163 lines
18 KiB
XML
163 lines
18 KiB
XML
<document id="9EABEF06D8E4D364AA80DE027799EEE5" ID-DOI="10.11646/zootaxa.4994.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="5043261" ID-ZooBank="107FCA64-345F-40A4-99D3-5C1441EEAD93" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1624993528766" checkinUser="plazi" docAuthor="Perkins, Philip D." docDate="2021" docId="03A187DEFFE8FFEB6194FF670737F35E" docLanguage="en" docName="zootaxa.4994.1.1.pdf" docOrigin="Zootaxa 4994 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Hydrochus colossus Perkins 2021, new species" docType="treatment" docVersion="3" lastPageNumber="21" masterDocId="FF98FFA6FFFCFFFF6103FFF0045FF707" masterDocTitle="Taxonomy of water beetles in the genus Hydrochus Leach, 1817, from Bolivia, Brazil and Paraguay (Coleoptera: Hydrochidae)" masterLastPageNumber="93" masterPageNumber="1" pageNumber="21" updateTime="1698978858223" updateUser="plazi">
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<mods:mods id="8F307EFC8C8A9FEE2C51ABD450D31AA2" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="1F25FCBC84FBB746361E4F9235E1D3F2">
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<mods:title id="DA6DDB6D2DF18078B316DF0DF72E4740">Taxonomy of water beetles in the genus Hydrochus Leach, 1817, from Bolivia, Brazil and Paraguay (Coleoptera: Hydrochidae)</mods:title>
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</mods:titleInfo>
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<mods:name id="9E27B3128C7D3804F3CD80CE29985B8B" type="personal">
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<mods:role id="82F4CA4C29D72A66B33A2A0304518833">
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<mods:roleTerm id="03B97704309C2998BC62AF65F02678E8">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="100B747F85C1D4EA4A9A6D8D8D2FFA31">Perkins, Philip D.</mods:namePart>
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<mods:typeOfResource id="98F1DB9DB7EA22C7D18CFFBED919C53C">text</mods:typeOfResource>
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<mods:title id="EB589E78B13FC407A20F412184AABE9D">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="9D4C65671F752BE2A1F2F4467CAA36F6">
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<mods:date id="1D2DDF8E47CE19A8769D27D6A27287D5">2021</mods:date>
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<mods:detail id="C9DEAD037AFCFE8A5E267D838F9F70A9" type="pubDate">
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<mods:number id="5416224541628CC56C9012C6724D2AF9">2021-06-28</mods:number>
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<mods:detail id="5EAF9B2BE0DE2F2381C94094AE826F7D" type="volume">
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<mods:number id="22462C0BF88D99C1681E94B0F07B05FF">4994</mods:number>
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</mods:detail>
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<mods:detail id="734F246E91AE4E3C20D929F5A6A10939" type="issue">
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<mods:number id="A0DA3CB80E015D7AAFEE0F3A370E3C1F">1</mods:number>
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<mods:start id="6BB82E5E3C35EBE56A4F41FBAF6D9184">1</mods:start>
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<mods:end id="6457D909C2A1866D80C44105FBF31E69">93</mods:end>
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<mods:classification id="88987246AB6CC26F04D1E4DBF56FDB02">journal article</mods:classification>
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<mods:identifier id="D42C2A73B7A6E9424F77A8B9CE118991" type="DOI">10.11646/zootaxa.4994.1.1</mods:identifier>
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<mods:identifier id="6E4B7C0982FFCF1C0B66EEE9F9110AD3" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="D673ED9743414A96D4B2F0A099DD9AC2" type="Zenodo-Dep">5043261</mods:identifier>
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<mods:identifier id="A12E6BCFB50AC57133E66A862BB1235A" type="ZooBank">107FCA64-345F-40A4-99D3-5C1441EEAD93</mods:identifier>
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</mods:mods>
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<treatment id="03A187DEFFE8FFEB6194FF670737F35E" LSID="urn:lsid:plazi:treatment:03A187DEFFE8FFEB6194FF670737F35E" httpUri="http://treatment.plazi.org/id/03A187DEFFE8FFEB6194FF670737F35E" lastPageNumber="21" pageId="20" pageNumber="21">
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<subSubSection id="C3126543FFE8FFEB6194FF670677F7B6" box="[151,552,151,178]" pageId="20" pageNumber="21" type="nomenclature">
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<paragraph id="8BB736C8FFE8FFEB6194FF670677F7B6" blockId="20.[151,552,151,214]" box="[151,552,151,178]" pageId="20" pageNumber="21">
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<heading id="D0FF81A4FFE8FFEB6194FF670677F7B6" bold="true" box="[151,552,151,178]" fontSize="11" level="2" pageId="20" pageNumber="21" reason="4">
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<emphasis id="B97CEADAFFE8FFEB6194FF670677F7B6" bold="true" box="[151,552,151,178]" pageId="20" pageNumber="21">
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||
<taxonomicName id="4C084D4BFFE8FFEB6194FF6705D6F7B5" ID-CoL="8Y4PN" authority="Perkins, 2021" authorityName="Perkins" authorityYear="2021" box="[151,393,151,178]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="colossus" status="sp. nov.">
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<emphasis id="B97CEADAFFE8FFEB6194FF6705D6F7B5" bold="true" box="[151,393,151,178]" italics="true" pageId="20" pageNumber="21">Hydrochus colossus</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="A24F57A1FFE8FFEB609BFF670677F7B6" box="[408,552,151,177]" pageId="20" pageNumber="21" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C3126543FFE8FFEB6194FF4B0568F7D1" box="[151,311,187,214]" pageId="20" pageNumber="21" type="description">
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||
<paragraph id="8BB736C8FFE8FFEB6194FF4B0568F7D1" blockId="20.[151,552,151,214]" box="[151,311,187,214]" pageId="20" pageNumber="21">
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<figureCitation id="13332A4DFFE8FFEB6194FF4B04BCF7D2" box="[151,227,187,213]" captionStart="FIGURE 1" captionStartId="6.[151,250,1943,1967]" captionTargetBox="[158,1402,188,1903]" captionTargetId="figure-28@6.[152,1435,181,1920]" captionTargetPageId="6" captionText="FIGURE 1. Dorsal habitus of Hydrochus species in study area." figureDoi="http://doi.org/10.5281/zenodo.5043263" httpUri="https://zenodo.org/record/5043263/files/figure.png" pageId="20" pageNumber="21">Figs. 1</figureCitation>
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,
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<figureCitation id="13332A4DFFE8FFEB61F2FF4B0550F7D1" box="[241,271,187,214]" captionStart="FIGURE 52" captionStartId="78.[151,250,1963,1987]" captionTargetBox="[239,1343,211,1896]" captionTargetId="figure-28@78.[177,1410,181,1939]" captionTargetPageId="78" captionText="FIGURE 52. Hydrochus colossus n. sp., habitus and female genital segments of holotype." figureDoi="http://doi.org/10.5281/zenodo.5043344" httpUri="https://zenodo.org/record/5043344/files/figure.png" pageId="20" pageNumber="21">52</figureCitation>
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,
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<figureCitation id="13332A4DFFE8FFEB6018FF4B0568F7D1" box="[283,311,187,214]" captionStart="FIGURES 53–54" captionStartId="79.[151,264,1964,1988]" captionTargetBox="[218,1368,181,1941]" captionTargetId="figure-15@79.[218,1368,181,1941]" captionTargetPageId="79" captionText="FIGURES 53–54. Geographical distributions. 53. H spanglerorum species group. 54. H collaris species group." figureDoi="http://doi.org/10.5281/zenodo.5043348" httpUri="https://zenodo.org/record/5043348/files/figure.png" pageId="20" pageNumber="21">53</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3126543FFE8FFEB6194FEF3052DF662" pageId="20" pageNumber="21" type="materials_examined">
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<paragraph id="8BB736C8FFE8FFEB6194FEF3052DF662" blockId="20.[151,1437,259,1114]" pageId="20" pageNumber="21">
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<emphasis id="B97CEADAFFE8FFEB6194FEF30513F61A" bold="true" box="[151,332,259,285]" pageId="20" pageNumber="21">Type Material.</emphasis>
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<materialsCitation id="3B603C95FFE8FFEB6054FEF3052DF662" collectingDate="2018-06-27" collectionCode="INPA" collectorName="Hamada" country="Brazil" elevation="225" latitude="-20.72281" location="Aquidauana" longLatPrecision="1" longitude="-55.69127" pageId="20" pageNumber="21" specimenCount="1" stateProvince="Mato Grosso do Sul" typeStatus="holotype">
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<emphasis id="B97CEADAFFE8FFEB6054FEF30665F61A" bold="true" box="[343,570,259,285]" pageId="20" pageNumber="21">
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<typeStatus id="54B3886AFFE8FFEB6054FEF3059BF61A" box="[343,452,259,285]" pageId="20" pageNumber="21" type="holotype">Holotype</typeStatus>
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(female):
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</emphasis>
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“
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<collectingCountry id="F31F7658FFE8FFEB634CFEF306E4F61A" box="[591,699,259,285]" name="Brazil" pageId="20" pageNumber="21">BRAZIL</collectingCountry>
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:
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<collectingRegion id="49CCF82AFFE8FFEB63C9FEF407E5F61A" box="[714,954,259,286]" country="Brazil" name="Mato Grosso do Sul" pageId="20" pageNumber="21">Mato Grosso do Sul</collectingRegion>
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,
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<location id="8ED76013FFE8FFEB62C4FEF30011F61A" LSID="urn:lsid:plazi:treatment:03A187DEFFE8FFEB6194FF670737F35E:8ED76013FFE8FFEB62C4FEF30011F61A" box="[967,1102,259,285]" country="Brazil" latitude="-20.72281" longLatPrecision="1" longitude="-55.69127" name="Aquidauana" pageId="20" pageNumber="21" stateProvince="Mato Grosso do Sul">Aquidauana</location>
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(c.
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<quantity id="4CF09B2DFFE8FFEB6581FEF30092F61A" box="[1154,1229,259,285]" metricMagnitude="4" metricUnit="m" metricValue="2.7" pageId="20" pageNumber="21" unit="km" value="27.0">27 km</quantity>
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S) on MS-174 (
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<geoCoordinate id="EE3C500FFFE8FFEB6490FEF3055CF646" degrees="20.72281" direction="south" orientation="latitude" pageId="20" pageNumber="21" precision="1" value="-20.72281">- 20.72281</geoCoordinate>
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,
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<geoCoordinate id="EE3C500FFFE8FFEB6012FED705D6F645" box="[273,393,295,322]" degrees="55.69127" direction="west" orientation="longitude" pageId="20" pageNumber="21" precision="1" value="-55.69127">-55.69127</geoCoordinate>
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);
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<quantity id="4CF09B2DFFE8FFEB609CFED705BCF645" box="[415,483,295,322]" metricMagnitude="2" metricUnit="m" metricValue="2.25" pageId="20" pageNumber="21" unit="m" value="225.0">
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<elevation id="0025D1FBFFE8FFEB609CFED705BCF645" box="[415,483,295,322]" metricMagnitude="2" metricUnit="m" metricValue="2.25" pageId="20" pageNumber="21" unit="m" value="225.0">225m</elevation>
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</quantity>
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; leg.
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<collectorName id="26FD531EFFE8FFEB6327FED706DEF646" box="[548,641,295,321]" pageId="20" pageNumber="21">Hamada</collectorName>
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and team;
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||
<date id="FFB61008FFE8FFEB6204FED707DEF646" box="[775,897,295,321]" pageId="20" pageNumber="21" value="2018-06-27">
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||
<collectingDate id="EFF2E9E0FFE8FFEB6204FED707DEF646" box="[775,897,295,321]" pageId="20" pageNumber="21" value="2018-06-27">27.vi.2018</collectingDate>
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</date>
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; seepage and debris nr. stream margin; BR18- 0627-01E” (
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<collectionCode id="ED19AE0DFFE8FFEB6022FEBB0538F662" box="[289,359,331,357]" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="20" pageNumber="21">INPA</collectionCode>
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).
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</materialsCitation>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C3126543FFE8FFEB61C4FE9F06B3F6F1" pageId="20" pageNumber="21" type="diagnosis">
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<paragraph id="8BB736C8FFE8FFEB61C4FE9F06B3F6F1" blockId="20.[151,1437,259,1114]" pageId="20" pageNumber="21">
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<emphasis id="B97CEADAFFE8FFEB61C4FE9F058CF68E" bold="true" box="[199,467,367,393]" pageId="20" pageNumber="21">Differential Diagnosis.</emphasis>
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A member of the
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<taxonomicName id="4C084D4BFFE8FFEB63A1FE80073EF68E" box="[674,865,367,393]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="spanglerorum">
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<emphasis id="B97CEADAFFE8FFEB63A1FE80073EF68E" box="[674,865,367,393]" italics="true" pageId="20" pageNumber="21">H. spanglerorum</emphasis>
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</taxonomicName>
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species group, differentiated from the other members (
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<taxonomicName id="4C084D4BFFE8FFEB61D4FE64053BF6AA" authorityName="Perkins" authorityYear="2021" box="[215,356,404,429]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="accinctus">
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<emphasis id="B97CEADAFFE8FFEB61D4FE64053BF6AA" box="[215,356,404,429]" italics="true" pageId="20" pageNumber="21">H. accinctus</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C084D4BFFE8FFEB6098FE640642F6AA" authorityName="Perkins" authorityYear="2021" box="[411,541,403,429]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="colossus">
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<emphasis id="B97CEADAFFE8FFEB6098FE640642F6AA" box="[411,541,403,429]" italics="true" pageId="20" pageNumber="21">H. colossus</emphasis>
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</taxonomicName>
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) by the combination of large, robust, strongly sclerotized body (ca. 5.10 x 2.00 mm), the densely punctate dorsum, the relatively wide elytral interstriae. Refer to the diagnosis of
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<taxonomicName id="4C084D4BFFE8FFEB65EBFE48012AF6D6" authorityName="Perkins" authorityYear="2021" box="[1256,1397,440,465]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="accinctus">
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<emphasis id="B97CEADAFFE8FFEB65EBFE48012AF6D6" box="[1256,1397,440,465]" italics="true" pageId="20" pageNumber="21">H. accinctus</emphasis>
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</taxonomicName>
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for characteristics of the
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<taxonomicName id="4C084D4BFFE8FFEB6085FE2C061BF6F2" box="[390,580,475,501]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="spanglerorum">
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<emphasis id="B97CEADAFFE8FFEB6085FE2C061BF6F2" box="[390,580,475,501]" italics="true" pageId="20" pageNumber="21">H. spanglerorum</emphasis>
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</taxonomicName>
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species group.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3126543FFE8FFEB61C4FE0F06BDF331" pageId="20" pageNumber="21" type="description">
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<paragraph id="8BB736C8FFE8FFEB61C4FE0F06CBF5CA" blockId="20.[151,1437,259,1114]" pageId="20" pageNumber="21">
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||
<emphasis id="B97CEADAFFE8FFEB61C4FE0F050EF51E" bold="true" box="[199,337,511,537]" pageId="20" pageNumber="21">Description</emphasis>
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||
. Size:
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<typeStatus id="54B3886AFFE8FFEB60A7FE0F0659F51E" box="[420,518,511,537]" pageId="20" pageNumber="21" type="holotype">holotype</typeStatus>
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(length/width, mm): body (length to elytral apices) 5.07/2.00; head width 1.17; pronotum l/w 1.22/1.29; PA 1.12; PB 1.07; elytra 3.46/2.00. Habitus as illustrated (
|
||
<figureCitation id="13332A4DFFE8FFEB6553FDD300F9F539" box="[1104,1190,547,574]" captionStart="FIGURE 52" captionStartId="78.[151,250,1963,1987]" captionTargetBox="[239,1343,211,1896]" captionTargetId="figure-28@78.[177,1410,181,1939]" captionTargetPageId="78" captionText="FIGURE 52. Hydrochus colossus n. sp., habitus and female genital segments of holotype." figureDoi="http://doi.org/10.5281/zenodo.5043344" httpUri="https://zenodo.org/record/5043344/files/figure.png" pageId="20" pageNumber="21">Fig. 52</figureCitation>
|
||
). Very large (ca.
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||
<quantity id="4CF09B2DFFE8FFEB6468FDD30498F565" metricMagnitude="-3" metricUnit="m" metricValue="5.07" pageId="20" pageNumber="21" unit="mm" value="5.07">5.07 mm</quantity>
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), elongate. Dorsum grey to black to very faint dull gold, with little iridescence; elytra with numerous black spots. Legs dark brown to black; all surfaces of femora densely coarsely punctate; tibiae with outer apical margin produced and bearing few short stout spines. Pronotum very densely, coarsely punctate. Elytral striae with deep round punctures; interstriae wider than striae.
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</paragraph>
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<paragraph id="8BB736C8FFE8FFEB61C4FD2700C1F411" blockId="20.[151,1437,259,1114]" pageId="20" pageNumber="21">Head densely, coarsely punctate; frons markedly raised in middle, with deep groove on each side of timidity, area between groove and eye also convex; frontoclypeal suture absent; clypeus disc convex.</paragraph>
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<paragraph id="8BB736C8FFE8FFEB61C4FCD007E1F486" blockId="20.[151,1437,259,1114]" pageId="20" pageNumber="21">Pronotum wider than long (L/W as ca. 1.22/1.29), widest at about distal 2/3; anterior margin bisinuate; narrowed at base; sides denticulate, sinuate; punctures on pronotum moderately sized, deep, very dense; depressions very deep, central depression margined laterally with rather sharp ridge.</paragraph>
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<paragraph id="8BB736C8FFE8FFEB61C4FC7B016AF4E9" blockId="20.[151,1437,259,1114]" pageId="20" pageNumber="21">
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||
Elytra with strial punctures very deep and ca. 2–3x those on pronotum; interstriae ca. 2x width of striae; 3
|
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<superScript id="7C7D9B80FFE8FFEB648CFC7B01C2F49E" attach="none" box="[1423,1437,907,921]" fontSize="6" pageId="20" pageNumber="21">rd</superScript>
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||
and 5
|
||
<superScript id="7C7D9B80FFE8FFEB61D6FC5F04BDF4BA" attach="left" box="[213,226,943,957]" fontSize="6" pageId="20" pageNumber="21">th</superScript>
|
||
interstriae raised, costate to subcostate, throughout length, not additionally raised at usual area of callus; 9
|
||
<superScript id="7C7D9B80FFE8FFEB6493FC5F01C2F4BA" attach="right" box="[1424,1437,943,957]" fontSize="6" pageId="20" pageNumber="21">th</superScript>
|
||
interstria raised and overhanging 10
|
||
<superScript id="7C7D9B80FFE8FFEB6329FC230668F4E6" attach="none" box="[554,567,979,993]" fontSize="6" pageId="20" pageNumber="21">th</superScript>
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||
; apices truncate, with submarginal row of large, elongate punctures.
|
||
</paragraph>
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||
<paragraph id="8BB736C8FFE8FFEB61C4FC0707C6F316" blockId="20.[151,1437,259,1114]" box="[199,921,1015,1042]" pageId="20" pageNumber="21">Ventral characters: Mentum and submentum each with 2 foveae.</paragraph>
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<paragraph id="8BB736C8FFE8FFEB61C4FBEB06BDF331" blockId="20.[151,1437,259,1114]" box="[199,738,1051,1078]" pageId="20" pageNumber="21">
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Female genital segments as illustrated (
|
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<figureCitation id="13332A4DFFE8FFEB6382FBEB068AF331" box="[641,725,1051,1078]" captionStart="FIGURE 52" captionStartId="78.[151,250,1963,1987]" captionTargetBox="[239,1343,211,1896]" captionTargetId="figure-28@78.[177,1410,181,1939]" captionTargetPageId="78" captionText="FIGURE 52. Hydrochus colossus n. sp., habitus and female genital segments of holotype." figureDoi="http://doi.org/10.5281/zenodo.5043344" httpUri="https://zenodo.org/record/5043344/files/figure.png" pageId="20" pageNumber="21">Fig. 52</figureCitation>
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).
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</paragraph>
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</subSubSection>
|
||
<subSubSection id="C3126543FFE8FFEB61C4FBCF0737F35E" box="[199,872,1087,1114]" pageId="20" pageNumber="21" type="etymology">
|
||
<paragraph id="8BB736C8FFE8FFEB61C4FBCF0737F35E" blockId="20.[151,1437,259,1114]" box="[199,872,1087,1114]" pageId="20" pageNumber="21">
|
||
<emphasis id="B97CEADAFFE8FFEB61C4FBCF0511F35E" bold="true" box="[199,334,1087,1113]" pageId="20" pageNumber="21">Etymology.</emphasis>
|
||
Named in reference to the very large body size.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |