treatments-xml/data/F1/6B/FB/F16BFB33FFFAFFA1FF6AF9E9FC51F995.xml
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<document id="0BF05C9FA2F48B776E823A73AA13424A" ID-CLB-Dataset="37321" ID-DOI="10.11646/zootaxa.4209.1.1" ID-GBIF-Dataset="2de5b7bf-3975-458a-b200-07ec69f4e013" ID-ISSN="1175-5326" ID-Zenodo-Dep="272646" ID-ZooBank="849BAB5C-464A-4B4A-A586-5742411EDC01" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1484407904318" checkinUser="plazi" docAuthor="Ng, Peter K. L. &amp; Castro, Peter" docDate="2016" docId="F16BFB33FFFAFFA1FF6AF9E9FC51F995" docLanguage="en" docName="zootaxa.4209.1.1.pdf.imf" docOrigin="Zootaxa 4209 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Megaesthesius migmus Ng &amp; Castro, 2016, n. sp." docType="treatment" docVersion="8" lastPageNumber="91" masterDocId="0D52834BFFA3FFFBFFFDFFD1FFCDFFB9" masterDocTitle="Revision of the family Chasmocarcinidae Serène, 1964 (Crustacea, Brachyura, Goneplacoidea)" masterLastPageNumber="182" masterPageNumber="1" pageNumber="90" updateTime="1698435566492" updateUser="ExternalLinkService">
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<mods:title id="431732E344F652B03FAB998926F916D0">Revision of the family Chasmocarcinidae Serène, 1964 (Crustacea, Brachyura, Goneplacoidea)</mods:title>
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<mods:namePart id="2DDB0DD01107EA4071526A0788797D91">Ng, Peter K. L.</mods:namePart>
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<mods:namePart id="D22C6535E3CFD763459BA05C33F4D5F9">Castro, Peter</mods:namePart>
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<taxonomicName id="BEC231A6FFFAFFA2FF6AF9E9FE7DF9EB" ID-CoL="3YTMY" box="[151,432,1592,1618]" class="Malacostraca" family="Chasmocarcinidae" genus="Megaesthesius" kingdom="Animalia" order="Decapoda" pageId="89" pageNumber="90" phylum="Arthropoda" rank="species" species="migmus" status="sp. nov.">
<emphasis id="4BB69637FFFAFFA2FF6AF9E9FE7DF9EB" bold="true" box="[151,432,1592,1618]" italics="true" pageId="89" pageNumber="90">Megaesthesius migmus</emphasis>
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(
<figureCitation id="E1F956A0FFFAFFA2FF62F98BFF34F9CA" box="[159,249,1626,1651]" captionStart="FIGURE 19" captionStartId="92.[151,250,1665,1687]" captionTargetBox="[203,1371,195,1618]" captionTargetId="figure@92.[203,1371,195,1632]" captionTargetPageId="92" captionText="FIGURE 19. Species of Megaesthesius and Alainthesius n. gen. A, B, Megaesthesius sagedae, Singapore: male 2.5 × 3.0 mm (ZRC 1995.883); C F, Megaesthesius yokoyai (after Takeda &amp; Miyake 1969 a: figs. 3 a, 4 f, 4 a, 4 e, respectively); G L, Alainthesius signatus n. sp., Madagascar: holotype male 2.5 × 3.3 mm (MNHN-IU- 2013 - 9472) (after Crosnier 1975: fig. 1 a, 1 k, 1 j, 1 b, 1 c, 1 i, respectively); M R, Megaesthesius westralia, Australia: holotype male 3.4 × 3.8 mm (after Davie 2013: figs. 1 A, B, 2 A, D, E, 3 E); S, W, Megaesthesius sagedae, Singapore: male 2.5 × 3.0 mm (ZRC 1995.885); T, X, Megaesthesius migmus n. sp., Philippines: holotype male 4.4 × 4.9 mm (NMCR); U, Y, Alainthesius bertrandi n. sp., New Caledonia: holotype male 3.3 × 4.1 mm (MNHN-IU- 2013 - 9073); V, Alainthesius bertrandi n. sp., Papua New Guinea: female 5.6 × 6.9 mm (MNHN-IU- 2013 - 9012). A, overall habitus; B, C, G, M, dorsal view of carapace; E, left P 4; F, I, frontal view of cephalothorax; H, O, frontal and anterolateral margin of carapace; N, ventral view of cephalothorax; J, right P 3; K, buccal cavity and posterior margin of epistome; Q, right P 4; R, right P 5; D, L, P, male pleon; S V, posterior margin of epistome; W Y, left thoracic sternites 7 and 8, “ supplementary plate ”, penis tube and penis. Abbreviations: cx 4 = coxa of P 4; cx 5 = coxa of P 5; p = penis; pt = calcified penis tube; sp = “ supplementary plate ”; st 7 = thoracic sternite 7; st 8 = thoracic sternite 8." httpUri="https://zenodo.org/record/272665/files/figure.png" pageId="89" pageNumber="90">Figs. 19</figureCitation>
T, X; 20A, B; 27C; 34D; 48D; 55D; 62C; 82DF; 88G; 92C; 103)
</paragraph>
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material
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.
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male (4.4 ×
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) [photographed],
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Sea, off
<collectorName id="D4372FF3FFFAFFA2FAF4F973FA5BF903" box="[1289,1430,1698,1723]" pageId="89" pageNumber="90">Pamilacan I.</collectorName>
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.
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Sea, off
<collectorName id="D4372FF3FFFAFFA2FAECF93BFA5BF8BB" box="[1297,1430,1770,1795]" pageId="89" pageNumber="90">Balicasag I.</collectorName>
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,
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Sea,
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.—
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) [photographed], northwestern
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2007, stn CP2711,
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<date id="0D7C6CE5FFFAFFA2FD1CF8ABFCAFF82A" box="[737,866,1914,1939]" pageId="89" pageNumber="90" value="2007-05-28">28.05.2007</date>
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</paragraph>
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<subSubSection id="31D819AEFFFAFFA1FF3AF84FFF1CFC1D" lastPageId="90" lastPageNumber="91" pageId="89" pageNumber="90" type="diagnosis">
<paragraph id="797D4A25FFFAFFA1FF3AF84FFEE7FCE4" blockId="89.[151,1436,1698,2011]" lastBlockId="90.[151,1437,151,1581]" lastPageId="90" lastPageNumber="91" pageId="89" pageNumber="90">
<emphasis id="4BB69637FFFAFFA2FF3AF84FFEF7F80E" bold="true" box="[199,314,1950,1975]" pageId="89" pageNumber="90">Diagnosis</emphasis>
. Carapace (
<figureCitation id="E1F956A0FFFAFFA2FE34F84FFDE7F80E" box="[457,554,1950,1975]" captionStart="FIGURE 20" captionStartId="93.[151,250,1585,1607]" captionTargetBox="[151,1436,193,1563]" captionTargetId="figure@93.[151,1436,193,1563]" captionTargetPageId="93" captionText="FIGURE 20. Overall dorsal view of species of Megaesthesius and Alainthesius n. gen. A, Megaesthesius migmus n. sp., Philippines: holotype male 4.4 × 4.9 mm (NMCR); B, Megaesthesius migmus n. sp., Philippines: paratype female 3.9 × 4.5 mm (ZRC 2015.245); C, Alainthesius signatus n. sp., Madagascar: holotype male 2.5 × 3.3 mm (MNHN-IU- 2013 - 9472); D, Alainthesius signatus n. sp., Madagascar: paratype female 2.7 × 3.0 mm (MNHN-IU- 2013 - 9471); E, F, Alainthesius bertrandi n. sp., New Caledonia: holotype male 3.3 × 4.1 mm (MNHN-IU- 2013 - 9073); G, Alainthesius bertrandi n. sp., Papua New Guinea: female 5.6 × 6.9 mm (MNHN-IU- 2013 - 9012); H, Alainthesius bertrandi n. sp., New Caledonia: female 3.4 × 4.0 mm (MNHN-IU- 2013 - 9007)." httpUri="https://zenodo.org/record/272666/files/figure.png" pageId="89" pageNumber="90">Figs. 20</figureCitation>
A, B; 103AC) subquadrate, 1.1 wider than long; front bilobed, with deep median cleft; dorsal surface with low, spherical tubercles; post-orbital margins sloping to angular anteroexternal angle marked by small tooth in some specimens. Lateral margins straight, parallel to each other, each with 3 straight, dented regions separated by deep notches, acute or obtuse tooth on posterior third of margin. Posterior margin nearly straight. Long orbits, eye peduncle (
<figureCitation id="E1F956A0FFF9FFA1FD00FF0EFCACFF41" box="[765,865,223,248]" captionStart="FIGURE 27" captionStartId="100.[151,250,1447,1469]" captionTargetBox="[151,1436,193,1426]" captionTargetId="figure@100.[151,1436,193,1426]" captionTargetPageId="100" captionText="FIGURE 27. Frontal view of species of Megaesthesius and Microtopsis. A, Microtopsis takedai, Japan: paratype female 3.6 × 2.7 mm (ZRC 2012.0146); B, Megaesthesius sagedae, Singapore: male 2.5 × 3.0 mm (ZRC 1995.883); C, Megaesthesius migmus n. sp., Philippines: holotype male 4.4 × 4.9 mm (NMCR); D, Alainthesius signatus n. sp., Madagascar: paratype female 2.7 × 3.0 mm (MNHN-IU- 2013 - 9471); E, Alainthesius bertrandi n. sp., New Caledonia: holotype male 3.3 × 4.1 mm (MNHN-IU- 2013 - 9073); F, Alainthesius bertrandi n. sp., Papua New Guinea: female 5.6 × 6.9 mm (MNHN-IU- 2013 - 9012)." httpUri="https://zenodo.org/record/272673/files/figure.png" pageId="90" pageNumber="91">Figs. 27</figureCitation>
C; 103D) conspicuously long, immobile, cornea reduced, non-pigmented. Epistome (
<figureCitation id="E1F956A0FFF9FFA1FDCFFED5FD4FFEA5" box="[562,642,260,285]" captionStart="FIGURE 19" captionStartId="92.[151,250,1665,1687]" captionTargetBox="[203,1371,195,1618]" captionTargetId="figure@92.[203,1371,195,1632]" captionTargetPageId="92" captionText="FIGURE 19. Species of Megaesthesius and Alainthesius n. gen. A, B, Megaesthesius sagedae, Singapore: male 2.5 × 3.0 mm (ZRC 1995.883); C F, Megaesthesius yokoyai (after Takeda &amp; Miyake 1969 a: figs. 3 a, 4 f, 4 a, 4 e, respectively); G L, Alainthesius signatus n. sp., Madagascar: holotype male 2.5 × 3.3 mm (MNHN-IU- 2013 - 9472) (after Crosnier 1975: fig. 1 a, 1 k, 1 j, 1 b, 1 c, 1 i, respectively); M R, Megaesthesius westralia, Australia: holotype male 3.4 × 3.8 mm (after Davie 2013: figs. 1 A, B, 2 A, D, E, 3 E); S, W, Megaesthesius sagedae, Singapore: male 2.5 × 3.0 mm (ZRC 1995.885); T, X, Megaesthesius migmus n. sp., Philippines: holotype male 4.4 × 4.9 mm (NMCR); U, Y, Alainthesius bertrandi n. sp., New Caledonia: holotype male 3.3 × 4.1 mm (MNHN-IU- 2013 - 9073); V, Alainthesius bertrandi n. sp., Papua New Guinea: female 5.6 × 6.9 mm (MNHN-IU- 2013 - 9012). A, overall habitus; B, C, G, M, dorsal view of carapace; E, left P 4; F, I, frontal view of cephalothorax; H, O, frontal and anterolateral margin of carapace; N, ventral view of cephalothorax; J, right P 3; K, buccal cavity and posterior margin of epistome; Q, right P 4; R, right P 5; D, L, P, male pleon; S V, posterior margin of epistome; W Y, left thoracic sternites 7 and 8, “ supplementary plate ”, penis tube and penis. Abbreviations: cx 4 = coxa of P 4; cx 5 = coxa of P 5; p = penis; pt = calcified penis tube; sp = “ supplementary plate ”; st 7 = thoracic sternite 7; st 8 = thoracic sternite 8." httpUri="https://zenodo.org/record/272665/files/figure.png" pageId="90" pageNumber="91">Fig. 19</figureCitation>
T) compressed, posterior margin with broad median lobe with fissure, narrow, nearly straight lateral margins without fissures. Antennule (
<figureCitation id="E1F956A0FFF9FFA1FC4CFEF6FBCAFEF9" box="[945,1031,295,320]" captionStart="FIGURE 27" captionStartId="100.[151,250,1447,1469]" captionTargetBox="[151,1436,193,1426]" captionTargetId="figure@100.[151,1436,193,1426]" captionTargetPageId="100" captionText="FIGURE 27. Frontal view of species of Megaesthesius and Microtopsis. A, Microtopsis takedai, Japan: paratype female 3.6 × 2.7 mm (ZRC 2012.0146); B, Megaesthesius sagedae, Singapore: male 2.5 × 3.0 mm (ZRC 1995.883); C, Megaesthesius migmus n. sp., Philippines: holotype male 4.4 × 4.9 mm (NMCR); D, Alainthesius signatus n. sp., Madagascar: paratype female 2.7 × 3.0 mm (MNHN-IU- 2013 - 9471); E, Alainthesius bertrandi n. sp., New Caledonia: holotype male 3.3 × 4.1 mm (MNHN-IU- 2013 - 9073); F, Alainthesius bertrandi n. sp., Papua New Guinea: female 5.6 × 6.9 mm (MNHN-IU- 2013 - 9012)." httpUri="https://zenodo.org/record/272673/files/figure.png" pageId="90" pageNumber="91">Fig. 27</figureCitation>
C) greatly enlarged, particularly in males. Third maxillipeds (
<figureCitation id="E1F956A0FFF9FFA1FE36FE9DFDECFEDD" box="[459,545,332,357]" captionStart="FIGURE 34" captionStartId="107.[151,250,1457,1479]" captionTargetBox="[188,1388,200,1429]" captionTargetId="figure@107.[151,1436,193,1456]" captionTargetPageId="107" captionText="FIGURE 34. Third maxilliped of species of Megaesthesius and Alainthesius n. gen. A, Megaesthesius sagedae, Singapore: male 2.5 × 3.0 mm (ZRC 1995.885); B, Megaesthesius yokoyai, Japan: male 3.5 × 3.8 mm (after Takeda &amp; Miyake 1969: fig. 4 c); C, Megaethesius westralia, Australia: holotype male 3.4 × 3.8 mm (after Davie 2013: fig. 2 C); D, Megaesthesius migmus n. sp., Philippines: paratype female 4.6 × 5.2 mm (ZRC 2015.243); E, Alainthesius signatus n. sp., Madagascar: holotype male 2.5 × 3.3 mm (after Crosnier 1975: fig. 1 e); F, Alainthesius bertrandi n. sp., New Caledonia: holotype male 3.3 × 4.1 mm (MNHN-IU- 2013 - 9073). Scales = 0.5 mm." httpUri="https://zenodo.org/record/272680/files/figure.png" pageId="90" pageNumber="91">Fig. 34</figureCitation>
D) nearly fill buccal cavern when closed; merus subtriangular, outer margin dentate; ischium subquadrate, dentate outer margin, longer than merus; outer margins of merus, ischium dentiform. Chelipeds (
<figureCitation id="E1F956A0FFF9FFA1FEEBFE45FEBEFE15" box="[278,371,404,429]" captionStart="FIGURE 20" captionStartId="93.[151,250,1585,1607]" captionTargetBox="[151,1436,193,1563]" captionTargetId="figure@93.[151,1436,193,1563]" captionTargetPageId="93" captionText="FIGURE 20. Overall dorsal view of species of Megaesthesius and Alainthesius n. gen. A, Megaesthesius migmus n. sp., Philippines: holotype male 4.4 × 4.9 mm (NMCR); B, Megaesthesius migmus n. sp., Philippines: paratype female 3.9 × 4.5 mm (ZRC 2015.245); C, Alainthesius signatus n. sp., Madagascar: holotype male 2.5 × 3.3 mm (MNHN-IU- 2013 - 9472); D, Alainthesius signatus n. sp., Madagascar: paratype female 2.7 × 3.0 mm (MNHN-IU- 2013 - 9471); E, F, Alainthesius bertrandi n. sp., New Caledonia: holotype male 3.3 × 4.1 mm (MNHN-IU- 2013 - 9073); G, Alainthesius bertrandi n. sp., Papua New Guinea: female 5.6 × 6.9 mm (MNHN-IU- 2013 - 9012); H, Alainthesius bertrandi n. sp., New Caledonia: female 3.4 × 4.0 mm (MNHN-IU- 2013 - 9007)." httpUri="https://zenodo.org/record/272666/files/figure.png" pageId="90" pageNumber="91">Figs. 20</figureCitation>
A, B; 103AC) subequal in length, nearly similar in both sexes; cutting margins of both chelae (
<figureCitation id="E1F956A0FFF9FFA1FF62FE66FF3FFE69" box="[159,242,439,464]" captionStart="FIGURE 48" captionStartId="121.[151,250,1288,1310]" captionTargetBox="[151,1436,217,1267]" captionTargetId="figure@121.[151,1436,193,1267]" captionTargetPageId="121" captionText="FIGURE 48. Chelae of species of Megaesthesius, Alainthesius n. gen. and Microtopsis. A, B, Microtopsis takedai, Japan: holotype male 2.4 × 3.1 mm (after Komai et al. 2012: fig. 3 B, D); C, Megaesthesius sagedae, Singapore: male 2.5 × 3.0 mm (ZRC 1995.883); D, Megaesthesius migmus n. sp., Philippines: holotype male 4.4 × 4.9 mm (NMCR); E, Megaesthesius yokoyai, Japan: male (after Takeda &amp; Miyake 1969: fig. 4 d); F, Alainthesius signatus n. sp., Madagascar: holotype male 3.5 × 3.8 mm (after Crosnier 1975: fig. 1 d); G, Alainthesius bertrandi n. sp., New Caledonia: holotype male 3.3 × 4.1 mm (MNHN- IU- 2013 - 9073); H, Alainthesius bertrandi n. sp., Papua New Guinea: female 5.6 × 6.9 mm (MNHN-IU- 2013 - 9012)." httpUri="https://zenodo.org/record/272694/files/figure.png" pageId="90" pageNumber="91">Fig. 48</figureCitation>
D) with broad, shallow teeth in both sexes; ventral margin of palm dentiform; ventral surface of cheliped merus smooth. Proportionally long ambulatory legs (
<figureCitation id="E1F956A0FFF9FFA1FCFEFE0DFC97FE4D" box="[771,858,476,501]" captionStart="FIGURE 20" captionStartId="93.[151,250,1585,1607]" captionTargetBox="[151,1436,193,1563]" captionTargetId="figure@93.[151,1436,193,1563]" captionTargetPageId="93" captionText="FIGURE 20. Overall dorsal view of species of Megaesthesius and Alainthesius n. gen. A, Megaesthesius migmus n. sp., Philippines: holotype male 4.4 × 4.9 mm (NMCR); B, Megaesthesius migmus n. sp., Philippines: paratype female 3.9 × 4.5 mm (ZRC 2015.245); C, Alainthesius signatus n. sp., Madagascar: holotype male 2.5 × 3.3 mm (MNHN-IU- 2013 - 9472); D, Alainthesius signatus n. sp., Madagascar: paratype female 2.7 × 3.0 mm (MNHN-IU- 2013 - 9471); E, F, Alainthesius bertrandi n. sp., New Caledonia: holotype male 3.3 × 4.1 mm (MNHN-IU- 2013 - 9073); G, Alainthesius bertrandi n. sp., Papua New Guinea: female 5.6 × 6.9 mm (MNHN-IU- 2013 - 9012); H, Alainthesius bertrandi n. sp., New Caledonia: female 3.4 × 4.0 mm (MNHN-IU- 2013 - 9007)." httpUri="https://zenodo.org/record/272666/files/figure.png" pageId="90" pageNumber="91">Fig. 20</figureCitation>
A, B), P5 merus not reaching front when folded. Dorsal, ventral margins of meri, dorsal margin of carpus of ambulatory legs with minute teeth. P5 merus 0.70.8 cl. Small, sharp tooth on inner margin of cheliped carpus, ventral surface of cheliped merus smooth in both sexes. Fused thoracic sternites 1, 2 (
<figureCitation id="E1F956A0FFF9FFA1FE0EFD96FD84FDD9" box="[499,585,583,608]" captionStart="FIGURE 55" captionStartId="128.[151,250,1514,1536]" captionTargetBox="[151,1436,179,1493]" captionTargetId="figure@128.[151,1436,179,1493]" captionTargetPageId="128" captionText="FIGURE 55. Anterior thoracic sternum and male pleon of species of Statommatia n. gen., Microtopsis and Megaesthesius. A, Statommatia granulosa n. sp., New Caledonia: holotype male 4.1 × 4.7 mm (MNHN-IU- 2013 - 9454); B, Microtopsis takedai, Japan: paratype male, carapace damaged (ZRC 2012.0146); C, Megaesthesius sagedae, Singapore: male 2.5 × 3.0 mm (ZRC 1995.885); D, Megaesthesius migmus n. sp., Philippines: holotype male 4.4 × 4.9 mm (NMCR); E, Alainthesius signatus n. sp., Madagascar: holotype male 2.5 × 3.3 mm (MNHN-IU- 2013 - 9472); F, Alainthesius bertrandi n. sp., New Caledonia: holotype male 3.3 × 4.1 mm (MNHN-IU- 2013 - 9073)." httpUri="https://zenodo.org/record/272701/files/figure.png" pageId="90" pageNumber="91">Fig. 55</figureCitation>
D) broadly triangular, proportionally wide, short. Male pleon with lateral margins of somite 6, fused somites 35 nearly straight; postero-lateral regions slightly swollen; telson proportionally short (
<figureCitation id="E1F956A0FFF9FFA1FE62FD5EFDC7FD11" box="[415,522,655,680]" captionStart="FIGURE 55" captionStartId="128.[151,250,1514,1536]" captionTargetBox="[151,1436,179,1493]" captionTargetId="figure@128.[151,1436,179,1493]" captionTargetPageId="128" captionText="FIGURE 55. Anterior thoracic sternum and male pleon of species of Statommatia n. gen., Microtopsis and Megaesthesius. A, Statommatia granulosa n. sp., New Caledonia: holotype male 4.1 × 4.7 mm (MNHN-IU- 2013 - 9454); B, Microtopsis takedai, Japan: paratype male, carapace damaged (ZRC 2012.0146); C, Megaesthesius sagedae, Singapore: male 2.5 × 3.0 mm (ZRC 1995.885); D, Megaesthesius migmus n. sp., Philippines: holotype male 4.4 × 4.9 mm (NMCR); E, Alainthesius signatus n. sp., Madagascar: holotype male 2.5 × 3.3 mm (MNHN-IU- 2013 - 9472); F, Alainthesius bertrandi n. sp., New Caledonia: holotype male 3.3 × 4.1 mm (MNHN-IU- 2013 - 9073)." httpUri="https://zenodo.org/record/272701/files/figure.png" pageId="90" pageNumber="91">Figs. 55</figureCitation>
D). Outer (ventral) surface of penis calcified, resembling plate between “supplementary plate”, sternite 8 (
<figureCitation id="E1F956A0FFF9FFA1FDEBFD65FDAAFD75" box="[534,615,692,717]" captionStart="FIGURE 19" captionStartId="92.[151,250,1665,1687]" captionTargetBox="[203,1371,195,1618]" captionTargetId="figure@92.[203,1371,195,1632]" captionTargetPageId="92" captionText="FIGURE 19. Species of Megaesthesius and Alainthesius n. gen. A, B, Megaesthesius sagedae, Singapore: male 2.5 × 3.0 mm (ZRC 1995.883); C F, Megaesthesius yokoyai (after Takeda &amp; Miyake 1969 a: figs. 3 a, 4 f, 4 a, 4 e, respectively); G L, Alainthesius signatus n. sp., Madagascar: holotype male 2.5 × 3.3 mm (MNHN-IU- 2013 - 9472) (after Crosnier 1975: fig. 1 a, 1 k, 1 j, 1 b, 1 c, 1 i, respectively); M R, Megaesthesius westralia, Australia: holotype male 3.4 × 3.8 mm (after Davie 2013: figs. 1 A, B, 2 A, D, E, 3 E); S, W, Megaesthesius sagedae, Singapore: male 2.5 × 3.0 mm (ZRC 1995.885); T, X, Megaesthesius migmus n. sp., Philippines: holotype male 4.4 × 4.9 mm (NMCR); U, Y, Alainthesius bertrandi n. sp., New Caledonia: holotype male 3.3 × 4.1 mm (MNHN-IU- 2013 - 9073); V, Alainthesius bertrandi n. sp., Papua New Guinea: female 5.6 × 6.9 mm (MNHN-IU- 2013 - 9012). A, overall habitus; B, C, G, M, dorsal view of carapace; E, left P 4; F, I, frontal view of cephalothorax; H, O, frontal and anterolateral margin of carapace; N, ventral view of cephalothorax; J, right P 3; K, buccal cavity and posterior margin of epistome; Q, right P 4; R, right P 5; D, L, P, male pleon; S V, posterior margin of epistome; W Y, left thoracic sternites 7 and 8, “ supplementary plate ”, penis tube and penis. Abbreviations: cx 4 = coxa of P 4; cx 5 = coxa of P 5; p = penis; pt = calcified penis tube; sp = “ supplementary plate ”; st 7 = thoracic sternite 7; st 8 = thoracic sternite 8." httpUri="https://zenodo.org/record/272665/files/figure.png" pageId="90" pageNumber="91">
Fig.
<date id="0D7C6CE5FFF9FFA1FDB7FD65FDAAFD75" box="[586,615,692,716]" pageId="90" pageNumber="91" value="1919-10">19</date>
</figureCitation>
X). G1 (
<figureCitation id="E1F956A0FFF9FFA1FD3EFD65FCD9FD75" box="[707,788,692,717]" captionStart="FIGURE 82" captionStartId="155.[151,250,1823,1845]" captionTargetBox="[177,1417,213,1802]" captionTargetId="figure@155.[151,1436,193,1802]" captionTargetPageId="155" captionText="FIGURE 82. Gonopods of species of Megaesthesius. A C, M. sagedae, Singapore: male 2.5 × 3.0 mm (ZRC 1995.885); D F, M. migmus n. sp., Philippines: holotype male 4.4 × 4.9 mm (NMCR); G I, M. yokoyai, Japan: male 3.5 × 3.8 mm (after Takeda &amp; Miyake 1969: fig. 4 h, i); J M, M. westralia, Australia: holotype male 3.4 × 3.8 mm (after Davie 2013: fig. 3 A D). A, D, G, H, J, K, ventral view of left G 1; B, E, I, K, L, dorsal view of left G 1; C, F, M, left G 2. Scales = 0.1 mm." httpUri="https://zenodo.org/record/272728/files/figure.png" pageId="90" pageNumber="91">Fig. 82</figureCitation>
D, E) slender, distal segment, straight, with spinules, apex slanted. G2 (
<figureCitation id="E1F956A0FFF9FFA1FED8FD06FEB8FD49" box="[293,373,727,752]" captionStart="FIGURE 82" captionStartId="155.[151,250,1823,1845]" captionTargetBox="[177,1417,213,1802]" captionTargetId="figure@155.[151,1436,193,1802]" captionTargetPageId="155" captionText="FIGURE 82. Gonopods of species of Megaesthesius. A C, M. sagedae, Singapore: male 2.5 × 3.0 mm (ZRC 1995.885); D F, M. migmus n. sp., Philippines: holotype male 4.4 × 4.9 mm (NMCR); G I, M. yokoyai, Japan: male 3.5 × 3.8 mm (after Takeda &amp; Miyake 1969: fig. 4 h, i); J M, M. westralia, Australia: holotype male 3.4 × 3.8 mm (after Davie 2013: fig. 3 A D). A, D, G, H, J, K, ventral view of left G 1; B, E, I, K, L, dorsal view of left G 1; C, F, M, left G 2. Scales = 0.1 mm." httpUri="https://zenodo.org/record/272728/files/figure.png" pageId="90" pageNumber="91">Fig. 82</figureCitation>
F) nearly as long as G1, straight; slender, distal segment long, curved, as long as basal segment, directed inward. Somites of female pleon (
<figureCitation id="E1F956A0FFF9FFA1FD73FD2DFD2EFCAD" box="[654,739,764,789]" captionStart="FIGURE 88" captionStartId="161.[151,250,1770,1792]" captionTargetBox="[151,1436,193,1749]" captionTargetId="figure@161.[151,1436,193,1749]" captionTargetPageId="161" captionText="FIGURE 88. Female thoracic sternum and pleon of species of Statommatia n. gen., Tenagopelta n. gen., Microtopsis and Megaesthesius. A, Statommatia apta, Philippines: female 8.5 × 10.6 mm (ZRC 2015.238); B, Statommatia malagasy n. sp., Madagascar: paratype female 9.4 × 8.1 mm (ZRC 2015.215); C, Tenagopelta pacifica n. sp., Philippines: female 9.2 × 10.4 mm (ZRC 2015.187); D, Tenagopelta brachyphallus n. sp., Australia: female 6.9 × 7.6 mm (NMV J 61680); E, Microtopsis takedai, Japan: paratype female 3.6 × 2.7 mm (ZRC 2012.0146); F, Megaesthesius sagedae, Singapore: female 3.0 × 3.6 mm (ZRC 2013.1804); G, Megaesthesius migmus, Philippines: paratype female 3.9 × 4.5 mm (ZRC 2015.245); H, Alainthesius bertrandi n. sp., Papua New Guinea: female 5.6 × 6.9 mm (MNHN-IU- 2013 - 9012). Vulvae exposed in E." httpUri="https://zenodo.org/record/272734/files/figure.png" pageId="90" pageNumber="91">Fig. 88</figureCitation>
G) with gently convex margins; telson proportionally short. Sterno-pleonal cavity of female (
<figureCitation id="E1F956A0FFF9FFA1FDF4FCCEFD96FC81" box="[521,603,799,824]" captionStart="FIGURE 92" captionStartId="165.[151,250,934,956]" captionTargetBox="[151,1436,193,912]" captionTargetId="figure@165.[151,1436,193,912]" captionTargetPageId="165" captionText="FIGURE 92. Vulvae of species of Microtopsis and Megaesthesius. A, Microtopsis takedai, Japan: paratype female 3.6 × 2.7 mm (ZRC 2012.0146); B, Megaesthesius sagedae, Singapore: female carapace length 2.9 mm, sides damaged (ZRC 1995.883); C, Megaesthesius migmus n. sp., Philippines: paratype female 3.9 × 4.5 mm (ZRC 2015.245); D, Alainthesius bertrandi n. sp., Papua New Guinea: female 5.6 × 6.9 mm (MNHN-IU- 2013 - 9012)." httpUri="https://zenodo.org/record/272738/files/figure.png" pageId="90" pageNumber="91">Fig. 92</figureCitation>
C) deep, vulvae far apart from each other on outer margins of cavity close to suture 5/6.
</paragraph>
<paragraph id="797D4A25FFF9FFA1FF3AFCB6FF1CFC1D" blockId="90.[151,1437,151,1581]" pageId="90" pageNumber="91">
<emphasis id="4BB69637FFF9FFA1FF3AFCB6FED7FC39" bold="true" box="[199,282,871,896]" pageId="90" pageNumber="91">Colour</emphasis>
. The carapace is white to pinkish-white beige in life, with white chelipeds and ambulatory legs (
<figureCitation id="E1F956A0FFF9FFA1FA92FCB6FF08FC1D" captionStart="FIGURE 103" captionStartId="175.[151,250,1974,1996]" captionTargetBox="[151,1436,1281,1952]" captionTargetId="figure@175.[151,1436,1281,1952]" captionTargetPageId="175" captionText="FIGURE 103 Megaesthesius migmus n. sp. A, Philippines: holotype male 4.4 × 4.9 mm (NMCR); B D, Philippines: paratype female 4.6 × 5.2 mm (ZRC 2015.243). [P. K. L. Ng]" httpUri="https://zenodo.org/record/272749/files/figure.png" pageId="90" pageNumber="91">Fig. 103</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="31D819AEFFF9FFA1FF3AFC7EFD1BFC54" pageId="90" pageNumber="91" type="etymology">
<paragraph id="797D4A25FFF9FFA1FF3AFC7EFD1BFC54" blockId="90.[151,1437,151,1581]" pageId="90" pageNumber="91">
<emphasis id="4BB69637FFF9FFA1FF3AFC7EFE8BFC71" bold="true" box="[199,326,943,968]" pageId="90" pageNumber="91">Etymology</emphasis>
. From the Greek
<emphasis id="4BB69637FFF9FFA1FDF1FC60FD95FC71" box="[524,600,945,968]" italics="true" pageId="90" pageNumber="91">migma</emphasis>
for “mixture” in reference to the mixture of characters of the other species of
<taxonomicName id="BEC231A6FFF9FFA1FF48FC05FE97FC54" box="[181,346,980,1005]" class="Malacostraca" family="Chasmocarcinidae" genus="Megaesthesius" kingdom="Animalia" order="Decapoda" pageId="90" pageNumber="91" phylum="Arthropoda" rank="genus">
<emphasis id="4BB69637FFF9FFA1FF48FC05FE97FC54" box="[181,346,980,1005]" italics="true" pageId="90" pageNumber="91">Megaesthesius</emphasis>
</taxonomicName>
that characterise the new species.
</paragraph>
</subSubSection>
<subSubSection id="31D819AEFFF9FFA1FF3AFC26FF22F9B1" pageId="90" pageNumber="91" type="discussion">
<paragraph id="797D4A25FFF9FFA1FF3AFC26FEC9FAB4" blockId="90.[151,1437,151,1581]" pageId="90" pageNumber="91">
<emphasis id="4BB69637FFF9FFA1FF3AFC26FEF9FBA9" bold="true" box="[199,308,1015,1040]" pageId="90" pageNumber="91">Remarks</emphasis>
.
<taxonomicName id="BEC231A6FFF9FFA1FEB9FC26FD87FBA9" box="[324,586,1015,1040]" class="Malacostraca" family="Chasmocarcinidae" genus="Megaesthesius" kingdom="Animalia" order="Decapoda" pageId="90" pageNumber="91" phylum="Arthropoda" rank="species" species="migmus" status="sp. nov.">
<emphasis id="4BB69637FFF9FFA1FEB9FC26FD87FBA9" box="[324,586,1015,1040]" italics="true" pageId="90" pageNumber="91">Megaesthesius migmus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="50852B4CFFF9FFA1FDAEFC26FD59FBA9" box="[595,660,1015,1040]" pageId="90" pageNumber="91" rank="species">
<emphasis id="4BB69637FFF9FFA1FDAEFC26FD59FBA9" bold="true" box="[595,660,1015,1040]" pageId="90" pageNumber="91">n. sp.</emphasis>
</taxonomicNameLabel>
is distinct from
<taxonomicName id="BEC231A6FFF9FFA1FCABFC28FC17FBA9" box="[854,986,1015,1040]" class="Malacostraca" family="Chasmocarcinidae" genus="Megaesthesius" kingdom="Animalia" order="Decapoda" pageId="90" pageNumber="91" phylum="Arthropoda" rank="species" species="sagedae">
<emphasis id="4BB69637FFF9FFA1FCABFC28FC17FBA9" box="[854,986,1015,1040]" italics="true" pageId="90" pageNumber="91">M. sagedae</emphasis>
</taxonomicName>
in having the carapace proportionately wider (
<figureCitation id="E1F956A0FFF9FFA1FF1BFBCDFEFAFB8D" box="[230,311,1052,1077]" captionStart="FIGURE 20" captionStartId="93.[151,250,1585,1607]" captionTargetBox="[151,1436,193,1563]" captionTargetId="figure@93.[151,1436,193,1563]" captionTargetPageId="93" captionText="FIGURE 20. Overall dorsal view of species of Megaesthesius and Alainthesius n. gen. A, Megaesthesius migmus n. sp., Philippines: holotype male 4.4 × 4.9 mm (NMCR); B, Megaesthesius migmus n. sp., Philippines: paratype female 3.9 × 4.5 mm (ZRC 2015.245); C, Alainthesius signatus n. sp., Madagascar: holotype male 2.5 × 3.3 mm (MNHN-IU- 2013 - 9472); D, Alainthesius signatus n. sp., Madagascar: paratype female 2.7 × 3.0 mm (MNHN-IU- 2013 - 9471); E, F, Alainthesius bertrandi n. sp., New Caledonia: holotype male 3.3 × 4.1 mm (MNHN-IU- 2013 - 9073); G, Alainthesius bertrandi n. sp., Papua New Guinea: female 5.6 × 6.9 mm (MNHN-IU- 2013 - 9012); H, Alainthesius bertrandi n. sp., New Caledonia: female 3.4 × 4.0 mm (MNHN-IU- 2013 - 9007)." httpUri="https://zenodo.org/record/272666/files/figure.png" pageId="90" pageNumber="91">Fig. 20</figureCitation>
A, B versus
<figureCitation id="E1F956A0FFF9FFA1FE3DFBCDFDC2FB8D" box="[448,527,1052,1077]" captionStart="FIGURE 19" captionStartId="92.[151,250,1665,1687]" captionTargetBox="[203,1371,195,1618]" captionTargetId="figure@92.[203,1371,195,1632]" captionTargetPageId="92" captionText="FIGURE 19. Species of Megaesthesius and Alainthesius n. gen. A, B, Megaesthesius sagedae, Singapore: male 2.5 × 3.0 mm (ZRC 1995.883); C F, Megaesthesius yokoyai (after Takeda &amp; Miyake 1969 a: figs. 3 a, 4 f, 4 a, 4 e, respectively); G L, Alainthesius signatus n. sp., Madagascar: holotype male 2.5 × 3.3 mm (MNHN-IU- 2013 - 9472) (after Crosnier 1975: fig. 1 a, 1 k, 1 j, 1 b, 1 c, 1 i, respectively); M R, Megaesthesius westralia, Australia: holotype male 3.4 × 3.8 mm (after Davie 2013: figs. 1 A, B, 2 A, D, E, 3 E); S, W, Megaesthesius sagedae, Singapore: male 2.5 × 3.0 mm (ZRC 1995.885); T, X, Megaesthesius migmus n. sp., Philippines: holotype male 4.4 × 4.9 mm (NMCR); U, Y, Alainthesius bertrandi n. sp., New Caledonia: holotype male 3.3 × 4.1 mm (MNHN-IU- 2013 - 9073); V, Alainthesius bertrandi n. sp., Papua New Guinea: female 5.6 × 6.9 mm (MNHN-IU- 2013 - 9012). A, overall habitus; B, C, G, M, dorsal view of carapace; E, left P 4; F, I, frontal view of cephalothorax; H, O, frontal and anterolateral margin of carapace; N, ventral view of cephalothorax; J, right P 3; K, buccal cavity and posterior margin of epistome; Q, right P 4; R, right P 5; D, L, P, male pleon; S V, posterior margin of epistome; W Y, left thoracic sternites 7 and 8, “ supplementary plate ”, penis tube and penis. Abbreviations: cx 4 = coxa of P 4; cx 5 = coxa of P 5; p = penis; pt = calcified penis tube; sp = “ supplementary plate ”; st 7 = thoracic sternite 7; st 8 = thoracic sternite 8." httpUri="https://zenodo.org/record/272665/files/figure.png" pageId="90" pageNumber="91">Fig. 19</figureCitation>
A, B), the lateral margin of the carapace having distinct lobiform teeth (
<figureCitation id="E1F956A0FFF9FFA1FACCFBCDFA4FFB8D" box="[1329,1410,1052,1077]" captionStart="FIGURE 20" captionStartId="93.[151,250,1585,1607]" captionTargetBox="[151,1436,193,1563]" captionTargetId="figure@93.[151,1436,193,1563]" captionTargetPageId="93" captionText="FIGURE 20. Overall dorsal view of species of Megaesthesius and Alainthesius n. gen. A, Megaesthesius migmus n. sp., Philippines: holotype male 4.4 × 4.9 mm (NMCR); B, Megaesthesius migmus n. sp., Philippines: paratype female 3.9 × 4.5 mm (ZRC 2015.245); C, Alainthesius signatus n. sp., Madagascar: holotype male 2.5 × 3.3 mm (MNHN-IU- 2013 - 9472); D, Alainthesius signatus n. sp., Madagascar: paratype female 2.7 × 3.0 mm (MNHN-IU- 2013 - 9471); E, F, Alainthesius bertrandi n. sp., New Caledonia: holotype male 3.3 × 4.1 mm (MNHN-IU- 2013 - 9073); G, Alainthesius bertrandi n. sp., Papua New Guinea: female 5.6 × 6.9 mm (MNHN-IU- 2013 - 9012); H, Alainthesius bertrandi n. sp., New Caledonia: female 3.4 × 4.0 mm (MNHN-IU- 2013 - 9007)." httpUri="https://zenodo.org/record/272666/files/figure.png" pageId="90" pageNumber="91">Fig. 20</figureCitation>
A, B) (only small spines in
<taxonomicName id="BEC231A6FFF9FFA1FE55FB90FDE4FBE1" box="[424,553,1087,1112]" class="Malacostraca" family="Chasmocarcinidae" genus="Megaesthesius" kingdom="Animalia" order="Decapoda" pageId="90" pageNumber="91" phylum="Arthropoda" rank="species" species="sagedae">
<emphasis id="4BB69637FFF9FFA1FE55FB90FDE4FBE1" box="[424,553,1087,1112]" italics="true" pageId="90" pageNumber="91">M. sagedae</emphasis>
</taxonomicName>
,
<figureCitation id="E1F956A0FFF9FFA1FDCAFBEEFD4BFBE1" box="[567,646,1087,1112]" captionStart="FIGURE 19" captionStartId="92.[151,250,1665,1687]" captionTargetBox="[203,1371,195,1618]" captionTargetId="figure@92.[203,1371,195,1632]" captionTargetPageId="92" captionText="FIGURE 19. Species of Megaesthesius and Alainthesius n. gen. A, B, Megaesthesius sagedae, Singapore: male 2.5 × 3.0 mm (ZRC 1995.883); C F, Megaesthesius yokoyai (after Takeda &amp; Miyake 1969 a: figs. 3 a, 4 f, 4 a, 4 e, respectively); G L, Alainthesius signatus n. sp., Madagascar: holotype male 2.5 × 3.3 mm (MNHN-IU- 2013 - 9472) (after Crosnier 1975: fig. 1 a, 1 k, 1 j, 1 b, 1 c, 1 i, respectively); M R, Megaesthesius westralia, Australia: holotype male 3.4 × 3.8 mm (after Davie 2013: figs. 1 A, B, 2 A, D, E, 3 E); S, W, Megaesthesius sagedae, Singapore: male 2.5 × 3.0 mm (ZRC 1995.885); T, X, Megaesthesius migmus n. sp., Philippines: holotype male 4.4 × 4.9 mm (NMCR); U, Y, Alainthesius bertrandi n. sp., New Caledonia: holotype male 3.3 × 4.1 mm (MNHN-IU- 2013 - 9073); V, Alainthesius bertrandi n. sp., Papua New Guinea: female 5.6 × 6.9 mm (MNHN-IU- 2013 - 9012). A, overall habitus; B, C, G, M, dorsal view of carapace; E, left P 4; F, I, frontal view of cephalothorax; H, O, frontal and anterolateral margin of carapace; N, ventral view of cephalothorax; J, right P 3; K, buccal cavity and posterior margin of epistome; Q, right P 4; R, right P 5; D, L, P, male pleon; S V, posterior margin of epistome; W Y, left thoracic sternites 7 and 8, “ supplementary plate ”, penis tube and penis. Abbreviations: cx 4 = coxa of P 4; cx 5 = coxa of P 5; p = penis; pt = calcified penis tube; sp = “ supplementary plate ”; st 7 = thoracic sternite 7; st 8 = thoracic sternite 8." httpUri="https://zenodo.org/record/272665/files/figure.png" pageId="90" pageNumber="91">Fig. 19</figureCitation>
A, B), the aesthetascs of the antennules are distinctly shorter (
<figureCitation id="E1F956A0FFF9FFA1FAC4FBEEFA44FBE1" box="[1337,1417,1087,1112]" captionStart="FIGURE 27" captionStartId="100.[151,250,1447,1469]" captionTargetBox="[151,1436,193,1426]" captionTargetId="figure@100.[151,1436,193,1426]" captionTargetPageId="100" captionText="FIGURE 27. Frontal view of species of Megaesthesius and Microtopsis. A, Microtopsis takedai, Japan: paratype female 3.6 × 2.7 mm (ZRC 2012.0146); B, Megaesthesius sagedae, Singapore: male 2.5 × 3.0 mm (ZRC 1995.883); C, Megaesthesius migmus n. sp., Philippines: holotype male 4.4 × 4.9 mm (NMCR); D, Alainthesius signatus n. sp., Madagascar: paratype female 2.7 × 3.0 mm (MNHN-IU- 2013 - 9471); E, Alainthesius bertrandi n. sp., New Caledonia: holotype male 3.3 × 4.1 mm (MNHN-IU- 2013 - 9073); F, Alainthesius bertrandi n. sp., Papua New Guinea: female 5.6 × 6.9 mm (MNHN-IU- 2013 - 9012)." httpUri="https://zenodo.org/record/272673/files/figure.png" pageId="90" pageNumber="91">Fig. 27</figureCitation>
C versus
<figureCitation id="E1F956A0FFF9FFA1FF18FBB5FEFAFBC5" box="[229,311,1124,1149]" captionStart="FIGURE 27" captionStartId="100.[151,250,1447,1469]" captionTargetBox="[151,1436,193,1426]" captionTargetId="figure@100.[151,1436,193,1426]" captionTargetPageId="100" captionText="FIGURE 27. Frontal view of species of Megaesthesius and Microtopsis. A, Microtopsis takedai, Japan: paratype female 3.6 × 2.7 mm (ZRC 2012.0146); B, Megaesthesius sagedae, Singapore: male 2.5 × 3.0 mm (ZRC 1995.883); C, Megaesthesius migmus n. sp., Philippines: holotype male 4.4 × 4.9 mm (NMCR); D, Alainthesius signatus n. sp., Madagascar: paratype female 2.7 × 3.0 mm (MNHN-IU- 2013 - 9471); E, Alainthesius bertrandi n. sp., New Caledonia: holotype male 3.3 × 4.1 mm (MNHN-IU- 2013 - 9073); F, Alainthesius bertrandi n. sp., Papua New Guinea: female 5.6 × 6.9 mm (MNHN-IU- 2013 - 9012)." httpUri="https://zenodo.org/record/272673/files/figure.png" pageId="90" pageNumber="91">Fig. 27</figureCitation>
B); the fingers of the chela are relatively more slender and longer (
<figureCitation id="E1F956A0FFF9FFA1FBE4FBB5FBA4FBC5" box="[1049,1129,1124,1149]" captionStart="FIGURE 48" captionStartId="121.[151,250,1288,1310]" captionTargetBox="[151,1436,217,1267]" captionTargetId="figure@121.[151,1436,193,1267]" captionTargetPageId="121" captionText="FIGURE 48. Chelae of species of Megaesthesius, Alainthesius n. gen. and Microtopsis. A, B, Microtopsis takedai, Japan: holotype male 2.4 × 3.1 mm (after Komai et al. 2012: fig. 3 B, D); C, Megaesthesius sagedae, Singapore: male 2.5 × 3.0 mm (ZRC 1995.883); D, Megaesthesius migmus n. sp., Philippines: holotype male 4.4 × 4.9 mm (NMCR); E, Megaesthesius yokoyai, Japan: male (after Takeda &amp; Miyake 1969: fig. 4 d); F, Alainthesius signatus n. sp., Madagascar: holotype male 3.5 × 3.8 mm (after Crosnier 1975: fig. 1 d); G, Alainthesius bertrandi n. sp., New Caledonia: holotype male 3.3 × 4.1 mm (MNHN- IU- 2013 - 9073); H, Alainthesius bertrandi n. sp., Papua New Guinea: female 5.6 × 6.9 mm (MNHN-IU- 2013 - 9012)." httpUri="https://zenodo.org/record/272694/files/figure.png" pageId="90" pageNumber="91">Fig. 48</figureCitation>
C versus
<figureCitation id="E1F956A0FFF9FFA1FB2DFBB5FAECFBC5" box="[1232,1313,1124,1149]" captionStart="FIGURE 48" captionStartId="121.[151,250,1288,1310]" captionTargetBox="[151,1436,217,1267]" captionTargetId="figure@121.[151,1436,193,1267]" captionTargetPageId="121" captionText="FIGURE 48. Chelae of species of Megaesthesius, Alainthesius n. gen. and Microtopsis. A, B, Microtopsis takedai, Japan: holotype male 2.4 × 3.1 mm (after Komai et al. 2012: fig. 3 B, D); C, Megaesthesius sagedae, Singapore: male 2.5 × 3.0 mm (ZRC 1995.883); D, Megaesthesius migmus n. sp., Philippines: holotype male 4.4 × 4.9 mm (NMCR); E, Megaesthesius yokoyai, Japan: male (after Takeda &amp; Miyake 1969: fig. 4 d); F, Alainthesius signatus n. sp., Madagascar: holotype male 3.5 × 3.8 mm (after Crosnier 1975: fig. 1 d); G, Alainthesius bertrandi n. sp., New Caledonia: holotype male 3.3 × 4.1 mm (MNHN- IU- 2013 - 9073); H, Alainthesius bertrandi n. sp., Papua New Guinea: female 5.6 × 6.9 mm (MNHN-IU- 2013 - 9012)." httpUri="https://zenodo.org/record/272694/files/figure.png" pageId="90" pageNumber="91">Fig. 48</figureCitation>
D); and the G1 is relatively longer and more slender (
<figureCitation id="E1F956A0FFF9FFA1FD95FB56FD74FB19" box="[616,697,1159,1184]" captionStart="FIGURE 82" captionStartId="155.[151,250,1823,1845]" captionTargetBox="[177,1417,213,1802]" captionTargetId="figure@155.[151,1436,193,1802]" captionTargetPageId="155" captionText="FIGURE 82. Gonopods of species of Megaesthesius. A C, M. sagedae, Singapore: male 2.5 × 3.0 mm (ZRC 1995.885); D F, M. migmus n. sp., Philippines: holotype male 4.4 × 4.9 mm (NMCR); G I, M. yokoyai, Japan: male 3.5 × 3.8 mm (after Takeda &amp; Miyake 1969: fig. 4 h, i); J M, M. westralia, Australia: holotype male 3.4 × 3.8 mm (after Davie 2013: fig. 3 A D). A, D, G, H, J, K, ventral view of left G 1; B, E, I, K, L, dorsal view of left G 1; C, F, M, left G 2. Scales = 0.1 mm." httpUri="https://zenodo.org/record/272728/files/figure.png" pageId="90" pageNumber="91">Fig. 82</figureCitation>
D, E in contrast to
<figureCitation id="E1F956A0FFF9FFA1FC76FB56FC16FB19" box="[907,987,1159,1184]" captionStart="FIGURE 82" captionStartId="155.[151,250,1823,1845]" captionTargetBox="[177,1417,213,1802]" captionTargetId="figure@155.[151,1436,193,1802]" captionTargetPageId="155" captionText="FIGURE 82. Gonopods of species of Megaesthesius. A C, M. sagedae, Singapore: male 2.5 × 3.0 mm (ZRC 1995.885); D F, M. migmus n. sp., Philippines: holotype male 4.4 × 4.9 mm (NMCR); G I, M. yokoyai, Japan: male 3.5 × 3.8 mm (after Takeda &amp; Miyake 1969: fig. 4 h, i); J M, M. westralia, Australia: holotype male 3.4 × 3.8 mm (after Davie 2013: fig. 3 A D). A, D, G, H, J, K, ventral view of left G 1; B, E, I, K, L, dorsal view of left G 1; C, F, M, left G 2. Scales = 0.1 mm." httpUri="https://zenodo.org/record/272728/files/figure.png" pageId="90" pageNumber="91">Fig. 82</figureCitation>
A, B). As discussed earlier, the calcified plate on the surface of the penis is mobile in the single male of
<taxonomicName id="BEC231A6FFF9FFA1FCA6FB7CFC15FB7D" box="[859,984,1197,1220]" class="Malacostraca" family="Chasmocarcinidae" genus="Megaesthesius" kingdom="Animalia" order="Decapoda" pageId="90" pageNumber="91" phylum="Arthropoda" rank="species" species="migmus">
<emphasis id="4BB69637FFF9FFA1FCA6FB7CFC15FB7D" box="[859,984,1197,1220]" italics="true" pageId="90" pageNumber="91">M. migmus</emphasis>
</taxonomicName>
examined (
<figureCitation id="E1F956A0FFF9FFA1FBA6FB7DFB67FB7D" box="[1115,1194,1196,1221]" captionStart="FIGURE 19" captionStartId="92.[151,250,1665,1687]" captionTargetBox="[203,1371,195,1618]" captionTargetId="figure@92.[203,1371,195,1632]" captionTargetPageId="92" captionText="FIGURE 19. Species of Megaesthesius and Alainthesius n. gen. A, B, Megaesthesius sagedae, Singapore: male 2.5 × 3.0 mm (ZRC 1995.883); C F, Megaesthesius yokoyai (after Takeda &amp; Miyake 1969 a: figs. 3 a, 4 f, 4 a, 4 e, respectively); G L, Alainthesius signatus n. sp., Madagascar: holotype male 2.5 × 3.3 mm (MNHN-IU- 2013 - 9472) (after Crosnier 1975: fig. 1 a, 1 k, 1 j, 1 b, 1 c, 1 i, respectively); M R, Megaesthesius westralia, Australia: holotype male 3.4 × 3.8 mm (after Davie 2013: figs. 1 A, B, 2 A, D, E, 3 E); S, W, Megaesthesius sagedae, Singapore: male 2.5 × 3.0 mm (ZRC 1995.885); T, X, Megaesthesius migmus n. sp., Philippines: holotype male 4.4 × 4.9 mm (NMCR); U, Y, Alainthesius bertrandi n. sp., New Caledonia: holotype male 3.3 × 4.1 mm (MNHN-IU- 2013 - 9073); V, Alainthesius bertrandi n. sp., Papua New Guinea: female 5.6 × 6.9 mm (MNHN-IU- 2013 - 9012). A, overall habitus; B, C, G, M, dorsal view of carapace; E, left P 4; F, I, frontal view of cephalothorax; H, O, frontal and anterolateral margin of carapace; N, ventral view of cephalothorax; J, right P 3; K, buccal cavity and posterior margin of epistome; Q, right P 4; R, right P 5; D, L, P, male pleon; S V, posterior margin of epistome; W Y, left thoracic sternites 7 and 8, “ supplementary plate ”, penis tube and penis. Abbreviations: cx 4 = coxa of P 4; cx 5 = coxa of P 5; p = penis; pt = calcified penis tube; sp = “ supplementary plate ”; st 7 = thoracic sternite 7; st 8 = thoracic sternite 8." httpUri="https://zenodo.org/record/272665/files/figure.png" pageId="90" pageNumber="91">
Fig.
<date id="0D7C6CE5FFF9FFA1FB72FB7DFB67FB7D" box="[1167,1194,1196,1220]" pageId="90" pageNumber="91" value="1919-10">19</date>
</figureCitation>
X) but is immobile in
<taxonomicName id="BEC231A6FFF9FFA1FF6AFB00FED6FB51" box="[151,283,1231,1256]" class="Malacostraca" family="Chasmocarcinidae" genus="Megaesthesius" kingdom="Animalia" order="Decapoda" pageId="90" pageNumber="91" phylum="Arthropoda" rank="species" species="sagedae">
<emphasis id="4BB69637FFF9FFA1FF6AFB00FED6FB51" box="[151,283,1231,1256]" italics="true" pageId="90" pageNumber="91">M. sagedae</emphasis>
</taxonomicName>
(
<figureCitation id="E1F956A0FFF9FFA1FED0FB1EFE4DFB51" box="[301,384,1231,1256]" captionStart="FIGURE 19" captionStartId="92.[151,250,1665,1687]" captionTargetBox="[203,1371,195,1618]" captionTargetId="figure@92.[203,1371,195,1632]" captionTargetPageId="92" captionText="FIGURE 19. Species of Megaesthesius and Alainthesius n. gen. A, B, Megaesthesius sagedae, Singapore: male 2.5 × 3.0 mm (ZRC 1995.883); C F, Megaesthesius yokoyai (after Takeda &amp; Miyake 1969 a: figs. 3 a, 4 f, 4 a, 4 e, respectively); G L, Alainthesius signatus n. sp., Madagascar: holotype male 2.5 × 3.3 mm (MNHN-IU- 2013 - 9472) (after Crosnier 1975: fig. 1 a, 1 k, 1 j, 1 b, 1 c, 1 i, respectively); M R, Megaesthesius westralia, Australia: holotype male 3.4 × 3.8 mm (after Davie 2013: figs. 1 A, B, 2 A, D, E, 3 E); S, W, Megaesthesius sagedae, Singapore: male 2.5 × 3.0 mm (ZRC 1995.885); T, X, Megaesthesius migmus n. sp., Philippines: holotype male 4.4 × 4.9 mm (NMCR); U, Y, Alainthesius bertrandi n. sp., New Caledonia: holotype male 3.3 × 4.1 mm (MNHN-IU- 2013 - 9073); V, Alainthesius bertrandi n. sp., Papua New Guinea: female 5.6 × 6.9 mm (MNHN-IU- 2013 - 9012). A, overall habitus; B, C, G, M, dorsal view of carapace; E, left P 4; F, I, frontal view of cephalothorax; H, O, frontal and anterolateral margin of carapace; N, ventral view of cephalothorax; J, right P 3; K, buccal cavity and posterior margin of epistome; Q, right P 4; R, right P 5; D, L, P, male pleon; S V, posterior margin of epistome; W Y, left thoracic sternites 7 and 8, “ supplementary plate ”, penis tube and penis. Abbreviations: cx 4 = coxa of P 4; cx 5 = coxa of P 5; p = penis; pt = calcified penis tube; sp = “ supplementary plate ”; st 7 = thoracic sternite 7; st 8 = thoracic sternite 8." httpUri="https://zenodo.org/record/272665/files/figure.png" pageId="90" pageNumber="91">Fig. 19</figureCitation>
W); but with one male of the former known, it is not clear if this is also a species-specific character.
</paragraph>
<paragraph id="797D4A25FFF9FFA1FF3AFAC6FF22F9B1" blockId="90.[151,1437,151,1581]" pageId="90" pageNumber="91">
Compared to
<taxonomicName id="BEC231A6FFF9FFA1FE99FAC8FE2EFA89" box="[356,483,1303,1328]" class="Malacostraca" family="Chasmocarcinidae" genus="Megaesthesius" kingdom="Animalia" order="Decapoda" pageId="90" pageNumber="91" phylum="Arthropoda" rank="species" species="yokoyai">
<emphasis id="4BB69637FFF9FFA1FE99FAC8FE2EFA89" box="[356,483,1303,1328]" italics="true" pageId="90" pageNumber="91">M. yokoyai</emphasis>
</taxonomicName>
,
<taxonomicName id="BEC231A6FFF9FFA1FE0EFAC8FDBFFA89" box="[499,626,1305,1328]" class="Malacostraca" family="Chasmocarcinidae" genus="Megaesthesius" kingdom="Animalia" order="Decapoda" pageId="90" pageNumber="91" phylum="Arthropoda" rank="species" species="migmus" status="sp. nov.">
<emphasis id="4BB69637FFF9FFA1FE0EFAC8FDBFFA89" box="[499,626,1305,1328]" italics="true" pageId="90" pageNumber="91">M. migmus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="50852B4CFFF9FFA1FD81FAC6FD70FA89" box="[636,701,1303,1328]" pageId="90" pageNumber="91" rank="species">
<emphasis id="4BB69637FFF9FFA1FD81FAC6FD70FA89" bold="true" box="[636,701,1303,1328]" pageId="90" pageNumber="91">n. sp.</emphasis>
</taxonomicNameLabel>
is easily distinguished by its proportionately longer ambulatory legs (
<figureCitation id="E1F956A0FFF9FFA1FF28FAEDFEE4FAED" box="[213,297,1340,1365]" captionStart="FIGURE 20" captionStartId="93.[151,250,1585,1607]" captionTargetBox="[151,1436,193,1563]" captionTargetId="figure@93.[151,1436,193,1563]" captionTargetPageId="93" captionText="FIGURE 20. Overall dorsal view of species of Megaesthesius and Alainthesius n. gen. A, Megaesthesius migmus n. sp., Philippines: holotype male 4.4 × 4.9 mm (NMCR); B, Megaesthesius migmus n. sp., Philippines: paratype female 3.9 × 4.5 mm (ZRC 2015.245); C, Alainthesius signatus n. sp., Madagascar: holotype male 2.5 × 3.3 mm (MNHN-IU- 2013 - 9472); D, Alainthesius signatus n. sp., Madagascar: paratype female 2.7 × 3.0 mm (MNHN-IU- 2013 - 9471); E, F, Alainthesius bertrandi n. sp., New Caledonia: holotype male 3.3 × 4.1 mm (MNHN-IU- 2013 - 9073); G, Alainthesius bertrandi n. sp., Papua New Guinea: female 5.6 × 6.9 mm (MNHN-IU- 2013 - 9012); H, Alainthesius bertrandi n. sp., New Caledonia: female 3.4 × 4.0 mm (MNHN-IU- 2013 - 9007)." httpUri="https://zenodo.org/record/272666/files/figure.png" pageId="90" pageNumber="91">Fig. 20</figureCitation>
A, B versus
<figureCitation id="E1F956A0FFF9FFA1FE4AFAEDFDC7FAED" box="[439,522,1340,1365]" captionStart="FIGURE 19" captionStartId="92.[151,250,1665,1687]" captionTargetBox="[203,1371,195,1618]" captionTargetId="figure@92.[203,1371,195,1632]" captionTargetPageId="92" captionText="FIGURE 19. Species of Megaesthesius and Alainthesius n. gen. A, B, Megaesthesius sagedae, Singapore: male 2.5 × 3.0 mm (ZRC 1995.883); C F, Megaesthesius yokoyai (after Takeda &amp; Miyake 1969 a: figs. 3 a, 4 f, 4 a, 4 e, respectively); G L, Alainthesius signatus n. sp., Madagascar: holotype male 2.5 × 3.3 mm (MNHN-IU- 2013 - 9472) (after Crosnier 1975: fig. 1 a, 1 k, 1 j, 1 b, 1 c, 1 i, respectively); M R, Megaesthesius westralia, Australia: holotype male 3.4 × 3.8 mm (after Davie 2013: figs. 1 A, B, 2 A, D, E, 3 E); S, W, Megaesthesius sagedae, Singapore: male 2.5 × 3.0 mm (ZRC 1995.885); T, X, Megaesthesius migmus n. sp., Philippines: holotype male 4.4 × 4.9 mm (NMCR); U, Y, Alainthesius bertrandi n. sp., New Caledonia: holotype male 3.3 × 4.1 mm (MNHN-IU- 2013 - 9073); V, Alainthesius bertrandi n. sp., Papua New Guinea: female 5.6 × 6.9 mm (MNHN-IU- 2013 - 9012). A, overall habitus; B, C, G, M, dorsal view of carapace; E, left P 4; F, I, frontal view of cephalothorax; H, O, frontal and anterolateral margin of carapace; N, ventral view of cephalothorax; J, right P 3; K, buccal cavity and posterior margin of epistome; Q, right P 4; R, right P 5; D, L, P, male pleon; S V, posterior margin of epistome; W Y, left thoracic sternites 7 and 8, “ supplementary plate ”, penis tube and penis. Abbreviations: cx 4 = coxa of P 4; cx 5 = coxa of P 5; p = penis; pt = calcified penis tube; sp = “ supplementary plate ”; st 7 = thoracic sternite 7; st 8 = thoracic sternite 8." httpUri="https://zenodo.org/record/272665/files/figure.png" pageId="90" pageNumber="91">Fig. 19</figureCitation>
E); the outer margins of the merus and ischium of the third maxilliped are lined with sharp teeth (
<figureCitation id="E1F956A0FFF9FFA1FEA1FA8EFE62FAC1" box="[348,431,1375,1400]" captionStart="FIGURE 34" captionStartId="107.[151,250,1457,1479]" captionTargetBox="[188,1388,200,1429]" captionTargetId="figure@107.[151,1436,193,1456]" captionTargetPageId="107" captionText="FIGURE 34. Third maxilliped of species of Megaesthesius and Alainthesius n. gen. A, Megaesthesius sagedae, Singapore: male 2.5 × 3.0 mm (ZRC 1995.885); B, Megaesthesius yokoyai, Japan: male 3.5 × 3.8 mm (after Takeda &amp; Miyake 1969: fig. 4 c); C, Megaethesius westralia, Australia: holotype male 3.4 × 3.8 mm (after Davie 2013: fig. 2 C); D, Megaesthesius migmus n. sp., Philippines: paratype female 4.6 × 5.2 mm (ZRC 2015.243); E, Alainthesius signatus n. sp., Madagascar: holotype male 2.5 × 3.3 mm (after Crosnier 1975: fig. 1 e); F, Alainthesius bertrandi n. sp., New Caledonia: holotype male 3.3 × 4.1 mm (MNHN-IU- 2013 - 9073). Scales = 0.5 mm." httpUri="https://zenodo.org/record/272680/files/figure.png" pageId="90" pageNumber="91">Fig. 34</figureCitation>
D) (unarmed in
<taxonomicName id="BEC231A6FFF9FFA1FD99FAB0FD2FFAC1" box="[612,738,1375,1400]" class="Malacostraca" family="Chasmocarcinidae" genus="Megaesthesius" kingdom="Animalia" order="Decapoda" pageId="90" pageNumber="91" phylum="Arthropoda" rank="species" species="yokoyai">
<emphasis id="4BB69637FFF9FFA1FD99FAB0FD2FFAC1" box="[612,738,1375,1400]" italics="true" pageId="90" pageNumber="91">M. yokoyai</emphasis>
</taxonomicName>
,
<figureCitation id="E1F956A0FFF9FFA1FD0CFA8EFC89FAC1" box="[753,836,1375,1400]" captionStart="FIGURE 34" captionStartId="107.[151,250,1457,1479]" captionTargetBox="[188,1388,200,1429]" captionTargetId="figure@107.[151,1436,193,1456]" captionTargetPageId="107" captionText="FIGURE 34. Third maxilliped of species of Megaesthesius and Alainthesius n. gen. A, Megaesthesius sagedae, Singapore: male 2.5 × 3.0 mm (ZRC 1995.885); B, Megaesthesius yokoyai, Japan: male 3.5 × 3.8 mm (after Takeda &amp; Miyake 1969: fig. 4 c); C, Megaethesius westralia, Australia: holotype male 3.4 × 3.8 mm (after Davie 2013: fig. 2 C); D, Megaesthesius migmus n. sp., Philippines: paratype female 4.6 × 5.2 mm (ZRC 2015.243); E, Alainthesius signatus n. sp., Madagascar: holotype male 2.5 × 3.3 mm (after Crosnier 1975: fig. 1 e); F, Alainthesius bertrandi n. sp., New Caledonia: holotype male 3.3 × 4.1 mm (MNHN-IU- 2013 - 9073). Scales = 0.5 mm." httpUri="https://zenodo.org/record/272680/files/figure.png" pageId="90" pageNumber="91">Fig. 34</figureCitation>
B); the fingers of the chela are longer and the ventral margin of the palm is lined with sharp teeth (
<figureCitation id="E1F956A0FFF9FFA1FD76FA55FD11FA25" box="[651,732,1412,1437]" captionStart="FIGURE 48" captionStartId="121.[151,250,1288,1310]" captionTargetBox="[151,1436,217,1267]" captionTargetId="figure@121.[151,1436,193,1267]" captionTargetPageId="121" captionText="FIGURE 48. Chelae of species of Megaesthesius, Alainthesius n. gen. and Microtopsis. A, B, Microtopsis takedai, Japan: holotype male 2.4 × 3.1 mm (after Komai et al. 2012: fig. 3 B, D); C, Megaesthesius sagedae, Singapore: male 2.5 × 3.0 mm (ZRC 1995.883); D, Megaesthesius migmus n. sp., Philippines: holotype male 4.4 × 4.9 mm (NMCR); E, Megaesthesius yokoyai, Japan: male (after Takeda &amp; Miyake 1969: fig. 4 d); F, Alainthesius signatus n. sp., Madagascar: holotype male 3.5 × 3.8 mm (after Crosnier 1975: fig. 1 d); G, Alainthesius bertrandi n. sp., New Caledonia: holotype male 3.3 × 4.1 mm (MNHN- IU- 2013 - 9073); H, Alainthesius bertrandi n. sp., Papua New Guinea: female 5.6 × 6.9 mm (MNHN-IU- 2013 - 9012)." httpUri="https://zenodo.org/record/272694/files/figure.png" pageId="90" pageNumber="91">Fig. 48</figureCitation>
D) (fingers short with the ventral margin of the palm smooth in
<taxonomicName id="BEC231A6FFF9FFA1FF6AFA78FEDBFA79" box="[151,278,1447,1472]" class="Malacostraca" family="Chasmocarcinidae" genus="Megaesthesius" kingdom="Animalia" order="Decapoda" pageId="90" pageNumber="91" phylum="Arthropoda" rank="species" species="yokoyai">
<emphasis id="4BB69637FFF9FFA1FF6AFA78FEDBFA79" box="[151,278,1447,1472]" italics="true" pageId="90" pageNumber="91">M. yokoyai</emphasis>
</taxonomicName>
,
<figureCitation id="E1F956A0FFF9FFA1FEDBFA76FEB7FA79" box="[294,378,1447,1472]" captionStart="FIGURE 48" captionStartId="121.[151,250,1288,1310]" captionTargetBox="[151,1436,217,1267]" captionTargetId="figure@121.[151,1436,193,1267]" captionTargetPageId="121" captionText="FIGURE 48. Chelae of species of Megaesthesius, Alainthesius n. gen. and Microtopsis. A, B, Microtopsis takedai, Japan: holotype male 2.4 × 3.1 mm (after Komai et al. 2012: fig. 3 B, D); C, Megaesthesius sagedae, Singapore: male 2.5 × 3.0 mm (ZRC 1995.883); D, Megaesthesius migmus n. sp., Philippines: holotype male 4.4 × 4.9 mm (NMCR); E, Megaesthesius yokoyai, Japan: male (after Takeda &amp; Miyake 1969: fig. 4 d); F, Alainthesius signatus n. sp., Madagascar: holotype male 3.5 × 3.8 mm (after Crosnier 1975: fig. 1 d); G, Alainthesius bertrandi n. sp., New Caledonia: holotype male 3.3 × 4.1 mm (MNHN- IU- 2013 - 9073); H, Alainthesius bertrandi n. sp., Papua New Guinea: female 5.6 × 6.9 mm (MNHN-IU- 2013 - 9012)." httpUri="https://zenodo.org/record/272694/files/figure.png" pageId="90" pageNumber="91">Fig. 48</figureCitation>
E); and the G1 is proportionately more slender with the distal part tapering and the opening large (
<figureCitation id="E1F956A0FFF9FFA1FF23FA1DFEE2FA5D" box="[222,303,1484,1509]" captionStart="FIGURE 82" captionStartId="155.[151,250,1823,1845]" captionTargetBox="[177,1417,213,1802]" captionTargetId="figure@155.[151,1436,193,1802]" captionTargetPageId="155" captionText="FIGURE 82. Gonopods of species of Megaesthesius. A C, M. sagedae, Singapore: male 2.5 × 3.0 mm (ZRC 1995.885); D F, M. migmus n. sp., Philippines: holotype male 4.4 × 4.9 mm (NMCR); G I, M. yokoyai, Japan: male 3.5 × 3.8 mm (after Takeda &amp; Miyake 1969: fig. 4 h, i); J M, M. westralia, Australia: holotype male 3.4 × 3.8 mm (after Davie 2013: fig. 3 A D). A, D, G, H, J, K, ventral view of left G 1; B, E, I, K, L, dorsal view of left G 1; C, F, M, left G 2. Scales = 0.1 mm." httpUri="https://zenodo.org/record/272728/files/figure.png" pageId="90" pageNumber="91">Fig. 82</figureCitation>
D, E) (G1 relatively stouter, with distal part not tapering and the opening rounded in
<taxonomicName id="BEC231A6FFF9FFA1FB18FA1CFAACFA5C" box="[1253,1377,1484,1509]" class="Malacostraca" family="Chasmocarcinidae" genus="Megaesthesius" kingdom="Animalia" order="Decapoda" pageId="90" pageNumber="91" phylum="Arthropoda" rank="species" species="yokoyai">
<emphasis id="4BB69637FFF9FFA1FB18FA1CFAACFA5C" box="[1253,1377,1484,1509]" italics="true" pageId="90" pageNumber="91">M. yokoyai</emphasis>
</taxonomicName>
,
<figureCitation id="E1F956A0FFF9FFA1FA92FA1DFF79F9B1" captionStart="FIGURE 82" captionStartId="155.[151,250,1823,1845]" captionTargetBox="[177,1417,213,1802]" captionTargetId="figure@155.[151,1436,193,1802]" captionTargetPageId="155" captionText="FIGURE 82. Gonopods of species of Megaesthesius. A C, M. sagedae, Singapore: male 2.5 × 3.0 mm (ZRC 1995.885); D F, M. migmus n. sp., Philippines: holotype male 4.4 × 4.9 mm (NMCR); G I, M. yokoyai, Japan: male 3.5 × 3.8 mm (after Takeda &amp; Miyake 1969: fig. 4 h, i); J M, M. westralia, Australia: holotype male 3.4 × 3.8 mm (after Davie 2013: fig. 3 A D). A, D, G, H, J, K, ventral view of left G 1; B, E, I, K, L, dorsal view of left G 1; C, F, M, left G 2. Scales = 0.1 mm." httpUri="https://zenodo.org/record/272728/files/figure.png" pageId="90" pageNumber="91">Fig. 82</figureCitation>
GI).
</paragraph>
</subSubSection>
<subSubSection id="31D819AEFFF9FFA1FF3AF9C5FC51F995" box="[199,924,1556,1581]" pageId="90" pageNumber="91" type="distribution">
<paragraph id="797D4A25FFF9FFA1FF3AF9C5FC51F995" blockId="90.[151,1437,151,1581]" box="[199,924,1556,1581]" pageId="90" pageNumber="91">
<emphasis id="4BB69637FFF9FFA1FF3AF9C5FE94F994" bold="true" box="[199,345,1556,1581]" pageId="90" pageNumber="91">Distribution</emphasis>
. Known only from
<collectingCountry id="01D50AB5FFF9FFA1FDCAF9C5FD74F994" box="[567,697,1556,1581]" name="Philippines" pageId="90" pageNumber="91">Philippines</collectingCountry>
. Depth:
<quantity id="BE3AE7C0FFF9FFA1FCEBF9C5FC55F995" box="[790,920,1556,1580]" metricMagnitude="2" metricUnit="m" metricValue="2.23" metricValueMax="3.18" metricValueMin="1.28" pageId="90" pageNumber="91" unit="m" value="223.0" valueMax="318.0" valueMin="128.0">
128
<specimenCount id="6FC481ACFFF9FFA1FCB0F9C5FC55F995" box="[845,920,1556,1580]" pageId="90" pageNumber="91" type="generic">318 m</specimenCount>
</quantity>
.
</paragraph>
</subSubSection>
</treatment>
</document>