125 lines
13 KiB
XML
125 lines
13 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1179.104037" ID-Pensoft-Pub="1313-2970-1179-243" ID-Pensoft-UUID="FD30B3684976524BBEFEED3F9C8679D1" ID-ZooBank="1C2AC35B27664077BA9B3EB4E8E8452A" ModsDocID="1313-2970-1179-243" checkinTime="1694565413728" checkinUser="pensoft" docAuthor="Jimenez-Ferbans, Larry, Maestre-Guerra, Ana, Villalba-Fuentes, Evelin, Barros-Barrios, Mayelis M. & Munoz-Montero, Jeison" docDate="2023" docId="0DC3CE5A646E541E8D9E650BE9185269" docLanguage="en" docName="ZooKeys 1179: 243-297" docOrigin="ZooKeys 1179" docPubDate="2023-09-12" docSource="http://dx.doi.org/10.3897/zookeys.1179.104037" docTitle="Passalus (Passalus) coniferus Eschscholtz 1829" docType="treatment" docVersion="1" id="FD30B3684976524BBEFEED3F9C8679D1" lastPageNumber="243" masterDocId="FD30B3684976524BBEFEED3F9C8679D1" masterDocTitle="Passalidae (Coleoptera, Scarabaeoidea) from the Caribbean coast of Colombia: synopsis, key, and new species description" masterLastPageNumber="297" masterPageNumber="243" pageNumber="243" updateTime="1694565413728" updateUser="pensoft">
|
|
<mods:mods id="A5C7F9A0E1C685AEF3F86633ADC9DA78" xmlns:mods="http://www.loc.gov/mods/v3">
|
|
<mods:titleInfo id="AC1553AEED5DDB8B84537A286E65F7D8">
|
|
<mods:title id="DA1D62AFBB249345366D3740A9FD0386">Passalidae (Coleoptera, Scarabaeoidea) from the Caribbean coast of Colombia: synopsis, key, and new species description</mods:title>
|
|
</mods:titleInfo>
|
|
<mods:name id="28DEC04166D3EA9EAC9EA382ADF73F5A" type="personal">
|
|
<mods:role id="ED027C5F274175F28BE24A4390B4ED24">
|
|
<mods:roleTerm id="8FDE1FD7A24364627D071895EB53D0EB">Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart id="44C2D1A8F56B4F713AFFE55C81AD3937">Jimenez-Ferbans, Larry</mods:namePart>
|
|
<mods:nameIdentifier id="F49BEAC7FF36FBDA4DFCF58352C84D52" type="ORCID">https://orcid.org/0000-0002-5710-2265</mods:nameIdentifier>
|
|
<mods:affiliation id="1F167FE69B70A797AC1F8D3CE711BB8A">Facultad de Ciencias Basicas. Universidad del Magdalena, carrera 32 # 22 - 08, Santa Marta, Zip code 470004, Colombia</mods:affiliation>
|
|
<mods:nameIdentifier id="5DC8ED4B54DD3F63F9DE2755351D43F0" type="email">larryjimenezferbans@gmail.com</mods:nameIdentifier>
|
|
</mods:name>
|
|
<mods:name id="8B0A3CD685621C398D7E3BEB20C90627" type="personal">
|
|
<mods:role id="4B9152A8FD50B78DAA3398146BFAEB25">
|
|
<mods:roleTerm id="DEED292911841B3FA0BF285EEB44FE15">Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart id="A87909555F76B099A4B9C015F44F09A4">Maestre-Guerra, Ana</mods:namePart>
|
|
<mods:nameIdentifier id="D7619889AD7577CA2EC0D7CD46B35979" type="ORCID">https://orcid.org/0000-0002-2046-8036</mods:nameIdentifier>
|
|
<mods:affiliation id="F1D1C9340CA79FA0B545759623A611A2">Facultad de Ciencias Basicas. Universidad del Magdalena, carrera 32 # 22 - 08, Santa Marta, Zip code 470004, Colombia</mods:affiliation>
|
|
</mods:name>
|
|
<mods:name id="CC733833C988EB925EC371CFE3B7281D" type="personal">
|
|
<mods:role id="A4BF44BBC7005450117513981D6DAEF0">
|
|
<mods:roleTerm id="A9D33D059751D558EDF290375A87C755">Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart id="3CD74E34056B685ED33E51046D3448F6">Villalba-Fuentes, Evelin</mods:namePart>
|
|
<mods:nameIdentifier id="9714821DC86BC581042F03B5B184E24E" type="ORCID">https://orcid.org/0000-0002-3332-5384</mods:nameIdentifier>
