142 lines
17 KiB
XML
142 lines
17 KiB
XML
<document id="1C21859C7654B1765CA4E2BF22DE7DD9" ID-DOI="10.11646/zootaxa.4994.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="5043261" ID-ZooBank="107FCA64-345F-40A4-99D3-5C1441EEAD93" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1624993528766" checkinUser="plazi" docAuthor="Perkins, Philip D." docDate="2021" docId="03A187DEFFE3FFE06194FC7A00E4F0DA" docLanguage="en" docName="zootaxa.4994.1.1.pdf" docOrigin="Zootaxa 4994 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Hydrochus pronotulus Perkins 2021, new species" docType="treatment" docVersion="4" lastPageNumber="32" masterDocId="FF98FFA6FFFCFFFF6103FFF0045FF707" masterDocTitle="Taxonomy of water beetles in the genus Hydrochus Leach, 1817, from Bolivia, Brazil and Paraguay (Coleoptera: Hydrochidae)" masterLastPageNumber="93" masterPageNumber="1" pageNumber="32" updateTime="1698978858223" updateUser="plazi">
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<mods:title id="5ADF1219848CAA57FFDCE4D19D6FECB0">Taxonomy of water beetles in the genus Hydrochus Leach, 1817, from Bolivia, Brazil and Paraguay (Coleoptera: Hydrochidae)</mods:title>
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<mods:name id="257CB6222B3A26463033531FB5645130" type="personal">
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<mods:roleTerm id="E8E416617FA1BE8E2FAE8E6EB08CCFF9">Author</mods:roleTerm>
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<mods:namePart id="D76C38F82C8A98724C67C4B889849861">Perkins, Philip D.</mods:namePart>
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<mods:typeOfResource id="14E5E080D23005283059D863F2D8DB87">text</mods:typeOfResource>
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<mods:title id="BF8D988BD654FEED02E34601CAE61E9A">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="55851B0BDB24B7901327E447FB514B24">
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<mods:date id="658DE45A37990151E1647C3D4C4EB67E">2021</mods:date>
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<mods:number id="52DAFEBA102B7C0917C3A54497348D50">2021-06-28</mods:number>
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<mods:detail id="A8B226D04060607A579BDBE52BD1A8E8" type="volume">
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<mods:number id="7B47BB35A4EA28CDB6FEADF166ED5D07">4994</mods:number>
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<mods:detail id="D72E748CF2C02558754EE0B2B27C41AA" type="issue">
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<mods:start id="49D1A6136793DAAB6FA9B9D465F3A941">1</mods:start>
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<mods:end id="C062019EB9741B97EB04F1271A9779FA">93</mods:end>
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<mods:classification id="A81FDEBBA1BC7F8282064B06C46E7AF7">journal article</mods:classification>
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<mods:identifier id="A09644E0CEB79F61739C213C418CE7F1" type="DOI">10.11646/zootaxa.4994.1.1</mods:identifier>
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<mods:identifier id="A171D78EF293A6521205017C828E825D" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="18C07F653EB8E78024597CC21CCF6244" type="Zenodo-Dep">5043261</mods:identifier>
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<mods:identifier id="72A745D0A17D9872BA567D4AE3A063D7" type="ZooBank">107FCA64-345F-40A4-99D3-5C1441EEAD93</mods:identifier>
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<treatment id="03A187DEFFE3FFE06194FC7A00E4F0DA" ID-DOI="http://doi.org/10.5281/zenodo.5043974" ID-Zenodo-Dep="5043974" LSID="urn:lsid:plazi:treatment:03A187DEFFE3FFE06194FC7A00E4F0DA" httpUri="http://treatment.plazi.org/id/03A187DEFFE3FFE06194FC7A00E4F0DA" lastPageNumber="32" pageId="31" pageNumber="32">
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<subSubSection id="C3126543FFE3FFE06194FC7A0618F4A3" box="[151,583,906,933]" pageId="31" pageNumber="32" type="nomenclature">
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<paragraph id="8BB736C8FFE3FFE06194FC7A0618F4A3" blockId="31.[151,583,906,969]" box="[151,583,906,933]" pageId="31" pageNumber="32">
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<heading id="D0FF81A4FFE3FFE06194FC7A0618F4A3" bold="true" box="[151,583,906,933]" fontSize="11" level="2" pageId="31" pageNumber="32" reason="4">
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<emphasis id="B97CEADAFFE3FFE06194FC7A0618F4A3" bold="true" box="[151,583,906,933]" pageId="31" pageNumber="32">
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<taxonomicName id="4C084D4BFFE3FFE06194FC7A05F7F4A2" ID-CoL="8Y4R6" authority="Perkins, 2021" authorityName="Perkins" authorityYear="2021" box="[151,424,906,933]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="pronotulus" status="sp. nov.">
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<emphasis id="B97CEADAFFE3FFE06194FC7A05F7F4A2" bold="true" box="[151,424,906,933]" italics="true" pageId="31" pageNumber="32">Hydrochus pronotulus</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="A24F57A1FFE3FFE060B4FC7B0618F4A3" box="[439,583,906,933]" pageId="31" pageNumber="32" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3126543FFE3FFE06194FC5F0568F4CE" box="[151,311,942,969]" pageId="31" pageNumber="32" type="description">
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<paragraph id="8BB736C8FFE3FFE06194FC5F0568F4CE" blockId="31.[151,583,906,969]" box="[151,311,942,969]" pageId="31" pageNumber="32">
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<figureCitation id="13332A4DFFE3FFE06194FC5F04BBF4CE" box="[151,228,943,969]" captionStart="FIGURE 3" captionStartId="8.[151,250,1943,1967]" captionTargetBox="[169,1397,189,1912]" captionTargetId="figure-28@8.[154,1433,181,1919]" captionTargetPageId="8" captionText="FIGURE 3. Dorsal habitus of Hydrochus species in study area." figureDoi="http://doi.org/10.5281/zenodo.5043269" httpUri="https://zenodo.org/record/5043269/files/figure.png" pageId="31" pageNumber="32">Figs. 3</figureCitation>
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,
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<figureCitation id="13332A4DFFE3FFE061F2FC5F0550F4CE" box="[241,271,943,969]" captionStart="FIGURES 18–19" captionStartId="61.[151,264,1944,1968]" captionTargetBox="[170,1425,205,1900]" captionTargetId="figure-15@61.[151,1435,181,1920]" captionTargetPageId="61" captionText="FIGURES 18–19. Habitus and male genitalia of holotypes. 18. Hydrochus pronotulus n. sp. 19. Hydrochus velatus n. sp." figureDoi="http://doi.org/10.5281/zenodo.5043296" httpUri="https://zenodo.org/record/5043296/files/figure.png" pageId="31" pageNumber="32">18</figureCitation>
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,
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<figureCitation id="13332A4DFFE3FFE06018FC5E0568F4CE" box="[283,311,942,969]" captionStart="FIGURES 55–56" captionStartId="80.[151,264,1943,1967]" captionTargetBox="[165,1422,181,1920]" captionTargetId="figure-28@80.[165,1422,181,1920]" captionTargetPageId="80" captionText="FIGURES 55–56. Geographical distributions. 55. Endemics and near endemics. 56. H. bicarinatus and H. merkli." figureDoi="http://doi.org/10.5281/zenodo.5043352" httpUri="https://zenodo.org/record/5043352/files/figure.png" pageId="31" pageNumber="32">55</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3126543FFE3FFE06194FC0705D7F332" pageId="31" pageNumber="32" type="materials_examined">
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<paragraph id="8BB736C8FFE3FFE06194FC0705D7F332" blockId="31.[151,1437,1014,2013]" pageId="31" pageNumber="32">
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<emphasis id="B97CEADAFFE3FFE06194FC070515F317" bold="true" box="[151,330,1014,1041]" pageId="31" pageNumber="32">Type Material.</emphasis>
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<materialsCitation id="3B603C95FFE3FFE06050FC0705D7F332" collectingDate="1956-05-01" collectionCode="NHMLAC" collectorName="N. Sao Joao da Alianca & F. S. Truxal" country="Brazil" location="Sao Joao da Alianca" pageId="31" pageNumber="32" specimenCount="1" stateProvince="Goias" typeStatus="holotype">
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<emphasis id="B97CEADAFFE3FFE06050FC070641F317" bold="true" box="[339,542,1014,1041]" pageId="31" pageNumber="32">
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<typeStatus id="54B3886AFFE3FFE06050FC07059FF316" box="[339,448,1015,1041]" pageId="31" pageNumber="32" type="holotype">Holotype</typeStatus>
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(male):
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</emphasis>
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<collectingCountry id="F31F7658FFE3FFE06324FC07062FF316" box="[551,624,1015,1041]" name="Brazil" pageId="31" pageNumber="32">Brazil</collectingCountry>
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:
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<collectingRegion id="49CCF82AFFE3FFE0637FFC07069FF316" box="[636,704,1015,1041]" country="Brazil" name="Goias" pageId="31" pageNumber="32">Goias</collectingRegion>
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, “20 kil
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<collectorName id="26FD531EFFE3FFE06226FC07006CF316" box="[805,1075,1015,1041]" pageId="31" pageNumber="32">
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N.
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<location id="8ED76013FFE3FFE0624AFC07006CF316" LSID="urn:lsid:plazi:treatment:03A187DEFFE3FFE06194FC7A00E4F0DA:8ED76013FFE3FFE0624AFC07006CF316" box="[841,1075,1015,1041]" country="Brazil" name="Sao Joao da Alianca" pageId="31" pageNumber="32" stateProvince="Goias">Sao Joao da Alianca</location>
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</collectorName>
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, Go.,
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<collectingCountry id="F31F7658FFE3FFE0657AFC07009FF316" box="[1145,1216,1015,1041]" name="Brazil" pageId="31" pageNumber="32">Brazil</collectingCountry>
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,
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<date id="FFB61008FFE3FFE065CDFC070103F316" box="[1230,1372,1014,1041]" pageId="31" pageNumber="32" value="1956-05-01">
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<collectingDate id="EFF2E9E0FFE3FFE065CDFC070103F316" box="[1230,1372,1014,1041]" pageId="31" pageNumber="32" value="1956-05-01">May 1 1956</collectingDate>
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</date>
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,
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<collectorName id="26FD531EFFE3FFE0646BFC0704BBF332" pageId="31" pageNumber="32">F. S. Truxal</collectorName>
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” (
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<collectionCode id="ED19AE0DFFE3FFE061FFFBEB0523F332" box="[252,380,1051,1077]" pageId="31" pageNumber="32">NHMLAC</collectionCode>
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).
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3126543FFE3FFE061C4FBCE069BF3A5" pageId="31" pageNumber="32" type="diagnosis">
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<paragraph id="8BB736C8FFE3FFE061C4FBCE069BF3A5" blockId="31.[151,1437,1014,2013]" pageId="31" pageNumber="32">
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<emphasis id="B97CEADAFFE3FFE061C4FBCE058BF35F" bold="true" box="[199,468,1086,1112]" pageId="31" pageNumber="32">Differential Diagnosis.</emphasis>
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Among Brazilian species, recognized by the combination of moderate size (ca.
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<quantity id="4CF09B2DFFE3FFE06468FBCF0498F37A" metricMagnitude="-3" metricUnit="m" metricValue="3.44" pageId="31" pageNumber="32" unit="mm" value="3.44">3.44 mm</quantity>
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), the black dorsal color, the small pronotum compared to the size of the elytra, the deep pronotal depressions, and the male genitalia (
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<figureCitation id="13332A4DFFE3FFE0609EFB7705AFF3A6" box="[413,496,1159,1185]" captionStart="FIGURES 18–19" captionStartId="61.[151,264,1944,1968]" captionTargetBox="[170,1425,205,1900]" captionTargetId="figure-15@61.[151,1435,181,1920]" captionTargetPageId="61" captionText="FIGURES 18–19. Habitus and male genitalia of holotypes. 18. Hydrochus pronotulus n. sp. 19. Hydrochus velatus n. sp." figureDoi="http://doi.org/10.5281/zenodo.5043296" httpUri="https://zenodo.org/record/5043296/files/figure.png" pageId="31" pageNumber="32">Fig. 18</figureCitation>
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; described below).
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</paragraph>
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</subSubSection>
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<subSubSection id="C3126543FFE3FFE061C4FB5A0106F0BE" pageId="31" pageNumber="32" type="description">
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<paragraph id="8BB736C8FFE3FFE061C4FB5A00B4F27E" blockId="31.[151,1437,1014,2013]" pageId="31" pageNumber="32">
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<emphasis id="B97CEADAFFE3FFE061C4FB5A050EF3C3" bold="true" box="[199,337,1194,1220]" pageId="31" pageNumber="32">Description</emphasis>
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. Size:
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<typeStatus id="54B3886AFFE3FFE060A7FB5B0659F3C2" box="[420,518,1195,1221]" pageId="31" pageNumber="32" type="holotype">holotype</typeStatus>
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(length/width, mm): body (length to elytral apices) 3.44/1.27; head width 0.80; pronotum l/w 0.71/0.75; PA 0.75; PB 0.63; elytra 2.24/1.27. Habitus as illustrated (
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<figureCitation id="13332A4DFFE3FFE06528FB3F0021F3EE" box="[1067,1150,1231,1257]" captionStart="FIGURES 18–19" captionStartId="61.[151,264,1944,1968]" captionTargetBox="[170,1425,205,1900]" captionTargetId="figure-15@61.[151,1435,181,1920]" captionTargetPageId="61" captionText="FIGURES 18–19. Habitus and male genitalia of holotypes. 18. Hydrochus pronotulus n. sp. 19. Hydrochus velatus n. sp." figureDoi="http://doi.org/10.5281/zenodo.5043296" httpUri="https://zenodo.org/record/5043296/files/figure.png" pageId="31" pageNumber="32">Fig. 18</figureCitation>
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). Body size moderate (ca.
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<quantity id="4CF09B2DFFE3FFE06194FB03055FF20A" box="[151,256,1267,1293]" metricMagnitude="-3" metricUnit="m" metricValue="3.44" pageId="31" pageNumber="32" unit="mm" value="3.44">3.44 mm</quantity>
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). Dorsum black, with slight or no iridescence, elytra without black spots. Legs brown to testaceous, with tibiofemoral joints darker. Punctation on elytra ca. 2–3x that of pronotum. Elytra interstriae ca. 0.5–1x strial puncture diameter. Interstria 9
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<superScript id="7C7D9B80FFE3FFE060ACFACA05E3F24F" attach="left" box="[431,444,1338,1352]" fontSize="6" pageId="31" pageNumber="32">th</superScript>
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more convex than others, overhanging 10
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<superScript id="7C7D9B80FFE3FFE0628EFACA07C5F24F" attach="left" box="[909,922,1338,1352]" fontSize="6" pageId="31" pageNumber="32">th</superScript>
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interstria. Usual area of callus on 5
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<superScript id="7C7D9B80FFE3FFE06423FACA0172F24F" attach="left" box="[1312,1325,1338,1352]" fontSize="6" pageId="31" pageNumber="32">th</superScript>
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small, ca. 3 punctures on each side. Another, very small callus on 7
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<superScript id="7C7D9B80FFE3FFE06216FAAE077DF26B" attach="none" box="[789,802,1374,1388]" fontSize="6" pageId="31" pageNumber="32">th</superScript>
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, located slightly anterior to callus on 5
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<superScript id="7C7D9B80FFE3FFE065D4FAAE00BBF26B" attach="none" box="[1239,1252,1374,1388]" fontSize="6" pageId="31" pageNumber="32">th</superScript>
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.
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</paragraph>
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<paragraph id="8BB736C8FFE3FFE061C4FA730553F2E2" blockId="31.[151,1437,1014,2013]" pageId="31" pageNumber="32">Pronotum very slightly wider than long (ratio ca. 42/40), widest at anterior margin, narrowed at base, sides sinuate, smooth; anterior margin slightly bisinuate; depressions moderately deep, punctation rather uniform over pronotum.</paragraph>
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<paragraph id="8BB736C8FFE3FFE061C4FA1F0052F12A" blockId="31.[151,1437,1014,2013]" pageId="31" pageNumber="32">Elytra with spaces between strial punctures narrow walls to ca. 0.5x their diameter; apices rather sharply conjointly rounded in dorsal view, in lateral view outer margin without angulation.</paragraph>
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<paragraph id="8BB736C8FFE3FFE061C4F9C70151F156" blockId="31.[151,1437,1014,2013]" box="[199,1294,1591,1617]" pageId="31" pageNumber="32">Ventral characters: Mentum punctate, with central depression; submentum with two small foveae.</paragraph>
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<paragraph id="8BB736C8FFE3FFE061C4F9AB0791F076" blockId="31.[151,1437,1014,2013]" pageId="31" pageNumber="32">
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Male genitalia (
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<figureCitation id="13332A4DFFE3FFE06074F9AB0596F172" box="[375,457,1627,1653]" captionStart="FIGURES 18–19" captionStartId="61.[151,264,1944,1968]" captionTargetBox="[170,1425,205,1900]" captionTargetId="figure-15@61.[151,1435,181,1920]" captionTargetPageId="61" captionText="FIGURES 18–19. Habitus and male genitalia of holotypes. 18. Hydrochus pronotulus n. sp. 19. Hydrochus velatus n. sp." figureDoi="http://doi.org/10.5281/zenodo.5043296" httpUri="https://zenodo.org/record/5043296/files/figure.png" pageId="31" pageNumber="32">Fig. 18</figureCitation>
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) general characters: genitalia moderately elongate, length ca. 4.8x greatest width, basal piece shorter than parameres (ratio ca. 1/1.5); parameres in dorsal/ventral views very narrow; paramere tips and aedeagus tip at same level, paramere tips arcing toward midline, tips widely separated, plsa absent, lateral margins of tips arcuate; in lateral view parameres narrow, very slightly sinuate, apex rounded; pseudoparameres wide in proximal 2/3, much narrower and tapering to pointed apex in distal 1/3, where medial margin with serrations; aedeagus in dorsal/ventral views very wide in distal 2/3, oval, much narrower in proximal 1/3, agw subapical; basal piece in dorsal/ventral views tapering from distal end to base, in lateral view slightly arcuate, greatest width in dorsal/ventral views greater than lateral width, orifice large, very slightly asymmetrical.
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</paragraph>
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<paragraph id="8BB736C8FFE3FFE061C4F88B00D1F092" blockId="31.[151,1437,1014,2013]" box="[199,1166,1915,1941]" pageId="31" pageNumber="32">Dorsal surface: adtl at ca. proximal 1/3; pdmm widely separated from alm; adbl small.</paragraph>
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<paragraph id="8BB736C8FFE3FFE061C4F86F0106F0BE" blockId="31.[151,1437,1014,2013]" box="[199,1369,1951,1977]" pageId="31" pageNumber="32">Ventral surface: avtl not apparent; pvmm similar to pdmm except at tips where parameres slightly wider.</paragraph>
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</subSubSection>
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<subSubSection id="C3126543FFE3FFE061C4F83300E4F0DA" box="[199,1211,1987,2013]" pageId="31" pageNumber="32" type="etymology">
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<paragraph id="8BB736C8FFE3FFE061C4F83300E4F0DA" blockId="31.[151,1437,1014,2013]" box="[199,1211,1987,2013]" pageId="31" pageNumber="32">
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<emphasis id="B97CEADAFFE3FFE061C4F8330517F0DA" bold="true" box="[199,328,1987,2013]" pageId="31" pageNumber="32">Etymology</emphasis>
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. Named in reference to the small pronotum, compared to the size of the elytra.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |