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<document id="54F7D89C199553C4A42C62441F21EBDC" ID-CLB-Dataset="24063" ID-DOI="10.1080/00222933.2019.1692088" ID-GBIF-Dataset="e6443eb6-bfcc-44a6-a547-7af9a94d4a7d" ID-ISSN="1464-5262" ID-Zenodo-Dep="3665519" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1581529324984" checkinUser="valdenar" docAuthor="Nunes, Rafael V. &amp; Vaz-de-Mello, Fernando Z." docDate="2019" docId="C740D6092A27BB03FE9AE1D1E606292B" docLanguage="en" docName="J_NAT_HIST.53.3738.22312351.pdf.imf" docOrigin="Journal of Natural History 53 (37)" docStyle="DocumentStyle:485A7191CD41D6261F41521A3380DC73.9:JNatHist.2017-.journal_article.1cover" docStyleId="485A7191CD41D6261F41521A3380DC73" docStyleName="JNatHist.2017-.journal_article.1cover" docStyleVersion="9" docTitle="Dichotomius (Cephagonus) fernandosilvai Nunes &amp; Vaz-de-Mello 2019, new species" docType="treatment" docVersion="21" lastPageNumber="2300" masterDocId="3B79AE712A62BB45FFA2E57EE46C2C23" masterDocTitle="Taxonomic revision of Dichotomius (Cephagonus) Luederwaldt 1929 and the taxonomic status of remaining Dichotomius Hope 1838 subgenera (Coleoptera: Scarabaeidae: Scarabaeinae: Dichotomiini)" masterLastPageNumber="2351" masterPageNumber="2231" pageNumber="2299" updateTime="1698782751744" updateUser="ExternalLinkService">
<mods:mods id="2F3044CCC4890A15F10CF88BC6C49C79" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="4F3DFD0508A55B479EEFD6869DC0572E">Taxonomic revision of Dichotomius (Cephagonus) Luederwaldt 1929 and the taxonomic status of remaining Dichotomius Hope 1838 subgenera (Coleoptera: Scarabaeidae: Scarabaeinae: Dichotomiini)</mods:title>
</mods:titleInfo>
<mods:name id="CF33D470EB89636B570E19EEF33904A8" type="personal">
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<mods:namePart id="B7DFC4CD6BE8EFE50827B63C235E1D60">Nunes, Rafael V.</mods:namePart>
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<mods:namePart id="4A455E60DAA633FA2786DB3F57A46255">Vaz-de-Mello, Fernando Z.</mods:namePart>
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<mods:title id="480BA5EA661C60FCB2DB07A44CDF778E">Journal of Natural History</mods:title>
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<mods:date id="99317E65EAB73C7539B9089ADA7B5780">2019</mods:date>
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<mods:number id="05514D503C0BFE98C16BC92528F5152D">2020-01-13</mods:number>
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<treatment id="C740D6092A27BB03FE9AE1D1E606292B" ID-DOI="http://doi.org/10.5281/zenodo.3671896" ID-GBIF-Taxon="162116287" ID-Zenodo-Dep="3671896" LSID="urn:lsid:plazi:treatment:C740D6092A27BB03FE9AE1D1E606292B" httpUri="http://treatment.plazi.org/id/C740D6092A27BB03FE9AE1D1E606292B" lastPageId="70" lastPageNumber="2300" pageId="69" pageNumber="2299">
<subSubSection id="07F334942A27BB00FE9AE1D1E6BD28C8" pageId="69" pageNumber="2299" type="nomenclature">
<paragraph id="4F56671F2A27BB00FE9AE1D1E78728EB" blockId="69.[312,1003,1199,1259]" box="[312,1003,1199,1224]" pageId="69" pageNumber="2299">
<heading id="141ED0732A27BB00FE9AE1D1E78728EB" box="[312,1003,1199,1224]" pageId="69" pageNumber="2299">
2.4.:
<taxonomicName id="88E91C9C2A27BB00FECCE1D1E72728EB" ID-CoL="35N9W" authority="Nunes &amp; Vaz-de-Mello, 2019" authorityName="Nunes &amp; Vaz-de-Mello" authorityYear="2019" box="[366,843,1199,1224]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="69" pageNumber="2299" phylum="Arthropoda" rank="species" species="fernandosilvai" status="sp. nov." subGenus="Cephagonus">
<emphasis id="7D9DBB0D2A27BB00FECCE1D1E72728EB" box="[366,843,1199,1224]" italics="true" pageId="69" pageNumber="2299">Dichotomius (Cephagonus) fernandosilvai</emphasis>
</taxonomicName>
<emphasis id="7D9DBB0D2A27BB00FCF6E1D1E78728EB" bold="true" box="[852,1003,1199,1224]" pageId="69" pageNumber="2299">
<taxonomicNameLabel id="66AE06762A27BB00FCF6E1D1E78728EB" box="[852,1003,1199,1224]" pageId="69" pageNumber="2299" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="4F56671F2A27BB00FDF0E1ACE6BD28C8" blockId="69.[312,1003,1199,1259]" box="[594,721,1234,1259]" pageId="69" pageNumber="2299">
<heading id="141ED0732A27BB00FDF0E1ACE6BD28C8" box="[594,721,1234,1259]" centered="true" fontSize="11" level="2" pageId="69" pageNumber="2299" reason="5">
(
<figureCitation id="D7D27B9A2A27BB00FDF8E1ACE6A528C8" box="[602,713,1234,1259]" captionStart="Figure 23" captionStartId="38.[160,225,1583,1606]" captionTargetBox="[163,1153,142,1546]" captionTargetId="figure@38.[163,1153,142,1551]" captionTargetPageId="38" captionText="Figure 23. D. fernandosilvai: (a) male dorsal habitus (Holotype); (b) female dorsal habitus; (c) female ventral habitus; (d) paramera dorsal view; (e) paramera ventral view; (f) paramera lateral view." figureDoi="http://doi.org/10.5281/zenodo.3665561" httpUri="https://zenodo.org/record/3665561/files/figure.png" pageId="69" pageNumber="2299">Figure 23</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="07F334942A27BB00FF02E057E6122A74" pageId="69" pageNumber="2299" type="diagnosis">
<paragraph id="4F56671F2A27BB00FF02E057E6122A74" blockId="69.[160,1156,1321,1623]" pageId="69" pageNumber="2299">
<emphasis id="7D9DBB0D2A27BB00FF02E057E5762961" bold="true" box="[160,282,1321,1346]" italics="true" pageId="69" pageNumber="2299">Diagnosis.</emphasis>
distinguished from the other species in the group by the following combination of characteristics: males
<emphasis id="7D9DBB0D2A27BB00FE42E032E58A2947" box="[480,486,1356,1380]" italics="true" pageId="69" pageNumber="2299"></emphasis>
cephalic carina strongly transverseı larger than tallı slightly emarginated at apex; anterior portion of pronotum bearing coarse punctuation (as in
<taxonomicName id="88E91C9C2A27BB00FF02E0ECE5442989" authorityName="Nunes &amp; Vaz-de-Mello" authorityYear="2019" box="[160,296,1426,1450]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="69" pageNumber="2299" phylum="Arthropoda" rank="species" species="filgueirasi">
<emphasis id="7D9DBB0D2A27BB00FF02E0ECE5442989" box="[160,296,1426,1450]" italics="true" pageId="69" pageNumber="2299">D. filgueirasi</emphasis>
</taxonomicName>
of
<taxonomicName id="88E91C9C2A27BB00FEEDE0ECE5E52989" authorityName="Haroldi" authorityYear="1867" box="[335,393,1426,1450]" class="Insecta" family="Scarabaeidae" genus="Pinotus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="69" pageNumber="2299" phylum="Arthropoda" rank="species" species="fissus">
<emphasis id="7D9DBB0D2A27BB00FEEDE0ECE5E52989" box="[335,393,1426,1450]" italics="true" pageId="69" pageNumber="2299">fissus</emphasis>
</taxonomicName>
group); pronotal margin serrate (as in
<taxonomicName id="88E91C9C2A27BB00FCE5E0ECE7DE2989" authorityName="Nunes &amp; Vaz-de-Mello" authorityYear="2019" box="[839,946,1426,1450]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="69" pageNumber="2299" phylum="Arthropoda" rank="species" species="baiano">
<emphasis id="7D9DBB0D2A27BB00FCE5E0ECE7DE2989" box="[839,946,1426,1450]" italics="true" pageId="69" pageNumber="2299">D. baiano</emphasis>
</taxonomicName>
and
<taxonomicName id="88E91C9C2A27BB00FC4CE0ECE01A2989" authorityName="Nunes &amp; Vaz-de-Mello" authorityYear="2019" box="[1006,1142,1426,1450]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="69" pageNumber="2299" phylum="Arthropoda" rank="species" species="filgueirasi">
<emphasis id="7D9DBB0D2A27BB00FC4CE0ECE01A2989" box="[1006,1142,1426,1450]" italics="true" pageId="69" pageNumber="2299">D. filgueirasi</emphasis>
</taxonomicName>
); males
<emphasis id="7D9DBB0D2A27BB00FF43E0CAE48B29EF" box="[225,231,1460,1484]" italics="true" pageId="69" pageNumber="2299"></emphasis>
pronotal disc with a pair of central knobs at anterior portion; posterior margin of elytra with elliptical punctures (
<figureCitation id="D7D27B9A2A27BB00FDB8E0A9E6E729CC" box="[538,651,1495,1519]" captionStart="Figure 14" captionStartId="29.[160,225,1611,1634]" captionTargetBox="[163,1153,142,1554]" captionTargetId="figure@30.[163,1153,142,1554]" captionTargetPageId="30" captionText="Figure 14. Types of punctures and head/pronotal disc surfaces: (a) smooth surface with fine puncture; (b) ocellate punctures type I; (c) elliptical punctures; (d) ocellate punctures type II; (e) fine punctures (pronotum) and coarse punctures (head, indicated by white arrow); (f) clypeus and gena surfacewrinkled, wrinkled." figureDoi="http://doi.org/10.5281/zenodo.3814761" httpUri="https://zenodo.org/record/3814761/files/figure.png" pageId="69" pageNumber="2299">Figure 14</figureCitation>
(d)) and elytra lacking microsculpture. This species resembles smaller males of
<taxonomicName id="88E91C9C2A27BB00FDC1E084E6982A31" authorityName="Nunes &amp; Vaz-de-Mello" authorityYear="2019" box="[611,756,1530,1554]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="69" pageNumber="2299" phylum="Arthropoda" rank="species" species="filgueirasi">
<emphasis id="7D9DBB0D2A27BB00FDC1E084E6982A31" box="[611,756,1530,1554]" italics="true" pageId="69" pageNumber="2299">D. filgueirasi</emphasis>
</taxonomicName>
(also occurring in the state of
<collectingRegion id="8D2DA9FD2A27BB00FF02E363E5582A16" box="[160,308,1565,1589]" country="Brazil" name="Pernambuco" pageId="69" pageNumber="2299">Pernambuco</collectingRegion>
)ı differences are concentrated on the pronotal knobs (
<taxonomicName id="88E91C9C2A27BB00FC13E363E0562A16" authorityName="Nunes &amp; Vaz-de-Mello" authorityYear="2019" box="[945,1082,1564,1589]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="69" pageNumber="2299" phylum="Arthropoda" rank="species" species="filgueirasi">
<emphasis id="7D9DBB0D2A27BB00FC13E363E0562A16" box="[945,1082,1564,1589]" italics="true" pageId="69" pageNumber="2299">D. filgueirasi</emphasis>
</taxonomicName>
tends to lack it) and differentiation by paramera.
</paragraph>
</subSubSection>
<subSubSection id="07F334942A27BB03FF1CE3FBE7682C84" lastPageId="70" lastPageNumber="2300" pageId="69" pageNumber="2299" type="materials_examined">
<paragraph id="4F56671F2A27BB00FF1CE3FBE6962A9C" blockId="69.[190,1155,1669,1727]" pageId="69" pageNumber="2299">
<emphasis id="7D9DBB0D2A27BB00FF1CE3FBE4A02ABA" bold="true" box="[190,204,1669,1689]" pageId="69" pageNumber="2299"></emphasis>
<materialsCitation id="FF816D422A27BB00FF7AE3FBE6962A9C" ID-GBIF-Occurrence="2570430603" collectingDate="2008-06-12" collectionCode="CEMT" country="Brazil" location="Silva" municipality="Caruaru. Parque Ecologico Joao Vasconcelos Sobrinho. Pitfall" pageId="69" pageNumber="2299" specimenCount="1" specimenCount-male="1" stateProvince="Pernambuco" typeStatus="holotype">
<typeStatus id="9052D9BD2A27BB00FF7AE3FBE53B2ABE" box="[216,343,1669,1693]" pageId="69" pageNumber="2299" type="holotype">HOLOTYPEı</typeStatus>
<specimenCount id="59EFAC962A27BB00FEFFE3FBE5FA2ABE" box="[349,406,1669,1693]" pageId="69" pageNumber="2299" type="male">male</specimenCount>
:
<collectingCountry id="37FE278F2A27BB00FE3CE3FBE5822ABE" box="[414,494,1669,1693]" name="Brazil" pageId="69" pageNumber="2299">BRASIL</collectingCountry>
:
<emphasis id="7D9DBB0D2A27BB00FE54E3FBE6E12ABE" bold="true" box="[502,653,1669,1693]" pageId="69" pageNumber="2299">
<collectingRegion id="8D2DA9FD2A27BB00FE54E3FBE6E12ABE" box="[502,653,1669,1693]" country="Brazil" name="Pernambuco" pageId="69" pageNumber="2299">Pernambuco</collectingRegion>
</emphasis>
:
<collectingMunicipality id="AF32FD652A27BB00FD3AE3FBE5E02A9C" pageId="69" pageNumber="2299">Caruaru. Parque Ecológico João Vasconcelos Sobrinho. Pitfall</collectingMunicipality>
.
<date id="3B5741DF2A27BB00FE35E3D9E6632A9C" box="[407,527,1703,1727]" pageId="69" pageNumber="2299" value="2008-06-12">
<collectingDate id="2B13B8372A27BB00FE35E3D9E6632A9C" box="[407,527,1703,1727]" pageId="69" pageNumber="2299" value="2008-06-12">12-vi-2008</collectingDate>
</date>
. FAB
<location id="4A3631C42A27BB00FDEEE3D9E6132A9C" LSID="urn:lsid:plazi:treatment:C740D6092A27BB03FE9AE1D1E606292B:4A3631C42A27BB00FDEEE3D9E6132A9C" box="[588,639,1703,1727]" country="Brazil" municipality="Caruaru. Parque Ecologico Joao Vasconcelos Sobrinho. Pitfall" name="Silva" pageId="69" pageNumber="2299" stateProvince="Pernambuco">Silva</location>
[at
<collectionCode id="29F8FFDA2A27BB00FD0EE3D9E69D2A9C" box="[684,753,1703,1727]" pageId="69" pageNumber="2299">CEMT</collectionCode>
].
</materialsCitation>
</paragraph>
<paragraph id="4F56671F2A24BB03FF1CE5F1E7682C84" blockId="70.[190,772,143,167]" box="[190,772,143,167]" pageId="70" pageNumber="2300">
<emphasis id="7D9DBB0D2A24BB03FF1CE5F1E4A02C80" bold="true" box="[190,204,143,163]" pageId="70" pageNumber="2300"></emphasis>
<typeStatus id="9052D9BD2A24BB03FF7AE5F1E5232C84" box="[216,335,143,167]" pageId="70" pageNumber="2300" type="paratype">PARATYPE</typeStatus>
: same data of
<typeStatus id="9052D9BD2A24BB03FE51E5F1E6362C84" box="[499,602,143,167]" pageId="70" pageNumber="2300" type="holotype">Holotype</typeStatus>
[
<specimenCount id="59EFAC962A24BB03FDC8E5F1E6FE2C84" box="[618,658,143,167]" pageId="70" pageNumber="2300" type="female">1 ♀</specimenCount>
at
<collectionCode id="29F8FFDA2A24BB03FD14E5F1E6972C84" box="[694,763,143,167]" pageId="70" pageNumber="2300">CEMT</collectionCode>
].
</paragraph>
</subSubSection>
<subSubSection id="07F334942A24BB03FF02E5A1E6E12860" pageId="70" pageNumber="2300" type="description">
<paragraph id="4F56671F2A24BB03FF02E5A1E6612F00" blockId="70.[160,1156,223,803]" pageId="70" pageNumber="2300">
<emphasis id="7D9DBB0D2A24BB03FF02E5A1E5DF2CDB" bold="true" box="[160,435,223,248]" italics="true" pageId="70" pageNumber="2300">
Description (
<typeStatus id="9052D9BD2A24BB03FE9EE5A1E5C82CDB" box="[316,420,223,248]" pageId="70" pageNumber="2300" type="holotype">holotype</typeStatus>
).
</emphasis>
BL: 19 mmı PW:
<quantity id="8811CAFA2A24BB03FD29E59EE68D2CDB" box="[651,737,224,248]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="70" pageNumber="2300" unit="mm" value="12.0">12 mm</quantity>
. Colour: blackı shiny.
<emphasis id="7D9DBB0D2A24BB03FC47E59EE04F2CDB" bold="true" box="[997,1059,224,248]" pageId="70" pageNumber="2300">Head</emphasis>
: frontoclypeal surface smooth. Genal surface strongly wrinkled. Inter-ocular surface with coarse punctures. Gena borders straight. Cephalic carina larger than tallerı emarginated at apexı with striations at the base.
<emphasis id="7D9DBB0D2A24BB03FE52E436E6042D43" bold="true" box="[496,616,328,352]" pageId="70" pageNumber="2300">Pronotum</emphasis>
: borders serrate. Pronotal disc shinyı lacking punctures and with a central pair of knobs at anterior portion. Anterior excavation and anterior angles filled with coarse puncture. Posterior margin with two rows of elliptical punctures.
<emphasis id="7D9DBB0D2A24BB03FEBDE4CEE5D32DEB" bold="true" box="[287,447,432,456]" pageId="70" pageNumber="2300">Metasternum</emphasis>
: sides and anterior portion of anterior lobe covered by dense red setae. Setigerous punctures equally spacedı where present. Elytra: striae weakly impressed with weak punctures. Punctures separated by twice their diameter. Interstriae flatı lacking microsculpture.
<emphasis id="7D9DBB0D2A24BB03FDFCE766E6BF2E13" bold="true" box="[606,723,536,560]" pageId="70" pageNumber="2300">Abdomen</emphasis>
: glabrous and lacking distinct row of continuous punctuation along anterior margin of the ventrite.
<emphasis id="7D9DBB0D2A24BB03FCF9E745E7BD2E70" bold="true" box="[859,977,571,595]" pageId="70" pageNumber="2300">Aedeagus</emphasis>
(
<figureCitation id="D7D27B9A2A24BB03FC7DE745E0262E70" box="[991,1098,571,595]" captionStart="Figure 23" captionStartId="38.[160,225,1583,1606]" captionTargetBox="[163,1153,142,1546]" captionTargetId="figure@38.[163,1153,142,1551]" captionTargetPageId="38" captionText="Figure 23. D. fernandosilvai: (a) male dorsal habitus (Holotype); (b) female dorsal habitus; (c) female ventral habitus; (d) paramera dorsal view; (e) paramera ventral view; (f) paramera lateral view." figureDoi="http://doi.org/10.5281/zenodo.3665561" httpUri="https://zenodo.org/record/3665561/files/figure.png" pageId="70" pageNumber="2300">Figure 23</figureCitation>
(d-f)): parameraı apexes rounded dorsallyı convergent; longitudinal excavation taking only 1/5 of paramera length and restricted to apex; basal excavations weakı transversalı reaching the sides of the paramera. Laterallyı with a subtle declivity towards the apex; excavation near to the posterior border occupying half of paramera length towards the base. Ventrallyı a pair of wideı round excavations; subgenital plate rounded at the base and widely emarginated at the apex.
</paragraph>
<paragraph id="4F56671F2A24BB03FF02E625E6E12860" blockId="70.[160,1155,859,1091]" pageId="70" pageNumber="2300">
<emphasis id="7D9DBB0D2A24BB03FF02E625E5AA2F57" bold="true" box="[160,454,859,884]" italics="true" pageId="70" pageNumber="2300">Morphological variation.</emphasis>
female BL: 19 mmı PW:
<quantity id="8811CAFA2A24BB03FD41E625E75E2F50" box="[739,818,859,883]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" pageId="70" pageNumber="2300" unit="mm" value="11.0">11 mm</quantity>
and distinguishable from the male by the following: Head: fronto-clypeal surface transversely wrinkled. Cephalic carina transversalı forming a weak pair of tubercles. Pronotum: punctuation coarser than that of malesı posterior margin with two defined rows of elliptical punctures. Pronotal borders strongly serrate. Elytra: striae deeply impressedı punctures separated by twice their diameter. Abdomen: sixth ventrite bearing a pair of triangular projections reaching the apical margin of the pygidium (
<figureCitation id="D7D27B9A2A24BB03FDA7E155E60E2860" box="[517,610,1067,1091]" captionStart="Figure 9" captionStartId="22.[160,225,1548,1571]" captionTargetBox="[163,1153,142,1516]" captionTargetId="figure@22.[163,1153,142,1516]" captionTargetPageId="22" captionText="Figure 9. (a) Abdominal ventrites of a male of D. spadiceus. (b) ventrites of a female of D. spadiceus. (c) abdominal ventrites of a female of D. itatiaiaensisı (d) detail of C; (e) abdominal ventrites of a female of D. fissus. (f) Adbominal ventrites of a female of D. reichei. (g) Abdominal ventrites of a female of D. lucianomourai." figureDoi="http://doi.org/10.5281/zenodo.3665537" httpUri="https://zenodo.org/record/3665537/files/figure.png" pageId="70" pageNumber="2300">Figure 9</figureCitation>
(b)).
</paragraph>
</subSubSection>
<subSubSection id="07F334942A24BB03FF02E105E07C28B7" box="[160,1040,1147,1172]" pageId="70" pageNumber="2300" type="distribution">
<paragraph id="4F56671F2A24BB03FF02E105E07C28B7" blockId="70.[160,1040,1147,1172]" box="[160,1040,1147,1172]" pageId="70" pageNumber="2300">
<emphasis id="7D9DBB0D2A24BB03FF02E105E55F28B7" bold="true" box="[160,307,1147,1172]" italics="true" pageId="70" pageNumber="2300">Distribution.</emphasis>
municipality of Caruaruı state of Pernambucoı
<collectingCountry id="37FE278F2A24BB03FCEDE102E7E728B7" box="[847,907,1148,1172]" name="Brazil" pageId="70" pageNumber="2300">Brazil</collectingCountry>
(
<figureCitation id="D7D27B9A2A24BB03FC38E102E06F28B7" box="[922,1027,1148,1172]" captionStart="Figure 20" captionStartId="35.[160,225,1019,1042]" captionTargetBox="[163,1152,142,987]" captionTargetId="figure@35.[163,1153,142,988]" captionTargetPageId="35" captionText="Figure 20. Distribution map of the species in the D. fissus group." figureDoi="http://doi.org/10.5281/zenodo.3665555" httpUri="https://zenodo.org/record/3665555/files/figure.png" pageId="70" pageNumber="2300">Figure 20</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="07F334942A24BB03FF02E1B3E606292B" pageId="70" pageNumber="2300" type="etymology">
<paragraph id="4F56671F2A24BB03FF02E1B3E606292B" blockId="70.[160,1156,1229,1288]" pageId="70" pageNumber="2300">
<emphasis id="7D9DBB0D2A24BB03FF02E1B3E54F28C5" bold="true" box="[160,291,1229,1254]" italics="true" pageId="70" pageNumber="2300">Etymology.</emphasis>
named after Dr. Fernando Augusto Barbosa Silva (Universidade Federal do Paráı
<collectingCountry id="37FE278F2A24BB03FF7CE18EE572292B" box="[222,286,1264,1288]" name="Brazil" pageId="70" pageNumber="2300">Brasil</collectingCountry>
)ı who collected the
<typeStatus id="9052D9BD2A24BB03FE5DE18EE60A292B" box="[511,614,1264,1288]" pageId="70" pageNumber="2300" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
</treatment>
</document>