|
|
<mods:affiliation id="514EF9F2CA5B9D906743610AEADD3991">Facultad de Ciencias Basicas. Universidad del Magdalena, carrera 32 # 22 - 08, Santa Marta, Zip code 470004, Colombia</mods:affiliation>
|
|
</mods:name>
|
|
<mods:name id="7D08C0F2C26BA9DD23394640F8465FC1" type="personal">
|
|
<mods:role id="9DBC0698E23E1B708C0C35CE4DC2B9D5">
|
|
<mods:roleTerm id="D5314BFBFD0CA9846D2A9440FC1C5E05">Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart id="03771453DEA0BDA4BF04FE5013D0C5B6">Barros-Barrios, Mayelis M.</mods:namePart>
|
|
<mods:nameIdentifier id="703CF4CE7F8B01E0BF45E5785AFE9E8A" type="ORCID">https://orcid.org/0000-0002-2634-5408</mods:nameIdentifier>
|
|
<mods:affiliation id="D078742C6302E3E3345938A3A1907793">Facultad de Ciencias Exactas y Naturales. Universidad de Caldas, Calle 65 # 26 - 10, Manizales, Zip code 170004, Colombia</mods:affiliation>
|
|
</mods:name>
|
|
<mods:name id="209F06D1B23E75DD5E173EC179F601B5" type="personal">
|
|
<mods:role id="D8E4984A9868749AB6A3F15BC02D9CA5">
|
|
<mods:roleTerm id="763A7F60F95520322FF0157623F7F422">Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart id="30166E48DC9C03270B68094BDE67834F">Munoz-Montero, Jeison</mods:namePart>
|
|
<mods:nameIdentifier id="F81EEC85CDD9B9DCC2ACBDB9D634EE48" type="ORCID">https://orcid.org/0009-0003-2563-9388</mods:nameIdentifier>
|
|
<mods:affiliation id="8E512C87021C4AC5B96F9A7CB97C445B">Facultad de Ciencias Basicas. Universidad del Magdalena, carrera 32 # 22 - 08, Santa Marta, Zip code 470004, Colombia</mods:affiliation>
|
|
</mods:name>
|
|
<mods:typeOfResource id="89569B2849D6A5B299D41C6BFEDA6784">text</mods:typeOfResource>
|
|
<mods:relatedItem id="AFF25B83B5C86F134DEAD5725305080E" type="host">
|
|
<mods:titleInfo id="E5D5FDCE5CC15221821015F0B350DD81">
|
|
<mods:title id="3B472433DF39B3417D73EFB058CB082F">ZooKeys</mods:title>
|
|
</mods:titleInfo>
|
|
<mods:part id="646774D31D0BC20FE6EA72B6A2DCD113">
|
|
<mods:date id="F9EF2C6610D898E6BDDBFC7F63874FE1">2023</mods:date>
|
|
<mods:detail id="6AC8E12CEBEE4DDB67F132804AC6D95F" type="pubDate">
|
|
<mods:number id="72D58A235C9D977235D2C8FBB1A07A72">2023-09-12</mods:number>
|
|
</mods:detail>
|
|
<mods:detail id="40C0AAAD606AED8A2A1F8774B0E92E80" type="volume">
|
|
<mods:number id="31B7971A2288DD4A1F5C146342D3F203">1179</mods:number>
|
|
</mods:detail>
|
|
<mods:extent id="2F5922D32E799D0DA2A2F1EC29BC4C7C" unit="page">
|
|
<mods:start id="494AAD05ADD0CB87FA22BDB179DC6F16">243</mods:start>
|
|
<mods:end id="192B6D117F72ABB7D7F03611EECC8FAA">297</mods:end>
|
|
</mods:extent>
|
|
</mods:part>
|
|
</mods:relatedItem>
|
|
<mods:location id="DCED141FB5C11D562A8EBF6CA902BA01">
|
|
<mods:url id="F8B955A170A507CE563224318AADEB73">http://dx.doi.org/10.3897/zookeys.1179.104037</mods:url>
|
|
</mods:location>
|
|
<mods:classification id="BEEFAC951751EBFEE2C412CD2EA8EF49">journal article</mods:classification>
|
|
<mods:identifier id="F1E3BFDCAD2CD74254046CDE5F5C1C80" type="DOI">http://dx.doi.org/10.3897/zookeys.1179.104037</mods:identifier>
|
|
<mods:identifier id="706E1BFC5C66008D7268D31806C8C7D7" type="Pensoft-Pub">1313-2970-1179-243</mods:identifier>
|
|
<mods:identifier id="E36C951057EAEE8DC487F9D5CC15F741" type="ZooBank">1C2AC35B27664077BA9B3EB4E8E8452A</mods:identifier>
|
|
<mods:identifier id="0FC287CFE5D924235A3D3A44F661A61D" type="Pensoft-UUID">FD30B3684976524BBEFEED3F9C8679D1</mods:identifier>
|
|
</mods:mods>
|
|
<treatment id="0DC3CE5A646E541E8D9E650BE9185269" LSID="urn:lsid:plazi:treatment:0DC3CE5A646E541E8D9E650BE9185269" httpUri="http://treatment.plazi.org/id/0DC3CE5A646E541E8D9E650BE9185269" lastPageNumber="243" pageId="0" pageNumber="243">
|
|
<subSubSection id="6130906B3E5D17057390A0CE9AE266A6" pageId="0" pageNumber="243" type="nomenclature">
|
|
<paragraph id="F1A357A2D7C46B0035110CB7E926A609" pageId="0" pageNumber="243">
|
|
1.
|
|
<taxonomicName id="EDFDEF08A1D800C94CD181C541872910" ID-CoL="6TSBG" ID-ENA="2652925" LSID="0DC3CE5A-646E-541E-8D9E-650BE9185269" authority="Eschscholtz, 1829" authorityName="Eschscholtz" authorityYear="1829" class="Insecta" family="Passalidae" genus="Passalus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Passalus (Passalus) coniferus" order="Coleoptera" pageId="0" pageNumber="243" phylum="Arthropoda" rank="species" species="coniferus" subGenus="Passalus">Passalus (Passalus) coniferus Eschscholtz, 1829</taxonomicName>
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="4C52F91EA8997DE4B0565E0218ABE61A" pageId="0" pageNumber="243" type="description">
|
|
<paragraph id="F89DF1EE630383715056F83162D338E9" pageId="0" pageNumber="243">
|
|
<figureCitation id="CED06914E6534CBEE012A942D5994C84" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Passalus (Passalus) coniferus Eschscholtz, 1829 A head and pronotum in dorsal view B habitus dorsal C habitus ventral D habitus lateral. Scale bars: 2.0 mm (A); 3.0 mm (B, C, D)." figureDoi="10.3897/zookeys.1179.104037.figure2" httpUri="https://binary.pensoft.net/fig/904566" pageId="0" pageNumber="243">Fig. 2</figureCitation>
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="BD06BEA017D7CCFC2EABE839EDA6457C" pageId="0" pageNumber="243" type="diagnosis">
|
|
<paragraph id="F0AB682AF8BA5267DE17948A5563B5E2" pageId="0" pageNumber="243">Diagnosis.</paragraph>
|
|
<paragraph id="27B309CD151A8668A3F607D659FF539B" pageId="0" pageNumber="243">39.6-46.5 mm total length. Body robust. Anterior border of the frons with two prominent secondary mediofrontal tubercles. Mediofrontal tubercles large, located on base of each laterofrontal tubercle. Central tubercle with apex free. Lateroposterior tubercles distinct. Eyes large. Antennal club tri-lamellate, with lamellae long. Lacinia with apex bidentate. Mediobasal area of mentum protruding and glabrous, sometimes with scarce setae on the posterior border. Marginal pronotal groove occupying 2/3 of the pronotum anterior border. Prosternellum rhomboidal, truncate. Mesosternum pubescent, with inconspicuous and elongated scars. Metasternum pubescent anterolaterally and in lateral groove; disc smooth, delimited posteriorly to middle by punctations. Humeri pubescent, epipleura pubescent in basal 2/3. Last abdominal sternite with complete marginal groove. Anterior ventral border of the profemur with well-developed groove. Meso- and metatibia with one or two small spines.</paragraph>
|
|
<caption id="B1B9D4094BC413657FBB4598363ED9C7" doi="10.3897/zookeys.1179.104037.figure2" httpUri="https://binary.pensoft.net/fig/904566" pageId="0" pageNumber="243" start="Figure 2" startId="F2">
|
|
<paragraph id="10F54BA1A0B2F4F4034516FF0EF45235" pageId="0" pageNumber="243">
|
|
<emphasis id="369A4ED0148C84E51C127C52B15572B0" bold="true" pageId="0" pageNumber="243">Figure 2.</emphasis>
|
|
<taxonomicName id="49A9F81A8CF88028CF96E79556557538" authorityName="Eschscholtz" authorityYear="1829" class="Insecta" family="Passalidae" genus="Passalus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Passalus (Passalus) coniferus" order="Coleoptera" pageId="0" pageNumber="243" phylum="Arthropoda" rank="species" species="coniferus" subGenus="Passalus">Passalus (Passalus) coniferus</taxonomicName>
|
|
Eschscholtz, 1829
|
|
<emphasis id="3F7C0068049EE7EC86288DC0CA1D9ED8" bold="true" pageId="0" pageNumber="243">A</emphasis>
|
|
head and pronotum in dorsal view
|
|
<emphasis id="A751F66632155DA39FB0817093A76A14" bold="true" pageId="0" pageNumber="243">B</emphasis>
|
|
habitus dorsal
|
|
<emphasis id="7F23AB93A72EA1AF813723EFC8947051" bold="true" pageId="0" pageNumber="243">C</emphasis>
|
|
habitus ventral
|
|
<emphasis id="9A8B672C4E32FB788F4776B4F68B400D" bold="true" pageId="0" pageNumber="243">D</emphasis>
|
|
habitus lateral. Scale bars: 2.0 mm (
|
|
<emphasis id="E52B18A56974415D886497AF96DD3B37" bold="true" pageId="0" pageNumber="243">A</emphasis>
|
|
); 3.0 mm (
|
|
<emphasis id="742E3F6EBF91F7C28322938060631F46" bold="true" pageId="0" pageNumber="243">B, C, D</emphasis>
|
|
).
|
|
</paragraph>
|
|
</caption>
|
|
</subSubSection>
|
|
<subSubSection id="0D9A246257404565BA7AB6EC090F1054" pageId="0" pageNumber="243" type="comments">
|
|
<paragraph id="DF096095405C9393B06D1569F47E9505" pageId="0" pageNumber="243">Comments.</paragraph>
|
|
<paragraph id="0476A38D40F731144A6D14EDF9F03515" pageId="0" pageNumber="243">
|
|
A South American species, with distribution in the Antilles (
|
|
<bibRefCitation id="0F5129207E8C96D0CE3DF1638174F8CF" author="Reyes-Castillo, P" journalOrPublisher="Bulletin du Museum National d'Histoire Naturelle" pageId="0" pageNumber="243" pagination="1541 - 1587" refId="B30" refString="Reyes-Castillo, P, 1973. Passalidae de la Guyana Francesa (Coleoptera, Lamellicornia). Bulletin du Museum National d'Histoire Naturelle 197: 1541 - 1587" title="Passalidae de la Guyana Francesa (Coleoptera, Lamellicornia)." volume="197" year="1973">Reyes-Castillo 1973</bibRefCitation>
|
|
). It is a species of highly variable size,
|
|
<bibRefCitation id="F4F8558164BAFD94111A5459B531DBDF" author="Reyes-Castillo, P" editor="Onore, G" journalOrPublisher="Monografias tercer milenio vol. III, Sociedad Entomologica Aragones (SEA), Zaragoza" pageId="0" pageNumber="243" pagination="35 - 50" refId="B31" refString="Reyes-Castillo, P, Amat-Garcia, GD, 2003. Passalidae (Coleoptera) de Colombia. In: Onore, G, Reyes-Castillo, P, Zunino, M, Eds., Escarabajos de Latinoamerica: estado del conocimiento. Monografias tercer milenio vol. III, Sociedad Entomologica Aragones (SEA), Zaragoza: 35 - 50" title="Passalidae (Coleoptera) de Colombia." volumeTitle="Escarabajos de Latinoamerica: estado del conocimiento." year="2003">
|
|
Reyes-Castillo and
|
|
<normalizedToken id="35CB0F688278CEB385AD2AA30E74C198" originalValue="Amat-García">Amat-Garcia</normalizedToken>
|
|
(2003)
|
|
</bibRefCitation>
|
|
notes that the specimens from the Sierra Nevada de Santa Marta are larger than those from the rest of the country, with the apex of the horn free and very long.
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |