150 lines
19 KiB
XML
150 lines
19 KiB
XML
<document id="86D6139BD991C493C42C67E2F324C171" ID-DOI="10.2307/3889334" ID-GBIF-Dataset="6f85aa0a-0368-41a5-83d3-21c33f40845e" ID-Zenodo-Dep="3739929" IM.tables_requiresApprovalFor="existingObjects,jeremy" IM.taxonomicNames_approvedBy="jeremy" approvalRequired="19" approvalRequired_for_bibRefs="2" approvalRequired_for_illustrations="7" approvalRequired_for_taxonomicNames="2" approvalRequired_for_textStreams="8" checkinTime="1585838636713" checkinUser="plazi" docAuthor="Christopher A. Brochu" docDate="2003" docId="9A3A87D00B5E0D67FAD5AA2438CFF7AD" docLanguage="en" docName="socVertebratePaleoMemoir_Brochu2003ABBYY_2.pdf" docOrigin="Journal of Vertebrate Paleontology 22" docStyle="DocumentStyle{}" docTitle="Albertosaurus Osborn 1905" docType="treatment" docVersion="12" lastPageNumber="4" masterDocId="6603FFA80B5D0D64FFF6AB533D39FF9A" masterDocTitle="Osteology of Tyrannosaurus rex: insights from a nearly complete skeleton and high-resolution computed tomographic analysis of the skull" masterLastPageNumber="138" masterPageNumber="1" pageNumber="4" updateTime="1689591936236" updateUser="jeremy">
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<mods:titleInfo id="B5334C3E34A166DA56F85E3560723F62">
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<mods:title id="9662EF0B43EB61BA49C75CD55AD63C4F">Osteology of Tyrannosaurus rex: insights from a nearly complete skeleton and high-resolution computed tomographic analysis of the skull</mods:title>
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<mods:name id="8E6F4EF1C4AE930369DB47E463722F3D" type="personal">
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<mods:roleTerm id="BF264D6E3B571252ADD24A33C64B6F15">Author</mods:roleTerm>
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<mods:namePart id="4829126F40F227099AA9DB57EE4376D4">Christopher A. Brochu</mods:namePart>
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<mods:affiliation id="52ABD0FACC2CC418E341D8EF7FF427D9">Department of Geology, Field Museum, 1400 S. Lake Shore Drive, Chicago, Illinois 60605</mods:affiliation>
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<mods:typeOfResource id="13501118E569C4878AE1BDDEA7F6687E">text</mods:typeOfResource>
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<mods:title id="E069BE13A1C71E2F3921946CC97E1B52">Journal of Vertebrate Paleontology</mods:title>
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</mods:titleInfo>
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<mods:part id="79360274137367B762F4EF9F7C4100C2">
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<mods:date id="75F1D8BFAADEE6A724C024079D46AD57">2003</mods:date>
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<mods:detail id="9B0E723D03FF8D6B14DE7A679EA20565" type="pubDate">
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<mods:number id="A2E590D7BF4F4A5B0CF1651AAE86E8E5">2003-12-31</mods:number>
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<mods:detail id="9B6B74A687CBD5FA897E9E0D50C5F9AD" type="volume">
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<mods:number id="EB53A2A43389C58FF4BCF6B890BA6454">22</mods:number>
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<mods:classification id="CE16B946F9616904C089B3AF9C709D75">journal article</mods:classification>
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<mods:identifier id="F9D24F6F31EAF8096CF45240B1E055F8" type="DOI">10.2307/3889334</mods:identifier>
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<treatment id="9A3A87D00B5E0D67FAD5AA2438CFF7AD" ID-DOI="http://doi.org/10.5281/zenodo.3810813" ID-GBIF-Taxon="190303346" ID-Zenodo-Dep="3810813" LSID="urn:lsid:plazi:treatment:9A3A87D00B5E0D67FAD5AA2438CFF7AD" httpUri="http://treatment.plazi.org/id/9A3A87D00B5E0D67FAD5AA2438CFF7AD" lastPageNumber="4" pageId="3" pageNumber="4">
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<subSubSection id="5A89654D0B5E0D67FAD5AA243B2BFE00" box="[1315,1554,375,410]" pageId="3" pageNumber="4" type="nomenclature">
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<paragraph id="122C36C60B5E0D67FAD5AA243B2BFE00" blockId="3.[1276,2332,280,3163]" box="[1315,1554,375,410]" pageId="3" pageNumber="4">
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<taxonomicName id="D5934D450B5E0D67FAD5AA243B2BFE00" authority="Osborn, 1905" authorityName="Osborn" authorityYear="1905" baseAuthorityName="Witmer" baseAuthorityYear="1997" box="[1315,1554,375,410]" class="Reptilia" family="Tyrannosauridae" genus="Albertosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="3" pageNumber="4" phylum="Chordata" rank="genus">
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<emphasis id="20E7EAD40B5E0D67FAD5AA243B2BFE00" box="[1315,1554,375,410]" italics="true" pageId="3" pageNumber="4">Albertosaurus</emphasis>
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</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="5A89654D0B5E0D67F9D3AA253AFDFD09" pageId="3" pageNumber="4" type="discussion">
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<paragraph id="122C36C60B5E0D67F9D3AA253AFDFD09" blockId="3.[1276,2332,280,3163]" pageId="3" pageNumber="4">
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and
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<emphasis id="20E7EAD40B5E0D67F983AA2635D7FE0F" box="[1653,2286,369,408]" italics="true" pageId="3" pageNumber="4">
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Gorgosaurus—
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<taxonomicName id="D5934D450B5E0D67F88EAA213558FE0C" authority="Osborn, 1905" authorityName="Osborn" authorityYear="1905" baseAuthorityName="Witmer" baseAuthorityYear="1997" box="[1912,2145,370,406]" class="Reptilia" family="Tyrannosauridae" genus="Albertosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="3" pageNumber="4" phylum="Chordata" rank="genus">Albertosaurus</taxonomicName>
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libratus
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</emphasis>
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is based on a nearly complete skeleton (NMC 2120) described by Lambe (1914a, b, 1917) as
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<emphasis id="20E7EAD40B5E0D67F93EAA9B3508FE71" box="[1736,2097,455,492]" italics="true" pageId="3" pageNumber="4">
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<taxonomicName id="D5934D450B5E0D67F93EAA9B3AA4FE76" box="[1736,1949,456,492]" class="Reptilia" family="Tyrannosauridae" genus="Gorgosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="3" pageNumber="4" phylum="Chordata" rank="genus">Gorgosaurus</taxonomicName>
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libratus.
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</emphasis>
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The holotype skull is crushed, but most of it is preserved. It hails from the Dinosaur Park Formation and is the best-sampled North American tyrannosaurid, with multiple well-preserved specimens having been collected since Lambe’s time.
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</paragraph>
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</subSubSection>
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<subSubSection id="5A89654D0B5E0D67FAD3A9CE38CFF7AD" pageId="3" pageNumber="4" type="discussion">
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<paragraph id="122C36C60B5E0D67FAD3A9CE3B22FC70" blockId="3.[1276,2332,280,3163]" pageId="3" pageNumber="4">
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<taxonomicName id="D5934D450B5E0D67FAD3A9CE3AE5FD24" authority="Osborn, 1905" authorityName="Osborn" authorityYear="1905" box="[1317,2012,666,705]" class="Reptilia" family="Tyrannosauridae" genus="Albertosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="sarcophagus">
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<emphasis id="20E7EAD40B5E0D67FAD3A9CE3BD4FD5A" box="[1317,1773,668,705]" italics="true" pageId="3" pageNumber="4">Albertosaurus sarcophagus</emphasis>
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<bibRefCitation id="76024B370B5E0D67F90BA9C83AE5FD24" author="Osborn, H. F." box="[1789,2012,666,703]" journalOrPublisher="Bulletin of the American Museum of Natural History" pageId="3" pageNumber="4" pagination="259 - 265" part="21" refId="ref76275" refString="Osborn, H. F. 1905. Tyrannosaurus and other Cretaceous carnivorous dinosaurs. Bulletin of the American Museum of Natural History 21: 259 - 265." title="Tyrannosaurus and other Cretaceous carnivorous dinosaurs" type="journal article" year="1905">Osborn, 1905</bibRefCitation>
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</taxonomicName>
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is based on a partial skeleton (NMC 5600) from the overlying Edmonton Formation of Alberta (
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<bibRefCitation id="76024B370B5E0D67F9BDA9A33A0CFC89" author="Osborn, H. F." box="[1611,1845,751,788]" journalOrPublisher="Bulletin of the American Museum of Natural History" pageId="3" pageNumber="4" pagination="259 - 265" part="21" refId="ref76275" refString="Osborn, H. F. 1905. Tyrannosaurus and other Cretaceous carnivorous dinosaurs. Bulletin of the American Museum of Natural History 21: 259 - 265." title="Tyrannosaurus and other Cretaceous carnivorous dinosaurs" type="journal article" year="1905">Osborn, 1905</bibRefCitation>
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). It is not as well preserved as the type of
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<emphasis id="20E7EAD40B5E0D67FA1DA8483B9AFCA4" box="[1515,1699,794,831]" italics="true" pageId="3" pageNumber="4">A. libratus.</emphasis>
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A partial skeleton with no skull from the Edmonton was later described by
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<bibRefCitation id="76024B370B5E0D67F888A810356AFCFD" author="Parks, W. A." box="[1918,2131,835,871]" journalOrPublisher="University of Toronto Studies (Geological Series)" pageId="3" pageNumber="4" pagination="1 - 24" part="26" refId="ref76516" refString="Parks, W. A. 1928. Albertosaurus arctunguis, a new species of theropodous dinosaur from the Edmonton Formation of Alberta. University of Toronto Studies (Geological Series) 26: 1 - 24." title="Albertosaurus arctunguis, a new species of theropodous dinosaur from the Edmonton Formation of Alberta" type="journal article" year="1928">Parks (1928)</bibRefCitation>
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as
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<emphasis id="20E7EAD40B5E0D67F779A8113B0CFC0E" italics="true" pageId="3" pageNumber="4">
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<taxonomicName id="D5934D450B5E0D67F779A8113855FC0F" authority="Osborn, 1905" authorityName="Osborn" authorityYear="1905" baseAuthorityName="Witmer" baseAuthorityYear="1997" class="Reptilia" family="Tyrannosauridae" genus="Albertosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="3" pageNumber="4" phylum="Chordata" rank="genus">Albertosaurus</taxonomicName>
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arctunguis',
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</emphasis>
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most authorities now regard
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<emphasis id="20E7EAD40B5E0D67F7C2A83F3421FC0A" box="[2100,2328,876,912]" italics="true" pageId="3" pageNumber="4">A. arctunguis</emphasis>
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to be a junior synonym of
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<taxonomicName id="D5934D450B5E0D67F941A8C93A81FC24" authorityName="Osborn" authorityYear="1905" box="[1719,1976,922,958]" class="Reptilia" family="Tyrannosauridae" genus="Albertosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="sarcophagus">
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<emphasis id="20E7EAD40B5E0D67F941A8C93A81FC24" box="[1719,1976,922,958]" italics="true" pageId="3" pageNumber="4">A. sarcophagus</emphasis>
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</taxonomicName>
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(Molnar et al., 1990; Carpenter, 1992).
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</paragraph>
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<paragraph id="122C36C60B5E0D67FADFA8A238F2FA0B" blockId="3.[1276,2332,280,3163]" pageId="3" pageNumber="4">
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<bibRefCitation id="76024B370B5E0D67FADFA8A23B2EFB8F" author="Russell, D. A." box="[1321,1559,1009,1045]" journalOrPublisher="National Museum of Natural Sciences Publications in Palaeontology" pageId="3" pageNumber="4" pagination="1 - 34" part="1" refId="ref77430" refString="Russell, D. A. 1970. Tyrannosaurs from the Late Cretaceous of western Canada. National Museum of Natural Sciences Publications in Palaeontology 1: 1 - 34." title="Tyrannosaurs from the Late Cretaceous of western Canada" type="journal article" year="1970">Russell (1970)</bibRefCitation>
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was the first to synonymize
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<taxonomicName id="D5934D450B5E0D67F807A8BE35FFFB8B" box="[2033,2246,1005,1041]" class="Reptilia" family="Tyrannosauridae" genus="Gorgosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="3" pageNumber="4" phylum="Chordata" rank="genus">
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<emphasis id="20E7EAD40B5E0D67F807A8BE35FFFB8B" box="[2033,2246,1005,1041]" italics="true" pageId="3" pageNumber="4">Gorgosaurus</emphasis>
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</taxonomicName>
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with
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<emphasis id="20E7EAD40B5E0D67FB0BAF4F38D7FBDA" box="[1277,1518,1052,1088]" italics="true" pageId="3" pageNumber="4">
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<taxonomicName id="D5934D450B5E0D67FB0BAF4F38D0FBDA" authority="Osborn, 1905" authorityName="Osborn" authorityYear="1905" baseAuthorityName="Witmer" baseAuthorityYear="1997" box="[1277,1513,1052,1088]" class="Reptilia" family="Tyrannosauridae" genus="Albertosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="3" pageNumber="4" phylum="Chordata" rank="genus">Albertosaurus</taxonomicName>
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.
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</emphasis>
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I agree with Holtz (2001a) that the character states diagnosing a more inclusive
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<taxonomicName id="D5934D450B5E0D67F8A4AF103502FBFD" authority="Osborn, 1905" authorityName="Osborn" authorityYear="1905" baseAuthorityName="Witmer" baseAuthorityYear="1997" box="[1874,2107,1091,1127]" class="Reptilia" family="Tyrannosauridae" genus="Albertosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="3" pageNumber="4" phylum="Chordata" rank="genus">
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<emphasis id="20E7EAD40B5E0D67F8A4AF103502FBFD" box="[1874,2107,1091,1127]" italics="true" pageId="3" pageNumber="4">Albertosaurus</emphasis>
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</taxonomicName>
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are plesiomorphic (and thus problematic), and the characters used by
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<bibRefCitation id="76024B370B5E0D67F73BAF3F388AFB25" author="Russell, D. A." journalOrPublisher="National Museum of Natural Sciences Publications in Palaeontology" pageId="3" pageNumber="4" pagination="1 - 34" part="1" refId="ref77430" refString="Russell, D. A. 1970. Tyrannosaurs from the Late Cretaceous of western Canada. National Museum of Natural Sciences Publications in Palaeontology 1: 1 - 34." title="Tyrannosaurs from the Late Cretaceous of western Canada" type="journal article" year="1970">Russell (1970)</bibRefCitation>
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to separate them as species rely on skull and limb proportions that might be subject to preservational or ontogenetic variation. Ongoing work by Philip Currie (P. Currie, pers. comm.) is clarifying the diagnoses of these fossils, but for purposes of this analysis, they will be considered species of
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<emphasis id="20E7EAD40B5E0D67F710AE6C38F2FA0B" italics="true" pageId="3" pageNumber="4">
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<taxonomicName id="D5934D450B5E0D67F710AE6C38FFFA0B" authority="Osborn, 1905" authorityName="Osborn" authorityYear="1905" baseAuthorityName="Witmer" baseAuthorityYear="1997" class="Reptilia" family="Tyrannosauridae" genus="Albertosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="3" pageNumber="4" phylum="Chordata" rank="genus">Albertosaurus</taxonomicName>
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.
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</emphasis>
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</paragraph>
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<paragraph id="122C36C60B5E0D67FADCAECA38CFF7AD" blockId="3.[1276,2332,280,3163]" pageId="3" pageNumber="4">
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Carpenter (1992) argued that the promaxillary fenestra lies close to the maxillary fenestra in
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<emphasis id="20E7EAD40B5E0D67F8ABAE923524FA7E" box="[1885,2077,1472,1509]" italics="true" pageId="3" pageNumber="4">A. libratus,</emphasis>
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but not in
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<emphasis id="20E7EAD40B5E0D67F700AEED38E3F989" italics="true" pageId="3" pageNumber="4">
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<taxonomicName id="D5934D450B5E0D67F700AEED38EDF989" authorityName="Osborn" authorityYear="1905" class="Reptilia" family="Tyrannosauridae" genus="Albertosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="sarcophagus">A. sarcophagus</taxonomicName>
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,
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</emphasis>
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and that the promaxillary fenestra of
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<emphasis id="20E7EAD40B5E0D67F79EAEB83423F995" box="[2152,2330,1515,1551]" italics="true" pageId="3" pageNumber="4">A. libratus</emphasis>
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is invisible in lateral view. In all mature
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<emphasis id="20E7EAD40B5E0D67F828AD4435ADF9A0" box="[2014,2196,1558,1595]" italics="true" pageId="3" pageNumber="4">A. libratus</emphasis>
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and
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<taxonomicName id="D5934D450B5E0D67F703AD4638E8F9F2" authorityName="Osborn" authorityYear="1905" class="Reptilia" family="Tyrannosauridae" genus="Albertosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="sarcophagus">
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<emphasis id="20E7EAD40B5E0D67F703AD4638E8F9F2" italics="true" pageId="3" pageNumber="4">A. sarcophagus</emphasis>
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</taxonomicName>
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maxillae available for this study, the morphology of the maxillary and promaxillary fenestrae was identical. The reconstruction of
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<emphasis id="20E7EAD40B5E0D67F9D6ADCB3BF6F926" box="[1568,1743,1688,1724]" italics="true" pageId="3" pageNumber="4">A. libratus</emphasis>
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figured by Carpenter (1992:fig. 2E) looks similar to that published by
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<bibRefCitation id="76024B370B5E0D67F8BFAD933508F97E" author="Russell, D. A." box="[1865,2097,1728,1764]" journalOrPublisher="National Museum of Natural Sciences Publications in Palaeontology" pageId="3" pageNumber="4" pagination="1 - 34" part="1" refId="ref77430" refString="Russell, D. A. 1970. Tyrannosaurs from the Late Cretaceous of western Canada. National Museum of Natural Sciences Publications in Palaeontology 1: 1 - 34." title="Tyrannosaurs from the Late Cretaceous of western Canada" type="journal article" year="1970">Russell (1970</bibRefCitation>
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:fig. 1), but in Russell’s reconstruction, the lateral morphological details are based on FMNH PR308. In that specimen, the promaxillary fenestra is not preserved on either side. In fact, based on the number of dentary and maxillary alveoli, this specimen may pertain to
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<emphasis id="20E7EAD40B5E0D67FA3CACC53BEAF820" box="[1482,1747,1942,1978]" italics="true" pageId="3" pageNumber="4">
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<taxonomicName id="D5934D450B5E0D67FA3CACC53BF4F820" baseAuthorityName="Russell" baseAuthorityYear="1970" box="[1482,1741,1942,1978]" class="Reptilia" family="Tyrannosauridae" genus="Daspletosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="3" pageNumber="4" phylum="Chordata" rank="genus">Daspletosaurus</taxonomicName>
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||
,
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</emphasis>
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not
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<taxonomicName id="D5934D450B5E0D67F8B6ACC73510F822" authority="Osborn, 1905" authorityName="Osborn" authorityYear="1905" baseAuthorityName="Witmer" baseAuthorityYear="1997" box="[1856,2089,1940,1976]" class="Reptilia" family="Tyrannosauridae" genus="Albertosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="3" pageNumber="4" phylum="Chordata" rank="genus">
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<emphasis id="20E7EAD40B5E0D67F8B6ACC73510F822" box="[1856,2089,1940,1976]" italics="true" pageId="3" pageNumber="4">Albertosaurus</emphasis>
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</taxonomicName>
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(
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<bibRefCitation id="76024B370B5E0D67F7A4ACC03428F82D" author="Carr, T. D." box="[2130,2321,1939,1975]" journalOrPublisher="Journal of Vertebrate Paleontology" pageId="3" pageNumber="4" pagination="497 - 520" part="19" refId="ref71985" refString="Carr, T. D. 1999. Craniofacial ontogeny in Tyrannosauridae (Dinosauria, Theropoda). Journal of Vertebrate Paleontology 19: 497 - 520." title="Craniofacial ontogeny in Tyrannosauridae (Dinosauria, Theropoda)" type="journal article" year="1999">Carr, 1999</bibRefCitation>
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||
; Holtz, 2001a)—and the promaxillary fenestra is not visible in lateral view in
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<emphasis id="20E7EAD40B5E0D67FA0AACB93A3CF794" box="[1532,1797,2026,2062]" italics="true" pageId="3" pageNumber="4">
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||
<taxonomicName id="D5934D450B5E0D67FA0AACB93BC6F794" baseAuthorityName="Russell" baseAuthorityYear="1970" box="[1532,1791,2026,2062]" class="Reptilia" family="Tyrannosauridae" genus="Daspletosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dinosauria" pageId="3" pageNumber="4" phylum="Chordata" rank="genus">Daspletosaurus</taxonomicName>
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,
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</emphasis>
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as implied by Carpenter (1992) for
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<emphasis id="20E7EAD40B5E0D67FACAA34038CFF7AD" box="[1340,1526,2067,2103]" italics="true" pageId="3" pageNumber="4">A. libratus.</emphasis>
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</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |