229 lines
44 KiB
XML
229 lines
44 KiB
XML
<document id="E3738F097948AC3869E953C6F8B7F4BF" ID-CLB-Dataset="6637" ID-DOI="10.5281/zenodo.4648646" ID-GBIF-Dataset="9797e168-d1b1-4c0a-8786-638b6f992159" ID-ISSN="1536-9307" ID-Zenodo-Dep="4648646" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1617136052780" checkinUser="carolina" docAuthor="Kovařík, František & Šťáhlavský, František" docDate="2020" docId="062F616DB930FF9727EEBA00FBA45898" docLanguage="en" docName="Euscorpius.315.1-37.pdf" docOrigin="Euscorpius 315" docStyle="DocumentStyle:5E96F6379CE20B0E8E4F9E7DF8E1C0DF.2:Euscorpius.2020-.journal_article" docStyleId="5E96F6379CE20B0E8E4F9E7DF8E1C0DF" docStyleName="Euscorpius.2020-.journal_article" docStyleVersion="2" docTitle="Euscorpius Thorell 1876" docType="treatment" docVersion="9" lastPageNumber="2" masterDocId="FA161915B932FF94240FBC31FF9A5E4A" masterDocTitle="Five new species of Euscorpius Thorell, 1876 (Scorpiones: Euscorpiidae) from Albania, Greece, North Macedonia, and Serbia" masterLastPageNumber="37" masterPageNumber="1" pageNumber="1" updateTime="1698935484995" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="FA75356DB0AB9F5FC0DEE20E5139A827">Five new species of Euscorpius Thorell, 1876 (Scorpiones: Euscorpiidae) from Albania, Greece, North Macedonia, and Serbia</mods:title>
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<mods:title id="5F3C664AE244A39EB0CCF8FF70806C67">Euscorpius</mods:title>
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<mods:date id="6BF4768EFABD495217ED3491B6EAB604">2020</mods:date>
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<emphasis id="BCF20C69B930FF9627EEBA00FBF05801" bold="true" box="[993,1130,1585,1611]" italics="true" pageId="2" pageNumber="1">Euscorpius</emphasis>
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<bibRefCitation id="EA17AD8AB930FF96207EBA00FA8B5801" author="THORELL" box="[1137,1297,1585,1611]" pageId="2" pageNumber="1" pagination="1 - 15" refId="ref13065" refString="THORELL, T. 1876. On the classification of scorpions. Annals and Magazine of Natural History, 4 (17): 1 - 15." type="journal article" year="1876">Thorell, 1876</bibRefCitation>
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captionStart-9="Figures 64–71" captionStartId-0="3.[115,192,1140,1162]" captionStartId-1="6.[115,192,1990,2012]" captionStartId-10="16.[116,193,1960,1982]" captionStartId-11="18.[113,190,1988,2010]" captionStartId-12="19.[116,193,1958,1980]" captionStartId-13="20.[113,190,1958,1980]" captionStartId-14="21.[116,193,1990,2012]" captionStartId-15="22.[116,193,1956,1978]" captionStartId-16="24.[113,190,1977,1999]" captionStartId-17="25.[113,190,1966,1988]" captionStartId-18="26.[115,192,1958,1980]" captionStartId-19="27.[118,195,1987,2009]" captionStartId-2="7.[115,192,1958,1980]" captionStartId-20="28.[115,192,1971,1993]" captionStartId-21="29.[116,184,1234,1256]" captionStartId-22="30.[113,190,1988,2010]" captionStartId-23="31.[113,190,1449,1471]" captionStartId-24="32.[113,190,1958,1980]" captionStartId-25="33.[115,192,1977,1999]" captionStartId-26="34.[113,190,1962,1984]" captionStartId-27="35.[116,184,1289,1311]" captionStartId-28="36.[818,895,925,947]" 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captionTargetId-16="figure-57@24.[113,1474,242,1946]" captionTargetId-17="figure-65@25.[113,1474,234,1906]" captionTargetId-18="figure-75@26.[113,1474,234,1934]" captionTargetId-19="figure-47@27.[113,1474,234,1977]" captionTargetId-2="figure-78@7.[113,1474,234,1939]" captionTargetId-20="figure-63@28.[113,1474,229,1964]" captionTargetId-21="figure-414@29.[113,1474,234,1226]" captionTargetId-22="figure-57@30.[113,1474,234,1975]" captionTargetId-23="figure-284@31.[113,1474,234,1424]" captionTargetId-24="figure-78@32.[113,1474,234,1947]" captionTargetId-25="figure-51@33.[113,1474,234,1960]" captionTargetId-26="figure-64@34.[113,1474,234,1955]" captionTargetId-27="figure-393@35.[113,1474,234,1281]" captionTargetId-28="figure-282@36.[818,1474,232,908]" captionTargetId-3="figure-66@8.[113,1474,234,1973]" captionTargetId-4="figure-50@9.[115,1476,229,1988]" captionTargetId-5="figure-62@10.[113,1474,229,1958]" captionTargetId-6="figure-53@12.[113,1474,234,1968]" captionTargetId-7="figure-65@13.[114,1475,234,1943]" captionTargetId-8="figure-75@14.[111,1472,234,1931]" captionTargetId-9="figure-48@15.[114,1475,234,1963]" captionTargetPageId-0="3" captionTargetPageId-1="6" captionTargetPageId-10="16" captionTargetPageId-11="18" captionTargetPageId-12="19" captionTargetPageId-13="20" captionTargetPageId-14="21" captionTargetPageId-15="22" captionTargetPageId-16="24" captionTargetPageId-17="25" captionTargetPageId-18="26" captionTargetPageId-19="27" captionTargetPageId-2="7" captionTargetPageId-20="28" captionTargetPageId-21="29" captionTargetPageId-22="30" captionTargetPageId-23="31" captionTargetPageId-24="32" captionTargetPageId-25="33" captionTargetPageId-26="34" captionTargetPageId-27="35" captionTargetPageId-28="36" captionTargetPageId-3="8" captionTargetPageId-4="9" captionTargetPageId-5="10" captionTargetPageId-6="12" captionTargetPageId-7="13" captionTargetPageId-8="14" captionTargetPageId-9="15" captionText-0="Figures 1–9: Euscorpius females (1–2) and males (3–9), pedipalp chela external. Figure 1. E. concinnus (C. L. Koch, 1837), Italy, Tuscany, Camporgiano, 44.1587131°N 10.3357447°E, FKCP. Figure 2. E. gocmeni Tropea et al., 2014, Turkey, Antalya Province, Akseki District, Elmali Yolu, 36º21'29.9"N 29º28'00.1"E, FKCP. Figure 3. E. tergestinus (C. L. Koch, 1837), Croatia, Posedarje, 20 km E of Zadar, FKCP. Figure 4. E. janstai sp. n., paratype. Figure 5. E. aquilejensis (C. L. Koch, 1837), Italy, Abruzzo, Silvi Marina, FKCP. Figure 6. E. yagmuri KovařÍk et al., 2014, holotype, Greece, Epirus, Preveza District, Mt. Zalongo, Cassope (Kassope, Kassopi), 39.145278ºN 20.675556ºE, FKCP. Figure 7. E. lycius Yağmur et al., 2013, Turkey, Antalya Province, 36º24'N 29º41'E, 976 m a. s. l., FKCP. Figure 8. E. solegladi Fet et al., 2014, paratype, Bulgaria, “Sandanski Pirin” (Blagoevgrad Province, Pirin Mts., Sandanski), NMPC. Figure 9. E. hadzii Di Caporiacco, 1950, Albania, Prokletije Mts., road Theti–Boge, FKCP." captionText-1="Figures 10–13: E. bonacinai sp. n. Figures 10–11. Male holotype in dorsal (10) and ventral (11) views. Figures 13–14. Female paratype in dorsal (12) and ventral (13) views. Scale bar: 10 mm." captionText-10="Figures 72–79. E. janstai sp. n., female paratype, pedipalp segments. Chela dorsal (72), external (73) and ventral (74) views. Patella dorsal (75), external (76) and ventral (77) views. Trochanter and femur dorsal (78) and ventral (79) views.Trichobothrial pattern is indicated by white circles." captionText-11="Figures 80–83: E. kabateki sp. n. Figures 80–81. Male holotype in dorsal (80) and ventral (81) views. Figures 82–83. Female paratopotype in dorsal (82) and ventral (83) views. Scale bar: 10 mm." captionText-12="Figures 84–90: E. kabateki sp. n. Figures 84, 86–87, 89–90. Male holotype, carapace and tergites I–IV (84), coxosternal area and sternites III–V (86), left legs III–IV, retrolateral aspect (89–90 respectively), and chelicerae and eyes with part of carapace (87). Figures 85, 88. Female paratopotype, carapace and tergites I–III (85), coxosternal area and sternites III–IV (88)." captionText-13="Figures 91–98: E. kabateki sp. n. Figures 91–94. Female paratopotype, telson lateral (91), and metasoma and telson lateral (92), dorsal (93), and ventral (94) views. Figures 95. Male paratype, telson lateral. Figures 96–98. Male holotype, metasoma and telson lateral (96), dorsal (97), and ventral (98) views. Scale bars: 5 mm (92–94, 96–98)." captionText-14="Figures 99–106. E. kabateki sp. n., male holotype, pedipalp segments. Chela dorsal (99), external (100) and ventral (101) views. Patella dorsal (102), external (103) and ventral (104) views. Trochanter and femur dorsal (105) and ventral (106) views." captionText-15="Figures 107–114. E. kabateki sp. n., female paratopotype, pedipalp segments. Chela dorsal (107), external (108) and ventral (109) views. Patella dorsal (110), external (111) and ventral (112) views. Trochanter and femur dorsal (113) and ventral (114) views. Trichobothrial pattern is indicated by white circles." captionText-16="Figures 115–118: E. sadileki sp. n. Figures 115–116. Male holotype in dorsal (115) and ventral (116) views. Figures 117–118. Female paratopotype in dorsal (117) and ventral (118) views. Scale bar: 10 mm." captionText-17="Figures 119–125: E. sadileki sp. n. Figures 119, 121, 123–125. Male holotype, carapace and tergites I–IV (119), coxosternal area and sternites (121), left legs II–IV, retrolateral aspect (123–125 respectively). Figures 120, 122. Female paratopotype, carapace and tergites I–III (120), coxosternal area and sternites III–IV (122)." captionText-18="Figures 126–133: E. sadileki sp. n. Figures 126–129. Female paratopotype, telson lateral (126), and metasoma and telson lateral (127), dorsal (128), and ventral (129) views. Figures 130–133. Male holotype, telson lateral (130), and metasoma and telson lateral (131), dorsal (132), and ventral (133) views. Scale bars: 5 mm (127–129, 131–133)." captionText-19="Figures 134–140. E. sadileki sp. n., male holotype, pedipalp segments. Chela dorsal (134), external (135) and ventral (136) views. Patella dorsal (137), external (138) and ventral (139) views. Trochanter and femur dorsal (140) view." captionText-2="Figures 14–23: E. bonacinai sp. n. Figures 14, 16, 18–21, 23. Male holotype, carapace and tergites I–III (14), coxosternal area and sternites III–V (16), left legs I–IV, retrolateral aspect (18–21 respectively), and telson lateral (23). Figures 15, 17, 22. Female paratype, carapace and tergites I–III (15), coxosternal area and sternites III–IV (17), and telson lateral (22)." captionText-20="Figures 141–148: E. sadileki sp. n., female paratopotype, pedipalp segments. Chela dorsal (141), external (142) and ventral (143) views. Patella dorsal (144), external (145) and ventral (146) views. Trochanter and femur dorsal (147) and ventral (148) views. Trichobothrial pattern is indicated by white circles." captionText-21="Figure 149. E. sadileki sp. n., female paratype in vivo habitus." captionText-22="Figures 150–153: E. scheraboni sp. n. Figures 150–151. Male holotype in dorsal (150) and ventral (151) views. Figures 152–153. Female paratype in dorsal (152) and ventral (153) views. Scale bar: 10 mm." captionText-23="Figures 154–159: E. scheraboni sp. n. Figures 154, 156, 158–159. Male holotype, carapace and tergites I–IV (154), coxosternal area and sternites III–V (156), left legs III–IV, retrolateral aspect (158–159 respectively). Figures 155, 157. Female paratype, carapace and tergites I–III (155), coxosternal area (157)." captionText-24="Figures 160–167: E. scheraboni sp. n. Figures 160–163. Female paratype, telson lateral (160), and metasoma and telson lateral (161), dorsal (162), and ventral (163) views. Figures 164, 165–167. Male holotype, telson lateral (164), and metasoma and telson lateral (165), dorsal (166), and ventral (167) views. Scale bars: 5 mm (161–163, 165–167)." captionText-25="Figures 168–175. E. scheraboni sp. n., male holotype, pedipalp segments. Chela dorsal (168), external (169) and ventral (170) views. Patella dorsal (171), external (172) and ventral (173) views. Trochanter and femur dorsal (174) and ventral (175) views." captionText-26="Figures 176–183. E. scheraboni sp. n., female paratype, pedipalp segments. Chela dorsal (176), external (177) and ventral (178) views. Patella dorsal (179), external (180) and ventral (181) views. Trochanter and femur dorsal (182) and ventral (183) views. Trichobothrial pattern is indicated by white circles." captionText-27="Figure 184. Map of distribution of five species described here." captionText-28="Figures 185–186. Male karyotypes of E. janstai sp. n. (185), and E. sadileki sp. n. (186) based on postpachytene. Scale bar: 10 μm." captionText-3="Figures 24–29: E. bonacinai sp. n. Figures 24–26. Female paratype, metasoma and telson lateral (24), dorsal (25), and ventral (26) views. Figures 27–29. Male holotype, metasoma and telson lateral (27), dorsal (28), and ventral (29) views. Scale bars: 5 mm (24–26, 27–29)." captionText-4="Figures 30–37. E. bonacinai sp. n., male holotype, pedipalp segments. Chela dorsal (30), external (31) and ventral (32) views. Patella dorsal (33), external (34) and ventral (35) views. Trochanter and femur dorsal (36) and ventral (37) views." captionText-5="Figures 38–45. E. bonacinai sp. n., female paratype, pedipalp segments. Chela dorsal (38), external (39) and ventral (40) views. Patella dorsal (41), external (42) and ventral (43) views. Trochanter and femur dorsal (44) and ventral (45) views. Trichobothrial pattern is indicated by white circles." captionText-6="Figures 46–49: E. janstai sp. n. Figures 46–47. Male holotype in dorsal (46) and ventral (47) views. Figures 48–49. Female paratype in dorsal (48) and ventral (49) views." captionText-7="Figures 50–55: E. janstai sp. n. Figures 50, 52, 54–55. Male holotype, carapace and tergites I–III (50), coxosternal area and sternites (52), left legs III–IV, retrolateral aspect (54–55 respectively). Figures 51, 52. Female paratype, carapace and tergites I–III (51), coxosternal area and sternites III–IV (52)." captionText-8="Figures 56–63: E. janstai sp. n. Figures 57–60. Female paratype, telson lateral (57), and metasoma and telson lateral (58), dorsal (59), and ventral (60) views. Figures 56, 61–63. Male holotype, telson lateral (56), and metasoma and telson lateral (61), dorsal (62), and ventral (63) views. Scale bar: 5 mm (58–63)." captionText-9="Figures 64–71. E. janstai sp. n., male holotype, pedipalp segments. Chela dorsal (64), external (65) and ventral (66) views. Patella dorsal (67), external (68) and ventral (69) views. Femur dorsal (70). Movable finger dentition (71)." figureDoi-0="http://doi.org/10.5281/zenodo.4648648" figureDoi-1="http://doi.org/10.5281/zenodo.4648650" figureDoi-10="http://doi.org/10.5281/zenodo.4648668" figureDoi-11="http://doi.org/10.5281/zenodo.4648670" figureDoi-12="http://doi.org/10.5281/zenodo.4648672" figureDoi-13="http://doi.org/10.5281/zenodo.4648674" figureDoi-14="http://doi.org/10.5281/zenodo.4648676" figureDoi-15="http://doi.org/10.5281/zenodo.4648678" figureDoi-16="http://doi.org/10.5281/zenodo.4648680" figureDoi-17="http://doi.org/10.5281/zenodo.4648682" figureDoi-18="http://doi.org/10.5281/zenodo.4648684" figureDoi-19="http://doi.org/10.5281/zenodo.4648686" figureDoi-2="http://doi.org/10.5281/zenodo.4648652" figureDoi-20="http://doi.org/10.5281/zenodo.4648688" figureDoi-21="http://doi.org/10.5281/zenodo.4648691" figureDoi-22="http://doi.org/10.5281/zenodo.4648693" figureDoi-23="http://doi.org/10.5281/zenodo.4648695" figureDoi-24="http://doi.org/10.5281/zenodo.4648697" figureDoi-25="http://doi.org/10.5281/zenodo.4648699" figureDoi-26="http://doi.org/10.5281/zenodo.4648701" figureDoi-27="http://doi.org/10.5281/zenodo.4648703" figureDoi-28="http://doi.org/10.5281/zenodo.4648705" figureDoi-3="http://doi.org/10.5281/zenodo.4648654" figureDoi-4="http://doi.org/10.5281/zenodo.4648656" figureDoi-5="http://doi.org/10.5281/zenodo.4648658" figureDoi-6="http://doi.org/10.5281/zenodo.4648660" figureDoi-7="http://doi.org/10.5281/zenodo.4648662" figureDoi-8="http://doi.org/10.5281/zenodo.4648664" figureDoi-9="http://doi.org/10.5281/zenodo.4648666" httpUri-0="https://zenodo.org/record/4648648/files/figure.png" httpUri-1="https://zenodo.org/record/4648650/files/figure.png" httpUri-10="https://zenodo.org/record/4648668/files/figure.png" httpUri-11="https://zenodo.org/record/4648670/files/figure.png" httpUri-12="https://zenodo.org/record/4648672/files/figure.png" httpUri-13="https://zenodo.org/record/4648674/files/figure.png" httpUri-14="https://zenodo.org/record/4648676/files/figure.png" httpUri-15="https://zenodo.org/record/4648678/files/figure.png" httpUri-16="https://zenodo.org/record/4648680/files/figure.png" httpUri-17="https://zenodo.org/record/4648682/files/figure.png" httpUri-18="https://zenodo.org/record/4648684/files/figure.png" httpUri-19="https://zenodo.org/record/4648686/files/figure.png" httpUri-2="https://zenodo.org/record/4648652/files/figure.png" httpUri-20="https://zenodo.org/record/4648688/files/figure.png" httpUri-21="https://zenodo.org/record/4648691/files/figure.png" httpUri-22="https://zenodo.org/record/4648693/files/figure.png" httpUri-23="https://zenodo.org/record/4648695/files/figure.png" httpUri-24="https://zenodo.org/record/4648697/files/figure.png" httpUri-25="https://zenodo.org/record/4648699/files/figure.png" httpUri-26="https://zenodo.org/record/4648701/files/figure.png" httpUri-27="https://zenodo.org/record/4648703/files/figure.png" httpUri-28="https://zenodo.org/record/4648705/files/figure.png" httpUri-3="https://zenodo.org/record/4648654/files/figure.png" httpUri-4="https://zenodo.org/record/4648656/files/figure.png" httpUri-5="https://zenodo.org/record/4648658/files/figure.png" httpUri-6="https://zenodo.org/record/4648660/files/figure.png" httpUri-7="https://zenodo.org/record/4648662/files/figure.png" httpUri-8="https://zenodo.org/record/4648664/files/figure.png" httpUri-9="https://zenodo.org/record/4648666/files/figure.png" pageId="2" pageNumber="1">Figures 1–186</figureCitation>
|
||
,
|
||
<tableCitation id="C304E5C0B930FF96209DBA62FA9D5821" box="[1170,1287,1619,1643]" captionStart-0="Table 1" captionStart-1="Table 2" captionStart-2="Table 3" captionStart-3="Table 4" captionStartId-0="5.[113,169,994,1015]" captionStartId-1="11.[113,169,1008,1029]" captionStartId-2="23.[114,170,983,1004]" captionStartId-3="36.[115,171,1931,1952]" captionTargetId-0="graphics-459@5.[114,1473,242,968]" captionTargetId-1="graphics-496@11.[120,1474,234,994]" captionTargetId-2="graphics-512@23.[114,1475,234,972]" captionTargetId-3="graphics-283@36.[113,776,232,1915]" captionTargetPageId-0="5" captionTargetPageId-1="11" captionTargetPageId-2="23" captionTargetPageId-3="36" captionText-0="Table 1. Comparative measurements of adults of Euscorpius bonacinai sp. n. Abbreviations: length (L), width (W, in carapace it corresponds to posterior width), depth (D)." captionText-1="Table 2. Comparative measurements of adults of Euscorpius janstai sp. n. and E. kabateki sp. n. Abbreviations: length (L), width (W, in carapace it corresponds to posterior width), depth (D)." captionText-2="Table 3. Comparative measurements of adults of Euscorpius sadileki sp. n. and E. scheraboni sp. n. Abbreviations: length (L), width (W, in carapace it corresponds to posterior width), depth (D)." captionText-3="Table 4. Measurements of the relative diploid set length (%DSL) and arm ratio (AR) of chromosomes (including standard deviation ±SD) of Euscorpius janstai sp. n. and E. sadileki sp. n. Abbreviations: M – metacentric,SM – submetacentric, ST – subtelocentric,T – telocentric." httpUri-0="http://table.plazi.org/id/DAF980F3B937FF91247EBFD3FE095A5F" httpUri-1="http://table.plazi.org/id/DAF980F3B939FF9F247EBFC1FDE15A69" httpUri-2="http://table.plazi.org/id/DAF980F3B925FF83247DBFE6FDE15A40" httpUri-3="http://table.plazi.org/id/DAF980F3B916FFB0247CBBBAFC9E59BD" pageId="2" pageNumber="1" tableUuid-0="DAF980F3B937FF91247EBFD3FE095A5F" tableUuid-1="DAF980F3B939FF9F247EBFC1FDE15A69" tableUuid-2="DAF980F3B925FF83247DBFE6FDE15A40" tableUuid-3="DAF980F3B916FFB0247CBBBAFC9E59BD">Tables 1–4</tableCitation>
