153 lines
18 KiB
XML
153 lines
18 KiB
XML
<document id="1FCFA93A19DC744D7D020D7F0E6C26BD" ID-DOI="10.11646/zootaxa.3894.1.11" ID-GBIF-Dataset="6c37cabe-0df5-462b-9f82-b5acfc8c37cc" ID-ISSN="1175-5326" ID-Zenodo-Dep="229191" ID-ZooBank="78A995DA-7D11-48CE-A547-C7653E505720" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460997608442" checkinUser="plazi" docAuthor="Golovatch, Sergei I. & Vandenspiegel, Didier" docDate="2014" docId="0389DE08734D255AFF35F9A5FA63C533" docLanguage="en" docName="zt03894p151.pdf" docOrigin="Zootaxa 3894 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Koponenius Golovatch & Vandenspiegel, 2014, gen. nov." docType="treatment" docUuid="50B2AE78-D7C8-4787-B2B2-678CA2601748" docUuidSource="ZooBank" docVersion="8" lastPageNumber="144" masterDocId="FFB0A670734C2559FFA2FD62FFEBC76C" masterDocTitle="Koponenius gen. nov., a new genus of the millipede family Haplodesmidae from the Himalayas of India and Nepal (Diplopoda: Polydesmida)" masterLastPageNumber="151" masterPageNumber="141" pageNumber="142" updateTime="1698563275390" updateUser="plazi">
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<mods:titleInfo id="B0A3FD7F16C6C490C4E6D15D42B20999">
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<mods:title id="A22A1E1065DDCA05A0788E483C45847D">Koponenius gen. nov., a new genus of the millipede family Haplodesmidae from the Himalayas of India and Nepal (Diplopoda: Polydesmida)</mods:title>
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<mods:namePart id="4CFFD84EF4E74137F1BFC75E0BAB104C">Golovatch, Sergei I.</mods:namePart>
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<mods:roleTerm id="CD2E9F7AFF09F1E0F3ED0413040383EF">Author</mods:roleTerm>
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<mods:namePart id="BF3B08E1C3E4FFAD23875A29E38658E2">Vandenspiegel, Didier</mods:namePart>
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<mods:title id="2A88D4C6C2AFE4EF3DF442A65F848ECF">Zootaxa</mods:title>
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<mods:date id="DA6BEAA2423F4FB5D635A4AAEF1C4275">2014</mods:date>
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<mods:detail id="CDE4CC27E71C74A1BF06720DFB8CBA16" type="volume">
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<mods:number id="F8EC26F618FF49862E94A5CAE6EBE46E">3894</mods:number>
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<mods:classification id="6F48182F546C29D693B58D3432C1DAD9">journal article</mods:classification>
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<mods:identifier id="7210362EABC948C1CF8D925F2A0FEDD8" type="DOI">10.11646/zootaxa.3894.1.11</mods:identifier>
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<treatment id="0389DE08734D255AFF35F9A5FA63C533" ID-DOI="http://doi.org/10.5281/zenodo.6126345" ID-GBIF-Taxon="119597639" ID-Zenodo-Dep="6126345" LSID="urn:lsid:zoobank.org:act:50B2AE78-D7C8-4787-B2B2-678CA2601748" httpUri="http://treatment.plazi.org/id/0389DE08734D255AFF35F9A5FA63C533" lastPageId="3" lastPageNumber="144" pageId="1" pageNumber="142">
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<subSubSection id="C33A3C95734D2558FF35F9A5FD23C221" pageId="1" pageNumber="142" type="nomenclature">
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<paragraph id="8B9F6F1E734D2558FF35F9A5FE76C38D" blockId="1.[151,413,1223,1249]" box="[151,413,1223,1249]" pageId="1" pageNumber="142">
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<heading id="D0D7D872734D2558FF35F9A5FE76C38D" bold="true" box="[151,413,1223,1249]" fontSize="11" level="1" pageId="1" pageNumber="142" reason="1">
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<emphasis id="B954B30C734D2558FF35F9A5FE76C38D" bold="true" box="[151,413,1223,1249]" pageId="1" pageNumber="142">
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<taxonomicName id="4C20149D734D2558FF35F9A5FEC3C38D" ID-CoL="7PM6S" box="[151,296,1223,1249]" class="Diplopoda" family="Haplodesmidae" genus="Koponenius" kingdom="Animalia" order="Polydesmida" pageId="1" pageNumber="142" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B954B30C734D2558FF35F9A5FEC3C38D" bold="true" box="[151,296,1223,1249]" italics="true" pageId="1" pageNumber="142">Koponenius</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2670E77734D2558FE8DF9A5FE76C38D" box="[303,413,1223,1249]" pageId="1" pageNumber="142" rank="genus">gen. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B9F6F1E734D2558FF35F86CFD23C221" blockId="1.[151,1437,1294,2006]" pageId="1" pageNumber="142">
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<emphasis id="B954B30C734D2558FF35F86CFF0DC24B" bold="true" box="[151,230,1294,1319]" pageId="1" pageNumber="142">Name:</emphasis>
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Honours Seppo Koponen, a prominent colleague arachnologist from Turku University,
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<collectingCountry id="F3372F8E734D2558FB49F86CFAA8C24B" box="[1259,1347,1294,1319]" name="Finland" pageId="1" pageNumber="142">Finland</collectingCountry>
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, on the occasion of his 70th birthday; masculine in gender.
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</paragraph>
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</subSubSection>
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<subSubSection id="C33A3C95734D2558FF65F83BFE03C16E" pageId="1" pageNumber="142" type="diagnosis">
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<paragraph id="8B9F6F1E734D2558FF65F83BFE03C16E" blockId="1.[151,1437,1294,2006]" pageId="1" pageNumber="142">
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<emphasis id="B954B30C734D2558FF65F83BFEA8C21E" bold="true" box="[199,323,1369,1394]" pageId="1" pageNumber="142">Diagnosis:</emphasis>
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Differs from the other 6 known haplodesmid genera by only 19 body segments in both sexes, combined with a rather pyrgodesmid-like body shape and a special ozopore formula (5, 7–18), ozopores being borne on porosteles, as well as 4 transverse rows of setigerous isostictic tubercles per postcollum metatergum and a clearly helicoid, twisted prefemoral portion of the gonopod so that the seminal groove runs mostly laterally, not mesally (see also Key below).
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</paragraph>
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</subSubSection>
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<subSubSection id="C33A3C95734D2558FF65FB6FFCDDC0DE" pageId="1" pageNumber="142" type="description">
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<paragraph id="8B9F6F1E734D2558FF65FB6FFC2BC1B6" blockId="1.[151,1437,1294,2006]" pageId="1" pageNumber="142">
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<emphasis id="B954B30C734D2558FF65FB6FFEB0C14A" bold="true" box="[199,347,1549,1574]" pageId="1" pageNumber="142">Description:</emphasis>
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Body with 19 segments in both sexes, subcylindrical, not capable of volvation. Head, collum and following metaterga clothed with a dense cerategument. Ozopores borne on simple porosteles, pore formula 5, 7–18. Collum relatively large, sometimes covering the head from above. Paraterga 2 considerably enlarged, but not so as to ensure complete volvation, instead subtending the head laterally. Each postcollum metatergum usually with 4 transverse rows of isostictic setigerous tubercles. Telson strongly flattened dorsoventrally, subtruncate at tip, fully concealing a small epiproct tip from above. Spiracles seem to be absent.
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</paragraph>
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<paragraph id="8B9F6F1E734D2558FF65FB87FCDDC0DE" blockId="1.[151,1437,1294,2006]" pageId="1" pageNumber="142">
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Gonopods of a facies rather typical of
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<taxonomicName id="4C20149D734D2558FD27FB87FCDDC192" box="[645,822,1765,1790]" class="Diplopoda" family="Haplodesmidae" kingdom="Animalia" order="Polydesmida" pageId="1" pageNumber="142" phylum="Arthropoda" rank="family">Haplodesmidae</taxonomicName>
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,
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<emphasis id="B954B30C734D2558FCE1FB84FC60C191" box="[835,907,1766,1789]" italics="true" pageId="1" pageNumber="142">in situ</emphasis>
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both held nearly parallel to each other: coxae elongate, non-globose, subcylindrical, both well fused anteromedially, each carrying an elongated cannula on mesal face; telopodites elongated, slender, each clearly subdivided into an elongated, clearly helicoid, twisted prefemoral (= densely setose) part on which the seminal groove runs mostly laterally, not mesally; acropodites curved caudad, slender, uni- or biramous; seminal groove borne on a long solenomere, terminating apically, devoid both of an accessory seminal chamber and a hairy pulvillus.
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</paragraph>
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</subSubSection>
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<subSubSection id="C33A3C95734D255BFF65FADFFCC7C0AB" lastPageId="2" lastPageNumber="143" pageId="1" pageNumber="142" type="materials_examined">
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<paragraph id="8B9F6F1E734D2558FF65FADFFD55C0B9" blockId="1.[151,1437,1294,2006]" box="[199,702,1981,2006]" pageId="1" pageNumber="142">
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<emphasis id="B954B30C734D2558FF65FADFFEB0C0BA" bold="true" box="[199,347,1981,2006]" pageId="1" pageNumber="142">
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<typeStatus id="549BD1BC734D2558FF65FADFFEE9C0BA" box="[199,258,1981,2006]" pageId="1" pageNumber="142">Type</typeStatus>
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species
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</emphasis>
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:
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<taxonomicName id="4C20149D734D2558FEC8FADCFD8BC0B9" box="[362,608,1982,2005]" class="Diplopoda" family="Haplodesmidae" genus="Koponenius" kingdom="Animalia" order="Polydesmida" pageId="1" pageNumber="142" phylum="Arthropoda" rank="species" species="unicornis" status="sp. nov.">
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<emphasis id="B954B30C734D2558FEC8FADCFD8BC0B9" box="[362,608,1982,2005]" italics="true" pageId="1" pageNumber="142">Koponenius unicornis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2670E77734D2558FDC5FADCFD55C0B9" box="[615,702,1982,2005]" pageId="1" pageNumber="142" rank="species">sp. nov.</taxonomicNameLabel>
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</paragraph>
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<caption id="DF5F3F96734E255BFF35FB90FDDBC0EF" httpUri="https://zenodo.org/record/229193/files/figure.png" pageId="2" pageNumber="143" targetBox="[151,1436,195,1753]" targetPageId="2">
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<paragraph id="8B9F6F1E734E255BFF35FB90FDDBC0EF" blockId="2.[151,1436,1778,1923]" pageId="2" pageNumber="143">
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<emphasis id="B954B30C734E255BFF35FB90FEF9C06B" bold="true" box="[151,274,1778,1800]" pageId="2" pageNumber="143">FIGURE 2.</emphasis>
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<taxonomicName id="4C20149D734E255BFEB5FB91FE1DC064" box="[279,502,1779,1800]" class="Diplopoda" family="Haplodesmidae" genus="Koponenius" kingdom="Animalia" order="Polydesmida" pageId="2" pageNumber="143" phylum="Arthropoda" rank="species" species="unicornis" status="sp. nov.">
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<emphasis id="B954B30C734E255BFEB5FB91FE1DC064" box="[279,502,1779,1800]" italics="true" pageId="2" pageNumber="143">Koponenius unicornis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2670E77734E255BFE59FB91FDA0C064" box="[507,587,1779,1800]" pageId="2" pageNumber="143" rank="species">sp. nov.</taxonomicNameLabel>
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, male paratype. A, D, E, H, Q: anterior body part, lateral, ventral, ventral, dorsal and ventral views, respectively. B, F, I: midbody segments, lateral, ventral and dorsal views, respectively. C, G, J, K: posterior body part, lateral, ventral, dorsal and dorsal views, respectively. L: head, ventral view. M: cross-section of a midbody segment. N–P: tergal texture, dorsal views. R: midbody leg, subventral view. Scale bars: B, C, F, G, I, M, 0.5 mm; A, D, E, H, J–L, 0.2 mm; P–R, 0.1 mm; N, 0.05 mm; O, 0.02 mm.
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</paragraph>
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</caption>
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<paragraph id="8B9F6F1E734E255BFF65FACDFCC7C0AB" blockId="2.[199,1436,1967,2028]" box="[199,812,1967,1992]" pageId="2" pageNumber="143">
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<emphasis id="B954B30C734E255BFF65FACDFE34C0A4" bold="true" box="[199,479,1967,1992]" pageId="2" pageNumber="143">Other species included:</emphasis>
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<taxonomicName id="4C20149D734E255BFE44FAD2FD24C0A4" box="[486,719,1968,1992]" class="Diplopoda" family="Haplodesmidae" genus="Koponenius" kingdom="Animalia" order="Polydesmida" pageId="2" pageNumber="143" phylum="Arthropoda" rank="species" species="biramus" status="sp. nov.">
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<emphasis id="B954B30C734E255BFE44FAD2FD24C0A4" box="[486,719,1968,1992]" italics="true" pageId="2" pageNumber="143">Koponenius biramus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2670E77734E255BFD77FAD2FCC7C0AB" box="[725,812,1968,1991]" pageId="2" pageNumber="143" rank="species">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="C33A3C95734E255AFF65FAB1FA63C533" lastPageId="3" lastPageNumber="144" pageId="2" pageNumber="143" type="discussion">
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<paragraph id="8B9F6F1E734E255AFF65FAB1FD1DC670" blockId="2.[199,1436,1967,2028]" lastBlockId="3.[151,1437,151,608]" lastPageId="3" lastPageNumber="144" pageId="2" pageNumber="143">
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<emphasis id="B954B30C734E255BFF65FAB1FED6C080" bold="true" box="[199,317,2003,2028]" pageId="2" pageNumber="143">Remarks:</emphasis>
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The family
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<taxonomicName id="4C20149D734E255BFE7BFAB1FD6CC080" box="[473,647,2003,2028]" class="Diplopoda" family="Haplodesmidae" kingdom="Animalia" order="Polydesmida" pageId="2" pageNumber="143" phylum="Arthropoda" rank="family">Haplodesmidae</taxonomicName>
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hitherto contained six genera. Most of them can be conveniently divided into two morphological groups: (1) The “haplodesmid”
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<typeStatus id="549BD1BC734F255AFC22FDFAFC5BC7DC" box="[896,944,152,176]" pageId="3" pageNumber="144">type</typeStatus>
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usually shows ozopores borne on special, mostly boletiform porosteles or tubercles, while paraterga are absent from body segment 3 onwards, and (2) The “doratodesmid”
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<typeStatus id="549BD1BC734F255AFEF0FD82FE69C794" box="[338,386,224,248]" pageId="3" pageNumber="144">type</typeStatus>
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is normally devoid of porosteles, whereas the paraterga are present on all segments, often ensuring complete volvation (
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<bibRefCitation id="EFB112EF734F255AFE46FC66FD01C670" author="Golovatch" box="[484,746,260,285]" pageId="3" pageNumber="144" refString="Golovatch, S. I., Geoffroy, J. - J., Mauries, J. - P. & VandenSpiegel, D. (2009 a) Review of the millipede family Haplodesmidae Cook, 1895, with descriptions of some new or poorly-known species (Diplopoda, Polydesmida). ZooKeys, 7, 1 - 53. http: // dx. doi. org / 10.3897 / zookeys. 7.117" type="journal article" year="2009" yearSuffix="a">
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Golovatch
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<emphasis id="B954B30C734F255AFDC0FC67FD71C670" box="[610,666,260,284]" italics="true" pageId="3" pageNumber="144">et al.</emphasis>
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2009a
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</bibRefCitation>
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).
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</paragraph>
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<paragraph id="8B9F6F1E734F255AFF65FC4AFA63C533" blockId="3.[151,1437,151,608]" pageId="3" pageNumber="144">
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<taxonomicName id="4C20149D734F255AFF65FC4AFEA6C653" box="[199,333,296,319]" class="Diplopoda" family="Haplodesmidae" genus="Koponenius" kingdom="Animalia" order="Polydesmida" pageId="3" pageNumber="144" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B954B30C734F255AFF65FC4AFEA6C653" box="[199,333,296,319]" italics="true" pageId="3" pageNumber="144">Koponenius</emphasis>
|
||
</taxonomicName>
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||
<taxonomicNameLabel id="A2670E77734F255AFEF9FC4AFE23C653" box="[347,456,296,320]" pageId="3" pageNumber="144" rank="genus">gen. nov.</taxonomicNameLabel>
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||
seems to be especially similar to
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<taxonomicName id="4C20149D734F255AFCD6FC4AFB0BC62C" authority="Silvestri, 1910" authorityName="Silvestri" authorityYear="1910" box="[884,1248,295,320]" class="Diplopoda" family="Haplodesmidae" genus="Prosopodesmus" kingdom="Animalia" order="Polydesmida" pageId="3" pageNumber="144" phylum="Arthropoda" rank="genus">
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||
<emphasis id="B954B30C734F255AFCD6FC4AFBCFC62C" box="[884,1060,296,320]" italics="true" pageId="3" pageNumber="144">Prosopodesmus</emphasis>
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Silvestri, 1910
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</taxonomicName>
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, likewise being intermediate between the two groups described above in showing quite strongly developed paraterga, coupled with the presence of porosteles. However, all 7 known
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<taxonomicName id="4C20149D734F255AFD61FC12FC98C6E4" box="[707,883,368,392]" class="Diplopoda" family="Haplodesmidae" genus="Prosopodesmus" kingdom="Animalia" order="Polydesmida" pageId="3" pageNumber="144" phylum="Arthropoda" rank="genus">
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<emphasis id="B954B30C734F255AFD61FC12FC98C6E4" box="[707,883,368,392]" italics="true" pageId="3" pageNumber="144">Prosopodesmus</emphasis>
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</taxonomicName>
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species (mostly native to tropical
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<collectingCountry id="F3372F8E734F255AFB53FC0DFAB0C6E4" box="[1265,1371,367,392]" name="Australia" pageId="3" pageNumber="144">Australia</collectingCountry>
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, but a few to southern
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<collectingCountry id="F3372F8E734F255AFEC1FCF6FE4EC6C0" box="[355,421,404,428]" name="Japan" pageId="3" pageNumber="144">Japan</collectingCountry>
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) are considerably more pyrgodesmid-like in body shape, the collum being strongly flabellate, its fore margin 12-lobulated, covering the head from above. In addition, they have 20 body segments, only 3 transverse rows of isostictic tubercles per postcollum metatergum, varying pore formulas, but not (5, 7–18), usually mushroom-shaped porosteles, and the seminal groove on falcate and unbranched gonopod telopodites which runs only on the mesal side and is typically supplied with a hairy pulvillus near its orifice (
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<bibRefCitation id="EFB112EF734F255AFB4AFF46FA7AC550" author="Mesibov" box="[1256,1425,548,573]" pageId="3" pageNumber="144" refString="Mesibov, R. E. (2012) New species of Prosopodesmus Silvestri, 1910 (Diplopoda, Polydesmida, Haplodesmidae) from Queensland, Australia. ZooKeys, 190, 33 - 54. http: // dx. doi. org / 10.3897 / zookeys. 190.3276" type="journal article" year="2012">Mesibov 2012</bibRefCitation>
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). The above similarities between these two genera are also reflected in an updated key which concludes this paper.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |