treatments-xml/data/03/89/00/0389006BFF815A19FF5CFC34C1F6FD89.xml

<document id="0631785F3ADD9AF85CEEC0069C25FF94" ID-CLB-Dataset="55493" ID-DOI="10.11646/phytotaxa.547.1.6" ID-GBIF-Dataset="7b100ae6-1e4b-4da3-bb71-1c1cd01ea70c" ID-ISSN="1179-3163" ID-Zenodo-Dep="6555995" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1652771493808" checkinUser="plazi" docAuthor="Lin, Che-Wei, Hsu, Tian-Chuan, Luu, Hong Truong, Yang, Tsung-Yu Aleck &amp; Li, Chia- Wei" docDate="2022" docId="0389006BFF815A19FF5CFC34C1F6FD89" docLanguage="en" docName="phytotaxa.547.1.6.pdf" docOrigin="Phytotaxa 547 (1)" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.7:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="7" docTitle="Nephoanthus C. W. Lin &amp; T. C. Hsu 2022, gen. nov." docType="treatment" docVersion="8" lastPageNumber="69" masterDocId="FFB07813FF805A1AFFD4FFC0C013FFE1" masterDocTitle="Nephoanthus (Melastomataceae: Sonerileae), a new genus segregated from Phyllagathis s. l., with a new species from Southern Vietnam" masterLastPageNumber="76" masterPageNumber="66" pageNumber="67" updateTime="1699333075086" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="EBCC28E54D00EB0871A6F4AD72A379D6">Nephoanthus (Melastomataceae: Sonerileae), a new genus segregated from Phyllagathis s. l., with a new species from Southern Vietnam</mods:title>
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<mods:namePart id="C692E0F5E7AF356DE13BD1E6D78283FF">Lin, Che-Wei</mods:namePart>
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<mods:affiliation id="792B4D1CB629282AE95C2DFAE5897021">Botanical Garden Division, Taiwan Forestry Research Institute, Taipei 10066, Taiwan &amp; varalba @ gmail. com; https: // orcid. org / 0000 - 0002 - 8564 - 5793</mods:affiliation>
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<mods:namePart id="499E2650AA753AE90C18686CC37AB681">Hsu, Tian-Chuan</mods:namePart>
<mods:affiliation id="EA16B4F096B55EF7FE22C66CFC6261B6">Botanical Garden Division, Taiwan Forestry Research Institute, Taipei 10066, Taiwan &amp; Department of Life Sciences, National Chung Hsing University, Taichung 40227, Taiwan</mods:affiliation>
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<mods:namePart id="063E565A41B09D80F4A83FD8C7B0411F">Luu, Hong Truong</mods:namePart>
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<mods:affiliation id="0426BD2CD7C838F0BF001AF494B176C1">Southern Institute of Ecology, Institute of Applied Materials Science &amp; Graduate University of Science and Technology, Vietnam Academy of Science and Technology, Ho Chi Minh City, Vietnam &amp; hongtruongluu @ gmail. com; https: // orcid. org / 0000 - 0002 - 7036 - 7081</mods:affiliation>
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<mods:namePart id="6EB60AE579A3D1F78A757C1AC3F16A4C">Yang, Tsung-Yu Aleck</mods:namePart>
<mods:affiliation id="D4DB507F96AE5C31B3B9EF94C1F853F2">Biology Department, National Museum of Natural Science, Taichung 40453, Taiwan &amp; Department of Life Sciences, National Chung Hsing University, Taichung 40227, Taiwan</mods:affiliation>
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<mods:namePart id="DDA84AD7D01FF4EBAAE3549141810523">Li, Chia- Wei</mods:namePart>
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<mods:affiliation id="AA55D5FDA3248B5D944DC8F22805664D">Institute of Molecular and Cellular Biology, National Tsing Hua University, Hsinchu 30013, Taiwan &amp; cwli @ life. nthu. edu. tw; https: // orcid. org / 0000 - 0002 - 6280 - 0877</mods:affiliation>
<mods:nameIdentifier id="F5F7CF4A7E0B41C6C65725D4FC16A7FF" type="email">cwli@life.nthu.edu.tw</mods:nameIdentifier>
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<mods:date id="6CB6AD4131F855EBD24DFFE3ED692D0B">2022</mods:date>
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<taxonomicName id="4C20CAFEFF815A1BFF5CFC34C218FBEF" ID-CoL="992CR" authority="C.W.Lin &amp; T.C.Hsu" authorityName="C. W. Lin &amp; T. C. Hsu" authorityYear="2022" box="[136,523,1012,1039]" class="Magnoliopsida" family="Melastomataceae" genus="Nephoanthus" kingdom="Plantae" order="Myrtales" pageId="1" pageNumber="67" phylum="Tracheophyta" rank="genus" status="gen. nov.">
<emphasis id="B9546D6FFF815A1BFF5CFC34C137FBEE" bold="true" box="[136,292,1012,1039]" italics="true" pageId="1" pageNumber="67">Nephoanthus</emphasis>
C.W.Lin &amp; T.C.Hsu
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<emphasis id="B9546D6FFF815A1BFDC5FC35C265FBEF" box="[529,630,1013,1038]" italics="true" pageId="1" pageNumber="67">
<taxonomicNameLabel id="A267D014FF815A1BFDC5FC35C265FBEF" box="[529,630,1013,1038]" pageId="1" pageNumber="67" rank="genus">gen. nov.</taxonomicNameLabel>
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<subSubSection id="C33AE2F6FF815A1BFF5CFB80C262FBB7" box="[136,625,1088,1110]" pageId="1" pageNumber="67" type="type_taxon">
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<emphasis id="B9546D6FFF815A1BFF5CFB80C0D1FBB4" bold="true" box="[136,194,1088,1109]" pageId="1" pageNumber="67">
<typeStatus id="549B0FDFFF815A1BFF5CFB80C0ADFBB4" box="[136,190,1088,1109]" pageId="1" pageNumber="67">Type</typeStatus>
:
</emphasis>
— 
<taxonomicName id="4C20CAFEFF815A1BFF0EFB80C27DFBB7" ID-CoL="4H32Y" authority="Hansen (1990: 39)" authorityName="Hansen" authorityPageNumber="39" authorityYear="1990" box="[218,622,1088,1110]" class="Magnoliopsida" family="Melastomataceae" genus="Phyllagathis" kingdom="Plantae" order="Myrtales" pageId="1" pageNumber="67" phylum="Tracheophyta" rank="species" species="prostrata">
<emphasis id="B9546D6FFF815A1BFF0EFB80C1A1FBB4" box="[218,434,1088,1109]" italics="true" pageId="1" pageNumber="67">Phyllagathis prostrata</emphasis>
<bibRefCitation id="EFB1CC8CFF815A1BFE6CFB80C27DFBB7" author="Hansen, C." box="[440,622,1088,1110]" pageId="1" pageNumber="67" pagination="37 - 41" refId="ref5171" refString="Hansen, C. (1990) New species and combinations in Allomorphia, Phyllagathis and Sporoxeia (Melastomataceae) in Indo-China. Bulletin du Museum National d'Histoire Naturelle, Section B, Adansonia, 4 e serie 12 (1): 37 - 41." type="journal article" year="1990">Hansen (1990: 39)</bibRefCitation>
</taxonomicName>
.
</paragraph>
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<subSubSection id="C33AE2F6FF815A1BFF5CFBA3C253FAEB" pageId="1" pageNumber="67" type="diagnosis">
<paragraph id="8B9FB17DFF815A1BFF5CFBA3C253FAEB" blockId="1.[136,1452,1088,1290]" pageId="1" pageNumber="67">
<emphasis id="B9546D6FFF815A1BFF5CFBA3C0E0FB98" bold="true" box="[136,243,1123,1145]" pageId="1" pageNumber="67">Diagnosis:</emphasis>
— 
<taxonomicName id="4C20CAFEFF815A1BFEDFFBA4C199FB98" box="[267,394,1124,1145]" class="Magnoliopsida" family="Melastomataceae" genus="Nephoanthus" kingdom="Plantae" order="Myrtales" pageId="1" pageNumber="67" phylum="Tracheophyta" rank="genus">
<emphasis id="B9546D6FFF815A1BFEDFFBA4C199FB98" box="[267,394,1124,1145]" italics="true" pageId="1" pageNumber="67">Nephoanthus</emphasis>
</taxonomicName>
resembles 
<taxonomicName id="4C20CAFEFF815A1BFE23FBA4C293FB98" authorityName="Chen" authorityYear="1979" box="[503,640,1124,1145]" class="Magnoliopsida" family="Melastomataceae" genus="Tigridiopalma" kingdom="Plantae" order="Myrtales" pageId="1" pageNumber="67" phylum="Tracheophyta" rank="genus">
<emphasis id="B9546D6FFF815A1BFE23FBA4C293FB98" box="[503,640,1124,1145]" italics="true" pageId="1" pageNumber="67">Tigridiopalma</emphasis>
</taxonomicName>
in the herbaceous habit, ovate leaves and stamens with the decurrent connective of anthers forming an spur dorsally and two erect lobes ventrally. However, it is distinct from 
<taxonomicName id="4C20CAFEFF815A1BFBCBFB48C4BBFB7C" authorityName="Chen" authorityYear="1979" box="[1055,1192,1160,1181]" class="Magnoliopsida" family="Melastomataceae" genus="Tigridiopalma" kingdom="Plantae" order="Myrtales" pageId="1" pageNumber="67" phylum="Tracheophyta" rank="genus">
<emphasis id="B9546D6FFF815A1BFBCBFB48C4BBFB7C" box="[1055,1192,1160,1181]" italics="true" pageId="1" pageNumber="67">Tigridiopalma</emphasis>
</taxonomicName>
in its much smaller foliage by 3–11 × 
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(vs. 20–51.8 × 
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), umbellate (vs. scorpioid) inflorescences, terete or slightly 8- or 10-ribbed (vs. 5-winged) hypanthia, isomorphic (vs. dimorphic) anthers, and old capsules with distally horned (vs. not horned) placental columns and thready (vs. non-thready) placentas.
</paragraph>
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<subSubSection id="C33AE2F6FF815A1BFF5CFAF8C465F872" pageId="1" pageNumber="67" type="description">
<paragraph id="8B9FB17DFF815A1BFF5CFAF8C465F872" blockId="1.[136,1452,1336,2047]" pageId="1" pageNumber="67">
<emphasis id="B9546D6FFF815A1BFF5CFAF8C0C2FAB3" bold="true" box="[136,209,1336,1362]" pageId="1" pageNumber="67">Herbs</emphasis>
caulescent, perennial. 
<emphasis id="B9546D6FFF815A1BFE1BFAF8C204FAB3" bold="true" box="[463,535,1336,1362]" pageId="1" pageNumber="67">Stems</emphasis>
slender, terete or ribbed, basally creeping, rooting at nodes and branched, apically creeping or ascending, velutinous or hirsute. 
<emphasis id="B9546D6FFF815A1BFDA7FA9CC2D6FA97" bold="true" box="[627,709,1372,1398]" pageId="1" pageNumber="67">Leaves</emphasis>
opposite, isomorphic, equal to subequal, 7-nerved, ovate to broadly ovate, velutinous or hirsute, trichomes 
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long, upper surface flat to rugose, base cordate, apex acuminate to rounded, margin denticulate or crenulate, ciliate. 
<emphasis id="B9546D6FFF815A1BFD61FA64C302FA5F" bold="true" box="[693,785,1444,1470]" pageId="1" pageNumber="67">Petioles</emphasis>
subterete and slightly grooved adaxially, pilose, trichomes 
<quantity id="4CD81C98FF815A1BFF5CFA08C0FAFA02" box="[136,233,1480,1507]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="5.0" metricValueMin="2.0" pageId="1" pageNumber="67" unit="mm" value="3.5" valueMax="5.0" valueMin="2.0">2–5 mm</quantity>
long. 
<emphasis id="B9546D6FFF815A1BFEECFA08C1F2FA03" bold="true" box="[312,481,1480,1506]" pageId="1" pageNumber="67">Inflorescences</emphasis>
terminal or rarely axillary, umbellate, nearly capitate, peduncles short, glabrous or sparsely hirsute, bracts persistent, deltoid, ovate to cordate, apex attenuate to obtuse, abaxial surface hairy, margin ciliate. 
<emphasis id="B9546D6FFF815A1BFF0DF9D0C12BF9CB" bold="true" box="[217,312,1552,1578]" pageId="1" pageNumber="67">Flowers</emphasis>
4- or 5-merous, shortly pedicellate. 
<emphasis id="B9546D6FFF815A1BFD1CF9D0C373F9CB" bold="true" box="[712,864,1552,1578]" pageId="1" pageNumber="67">Hypanthium</emphasis>
cup-shaped, terete or slightly 8- or 10-ribbed, pilose. 
<emphasis id="B9546D6FFF815A1BFF5CF9F4C0C7F9AF" bold="true" box="[136,212,1588,1614]" pageId="1" pageNumber="67">Sepals</emphasis>
triangular and keeled, apex aristate, each with a subapical fleshy trichome, trichomes multi-branched, 
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long; sepals connate at the base and forming a ring, hirsute; perishing with the tubular part of hypanthium shortly after flowering. 
<emphasis id="B9546D6FFF815A1BFE95F9BCC19AF977" bold="true" box="[321,393,1660,1686]" pageId="1" pageNumber="67">Petals</emphasis>
oblique, elliptic, obovate to oblong, rosy pink, apex mucronate-apiculate, glabrous or with a few apical glandular trichomes dorsally. 
<emphasis id="B9546D6FFF815A1BFD8EF960C2ACF95B" bold="true" box="[602,703,1696,1722]" pageId="1" pageNumber="67">Stamens</emphasis>
isomorphic, filaments slightly flat, white to purplish-red, anthers subulate, apex attenuate, slightly curved ventrally, pale pink to purplish, pore 1, connective extended below anther sacs and forming an ovate, yellow or orangish-red spur dorsally and two erect narrowly triangular, yellow or orangishred lobes ventrally. 
<emphasis id="B9546D6FFF815A1BFEB2F8CCC1B2F8C7" bold="true" box="[358,417,1804,1830]" pageId="1" pageNumber="67">Style</emphasis>
white to pink, stigma capitulate; Ovary 
<emphasis id="B9546D6FFF815A1BFCB3F8CDC39AF8C7" box="[871,905,1805,1830]" italics="true" pageId="1" pageNumber="67">ca.</emphasis>
1/2 as long as the hypanthium, crown with fully connate lobes, margin entire or denticulate with some minute uni-seriate glandular trichomes, anther pockets shallow, placentae stalked. 
<emphasis id="B9546D6FFF815A1BFE88F894C1D4F88F" bold="true" box="[348,455,1876,1902]" pageId="1" pageNumber="67">Capsules</emphasis>
shortly pedicellate, hypanthium cup-shaped, 8 or 10-ribbed, placental column distally horned, placentas thready. 
<emphasis id="B9546D6FFF815A1BFE67F8B8C1E5F873" bold="true" box="[435,502,1912,1938]" pageId="1" pageNumber="67">Seeds</emphasis>
numerous, pale brown, cuneate, 0.3–0.4 × 
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.
</paragraph>
</subSubSection>
<subSubSection id="C33AE2F6FF815A1BFF69F85CC288F83A" pageId="1" pageNumber="67" type="etymology">
<paragraph id="8B9FB17DFF815A1BFF69F85CC288F83A" blockId="1.[136,1452,1336,2047]" pageId="1" pageNumber="67">
<emphasis id="B9546D6FFF815A1BFF69F85CC15BF857" bold="true" box="[189,328,1948,1974]" pageId="1" pageNumber="67">Etymology:</emphasis>
—The generic epithet is derived from Greek 
<emphasis id="B9546D6FFF815A1BFCE8F85CC398F857" box="[828,907,1948,1974]" italics="true" pageId="1" pageNumber="67">nephos</emphasis>
, cloud, and 
<emphasis id="B9546D6FFF815A1BFBC7F85CC44DF857" box="[1043,1118,1948,1974]" italics="true" pageId="1" pageNumber="67">anthus</emphasis>
, flower, referring to the cloud forest habitat where species of this genus grow.
</paragraph>
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<subSubSection id="C33AE2F6FF815A1BFF69F824C376F81F" box="[189,869,2020,2047]" pageId="1" pageNumber="67" type="distribution">
<paragraph id="8B9FB17DFF815A1BFF69F824C376F81F" blockId="1.[136,1452,1336,2047]" box="[189,869,2020,2047]" pageId="1" pageNumber="67">
<emphasis id="B9546D6FFF815A1BFF69F824C14AF81F" bold="true" box="[189,345,2020,2046]" pageId="1" pageNumber="67">Distribution:</emphasis>
— 
<collectingCountry id="F337F1EDFF815A1BFEA1F824C1C7F81F" box="[373,468,2020,2046]" name="Vietnam" pageId="1" pageNumber="67">Vietnam</collectingCountry>
and 
<collectingCountry id="F337F1EDFF815A1BFDDEF824C25EF81F" box="[522,589,2020,2046]" name="China" pageId="1" pageNumber="67">China</collectingCountry>
(
<collectingRegion id="49E47F9FFF815A1BFD88F824C2BCF81F" box="[604,687,2020,2046]" country="China" name="Hainan" pageId="1" pageNumber="67">Hainan</collectingRegion>
Island) (
<figureCitation id="131BADF8FF815A1BFCC6F824C347F81F" box="[786,852,2020,2046]" captionStart="FIGURE 1" captionStartId="2.[136,229,879,901]" captionTargetBox="[195,1391,175,855]" captionTargetId="figure-18@2.[195,1391,175,855]" captionTargetPageId="2" captionText="FIGURE 1. Distribution map of Nephoanthus nubigenus (yellow star) and N. prostratus (pink circle)." figureDoi="http://doi.org/10.5281/zenodo.6556004" httpUri="https://zenodo.org/record/6556004/files/figure.png" pageId="1" pageNumber="67">Fig. 1</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="DF5FE1F5FF825A18FF5CFCAFC47EFC64" ID-DOI="http://doi.org/10.5281/zenodo.6556004" ID-Zenodo-Dep="6556004" box="[136,1133,879,901]" httpUri="https://zenodo.org/record/6556004/files/figure.png" pageId="2" pageNumber="68" startId="2.[136,229,879,901]" targetBox="[195,1391,175,855]" targetPageId="2">
<paragraph id="8B9FB17DFF825A18FF5CFCAFC47EFC64" blockId="2.[136,1133,879,901]" box="[136,1133,879,901]" pageId="2" pageNumber="68">
<emphasis id="B9546D6FFF825A18FF5CFCAFC0EDFC65" bold="true" box="[136,254,879,901]" pageId="2" pageNumber="68">FIGURE 1.</emphasis>
Distribution map of 
<taxonomicName id="4C20CAFEFF825A18FE13FCAFC2A9FC64" authorityName="C. W. Lin, Luu &amp; T. C. Hsu. A. Habit and Habitat" authorityYear="2022" box="[455,698,879,901]" class="Magnoliopsida" family="Melastomataceae" genus="Nephoanthus" kingdom="Plantae" order="Myrtales" pageId="2" pageNumber="68" phylum="Tracheophyta" rank="species" species="nubigenus">
<emphasis id="B9546D6FFF825A18FE13FCAFC2A9FC64" bold="true" box="[455,698,879,901]" italics="true" pageId="2" pageNumber="68">Nephoanthus nubigenus</emphasis>
</taxonomicName>
(yellow star) and 
<taxonomicName id="4C20CAFEFF825A18FCBDFCAFC3F9FC65" authorityName="C. W. Lin &amp; T. C. Hsu. A, B. Habit" authorityYear="2022" baseAuthorityName="C. W. Lin &amp; T. C. Hsu. A, B. Habit" baseAuthorityYear="2022" box="[873,1002,879,900]" class="Magnoliopsida" family="Melastomataceae" genus="Nephoanthus" kingdom="Plantae" order="Myrtales" pageId="2" pageNumber="68" phylum="Tracheophyta" rank="species" species="prostratus">
<emphasis id="B9546D6FFF825A18FCBDFCAFC3F9FC65" bold="true" box="[873,1002,879,900]" italics="true" pageId="2" pageNumber="68">N. prostratus</emphasis>
</taxonomicName>
(pink circle).
</paragraph>
</caption>
<caption id="DF5FE1F5FF825A18FF5CF82FC4E3F7C8" ID-DOI="http://doi.org/10.5281/zenodo.6556006" ID-Zenodo-Dep="6556006" httpUri="https://zenodo.org/record/6556006/files/figure.png" pageId="2" pageNumber="68" startId="2.[136,229,2031,2053]" targetBox="[195,1391,927,2007]" targetPageId="2">
<paragraph id="8B9FB17DFF825A18FF5CF82FC4E3F7C8" blockId="2.[136,1452,2031,2089]" pageId="2" pageNumber="68">
<emphasis id="B9546D6FFF825A18FF5CF82FC0EDF7E5" bold="true" box="[136,254,2031,2053]" pageId="2" pageNumber="68">FIGURE 2.</emphasis>
Two taxa morphologically similar to 
<taxonomicName id="4C20CAFEFF825A18FDBDF82FC2FBF7E5" box="[617,744,2031,2052]" class="Magnoliopsida" family="Melastomataceae" genus="Nephoanthus" kingdom="Plantae" order="Myrtales" pageId="2" pageNumber="68" phylum="Tracheophyta" rank="genus">
<emphasis id="B9546D6FFF825A18FDBDF82FC2FBF7E5" box="[617,744,2031,2052]" italics="true" pageId="2" pageNumber="68">Nephoanthus</emphasis>
</taxonomicName>
: 
<taxonomicName id="4C20CAFEFF825A18FD20F82FC3FEF7E4" authorityName="Blume" authorityYear="1831" baseAuthorityName="Jack" baseAuthorityYear="1823" box="[756,1005,2031,2053]" class="Magnoliopsida" family="Melastomataceae" genus="Phyllagathis" kingdom="Plantae" order="Myrtales" pageId="2" pageNumber="68" phylum="Tracheophyta" rank="species" species="rotundifolia">
<emphasis id="B9546D6FFF825A18FD20F82FC3FEF7E4" bold="true" box="[756,1005,2031,2053]" italics="true" pageId="2" pageNumber="68">Phyllagathis rotundifolia</emphasis>
</taxonomicName>
(A &amp; B); 
<taxonomicName id="4C20CAFEFF825A18FB9BF82FC555F7E4" box="[1103,1350,2031,2053]" class="Magnoliopsida" family="Melastomataceae" genus="Tigridiopalma" kingdom="Plantae" order="Myrtales" pageId="2" pageNumber="68" phylum="Tracheophyta" rank="species" species="magnifica">
<emphasis id="B9546D6FFF825A18FB9BF82FC555F7E4" bold="true" box="[1103,1350,2031,2053]" italics="true" pageId="2" pageNumber="68">Tigridiopalma magnifica</emphasis>
</taxonomicName>
(C−E). 
<emphasis id="B9546D6FFF825A18FA40F82FC5BFF7E5" bold="true" box="[1428,1452,2031,2052]" pageId="2" pageNumber="68">A.</emphasis>
Habit; 
<emphasis id="B9546D6FFF825A18FF1EF7D4C0F3F7C8" bold="true" box="[202,224,2068,2089]" pageId="2" pageNumber="68">B.</emphasis>
Inflorescence; 
<emphasis id="B9546D6FFF825A18FEA0F7D3C198F7C8" bold="true" box="[372,395,2067,2089]" pageId="2" pageNumber="68">C.</emphasis>
Habit; 
<emphasis id="B9546D6FFF825A18FE07F7D4C214F7C8" bold="true" box="[467,519,2068,2089]" pageId="2" pageNumber="68">D, E.</emphasis>
Flower, face and side views. A &amp; B by Che-Wei Lin; C−E by Wei-Yen Chen.
</paragraph>
</caption>
<subSubSection id="C33AE2F6FF835A19FF69FF5EC1F6FD89" pageId="3" pageNumber="69" type="discussion">
<paragraph id="8B9FB17DFF835A19FF69FF5EC1F6FD89" blockId="3.[136,1452,158,616]" pageId="3" pageNumber="69">
<emphasis id="B9546D6FFF835A19FF69FF5EC1A6FF59" bold="true" box="[189,437,158,184]" pageId="3" pageNumber="69">Taxonomic remarks:</emphasis>
—The gross morphology of 
<taxonomicName id="4C20CAFEFF835A19FD25FF5EC396FF59" box="[753,901,158,184]" class="Magnoliopsida" family="Melastomataceae" genus="Nephoanthus" kingdom="Plantae" order="Myrtales" pageId="3" pageNumber="69" phylum="Tracheophyta" rank="genus">
<emphasis id="B9546D6FFF835A19FD25FF5EC396FF59" box="[753,901,158,184]" italics="true" pageId="3" pageNumber="69">Nephoanthus</emphasis>
</taxonomicName>
is somewhat similar to 
<taxonomicName id="4C20CAFEFF835A19FB58FF5EC5B6FF59" authorityName="Blume" authorityYear="1831" baseAuthorityName="Jack" baseAuthorityYear="1823" box="[1164,1445,158,184]" class="Magnoliopsida" family="Melastomataceae" genus="Phyllagathis" kingdom="Plantae" order="Myrtales" pageId="3" pageNumber="69" phylum="Tracheophyta" rank="species" species="rotundifolia">
<emphasis id="B9546D6FFF835A19FB58FF5EC5B6FF59" box="[1164,1445,158,184]" italics="true" pageId="3" pageNumber="69">Phyllagathis rotundifolia</emphasis>
</taxonomicName>
, the 
<typeStatus id="549B0FDFFF835A19FF67FF03C0F0FF3C" box="[179,227,195,221]" pageId="3" pageNumber="69">type</typeStatus>
species of 
<taxonomicName id="4C20CAFEFF835A19FEBCFF02C1E5FF3D" authorityName="Lin &amp; Hsu &amp; Luu &amp; Yang &amp; Li" authorityYear="2022" box="[360,502,194,220]" class="Magnoliopsida" family="Melastomataceae" genus="Phyllagathis" kingdom="Plantae" order="Myrtales" pageId="3" pageNumber="69" phylum="Tracheophyta" rank="genus">
<emphasis id="B9546D6FFF835A19FEBCFF02C1E5FF3D" box="[360,502,194,220]" italics="true" pageId="3" pageNumber="69">Phyllagathis</emphasis>
</taxonomicName>
, as they share dwarf herbaceous habits, basally creeping stems and nearly capitate inflorescences embraced by large persistent bracts (
<figureCitation id="131BADF8FF835A19FD05FF26C304FEE1" box="[721,791,230,256]" captionStart="FIGURE 2" captionStartId="2.[136,229,2031,2053]" captionTargetBox="[195,1391,927,2007]" captionTargetId="figure-38@2.[195,1391,927,2007]" captionTargetPageId="2" captionText="FIGURE 2. Two taxa morphologically similar to Nephoanthus: Phyllagathis rotundifolia (A &amp; B); Tigridiopalma magnifica (C−E). A. Habit; B. Inflorescence; C. Habit; D, E. Flower, face and side views. A &amp; B by Che-Wei Lin; C−E by Wei-Yen Chen." figureDoi="http://doi.org/10.5281/zenodo.6556006" httpUri="https://zenodo.org/record/6556006/files/figure.png" pageId="3" pageNumber="69">Fig. 2</figureCitation>
, A &amp; B). However, 
<taxonomicName id="4C20CAFEFF835A19FC28FF26C483FEE1" box="[1020,1168,230,256]" class="Magnoliopsida" family="Melastomataceae" genus="Nephoanthus" kingdom="Plantae" order="Myrtales" pageId="3" pageNumber="69" phylum="Tracheophyta" rank="genus">
<emphasis id="B9546D6FFF835A19FC28FF26C483FEE1" box="[1020,1168,230,256]" italics="true" pageId="3" pageNumber="69">Nephoanthus</emphasis>
</taxonomicName>
is distinguishable by the absence of stellate minute brown glands on vegetative organs, pink to purplish anthers, and also by the presence of a pair of ventral appendages at the base of the anther sac. Because molecular data (
<bibRefCitation id="EFB1CC8CFF835A19FBE5FEEEC4EEFEA9" author="Zhou, Q. J. &amp; Lin, C. W. &amp; Dai, J. H. &amp; Zhou, R. C. &amp; Liu, Y." box="[1073,1277,302,328]" pageId="3" pageNumber="69" pagination="256 - 267" refId="ref5535" refString="Zhou, Q. J., Lin, C. W., Dai, J. H., Zhou, R. C. &amp; Liu, Y. (2019 a) Exploring the generic delimitation of Phyllagathis and Bredia (Melastomataceae): A combined nuclear and chloroplast DNA analysis. Journal of Systematics and Evolution 57 (3): 256 - 267. https: // doi. org / 10.1111 / jse. 12451" type="journal article" year="2019">
Zhou 
<emphasis id="B9546D6FFF835A19FBA1FEEFC4BDFEA9" box="[1141,1198,302,328]" italics="true" pageId="3" pageNumber="69">et al.</emphasis>
2019a
</bibRefCitation>
; 
<bibRefCitation id="EFB1CC8CFF835A19FADEFEEEC540FEA9" author="Zhou, Q. J. &amp; Dai, J. H. &amp; Lin, C. W. &amp; Denda, T. &amp; Zhou, R. C. &amp; Liu, Y." box="[1290,1363,302,328]" pageId="3" pageNumber="69" pagination="121 - 150" refId="ref5617" refString="Zhou, Q. J., Dai, J. H., Lin, C. W., Denda, T., Zhou, R. C. &amp; Liu, Y. (2019 b) Recircumscription of Bredia and resurrection of Tashiroea (Sonerileae, Melastomataceae) with description of a new species T. villosa. Phytokeys 127: 121 - 150. https: // doi. org / 10.3897 / phytokeys. 127.36608" type="journal article" year="2019">2019b</bibRefCitation>
; 
<bibRefCitation id="EFB1CC8CFF835A19FA8BFEEEC5B4FEA9" author="Zhou, Q. &amp; Lin, C. W. &amp; Ng, W. L. &amp; Dai, J. &amp; Denda, T. &amp; Zhou, R. &amp; Liu, Y." box="[1375,1447,302,328]" pageId="3" pageNumber="69" pagination="1477" refId="ref5699" refString="Zhou, Q., Lin, C. W., Ng, W. L., Dai, J., Denda, T., Zhou, R. &amp; Liu, Y. (2019 c) Analyses of plastome sequences improve phylogenetic resolution and provide new insight into the evolutionary history of Asian Sonerileae / Dissochaeteae. Frontiers in Plant Science 10: 1477. https: // doi. org / 10.3389 / fpls. 2019.01477" type="journal article" year="2019">2019c</bibRefCitation>
; 
<bibRefCitation id="EFB1CC8CFF835A19FF5CFE93C150FE8D" author="Zeng, S. J. &amp; Xu, Y. C. &amp; Wang, G. T. &amp; Jia, P. &amp; Cui, D. F." box="[136,323,338,365]" pageId="3" pageNumber="69" pagination="33 - 42" refId="ref5468" refString="Zeng, S. J., Xu, Y. C., Wang, G. T., Jia, P. &amp; Cui, D. F. (2021) Tigridiopalma exaltata, a new and endangered species of Melastomataceae in China. Phytokeys 176: 33 - 42. https: // doi. org / 10.3897 / phytokeys. 176.63619" type="journal article" year="2021">
Zeng 
<emphasis id="B9546D6FFF835A19FF1DFE93C111FE8D" box="[201,258,338,364]" italics="true" pageId="3" pageNumber="69">et al.</emphasis>
2021
</bibRefCitation>
) strongly supports the fact that 
<taxonomicName id="4C20CAFEFF835A19FD7EFE92C32DFE8D" box="[682,830,338,364]" class="Magnoliopsida" family="Melastomataceae" genus="Nephoanthus" kingdom="Plantae" order="Myrtales" pageId="3" pageNumber="69" phylum="Tracheophyta" rank="genus">
<emphasis id="B9546D6FFF835A19FD7EFE92C32DFE8D" box="[682,830,338,364]" italics="true" pageId="3" pageNumber="69">Nephoanthus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C20CAFEFF835A19FCA2FE93C404FE8D" authorityName="Blume" authorityYear="1831" baseAuthorityName="Jack" baseAuthorityYear="1823" box="[886,1047,338,364]" class="Magnoliopsida" family="Melastomataceae" genus="Phyllagathis" kingdom="Plantae" order="Myrtales" pageId="3" pageNumber="69" phylum="Tracheophyta" rank="species" species="rotundifolia">
<emphasis id="B9546D6FFF835A19FCA2FE93C404FE8D" box="[886,1047,338,364]" italics="true" pageId="3" pageNumber="69">P. rotundifolia</emphasis>
</taxonomicName>
are in different major clades within 
<taxonomicName id="4C20CAFEFF835A19FF5CFEB6C112FE71" authorityName="Triana" authorityYear="1866" box="[136,257,374,400]" class="Magnoliopsida" family="Melastomataceae" kingdom="Plantae" order="Myrtales" pageId="3" pageNumber="69" phylum="Tracheophyta" rank="tribe" tribe="Sonerileae">Sonerileae</taxonomicName>
, their morphological similarities are either symplesiomorphies or the result of parallel evolution. Within the superclade recognized in 
<bibRefCitation id="EFB1CC8CFF835A19FE06FE5AC2B5FE55" author="Zhou, Q. &amp; Lin, C. W. &amp; Ng, W. L. &amp; Dai, J. &amp; Denda, T. &amp; Zhou, R. &amp; Liu, Y." box="[466,678,410,436]" pageId="3" pageNumber="69" pagination="1477" refId="ref5699" refString="Zhou, Q., Lin, C. W., Ng, W. L., Dai, J., Denda, T., Zhou, R. &amp; Liu, Y. (2019 c) Analyses of plastome sequences improve phylogenetic resolution and provide new insight into the evolutionary history of Asian Sonerileae / Dissochaeteae. Frontiers in Plant Science 10: 1477. https: // doi. org / 10.3389 / fpls. 2019.01477" type="journal article" year="2019">
Zhou 
<emphasis id="B9546D6FFF835A19FDC0FE5BC25EFE55" box="[532,589,410,436]" italics="true" pageId="3" pageNumber="69">et al.</emphasis>
(2019c
</bibRefCitation>
; also see INTRODUCTION), 
<taxonomicName id="4C20CAFEFF835A19FC2AFE5AC481FE55" box="[1022,1170,410,436]" class="Magnoliopsida" family="Melastomataceae" genus="Nephoanthus" kingdom="Plantae" order="Myrtales" pageId="3" pageNumber="69" phylum="Tracheophyta" rank="genus">
<emphasis id="B9546D6FFF835A19FC2AFE5AC481FE55" box="[1022,1170,410,436]" italics="true" pageId="3" pageNumber="69">Nephoanthus</emphasis>
</taxonomicName>
is morphologically most similar to 
<taxonomicName id="4C20CAFEFF835A19FF28FE7EC18FFE39" authorityName="Chen" authorityYear="1979" box="[252,412,446,472]" class="Magnoliopsida" family="Melastomataceae" genus="Tigridiopalma" kingdom="Plantae" order="Myrtales" pageId="3" pageNumber="69" phylum="Tracheophyta" rank="genus">
<emphasis id="B9546D6FFF835A19FF28FE7EC18FFE39" box="[252,412,446,472]" italics="true" pageId="3" pageNumber="69">Tigridiopalma</emphasis>
</taxonomicName>
(
<figureCitation id="131BADF8FF835A19FE79FE7EC1E1FE39" box="[429,498,446,472]" captionStart="FIGURE 2" captionStartId="2.[136,229,2031,2053]" captionTargetBox="[195,1391,927,2007]" captionTargetId="figure-38@2.[195,1391,927,2007]" captionTargetPageId="2" captionText="FIGURE 2. Two taxa morphologically similar to Nephoanthus: Phyllagathis rotundifolia (A &amp; B); Tigridiopalma magnifica (C−E). A. Habit; B. Inflorescence; C. Habit; D, E. Flower, face and side views. A &amp; B by Che-Wei Lin; C−E by Wei-Yen Chen." figureDoi="http://doi.org/10.5281/zenodo.6556006" httpUri="https://zenodo.org/record/6556006/files/figure.png" pageId="3" pageNumber="69">Fig. 2</figureCitation>
, C–E) but it is easily distinguished by its much smaller foliage by 3–11 × 
<quantity id="4CD81C98FF835A19FA96FE7EC5B8FE38" box="[1346,1451,446,473]" metricMagnitude="-2" metricUnit="m" metricValue="4.35" metricValueMax="7.0" metricValueMin="1.7" pageId="3" pageNumber="69" unit="cm" value="4.35" valueMax="7.0" valueMin="1.7">1.7–7 cm</quantity>
(vs. 20–51.8 × 
<quantity id="4CD81C98FF835A19FEF9FE22C1DAFE1C" box="[301,457,482,509]" metricMagnitude="-1" metricUnit="m" metricValue="2.925" metricValueMax="4.49" metricValueMin="1.3599999999999999" pageId="3" pageNumber="69" unit="cm" value="29.25" valueMax="44.9" valueMin="13.6">13.6–44.9 cm</quantity>
), umbellate (vs. scorpioid) inflorescences, terete or slightly 8- or 10-ribbed (vs. 5-winged) hypanthia, isomorphic (vs. dimorphic) anthers, and old capsules with distally horned (vs. not horned) placental columns and thready (vs. non-thready) placentas. A detailed comparison between 
<taxonomicName id="4C20CAFEFF835A19FC7BFDEAC450FDA5" box="[943,1091,554,580]" class="Magnoliopsida" family="Melastomataceae" genus="Nephoanthus" kingdom="Plantae" order="Myrtales" pageId="3" pageNumber="69" phylum="Tracheophyta" rank="genus">
<emphasis id="B9546D6FFF835A19FC7BFDEAC450FDA5" box="[943,1091,554,580]" italics="true" pageId="3" pageNumber="69">Nephoanthus</emphasis>
</taxonomicName>
and phylogenetically allied taxa (
<figureCitation id="131BADF8FF835A19FF44FD8EC0C0FD89" box="[144,211,590,616]" captionStart="FIGURE 3" captionStartId="5.[136,229,1732,1754]" captionTargetBox="[227,1360,190,1708]" captionTargetId="figure-20@5.[227,1360,190,1708]" captionTargetPageId="5" captionText="FIGURE 3. A selection of taxa within the superclade recognized by Zhou et al. (2019c) that contains Nephoanthus, showing varied habits and morphological characters: Driessenia inaequalifolia (M), D. phasmolacuna (K &amp; L); Medinilla laxiflora (B), M. speciosa (G &amp; H), M. sp. (I &amp; J); Phyllagathis gymnantha (N), P. rajah (A, C &amp; D), P. stellata (E &amp; F). A. Terrestrial in lowland mixed dipterocarp forest; B. Understory epiphyte on tree trunks and branches; C. Pleiochasium; D. Side view of flower, appendages firmly connected with the anther sacs, concolorous, hypanthium 10-ribbed; E. Umbelliform inflorescence; F. Side view of flower, anthers with concolorous appendages, connective slightly prolonged below anther sacs, hypanthium 10-ribbed; G. Paniculate inflorescence; H. Face view of flower, appendages firmly connected with the anther sacs, discolorous; I. Paniculate inflorescence; J. Face view of flower, appendages firmly connected with the anther sacs, concolorous; K. Thyrse, also showing the old capsule as empty cups; L. Face view of flower, anthers with concolorous appendages, connective slightly prolonged below anther sacs; M. Old capsule as empty cups; N. Old capsules, ovary with 4-horned placental column, placentas thready.All photos by Che-Wei Lin." figureDoi="http://doi.org/10.5281/zenodo.6556011" httpUri="https://zenodo.org/record/6556011/files/figure.png" pageId="3" pageNumber="69">Fig. 3</figureCitation>
) is presented in 
<tableCitation id="C6A284C6FF835A19FE5FFD8EC1F6FD89" box="[395,485,590,616]" captionStart="TABLE 1" captionStartId="3.[136,224,664,688]" captionTargetPageId="3" captionText="TABLE 1. Morphological comparison of Nephoanthus and phylogenetically allied taxa revealed in Zhou et al. (2019c)." httpUri="http://table.plazi.org/id/DF5FE1F5FF835A19FF5CFD58C595FD51" pageId="3" pageNumber="69" tableUuid="DF5FE1F5FF835A19FF5CFD58C595FD51">Table 1.</tableCitation>
</paragraph>
</subSubSection>
</treatment>
</document>