|
||
)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C69C83F0B930FF96273FBAA2FAA85881" pageId="2" pageNumber="1" type="reference_group">
|
||
<paragraph id="8E39D07BB930FF96273FBAA2FAA85881" blockId="2.[816,1474,1683,1739]" pageId="2" pageNumber="1">
|
||
<treatmentCitationGroup id="AE96F755B930FF96273FBAA2FAA85881" pageId="2" pageNumber="1">
|
||
<treatmentCitation id="0F27F66AB930FF96273FBAA2FBF558E1" author="THORELL" box="[816,1135,1683,1707]" page="15" pageId="2" pageNumber="1" year="1876">
|
||
<taxonomicName id="4986ABF8B930FF96273FBAA2FBF558E1" ID-CoL="4H4X" authority="Thorell, 1876: 15" authorityName="Thorell" authorityPageNumber="15" authorityYear="1876" box="[816,1135,1683,1707]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="1" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="BCF20C69B930FF96273FBAA2FC3258E0" box="[816,936,1683,1706]" italics="true" pageId="2" pageNumber="1">Euscorpius</emphasis>
|
||
<bibRefCitation id="EA17AD8AB930FF9627BFBAA2FBF558E1" author="THORELL" box="[944,1135,1683,1707]" pageId="2" pageNumber="1" pagination="1 - 15" refId="ref13065" refString="THORELL, T. 1876. On the classification of scorpions. Annals and Magazine of Natural History, 4 (17): 1 - 15." type="journal article" year="1876">Thorell, 1876: 15</bibRefCitation>
|
||
</taxonomicName>
|
||
</treatmentCitation>
|
||
; Fet & Sissom, 2000: 357–377 (in part, complete reference list until 1998).
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C69C83F0B930FF96273FBAC2FAD05941" box="[816,1354,1779,1803]" pageId="2" pageNumber="1" type="materials_examined">
|
||
<paragraph id="8E39D07BB930FF96273FBAC2FAD05941" blockId="2.[816,1354,1779,1803]" box="[816,1354,1779,1803]" pageId="2" pageNumber="1">
|
||
<materialsCitation id="3EEEDA26B930FF96273FBAC2FAD05941" ID-GBIF-Occurrence="3114118309" box="[816,1354,1779,1803]" collectionCode="SPECIES" pageId="2" pageNumber="1" specimenCount="1" typeStatus="holotype">
|
||
<typeStatus id="513D6ED9B930FF96273FBAC2FCFC5940" box="[816,870,1779,1802]" pageId="2" pageNumber="1">TYPE</typeStatus>
|
||
<collectionCode id="E89748BEB930FF962762BAC2FC245941" box="[877,958,1779,1803]" pageId="2" pageNumber="1">SPECIES</collectionCode>
|
||
.
|
||
<taxonomicName id="4986ABF8B930FF9627C8BAC2FAD05941" authority="Linnaeus, 1767" authorityName="Linnaeus" authorityYear="1767" box="[967,1354,1779,1803]" class="Arachnida" family="Scorpionidae" genus="Scorpio" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="1" phylum="Arthropoda" rank="species" species="carpathicus">
|
||
<emphasis id="BCF20C69B930FF9627C8BAC2FB055941" box="[967,1183,1779,1803]" italics="true" pageId="2" pageNumber="1">Scorpio carpathicus</emphasis>
|
||
Linnaeus, 1767
|
||
</taxonomicName>
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C69C83F0B930FF97273FBB02FBA45898" lastPageId="3" lastPageNumber="2" pageId="2" pageNumber="1" type="diagnosis">
|
||
<paragraph id="8E39D07BB930FF97273FBB02FE6E5878" blockId="2.[816,1475,1843,2027]" lastBlockId="3.[113,772,1402,1586]" lastPageId="3" lastPageNumber="2" pageId="2" pageNumber="1">
|
||
DIAGNOSIS. Total length
|
||
<quantity id="497E7D9EB930FF96205EBB02FB4B5900" box="[1105,1233,1843,1867]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" metricValueMax="5.0" metricValueMin="2.0" pageId="2" pageNumber="1" unit="mm" value="35.0" valueMax="50.0" valueMin="20.0">20–50 mm</quantity>
|
||
. Movable fingers of pedipalp chela with MD aligned in a single straight row in entire length of chelal fingers; ID, IAD (
|
||
<quantity id="497E7D9EB930FF9620F1BB42FAB159C1" box="[1278,1323,1907,1931]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="2" pageNumber="1" unit="in" value="5.0">5 in</quantity>
|
||
number) and OD present. OD are displaced to outer aspect of fingers. Pedipalp chela flat in appearance. Trichobothrial pattern
|
||
<typeStatus id="513D6ED9B930FF96219BBB82FA585981" box="[1428,1474,1971,1995]" pageId="2" pageNumber="1">type</typeStatus>
|
||
C. Two subdistal denticles present on cheliceral movable finger dorsal edge. Ventral edge of cheliceral movable finger smooth; serrula absent. Two pedal spurs present on legs. Tarsal spurs on legs absent. Sternum pentagonal in shape. Hemispermatophore lamelliform in shape with broad trunks. Telson without subaculear tubercle. Metasoma I–IV with single median carina usually absent.
|
||
</paragraph>
|
||
<caption id="DAF980F3B931FF97247CB845FDCF5B1C" ID-DOI="http://doi.org/10.5281/zenodo.4648648" ID-Zenodo-Dep="4648648" httpUri="https://zenodo.org/record/4648648/files/figure.png" pageId="3" pageNumber="2" startId="3.[115,192,1140,1162]" targetBox="[122,1453,277,1110]" targetPageId="3">
|
||
<paragraph id="8E39D07BB931FF97247CB845FDCF5B1C" blockId="3.[115,1474,1140,1367]" pageId="3" pageNumber="2">
|
||
<emphasis id="BCF20C69B931FF97247CB845FF705AC3" bold="true" box="[115,234,1140,1162]" pageId="3" pageNumber="2">Figures 1–9</emphasis>
|
||
:
|
||
<taxonomicName id="4986ABF8B931FF9724F8B845FEF95AC3" authorityName="Thorell" authorityYear="1876" box="[247,355,1140,1161]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="BCF20C69B931FF9724F8B845FEF95AC3" box="[247,355,1140,1161]" italics="true" pageId="3" pageNumber="2">Euscorpius</emphasis>
|
||
</taxonomicName>
|
||
females (1–2) and males (3–9), pedipalp chela external.
|
||
<emphasis id="BCF20C69B931FF972785B845FC7B5AC3" bold="true" box="[906,993,1140,1162]" pageId="3" pageNumber="2">Figure 1</emphasis>
|
||
.
|
||
<taxonomicName id="4986ABF8B931FF9727E2B844FAB15AC0" authority="(C. L. Koch, 1837)" baseAuthorityName="C. L. Koch" baseAuthorityYear="1837" box="[1005,1323,1140,1162]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="concinnus">
|
||
<emphasis id="BCF20C69B931FF9727E2B844FBF35AC3" box="[1005,1129,1140,1162]" italics="true" pageId="3" pageNumber="2">E. concinnus</emphasis>
|
||
(C. L. Koch, 1837)
|
||
</taxonomicName>
|
||
, Italy, Tuscany, Camporgiano, 44.1587131°N 10.3357447°E, FKCP.
|
||
<emphasis id="BCF20C69B931FF972676B8A1FD4B5AEF" bold="true" box="[633,721,1168,1190]" pageId="3" pageNumber="2">Figure 2</emphasis>
|
||
.
|
||
<taxonomicName id="4986ABF8B931FF9726EFB8A0FB895AEC" authority="Tropea et al., 2014" authorityName="Tropea" authorityYear="2014" box="[736,1043,1169,1191]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="gocmeni">
|
||
<emphasis id="BCF20C69B931FF9726EFB8A0FCD45AEC" box="[736,846,1169,1190]" italics="true" pageId="3" pageNumber="2">E. gocmeni</emphasis>
|
||
Tropea et al., 2014
|
||
</taxonomicName>
|
||
, Turkey, Antalya Province, Akseki District, Elmali Yolu, 36º21'29.9"N 29º28'00.1"E, FKCP.
|
||
<emphasis id="BCF20C69B931FF972640B89CFD3C5A88" bold="true" box="[591,678,1197,1219]" pageId="3" pageNumber="2">Figure 3</emphasis>
|
||
.
|
||
<taxonomicName id="4986ABF8B931FF9726BCB89FFC665A8E" authority="(C. L. Koch, 1837)" baseAuthorityName="C. L. Koch" baseAuthorityYear="1837" box="[691,1020,1197,1220]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="tergestinus">
|
||
<emphasis id="BCF20C69B931FF9726BCB89FFCAD5A89" box="[691,823,1198,1219]" italics="true" pageId="3" pageNumber="2">E. tergestinus</emphasis>
|
||
(C. L. Koch, 1837)
|
||
</taxonomicName>
|
||
, Croatia, Posedarje, 20 km E of Zadar, FKCP.
|
||
<emphasis id="BCF20C69B931FF97247CB8FBFF535A95" bold="true" box="[115,201,1226,1248]" pageId="3" pageNumber="2">Figure 4</emphasis>
|
||
.
|
||
<taxonomicName id="4986ABF8B931FF9724DAB8FAFEAB5AAA" authority="Kovařík & Šťáhlavský, 2020" authorityName="Kovařík & Šťáhlavský" authorityYear="2020" box="[213,305,1227,1248]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="janstai" status="sp. nov.">
|
||
<emphasis id="BCF20C69B931FF9724DAB8FAFEAB5AAA" box="[213,305,1227,1248]" italics="true" pageId="3" pageNumber="2">E. janstai</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A7C1B112B931FF972537B8FBFEF75AAA" box="[312,365,1226,1248]" pageId="3" pageNumber="2" rank="species">
|
||
<emphasis id="BCF20C69B931FF972537B8FBFED55A95" bold="true" box="[312,335,1226,1247]" pageId="3" pageNumber="2">sp</emphasis>
|
||
.
|
||
<emphasis id="BCF20C69B931FF972554B8FBFEF25A95" bold="true" box="[347,360,1226,1247]" pageId="3" pageNumber="2">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
, paratype.
|
||
<emphasis id="BCF20C69B931FF9725D6B8FBFDB55A95" bold="true" box="[473,559,1226,1248]" pageId="3" pageNumber="2">Figure 5</emphasis>
|
||
.
|
||
<taxonomicName id="4986ABF8B931FF972633B8FAFC105AAA" authority="(C. L. Koch, 1837)" baseAuthorityName="C. L. Koch" baseAuthorityYear="1837" box="[572,906,1226,1248]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="aquilejensis">
|
||
<emphasis id="BCF20C69B931FF972633B8FAFD505A95" box="[572,714,1226,1248]" italics="true" pageId="3" pageNumber="2">E. aquilejensis</emphasis>
|
||
(C. L. Koch, 1837)
|
||
</taxonomicName>
|
||
, Italy, Abruzzo, Silvi Marina, FKCP.
|
||
<emphasis id="BCF20C69B931FF9720FCB8FBFAD35A95" bold="true" box="[1267,1353,1226,1248]" pageId="3" pageNumber="2">Figure 6</emphasis>
|
||
.
|
||
<taxonomicName id="4986ABF8B931FF972159B8FAFEA85AB6" authority="KovarIk et al., 2014" authorityName="KovarIk" authorityYear="2014" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="yagmuri">
|
||
<emphasis id="BCF20C69B931FF972159B8FAFA5B5AAA" box="[1366,1473,1227,1248]" italics="true" pageId="3" pageNumber="2">E. yagmuri</emphasis>
|
||
KovařÍk et al., 2014
|
||
</taxonomicName>
|
||
, holotype, Greece, Epirus, Preveza District, Mt. Zalongo, Cassope (Kassope, Kassopi), 39.145278ºN 20.675556ºE, FKCP.
|
||
<emphasis id="BCF20C69B931FF97247CB935FF505B53" bold="true" box="[115,202,1284,1306]" pageId="3" pageNumber="2">Figure 7</emphasis>
|
||
.
|
||
<taxonomicName id="4986ABF8B931FF9724D8B934FE6E5B53" authority="Yagmur et al., 2013" authorityName="Yagmur" authorityYear="2013" box="[215,500,1284,1306]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="lycius">
|
||
<emphasis id="BCF20C69B931FF9724D8B934FEB15B53" box="[215,299,1284,1306]" italics="true" pageId="3" pageNumber="2">E. lycius</emphasis>
|
||
Yağmur et al., 2013
|
||
</taxonomicName>
|
||
, Turkey, Antalya Province, 36º24'N 29º41'E, 976 m a. s. l., FKCP.
|
||
<emphasis id="BCF20C69B931FF97208AB935FB465B53" bold="true" box="[1157,1244,1284,1306]" pageId="3" pageNumber="2">Figure 8</emphasis>
|
||
.
|
||
<taxonomicName id="4986ABF8B931FF9720E6B934FF3F5B7C" authority="Fet et al., 2014" authorityName="Fet" authorityYear="2014" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="solegladi">
|
||
<emphasis id="BCF20C69B931FF9720E6B934FAC75B53" box="[1257,1373,1284,1306]" italics="true" pageId="3" pageNumber="2">E. solegladi</emphasis>
|
||
Fet et al., 2014
|
||
</taxonomicName>
|
||
, paratype, Bulgaria, “Sandanski Pirin” (Blagoevgrad Province, Pirin Mts., Sandanski), NMPC.
|
||
<emphasis id="BCF20C69B931FF972037B911FB145B7F" bold="true" box="[1080,1166,1312,1334]" pageId="3" pageNumber="2">Figure 9</emphasis>
|
||
.
|
||
<taxonomicName id="4986ABF8B931FF972096B910FA275B7C" authority="Di Caporiacco, 1950" authorityName="Di Caporiacco" authorityYear="1950" box="[1177,1469,1312,1335]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="hadzii">
|
||
<emphasis id="BCF20C69B931FF972096B910FB755B7C" box="[1177,1263,1313,1334]" italics="true" pageId="3" pageNumber="2">E. hadzii</emphasis>
|
||
Di Caporiacco, 1950
|
||
</taxonomicName>
|
||
, Albania, Prokletije Mts., road Theti–Boge, FKCP.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8E39D07BB931FF97247EBA6DFE17583C" blockId="3.[113,397,1628,1654]" box="[113,397,1628,1654]" pageId="3" pageNumber="2">
|
||
<emphasis id="BCF20C69B931FF97247EBA6DFE1C583C" bold="true" box="[113,390,1628,1654]" pageId="3" pageNumber="2">Taxonomic characters</emphasis>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8E39D07BB931FF97247EBAAFFBA45898" blockId="3.[113,772,1694,2038]" lastBlockId="3.[816,1474,1402,1747]" pageId="3" pageNumber="2">
|
||
Characters important for distinguishing species in the genus
|
||
<taxonomicName id="4986ABF8B931FF9724B5BA8FFEA8589F" authorityName="Thorell" authorityYear="1876" box="[186,306,1726,1749]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="BCF20C69B931FF9724B5BA8FFEA8589F" box="[186,306,1726,1749]" italics="true" pageId="3" pageNumber="2">Euscorpius</emphasis>
|
||
</taxonomicName>
|
||
are: (1) trichobothriotaxy of the pedipalp patella, i. e., number of trichobothria in the external series
|
||
<emphasis id="BCF20C69B931FF97247EBACFFF10595C" box="[113,138,1790,1814]" italics="true" pageId="3" pageNumber="2">eb</emphasis>
|
||
,
|
||
<emphasis id="BCF20C69B931FF972496BACFFF215956" box="[153,187,1790,1820]" italics="true" pageId="3" pageNumber="2">
|
||
eb
|
||
<subScript id="1202D23EB931FF9724BCBB3FFF215956" attach="right" box="[179,187,1806,1820]" fontSize="6" pageId="3" pageNumber="2">a</subScript>
|
||
</emphasis>
|
||
,
|
||
<emphasis id="BCF20C69B931FF9724C6BACFFF77595C" box="[201,237,1790,1814]" italics="true" pageId="3" pageNumber="2">esb</emphasis>
|
||
,
|
||
<emphasis id="BCF20C69B931FF9724F3BACFFE81595F" box="[252,283,1790,1813]" italics="true" pageId="3" pageNumber="2">em</emphasis>
|
||
,
|
||
<emphasis id="BCF20C69B931FF972525BACFFED2595F" box="[298,328,1790,1813]" italics="true" pageId="3" pageNumber="2">est</emphasis>
|
||
, and
|
||
<emphasis id="BCF20C69B931FF97258ABACFFE02595F" box="[389,408,1790,1813]" italics="true" pageId="3" pageNumber="2">et</emphasis>
|
||
, in combination with the number of ventral trichobothria; (2) shape and dimensions of the carapace, pedipalp and metasomal segments; (3) granulation and carination of metasomal segments; (4) coloration; and (5) pectinal tooth count. Two currently valid subgenera can be separated by the number of ventral trichobothria on the pedipalp chela manus, i. e.,
|
||
<quantity id="497E7D9EB931FF9725BABB8FFE79599C" box="[437,483,1982,2006]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="3" pageNumber="2" unit="in" value="4.0">4 in</quantity>
|
||
<taxonomicName id="4986ABF8B931FF9725E0BB8FFDFD599F" authorityName="Thorell" authorityYear="1876" box="[495,615,1982,2005]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="genus" sensu="stricto">
|
||
<emphasis id="BCF20C69B931FF9725E0BB8FFDFD599F" box="[495,615,1982,2005]" italics="true" pageId="3" pageNumber="2">Euscorpius</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A7C1B112B931FF97267CBB8FFD65599C" box="[627,767,1982,2006]" pageId="3" pageNumber="2" sensu="stricto">sensu stricto</taxonomicNameLabel>
|
||
, and more than
|
||
<quantity id="497E7D9EB931FF972512BBEFFED059BC" box="[285,330,2014,2038]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" pageId="3" pageNumber="2" unit="in" value="6.0">6 in</quantity>
|
||
<taxonomicName id="4986ABF8B931FF97255ABBEFFD2C59BC" authority="Birula, 1917" authorityName="Birula" authorityYear="1917" box="[341,694,2014,2038]" class="Arachnida" family="Euscorpiidae" genus="Polytrichobothrius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="BCF20C69B931FF97255ABBEFFD8459BC" box="[341,542,2014,2038]" italics="true" pageId="3" pageNumber="2">Polytrichobothrius</emphasis>
|
||
Birula, 1917
|
||
</taxonomicName>
|
||
. These characters alone are not always sufficient for distinguishing a total of ca. 60 species, most of which were described or revalidated in the last two decades. Taxonomy of this genus remains complex and challenging. Recently, DNA analysis was suggested to be the only solution to this problem (see e. g.
|
||
<bibRefCitation id="EA17AD8AB931FF97273FBA2BFBBD5879" author="PARMAKELIS" box="[816,1063,1562,1587]" pageId="3" pageNumber="2" pagination="728 - 748" refId="ref12658" refString="PARMAKELIS, A., I. STATHI, P. KOTSAKIOZI, S. POULIKARAKOU & V. FET. 2013. Hidden diversity of Euscorpius (Scorpiones: Euscorpiidae) in Greece revealed by multilocus species-delimitation approaches. Biological Journal of the Linnean Society, 110: 728 - 748." type="journal article" year="2013">Parmakelis et al., 2013</bibRefCitation>
|
||
; Fet et al., 2018). Based on our studies of this genus and the application of diagnostic characters to the related genus
|
||
<taxonomicName id="4986ABF8B931FF9727C6BA6AFA095838" authority="Karsch, 1881" authorityName="Karsch" authorityYear="1881" box="[969,1427,1626,1651]" class="Arachnida" family="Euscorpiidae" genus="Megacormus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="BCF20C69B931FF9727C6BA6AFBCE5838" box="[969,1108,1627,1650]" italics="true" pageId="3" pageNumber="2">Megacormus</emphasis>
|
||
Karsch, 1881 (Euscorpiidae)
|
||
</taxonomicName>
|
||
and to the related family
|
||
<taxonomicName id="4986ABF8B931FF972017BA4BFAC558D8" authority="Kraepelin, 1905" authorityName="Kraepelin" authorityYear="1905" box="[1048,1375,1658,1682]" class="Arachnida" family="Scorpiopidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="family">Scorpiopidae Kraepelin, 1905</taxonomicName>
|
||
, we here propose application of two additional characters to elucidate taxonomy of
|
||
<taxonomicName id="4986ABF8B931FF9727B0BA8AFBAD5898" authorityName="Thorell" authorityYear="1876" box="[959,1079,1723,1746]" class="Arachnida" family="Euscorpiidae" genus="Euscorpius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="BCF20C69B931FF9727B0BA8AFBAD5898" box="[959,1079,1723,1746]" italics="true" pageId="3" pageNumber="2">Euscorpius</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |