treatments-xml/data/03/A1/87/03A187CAFFDD943D1FEAFE38904AFAD8.xml

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<mods:title id="23B393EEAD46272F074842F3B803E182">On the taxonomy and zoogeography of some West Palaearctic Quedius species, with a focus on the East Mediterranean and the species allied to Quedius umbrinus and Q. nivicola (Coleoptera: Staphylinidae: Staphylininae)</mods:title>
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<mods:date id="2B0CDBF80C30B81CDC700D675F337832">2018</mods:date>
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<treatment id="03A187CAFFDD943D1FEAFE38904AFAD8" ID-DOI="http://doi.org/10.5281/zenodo.4334237" ID-GBIF-Taxon="167288458" ID-Zenodo-Dep="4334237" LSID="urn:lsid:plazi:treatment:03A187CAFFDD943D1FEAFE38904AFAD8" httpUri="http://treatment.plazi.org/id/03A187CAFFDD943D1FEAFE38904AFAD8" lastPageId="8" lastPageNumber="157" pageId="5" pageNumber="154">
<subSubSection id="C3126557FFDD94301FEAFE38913AFDBD" box="[178,893,496,519]" pageId="5" pageNumber="154" type="nomenclature">
<paragraph id="8BB736DCFFDD94301FEAFE38913AFDBD" blockId="5.[178,893,496,519]" box="[178,893,496,519]" pageId="5" pageNumber="154">
<heading id="D0FF81B0FFDD94301FEAFE38913AFDBD" box="[178,893,496,519]" fontSize="8" level="2" pageId="5" pageNumber="154" reason="2">
<emphasis id="B97CEACEFFDD94301FEAFE38903EFDBF" bold="true" box="[178,633,496,518]" pageId="5" pageNumber="154">
<taxonomicName id="4C084D5FFFDD94301FEAFE389049FDBC" ID-CoL="4R35C" authorityName="Coiffait" authorityYear="1963" box="[178,526,496,518]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="154" phylum="Arthropoda" rank="species" species="hermonensis" subGenus="Raphirus">
<emphasis id="B97CEACEFFDD94301FEAFE38934EFDBC" bold="true" box="[178,265,496,518]" italics="true" pageId="5" pageNumber="154">Quedius</emphasis>
(
<emphasis id="B97CEACEFFDD94301E40FE38933EFDBC" bold="true" box="[280,377,496,518]" italics="true" pageId="5" pageNumber="154">Raphirus</emphasis>
) 
<emphasis id="B97CEACEFFDD94301ED0FE389049FDBC" bold="true" box="[392,526,496,518]" italics="true" pageId="5" pageNumber="154">hermonensis</emphasis>
</taxonomicName>
<bibRefCitation id="EF994B2DFFDD94301D4CFE38903EFDBF" author="COIFFAIT H" box="[532,633,496,518]" pageId="5" pageNumber="154" pagination="372 - 420" refId="ref20207" refString="COIFFAIT H. (1963): Les Quedius du sous-genre Sauridus de la region palearctique occidentale (avec description de fomes nouvelles). - Bulletin d e la Societe d'Histoire Naturelle de Toulouse 98 (3 - 4): 372 - 420." type="journal article" year="1963">COIFFAIT</bibRefCitation>
</emphasis>
, 
<emphasis id="B97CEACEFFDD94301DDDFE3890FFFDBF" bold="true" box="[645,696,496,517]" pageId="5" pageNumber="154">1963</emphasis>
(
<figureCitation id="13332A59FFDD94301D9DFE38916EFDBC" box="[709,809,496,518]" captionStart-0="Figs 1-12" captionStart-1="Figs 13-24" captionStartId-0="6.[178,218,1506,1528]" captionStartId-1="7.[178,217,1506,1528]" captionTargetBox-0="[182,1076,226,1483]" captionTargetBox-1="[187,1079,217,1465]" captionTargetId-0="figure@6.[178,1085,212,1497]" captionTargetId-1="figure@7.[178,1085,212,1497]" captionTargetPageId-1="7" captionText-0="Figs 1-12: Quedius hermonensis from Samos (1-3), Israel (4-6), and Cyprus (7-12): (1, 4, 7, 10) aedeagus in lateral view; (2, 5, 8, 11) median lobe of aedeagus in ventral view; (3, 6, 9, 12) paramere. Scale bar: 0.5 mm." captionText-1="Figs 13-24: Quedius hermonensis from Cyprus (13-14), Q. orientalis (15-17), and Q. microcapillatus (18-24) from Iraq (18-20), Turkey (21), and Iran (22-24): (13, 15, 22) median lobe of aedeagus in lateral view; (14, 17, 20, 24) paramere; (16, 19, 23) median lobe of aedeagus in ventral view; (18, 21) aedeagus in lateral view. Scale bar: 0.5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.3995001" figureDoi-1="http://doi.org/10.5281/zenodo.3985370" httpUri-0="https://zenodo.org/record/3995001/files/figure.png" httpUri-1="https://zenodo.org/record/3985370/files/figure.png" pageId="5" pageNumber="154">Figs 1-14</figureCitation>
, 
<figureCitation id="13332A59FFDD94301C6CFE399134FDBD" box="[820,883,497,519]" captionStart="Map 1" captionStartId="9.[178,223,1266,1287]" captionTargetBox="[178,1085,639,1254]" captionTargetId="figure@9.[178,1085,639,1254]" captionTargetPageId="9" captionText="Map 1: Distributions of Quedius hermonensis (black circles), Q. microcapillatus (white circles), and Q. orientalis (black star), based on revised records." figureDoi="http://doi.org/10.5281/zenodo.3985374" httpUri="https://zenodo.org/record/3985374/files/figure.png" pageId="5" pageNumber="154">Map 1</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3126557FFDD94301FEAFDD4966CFD08" pageId="5" pageNumber="154" type="reference_group">
<paragraph id="8BB736DCFFDD94301FEAFDD4900BFD88" blockId="5.[178,1087,540,966]" box="[178,588,540,562]" pageId="5" pageNumber="154">
<treatmentCitationGroup id="AB1811F2FFDD94301FEAFDD4900BFD88" box="[178,588,540,562]" pageId="5" pageNumber="154">
<treatmentCitation id="0AA910CDFFDD94301FEAFDD4907FFD8B" author="COIFFAIT H" box="[178,568,540,562]" page="417" pageId="5" pageNumber="154" year="1963">
<taxonomicName id="4C084D5FFFDD94301FEAFDD4907FFD8B" ID-CoL="4R35C" authority="COIFFAIT, 1963: 417" authorityName="Coiffait" authorityYear="1963" box="[178,568,540,562]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="154" phylum="Arthropoda" rank="species" species="hermonensis">
<emphasis id="B97CEACEFFDD94301FEAFDD49332FD8B" box="[178,373,540,561]" italics="true" pageId="5" pageNumber="154">Quedius hermonensis</emphasis>
<bibRefCitation id="EF994B2DFFDD94301E23FDD4907FFD8B" author="COIFFAIT H" box="[379,568,540,562]" pageId="5" pageNumber="154" pagination="372 - 420" refId="ref20207" refString="COIFFAIT H. (1963): Les Quedius du sous-genre Sauridus de la region palearctique occidentale (avec description de fomes nouvelles). - Bulletin d e la Societe d'Histoire Naturelle de Toulouse 98 (3 - 4): 372 - 420." type="journal article" year="1963">
COIFFAIT, 1963: 
<specimenCount id="9D0EFD55FFDD94301D4EFDD4907FFD8B" box="[534,568,540,561]" pageId="5" pageNumber="154" type="generic">417</specimenCount>
</bibRefCitation>
</taxonomicName>
</treatmentCitation>
f.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BB736DCFFDD94301FEAFDFE90D5FDF1" blockId="5.[178,1087,540,966]" box="[178,658,566,588]" pageId="5" pageNumber="154">
<treatmentCitationGroup id="AB1811F2FFDD94301FEAFDFE90D5FDF1" box="[178,658,566,588]" pageId="5" pageNumber="154">
<emphasis id="B97CEACEFFDD94301FEAFDFE9333FDF1" box="[178,372,566,587]" italics="true" pageId="5" pageNumber="154">
<taxonomicName id="4C084D5FFFDD94301FEAFDFE92B9FDF1" ID-CoL="8VS2M" box="[178,254,566,587]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="177" phylum="Arthropoda" rank="genus">Quedius</taxonomicName>
coiffaitianus
</emphasis>
<treatmentCitation id="0AA910CDFFDD94301E22FDFE905AFDF1" author="FAGEL G" box="[378,541,566,588]" page="11" pageId="5" pageNumber="154" year="1968">
<bibRefCitation id="EF994B2DFFDD94301E22FDFE905AFDF1" author="FAGEL G" box="[378,541,566,588]" pageId="5" pageNumber="154" pagination="1 - 13" refId="ref20496" refString="FAGEL G. (1968 a): Contribution a la connaissance des Staphylinidae CI. - Remarques diverses sur quelques Quedius de Mediterranee orientale. - B ulletin Institut Royale des Sciences Naturelles de Belgique 44 (12): 1 - 13." type="journal article" year="1968">FAGEL, 1968a: 11</bibRefCitation>
</treatmentCitation>
ff.; 
<emphasis id="B97CEACEFFDD94301D1BFDFE90D5FDF1" bold="true" box="[579,658,566,587]" pageId="5" pageNumber="154">nov.syn.</emphasis>
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BB736DCFFDD94301FEAFD9890D0FDDF" blockId="5.[178,1087,540,966]" box="[178,663,592,614]" pageId="5" pageNumber="154">
<treatmentCitationGroup id="AB1811F2FFDD94301FEAFD9890D0FDDF" box="[178,663,592,614]" pageId="5" pageNumber="154">
<emphasis id="B97CEACEFFDD94301FEAFD98933FFDDC" box="[178,376,592,614]" italics="true" pageId="5" pageNumber="154">
<taxonomicName id="4C084D5FFFDD94301FEAFD9892B9FDDF" ID-CoL="8VS2M" box="[178,254,592,613]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="177" phylum="Arthropoda" rank="genus">Quedius</taxonomicName>
rugosipennis
</emphasis>
<treatmentCitation id="0AA910CDFFDD94301E26FD989065FDDC" author="FAGEL G" box="[382,546,592,614]" page="115" pageId="5" pageNumber="154" year="1969">
<bibRefCitation id="EF994B2DFFDD94301E26FD989065FDDC" author="FAGEL G" box="[382,546,592,614]" pageId="5" pageNumber="154" pagination="103 - 125" refId="ref20632" refString="FAGEL G. (1969): Contribution a la connaissance des Staphylinidae CVIII. - Sur quelques especes inedites provenant du bassin mediterraneen. - Bulletin et Annales de la Societe Royale d'Entomologie de Belgique 105: 103 - 125." type="journal article" year="1969">FAGEL, 1969: 115</bibRefCitation>
</treatmentCitation>
ff.; 
<emphasis id="B97CEACEFFDD94301D10FD9890D0FDDF" bold="true" box="[584,663,592,613]" pageId="5" pageNumber="154">nov.syn.</emphasis>
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BB736DCFFDD94301FEAFDA7966CFD08" blockId="5.[178,1087,540,966]" pageId="5" pageNumber="154">
T y p e m a t e r i a l e x a m i n e d: 
<emphasis id="B97CEACEFFDD94301D0CFDA790A4FD3E" box="[596,739,623,644]" italics="true" pageId="5" pageNumber="154">Q. coiffaitianus</emphasis>
: 
<typeStatus id="54B3887EFFDD94301DA8FDA7910BFD3E" box="[752,844,623,644]" pageId="5" pageNumber="154" type="paratype">Paratypes</typeStatus>
: 
<specimenCount id="9D0EFD55FFDD94301C0EFDA791DBFD3E" box="[854,924,623,645]" pageId="5" pageNumber="154" type="generic">15 exs.</specimenCount>
: &quot; 
<collectingCountry id="F31F764CFFDD94301CF5FDA791B0FD3E" box="[941,1015,623,644]" name="Lebanon" pageId="5" pageNumber="154">LIBAN</collectingCountry>
: Nabeh Safa, mousses gorges d'eau [
<specimenCount id="9D0EFD55FFDD94301EE3FD4E9062FD26" box="[443,549,646,668]" pageId="5" pageNumber="154" type="generic">1 specimen</specimenCount>
: &quot;bois marécageux&quot;], 
<quantity id="4CF09B39FFDD94301DB7FD4E9170FD21" box="[751,823,646,667]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="154" unit="m" value="1000.0">1000 m</quantity>
, 
<date id="FFB6101CFFDD94301C19FD4E91C3FD26" box="[833,900,646,668]" pageId="5" pageNumber="154" value="1966-05">V.1966</date>
G. Fagel / G. Fagel det., 
<emphasis id="B97CEACEFFDD94301F87FD549317FD0B" box="[223,336,668,689]" italics="true" pageId="5" pageNumber="154">coiffaitianus</emphasis>
n.sp. / 
<typeStatus id="54B3887EFFDD94301ECAFD5593A6FD08" box="[402,481,669,690]" pageId="5" pageNumber="154" type="paratype">Paratype</typeStatus>
/ 
<taxonomicName id="4C084D5FFFDD94301EABFD549142FD08" ID-CoL="4R35C" authority="Coiffait" authorityName="Coiffait" authorityYear="1963" box="[499,773,668,690]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="154" phylum="Arthropoda" rank="species" species="hermonensis">
<emphasis id="B97CEACEFFDD94301EABFD5490F1FD0B" box="[499,694,668,689]" italics="true" pageId="5" pageNumber="154">Quedius hermonensis</emphasis>
Coiffait
</taxonomicName>
, det. V. 
<bibRefCitation id="EF994B2DFFDD94301C09FD549182FD0B" author="ASSING V &amp; SCHULKE M &amp; BRACHAT V" box="[849,965,668,690]" pageId="5" pageNumber="154" pagination="1" refId="ref19985" refString="ASSING V., SCHULKE M., BRACHAT V. &amp; H. MEYBOHM (in press): On the Staphylinidae of the Greek island Corfu (Insecta: Coleoptera). - Contributions to En tomology 68 (1) (2018)." type="journal article" year="2018">Assing 2018</bibRefCitation>
&quot; (IRSNB).
</paragraph>
</subSubSection>
<subSubSection id="C3126557FFDD943D1FEAFD73904AFAD8" lastPageId="8" lastPageNumber="157" pageId="5" pageNumber="154" type="description">
<paragraph id="8BB736DCFFDD94301FEAFD7391F9FD44" blockId="5.[178,1087,540,966]" pageId="5" pageNumber="154">
<emphasis id="B97CEACEFFDD94301FEAFD739302FD6B" box="[178,325,699,721]" italics="true" pageId="5" pageNumber="154">Q. rugosipennis</emphasis>
: 
<materialsCitation id="3B603C81FFDD94301E0BFD73938DFD44" ID-GBIF-Occurrence="2846796350" collectingDate="1968-05" collectionCode="IRSNB" collectorName="G. Fagel" determinerName="G. Fagel" location="Anatolie" pageId="5" pageNumber="154" specimenCount="1" specimenCount-male="1" stateProvince="Antalya" typeStatus="holotype">
<typeStatus id="54B3887EFFDD94301E0BFD7393EEFD6B" box="[339,425,699,721]" pageId="5" pageNumber="154" type="holotype">Holotype</typeStatus>
<specimenCount id="9D0EFD55FFDD94301EF1FD7393FFFD6B" box="[425,440,699,721]" pageId="5" pageNumber="154" type="male">♂</specimenCount>
[dissected prior to present study; aedeagus missing]: &quot; 
<location id="8ED76007FFDD94301CEEFD739641FD6B" LSID="urn:lsid:plazi:treatment:03A187CAFFDD943D1FEAFE38904AFAD8:8ED76007FFDD94301CEEFD739641FD6B" box="[950,1030,699,721]" name="Anatolie" pageId="5" pageNumber="154" stateProvince="Antalya">Anatolie</location>
mér., 
<collectingRegion id="49CCF83EFFDD94301FEAFD1A92BBFD52" box="[178,252,722,744]" country="Turkey" name="Antalya" pageId="5" pageNumber="154">Antalya</collectingRegion>
, 
<date id="FFB6101CFFDD94301E5DFD1A930FFD52" box="[261,328,722,744]" pageId="5" pageNumber="154" value="1968-05">
<collectingDate id="EFF2E9F4FFDD94301E5DFD1A930FFD52" box="[261,328,722,744]" pageId="5" pageNumber="154" value="1968-05">V.1968</collectingDate>
</date>
<collectorName id="26FD530AFFDD94301E16FD1A93DBFD52" box="[334,412,722,744]" pageId="5" pageNumber="154">G. Fagel</collectorName>
/ 
<determinerName id="7E6BDA18FFDD94301EF6FD1A93BBFD52" box="[430,508,722,744]" pageId="5" pageNumber="154">G. Fagel</determinerName>
det., 
<emphasis id="B97CEACEFFDD94301D77FD1A90E3FD5D" box="[559,676,722,743]" italics="true" pageId="5" pageNumber="154">rugosipennis</emphasis>
n.sp. / 
<typeStatus id="54B3887EFFDD94301DBFFD1A9151FD5D" box="[743,790,722,743]" pageId="5" pageNumber="154">Type</typeStatus>
/ 
<taxonomicName id="4C084D5FFFDD94301C7FFD1A967DFD52" authority="Coiffait" authorityName="Coiffait" authorityYear="1963" box="[807,1082,722,744]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="154" phylum="Arthropoda" rank="species" species="hermonensis">
<emphasis id="B97CEACEFFDD94301C7FFD1A91ACFD5D" box="[807,1003,722,743]" italics="true" pageId="5" pageNumber="154">Quedius hermonensis</emphasis>
Coiffait
</taxonomicName>
, det. V. 
