treatments-xml/data/C7/40/D6/C740D6092A0BBB2FFE9BE64CE6802845.xml
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<document id="DD4623FF213271A03D262ED4D730717D" ID-CLB-Dataset="24063" ID-DOI="10.1080/00222933.2019.1692088" ID-GBIF-Dataset="e6443eb6-bfcc-44a6-a547-7af9a94d4a7d" ID-ISSN="1464-5262" ID-Zenodo-Dep="3665519" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1581529324984" checkinUser="valdenar" docAuthor="Nunes, Rafael V. &amp; Vaz-de-Mello, Fernando Z." docDate="2019" docId="C740D6092A0BBB2FFE9BE64CE6802845" docLanguage="en" docName="J_NAT_HIST.53.3738.22312351.pdf.imf" docOrigin="Journal of Natural History 53 (37)" docStyle="DocumentStyle:485A7191CD41D6261F41521A3380DC73.9:JNatHist.2017-.journal_article.1cover" docStyleId="485A7191CD41D6261F41521A3380DC73" docStyleName="JNatHist.2017-.journal_article.1cover" docStyleVersion="9" docTitle="Dichotomius (Cephagonus) rondoniaensis Nunes &amp; Vaz-de-Mello 2019, new species" docType="treatment" docVersion="18" lastPageNumber="2336" masterDocId="3B79AE712A62BB45FFA2E57EE46C2C23" masterDocTitle="Taxonomic revision of Dichotomius (Cephagonus) Luederwaldt 1929 and the taxonomic status of remaining Dichotomius Hope 1838 subgenera (Coleoptera: Scarabaeidae: Scarabaeinae: Dichotomiini)" masterLastPageNumber="2351" masterPageNumber="2231" pageNumber="2335" updateTime="1698782751744" updateUser="ExternalLinkService">
<mods:mods id="DB1E953EDFB34CA7508842CB074DCD75" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="1022A593FBE55FD18EE245F593E1804D">Taxonomic revision of Dichotomius (Cephagonus) Luederwaldt 1929 and the taxonomic status of remaining Dichotomius Hope 1838 subgenera (Coleoptera: Scarabaeidae: Scarabaeinae: Dichotomiini)</mods:title>
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<mods:namePart id="DAA5147FF972FC87C6A0C23F2925F729">Nunes, Rafael V.</mods:namePart>
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<mods:namePart id="18DC12E03F3205744447E42668E30040">Vaz-de-Mello, Fernando Z.</mods:namePart>
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<mods:title id="BB2CEA00CBB40E00D61E4DF9841AACED">Journal of Natural History</mods:title>
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<mods:date id="FDAD6C50BB4FF57F91568FF4073EE65C">2019</mods:date>
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<mods:number id="C7A6CDC1AA455C7E77A9AEAE394132FF">2020-01-13</mods:number>
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<treatment id="C740D6092A0BBB2FFE9BE64CE6802845" ID-DOI="http://doi.org/10.5281/zenodo.3671881" ID-GBIF-Taxon="162116292" ID-Zenodo-Dep="3671881" LSID="urn:lsid:plazi:treatment:C740D6092A0BBB2FFE9BE64CE6802845" httpUri="http://treatment.plazi.org/id/C740D6092A0BBB2FFE9BE64CE6802845" lastPageId="106" lastPageNumber="2336" pageId="105" pageNumber="2335">
<subSubSection id="07F334942A0BBB2CFE9BE64CE6802F4D" pageId="105" pageNumber="2335" type="nomenclature">
<paragraph id="4F56671F2A0BBB2CFE9BE64CE7862F68" blockId="105.[313,1002,818,878]" box="[313,1002,818,843]" pageId="105" pageNumber="2335">
<heading id="141ED0732A0BBB2CFE9BE64CE7862F68" box="[313,1002,818,843]" pageId="105" pageNumber="2335">
6.1.:
<taxonomicName id="88E91C9C2A0BBB2CFECDE64CE7272F68" ID-CoL="6D43C" authority="Nunes &amp; Vaz-de-Mello, 2019" authorityName="Nunes &amp; Vaz-de-Mello" authorityYear="2019" box="[367,843,818,843]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="105" pageNumber="2335" phylum="Arthropoda" rank="species" species="rondoniaensis" status="sp. nov." subGenus="Cephagonus">
<emphasis id="7D9DBB0D2A0BBB2CFECDE64CE7272F68" box="[367,843,818,843]" italics="true" pageId="105" pageNumber="2335">Dichotomius (Cephagonus) rondoniaensis</emphasis>
</taxonomicName>
<emphasis id="7D9DBB0D2A0BBB2CFCF1E64CE7862F68" bold="true" box="[851,1002,818,843]" pageId="105" pageNumber="2335">
<taxonomicNameLabel id="66AE06762A0BBB2CFCF1E64CE7862F68" box="[851,1002,818,843]" pageId="105" pageNumber="2335" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="4F56671F2A0BBB2CFD95E62BE6802F4D" blockId="105.[313,1002,818,878]" box="[567,748,853,878]" pageId="105" pageNumber="2335">
<heading id="141ED0732A0BBB2CFD95E62BE6802F4D" box="[567,748,853,878]" centered="true" fontSize="11" level="2" pageId="105" pageNumber="2335" reason="5">
(
<figureCitation id="D7D27B9A2A0BBB2CFD9DE62BE6C32F4D" box="[575,687,853,878]" captionStart="Figure 47" captionStartId="62.[160,225,1562,1585]" captionTargetBox="[163,1151,142,1530]" captionTargetId="figure@62.[163,1153,142,1530]" captionTargetPageId="62" captionText="Figure 47. D. rondoniaensis: (a) male dorsal habitus; (b) paramera dorsal view; (c) paramera ventral view; (d) paramera lateral view. D. vargasae: (e) male dorsal habitus; (f) paramera dorsal view; (g) paramera ventral view; (h) paramera lateral view." figureDoi="http://doi.org/10.5281/zenodo.3666756" httpUri="https://zenodo.org/record/3666756/files/figure.png" pageId="105" pageNumber="2335">Figure 47</figureCitation>
(a-d))
</heading>
</paragraph>
</subSubSection>
<subSubSection id="07F334942A0BBB2CFF02E6D5E62D286C" pageId="105" pageNumber="2335" type="diagnosis">
<paragraph id="4F56671F2A0BBB2CFF02E6D5E62D286C" blockId="105.[160,1155,939,1103]" pageId="105" pageNumber="2335">
<emphasis id="7D9DBB0D2A0BBB2CFF02E6D5E5762FE7" bold="true" box="[160,282,939,964]" italics="true" pageId="105" pageNumber="2335">Diagnosis.</emphasis>
separated from other species in the
<taxonomicName id="88E91C9C2A0BBB2CFD79E6D2E70B2FE7" authorityName="Felsche" authorityYear="1901" box="[731,871,940,964]" class="Insecta" family="Scarabaeidae" genus="Pinotus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="105" pageNumber="2335" phylum="Arthropoda" rank="species" species="quadraticeps">
<emphasis id="7D9DBB0D2A0BBB2CFD79E6D2E70B2FE7" box="[731,871,940,964]" italics="true" pageId="105" pageNumber="2335">quadraticeps</emphasis>
</taxonomicName>
group by the following combined characters: larger male cephalic carina bifurcated at apex (as
<taxonomicName id="88E91C9C2A0BBB2CFC70E6B1E03C2FC4" box="[978,1104,975,999]" class="Insecta" family="Scarabaeidae" genus="Pinotus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="105" pageNumber="2335" phylum="Arthropoda" rank="species" species="fonsecae">
<emphasis id="7D9DBB0D2A0BBB2CFC70E6B1E03C2FC4" box="[978,1104,975,999]" italics="true" pageId="105" pageNumber="2335">D. fonsecae</emphasis>
</taxonomicName>
and
<taxonomicName id="88E91C9C2A0BBB2CFF02E68CE52C2829" authorityName="Nunes &amp; Vaz-de-Mello" authorityYear="2019" box="[160,320,1010,1034]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="105" pageNumber="2335" phylum="Arthropoda" rank="species" species="ricardosilvai">
<emphasis id="7D9DBB0D2A0BBB2CFF02E68CE52C2829" box="[160,320,1010,1034]" italics="true" pageId="105" pageNumber="2335">D. ricardosilvai</emphasis>
</taxonomicName>
); larger male pronotum with a central pair of rounded lobe on the anterior portion of the disc (as
<taxonomicName id="88E91C9C2A0BBB2CFE06E16AE665280F" authorityName="Harold" authorityYear="1869" box="[420,521,1044,1068]" class="Insecta" family="Scarabaeidae" genus="Pinotus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="105" pageNumber="2335" phylum="Arthropoda" rank="species" species="reichei">
<emphasis id="7D9DBB0D2A0BBB2CFE06E16AE665280F" box="[420,521,1044,1068]" italics="true" pageId="105" pageNumber="2335">D. reichei</emphasis>
</taxonomicName>
); female sixth ventrite having a central excavation with a central pair of knobs (
<figureCitation id="D7D27B9A2A0BBB2CFE0EE149E67B286C" box="[428,535,1079,1103]" captionStart="Figure 10" captionStartId="23.[160,225,1487,1510]" captionTargetBox="[163,1153,142,1456]" captionTargetId="figure@23.[163,1153,142,1456]" captionTargetPageId="23" captionText="Figure 10. (a) Abdominal ventrites of a male of D. vargasae. (b) Detail of a sixth ventrite (white arrow) of a female of D. vargasae. (c) Sixth ventrite of female of D. vargasae. (d) Abdominal ventrites of a female of D. rondoniaensis. Pyg pygidium." figureDoi="http://doi.org/10.5281/zenodo.3665539" httpUri="https://zenodo.org/record/3665539/files/figure.png" pageId="105" pageNumber="2335">Figure 10</figureCitation>
(d)).
</paragraph>
</subSubSection>
<subSubSection id="07F334942A0BBB2CFF1CE103E05F28DF" pageId="105" pageNumber="2335" type="materials_examined">
<paragraph id="4F56671F2A0BBB2CFF1CE103E74A2894" blockId="105.[190,1156,1148,1276]" pageId="105" pageNumber="2335">
<emphasis id="7D9DBB0D2A0BBB2CFF1CE103E4A028B2" bold="true" box="[190,204,1149,1169]" pageId="105" pageNumber="2335"></emphasis>
<materialsCitation id="FF816D422A0BBB2CFF7AE102E7722894" ID-GBIF-Occurrence="2570409567" collectingDate="2011-12-05" collectingDateMax="2011-12-07" collectingDateMin="2011-12-05" collectionCode="CEMT" country="Brazil" location="Pitfall Banana" municipality="Porto Velho" pageId="105" pageNumber="2335" specimenCount="1" specimenCount-male="1" stateProvince="Rondonia" typeStatus="holotype">
<typeStatus id="9052D9BD2A0BBB2CFF7AE102E53E28B7" box="[216,338,1148,1172]" pageId="105" pageNumber="2335" type="holotype">HOLOTYPE</typeStatus>
(
<specimenCount id="59EFAC962A0BBB2CFEFCE102E5F528B7" box="[350,409,1148,1172]" pageId="105" pageNumber="2335" type="male">male</specimenCount>
):
<collectingCountry id="37FE278F2A0BBB2CFE0BE102E59528B7" box="[425,505,1148,1172]" name="Brazil" pageId="105" pageNumber="2335">BRASIL</collectingCountry>
:
<emphasis id="7D9DBB0D2A0BBB2CFDA0E102E61928B7" bold="true" box="[514,629,1148,1172]" pageId="105" pageNumber="2335">
<collectingRegion id="8D2DA9FD2A0BBB2CFDA0E102E61928B7" box="[514,629,1148,1172]" country="Brazil" name="Rondonia" pageId="105" pageNumber="2335">Rondônia</collectingRegion>
</emphasis>
.
<collectingMunicipality id="AF32FD652A0BBB2CFD23E102E76B28B7" box="[641,775,1148,1172]" pageId="105" pageNumber="2335">Porto Velho</collectingMunicipality>
. ESEC-Cuniã
<geoCoordinate id="2ADD01D82A0BBB2CFC36E102E0EC28B7" box="[916,1152,1148,1172]" pageId="105" pageNumber="2335">
8³4
<emphasis id="7D9DBB0D2A0BBB2CFC1BE102E7D328B7" bold="true" box="[953,959,1148,1172]" italics="true" pageId="105" pageNumber="2335">ʹ</emphasis>
11
<emphasis id="7D9DBB0D2A0BBB2CFC78E102E78F28B7" box="[986,995,1148,1172]" italics="true" pageId="105" pageNumber="2335"></emphasis>
Sı 63³28
<emphasis id="7D9DBB0D2A0BBB2CFB9EE102E02E28B7" bold="true" box="[1084,1090,1148,1172]" italics="true" pageId="105" pageNumber="2335">ʹ</emphasis>
64
<emphasis id="7D9DBB0D2A0BBB2CFBFCE102E00B28B7" box="[1118,1127,1148,1172]" italics="true" pageId="105" pageNumber="2335"></emphasis>
W
</geoCoordinate>
.
<date id="3B5741DF2A0BBB2CFF7AE1E1E50D2894" box="[216,353,1183,1207]" pageId="105" pageNumber="2335" value="2011-12-05" valueMax="2011-12-07" valueMin="2011-12-05">
<collectingDate id="2B13B8372A0BBB2CFF7AE1E1E50D2894" box="[216,353,1183,1207]" pageId="105" pageNumber="2335" value="2011-12-05" valueMax="2011-12-07" valueMin="2011-12-05">5-7-XII-2011</collectingDate>
</date>
.
<location id="4A3631C42A0BBB2CFECEE1E1E66B2894" LSID="urn:lsid:plazi:treatment:C740D6092A0BBB2FFE9BE64CE6802845:4A3631C42A0BBB2CFECEE1E1E66B2894" box="[364,519,1183,1207]" country="Brazil" municipality="Porto Velho" name="Pitfall Banana" pageId="105" pageNumber="2335" stateProvince="Rondonia">Pitfall Banana</location>
. MAPA
<location id="4A3631C42A0BBB2CFDFFE1E1E6C12894" LSID="urn:lsid:plazi:treatment:C740D6092A0BBB2FFE9BE64CE6802845:4A3631C42A0BBB2CFDFFE1E1E6C12894" box="[605,685,1183,1207]" country="Brazil" municipality="Porto Velho" name="Silveira" pageId="105" pageNumber="2335" stateProvince="Rondonia">Silveira</location>
[at
<collectionCode id="29F8FFDA2A0BBB2CFD7BE1E1E7722894" box="[729,798,1183,1207]" pageId="105" pageNumber="2335">CEMT</collectionCode>
</materialsCitation>
].
</paragraph>
<paragraph id="4F56671F2A0BBB2CFF1CE1BCE05F28DF" blockId="105.[190,1156,1148,1276]" pageId="105" pageNumber="2335">
<emphasis id="7D9DBB0D2A0BBB2CFF1CE1BCE4A028F5" bold="true" box="[190,204,1218,1238]" pageId="105" pageNumber="2335"></emphasis>
<typeStatus id="9052D9BD2A0BBB2CFF7AE1BCE53028F9" box="[216,348,1218,1242]" pageId="105" pageNumber="2335" type="paratype">PARATYPES</typeStatus>
: same data of
<typeStatus id="9052D9BD2A0BBB2CFDABE1BCE61C28F9" box="[521,624,1218,1242]" pageId="105" pageNumber="2335" type="holotype">Holotype</typeStatus>
[4 ƋƋı
<specimenCount id="59EFAC962A0BBB2CFD72E1BCE76328F9" box="[720,783,1218,1242]" pageId="105" pageNumber="2335" type="female">4 ♀♀</specimenCount>
at
<collectionCode id="29F8FFDA2A0BBB2CFC9BE1BCE71228F9" box="[825,894,1218,1242]" pageId="105" pageNumber="2335">CEMT</collectionCode>
];
<collectingCountry id="37FE278F2A0BBB2CFC2DE1BCE7B328F9" box="[911,991,1218,1242]" name="Brazil" pageId="105" pageNumber="2335">BRASIL</collectingCountry>
.
<emphasis id="7D9DBB0D2A0BBB2CFC4EE1BCE04E28F9" bold="true" box="[1004,1058,1218,1242]" pageId="105" pageNumber="2335">Pará</emphasis>
. [
<specimenCount id="59EFAC962A0BBB2CFB98E1BCE00828F9" box="[1082,1124,1218,1242]" pageId="105" pageNumber="2335" type="female">1 ♀</specimenCount>
at
<collectionCode id="29F8FFDA2A0BBB2CFF7AE19AE57128DF" box="[216,285,1252,1276]" pageId="105" pageNumber="2335">CEMT</collectionCode>
];
<emphasis id="7D9DBB0D2A0BBB2CFE8EE19AE5CA28DF" bold="true" box="[300,422,1252,1276]" pageId="105" pageNumber="2335">
<collectingRegion id="8D2DA9FD2A0BBB2CFE8EE19AE5CA28DF" box="[300,422,1252,1276]" country="Brazil" name="Maranhao" pageId="105" pageNumber="2335">Maranhão</collectingRegion>
</emphasis>
. [
<specimenCount id="59EFAC962A0BBB2CFE19E19AE58F28DF" box="[443,483,1252,1276]" pageId="105" pageNumber="2335" type="female">1 ♀</specimenCount>
at
<collectionCode id="29F8FFDA2A0BBB2CFDAAE19AE62128DF" box="[520,589,1252,1276]" pageId="105" pageNumber="2335">CEMT</collectionCode>
].
<collectingCountry id="37FE278F2A0BBB2CFDFEE19AE6F628DF" box="[604,666,1252,1276]" name="Peru" pageId="105" pageNumber="2335">PERU</collectingCountry>
.
<emphasis id="7D9DBB0D2A0BBB2CFD01E19AE73E28DF" bold="true" box="[675,850,1252,1276]" pageId="105" pageNumber="2335">
<collectingRegion id="8D2DA9FD2A0BBB2CFD01E19AE73E28DF" box="[675,850,1252,1276]" country="Peru" name="Madre de Dios" pageId="105" pageNumber="2335">Madre de Dios</collectingRegion>
</emphasis>
[1 Ƌı
<specimenCount id="59EFAC962A0BBB2CFC35E19AE7AC28DF" box="[919,960,1252,1276]" pageId="105" pageNumber="2335" type="female">1 ♀</specimenCount>
at
<collectionCode id="29F8FFDA2A0BBB2CFC47E19AE04628DF" box="[997,1066,1252,1276]" pageId="105" pageNumber="2335">CEMT</collectionCode>
].
</paragraph>
</subSubSection>
<subSubSection id="07F334942A0BBB2FFF02E057E5812E97" lastPageId="106" lastPageNumber="2336" pageId="105" pageNumber="2335" type="description">
<paragraph id="4F56671F2A0BBB2FFF02E057E63E2D77" blockId="105.[160,1156,1321,1727]" lastBlockId="106.[160,1155,142,340]" lastPageId="106" lastPageNumber="2336" pageId="105" pageNumber="2335">
<emphasis id="7D9DBB0D2A0BBB2CFF02E057E59B2961" bold="true" box="[160,503,1321,1346]" italics="true" pageId="105" pageNumber="2335">Description (Holotypeı male).</emphasis>
BL: 22 mmı PW:
<quantity id="8811CAFA2A0BBB2CFD1AE054E7642961" box="[696,776,1322,1346]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="105" pageNumber="2335" unit="mm" value="12.0">12 mm</quantity>
. Colour: blackı shiny.
<emphasis id="7D9DBB0D2A0BBB2CFC57E054E05F2961" bold="true" box="[1013,1075,1322,1346]" pageId="105" pageNumber="2335">Head</emphasis>
: dorsal surface smooth. Clypeo-genal angle obtuse (100³). Cephalic carina longer than wideı bifurcated at apex producing a tubercle on each end.
<emphasis id="7D9DBB0D2A0BBB2CFCACE011E7EA29A4" bold="true" box="[782,902,1391,1415]" pageId="105" pageNumber="2335">Pronotum</emphasis>
: anterior portion with fine punctures evenly spaced; group of ocellate punctures present beneath mid-lateral foveae; posterior margin with 1
<emphasis id="7D9DBB0D2A0BBB2CFE5EE0CAE66529EF" box="[508,521,1460,1484]" italics="true" pageId="105" pageNumber="2335"></emphasis>
2 rows of ocellate punctures. Disc smoothı shinyı anterior portion with four tubercles aligned transversely.
<emphasis id="7D9DBB0D2A0BBB2CFD7EE0A9E71029CC" bold="true" box="[732,892,1495,1519]" pageId="105" pageNumber="2335">Metasternum</emphasis>
: sides and margins of anterior lobe covered by setigerous ocellate punctures. Middle portion of anterior lobe glabrous and lacking punctures. Setigerous punctures equally spaced at sides.
<emphasis id="7D9DBB0D2A0BBB2CFB9AE363E0112A16" bold="true" box="[1080,1149,1565,1589]" pageId="105" pageNumber="2335">Elytra</emphasis>
: striae deeply impressed having fine punctures spaced by 1-three times their diameter. Interstriae convexı disc with dark brown colourı lacking microsculpture with very fine punctures (20 x).
<emphasis id="7D9DBB0D2A0BBB2CFECBE3FBE5B22ABE" bold="true" box="[361,478,1669,1693]" pageId="105" pageNumber="2335">Abdomen</emphasis>
: sides of each ventrite covered by ocellate punctures with very short setae (35x) and excavated near anterior margin. Ventrite 1 having 1
<emphasis id="7D9DBB0D2A0BBB2CFBB5E3D9E0482A9C" box="[1047,1060,1703,1727]" italics="true" pageId="105" pageNumber="2335"></emphasis>
2 longer setae on each side. Puncture of each ventrite interrupted medially as well as groove along the posterior margin of sixth ventrite.
<emphasis id="7D9DBB0D2A08BB2FFDCBE5CFE6B32CEA" bold="true" box="[617,735,177,201]" pageId="106" pageNumber="2336">Aedeagus</emphasis>
(
<figureCitation id="D7D27B9A2A08BB2FFD51E5CFE7082CEA" box="[755,868,177,201]" captionStart="Figure 47" captionStartId="62.[160,225,1562,1585]" captionTargetBox="[163,1151,142,1530]" captionTargetId="figure@62.[163,1153,142,1530]" captionTargetPageId="62" captionText="Figure 47. D. rondoniaensis: (a) male dorsal habitus; (b) paramera dorsal view; (c) paramera ventral view; (d) paramera lateral view. D. vargasae: (e) male dorsal habitus; (f) paramera dorsal view; (g) paramera ventral view; (h) paramera lateral view." figureDoi="http://doi.org/10.5281/zenodo.3666756" httpUri="https://zenodo.org/record/3666756/files/figure.png" pageId="106" pageNumber="2336">Figure 47</figureCitation>
(b-d)): dorsallyı paramere apexes roundedı convergent; longitudinal excavation present taking whole paramera length; basal-rounded excavation present. Laterallyı abrupt declivity from basis towards apex. Ventrallyı each paramera with a lateral-rounded excavation; subgenital plate widely emarginated basallyı rounded at apex.
</paragraph>
<paragraph id="4F56671F2A08BB2FFF02E4F8E5812E97" blockId="106.[160,1156,390,692]" pageId="106" pageNumber="2336">
<emphasis id="7D9DBB0D2A08BB2FFF02E4F8E5A92DBC" bold="true" box="[160,453,390,415]" italics="true" pageId="106" pageNumber="2336">Morphological variation.</emphasis>
male BL: 19 to 22 mmı PW:
<quantity id="8811CAFA2A08BB2FFCA7E4F9E7122DBC" box="[773,894,391,415]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" metricValueMax="1.2" metricValueMin="1.1" pageId="106" pageNumber="2336" unit="mm" value="11.5" valueMax="12.0" valueMin="11.0">
11
<emphasis id="7D9DBB0D2A08BB2FFC82E4F9E7412DBC" box="[800,813,391,415]" italics="true" pageId="106" pageNumber="2336"></emphasis>
12 mm
</quantity>
. Female BL: 18
<emphasis id="7D9DBB0D2A08BB2FFB80E4F9E0432DBC" box="[1058,1071,391,415]" italics="true" pageId="106" pageNumber="2336"></emphasis>
22 mmı PW: 11
<emphasis id="7D9DBB0D2A08BB2FFF4EE4D7E4952DE2" box="[236,249,425,449]" italics="true" pageId="106" pageNumber="2336"></emphasis>
13 mmı distinguished from males as follows: Head: fronto-clypeal surface striated. Middle portion of interocular surface with chagrinated microsculptureı coarse puncture present near eyes. Cephalic carina not bifurcated apically. Pronotum: simply convexı lacking lobes. Posterior margin with strong ocellate punctures. Coarse puncture beneath mid-lateral pit. Abdomen: female sixth ventrite having a single shape in
<emphasis id="7D9DBB0D2A08BB2FFC55E74AE0122E6F" box="[1015,1150,564,588]" italics="true" pageId="106" pageNumber="2336">selenocopris</emphasis>
: composed by a pair of central tubercles that do not advance beneath pygidium apexı lateral to each tubercleı a triangular projection of the posterior margin advances beneath pygidium apex (
<figureCitation id="D7D27B9A2A08BB2FFEF5E7E2E5AF2E97" box="[343,451,668,692]" captionStart="Figure 10" captionStartId="23.[160,225,1487,1510]" captionTargetBox="[163,1153,142,1456]" captionTargetId="figure@23.[163,1153,142,1456]" captionTargetPageId="23" captionText="Figure 10. (a) Abdominal ventrites of a male of D. vargasae. (b) Detail of a sixth ventrite (white arrow) of a female of D. vargasae. (c) Sixth ventrite of female of D. vargasae. (d) Abdominal ventrites of a female of D. rondoniaensis. Pyg pygidium." figureDoi="http://doi.org/10.5281/zenodo.3665539" httpUri="https://zenodo.org/record/3665539/files/figure.png" pageId="106" pageNumber="2336">Figure 10</figureCitation>
(d)).
</paragraph>
</subSubSection>
<subSubSection id="07F334942A08BB2FFF02E798E7792EDC" box="[160,789,742,767]" pageId="106" pageNumber="2336" type="distribution">
<paragraph id="4F56671F2A08BB2FFF02E798E7792EDC" blockId="106.[160,789,742,767]" box="[160,789,742,767]" pageId="106" pageNumber="2336">
<emphasis id="7D9DBB0D2A08BB2FFF02E798E55F2EDC" bold="true" box="[160,307,742,767]" italics="true" pageId="106" pageNumber="2336">Distribution.</emphasis>
<collectingCountry id="37FE278F2A08BB2FFEE1E799E5132EDC" box="[323,383,743,767]" name="Brazil" pageId="106" pageNumber="2336">Brazil</collectingCountry>
(PAı MAı RO)ı
<collectingCountry id="37FE278F2A08BB2FFD83E799E6382EDC" box="[545,596,743,767]" name="Peru" pageId="106" pageNumber="2336">Peru</collectingCountry>
(
<collectingRegion id="8D2DA9FD2A08BB2FFDC0E799E7672EDC" box="[610,779,743,767]" country="Peru" name="Madre de Dios" pageId="106" pageNumber="2336">Madre de Dios</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="07F334942A08BB2FFF02E64CE7722FDB" pageId="106" pageNumber="2336" type="discussion">
<paragraph id="4F56671F2A08BB2FFF02E64CE7722FDB" blockId="106.[160,1156,818,1016]" pageId="106" pageNumber="2336">
<emphasis id="7D9DBB0D2A08BB2FFF02E64CE5FD2F68" bold="true" box="[160,401,818,843]" italics="true" pageId="106" pageNumber="2336">Systematic remarks.</emphasis>
males of this species are similar to those of
<taxonomicName id="88E91C9C2A08BB2FFC01E64DE0432F68" authorityName="Felsche" authorityYear="1901" box="[931,1071,819,843]" class="Insecta" family="Scarabaeidae" genus="Pinotus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="106" pageNumber="2336" phylum="Arthropoda" rank="species" species="quadraticeps">
<emphasis id="7D9DBB0D2A08BB2FFC01E64DE0432F68" box="[931,1071,819,843]" italics="true" pageId="106" pageNumber="2336">quadraticeps</emphasis>
</taxonomicName>
group; howeverı females have a different 6th abdominal sternite among
<taxonomicName id="88E91C9C2A08BB2FFC61E62BE0262F4E" box="[963,1098,853,877]" class="Insecta" family="Scarabaeidae" genus="Pinotus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="106" pageNumber="2345" phylum="Arthropoda" rank="subGenus" subGenus="Cephagonus">
<emphasis id="7D9DBB0D2A08BB2FFC61E62BE0262F4E" box="[963,1098,853,877]" italics="true" pageId="106" pageNumber="2336">Cephagonus</emphasis>
</taxonomicName>
and
<taxonomicName id="88E91C9C2A08BB2FFF02E606E5452FB3" authorityName="Hope" authorityYear="1838" box="[160,297,888,912]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="106" pageNumber="2336" phylum="Arthropoda" rank="genus">
<emphasis id="7D9DBB0D2A08BB2FFF02E606E5452FB3" box="[160,297,888,912]" italics="true" pageId="106" pageNumber="2336">Dichotomius</emphasis>
</taxonomicName>
(
<figureCitation id="D7D27B9A2A08BB2FFE98E606E5CB2FB3" box="[314,423,888,912]" captionStart="Figure 10" captionStartId="23.[160,225,1487,1510]" captionTargetBox="[163,1153,142,1456]" captionTargetId="figure@23.[163,1153,142,1456]" captionTargetPageId="23" captionText="Figure 10. (a) Abdominal ventrites of a male of D. vargasae. (b) Detail of a sixth ventrite (white arrow) of a female of D. vargasae. (c) Sixth ventrite of female of D. vargasae. (d) Abdominal ventrites of a female of D. rondoniaensis. Pyg pygidium." figureDoi="http://doi.org/10.5281/zenodo.3665539" httpUri="https://zenodo.org/record/3665539/files/figure.png" pageId="106" pageNumber="2336">Figure 10</figureCitation>
(d)). This species has a wide distribution however two localitiesı in
<collectingRegion id="8D2DA9FD2A08BB2FFF1EE6E5E5412F90" box="[188,301,923,947]" country="Brazil" name="Maranhao" pageId="106" pageNumber="2336">Maranhão</collectingRegion>
and Paráı are represented by very similar females. We do not know what to expectı but despite the very localised endemisms found in
<taxonomicName id="88E91C9C2A08BB2FFC8DE6C3E7DA2FF6" box="[815,950,957,981]" class="Insecta" family="Scarabaeidae" genus="Pinotus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="106" pageNumber="2345" phylum="Arthropoda" rank="subGenus" subGenus="Cephagonus">
<emphasis id="7D9DBB0D2A08BB2FFC8DE6C3E7DA2FF6" box="[815,950,957,981]" italics="true" pageId="106" pageNumber="2336">Cephagonus</emphasis>
</taxonomicName>
ı some species like
<taxonomicName id="88E91C9C2A08BB2FFF02E69EE5662FDB" authorityName="Nunes &amp; Vaz-de-Mello" authorityYear="2019" box="[160,266,992,1016]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="106" pageNumber="2336" phylum="Arthropoda" rank="species" species="asenjoi">
<emphasis id="7D9DBB0D2A08BB2FFF02E69EE5662FDB" box="[160,266,992,1016]" italics="true" pageId="106" pageNumber="2336">D. asenjoi</emphasis>
</taxonomicName>
have also have a wide and similar distribution.
</paragraph>
</subSubSection>
<subSubSection id="07F334942A08BB2FFF02E154E6802845" pageId="106" pageNumber="2336" type="etymology">
<paragraph id="4F56671F2A08BB2FFF02E154E6802845" blockId="106.[160,1155,1066,1126]" pageId="106" pageNumber="2336">
<emphasis id="7D9DBB0D2A08BB2FFF02E154E54F2860" bold="true" box="[160,291,1066,1091]" italics="true" pageId="106" pageNumber="2336">Etymology.</emphasis>
specific name refers to the state of Rondôniaı
<collectingCountry id="37FE278F2A08BB2FFCFAE155E7FF2860" box="[856,915,1067,1091]" name="Brazil" pageId="106" pageNumber="2336">Brasil</collectingCountry>
while the state was named after Marshall Cândido Rondon (
<date id="3B5741DF2A08BB2FFDC3E130E68E2845" box="[609,738,1102,1126]" pageId="106" pageNumber="2336" value="1865" valueMax="1958">
1865
<emphasis id="7D9DBB0D2A08BB2FFD38E130E6CB2845" box="[666,679,1102,1126]" italics="true" pageId="106" pageNumber="2336"></emphasis>
1958
</date>
).
</paragraph>
</subSubSection>
</treatment>
</document>