347 lines
35 KiB
XML
347 lines
35 KiB
XML
<document id="C52420487A69F0F2DD2CCB3F9F4253FC" ID-DOI="10.1111/jfb.14558" ID-ISSN="1095-8649" ID-PMC="PMC7821206" ID-PubMed="32985685" ID-Zenodo-Dep="10928926" IM.bibliography_approvedBy="julia" IM.illustrations_approvedBy="julia" IM.materialsCitations_approvedBy="julia" IM.tables_approvedBy="julia" IM.taxonomicNames_approvedBy="julia" IM.treatmentCitations_approvedBy="julia" IM.treatments_approvedBy="julia" checkinTime="1712261492542" checkinUser="felipe" docAuthor="Wagner, Maximilian, Kovac, Marcelo & Koblmüller, Stephan" docDate="2021" docId="195D87A2FFB55B2C521DFE940D005BD7" docLanguage="en" docName="JFishBiol.98.1.64-88.pdf" docOrigin="Journal of Fish Biology 98 (1)" docSource="http://dx.doi.org/10.1111/jfb.14558" docStyle="DocumentStyle:E312FF628AF650F200AD49ACE691BDEB.1:JFishBiol.2018-.journal_article.open" docStyleId="E312FF628AF650F200AD49ACE691BDEB" docStyleName="JFishBiol.2018-.journal_article.open" docStyleVersion="1" docTitle="Gouania Nardo 1833" docType="treatment" docVersion="3" lastPageNumber="70" masterDocId="E564FFDAFFB05B2A526AFF810C6A5F03" masterDocTitle="Unravelling the taxonomy of an interstitial fish radiation: Three new species of Gouania (Teleostei: Gobiesocidae) from the Mediterranean Sea and redescriptions of G. willdenowi and G. pigra" masterLastPageNumber="88" masterPageNumber="64" pageNumber="69" updateTime="1712674921547" updateUser="julia" zenodo-license-document="CC-BY-NC-ND-4.0">
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<mods:title id="FAD4C6B544EF41015BC1C6C7263FD409">Unravelling the taxonomy of an interstitial fish radiation: Three new species of Gouania (Teleostei: Gobiesocidae) from the Mediterranean Sea and redescriptions of G. willdenowi and G. pigra</mods:title>
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<mods:namePart id="0FF9763357C99D00090216EAA806024F">Wagner, Maximilian</mods:namePart>
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<mods:affiliation id="675E0CC4BE0A629300D38D75AF0C851F">Institute of Biology, University of Graz, Graz, Austria & Department of Biology, University of Antwerp, Antwerp, Belgium</mods:affiliation>
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<mods:namePart id="47CE8FD8BFBCE46372FD1A021DEF0E26">Kovac, Marcelo</mods:namePart>
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<mods:affiliation id="7AA2B0884748F56D7EF49E20FA551F52">Natural History Museum Rijeka, Rijeka, Croatia</mods:affiliation>
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<mods:namePart id="739E06FE4955C1BFD92EBC9C07C35FB0">Koblmüller, Stephan</mods:namePart>
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<mods:affiliation id="535B7D171C76F2AE1BC3590F52389E7C">Institute of Biology, University of Graz, Graz, Austria</mods:affiliation>
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<mods:title id="26697E470A4923A5EC1A4739640F56A8">Journal of Fish Biology</mods:title>
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<mods:date id="1EB4D3EC33D669079ADA92DD611BC21D">2021</mods:date>
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<mods:number id="D8B5CB3E436B59A9667C2213B6C1B057">2020-11-03</mods:number>
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<heading id="CA0381D8FFB55B2F521DFE940D9B5E2D" box="[119,497,276,302]" fontSize="10" level="3" pageId="5" pageNumber="69" reason="9">
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<emphasis id="A380EAA6FFB55B2F521DFE940CDB5E2D" bold="true" box="[119,177,277,302]" pageId="5" pageNumber="69">3.1.1</emphasis>
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<taxonomicName id="56F44D37FFB55B2F5292FE950D9B5E2D" ID-CoL="4QGL" ID-ENA="106678" authority="Nardo, 1833" authorityName="Nardo" authorityYear="1833" box="[248,497,276,302]" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="5" pageNumber="69" phylum="Chordata" rank="genus">
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<emphasis id="A380EAA6FFB55B2F5292FE950D9B5E2D" bold="true" box="[248,497,276,302]" pageId="5" pageNumber="69">
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<emphasis id="A380EAA6FFB55B2F5292FE950D3C5E2D" bold="true" box="[248,342,276,302]" italics="true" pageId="5" pageNumber="69">Gouania</emphasis>
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<bibRefCitation id="F5654B45FFB55B2F5334FE940D9B5E2D" author="Nardo, G. D." box="[350,497,277,302]" pageId="5" pageNumber="69" pagination="548 - 549" refId="ref28068" refString="Nardo, G. D. (1833). Eine neue Art von Lepadogaster (L. piger). Isis (Oken), 26, 548 - 549." type="journal article" year="1833">Nardo, 1833</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<taxonomicName id="56F44D37FFB55B2F521DFEDF0DFB5E71" ID-CoL="4QGL" authority="Nardo 1833: 548" authorityName="Nardo" authorityPageNumber="548" authorityYear="1833" box="[119,401,350,370]" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="5" pageNumber="69" phylum="Chordata" rank="genus">
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<emphasis id="A380EAA6FFB55B2F521DFEDF0CAB5E71" box="[119,193,350,370]" italics="true" pageId="5" pageNumber="69">Gouania</emphasis>
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<bibRefCitation id="F5654B45FFB55B2F52BFFEDF0DFB5E71" author="Nardo, G. D." box="[213,401,350,370]" pageId="5" pageNumber="69" pagination="548 - 549" refId="ref28068" refString="Nardo, G. D. (1833). Eine neue Art von Lepadogaster (L. piger). Isis (Oken), 26, 548 - 549." type="journal article" year="1833">Nardo 1833: 548</bibRefCitation>
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</taxonomicName>
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</paragraph>
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(
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<typeStatus id="4E4F8816FFB55B2F53C3FEDF0DB95E71" box="[425,467,350,370]" pageId="5" pageNumber="69">type</typeStatus>
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species:
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<taxonomicName id="56F44D37FFB55B2F5028FEDF0C835E96" ID-CoL="3H4Q7" authority="Nardo 1833" authorityName="Nardo" authorityYear="1833" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="5" pageNumber="69" phylum="Chordata" rank="species" species="prototypus">
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<emphasis id="A380EAA6FFB55B2F5028FEDF0E955E71" box="[578,767,350,370]" italics="true" pageId="5" pageNumber="69">Gouania prototypus</emphasis>
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<bibRefCitation id="F5654B45FFB55B2F521DFE000C835E96" author="Nardo, G. D." box="[119,233,385,405]" pageId="5" pageNumber="69" pagination="548 - 549" refId="ref28068" refString="Nardo, G. D. (1833). Eine neue Art von Lepadogaster (L. piger). Isis (Oken), 26, 548 - 549." type="journal article" year="1833">Nardo 1833</bibRefCitation>
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</taxonomicName>
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by original designation. Genus appeared first as
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<taxonomicName id="56F44D37FFB55B2F50D9FE000E915E96" authorityName="Nardo" authorityYear="1833" box="[691,763,385,405]" family="Gobiesocidae" genus="Covania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Gobiesociformes" pageId="5" pageNumber="69" phylum="Chordata" rank="genus">
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<emphasis id="A380EAA6FFB55B2F50D9FE000E915E96" box="[691,763,385,405]" italics="true" pageId="5" pageNumber="69">Covania</emphasis>
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</taxonomicName>
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, name corrected by
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<treatmentCitation id="105510A5FFB55B2F5340FE220D875EB4" author="Canestrini, G." box="[298,493,419,439]" page="181" pageId="5" pageNumber="69" year="1864">
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<bibRefCitation id="F5654B45FFB55B2F5340FE220D875EB4" author="Canestrini, G." box="[298,493,419,439]" pageId="5" pageNumber="69" pagination="177 - 196" refId="ref25110" refString="Canestrini, G. (1864). Studi sui Lepadogaster del Mediterraneo. Archivo per la Zoologia, l'anatomia e la Fisiologia, 3, 177 - 196." type="journal article" year="1864">Canestrini, 1864:181</bibRefCitation>
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</treatmentCitation>
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).
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</paragraph>
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||
</subSubSection>
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<subSubSection id="D9EE653FFFB55B2C52F5FE470D785E09" lastPageId="6" lastPageNumber="70" pageId="5" pageNumber="69" type="diagnosis">
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<paragraph id="914B36B4FFB55B2C52F5FE470D785E09" blockId="5.[119,767,350,509]" lastBlockId="6.[124,772,142,1237]" lastPageId="6" lastPageNumber="70" pageId="5" pageNumber="69">
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<emphasis id="A380EAA6FFB55B2F52F5FE470D6B5ED9" bold="true" box="[159,257,454,474]" pageId="5" pageNumber="69">Diagnosis.</emphasis>
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The genus diagnosis is based on the genus description by
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<bibRefCitation id="F5654B45FFB55B2F52FFFE680D7B5EFE" author="Briggs, J. C." box="[149,273,489,509]" pageId="5" pageNumber="69" refId="ref24998" refString="Briggs, J. C. (1955). A monograph of the clingfishes (order Xenopterygii). Stanford, CA: Stanford Ichthyological Bulletin." type="book" year="1955">Briggs (1955)</bibRefCitation>
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and adjusted to fit the new species: dorsal and anal fins reduced to low ridges with very weak rays, connected to the caudal fin. Ventral adhesive disc of
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double
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<emphasis id="A380EAA6FFB55B2F56DDFF3008D55FC7" bold="true" box="[1207,1215,177,196]" pageId="5" pageNumber="69">„</emphasis>
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<typeStatus id="4E4F8816FFB55B2F56A3FF30089E5FC6" box="[1225,1268,177,197]" pageId="5" pageNumber="69">type</typeStatus>
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, with no papillae in region A and flattened papillae in regions B and C. Disc small, 5.0
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<emphasis id="A380EAA6FFB55B2F57E4FF5509F05FE4" bold="true" box="[1422,1434,212,231]" pageId="5" pageNumber="69">–</emphasis>
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standard length. Body slender and elongated, posteriorly laterally compressed. Head rounded in dorsal outline; snout not produced. Upper jaw with outer row of medium-sized caniniforms frontally. Behind them irregularly scattered small conical inner teeth. Outer row continues laterally as large caniniforms, followed behind by medium-sized caniniforms. Lower jaw with outer row of medium-sized caniniforms frontally. Behind them irregularly scattered small conical inner teeth. The single row of larger caniniforms continuous laterally. Vertebrae 35
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<emphasis id="A380EAA6FFB65B2C5292FF0E0D6E5FA1" bold="true" box="[248,260,143,162]" pageId="6" pageNumber="70">–</emphasis>
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40. The first gill arch with hemibranch, the 2nd to 4th gill arches with holobranchs. No fleshy pad present on lower pectoral base. No subopercular spine. Gill membranes attached to isthmus. Six branchiostegals.
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</paragraph>
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<emphasis id="A380EAA6FFB55B2F521DFD480C8D5DDF" bold="true" box="[119,231,713,732]" pageId="5" pageNumber="69">
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F I G U
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<collectionCode id="F7E5AE71FFB55B2F52DFFD480CA95DDF" box="[181,195,713,732]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="5" pageNumber="69">R</collectionCode>
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E 2
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</emphasis>
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Comparative morphological overview and lateral line system. (a) Main morphological characteristics of head region of (a)
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<taxonomicName id="56F44D37FFB55B2F5374FCBF0DEE5C51" authority="Wagner & Kovac & Koblmüller, 2021" authorityName="Wagner & Kovac & Koblmüller" authorityYear="2021" box="[286,388,830,850]" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="5" pageNumber="69" phylum="Chordata" rank="species" species="adriatica" status="sp.">
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<emphasis id="A380EAA6FFB55B2F5374FCBF0DEE5C51" box="[286,388,830,850]" italics="true" pageId="5" pageNumber="69">G. adriatica</emphasis>
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</taxonomicName>
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</emphasis>
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<emphasis id="A380EAA6FFB55B2F521DFCDA0CCA5C6C" bold="true" box="[119,160,859,879]" pageId="5" pageNumber="69">nov.</emphasis>
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(
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<collectionCode id="F7E5AE71FFB55B2F52C0FCDA0CB25C6C" box="[170,216,859,879]" country="0" httpUri="http://grbio.org/cool/xq4z-q5gq" name="Prirodoslovni muzej Rijeka" pageId="5" pageNumber="69">PMR</collectionCode>
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<specimenCode id="C1529ECFFFB55B2F52B4FCDA0D435C6C" box="[222,297,859,879]" collectionCode="VP" pageId="5" pageNumber="69">VP4618</specimenCode>
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<emphasis id="A380EAA6FFB55B2F5345FCDD0D515C6C" bold="true" box="[303,315,860,879]" pageId="5" pageNumber="69">–</emphasis>
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<typeStatus id="4E4F8816FFB55B2F532BFCDA0DF05C6C" box="[321,410,859,879]" pageId="5" pageNumber="69" type="holotype">Holotype</typeStatus>
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)
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</materialsCitation>
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, (b)
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||
<taxonomicName id="56F44D37FFB55B2F52FEFCF90C975C8F" authority="Wagner & Kovac & Koblmüller, 2021" authorityName="Wagner & Kovac & Koblmüller" authorityYear="2021" box="[148,253,888,908]" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="5" pageNumber="69" phylum="Chordata" rank="species" species="orientalis" status="sp. nov.">
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<emphasis id="A380EAA6FFB55B2F52FEFCF90C975C8F" box="[148,253,888,908]" italics="true" pageId="5" pageNumber="69">G. orientalis</emphasis>
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||
</taxonomicName>
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<taxonomicNameLabel id="B8B357DDFFB55B2F5368FCF90D215C8F" box="[258,331,888,908]" pageId="5" pageNumber="69" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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||
(
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||
<collectionCode id="F7E5AE71FFB55B2F533EFCF90DE85C8F" box="[340,386,888,908]" country="0" httpUri="http://grbio.org/cool/xq4z-q5gq" name="Prirodoslovni muzej Rijeka" pageId="5" pageNumber="69">PMR</collectionCode>
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|
||
<specimenCode id="C1529ECFFFB55B2F521DFC170CA95CA9" box="[119,195,918,938]" collectionCode="VP" pageId="5" pageNumber="69">VP4585</specimenCode>
|
||
<emphasis id="A380EAA6FFB55B2F52A2FC170CBE5CAA" bold="true" box="[200,212,918,937]" pageId="5" pageNumber="69">–</emphasis>
|
||
<typeStatus id="4E4F8816FFB55B2F52B0FC170D595CA9" box="[218,307,918,938]" pageId="5" pageNumber="69" type="holotype">Holotype</typeStatus>
|
||
), (c)
|
||
<taxonomicName id="56F44D37FFB55B2F5331FC170D055CA9" authority="Wagner & Kovac & Koblmüller, 2021" authorityName="Wagner & Kovac & Koblmüller" authorityYear="2021" box="[347,367,918,938]" pageId="5" pageNumber="69" status="sp. nov.">
|
||
<emphasis id="A380EAA6FFB55B2F5331FC170D055CA9" box="[347,367,918,938]" italics="true" pageId="5" pageNumber="69">G.</emphasis>
|
||
</taxonomicName>
|
||
</materialsCitation>
|
||
<emphasis id="A380EAA6FFB55B2F521DFC320CB85CC4" box="[119,210,947,967]" italics="true" pageId="5" pageNumber="69">hofrichteri</emphasis>
|
||
<emphasis id="A380EAA6FFB55B2F52B2FC320D4A5CC4" bold="true" box="[216,288,947,967]" pageId="5" pageNumber="69">
|
||
<taxonomicNameLabel id="B8B357DDFFB55B2F52B2FC320D4A5CC4" box="[216,288,947,967]" pageId="5" pageNumber="69" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<collectionCode id="F7E5AE71FFB55B2F5340FC320D325CC4" box="[298,344,947,967]" country="0" httpUri="http://grbio.org/cool/xq4z-q5gq" name="Prirodoslovni muzej Rijeka" pageId="5" pageNumber="69">PMR</collectionCode>
|
||
<collectionCode id="F7E5AE71FFB55B2F5334FC320D105CC4" box="[350,378,947,967]" pageId="5" pageNumber="69">VP</collectionCode>
|
||
4595
|
||
<emphasis id="A380EAA6FFB55B2F521DFC500CE95CE7" bold="true" box="[119,131,977,996]" pageId="5" pageNumber="69">–</emphasis>
|
||
<typeStatus id="4E4F8816FFB55B2F52E3FC510C8A5CE7" box="[137,224,976,996]" pageId="5" pageNumber="69" type="holotype">Holotype</typeStatus>
|
||
) and (d)
|
||
<taxonomicName id="56F44D37FFB55B2F5344FC510DD35CE7" box="[302,441,976,996]" pageId="5" pageNumber="69">
|
||
<emphasis id="A380EAA6FFB55B2F5344FC510D195CE7" box="[302,371,976,996]" italics="true" pageId="5" pageNumber="69">G. pigra</emphasis>
|
||
(Nardo
|
||
</taxonomicName>
|
||
<materialsCitation id="219C3CE9FFB55B2F521DFC6F0DCD5B01" box="[119,423,1006,1026]" collectionCode="PMR" pageId="5" pageNumber="69" specimenCode="VP3529" specimenCount="1" typeStatus="neotype">
|
||
1827) (
|
||
<collectionCode id="F7E5AE71FFB55B2F52D2FC6F0C8C5B01" box="[184,230,1006,1026]" country="0" httpUri="http://grbio.org/cool/xq4z-q5gq" name="Prirodoslovni muzej Rijeka" pageId="5" pageNumber="69">PMR</collectionCode>
|
||
<specimenCode id="C1529ECFFFB55B2F5286FC6F0D525B01" box="[236,312,1006,1026]" collectionCode="VP" pageId="5" pageNumber="69">VP3529</specimenCode>
|
||
<emphasis id="A380EAA6FFB55B2F5357FC6F0D235B02" bold="true" box="[317,329,1006,1025]" pageId="5" pageNumber="69">–</emphasis>
|
||
<typeStatus id="4E4F8816FFB55B2F5325FC6F0DC85B01" box="[335,418,1006,1026]" pageId="5" pageNumber="69" type="neotype">Neotype</typeStatus>
|
||
)
|
||
</materialsCitation>
|
||
. (e) Position of pores (bold) and neuromasts (italic) shown on the example of
|
||
<taxonomicName id="56F44D37FFB55B2F5289FBC70D325B59" box="[227,344,1094,1114]" pageId="5" pageNumber="69">
|
||
<emphasis id="A380EAA6FFB55B2F5289FBC70D325B59" box="[227,344,1094,1114]" italics="true" pageId="5" pageNumber="69">G. willdenowi</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="F5654B45FFB55B2F5216FBE20C8E5B74" author="Risso, A." box="[124,228,1123,1143]" pageId="5" pageNumber="69" refId="ref28299" refString="Risso, A. (1810). Ichthyologie de Nice, ou histoire naturelle des poissons du Departement des Alpes Maritimes. Paris, France: F. Schoell." type="book" year="1810">Risso 1810</bibRefCitation>
|
||
) (
|
||
<collectionCode id="F7E5AE71FFB55B2F529EFBE20D485B74" box="[244,290,1123,1143]" country="0" httpUri="http://grbio.org/cool/xq4z-q5gq" name="Prirodoslovni muzej Rijeka" pageId="5" pageNumber="69">PMR</collectionCode>
|
||
<collectionCode id="F7E5AE71FFB55B2F5342FBE20D2E5B74" box="[296,324,1123,1143]" pageId="5" pageNumber="69">VP</collectionCode>
|
||
4574
|
||
<emphasis id="A380EAA6FFB55B2F5313FBE50DEF5B74" bold="true" box="[377,389,1124,1143]" pageId="5" pageNumber="69">–</emphasis>
|
||
<typeStatus id="4E4F8816FFB55B2F521DFB010CA25B97" box="[119,200,1152,1172]" pageId="5" pageNumber="69" type="neotype">Neotype</typeStatus>
|
||
) and the main morphological characteristics in the head region of this species. (f) Longitudinal infralateral and suborbital transversal rows of superficial neuromasts can be placed on the well-defined bottom of a deep (+) or shallow (
|
||
<emphasis id="A380EAA6FFB55B2F5330FACF0D0C5A62" bold="true" box="[346,358,1358,1377]" pageId="5" pageNumber="69">–</emphasis>
|
||
) groove. U, upper opercular tip; Abbreviations: L, lower opercular tip; SR, supralabial row; NR, nasal row; LIR, longitudinal infralateral; STR, suborbital transversal row; POR, postorbital transversal row; PTR, preopercular transversal row; SLR, subopercular longitudinal row; MR, mandibular row; AVR, anterior ventral row; PVR, posterior ventral row; ADR, anterior dorsal row; PDR, posterior dorsal row; HR, hyomandibular row; SR1, supraopercular row; SR2, suprapectoral row; DLR, dorsolateral longitudinal row; VLR, ventrolateral longitudinal row. Photographs by M. Wagner and M. Kovacˇic
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="D9EE653FFFB65B2C52CEFE990D005BD7" pageId="6" pageNumber="70" type="key">
|
||
<paragraph id="914B36B4FFB65B2C52CEFE990E745E2E" blockId="6.[124,772,142,1237]" box="[164,542,280,301]" pageId="6" pageNumber="70">
|
||
<emphasis id="A380EAA6FFB65B2C52CEFE990DD15E2F" bold="true" box="[164,443,280,300]" pageId="6" pageNumber="70">
|
||
Key to the species of
|
||
<taxonomicName id="56F44D37FFB65B2C531AFE990DD15E2F" ID-CoL="4QGL" authorityName="Nardo" authorityYear="1833" box="[368,443,280,300]" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="6" pageNumber="70" phylum="Chordata" rank="genus">
|
||
<emphasis id="A380EAA6FFB65B2C531AFE990DD15E2F" bold="true" box="[368,443,280,300]" italics="true" pageId="6" pageNumber="70">Gouania</emphasis>
|
||
</taxonomicName>
|
||
</emphasis>
|
||
(Figure 2).
|
||
</paragraph>
|
||
<key id="E5656185FFB65B2C52CEFEBA0D0C5BD7" pageId="6" pageNumber="70">
|
||
<keyStep id="2A002E11FFB65B2C52CEFEBA0EEC5EFE" pageId="6" pageNumber="70">
|
||
<paragraph id="914B36B4FFB65B2C52CEFEBA0EE75E96" blockId="6.[124,772,142,1237]" pageId="6" pageNumber="70">
|
||
<keyLead id="2A059581FFB65B2C52CEFEBA0EE75E96" pageId="6" pageNumber="70">
|
||
1a. Dorsal head profile concave above eye (Figure 2c,d), caudal-fin length 11.1
|
||
<emphasis id="A380EAA6FFB65B2C5367FEDE0D735E71" bold="true" box="[269,281,351,370]" pageId="6" pageNumber="70">–</emphasis>
|
||
13.0% of standard length, pectoral-fin length 5.4
|
||
<emphasis id="A380EAA6FFB65B2C509DFEDE0F695E71" bold="true" box="[759,771,351,370]" pageId="6" pageNumber="70">–</emphasis>
|
||
7.5% of standard length,
|
||
<emphasis id="A380EAA6FFB65B2C5308FE000D005E97" bold="true" box="[354,362,385,404]" pageId="6" pageNumber="70">“</emphasis>
|
||
slender
|
||
<emphasis id="A380EAA6FFB65B2C53C7FE000DDF5E97" bold="true" box="[429,437,385,404]" pageId="6" pageNumber="70">”</emphasis>
|
||
species of
|
||
<taxonomicName id="56F44D37FFB65B2C5076FE000E0C5E96" ID-CoL="4QGL" authorityName="Nardo" authorityYear="1833" box="[540,614,385,405]" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="6" pageNumber="70" phylum="Chordata" rank="genus">
|
||
<emphasis id="A380EAA6FFB65B2C5076FE000E0C5E96" box="[540,614,385,405]" italics="true" pageId="6" pageNumber="70">Gouania</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A380EAA6FFB65B2C5001FE000E115E97" bold="true" box="[619,635,385,404]" pageId="6" pageNumber="70">…</emphasis>
|
||
2
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="914B36B4FFB65B2C52CEFE220EEC5EFE" blockId="6.[124,772,142,1237]" pageId="6" pageNumber="70">
|
||
<keyLead id="2A059581FFB65B2C52CEFE220EEC5EFE" pageId="6" pageNumber="70">
|
||
1b. Dorsal head profile straight above eye (Figure 2a,b,e), caudal-fin length 13.5
|
||
<emphasis id="A380EAA6FFB65B2C5367FE460D735ED9" bold="true" box="[269,281,455,474]" pageId="6" pageNumber="70">–</emphasis>
|
||
17.5% of standard length, pectoral-fin length 8.2
|
||
<emphasis id="A380EAA6FFB65B2C509DFE460F695ED9" bold="true" box="[759,771,455,474]" pageId="6" pageNumber="70">–</emphasis>
|
||
11.3% of standard length,
|
||
<emphasis id="A380EAA6FFB65B2C5304FE680D1C5EFF" bold="true" box="[366,374,489,508]" pageId="6" pageNumber="70">“</emphasis>
|
||
stout
|
||
<emphasis id="A380EAA6FFB65B2C53CCFE680DC45EFF" bold="true" box="[422,430,489,508]" pageId="6" pageNumber="70">”</emphasis>
|
||
species of
|
||
<taxonomicName id="56F44D37FFB65B2C507FFE680E355EFE" ID-CoL="4QGL" authorityName="Nardo" authorityYear="1833" box="[533,607,489,509]" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="6" pageNumber="70" phylum="Chordata" rank="genus">
|
||
<emphasis id="A380EAA6FFB65B2C507FFE680E355EFE" box="[533,607,489,509]" italics="true" pageId="6" pageNumber="70">Gouania</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A380EAA6FFB65B2C500EFE680E1E5EFF" bold="true" box="[612,628,489,508]" pageId="6" pageNumber="70">…</emphasis>
|
||
3
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="2A002E11FFB65B2C52CEFD8A0EF55C9F" pageId="6" pageNumber="70">
|
||
<paragraph id="914B36B4FFB65B2C52CEFD8A0C9D5DA9" blockId="6.[124,772,142,1237]" pageId="6" pageNumber="70">
|
||
<keyLead id="2A059581FFB65B2C52CEFD8A0C9D5DA9" pageId="6" pageNumber="70">
|
||
2a. Posterior angle of jaws extends to, or close to, a vertical line drawn through the anterior edge of the anterior nostril; longitudinal infralateral and suborbital transversal rows of superficial neuromasts placed in the well-defined deep groove (Figure 2d,f); Adriatic Sea
|
||
<emphasis id="A380EAA6FFB65B2C5099FDF50F695D84" bold="true" box="[755,771,628,647]" pageId="6" pageNumber="70">…</emphasis>
|
||
<taxonomicName id="56F44D37FFB65B2C5216FD170C9D5DA9" ID-CoL="3H4M9" baseAuthorityName="Nardo" baseAuthorityYear="1827" box="[124,247,662,682]" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="6" pageNumber="70" phylum="Chordata" rank="species" species="pigra">
|
||
<emphasis id="A380EAA6FFB65B2C5216FD170C9D5DA9" box="[124,247,662,682]" italics="true" pageId="6" pageNumber="70">Gouania pigra</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="914B36B4FFB65B2C52CEFD380EF55C9F" blockId="6.[124,772,142,1237]" pageId="6" pageNumber="70">
|
||
<keyLead id="2A059581FFB65B2C52CEFD380EF55C9F" pageId="6" pageNumber="70">
|
||
2b. Posterior angle of jaws extends to between a vertical line drawn through posterior edge of anterior nostril and a vertical line drawn through anterior edge of eye; longitudinal infralateral and suborbital transversal rows of superficial neuromasts placed in shallow groove disappearing in posterior part of longitudinal infralateral row (Figure 2c,f); Aegean Sea, rare in the south Adriatic Sea (a single record from Pelješac,
|
||
<collectingCountry id="E9E37624FFB65B2C5329FC080DED5C9E" box="[323,391,905,925]" name="Croatia" pageId="6" pageNumber="70">Croatia</collectingCountry>
|
||
)
|
||
<emphasis id="A380EAA6FFB65B2C53F8FC080DC85C9F" bold="true" box="[402,418,905,924]" pageId="6" pageNumber="70">…</emphasis>
|
||
<taxonomicName id="56F44D37FFB65B2C53CDFC080E385C9E" authority="Wagner & Kovac & Koblmüller, 2021" authorityName="Wagner & Kovac & Koblmüller" authorityYear="2021" box="[423,594,905,925]" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="6" pageNumber="70" phylum="Chordata" rank="species" species="hofrichteri" status="sp. nov.">
|
||
<emphasis id="A380EAA6FFB65B2C53CDFC080E385C9E" box="[423,594,905,925]" italics="true" pageId="6" pageNumber="70">Gouania hofrichteri</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A380EAA6FFB65B2C503DFC090EF55C9F" bold="true" box="[599,671,904,924]" pageId="6" pageNumber="70">
|
||
<taxonomicNameLabel id="B8B357DDFFB65B2C503DFC090EF55C9F" box="[599,671,904,924]" pageId="6" pageNumber="70" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<caption id="C58B663CFFB65B2C515FFD63097E5C4E" ID-DOI="http://doi.org/10.5281/zenodo.10928930" ID-Zenodo-Dep="10928930" httpUri="https://zenodo.org/record/10928930/files/figure.png" pageId="6" pageNumber="70" startId="6.[821,833,738,757]" targetBox="[826,1463,141,694]" targetPageId="6" targetType="figure">
|
||
<paragraph id="914B36B4FFB65B2C515FFD63097E5C4E" blockId="6.[821,1466,737,845]" pageId="6" pageNumber="70">
|
||
<emphasis id="A380EAA6FFB65B2C515FFD630FCE5DF6" bold="true" box="[821,932,738,757]" pageId="6" pageNumber="70">F I G U R E 3</emphasis>
|
||
<taxonomicName id="56F44D37FFB65B2C51A8FD60080A5DF6" authority="Wagner & Kovac & Koblmüller, 2021" authorityName="Wagner & Kovac & Koblmüller" authorityYear="2021" box="[962,1120,737,757]" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="6" pageNumber="70" phylum="Chordata" rank="species" species="adriatica" status="sp. nov.">
|
||
<emphasis id="A380EAA6FFB65B2C51A8FD60080A5DF6" box="[962,1120,737,757]" italics="true" pageId="6" pageNumber="70">Gouania adriatica</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A380EAA6FFB65B2C560FFD6008D85DF6" bold="true" box="[1125,1202,737,757]" pageId="6" pageNumber="70">
|
||
<taxonomicNameLabel id="B8B357DDFFB65B2C560FFD6008C55DF6" box="[1125,1199,737,757]" pageId="6" pageNumber="70" rank="species">sp. nov.</taxonomicNameLabel>
|
||
,
|
||
</emphasis>
|
||
PMR VP4618, holotype, male, 41.41+6.77 mm, Stoja, Pula. Lateral view of specimen preserved in 4% formaldehyde (top). Lateral, dorsal and ventral view, alive (below). Photographs by M. Wagner and M. Kovacˇic
|
||
</paragraph>
|
||
</caption>
|
||
<keyStep id="2A002E11FFB65B2C52CEFC2A0D7F5B49" pageId="6" pageNumber="70">
|
||
<paragraph id="914B36B4FFB65B2C52CEFC2A0E7C5B07" blockId="6.[124,772,142,1237]" pageId="6" pageNumber="70">
|
||
<keyLead id="2A059581FFB65B2C52CEFC2A0E7C5B07" pageId="6" pageNumber="70">
|
||
3a. Posterior opercular edge with pointed upper tip and rounded lower posterior edge (Figure 2a); Adriatic Sea, northern Ionian Sea (Island of Corfu)
|
||
<emphasis id="A380EAA6FFB65B2C537FFC700D4F5B07" bold="true" box="[277,293,1009,1028]" pageId="6" pageNumber="70">…</emphasis>
|
||
<taxonomicName id="56F44D37FFB65B2C5341FC700DA25B06" authority="Wagner & Kovac & Koblmüller, 2021" authorityName="Wagner & Kovac & Koblmüller" authorityYear="2021" box="[299,456,1009,1029]" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="6" pageNumber="70" phylum="Chordata" rank="species" species="adriatica" status="sp. nov.">
|
||
<emphasis id="A380EAA6FFB65B2C5341FC700DA25B06" box="[299,456,1009,1029]" italics="true" pageId="6" pageNumber="70">Gouania adriatica</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A380EAA6FFB65B2C53A7FC710E7C5B07" bold="true" box="[461,534,1008,1028]" pageId="6" pageNumber="70">
|
||
<taxonomicNameLabel id="B8B357DDFFB65B2C53A7FC710E7C5B07" box="[461,534,1008,1028]" pageId="6" pageNumber="70" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="914B36B4FFB65B2C52CEFB920D7F5B49" blockId="6.[124,772,142,1237]" pageId="6" pageNumber="70">
|
||
<keyLead id="2A059581FFB65B2C52CEFB920D7F5B49" pageId="6" pageNumber="70">
|
||
3b. Posterior opercular edge with two equally long tips (Figure 2b,e)
|
||
<emphasis id="A380EAA6FFB65B2C529EFBB60D6E5B49" bold="true" box="[244,260,1079,1098]" pageId="6" pageNumber="70">…</emphasis>
|
||
4
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="2A002E11FFB65B2C52CEFBD80D005BD7" pageId="6" pageNumber="70">
|
||
<paragraph id="914B36B4FFB65B2C52CEFBD80CB35B8C" blockId="6.[124,772,142,1237]" pageId="6" pageNumber="70">
|
||
<keyLead id="2A059581FFB65B2C52CEFBD80CB35B8C" pageId="6" pageNumber="70">
|
||
4a. Vertebrae 37
|
||
<emphasis id="A380EAA6FFB65B2C532AFBD80D265B6F" bold="true" box="[320,332,1113,1132]" pageId="6" pageNumber="70">–</emphasis>
|
||
38; West Mediterranean to Messina
|
||
<emphasis id="A380EAA6FFB65B2C50C9FBD80ED95B6F" bold="true" box="[675,691,1113,1132]" pageId="6" pageNumber="70">…</emphasis>
|
||
<taxonomicName id="56F44D37FFB65B2C50D3FBD80CB35B8C" ID-CoL="3H4RC" baseAuthorityName="Risso" baseAuthorityYear="1810" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="6" pageNumber="70" phylum="Chordata" rank="species" species="willdenowi">
|
||
<emphasis id="A380EAA6FFB65B2C50D3FBD80CB35B8C" italics="true" pageId="6" pageNumber="70">Gouania willdenowi</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="914B36B4FFB65B2C52CEFB1F0D005BD7" blockId="6.[124,772,142,1237]" pageId="6" pageNumber="70">
|
||
<keyLead id="2A059581FFB65B2C52CEFB1F0D005BD7" pageId="6" pageNumber="70">
|
||
4b. Vertebrae 35
|
||
<emphasis id="A380EAA6FFB65B2C5320FB1E0D3C5BB1" bold="true" box="[330,342,1183,1202]" pageId="6" pageNumber="70">–</emphasis>
|
||
36; southern Ionian Sea and
|
||
<collectingRegion id="5330F856FFB65B2C5012FB1F0ED75BB1" box="[632,701,1182,1202]" country="Greece" name="Voreio Aigaio" pageId="6" pageNumber="70">Aegean</collectingRegion>
|
||
Sea
|
||
<emphasis id="A380EAA6FFB65B2C5099FB1E0F695BB1" bold="true" box="[755,771,1183,1202]" pageId="6" pageNumber="70">…</emphasis>
|
||
<taxonomicName id="56F44D37FFB65B2C5216FB400D715BD6" authority="Wagner & Kovac & Koblmüller, 2021" authorityName="Wagner & Kovac & Koblmüller" authorityYear="2021" box="[124,283,1217,1237]" family="Gobiesocidae" genus="Gouania" kingdom="Animalia" order="Gobiesociformes" pageId="6" pageNumber="70" phylum="Chordata" rank="species" species="orientalis" status="sp. nov.">
|
||
<emphasis id="A380EAA6FFB65B2C5216FB400D715BD6" box="[124,283,1217,1237]" italics="true" pageId="6" pageNumber="70">Gouania orientalis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A380EAA6FFB65B2C534BFB410D005BD7" bold="true" box="[289,362,1216,1236]" pageId="6" pageNumber="70">
|
||
<taxonomicNameLabel id="B8B357DDFFB65B2C534BFB410D005BD7" box="[289,362,1216,1236]" pageId="6" pageNumber="70" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
</key>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |