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<document id="6E880ADDEA761C7831D0460ED73D0608" ID-DOI="10.11646/zootaxa.3889.1.3" ID-GBIF-Dataset="7af6165c-b6fb-4235-be95-7addbfc8e653" ID-ISSN="1175-5326" ID-Zenodo-Dep="225443" ID-ZooBank="6D07A7C6-C941-4093-95C8-19E79D639BB4" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460987823067" checkinUser="plazi" docAuthor="Coelho, Beatriz W. T., De Seixas Felizardo, Sherlem P. &amp; Engel, Michael S." docDate="2014" docId="A81ECC6A0377FFA8FF03FA6BB0917A3F" docLanguage="en" docName="zt03889p070.pdf" docOrigin="Zootaxa 3889 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ruizantheda inca Coelho, Felizardo &amp; Engel, 2014, n. sp." docType="treatment" docUuid="D631BC6B-E783-49C7-B9C7-0243605A8707" docUuidSource="ZooBank" docVersion="10" masterDocId="5427B412037DFFA3FF94FFEFB3047C4A" masterDocTitle="Three new species of the bee genus Ruizantheda sensu lato (Hymenoptera: Halictidae: Caenohalictina)" originalUpdateDomain="Boston" originalUpdateTime="1545214577784" originalUpdateUser="gglab" updateTime="1698560485211" updateUser="plazi">
<mods:mods id="579B367ACAA2D2C49E0883FA6AC66E23" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="25D13E38D03E1A351B35A364B551A95D">
<mods:title id="2A68F293B193E475766F3F679468F358">Three new species of the bee genus Ruizantheda sensu lato (Hymenoptera: Halictidae: Caenohalictina)</mods:title>
</mods:titleInfo>
<mods:name id="96F0445DF68415D54070E879DC3B7FA2" type="personal">
<mods:role id="45162EE6ED627F39A2B890FF1D1C05D8">
<mods:roleTerm id="17D0F800314D3A98F41A313F17EC8B80">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="75D578685D33028561449D6BF215D945">Coelho, Beatriz W. T.</mods:namePart>
</mods:name>
<mods:name id="B08111952B97F96944FC5ECDF2E1695B" type="personal">
<mods:role id="A594EFF350A2796FBB30BF4F3554C513">
<mods:roleTerm id="8D7E3AFDD13CF24891131CEC42DA380D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B314DED077168E3BF3D508C3FC66777C">De Seixas Felizardo, Sherlem P.</mods:namePart>
</mods:name>
<mods:name id="C4E0AD1C46F7543D26CBABCE90E35C60" type="personal">
<mods:role id="8B7AAD978BACC0EF18FC698480FD17C3">
<mods:roleTerm id="1387984E1D705CCFE970BEE5D9B7584A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C5559E7B0AADA9ED48C5EAB5E054B0B0">Engel, Michael S.</mods:namePart>
</mods:name>
<mods:typeOfResource id="7738D8C2AC64DFFB2DCD01DF63712EF4">text</mods:typeOfResource>
<mods:relatedItem id="4312EBF47DA551898249B152C6717C58" type="host">
<mods:titleInfo id="CAAFB7BBC2E001B4440514C6A58F0E7B">
<mods:title id="A1E72CD32B00E5B115E3B840A232EA22">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="C8A2BAB0198D89E4639B104380553880">
<mods:date id="7A361EB5364956860A3973033F337815">2014</mods:date>
<mods:detail id="1ABEC15734117BEFEEB0E30A40736271" type="volume">
<mods:number id="B65D37DE7DEDA3100AF3CA1CE287B86B">3889</mods:number>
</mods:detail>
<mods:detail id="5991F38D9E8ADAC0417A19612C0D06A2" type="issue">
<mods:number id="84FEA173C3A4DEDCB627220147471C60">1</mods:number>
</mods:detail>
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<mods:classification id="226B4026B410C25582E3DFD296125F1E">journal volume</mods:classification>
<mods:identifier id="ABBC7B20EC5E39B85D795DC117EC5D52" type="DOI">10.11646/zootaxa.3889.1.3</mods:identifier>
<mods:identifier id="A8376D9F5B593FD02B6C5202B4E40345" type="GBIF-Dataset">7af6165c-b6fb-4235-be95-7addbfc8e653</mods:identifier>
<mods:identifier id="3F66891764CE925E8FF00E4EC18D9D15" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="6442FDB76667910F373166A5B9586127" type="Zenodo-Dep">225443</mods:identifier>
<mods:identifier id="06A0AD7EF7845D2F01813F62BDDF969C" type="ZooBank">6D07A7C6-C941-4093-95C8-19E79D639BB4</mods:identifier>
</mods:mods>
<treatment id="A81ECC6A0377FFA8FF03FA6BB0917A3F" ID-DOI="http://doi.org/10.5281/zenodo.6125102" ID-GBIF-Taxon="119594930" ID-Zenodo-Dep="6125102" LSID="urn:lsid:zoobank.org:act:D631BC6B-E783-49C7-B9C7-0243605A8707" httpUri="http://treatment.plazi.org/id/A81ECC6A0377FFA8FF03FA6BB0917A3F" lastPageId="11" pageId="10">
<subSubSection id="68AD2EF70377FFA9FF03FA6BB1EB798A" pageId="10" type="nomenclature">
<paragraph id="20087D7C0377FFA9FF03FA6BB2B179D4" blockId="10.[151,751,1412,1472]" box="[151,437,1412,1438]" pageId="10">
<heading id="7B40CA100377FFA9FF03FA6BB2B179D4" bold="true" box="[151,437,1412,1438]" fontSize="11" level="1" pageId="10" reason="1">
<emphasis id="12C3A16E0377FFA9FF03FA6BB2B179D4" bold="true" box="[151,437,1412,1438]" pageId="10">
<taxonomicName id="E7B706FF0377FFA9FF03FA6BB26F79D4" box="[151,363,1412,1438]" class="Insecta" family="Halictidae" genus="Ruizantheda" kingdom="Animalia" order="Hymenoptera" pageId="10" phylum="Arthropoda" rank="species" species="inca" status="sp. nov.">
<emphasis id="12C3A16E0377FFA9FF03FA6BB26F79D4" bold="true" box="[151,363,1412,1438]" italics="true" pageId="10">Ruizantheda inca</emphasis>
</taxonomicName>
<taxonomicNameLabel id="09F01C150377FFA9FEE7FA6BB2B179D4" box="[371,437,1412,1438]" pageId="10" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="20087D7C0377FFA9FF03FA48B1EB798A" blockId="10.[151,751,1412,1472]" box="[151,751,1447,1472]" pageId="10">
(
<figureCitation id="B88C61F90377FFA9FF0AFA48B247798A" box="[158,323,1447,1472]" captionStart="FIGURES 10 13" captionStartId="7.[151,264,1179,1202]" captionTargetBox="[152,1436,193,1157]" captionTargetId="figure@7.[151,1436,193,1158]" captionTargetPageId="7" captionText="FIGURES 10 13. Male. 10 12, habit of males, lateral view; 13, labrum, frontal view. 10, Ruizantheda kallos n. sp.; 11, R. aerugineus n. sp.; 12 13, R. inca n. sp." httpUri="https://zenodo.org/record/225446/files/figure.png" pageId="10">Figures 12, 13</figureCitation>
, 16, 19, 22, 25, 28, 31, 34, 37, 39, 40)
</paragraph>
</subSubSection>
<subSubSection id="68AD2EF70377FFA9FF03FA00B7147AD2" pageId="10" type="diagnosis">
<paragraph id="20087D7C0377FFA9FF03FA00B7147AD2" blockId="10.[151,1436,1519,2012]" pageId="10">
<emphasis id="12C3A16E0377FFA9FF03FA00B20E7A42" bold="true" box="[151,266,1519,1544]" pageId="10">Diagnosis</emphasis>
. Male: head longer than wide; malar area as long as 1/3 of mandibular width; clypeus projecting about 2/ 3 of its total length below the lower tangent of compound eyes; surface between compound eye and lateral ocelli slightly concave; vertex slightly expanded behind ocelli; metapostnotum with dense granulation and one unique, coarse carina medially; metasomal terga amber with metallic green bands; T1T4 with fine punctures, separated by 12 puncture widths in central area, surface between punctures microreticulate.
</paragraph>
</subSubSection>
<subSubSection id="68AD2EF70377FFA8FF53F94CB2E3798A" lastPageId="11" pageId="10" type="description">
<paragraph id="20087D7C0377FFA8FF53F94CB2E3798A" blockId="10.[151,1436,1519,2012]" lastBlockId="11.[151,1436,151,1653]" lastPageId="11" pageId="10">
<emphasis id="12C3A16E0377FFA9FF53F94CB25D7AF6" bold="true" box="[199,345,1699,1724]" pageId="10">Description.</emphasis>
Male.
<emphasis id="12C3A16E0377FFA9FE33F94BB2D07AF1" box="[423,468,1700,1723]" italics="true" pageId="10">Size</emphasis>
. Total body length
<quantity id="E74FD0990377FFA9FD3AF94BB1F67AF1" box="[686,754,1700,1723]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="10" unit="mm" value="7.0">7 mm</quantity>
; forewing length 1.471.48 mm.
<emphasis id="12C3A16E0377FFA9FBF3F94BB7C87AF1" box="[1127,1228,1700,1723]" italics="true" pageId="10">Structure</emphasis>
. Head longer than wide, head length to width ratio 1.06; malar area as long as 1/3 of mandibular width; mandible simple; labral distal process in the form of a minute inverted triangle; epistomal sulcus obtuse and slightly rounded; clypeus projecting about 2/3 of its length below lower tangent of compound eyes; compound eyes emarginated above level of antennal torulus; surface between compound eye and lateral ocellus slightly concave; vertex slightly expanded behind ocelli; preoccipital ridge rounded; antennal flagellum weakly sinuate, F2 slightly more than
<date id="54095BBC0377FFA9FB4EF8B7B7F27B3A" box="[1242,1270,1880,1904]" pageId="10">2x</date>
length of F1; pronotal lateral angle obtuse; anterior border of mesoscutum rounded; metapostnotum about 1.3x length of metanotum medially; legs unmodified; metabasitibial plate rounded, margin weakly defined, flat; S4 and S5 unmodified; gradulus of S6 greatly invaginated medially, covered by membrane, pygidial plate large with widely rounded margin; S7 with median apical process short, without setae; median apical process of S8 broad with apical margin bearing lateral peaks, without setae, separated by weak concavity, but apical edge of S8 with lateral lobes extending ventrally separated by a thin and slightly elongate medial surface in posterior view.
<emphasis id="12C3A16E0376FFA8FB38FF0FB6917CB2" box="[1196,1429,224,248]" italics="true" pageId="11">Genitalia. Gonobase</emphasis>
. Length nearly 1/3 that of gonocoxite.
<emphasis id="12C3A16E0376FFA8FDDFFEEBB1D77D51" box="[587,723,260,283]" italics="true" pageId="11">Gonocoxite.</emphasis>
Outer margins slightly convex apically, surface smooth; basal portion of inner dorsal margins parallel with large and rectangular excavated area; apical portion of inner dorsal margins strongly concave.
<emphasis id="12C3A16E0376FFA8FE50FEA3B1467D2E" box="[452,578,332,356]" italics="true" pageId="11">Gonostylus</emphasis>
. Basal region with
<emphasis id="12C3A16E0376FFA8FC8AFEA3B0447D2F" bold="true" box="[798,832,332,357]" pageId="11">rvl</emphasis>
membranous, short, about ½ of length of gonocoxite, rounded apically, with a few short setae at extremity and band of short setae on inner margin of dorsal surface;
<emphasis id="12C3A16E0376FFA8FAFBFE80B6987DC2" bold="true" box="[1391,1436,367,392]" pageId="11">mgl</emphasis>
a crescent-shaped lobe with scattered short setae;
<emphasis id="12C3A16E0376FFA8FD44FE7BB1F87DE7" bold="true" box="[720,764,404,429]" pageId="11">ogp</emphasis>
large and nearly entirely membranous, ending before
<emphasis id="12C3A16E0376FFA8FAFCFE7BB6917DE7" bold="true" box="[1384,1429,404,429]" pageId="11">mgl</emphasis>
, with many long setae near apex and free margin extending to ventral region; clump of short setae at base of
<emphasis id="12C3A16E0376FFA8FAFDFE58B6917D9A" bold="true" box="[1385,1429,439,464]" pageId="11">ogp</emphasis>
.
<emphasis id="12C3A16E0376FFA8FF03FE32B21D7DBE" box="[151,281,476,500]" italics="true" pageId="11">Penis valve</emphasis>
. Rounded in lateral view; main dorsal ridge central; apex narrow and parallel-sided, pointed at apex; outer lateral expansion with extremity bilobed, outer lamella lobed; ventral surface with prong narrow and parallelsided, slightly pointed at apex, extending past volsella posteriorly.
<emphasis id="12C3A16E0376FFA8FC01FDCBB0EF7E76" box="[917,1003,548,572]" italics="true" pageId="11">Volsella</emphasis>
. Inner apical corner rounded; medioapical margin convex.
<emphasis id="12C3A16E0376FFA8FE08FDA7B1027E2A" box="[412,518,584,608]" italics="true" pageId="11">Sculpture</emphasis>
. Upper paraocular area and frons densely punctuate, with very fine, contiguous punctures and scattered coarser setal bases intermixed, surface finely and irregularly roughened; lower paraocular area with coarse oblique punctures separated by 12 puncture widths, surface between punctures microreticulate; supraclypeal area with punctures finer than those of lower paraocular area and irregular in spacing in lower half, surface between punctures strongly microreticulate; clypeus with coarse punctures separated by 13 puncture widths, surface between punctures strongly microreticulate; mesoscutum densely punctuate, with fine, contiguous punctures that becoming oblique in central region, and coarser setal bases intermixed more concentrated on laterals; mesoscutellum with fine punctures irregular in size and spacing, surface between punctures smooth; metanotum more densely punctate than mesoscutellum, posterior margin roughened, surface between punctures smooth and shiny; mesepisternum and metepisternum densely punctate, with fine, contiguous punctures and several scattered coarser setal bases intermixed; dorsal surface of metapostnotum with triangular area defined by denser granulation than elsewhere, punctures fine and oblique, and with one unique, coarse carina not reaching edge medially; propodeal lateral and posterior surfaces microareolate, and scattered coarser setal bases intermixed, surfaces microstriate; dorsal surface of T1T4 with fine punctures, separated by 12 puncture widths in central area, surface between punctures microreticulate.
<emphasis id="12C3A16E0376FFA8FD57FBAFB0077812" box="[707,771,1088,1112]" italics="true" pageId="11">Color</emphasis>
. Bright green, except: mandible brown with reddish apex; about apical third of clypeus with yellow transverse band, sometimes small triangular area medially in its upper margin; labral basal process yellow; labral distal process brown; scape, pedicel and flagellum dark brown, flagellum much lighter below than above, sometimes nearly yellow; legs brown, coxae and femora with faint metallic green highlights, inner surface of protibia yellowish; inner surface of meso- and metatarsus light brown; T1T2 brown with metallic green highlights, posterior marginal zones amber, T3T5 amber with bands medially or only disc brown with faint metallic green highlights; sterna amber, except S1 with green highlights; pygidial plate yellow.
<emphasis id="12C3A16E0376FFA8FF7BFAD3B277791E" box="[239,371,1340,1364]" italics="true" pageId="11">Pubescence</emphasis>
. Dorsal surface with very short, fine, plumose whitish setae; long, simple or branched, white or light brown setae on nearly all body parts. Compound eyes with brown and long setae, longer than ocellar diameter; strip of short, plumose white setae along inner margin of compound eye; triangular area of metapostnotum without setae.
</paragraph>
</subSubSection>
<subSubSection id="68AD2EF70376FFA8FF53FA23B0B679AE" box="[199,946,1484,1509]" pageId="11" type="distribution">
<paragraph id="20087D7C0376FFA8FF53FA23B0B679AE" blockId="11.[151,1436,151,1653]" box="[199,946,1484,1509]" pageId="11">
<emphasis id="12C3A16E0376FFA8FF53FA23B26479AF" bold="true" box="[199,352,1484,1509]" pageId="11">Distribution.</emphasis>
This species is known at present from
<collectingRegion id="E273B39E0376FFA8FC81FA23B06879AF" box="[789,876,1484,1509]" pageId="11">Ancash</collectingRegion>
,
<collectingCountry id="58A03DEC0376FFA8FCE3FA22B0AA79AE" box="[887,942,1485,1508]" name="Peru" pageId="11">Peru</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="68AD2EF70376FFA8FF53FA00B0917A3F" pageId="11" type="materials_examined">
<paragraph id="20087D7C0376FFA8FF53FA00B64F7A66" blockId="11.[151,1436,151,1653]" pageId="11">
<materialsCitation id="90DF77210376FFA8FF53FA00B2037A66" ID-GBIF-Occurrence="1949444654" collectingDate="1996-05-15" collectionCode="AMNH" collectorName="J.G. Rozen &amp; A. Ugarte" country="Peru" location="vicinity Carhuaz" pageId="11" specimenCount="1" specimenCount-male="1" stateProvince="Ancash" typeStatus="holotype">
<emphasis id="12C3A16E0376FFA8FF53FA00B2A97A42" bold="true" box="[199,429,1519,1544]" pageId="11">Material examined</emphasis>
.
<emphasis id="12C3A16E0376FFA8FE2AFA1FB1217A42" box="[446,549,1520,1544]" italics="true" pageId="11">
<typeStatus id="FF0CC3DE0376FFA8FE2AFA1FB1217A42" box="[446,549,1520,1544]" pageId="11" type="holotype">Holotype</typeStatus>
</emphasis>
, male, PERU: Ancash: vicinity Carhuaz,
<date id="54095BBC0376FFA8FB99FA00B7787A42" box="[1037,1148,1519,1544]" pageId="11" value="1996-05-15">15.v.1996</date>
(J.G.Rozen &amp; A.Ugarte) (
<collectionCode id="46A6E5B90376FFA8FF0BF9FBB3FF7A66" LSID="urn:lsid:biocol.org:col:34925" box="[159,251,1556,1580]" collectionName="USA, New York, New York, American Museum of Natural History" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="11">AMNH</collectionCode>
).
</materialsCitation>
<materialsCitation id="90DF77210376FFA8FE99F9FAB64F7A66" ID-GBIF-Occurrence="1949444655" box="[269,1355,1556,1581]" collectingDate="1996-05-15" collectingRegion="Carhuaz" collectionCode="MPEG" collectorName="J.G. Rozen &amp; A. Ugarte" country="Peru" location="vicinity Carhuaz" pageId="11" specimenCount="1" specimenCount-male="1" stateProvince="Ancash" typeStatus="paratype">
<typeStatus id="FF0CC3DE0376FFA8FE99F9FAB2767A67" box="[269,370,1557,1581]" pageId="11" type="paratype">Paratype</typeStatus>
, PERU: Ancash:
<specimenCount id="36B1B6F50376FFA8FDA2F9FBB1807A67" box="[566,644,1556,1581]" pageId="11" type="male">1 male</specimenCount>
, vicinity Carhuaz,
<date id="54095BBC0376FFA8FCCCF9FBB0C27A67" box="[856,966,1556,1581]" pageId="11" value="1996-05-15">15.v.1996</date>
(J.G.Rozen &amp; A.Ugarte) (
<collectionCode id="46A6E5B90376FFA8FB7FF9FBB63B7A66" box="[1259,1343,1556,1580]" collectionName="Brazil, Para, Belem, Museu Paraense Emilio Goeldi" httpUri="http://grbio.org/cool/90vz-swjg" name="Museu Paraense Emilio Goeldi" pageId="11">MPEG</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="20087D7C0376FFA8FF53F9D8B0917A3F" blockId="11.[151,1436,151,1653]" pageId="11">
<emphasis id="12C3A16E0376FFA8FF53F9D8B23C7A1A" bold="true" box="[199,312,1591,1616]" pageId="11">Etmology</emphasis>
. The specific epithet honors the Incan Civilization that once lived widely in the Andean region, but particularly within Peru.The name is treated as a noun in apposition.
</paragraph>
</subSubSection>
</treatment>
</document>