pageNumber="71">–</emphasis>
12. STR and LIR rows of superficial neuromasts placed in a well-defined deep groove. DLR row of superficial neuromasts anteriorly starts above pectoral fin, continuous dorsolateral and ends posteriorly downwards at midlateral level above anus or behind it. VLR anteriorly starts behind pectoral fin base, continuous ventrolateral and ends posteriorly upwards with last papilla nearly at midlateral level at caudal fin base. Mouth terminal, upper and lower lips end about equally, lips fleshy, upper lip larger than the lower lip. Posterior angle of jaws extends to between vertical line drawn through posterior edge of anterior nostril and vertical line drawn through anterior edge of eye. Chin with bilobed or slightly bilobed fold at anterior edge covering MR row of superficial neuromasts. Gill membrane attached to isthmus, gill opening starting at the base of pectoral fin, with upper attachment of gill membrane opposite to 5th to 6th pectoral ray. Pointed upper and rounded lower posterior opercular edge. No subopercular spine. No fleshy pad present on lower pectoral base. Urogenital papilla present. Preanus length in postanus length 0.6
<emphasis id="A380EAA6FFB75B2D52FFF88E0CCB5821" bold="true" box="[149,161,1807,1826]" pageId="7" pageNumber="71">–</emphasis>
0.7. Anal papillae absent, area around anus only wrinkled.
</paragraph>
<paragraph id="914B36B4FFB75B2D52F5F8B00FE65D1C" blockId="7.[119,769,142,1965]" lastBlockId="7.[816,1465,142,1965]" pageId="7" pageNumber="71">
<emphasis id="A380EAA6FFB75B2D52F5F8B00CA85846" box="[159,194,1841,1861]" italics="true" pageId="7" pageNumber="71">Fins</emphasis>
. Rudimentary dorsal and anal fins located well posteriorly and short, reduced to low ridges with very weak rays, connected to caudal fin. Pectoral rays 15
<emphasis id="A380EAA6FFB75B2D5354F8F60D205889" bold="true" box="[318,330,1911,1930]" pageId="7" pageNumber="71">–</emphasis>
17. Caudal fin rounded, principal caudal rays 12
<emphasis id="A380EAA6FFB75B2D52FAF8180CF658AF" bold="true" box="[144,156,1945,1964]" pageId="7" pageNumber="71">–</emphasis>
13. Ventral adhesive disc (Figure 4a) of
<emphasis id="A380EAA6FFB75B2D504CF8180E4458AF" bold="true" box="[550,558,1945,1964]" pageId="7" pageNumber="71">“</emphasis>
double
<emphasis id="A380EAA6FFB75B2D5007F8180E1F58AF" bold="true" box="[621,629,1945,1964]" pageId="7" pageNumber="71">”</emphasis>
<typeStatus id="4E4F8816FFB75B2D5014F8180EC358AE" box="[638,681,1945,1965]" pageId="7" pageNumber="71">type</typeStatus>
, anterior margin crenate with large invagination on each lateral side and in some specimen central invagination at midventral visible; posterior margin crenate or villous. Disc small, disc length 5.0
<emphasis id="A380EAA6FFB75B2D5744FF5509505FE4" bold="true" box="[1326,1338,212,231]" pageId="7" pageNumber="71">–</emphasis>
<quantity id="560C9B51FFB75B2D5750FF5209195FE4" box="[1338,1395,211,231]" metricMagnitude="-1" metricUnit="m" metricValue="1.6001999999999998" pageId="7" pageNumber="71" unit="in" value="6.3">6.3 in</quantity>
SL, its width slightly larger than its length, width in length 0.9
<emphasis id="A380EAA6FFB75B2D5747FF7609535E09" bold="true" box="[1325,1337,247,266]" pageId="7" pageNumber="71">–</emphasis>
1.0. No papillae in region A and flattened papillae in regions B and C. In region B one or two rows of papillae with total papillae count 10
<emphasis id="A380EAA6FFB75B2D5725FEBD09315E4C" bold="true" box="[1359,1371,316,335]" pageId="7" pageNumber="71">–</emphasis>
30 and in region C two rows of papillae with total papillae count 9
<emphasis id="A380EAA6FFB75B2D575DFEDE09295E71" bold="true" box="[1335,1347,351,370]" pageId="7" pageNumber="71">–</emphasis>
15. No inner row of papillae on lateral sides of the central part of the anterior disc. Upper attachment of disc membrane attaching to base of pectoral fin at 15th
<emphasis id="A380EAA6FFB75B2D511CFE460FE85ED9" bold="true" box="[886,898,455,474]" pageId="7" pageNumber="71">–</emphasis>
17th pectoral ray,
<emphasis id="A380EAA6FFB75B2D5658FE47082F5ED9" box="[1074,1093,454,474]" italics="true" pageId="7" pageNumber="71">i.e</emphasis>
., at the ultimate or penultimate ray. In males, parts of disc region A appear to be perfused (see Figure 3, ventral view).
</paragraph>
<paragraph id="914B36B4FFB75B2D5132FDAF0FF45C54" blockId="7.[816,1465,142,1965]" pageId="7" pageNumber="71">
<emphasis id="A380EAA6FFB75B2D5132FDAF0FD55D41" box="[856,959,558,578]" italics="true" pageId="7" pageNumber="71">Colouration</emphasis>
. Background colouration in life flesh-coloured to yellow, slightly transparent and head pigmentation prominent, with a star-like pattern around eyes (Figure 3). Body without (especially in juveniles) or with irregular melanocytes that are decreasing in density towards the posterior part of body or dotted (
<emphasis id="A380EAA6FFB75B2D569CFD3809655DCE" box="[1270,1295,697,717]" italics="true" pageId="7" pageNumber="71">e.g</emphasis>
., specimens from
<collectingRegion id="5330F856FFB75B2D515AFD5A0F085DEC" box="[816,866,731,751]" country="Albania" name="Vlore" pageId="7" pageNumber="71">Vlorë</collectingRegion>
) in life. Formaldehyde fixed specimens white to yellow and without pigments. In ethanol yellow or skin-coloured with pigments still present. For more pictures of life colouration see Supporting Information File S1.
</paragraph>
<paragraph id="914B36B4FFB75B2D5132FCE7087B5AE9" blockId="7.[816,1465,142,1965]" pageId="7" pageNumber="71">
<emphasis id="A380EAA6FFB75B2D5132FCE708425C79" box="[856,1064,870,890]" italics="true" pageId="7" pageNumber="71">Dentition and osteology</emphasis>
. Upper jaw with outer row of about eight (one side) medium-sized caniniforms frontally. Behind them inner small conical teeth irregularly scattered in two separate (left and right) drop-like patches medially wide about five teeth, becoming narrowed to a single row of teeth laterally. Outer row continues laterally as two large caniniforms, followed behind by four or five medium-sized caniniforms. Lower jaw with outer row of about 15 (one side) medium-sized caniniforms frontally. Behind them single broad patch of small conical inner teeth medially wide about 5
<emphasis id="A380EAA6FFB75B2D5634FBFD08005B8C" bold="true" box="[1118,1130,1148,1167]" pageId="7" pageNumber="71">–</emphasis>
6 teeth, becoming narrowed to a single row of teeth laterally. The single row of about eight larger caniniforms continuous laterally. Pharyngeal jaws with small ceratobranchial 5, having several (about 5) small, conical teeth (Figure 5a), pharyngobranchial 3 toothplate not visible on 3D models from microCT images. Number of vertebrae 35, abdominal 15 and caudal 20. The first gill arch with hemibranch,the 2nd to 4th gill arches with holobranchs. Subopercle indistinguishable from opercle, shaped as its posterior elongated extension, not forming or having subopercular spine. Six branchiostegals. Maxillary, premaxillary, nasal and ceratobranchial 5 bones shaped as in Figure 5a. Nasal bones club-shaped.
</paragraph>
</subSubSection>
<subSubSection id="D9EE653FFFB75B2D5132FA7908D45976" pageId="7" pageNumber="71" type="etymology">
<paragraph id="914B36B4FFB75B2D5132FA7908D45976" blockId="7.[816,1465,142,1965]" pageId="7" pageNumber="71">
<emphasis id="A380EAA6FFB75B2D5132FA790FA8590F" bold="true" box="[856,962,1528,1548]" pageId="7" pageNumber="71">Etymology.</emphasis>
Named
<taxonomicName id="56F44D37FFB75B2D567CFA78080E590E" authorityName="Wagner &amp; Kovac &amp; Koblmüller" authorityYear="2021" box="[1046,1124,1529,1549]" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="7" pageNumber="71" phylum="Chordata" rank="species" species="adriatica">
<emphasis id="A380EAA6FFB75B2D567CFA78080E590E" box="[1046,1124,1529,1549]" italics="true" pageId="7" pageNumber="71">adriatica</emphasis>
</taxonomicName>
, meaning belonging to the Adriatic Sea,
<emphasis id="A380EAA6FFB75B2D5134F99D0F0C592C" bold="true" box="[862,870,1564,1583]" pageId="7" pageNumber="71">“</emphasis>
Mare Adriaticum
<emphasis id="A380EAA6FFB75B2D5669F99D0861592C" bold="true" box="[1027,1035,1564,1583]" pageId="7" pageNumber="71">”</emphasis>
or
<emphasis id="A380EAA6FFB75B2D5645F99D085D592C" bold="true" box="[1071,1079,1564,1583]" pageId="7" pageNumber="71">“</emphasis>
Mare Hadriaticum
<emphasis id="A380EAA6FFB75B2D568AF99D0882592C" bold="true" box="[1248,1256,1564,1583]" pageId="7" pageNumber="71">”</emphasis>
in Latin, which is the
<typeStatus id="4E4F8816FFB75B2D515AF9BF0F335951" box="[816,857,1598,1618]" pageId="7" pageNumber="71">type</typeStatus>
locality of this species. The name is an adjective in the nominative singular (Article 11.9.1.1.,
<collectionCode id="F7E5AE71FFB75B2D562DF9E008105976" box="[1095,1146,1633,1653]" pageId="7" pageNumber="71">ICZN</collectionCode>
, 1999).
</paragraph>
</subSubSection>
<subSubSection id="D9EE653FFFB75B225132F9020C965B24" lastPageId="8" lastPageNumber="72" pageId="7" pageNumber="71" type="biology_ecology">
<paragraph id="914B36B4FFB75B225132F9020C965B24" blockId="7.[816,1465,142,1965]" lastBlockId="8.[124,773,1009,1965]" lastPageId="8" lastPageNumber="72" pageId="7" pageNumber="71">
<emphasis id="A380EAA6FFB75B2D5132F90209255994" bold="true" box="[856,1359,1667,1687]" pageId="7" pageNumber="71">Ecology and geographical distribution (Figure 1a).</emphasis>
The geographic distribution ranges from the northern Adriatic Sea to the northern Ionian Sea (Island Corfu). In the Adriatic basin, the species is broadly sympatric with
<taxonomicName id="56F44D37FFB75B2D5662F96A082659FC" baseAuthorityName="Nardo" baseAuthorityYear="1827" box="[1032,1100,1771,1791]" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="7" pageNumber="71" phylum="Chordata" rank="species" species="pigra">
<emphasis id="A380EAA6FFB75B2D5662F96A082659FC" box="[1032,1100,1771,1791]" italics="true" pageId="7" pageNumber="71">G. pigra</emphasis>
</taxonomicName>
. Quantitative data on ecology is largely lacking. At one site in the Adriatic basin (Pelješac) it was found not just in sympatry, but even syntopy, also with
<taxonomicName id="56F44D37FFB75B2D56ABF8B009535846" authority="Wagner &amp; Kovac &amp; Koblmüller, 2021" authorityName="Wagner &amp; Kovac &amp; Koblmüller" authorityYear="2021" box="[1217,1337,1841,1861]" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="7" pageNumber="71" phylum="Chordata" rank="species" species="hofrichteri" status="sp. nov.">
<emphasis id="A380EAA6FFB75B2D56ABF8B009535846" box="[1217,1337,1841,1861]" italics="true" pageId="7" pageNumber="71">G. hofrichteri</emphasis>
</taxonomicName>
<emphasis id="A380EAA6FFB75B2D572BF8B109E75847" bold="true" box="[1345,1421,1840,1860]" pageId="7" pageNumber="71">
<taxonomicNameLabel id="B8B357DDFFB75B2D572BF8B109E75847" box="[1345,1421,1840,1860]" pageId="7" pageNumber="71" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<emphasis id="A380EAA6FFB75B2D57F0F8B009C45846" box="[1434,1454,1841,1861]" italics="true" pageId="7" pageNumber="71">i.e</emphasis>
., with both species in the same habitat (Figure 1c). Fish were found in the intertidal and sometimes syntopic with
<taxonomicName id="56F44D37FFB75B2D56B2F8F709D95889" baseAuthorityName="Bonnaterre" baseAuthorityYear="1788" box="[1240,1459,1910,1930]" family="Gobiesocidae" genus="Lepadogaster" kingdom="Animalia" order="Gobiesociformes" pageId="7" pageNumber="71" phylum="Chordata" rank="species" species="lepagaster">
<emphasis id="A380EAA6FFB75B2D56B2F8F709D95889" box="[1240,1459,1910,1930]" italics="true" pageId="7" pageNumber="71">Lepadogaster lepagaster</emphasis>
</taxonomicName>
. During extreme low tide (late winter and early spring tides are the most extreme) this species was also found in layers of pebbles above the waterline.
</paragraph>
</subSubSection>
<caption id="C58B663CFFB85B225216FCC40F9A5CB2" ID-DOI="http://doi.org/10.5281/zenodo.10928932" ID-Zenodo-Dep="10928932" httpUri="https://zenodo.org/record/10928932/files/figure.png" pageId="8" pageNumber="72" startId="8.[124,136,837,856]" targetBox="[270,1320,139,795]" targetPageId="8" targetType="figure">
<paragraph id="914B36B4FFB85B225216FCC40F9A5CB2" blockId="8.[124,1466,836,945]" pageId="8" pageNumber="72">
<emphasis id="A380EAA6FFB85B225216FCC40C865C5B" bold="true" box="[124,236,837,856]" pageId="8" pageNumber="72">F I G U R E 4</emphasis>
Sucking discs and disc-papillae of
<taxonomicName id="56F44D37FFB85B22502BFCC40EB05C5A" box="[577,730,837,857]" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="8" pageNumber="72" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="A380EAA6FFB85B22502BFCC40EE15C5A" box="[577,651,837,857]" italics="true" pageId="8" pageNumber="72">Gouania</emphasis>
species.
</taxonomicName>
(a)
<taxonomicName id="56F44D37FFB85B225093FCC40F0A5C5A" authority="Wagner &amp; Kovac &amp; Koblmüller, 2021" authorityName="Wagner &amp; Kovac &amp; Koblmüller" authorityYear="2021" box="[761,864,837,857]" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="8" pageNumber="72" phylum="Chordata" rank="species" species="adriatica" status="sp. nov.">
<emphasis id="A380EAA6FFB85B225093FCC40F0A5C5A" box="[761,864,837,857]" italics="true" pageId="8" pageNumber="72">G. adriatica</emphasis>
</taxonomicName>
<emphasis id="A380EAA6FFB85B22510FFCC50FC45C5B" bold="true" box="[869,942,836,856]" pageId="8" pageNumber="72">
<taxonomicNameLabel id="B8B357DDFFB85B22510FFCC50FC45C5B" box="[869,942,836,856]" pageId="8" pageNumber="72" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(PMR VP4618
<emphasis id="A380EAA6FFB85B225656FCC408225C5B" bold="true" box="[1084,1096,837,856]" pageId="8" pageNumber="72">–</emphasis>
Holotype), (b)
<taxonomicName id="56F44D37FFB85B2256BAFCC409535C5A" authority="Wagner &amp; Kovac &amp; Koblmüller, 2021" authorityName="Wagner &amp; Kovac &amp; Koblmüller" authorityYear="2021" box="[1232,1337,837,857]" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="8" pageNumber="72" phylum="Chordata" rank="species" species="orientalis" status="sp. nov.">
<emphasis id="A380EAA6FFB85B2256BAFCC409535C5A" box="[1232,1337,837,857]" italics="true" pageId="8" pageNumber="72">G. orientalis</emphasis>
</taxonomicName>
<emphasis id="A380EAA6FFB85B225754FCC509ED5C5B" bold="true" box="[1342,1415,836,856]" pageId="8" pageNumber="72">
<taxonomicNameLabel id="B8B357DDFFB85B225754FCC509ED5C5B" box="[1342,1415,836,856]" pageId="8" pageNumber="72" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(PMR VP4585
<emphasis id="A380EAA6FFB85B22536CFCE20D785C75" bold="true" box="[262,274,867,886]" pageId="8" pageNumber="72">–</emphasis>
Holotype), (c)
<taxonomicName id="56F44D37FFB85B2253F2FCE30E665C75" authority="Wagner &amp; Kovac &amp; Koblmüller, 2021" authorityName="Wagner &amp; Kovac &amp; Koblmüller" authorityYear="2021" box="[408,524,866,886]" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="8" pageNumber="72" phylum="Chordata" rank="species" species="hofrichteri" status="sp. nov.">
<emphasis id="A380EAA6FFB85B2253F2FCE30E665C75" box="[408,524,866,886]" italics="true" pageId="8" pageNumber="72">G. hofrichteri</emphasis>
</taxonomicName>
<emphasis id="A380EAA6FFB85B22507BFCE30E305C75" bold="true" box="[529,602,866,886]" pageId="8" pageNumber="72">
<taxonomicNameLabel id="B8B357DDFFB85B22507BFCE30E305C75" box="[529,602,866,886]" pageId="8" pageNumber="72" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(PMR VP4595
<emphasis id="A380EAA6FFB85B225082FCE20E9E5C75" bold="true" box="[744,756,867,886]" pageId="8" pageNumber="72">–</emphasis>
Holotype), (d)
<taxonomicName id="56F44D37FFB85B225116FCE308285C75" authority="(Nardo 1827)" baseAuthorityName="Nardo" baseAuthorityYear="1827" box="[892,1090,866,886]" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="8" pageNumber="72" phylum="Chordata" rank="species" species="pigra">
<emphasis id="A380EAA6FFB85B225116FCE30FAB5C75" box="[892,961,866,886]" italics="true" pageId="8" pageNumber="72">G. pigra</emphasis>
(Nardo 1827)
</taxonomicName>
(ZSM-PIS-047649
<emphasis id="A380EAA6FFB85B22569FFCE2096B5C75" bold="true" box="[1269,1281,867,886]" pageId="8" pageNumber="72">–</emphasis>
Other material) and (e)
<taxonomicName id="56F44D37FFB85B2252F2FCFE0DEE5C90" authority="(Risso 1810)" baseAuthorityName="Risso" baseAuthorityYear="1810" box="[152,388,895,915]" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="8" pageNumber="72" phylum="Chordata" rank="species" species="willdenowi">
<emphasis id="A380EAA6FFB85B2252F2FCFE0D675C90" box="[152,269,895,915]" italics="true" pageId="8" pageNumber="72">G. willdenowi</emphasis>
(
<bibRefCitation id="F5654B45FFB85B22537DFCFE0D155C90" author="Risso, A." box="[279,383,895,915]" pageId="8" pageNumber="72" refId="ref28299" refString="Risso, A. (1810). Ichthyologie de Nice, ou histoire naturelle des poissons du Departement des Alpes Maritimes. Paris, France: F. Schoell." type="book" year="1810">Risso 1810</bibRefCitation>
)
</taxonomicName>
(PMR VP4574
<emphasis id="A380EAA6FFB85B22507EFC010E4A5C90" bold="true" box="[532,544,896,915]" pageId="8" pageNumber="72">–</emphasis>
Neotype). (f) Males of
<taxonomicName id="56F44D37FFB85B22509FFCFE0F555C90" authorityName="Nardo" authorityYear="1833" box="[757,831,895,915]" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="8" pageNumber="72" phylum="Chordata" rank="genus">
<emphasis id="A380EAA6FFB85B22509FFCFE0F555C90" box="[757,831,895,915]" italics="true" pageId="8" pageNumber="72">Gouania</emphasis>
</taxonomicName>
can have seemingly perfused prominent finger-like extensions on sucking disc edge in region A.A, B and C correspond to disc regions. Photographs by M. Wagner
</paragraph>
</caption>
<subSubSection id="D9EE653FFFB85B2252CEFBB708EA590E" pageId="8" pageNumber="72" type="discussion">
<paragraph id="914B36B4FFB85B2252CEFBB708EA590E" blockId="8.[124,773,1009,1965]" lastBlockId="8.[821,1469,1008,1549]" pageId="8" pageNumber="72">
<emphasis id="A380EAA6FFB85B2252CEFBB70C915B49" bold="true" box="[164,251,1078,1098]" pageId="8" pageNumber="72">Remarks.</emphasis>
<taxonomicName id="56F44D37FFB85B22536DFBB70DC95B49" authority="Wagner &amp; Kovac &amp; Koblmüller, 2021" authorityName="Wagner &amp; Kovac &amp; Koblmüller" authorityYear="2021" box="[263,419,1078,1098]" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="8" pageNumber="72" phylum="Chordata" rank="species" species="adriatica" status="sp. nov.">
<emphasis id="A380EAA6FFB85B22536DFBB70DC95B49" box="[263,419,1078,1098]" italics="true" pageId="8" pageNumber="72">Gouania adriatica</emphasis>
</taxonomicName>
<emphasis id="A380EAA6FFB85B2253C5FBB70D915B49" bold="true" box="[431,507,1078,1098]" pageId="8" pageNumber="72">
<taxonomicNameLabel id="B8B357DDFFB85B2253C5FBB70D915B49" box="[431,507,1078,1098]" pageId="8" pageNumber="72" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from slender-bodied
<taxonomicName id="56F44D37FFB85B225216FBD80D635B6E" box="[124,265,1113,1133]" pageId="8" pageNumber="72">
<emphasis id="A380EAA6FFB85B225216FBD80CA95B6E" box="[124,195,1113,1133]" italics="true" pageId="8" pageNumber="72">Gouania</emphasis>
species
</taxonomicName>
(
<taxonomicName id="56F44D37FFB85B225379FBD80D3C5B6E" baseAuthorityName="Nardo" baseAuthorityYear="1827" box="[275,342,1113,1133]" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="8" pageNumber="72" phylum="Chordata" rank="species" species="pigra">
<emphasis id="A380EAA6FFB85B225379FBD80D3C5B6E" box="[275,342,1113,1133]" italics="true" pageId="8" pageNumber="72">G. pigra</emphasis>
</taxonomicName>
and
<taxonomicName id="56F44D37FFB85B2253E8FBD80D9A5B6E" authority="Wagner &amp; Kovac &amp; Koblmüller, 2021" authorityName="Wagner &amp; Kovac &amp; Koblmüller" authorityYear="2021" box="[386,496,1113,1133]" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="8" pageNumber="72" phylum="Chordata" rank="species" species="hofrichteri" status="sp. nov.">
<emphasis id="A380EAA6FFB85B2253E8FBD80D9A5B6E" box="[386,496,1113,1133]" italics="true" pageId="8" pageNumber="72">G. hofrichteri</emphasis>
</taxonomicName>
<emphasis id="A380EAA6FFB85B22539CFBD90E565B6F" bold="true" box="[502,572,1112,1132]" pageId="8" pageNumber="72">
<taxonomicNameLabel id="B8B357DDFFB85B22539CFBD90E565B6F" box="[502,572,1112,1132]" pageId="8" pageNumber="72" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
) by a dorsal head profile forming a straight line between nape above eye and upper lip tip (
<emphasis id="A380EAA6FFB85B225087FBFA0E945B8C" box="[749,766,1147,1167]" italics="true" pageId="8" pageNumber="72">vs</emphasis>
. dorsal head profile in lateral view
<emphasis id="A380EAA6FFB85B2253C9FB1E0DC05BB1" bold="true" box="[419,426,1183,1202]" pageId="8" pageNumber="72">“</emphasis>
S
<emphasis id="A380EAA6FFB85B2253DFFB1E0DD75BB1" bold="true" box="[437,445,1183,1202]" pageId="8" pageNumber="72">”</emphasis>
curved, concave above eye and convex at nape in slender-bodied
<taxonomicName id="56F44D37FFB85B2253EEFB400E785BD6" box="[388,530,1217,1237]" pageId="8" pageNumber="72">
<emphasis id="A380EAA6FFB85B2253EEFB400DA15BD6" box="[388,459,1217,1237]" italics="true" pageId="8" pageNumber="72">Gouania</emphasis>
species
</taxonomicName>
), by a low number of vertebrae (Supporting Information Table S2; 35
<emphasis id="A380EAA6FFB85B225399FB620E6E5BF4" box="[499,516,1251,1271]" italics="true" pageId="8" pageNumber="72">vs</emphasis>
. 38
<emphasis id="A380EAA6FFB85B22504FFB650E5B5BF4" bold="true" box="[549,561,1252,1271]" pageId="8" pageNumber="72">–</emphasis>
40) and in life a star-like pigmentation around eyes (
<emphasis id="A380EAA6FFB85B225301FA870D165A19" box="[363,380,1286,1306]" italics="true" pageId="8" pageNumber="72">vs</emphasis>
. no star-like pigmentation around eyes). Ten morphometric characters, as percentages of standard length, of
<taxonomicName id="56F44D37FFB85B22509BFAA80CAF5A5C" authority="Wagner &amp; Kovac &amp; Koblmüller, 2021" authorityName="Wagner &amp; Kovac &amp; Koblmüller" authorityYear="2021" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="8" pageNumber="72" phylum="Chordata" rank="species" species="adriatica" status="sp. nov">
<emphasis id="A380EAA6FFB85B22509BFAA80CAF5A5C" italics="true" pageId="8" pageNumber="72">G. adriatica</emphasis>
</taxonomicName>
<emphasis id="A380EAA6FFB85B2252A7FACA0D7A5A5C" bold="true" box="[205,272,1355,1375]" pageId="8" pageNumber="72">
<taxonomicNameLabel id="B8B357DDFFB85B2252A7FACA0D7A5A5C" box="[205,272,1355,1375]" pageId="8" pageNumber="72" rank="species">sp. nov</taxonomicNameLabel>
</emphasis>
are nonoverlapping in range with both slender-bodied
<taxonomicName id="56F44D37FFB85B225216FAEF0CA85A81" authorityName="Nardo" authorityYear="1833" box="[124,194,1390,1410]" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="8" pageNumber="72" phylum="Chordata" rank="genus">
<emphasis id="A380EAA6FFB85B225216FAEF0CA85A81" box="[124,194,1390,1410]" italics="true" pageId="8" pageNumber="72">Gouania</emphasis>
</taxonomicName>
: head length, head width at head invagination, vertical and horizontal eye diameter, body width at pectoral fins, pectoral-fin length, prepectoral distance, ventral adhesive disc length, predisc length and caudal-fin length (values in the Table 1). There are also morphometric characters nonoverlapping in range with only one of the two slender-bodied
<taxonomicName id="56F44D37FFB85B2252D5F99A0D6C592C" authorityName="Nardo" authorityYear="1833" box="[191,262,1563,1583]" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="8" pageNumber="72" phylum="Chordata" rank="genus">
<emphasis id="A380EAA6FFB85B2252D5F99A0D6C592C" box="[191,262,1563,1583]" italics="true" pageId="8" pageNumber="72">Gouania</emphasis>
</taxonomicName>
(Table 1). In addition,
<taxonomicName id="56F44D37FFB85B2253A7F99A0E45592C" authority="Wagner &amp; Kovac &amp; Koblmüller, 2021" authorityName="Wagner &amp; Kovac &amp; Koblmüller" authorityYear="2021" box="[461,559,1563,1583]" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="8" pageNumber="72" phylum="Chordata" rank="species" species="adriatica" status="sp. nov.">
<emphasis id="A380EAA6FFB85B2253A7F99A0E45592C" box="[461,559,1563,1583]" italics="true" pageId="8" pageNumber="72">G. adriatica</emphasis>
</taxonomicName>
<emphasis id="A380EAA6FFB85B22505FF99A0E16592C" bold="true" box="[565,636,1563,1583]" pageId="8" pageNumber="72">
<taxonomicNameLabel id="B8B357DDFFB85B22505FF99A0E16592C" box="[565,636,1563,1583]" pageId="8" pageNumber="72" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from
<taxonomicName id="56F44D37FFB85B22509BF99A0CCC5951" baseAuthorityName="Nardo" baseAuthorityYear="1827" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="8" pageNumber="72" phylum="Chordata" rank="species" species="pigra">
<emphasis id="A380EAA6FFB85B22509BF99A0CCC5951" italics="true" pageId="8" pageNumber="72">G. pigra</emphasis>
</taxonomicName>
by the posterior angle of jaws extending to between a vertical line drawn through the posterior edge of the anterior nostril and a vertical line drawn through the anterior edge of the eye (
<emphasis id="A380EAA6FFB85B22507DF9020E425994" box="[535,552,1667,1687]" italics="true" pageId="8" pageNumber="72">vs</emphasis>
. posterior angle of jaws extending to, or close to, a vertical line drawn through the anterior edge of the anterior nostril) and principal caudal-fin rays 12
<emphasis id="A380EAA6FFB85B225000F9480E1C59DF" bold="true" box="[618,630,1737,1756]" pageId="8" pageNumber="72">–</emphasis>
13 (
<emphasis id="A380EAA6FFB85B2250F1F9480EC659DE" box="[667,684,1737,1757]" italics="true" pageId="8" pageNumber="72">vs</emphasis>
. principal caudal rays 10
<emphasis id="A380EAA6FFB85B225291F96D0D6D59FC" bold="true" box="[251,263,1772,1791]" pageId="8" pageNumber="72">–</emphasis>
11).
<taxonomicName id="56F44D37FFB85B225344F96A0DFA59FC" authority="Wagner &amp; Kovac &amp; Koblmüller, 2021" authorityName="Wagner &amp; Kovac &amp; Koblmüller" authorityYear="2021" box="[302,400,1771,1791]" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="8" pageNumber="72" phylum="Chordata" rank="species" species="adriatica" status="sp. nov.">
<emphasis id="A380EAA6FFB85B225344F96A0DFA59FC" box="[302,400,1771,1791]" italics="true" pageId="8" pageNumber="72">G. adriatica</emphasis>
</taxonomicName>
<emphasis id="A380EAA6FFB85B2253FFF96A0DB659FC" bold="true" box="[405,476,1771,1791]" pageId="8" pageNumber="72">
<taxonomicNameLabel id="B8B357DDFFB85B2253FFF96A0DB659FC" box="[405,476,1771,1791]" pageId="8" pageNumber="72" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
also differs from
<taxonomicName id="56F44D37FFB85B22501DF96A0E8E59FC" authority="Wagner &amp; Kovac &amp; Koblmüller, 2021" authorityName="Wagner &amp; Kovac &amp; Koblmüller" authorityYear="2021" box="[631,740,1771,1791]" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="8" pageNumber="72" phylum="Chordata" rank="species" species="hofrichteri" status="sp. nov.">
<emphasis id="A380EAA6FFB85B22501DF96A0E8E59FC" box="[631,740,1771,1791]" italics="true" pageId="8" pageNumber="72">G. hofrichteri</emphasis>
</taxonomicName>
<emphasis id="A380EAA6FFB85B225080F96A0CC95821" bold="true" pageId="8" pageNumber="72">
<taxonomicNameLabel id="B8B357DDFFB85B225080F96A0CC95821" pageId="8" pageNumber="72" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
by longitudinal infralateral and suborbital transversal rows of superficial neuromasts placed in a well-defined deep groove (
<emphasis id="A380EAA6FFB85B225014F8B00EE55846" box="[638,655,1841,1861]" italics="true" pageId="8" pageNumber="72">vs</emphasis>
. longitudinal infralateral and suborbital transversal rows of superficial neuromasts placed in shallow groove disappearing in posterior part of longitudinal infralateral row), body cross-section behind pectoral fin base half oval with straight ventral side (
<emphasis id="A380EAA6FFB85B225676FC7108475B07" box="[1052,1069,1008,1028]" italics="true" pageId="8" pageNumber="72">vs</emphasis>
. body cross-section behind pectoral fin base triangular with ventral flat and dorsal pointed), upper attachment of gill membrane opposite to 5th to 6th pectoral ray (
<emphasis id="A380EAA6FFB85B22568DFBB708925B49" box="[1255,1272,1078,1098]" italics="true" pageId="8" pageNumber="72">vs</emphasis>
. opposite to 3rd
<emphasis id="A380EAA6FFB85B2257F8FBB609F45B49" bold="true" box="[1426,1438,1079,1098]" pageId="8" pageNumber="72">–</emphasis>
4th pectoral ray), the granules on body shallow and inconspicuous (
<emphasis id="A380EAA6FFB85B22571AFBD809EB5B6E" box="[1392,1409,1113,1133]" italics="true" pageId="8" pageNumber="72">vs</emphasis>
. granules on body, at least on posterior part and nape, large and prominent).
<taxonomicName id="56F44D37FFB85B2257C3FBFA0F145BB1" authority="Wagner &amp; Kovac &amp; Koblmüller, 2021" authorityName="Wagner &amp; Kovac &amp; Koblmüller" authorityYear="2021" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="8" pageNumber="72" phylum="Chordata" rank="species" species="adriatica" status="sp. nov.">
<emphasis id="A380EAA6FFB85B2257C3FBFA0F145BB1" italics="true" pageId="8" pageNumber="72">G. adriatica</emphasis>
</taxonomicName>
<emphasis id="A380EAA6FFB85B2251EEFB1F0FA15BB1" bold="true" box="[900,971,1182,1202]" pageId="8" pageNumber="72">
<taxonomicNameLabel id="B8B357DDFFB85B2251EEFB1F0FA15BB1" box="[900,971,1182,1202]" pageId="8" pageNumber="72" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from other stout-bodied species (
<taxonomicName id="56F44D37FFB85B225752FB1F09F65BB1" authority="Wagner &amp; Kovac &amp; Koblmüller, 2021" authorityName="Wagner &amp; Kovac &amp; Koblmüller" authorityYear="2021" box="[1336,1436,1182,1202]" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="8" pageNumber="72" phylum="Chordata" rank="species" species="orientalis" status="sp. nov.">
<emphasis id="A380EAA6FFB85B225752FB1F09F65BB1" box="[1336,1436,1182,1202]" italics="true" pageId="8" pageNumber="72">G. orientalis</emphasis>
</taxonomicName>
<emphasis id="A380EAA6FFB85B2257C9FB1F0F365BD7" bold="true" pageId="8" pageNumber="72">
<taxonomicNameLabel id="B8B357DDFFB85B2257C9FB1F0F365BD7" pageId="8" pageNumber="72" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="56F44D37FFB85B2251E2FB400F925BD6" baseAuthorityName="Risso" baseAuthorityYear="1810" box="[904,1016,1217,1237]" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="8" pageNumber="72" phylum="Chordata" rank="species" species="willdenowi">
<emphasis id="A380EAA6FFB85B2251E2FB400F925BD6" box="[904,1016,1217,1237]" italics="true" pageId="8" pageNumber="72">G. willdenowi</emphasis>
</taxonomicName>
) by a posterior opercular edge with pointed upper tip and rounded lower edge (
<emphasis id="A380EAA6FFB85B225650FB6208215BF4" box="[1082,1099,1251,1271]" italics="true" pageId="8" pageNumber="72">vs</emphasis>
. posterior opercular edge w-shaped with two equally long tips) and a reduced pigmentation. In addition, it differs from
<taxonomicName id="56F44D37FFB85B22510FFAA80FA35A3E" authority="Wagner &amp; Kovac &amp; Koblmüller, 2021" authorityName="Wagner &amp; Kovac &amp; Koblmüller" authorityYear="2021" box="[869,969,1321,1341]" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="8" pageNumber="72" phylum="Chordata" rank="species" species="orientalis" status="sp. nov.">
<emphasis id="A380EAA6FFB85B22510FFAA80FA35A3E" box="[869,969,1321,1341]" italics="true" pageId="8" pageNumber="72">G. orientalis</emphasis>
</taxonomicName>
<emphasis id="A380EAA6FFB85B2251BAFAA9087D5A3F" bold="true" box="[976,1047,1320,1340]" pageId="8" pageNumber="72">
<taxonomicNameLabel id="B8B357DDFFB85B2251BAFAA9087D5A3F" box="[976,1047,1320,1340]" pageId="8" pageNumber="72" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
by principal caudal-fin rays 12
<emphasis id="A380EAA6FFB85B22574DFAA809595A3F" bold="true" box="[1319,1331,1321,1340]" pageId="8" pageNumber="72">–</emphasis>
13 (
<emphasis id="A380EAA6FFB85B22573CFAA8090D5A3E" box="[1366,1383,1321,1341]" italics="true" pageId="8" pageNumber="72">vs</emphasis>
. principal caudal rays 10
<emphasis id="A380EAA6FFB85B2251D5FACD0FA15A5C" bold="true" box="[959,971,1356,1375]" pageId="8" pageNumber="72">–</emphasis>
11) and from
<taxonomicName id="56F44D37FFB85B22563FFACA08A15A5C" baseAuthorityName="Risso" baseAuthorityYear="1810" box="[1109,1227,1355,1375]" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="8" pageNumber="72" phylum="Chordata" rank="species" species="willdenowi">
<emphasis id="A380EAA6FFB85B22563FFACA08A15A5C" box="[1109,1227,1355,1375]" italics="true" pageId="8" pageNumber="72">G. willdenowi</emphasis>
</taxonomicName>
by vertical eye diameter 3.4
<emphasis id="A380EAA6FFB85B22513AFAEE0F365A81" bold="true" box="[848,860,1391,1410]" pageId="8" pageNumber="72">–</emphasis>
4.3% and horizontal eye diameter 3.0
<emphasis id="A380EAA6FFB85B2256D8FAEE08D45A81" bold="true" box="[1202,1214,1391,1410]" pageId="8" pageNumber="72">–</emphasis>
3.7% of standard length (
<emphasis id="A380EAA6FFB85B2257CCFAEF09DD5A81" box="[1446,1463,1390,1410]" italics="true" pageId="8" pageNumber="72">vs</emphasis>
. vertical eye diameter 2.6
<emphasis id="A380EAA6FFB85B22567BFA1008775AA7" bold="true" box="[1041,1053,1425,1444]" pageId="8" pageNumber="72">–</emphasis>
3.3% and horizontal eye diameter 2.3
<emphasis id="A380EAA6FFB85B225701FA10091D5AA7" bold="true" box="[1387,1399,1425,1444]" pageId="8" pageNumber="72">–</emphasis>
2.9% of standard length).
<taxonomicName id="56F44D37FFB85B2251BAFA3208595AC4" authority="Wagner &amp; Kovac &amp; Koblmüller, 2021" authorityName="Wagner &amp; Kovac &amp; Koblmüller" authorityYear="2021" box="[976,1075,1459,1479]" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="8" pageNumber="72" phylum="Chordata" rank="species" species="adriatica" status="sp. nov.">
<emphasis id="A380EAA6FFB85B2251BAFA3208595AC4" box="[976,1075,1459,1479]" italics="true" pageId="8" pageNumber="72">G. adriatica</emphasis>
</taxonomicName>
<emphasis id="A380EAA6FFB85B225650FA3208E85AC4" bold="true" box="[1082,1154,1459,1479]" pageId="8" pageNumber="72">
<taxonomicNameLabel id="B8B357DDFFB85B225650FA3208E85AC4" box="[1082,1154,1459,1479]" pageId="8" pageNumber="72" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is known from the Adriatic Sea as well as Corfu island and has no overlapping geographic records with
<taxonomicName id="56F44D37FFB85B22515FFA780FFD590E" authority="Wagner &amp; Kovac &amp; Koblmüller, 2021" authorityName="Wagner &amp; Kovac &amp; Koblmüller" authorityYear="2021" box="[821,919,1529,1549]" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="8" pageNumber="72" phylum="Chordata" rank="species" species="orientalis" status="sp. nov.">
<emphasis id="A380EAA6FFB85B22515FFA780FFD590E" box="[821,919,1529,1549]" italics="true" pageId="8" pageNumber="72">G. orientalis</emphasis>
</taxonomicName>
<emphasis id="A380EAA6FFB85B2251F7FA790F88590F" bold="true" box="[925,994,1528,1548]" pageId="8" pageNumber="72">
<taxonomicNameLabel id="B8B357DDFFB85B2251F7FA790F88590F" box="[925,994,1528,1548]" pageId="8" pageNumber="72" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="56F44D37FFB85B225666FA780811590E" baseAuthorityName="Risso" baseAuthorityYear="1810" box="[1036,1147,1529,1549]" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="8" pageNumber="72" phylum="Chordata" rank="species" species="willdenowi">
<emphasis id="A380EAA6FFB85B225666FA780811590E" box="[1036,1147,1529,1549]" italics="true" pageId="8" pageNumber="72">G. willdenowi</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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