<bibRefCitation id="EF994B2DFFDD94301FADFD20932EFD47" author="ASSING V &amp; SCHULKE M &amp; BRACHAT V" box="[245,361,744,766]" pageId="5" pageNumber="154" pagination="1" refId="ref19985" refString="ASSING V., SCHULKE M., BRACHAT V. &amp; H. MEYBOHM (in press): On the Staphylinidae of the Greek island Corfu (Insecta: Coleoptera). - Contributions to En tomology 68 (1) (2018)." type="journal article" year="2018">Assing 2018</bibRefCitation>
&quot; (
<collectionCode id="ED19AE19FFDD94301E24FD209383FD44" box="[380,452,744,766]" collectionName="Belgium, Brussels, Institut Royal des Sciences Naturelles de Belgique" country="Belgium" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="5" pageNumber="154">IRSNB</collectionCode>
)
</materialsCitation>
. 
<typeStatus id="54B3887EFFDD94301E8BFD219068FD44" box="[467,559,745,766]" pageId="5" pageNumber="154" type="paratype">Paratypes</typeStatus>
: 
<specimenCount id="9D0EFD55FFDD94301D6FFD209013FD47" box="[567,596,744,765]" pageId="5" pageNumber="154" type="male">1♂</specimenCount>
, 
<specimenCount id="9D0EFD55FFDD94301D07FD2090CBFCBA" box="[607,652,744,768]" pageId="5" pageNumber="154" type="female">3♀♀</specimenCount>
: same data as 
<typeStatus id="54B3887EFFDD94301C48FD209118FD44" box="[784,863,744,766]" pageId="5" pageNumber="154" type="holotype">holotype</typeStatus>
(IRSNB).
</paragraph>
<paragraph id="8BB736DCFFDD94301FEAFCC09132FC7C" blockId="5.[178,1087,540,966]" pageId="5" pageNumber="154">
<materialsCitation id="3B603C81FFDD94301FEAFCC09679FC8C" ID-GBIF-Occurrence="2846796312" collectingDate="2008-04-24" collectingDateMax="2008-04-24" collectingDateMin="2005-04-19" collectorName="Brachat &amp; Meybohm" country="Turkey" county="Gazipasa" elevation="40" latitude="32.883335" location="Kalandere" longLatPrecision="1208" longitude="35.633335" municipality="Kalandere" pageId="5" pageNumber="154" specimenCount="2" specimenCount-male="2" stateProvince="Antalya">
A d d i t i o n a l m a t e r i a l e x a m i n e d: 
<collectingCountry id="F31F764CFFDD94301DEEFCCF9145FCA7" box="[694,770,775,797]" name="Turkey" pageId="5" pageNumber="154">Turkey</collectingCountry>
: 
<specimenCount id="9D0EFD55FFDD94301C5DFCCF9173FCA7" box="[773,820,775,797]" pageId="5" pageNumber="154" type="male">2♂♂</specimenCount>
[1teneral], 
<collectingRegion id="49CCF83EFFDD94301CC9FCCF91A4FCA7" box="[913,995,775,797]" country="Turkey" name="Antalya" pageId="5" pageNumber="154">Antalya</collectingRegion>
, Anamur- 
<collectingCounty id="62D64E50FFDD94301F87FCE89376FC8C" box="[223,305,800,822]" pageId="5" pageNumber="154">Gazipaşa</collectingCounty>
, 
<collectingMunicipality id="6BD3ACA6FFDD94301E66FCE893DDFC8C" box="[318,410,800,822]" pageId="5" pageNumber="154">Kalandere</collectingMunicipality>
, 
<geoCoordinate id="EE3C501BFFDD94301EF0FCE893B5FC8C" box="[424,498,800,822]" degrees="36" direction="north" minutes="07" orientation="latitude" pageId="5" pageNumber="154" precision="925" value="36.116665">36°07'N</geoCoordinate>
, 
<geoCoordinate id="EE3C501BFFDD94301EA7FCE89000FC8F" box="[511,583,800,821]" degrees="32" direction="east" minutes="34" orientation="longitude" pageId="5" pageNumber="154" precision="925" value="32.566666">32°34'E</geoCoordinate>
, 
<quantity id="4CF09B39FFDD94301D0CFCE890C0FC8F" box="[596,647,800,821]" metricMagnitude="1" metricUnit="m" metricValue="4.0" pageId="5" pageNumber="154" unit="m" value="40.0">
<elevation id="0025D1EFFFDD94301D0CFCE890C0FC8F" box="[596,647,800,821]" metricMagnitude="1" metricUnit="m" metricValue="4.0" pageId="5" pageNumber="154" unit="m" value="40.0">40 m</elevation>
</quantity>
, 
<date id="FFB6101CFFDD94301DCDFCE890BEFC8F" box="[661,761,800,821]" pageId="5" pageNumber="154" value="2008-04-24">
<collectingDate id="EFF2E9F4FFDD94301DCDFCE890BEFC8F" box="[661,761,800,821]" pageId="5" pageNumber="154" value="2008-04-24">24.IV.2008</collectingDate>
</date>
, leg. 
<collectorName id="26FD530AFFDD94301C69FCE89132FC8C" box="[817,885,800,822]" pageId="5" pageNumber="154">Brachat</collectorName>
&amp; 
<collectorName id="26FD530AFFDD94301CC2FCE891B3FC8C" box="[922,1012,800,822]" pageId="5" pageNumber="154">Meybohm</collectorName>
(cAss).
</materialsCitation>
<materialsCitation id="3B603C81FFDD94301F87FCF091D8FCF4" ID-GBIF-Occurrence="2846796348" box="[223,927,824,846]" collectingDate="2005-04-19" collectingDateMax="2008-04-24" collectingDateMin="2005-04-19" collectorName="Brachat &amp; Meybohm &amp; Lompe &amp; Assmann" country="Turkey" county="Gazipasa" elevation="40" latitude="32.883335" location="Troodos" longLatPrecision="1208" longitude="35.633335" municipality="Kalandere" pageId="5" pageNumber="154" specimenCount="4" specimenCount-male="4" stateProvince="Antalya">
<collectingCountry id="F31F764CFFDD94301F87FCF09366FCF7" box="[223,289,824,845]" name="Cyprus" pageId="5" pageNumber="154">Cyprus</collectingCountry>
: 
<specimenCount id="9D0EFD55FFDD94301E72FCF09300FCF7" box="[298,327,824,845]" pageId="5" pageNumber="154" type="male">1♂</specimenCount>
, 
<location id="8ED76007FFDD94301E08FCF093DCFCF4" LSID="urn:lsid:plazi:treatment:03A187CAFFDD943D1FEAFE38904AFAD8:8ED76007FFDD94301E08FCF093DCFCF4" box="[336,411,824,846]" country="Turkey" county="Gazipasa" latitude="32.883335" longLatPrecision="1208" longitude="35.633335" municipality="Kalandere" name="Troodos" pageId="5" pageNumber="154" stateProvince="Antalya">Troodos</location>
, 
<geoCoordinate id="EE3C501BFFDD94301EFCFCF093A9FCF4" box="[420,494,824,846]" degrees="34" direction="north" minutes="55" orientation="latitude" pageId="5" pageNumber="154" precision="925" value="34.916668">34°55'N</geoCoordinate>
, 
<geoCoordinate id="EE3C501BFFDD94301EAEFCF09079FCF4" box="[502,574,824,846]" degrees="32" direction="east" minutes="52" orientation="longitude" pageId="5" pageNumber="154" precision="925" value="32.866665">32°52'E</geoCoordinate>
, 
<quantity id="4CF09B39FFDD94301D1FFCF090CCFCF7" box="[583,651,824,846]" metricMagnitude="3" metricUnit="m" metricValue="1.55" pageId="5" pageNumber="154" unit="m" value="1550.0">
<elevation id="0025D1EFFFDD94301D1FFCF090CCFCF7" box="[583,651,824,846]" metricMagnitude="3" metricUnit="m" metricValue="1.55" pageId="5" pageNumber="154" unit="m" value="1550.0">1550 m</elevation>
</quantity>
, 
<date id="FFB6101CFFDD94301DCCFCF090A9FCF7" box="[660,750,824,845]" pageId="5" pageNumber="154" value="2008-04-09">
<collectingDate id="EFF2E9F4FFDD94301DCCFCF090A9FCF7" box="[660,750,824,845]" pageId="5" pageNumber="154" value="2008-04-09">9.IV.2008</collectingDate>
</date>
, leg. 
<collectorName id="26FD530AFFDD94301C45FCF0911CFCF7" box="[797,859,824,845]" pageId="5" pageNumber="154">Lompe</collectorName>
(cAss).
</materialsCitation>
<materialsCitation id="3B603C81FFDD94301CFCFCF09193FCDC" ID-GBIF-Occurrence="2846796357" collectingDate="2005-04-19" collectingDateMax="2008-04-24" collectingDateMin="2005-04-19" collectorName="Brachat &amp; Meybohm &amp; Lompe &amp; Assmann" country="Turkey" county="Gazipasa" elevation="40" latitude="32.883335" location="Troodos" longLatPrecision="1208" longitude="35.633335" municipality="Kalandere" pageId="5" pageNumber="154" specimenCount="4" specimenCount-male="4" stateProvince="Antalya">
<collectingCountry id="F31F764CFFDD94301CFCFCF09190FCF4" box="[932,983,824,846]" name="Israel" pageId="5" pageNumber="154">Israel</collectingCountry>
: 
<specimenCount id="9D0EFD55FFDD94301C87FCF091BBFCF7" box="[991,1020,824,845]" pageId="5" pageNumber="154" type="male">1♂</specimenCount>
, 
<location id="8ED76007FFDD94301B5EFCF09367FCDC" LSID="urn:lsid:plazi:treatment:03A187CAFFDD943D1FEAFE38904AFAD8:8ED76007FFDD94301B5EFCF09367FCDC" country="Turkey" county="Gazipasa" latitude="32.883335" longLatPrecision="1208" longitude="35.633335" municipality="Kalandere" name="Sea of Galilee" pageId="5" pageNumber="154" stateProvince="Antalya">Sea of Galilee</location>
, 
<location id="8ED76007FFDD94301E71FC98939BFCDF" LSID="urn:lsid:plazi:treatment:03A187CAFFDD943D1FEAFE38904AFAD8:8ED76007FFDD94301E71FC98939BFCDF" box="[297,476,848,870]" country="Turkey" county="Gazipasa" latitude="32.883335" longLatPrecision="1208" longitude="35.633335" municipality="Kalandere" name="Beit Tsaida Reserve" pageId="5" pageNumber="154" stateProvince="Antalya">Beit Tsaida Reserve</location>
, 
<geoCoordinate id="EE3C501BFFDD94301EBDFC989077FCDC" box="[485,560,848,870]" degrees="32" direction="north" minutes="53" orientation="latitude" pageId="5" pageNumber="154" precision="925" value="32.883335">32°53'N</geoCoordinate>
, 
<geoCoordinate id="EE3C501BFFDD94301D61FC9890C7FCDC" box="[569,640,848,870]" degrees="35" direction="east" minutes="38" orientation="longitude" pageId="5" pageNumber="154" precision="925" value="35.633335">35°38'E</geoCoordinate>
, 0 m, 
<date id="FFB6101CFFDD94301DE0FC98915CFCDC" box="[696,795,848,870]" pageId="5" pageNumber="154" value="2005-04-19">
<collectingDate id="EFF2E9F4FFDD94301DE0FC98915CFCDC" box="[696,795,848,870]" pageId="5" pageNumber="154" value="2005-04-19">19.IV.2005</collectingDate>
</date>
, leg. 
<collectorName id="26FD530AFFDD94301C15FC9891DEFCDF" box="[845,921,848,869]" pageId="5" pageNumber="154">Assmann</collectorName>
(cFel)
</materialsCitation>
; 
<materialsCitation id="3B603C81FFDD94301C86FC9891D1FCC4" ID-GBIF-Occurrence="2846796364" collectingDate="2008-03-26" collectorName="Assmann" country="Jordan" location="Capernaum" municipality="Sea of Galilee" pageId="5" pageNumber="154" specimenCount="1" specimenCount-female="1">
<specimenCount id="9D0EFD55FFDD94301C86FC9891BDFCD2" box="[990,1018,848,872]" pageId="5" pageNumber="154" type="female">1♀</specimenCount>
, 
<collectingMunicipality id="6BD3ACA6FFDD94301B5DFC989367FCC4" pageId="5" pageNumber="154">Sea of Galilee</collectingMunicipality>
, 
<location id="8ED76007FFDD94301E75FCA093D3FCC7" LSID="urn:lsid:plazi:treatment:03A187CAFFDD943D1FEAFE38904AFAD8:8ED76007FFDD94301E75FCA093D3FCC7" box="[301,404,872,893]" country="Jordan" municipality="Sea of Galilee" name="Capernaum" pageId="5" pageNumber="154">Capernaum</location>
, bank of 
<collectingCountry id="F31F764CFFDD94301EABFCA0906AFCC4" box="[499,557,872,894]" name="Jordan" pageId="5" pageNumber="154">Jordan</collectingCountry>
river, 
<date id="FFB6101CFFDD94301D28FCA09094FCC4" box="[624,723,872,894]" pageId="5" pageNumber="154" value="2008-03-26">
<collectingDate id="EFF2E9F4FFDD94301D28FCA09094FCC4" box="[624,723,872,894]" pageId="5" pageNumber="154" value="2008-03-26">26.III.2008</collectingDate>
</date>
, leg. 
<collectorName id="26FD530AFFDD94301C53FCA09110FCC7" box="[779,855,872,893]" pageId="5" pageNumber="154">Assmann</collectorName>
(cFel)
</materialsCitation>
; 
<materialsCitation id="3B603C81FFDD94301CFCFCA09108FC2C" ID-GBIF-Occurrence="2846796391" collectingDate="2010-11-13" collectorName="Drees" country="Jordan" elevation="65" latitude="33.066666" location="Hula" longLatPrecision="1207" longitude="35.6" pageId="5" pageNumber="154" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D0EFD55FFDD94301CFCFCA09186FCC7" box="[932,961,872,893]" pageId="5" pageNumber="154" type="male">1♂</specimenCount>
, 
<location id="8ED76007FFDD94301C97FCA091BDFCC4" LSID="urn:lsid:plazi:treatment:03A187CAFFDD943D1FEAFE38904AFAD8:8ED76007FFDD94301C97FCA091BDFCC4" box="[975,1018,872,894]" country="Jordan" latitude="33.066666" longLatPrecision="1207" longitude="35.6" name="Hula" pageId="5" pageNumber="154">Hula</location>
N. R., 
<geoCoordinate id="EE3C501BFFDD94301F87FC48936EFC2F" box="[223,297,896,917]" degrees="33" direction="north" minutes="04" orientation="latitude" pageId="5" pageNumber="154" precision="925" value="33.066666">33°04'N</geoCoordinate>
, 
<geoCoordinate id="EE3C501BFFDD94301E6CFC48933CFC2C" box="[308,379,896,918]" degrees="35" direction="east" minutes="36" orientation="longitude" pageId="5" pageNumber="154" precision="925" value="35.6">35°36'E</geoCoordinate>
, 
<quantity id="4CF09B39FFDD94301EDEFC4893F1FC2F" box="[390,438,896,918]" metricMagnitude="1" metricUnit="m" metricValue="6.5" pageId="5" pageNumber="154" unit="m" value="65.0">
<elevation id="0025D1EFFFDD94301EDEFC4893F1FC2F" box="[390,438,896,918]" metricMagnitude="1" metricUnit="m" metricValue="6.5" pageId="5" pageNumber="154" unit="m" value="65.0">65 m</elevation>
</quantity>
, shore of pond, 
<date id="FFB6101CFFDD94301D10FC4890EAFC2F" box="[584,685,896,917]" pageId="5" pageNumber="154" value="2010-11-13">
<collectingDate id="EFF2E9F4FFDD94301D10FC4890EAFC2F" box="[584,685,896,917]" pageId="5" pageNumber="154" value="2010-11-13">13.XI.2010</collectingDate>
</date>
, leg. 
<collectorName id="26FD530AFFDD94301D87FC489155FC2F" box="[735,786,896,917]" pageId="5" pageNumber="154">Drees</collectorName>
(cFel)
</materialsCitation>
; 
<materialsCitation id="3B603C81FFDD94301C02FC489133FC14" ID-GBIF-Occurrence="2846796362" collectingDate="2013-04-12" collectorName="S Ashdod &amp; Buse" country="Jordan" latitude="31.75" location="Nitzanim" longLatPrecision="1214" longitude="34.616665" pageId="5" pageNumber="154" specimenCount="1" specimenCount-female="1">
<specimenCount id="9D0EFD55FFDD94301C02FC489131FC22" box="[858,886,896,920]" pageId="5" pageNumber="154" type="female">1♀</specimenCount>
, 
<collectorName id="26FD530AFFDD94301CDAFC48919CFC2C" box="[898,987,896,918]" pageId="5" pageNumber="154">S Ashdod</collectorName>
, 
<location id="8ED76007FFDD94301CBEFC48967DFC2C" LSID="urn:lsid:plazi:treatment:03A187CAFFDD943D1FEAFE38904AFAD8:8ED76007FFDD94301CBEFC48967DFC2C" box="[998,1082,896,918]" country="Jordan" latitude="31.75" longLatPrecision="1214" longitude="34.616665" name="Nitzanim" pageId="5" pageNumber="154">Nitzanim</location>
, 
<geoCoordinate id="EE3C501BFFDD94301F87FC50936EFC14" box="[223,297,920,942]" degrees="31" direction="north" minutes="45" orientation="latitude" pageId="5" pageNumber="154" precision="925" value="31.75">31°45'N</geoCoordinate>
, 
<geoCoordinate id="EE3C501BFFDD94301E6EFC50933AFC17" box="[310,381,920,941]" degrees="34" direction="east" minutes="37" orientation="longitude" pageId="5" pageNumber="154" precision="925" value="34.616665">34°37'E</geoCoordinate>
, sand dunes, pond shore, 
<date id="FFB6101CFFDD94301D37FC509093FC17" box="[623,724,920,941]" pageId="5" pageNumber="154" value="2013-04-12">
<collectingDate id="EFF2E9F4FFDD94301D37FC509093FC17" box="[623,724,920,941]" pageId="5" pageNumber="154" value="2013-04-12">12.IV.2013</collectingDate>
</date>
, leg. 
<collectorName id="26FD530AFFDD94301C52FC509171FC17" box="[778,822,920,941]" pageId="5" pageNumber="154">Buse</collectorName>
(cFel)
</materialsCitation>
; 
<materialsCitation id="3B603C81FFDD94301CD9FC509137FC7C" ID-GBIF-Occurrence="2846796310" collectingDate="2011-03-19" collectorName="Meybohm" country="Jordan" elevation="215" latitude="32.9" location="River" longLatPrecision="1208" longitude="35.616665" municipality="Sea Genezareth" pageId="5" pageNumber="154" specimenCount="1" specimenCount-female="1">
<specimenCount id="9D0EFD55FFDD94301CD9FC5091DAFC0A" box="[897,925,920,944]" pageId="5" pageNumber="154" type="female">1♀</specimenCount>
, 
<collectingMunicipality id="6BD3ACA6FFDD94301CF3FC50967DFC14" box="[939,1082,920,942]" pageId="5" pageNumber="154">Sea Genezareth</collectingMunicipality>
, 
<collectingCountry id="F31F764CFFDD94301F87FC78935EFC7C" box="[223,281,944,966]" name="Jordan" pageId="5" pageNumber="154">Jordan</collectingCountry>
<location id="8ED76007FFDD94301E47FC789316FC7C" LSID="urn:lsid:plazi:treatment:03A187CAFFDD943D1FEAFE38904AFAD8:8ED76007FFDD94301E47FC789316FC7C" box="[287,337,944,966]" country="Jordan" latitude="32.9" longLatPrecision="1208" longitude="35.616665" municipality="Sea Genezareth" name="River" pageId="5" pageNumber="154">River</location>
, 
<geoCoordinate id="EE3C501BFFDD94301E01FC7893E4FC7C" box="[345,419,944,966]" degrees="32" direction="north" minutes="54" orientation="latitude" pageId="5" pageNumber="154" precision="925" value="32.9">32°54'N</geoCoordinate>
, 
<geoCoordinate id="EE3C501BFFDD94301EF4FC7893B4FC7C" box="[428,499,944,966]" degrees="35" direction="east" minutes="37" orientation="longitude" pageId="5" pageNumber="154" precision="925" value="35.616665">35°37'E</geoCoordinate>
, - 
<quantity id="4CF09B39FFDD94301D5AFC78907BFC7F" box="[514,572,944,966]" metricMagnitude="2" metricUnit="m" metricValue="2.15" pageId="5" pageNumber="154" unit="m" value="215.0">
<elevation id="0025D1EFFFDD94301D5AFC78907BFC7F" box="[514,572,944,966]" metricMagnitude="2" metricUnit="m" metricValue="2.15" pageId="5" pageNumber="154" unit="m" value="215.0">215 m</elevation>
</quantity>
, 
<date id="FFB6101CFFDD94301D1DFC7890EFFC7F" box="[581,680,944,965]" pageId="5" pageNumber="154" value="2011-03-19">
<collectingDate id="EFF2E9F4FFDD94301D1DFC7890EFFC7F" box="[581,680,944,965]" pageId="5" pageNumber="154" value="2011-03-19">19.III.2011</collectingDate>
</date>
, leg. 
<collectorName id="26FD530AFFDD94301D8EFC789177FC7C" box="[726,816,944,966]" pageId="5" pageNumber="154">Meybohm</collectorName>
(cAss)
</materialsCitation>
.
</paragraph>
<paragraph id="8BB736DCFFDD94301FEAFC1C91C8FB48" blockId="5.[178,1085,980,1266]" pageId="5" pageNumber="154">
C o m m e n t: The original description of 
<taxonomicName id="4C084D5FFFDD94301D23FC1D915DFC53" authorityName="Coiffait" authorityYear="1963" box="[635,794,980,1002]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="154" phylum="Arthropoda" rank="species" species="hermonensis">
<emphasis id="B97CEACEFFDD94301D23FC1D915DFC53" box="[635,794,980,1002]" italics="true" pageId="5" pageNumber="154">Q. hermonensis</emphasis>
</taxonomicName>
is based on a male 
<typeStatus id="54B3887EFFDD94301CBCFC1C967AFC50" box="[996,1085,980,1002]" pageId="5" pageNumber="154" type="holotype">holotype</typeStatus>
and a female 
<typeStatus id="54B3887EFFDD94301E1AFC3A93DEFBBD" box="[322,409,1010,1031]" pageId="5" pageNumber="154" type="paratype">paratype</typeStatus>
from &quot; 
<collectingCountry id="F31F764CFFDD94301EBEFC3A9060FBB2" box="[486,551,1010,1032]" name="Lebanon" pageId="5" pageNumber="154">Liban</collectingCountry>
: Hasbaya&quot; and one female 
<typeStatus id="54B3887EFFDD94301C1FFC3A91D9FBBD" box="[839,926,1010,1031]" pageId="5" pageNumber="154" type="paratype">paratype</typeStatus>
from &quot;Nahr es Safa&quot; (
<bibRefCitation id="EF994B2DFFDD94301FAFFBC793D5FB9F" author="COIFFAIT H" box="[247,402,1039,1061]" pageId="5" pageNumber="154" pagination="372 - 420" refId="ref20207" refString="COIFFAIT H. (1963): Les Quedius du sous-genre Sauridus de la region palearctique occidentale (avec description de fomes nouvelles). - Bulletin d e la Societe d'Histoire Naturelle de Toulouse 98 (3 - 4): 372 - 420." type="journal article" year="1963">COIFFAIT 1963</bibRefCitation>
), that of 
<emphasis id="B97CEACEFFDD94301EB4FBC790CCFB9E" box="[492,651,1039,1060]" italics="true" pageId="5" pageNumber="154">Q. coiffaitianus</emphasis>
on a male 
<typeStatus id="54B3887EFFDD94301DA2FBC79114FB9F" box="[762,851,1039,1061]" pageId="5" pageNumber="154" type="holotype">holotype</typeStatus>
and 
<specimenCount id="9D0EFD55FFDD94301CDCFBC79642FB9F" box="[900,1029,1039,1061]" pageId="5" pageNumber="154" type="generic" typeStatus="paratypes">27 paratypes</specimenCount>
from &quot;Nabeh Safa&quot; (
<bibRefCitation id="EF994B2DFFDD94301E14FBE49391FBF8" author="FAGEL G" box="[332,470,1068,1090]" pageId="5" pageNumber="154" pagination="1 - 13" refId="ref20496" refString="FAGEL G. (1968 a): Contribution a la connaissance des Staphylinidae CI. - Remarques diverses sur quelques Quedius de Mediterranee orientale. - B ulletin Institut Royale des Sciences Naturelles de Belgique 44 (12): 1 - 13." type="journal article" year="1968">FAGEL 1968a</bibRefCitation>
) (i.e., from the same locality as that of one of the 
<typeStatus id="54B3887EFFDD94301C84FBE5967AFBF8" box="[988,1085,1069,1090]" pageId="5" pageNumber="154" type="paratype">paratypes</typeStatus>
of 
<taxonomicName id="4C084D5FFFDD94301F88FB829336FBE5" authorityName="Coiffait" authorityYear="1963" box="[208,369,1098,1119]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="154" phylum="Arthropoda" rank="species" species="hermonensis">
<emphasis id="B97CEACEFFDD94301F88FB829336FBE5" box="[208,369,1098,1119]" italics="true" pageId="5" pageNumber="154">Q. hermonensis</emphasis>
</taxonomicName>
), and that of 
<emphasis id="B97CEACEFFDD94301D58FB8290E3FBE5" box="[512,676,1098,1119]" italics="true" pageId="5" pageNumber="154">Q. rugosipennis</emphasis>
on a male 
<typeStatus id="54B3887EFFDD94301C43FB829133FBDA" box="[795,884,1098,1120]" pageId="5" pageNumber="154" type="holotype">holotype</typeStatus>
and four 
<typeStatus id="54B3887EFFDD94301C84FB82967AFBE5" box="[988,1085,1098,1119]" pageId="5" pageNumber="154" type="paratype">paratypes</typeStatus>
(one male and three females) from &quot;Anatolie méridionale: Antalya, route de Lara&quot; (
<bibRefCitation id="EF994B2DFFDD94301FE1FB4C9370FB20" author="FAGEL G" box="[185,311,1156,1178]" pageId="5" pageNumber="154" pagination="103 - 125" refId="ref20632" refString="FAGEL G. (1969): Contribution a la connaissance des Staphylinidae CVIII. - Sur quelques especes inedites provenant du bassin mediterraneen. - Bulletin et Annales de la Societe Royale d'Entomologie de Belgique 105: 103 - 125." type="journal article" year="1969">FAGEL 1969</bibRefCitation>
). An examination of the type material of 
<emphasis id="B97CEACEFFDD94301D8AFB4D912EFB23" box="[722,873,1156,1178]" italics="true" pageId="5" pageNumber="154">Q. coiffaitianus</emphasis>
and 
<emphasis id="B97CEACEFFDD94301CC3FB4D9671FB20" box="[923,1078,1157,1178]" italics="true" pageId="5" pageNumber="154">Q. rugosipennis</emphasis>
, as well as of additional material from various regions revealed that in both external and the male sexual characters they are within the range of intraspecific variation of 
<taxonomicName id="4C084D5FFFDD94301B7DFB779375FB4B" authorityName="Coiffait" authorityYear="1963" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="154" phylum="Arthropoda" rank="species" species="hermonensis">
<emphasis id="B97CEACEFFDD94301B7DFB779375FB4B" italics="true" pageId="5" pageNumber="154">Q. hermonensis</emphasis>
</taxonomicName>
by 
<bibRefCitation id="EF994B2DFFDD94301E00FB149044FB48" author="COIFFAIT H" box="[344,515,1244,1266]" pageId="5" pageNumber="154" pagination="372 - 420" refId="ref20207" refString="COIFFAIT H. (1963): Les Quedius du sous-genre Sauridus de la region palearctique occidentale (avec description de fomes nouvelles). - Bulletin d e la Societe d'Histoire Naturelle de Toulouse 98 (3 - 4): 372 - 420." type="journal article" year="1963">COIFFAIT (1963)</bibRefCitation>
. Hence the synonymy proposed above.
</paragraph>
<paragraph id="8BB736DCFFDD94301FEAFACA91F8F99A" blockId="5.[178,1085,1282,1568]" pageId="5" pageNumber="154">
Based on similar external characters (coloration, punctation, habitus, etc.), on the similar general morphology of the aedeagus, and a similar ecology, 
<taxonomicName id="4C084D5FFFDD94301C68FAD79195FA8E" authorityName="Coiffait" authorityYear="1963" box="[816,978,1311,1332]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="154" phylum="Arthropoda" rank="species" species="hermonensis">
<emphasis id="B97CEACEFFDD94301C68FAD79195FA8E" box="[816,978,1311,1332]" italics="true" pageId="5" pageNumber="154">Q. hermonensis</emphasis>
</taxonomicName>
is closely allied to 
<taxonomicName id="4C084D5FFFDD94301E4DFAF593DEFAEB" box="[277,409,1340,1362]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="154" phylum="Arthropoda" rank="species" species="umbrinus">
<emphasis id="B97CEACEFFDD94301E4DFAF593DEFAEB" box="[277,409,1340,1362]" italics="true" pageId="5" pageNumber="154">Q. umbrinus</emphasis>
</taxonomicName>
, not to 
<emphasis id="B97CEACEFFDD94301EAEFAF5903EFAE8" box="[502,633,1341,1362]" italics="true" pageId="5" pageNumber="154">Q. nigriceps</emphasis>
<bibRefCitation id="EF994B2DFFDD94301DDDFAF59158FAE8" author="KRAATZ G" box="[645,799,1340,1362]" pageId="5" pageNumber="154" pagination="377 - 1080" refId="ref21018" refString="KRAATZ G. (1857): Naturgeschichte der Insecten Deutschlands. Erste Abtheilung Coleoptera. Zweiter Band. Lieferung 3 - 6. - Nicolai, Berlin: 377 - 1080." type="book chapter" year="1857">KRAATZ, 1857</bibRefCitation>
as suggested by 
<bibRefCitation id="EF994B2DFFDD94301C87FAF49343FACA" author="COIFFAIT H" pageId="5" pageNumber="154" refId="ref20382" refString="COIFFAIT H. (1978 a): Coleopteres staphylinides de la region paleartique [sic] occidentale. III. Sous famille Staphylininae, Tribu Quediini. Sous famille Paederinae, Tribu Pinophilini. - Supplement a la Nouvell e Revue d'Entomologie 8 (4): 364 pp." type="book" year="1978">COIFFAIT (1978a)</bibRefCitation>
. Like 
<taxonomicName id="4C084D5FFFDD94301E1CFA929383FAD5" box="[324,452,1370,1391]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="154" phylum="Arthropoda" rank="species" species="umbrinus">
<emphasis id="B97CEACEFFDD94301E1CFA929383FAD5" box="[324,452,1370,1391]" italics="true" pageId="5" pageNumber="154">Q. umbrinus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C084D5FFFDD94301E89FA929036FAD5" authorityName="Coiffait" authorityYear="1963" box="[465,625,1370,1391]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="154" phylum="Arthropoda" rank="species" species="hermonensis">
<emphasis id="B97CEACEFFDD94301E89FA929036FAD5" box="[465,625,1370,1391]" italics="true" pageId="5" pageNumber="154">Q. hermonensis</emphasis>
</taxonomicName>
is subject to pronounced variation of coloration and of the shape of the aedeagus (
<figureCitation id="13332A59FFDD94301D0CFABF90FCFA37" box="[596,699,1399,1421]" captionStart-0="Figs 1-12" captionStart-1="Figs 13-24" captionStartId-0="6.[178,218,1506,1528]" captionStartId-1="7.[178,217,1506,1528]" captionTargetBox-0="[182,1076,226,1483]" captionTargetBox-1="[187,1079,217,1465]" captionTargetId-0="figure@6.[178,1085,212,1497]" captionTargetId-1="figure@7.[178,1085,212,1497]" captionTargetPageId-1="7" captionText-0="Figs 1-12: Quedius hermonensis from Samos (1-3), Israel (4-6), and Cyprus (7-12): (1, 4, 7, 10) aedeagus in lateral view; (2, 5, 8, 11) median lobe of aedeagus in ventral view; (3, 6, 9, 12) paramere. Scale bar: 0.5 mm." captionText-1="Figs 13-24: Quedius hermonensis from Cyprus (13-14), Q. orientalis (15-17), and Q. microcapillatus (18-24) from Iraq (18-20), Turkey (21), and Iran (22-24): (13, 15, 22) median lobe of aedeagus in lateral view; (14, 17, 20, 24) paramere; (16, 19, 23) median lobe of aedeagus in ventral view; (18, 21) aedeagus in lateral view. Scale bar: 0.5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.3995001" figureDoi-1="http://doi.org/10.5281/zenodo.3985370" httpUri-0="https://zenodo.org/record/3995001/files/figure.png" httpUri-1="https://zenodo.org/record/3985370/files/figure.png" pageId="5" pageNumber="154">Figs 1-14</figureCitation>
). The colour of the forebody ranges from brown to black, often with the anterior and posterior margins of the elytra narrowly yellowish to reddish (especially in material from 
<collectingCountry id="F31F764CFFDD94301DE5FA7A914CFA72" box="[701,779,1458,1480]" name="Cyprus" pageId="5" pageNumber="154">Cyprus</collectingCountry>
). The length of the aedeagus ranges from 1.0- 
<quantity id="4CF09B39FFDD94301E07FA0793F5FA5F" box="[351,434,1487,1509]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="5" pageNumber="154" unit="mm" value="1.3">1.3 mm</quantity>
. The width of the paramere is rather variable, too; in material from 
<collectingCountry id="F31F764CFFDD94301FB5FA249371F9BB" box="[237,310,1516,1537]" name="Cyprus" pageId="5" pageNumber="154">Cyprus</collectingCountry>
it is more slender and basally sometimes slightly constricted, whereas in material seen from other regions it is less slender and basally not constricted.
</paragraph>
<caption id="DF776654FFDE94331FEAFA2A93D1F99F" ID-DOI="http://doi.org/10.5281/zenodo.3995001" ID-Zenodo-Dep="3995001" httpUri="https://zenodo.org/record/3995001/files/figure.png" pageId="6" pageNumber="155" startId="6.[178,218,1506,1528]" targetBox="[182,1076,226,1483]" targetPageId="6">
<paragraph id="8BB736DCFFDE94331FEAFA2A93D1F99F" blockId="6.[178,1085,1506,1574]" pageId="6" pageNumber="155">
<emphasis id="B97CEACEFFDE94331FEAFA2A934AFA4D" bold="true" box="[178,269,1506,1528]" pageId="6" pageNumber="155">Figs 1-12</emphasis>
: 
<taxonomicName id="4C084D5FFFDE94331E43FA2A93A7FA4D" authorityName="Coiffait" authorityYear="1963" box="[283,480,1506,1527]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="155" phylum="Arthropoda" rank="species" species="hermonensis">
<emphasis id="B97CEACEFFDE94331E43FA2A93A7FA4D" box="[283,480,1506,1527]" italics="true" pageId="6" pageNumber="155">Quedius hermonensis</emphasis>
</taxonomicName>
from Samos (
<emphasis id="B97CEACEFFDE94331D31FA2A90C0FA4D" bold="true" box="[617,647,1506,1527]" pageId="6" pageNumber="155">1-3</emphasis>
), 
<collectingCountry id="F31F764CFFDE94331DC4FA2A9089FA42" box="[668,718,1506,1528]" name="Israel" pageId="6" pageNumber="155">Israel</collectingCountry>
(
<emphasis id="B97CEACEFFDE94331D86FA2A90BBFA4D" bold="true" box="[734,764,1506,1527]" pageId="6" pageNumber="155">4-6</emphasis>
), and 
<collectingCountry id="F31F764CFFDE94331C62FA2B913BFA42" box="[826,892,1507,1528]" name="Cyprus" pageId="6" pageNumber="155">Cyprus</collectingCountry>
(
<emphasis id="B97CEACEFFDE94331CD3FA2A91F3FA4D" bold="true" box="[907,948,1506,1527]" pageId="6" pageNumber="155">7-12</emphasis>
): (
<emphasis id="B97CEACEFFDE94331C8AFA2A9671FA4D" bold="true" box="[978,1078,1506,1528]" pageId="6" pageNumber="155">1, 4, 7, 10</emphasis>
) aedeagus in lateral view; (
<emphasis id="B97CEACEFFDE94331EFDFA319041F9B4" bold="true" box="[421,518,1529,1550]" pageId="6" pageNumber="155">2, 5, 8, 11</emphasis>
) median lobe of aedeagus in ventral view; (
<emphasis id="B97CEACEFFDE94331CC7FA319647F9B4" bold="true" box="[927,1024,1529,1550]" pageId="6" pageNumber="155">3, 6, 9, 12</emphasis>
) paramere. Scale bar: 
<quantity id="4CF09B39FFDE94331E12F9D893D5F99F" box="[330,402,1552,1574]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="6" pageNumber="155" unit="mm" value="0.5">0.5 mm</quantity>
.
</paragraph>
</caption>
<caption id="DF776654FFDF94321FEAFA2A902FF986" ID-DOI="http://doi.org/10.5281/zenodo.3985370" ID-Zenodo-Dep="3985370" httpUri="https://zenodo.org/record/3985370/files/figure.png" pageId="7" pageNumber="156" startId="7.[178,217,1506,1528]" targetBox="[187,1079,217,1465]" targetPageId="7">
<paragraph id="8BB736DCFFDF94321FEAFA2A902FF986" blockId="7.[178,1085,1506,1596]" pageId="7" pageNumber="156">
<emphasis id="B97CEACEFFDF94321FEAFA2A9354FA4D" bold="true" box="[178,275,1506,1528]" pageId="7" pageNumber="156">Figs 13-24</emphasis>
: 
<taxonomicName id="4C084D5FFFDF94321E78FA2A93A7FA4D" authorityName="Coiffait" authorityYear="1963" box="[288,480,1506,1527]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="156" phylum="Arthropoda" rank="species" species="hermonensis">
<emphasis id="B97CEACEFFDF94321E78FA2A93A7FA4D" box="[288,480,1506,1527]" italics="true" pageId="7" pageNumber="156">Quedius hermonensis</emphasis>
</taxonomicName>
from Cyprus (
<emphasis id="B97CEACEFFDF94321D3DFA2A90DEFA4D" bold="true" box="[613,665,1506,1527]" pageId="7" pageNumber="156">13-14</emphasis>
), 
<taxonomicName id="4C084D5FFFDF94321DF4FA2B915BFA4D" authorityName="Korge" authorityYear="1971" box="[684,796,1506,1528]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="156" phylum="Arthropoda" rank="species" species="orientalis">
<emphasis id="B97CEACEFFDF94321DF4FA2B915BFA4D" box="[684,796,1506,1528]" italics="true" pageId="7" pageNumber="156">Q. orientalis</emphasis>
</taxonomicName>
(
<emphasis id="B97CEACEFFDF94321C72FA2A911AFA4D" bold="true" box="[810,861,1506,1527]" pageId="7" pageNumber="156">15-17</emphasis>
), and 
<taxonomicName id="4C084D5FFFDF94321CCFFA2B967AFA4D" authorityName="Korge" authorityYear="1971" box="[919,1085,1506,1528]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="156" phylum="Arthropoda" rank="species" species="microcapillatus">
<emphasis id="B97CEACEFFDF94321CCFFA2B967AFA4D" box="[919,1085,1506,1528]" italics="true" pageId="7" pageNumber="156">Q. microcapillatus</emphasis>
</taxonomicName>
(
<emphasis id="B97CEACEFFDF94321FE2FA3192A9F9B4" bold="true" box="[186,238,1529,1550]" pageId="7" pageNumber="156">18-24</emphasis>
) from Iraq (
<emphasis id="B97CEACEFFDF94321E3BFA3193DFF9B4" bold="true" box="[355,408,1529,1550]" pageId="7" pageNumber="156">18-20</emphasis>
), Turkey (
<emphasis id="B97CEACEFFDF94321EA4FA319054F9B4" bold="true" box="[508,531,1529,1550]" pageId="7" pageNumber="156">21</emphasis>
), and Iran (
<emphasis id="B97CEACEFFDF94321DDAFA3190F1F9B4" bold="true" box="[642,694,1529,1550]" pageId="7" pageNumber="156">22-24</emphasis>
): (
<emphasis id="B97CEACEFFDF94321D8BFA319176F9B4" bold="true" box="[723,817,1529,1550]" pageId="7" pageNumber="156">13, 15, 22</emphasis>
) median lobe of aedeagus in lateral view; (
<emphasis id="B97CEACEFFDF94321E6AF9D893F3F99F" bold="true" box="[306,436,1552,1573]" pageId="7" pageNumber="156">14, 17, 20, 24</emphasis>
) paramere; (
<emphasis id="B97CEACEFFDF94321D73F9D890CFF99F" bold="true" box="[555,648,1552,1573]" pageId="7" pageNumber="156">16, 19, 23</emphasis>
) median lobe of aedeagus in ventral view; (
<emphasis id="B97CEACEFFDF94321B79F9D8928EF981" bold="true" pageId="7" pageNumber="156">18, 21</emphasis>
) aedeagus in lateral view. Scale bar: 0.5 mm.
</paragraph>
</caption>
<caption id="DF776654FFD0943D1FEAFC2C90BAFBAA" ID-DOI="http://doi.org/10.5281/zenodo.3985372" ID-Zenodo-Dep="3985372" httpUri="https://zenodo.org/record/3985372/files/figure.png" pageId="8" pageNumber="157" startId="8.[178,218,996,1018]" targetBox="[187,1085,212,973]" targetPageId="8">
<paragraph id="8BB736DCFFD0943D1FEAFC2C90BAFBAA" blockId="8.[178,1084,996,1041]" pageId="8" pageNumber="157">
<emphasis id="B97CEACEFFD0943D1FEAFC2C9350FC43" bold="true" box="[178,279,996,1018]" pageId="8" pageNumber="157">Figs 25-27</emphasis>
: 
<taxonomicName id="4C084D5FFFD0943D1E7CFC2C9389FC43" authorityName="Korge" authorityYear="1971" box="[292,462,996,1017]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="157" phylum="Arthropoda" rank="species" species="orientalis">
<emphasis id="B97CEACEFFD0943D1E7CFC2C9389FC43" box="[292,462,996,1017]" italics="true" pageId="8" pageNumber="157">Quedius orientalis</emphasis>
</taxonomicName>
(
<emphasis id="B97CEACEFFD0943D1E84FC2C93B4FC43" bold="true" box="[476,499,996,1017]" pageId="8" pageNumber="157">25</emphasis>
) and 
<taxonomicName id="4C084D5FFFD0943D1D71FC2C9093FC43" authorityName="Korge" authorityYear="1971" box="[553,724,996,1017]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="157" phylum="Arthropoda" rank="species" species="microcapillatus">
<emphasis id="B97CEACEFFD0943D1D71FC2C9093FC43" box="[553,724,996,1017]" italics="true" pageId="8" pageNumber="157">Q. microcapillatus</emphasis>
</taxonomicName>
(
<emphasis id="B97CEACEFFD0943D1DBAFC2C915FFC43" bold="true" box="[738,792,996,1017]" pageId="8" pageNumber="157">26-27</emphasis>
): (
<emphasis id="B97CEACEFFD0943D1C6CFC2C912EFC43" bold="true" box="[820,873,996,1017]" pageId="8" pageNumber="157">25-26</emphasis>
) habitus; (
<emphasis id="B97CEACEFFD0943D1C95FC2C91A3FC43" bold="true" box="[973,996,996,1017]" pageId="8" pageNumber="157">27</emphasis>
) posterosutural portion of elytra. Scale bars: 25-26: 1.0 mm; 27: 0.1 mm.
</paragraph>
</caption>
<paragraph id="8BB736DCFFD0943D1FEAFBD790CFFBD5" blockId="8.[178,1085,1055,1135]" pageId="8" pageNumber="157">
The habitat is similar to that of 
<taxonomicName id="4C084D5FFFD0943D1D5DFBD790C2FB8E" box="[517,645,1055,1076]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="157" phylum="Arthropoda" rank="species" species="umbrinus">
<emphasis id="B97CEACEFFD0943D1D5DFBD790C2FB8E" box="[517,645,1055,1076]" italics="true" pageId="8" pageNumber="157">Q. umbrinus</emphasis>
</taxonomicName>
. The examined specimens were primarily collected in moist substrates: in moist leaf litter, swamps, on the shores of lakes and ponds, and on the banks of rivers and streams.
</paragraph>
<paragraph id="8BB736DCFFD0943D1FEAFBB7904AFAD8" blockId="8.[178,1085,1151,1378]" pageId="8" pageNumber="157">
The revised distribution (
<figureCitation id="13332A59FFD0943D1E99FBB7904FFB2F" box="[449,520,1151,1173]" captionStart="Map 1" captionStartId="9.[178,223,1266,1287]" captionTargetBox="[178,1085,639,1254]" captionTargetId="figure@9.[178,1085,639,1254]" captionTargetPageId="9" captionText="Map 1: Distributions of Quedius hermonensis (black circles), Q. microcapillatus (white circles), and Q. orientalis (black star), based on revised records." figureDoi="http://doi.org/10.5281/zenodo.3985374" httpUri="https://zenodo.org/record/3985374/files/figure.png" pageId="8" pageNumber="157">Map 1</figureCitation>
) ranges from the Aegean Island Samos in the west across 
<collectingCountry id="F31F764CFFD0943D1FA4FB549302FB08" box="[252,325,1180,1202]" name="Turkey" pageId="8" pageNumber="157">Turkey</collectingCountry>
and 
<collectingCountry id="F31F764CFFD0943D1E26FB549380FB0B" box="[382,455,1180,1201]" name="Cyprus" pageId="8" pageNumber="157">Cyprus</collectingCountry>
to the Middle East (
<collectingCountry id="F31F764CFFD0943D1DF2FB549140FB08" box="[682,775,1180,1202]" name="Lebanon" pageId="8" pageNumber="157">Lebanon</collectingCountry>
, 
<collectingCountry id="F31F764CFFD0943D1C4DFB549115FB08" box="[789,850,1180,1202]" name="Israel" pageId="8" pageNumber="157">Israel</collectingCountry>
). While the species is fairly common in 
<collectingCountry id="F31F764CFFD0943D1E34FB7293F2FB75" box="[364,437,1210,1231]" name="Cyprus" pageId="8" pageNumber="157">Cyprus</collectingCountry>
(ASSING &amp; WUNDERLE 2001), it appears to be rather rare elsewhere. For previous records from 
<collectingCountry id="F31F764CFFD0943D1D1EFB1F90C8FB56" box="[582,655,1239,1260]" name="Cyprus" pageId="8" pageNumber="157">Cyprus</collectingCountry>
(as 
<emphasis id="B97CEACEFFD0943D1D89FB1F9138FB56" box="[721,895,1239,1260]" italics="true" pageId="8" pageNumber="157">Q. rugosipennis</emphasis>
), Samos (as 
<emphasis id="B97CEACEFFD0943D1B7CFB1F9372FAB0" italics="true" pageId="8" pageNumber="157">Q. rugosipennis</emphasis>
), and 
<collectingCountry id="F31F764CFFD0943D1E25FB3C93F2FAB0" box="[381,437,1268,1290]" name="Israel" pageId="8" pageNumber="157">Israel</collectingCountry>
see ASSING &amp; WUNDERLE (2001), 
<bibRefCitation id="EF994B2DFFD0943D1C76FB3C918CFAB0" author="ASSING V" box="[814,971,1268,1290]" pageId="8" pageNumber="157" pagination="81 - 102" refId="ref19575" refString="ASSING V. (2015): On the Staphylinidae of the Greek island Samos (Coleoptera: Staphylinidae). - Koleopt erologische Rundschau 85: 81 - 102." type="journal article" year="2015">ASSING (2015)</bibRefCitation>
, 
<bibRefCitation id="EF994B2DFFD0943D1C81FB3C92BFFA92" author="SMETANA A" pageId="8" pageNumber="157" pagination="115 - 126" refId="ref21678" refString="SMETANA A. (1978): Staphylinini and Quediini from Israel (Coleoptera, Staphylinidae). - Revue Suisse de Zoologie 85 (1): 115 - 126." type="journal article" year="1978">SMETANA (1978)</bibRefCitation>
, and 
<bibRefCitation id="EF994B2DFFD0943D1E77FADA9380FA92" author="ASSING V" box="[303,455,1298,1320]" pageId="8" pageNumber="157" pagination="1179 - 1210" refId="ref19530" refString="ASSING V. (2014): On the Staphylinidae of Israel II, with a revision of some species of Dinusa SAULCY (Coleoptera: Staphylinidae: Aleocharinae). - Linzer Biologisch e Beitrage 46 (2): 1179 - 1210." type="journal article" year="2014">ASSING (2014)</bibRefCitation>
, respectively. A previous record of 
<taxonomicName id="4C084D5FFFD0943D1C6DFADA9192FA9C" authorityName="Coiffait" authorityYear="1963" box="[821,981,1297,1319]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="157" phylum="Arthropoda" rank="species" species="hermonensis">
<emphasis id="B97CEACEFFD0943D1C6DFADA9192FA9C" box="[821,981,1297,1319]" italics="true" pageId="8" pageNumber="157">Q. hermonensis</emphasis>
</taxonomicName>
from 
<collectingCountry id="F31F764CFFD0943D1B4CFADA967AFA9D" box="[1044,1085,1298,1319]" name="Iraq" pageId="8" pageNumber="157">Iraq</collectingCountry>
(
<bibRefCitation id="EF994B2DFFD0943D1FE1FAE79303FAFF" author="ASSING V" box="[185,324,1327,1349]" pageId="8" pageNumber="157" pagination="1179 - 1210" refId="ref19530" refString="ASSING V. (2014): On the Staphylinidae of Israel II, with a revision of some species of Dinusa SAULCY (Coleoptera: Staphylinidae: Aleocharinae). - Linzer Biologisch e Beitrage 46 (2): 1179 - 1210." type="journal article" year="2014">ASSING 2014</bibRefCitation>
), however, which was overlooked by SCHÜLKE &amp; SMETANA (2015), refers to 
<taxonomicName id="4C084D5FFFD0943D1F94FA8493CDFADB" authorityName="Korge" authorityYear="1971" box="[204,394,1356,1377]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="157" phylum="Arthropoda" rank="species" species="microcapillatus">
<emphasis id="B97CEACEFFD0943D1F94FA8493CDFADB" box="[204,394,1356,1377]" italics="true" pageId="8" pageNumber="157">Q. microcapillatus</emphasis>
</taxonomicName>
(see below).
</paragraph>
</subSubSection>
</treatment>
</document>