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<document id="A7756E3A4EA84991751C7FD52F4B7AAD" ID-CLB-Dataset="301495" ID-DOI="10.3897/zookeys.1209.127951" ID-GBIF-Dataset="260efa91-98a5-4abe-96de-636bdebff8e8" ID-ZooBank="A07CCEE1-C2D5-4993-9437-55E3040E1F79" ID-publisher-id="127951" URI-arpha="64DE4B4F-BF17-5208-91E2-23C3DD2B612B" URI-zoobank="http://zoobank.org/A07CCEE1-C2D5-4993-9437-55E3040E1F79" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" XM.treatments_approvedBy="admin" article-type="research-article" checkinTime="1723647139662" checkinUser="pensoft" docAuthor="Zhang, Ludan, Wang, Ying, Li, Shuqiang, Zhang, Xiaoqing &amp; Yao, Zhiyuan" docDate="2024" docId="370E6028657B58B5B22C14AC2195FE63" docLanguage="en" docName="ZooKeys 1209: 315-330" docOrigin="ZooKeys 1209" docSource="https://zookeys.pensoft.net/article/127951/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Belisana tianyang Zhang, Li &amp; Yao 2024, sp. nov." docType="treatment" docUuid="4D5E7BF7-2704-4467-9EE5-056C1FFE1D33" docUuidSource="ZooBank" docVersion="3" dtd-version="3.0" lastPageNumber="330" masterDocId="64DE4B4FBF17520891E223C3DD2B612B" masterDocTitle="Three new spider species of Belisana Thorell, 1898 (Araneae, Pholcidae) from karst caves, with a list of Belisana species from Guangxi, China" masterLastPageNumber="330" masterPageNumber="315" pageNumber="315" updateTime="1723650954597" updateUser="admin">
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<mods:title id="63CD0F9A3C8633BCEB19169FEA829EB0">Three new spider species of Belisana Thorell, 1898 (Araneae, Pholcidae) from karst caves, with a list of Belisana species from Guangxi, China</mods:title>
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<mods:namePart id="74B933AD35782447244EECFAC22AE272">Zhang, Ludan</mods:namePart>
<mods:nameIdentifier id="420A6539CABE1EE1A5D1BC953D8A8774" type="ORCID">0000-0001-5657-751X</mods:nameIdentifier>
<mods:affiliation id="8CB903379088117028D87905214DCC38">College of Life Science, Shenyang Normal University, Shenyang 110034, Liaoning, China</mods:affiliation>
</mods:name>
<mods:name id="70D6B89E9FFC8FB2F6659ECF578FA53C" type="personal">
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<mods:namePart id="7761B1384CE55C4EB2221CFC715F723C">Wang, Ying</mods:namePart>
<mods:affiliation id="81A73221081D46E821507DB38DBEBAFB">College of Life Science, Shenyang Normal University, Shenyang 110034, Liaoning, China</mods:affiliation>
</mods:name>
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<mods:namePart id="DA34C0E7574951BEDFB87934BA76FAA1">Li, Shuqiang</mods:namePart>
<mods:nameIdentifier id="C7CD5DD80FA386719B9F710D23006CA3" type="ORCID">0000-0002-3290-5416</mods:nameIdentifier>
<mods:affiliation id="13FA03558EE993AC95936FDD143C22F4">Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China</mods:affiliation>
</mods:name>
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<mods:namePart id="4AC8D4580E3DDE02B9CDD657DE65F3CA">Zhang, Xiaoqing</mods:namePart>
<mods:affiliation id="B1E9ADB6836608D0D85C915E7E8EF20E">College of Life Science, Shenyang Normal University, Shenyang 110034, Liaoning, China</mods:affiliation>
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<mods:namePart id="89B5801A961FC02C504C656917572A51">Yao, Zhiyuan</mods:namePart>
<mods:nameIdentifier id="C1DD1AD60B832C6A713B018CD53E2D75" type="ORCID">0000-0002-1631-0949</mods:nameIdentifier>
<mods:affiliation id="8F778DB7D3F3BF2C70C4CE2DE3C97C0B">College of Life Science, Shenyang Normal University, Shenyang 110034, Liaoning, China</mods:affiliation>
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<mods:date id="3F7F02D880910B958ACC1856983FD536">2024</mods:date>
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<mods:number id="66343F585E39A7A23ABD7983885E9A77">2024-08-14</mods:number>
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<mods:detail id="D06D2B37B4AF54D50C8267E5738127BB" type="volume">
<mods:number id="D21E74195B6DE1853F36DD81BE4F0CC9">1209</mods:number>
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<treatment id="370E6028657B58B5B22C14AC2195FE63" ID-DOI="http://doi.org/10.5281/zenodo.13323012" ID-Zenodo-Dep="13323012" ID-arpha="370E6028-657B-58B5-B22C-14AC2195FE63" ID-zoobank="https://zoobank.org/4D5E7BF7-2704-4467-9EE5-056C1FFE1D33" LSID="urn:lsid:zoobank.org:act:4D5E7BF7-2704-4467-9EE5-056C1FFE1D33" LSID-ZBK="urn:lsid:zoobank.org:act:4D5E7BF7-2704-4467-9EE5-056C1FFE1D33" httpUri="http://treatment.plazi.org/id/370E6028657B58B5B22C14AC2195FE63">
<subSubSection id="0B560B44284830CA38C66C7874857731" type="nomenclature">
<paragraph id="91B39D8542E6312D1A15F5109EA104E5">
<taxonomicName id="7897E8C6BC8615187572CF13A4810AF8" authority="Zhang, Li &amp; Yao" authorityName="Zhang, Li &amp; Yao" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="tianyang" status="sp. nov.">
<emphasis id="6C3C76C49E1FF9627276871A50AF3313" italics="true">Belisana tianyang</emphasis>
Zhang, Li &amp; Yao
</taxonomicName>
<taxonomicNameLabel id="9E5DF5A2FFBF031698737F3BCE17E29C" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="B7D5FED580AEB3EE76B474080455DDEF" type="description">
<paragraph id="8980110535AF1559BC5669715B2B86CE">
<figureCitation id="97434D38AB5655B2041D81D49784D8A5" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Belisana tianyang sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at prolatero-subdistal sclerite, arrow 2 points at prolatero-ventral lamella, arrow 3 points at distal membranous lamella, arrow 4 points at distal spine D retrolateral view, arrow points at dorso-subdistal apophysis). Abbreviations: b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.10 mm (A, B); 0.02 mm (C, D)." figureDoi="10.3897/zookeys.1209.127951.figure6" httpUri="https://binary.pensoft.net/fig/1113530">Figs 6</figureCitation>
,
<figureCitation id="C88D576453450129E80E71246FFB71D2" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Belisana tianyang sp. nov., holotype male (C – F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E – H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 mm (A – D); 0.50 mm (E – H)." figureDoi="10.3897/zookeys.1209.127951.figure7" httpUri="https://binary.pensoft.net/fig/1113531">7</figureCitation>
,
<figureCitation id="51276ED3B157D8D21F63E7FAA6F75A8A" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species) A, B Belisana langping sp. nov. C, D B. lingui sp. nov. E, F B. tianyang sp. nov. Abbreviation: f = flap. Scale bars: 0.10 mm." figureDoi="10.3897/zookeys.1209.127951.figure8" httpUri="https://binary.pensoft.net/fig/1113532">8 E, F</figureCitation>
,
<figureCitation id="E85A436978586A90E8F428AC0A8B0F89" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Female genitalia in ventral and dorsal views A, B Belisana langping sp. nov. C, D B. lingui sp. nov. E, F B. tianyang sp. nov. Abbreviations: aa = anterior arch, ep = epigynal pocket, pp = pore plate. Scale bars: 0.10 mm." figureDoi="10.3897/zookeys.1209.127951.figure9" httpUri="https://binary.pensoft.net/fig/1113533">9 E, F</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0E6QAG" type="type material">
<paragraph id="EA38816606017F91CCE1CA13111DC699">
<heading id="2C6459703BCFDC07B1944173A3996C7B" reason="title">Type material.</heading>
</paragraph>
<paragraph id="0DF10530334FB1BD4E0CF1FF88E6856C">
<materialsCitation id="DE8A3F365D549CF837E6A6735D6B99B2" collectingDate="2011-08-07" collectionCode="IZCAS" collectorName="C Wang" country="China" county="Tianyang County" elevation="467" latitude="23.668716" location="Dongjing Town" longLatPrecision="1" longitude="106.56593" municipality="Baise" specimenCode="Ar 44997" specimenCount="1" specimenCount-male="1" stateProvince="Guangxi" typeStatus="holotype">
<emphasis id="8DF7B31EBAB8F497DA80ED0CA34959D8" bold="true">
<emphasis id="062328F1FD32CED0966C40595CBCF523" bold="true" italics="true">
<typeStatus id="2A504AEFC8D454BE3B76C4FECA5460B0" type="holotype">Holotype</typeStatus>
</emphasis>
:
<collectingCountry id="4D556A932DDBE4292306E0670D5FC808" name="China">China</collectingCountry>
</emphasis>
•
<specimenCount id="71DA15389087CBA9958217838DF5B657" count="1" type="male">♂</specimenCount>
;
<collectingRegion id="1DF842E1639D5A37830A5BF467614FDB" country="China" name="Guangxi">Guangxi</collectingRegion>
,
<collectingMunicipality id="74AD9100FE9C78204485E5B1F0B9E40D">Baise</collectingMunicipality>
,
<collectingCounty id="064EBCA0FFBB0A0C0AFCCEF336221165">Tianyang County</collectingCounty>
,
<location id="4808E0DEF613251CB761042E2761BFB1" LSID="urn:lsid:plazi:treatment:370E6028657B58B5B22C14AC2195FE63:4808E0DEF613251CB761042E2761BFB1" country="China" county="Tianyang County" latitude="23.668716" longLatPrecision="1" longitude="106.56593" municipality="Baise" name="Dongjing Town" stateProvince="Guangxi">Dongjing Town</location>
,
<location id="73703453CB23FDE70F82B682F74B46CB" LSID="urn:lsid:plazi:treatment:370E6028657B58B5B22C14AC2195FE63:73703453CB23FDE70F82B682F74B46CB" country="China" county="Tianyang County" latitude="23.668716" longLatPrecision="1" longitude="106.56593" municipality="Baise" name="Liangdongyan Cave" stateProvince="Guangxi">Liangdongyan Cave</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ENRAG" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[106.565933,23.668717]}">
<geoCoordinate id="5848A28B94CC884977D74F5CF7F23F05" degrees="23" direction="north" minutes="40.123" orientation="latitude" precision="1" value="23.668716">23 ° 40.123 ' N</geoCoordinate>
,
<geoCoordinate id="A90DAA6574999D800EE7930BCEE3162F" degrees="106" direction="east" minutes="33.956" orientation="longitude" precision="1" value="106.56593">106 ° 33.956 ' E</geoCoordinate>
</named-content>
; alt.
<quantity id="2BE78F00B2BE2156878438A46E650929" metricMagnitude="2" metricUnit="m" metricValue="4.67" unit="m" value="467.0">
<elevation id="29B24F5E53159216C80410B129F7524D" metricMagnitude="2" metricUnit="m" metricValue="4.67" unit="m" value="467.0">467 m</elevation>
</quantity>
;
<date id="B2025C46D6A75E3EECFFB17230CDB480" value="2011-08-07">
<collectingDate id="CD21757F56B37BC04764006D25F6F801" value="2011-08-07">7 Aug. 2011</collectingDate>
</date>
;
<collectorName id="B708B2AB9898E12C2344A738DEBE690D">C Wang</collectorName>
leg.;
<named-content id="DC9173297200DBB0A6431CE91094FD07" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/institute-zoology-chinese-academy-sciences" xlink_title="Institute of Zoology, Chinese Academy of Sciences">
<collectionCode id="2D3E8BB0CE7B04A65459EE91732689D3" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" type="">IZCAS</collectionCode>
</named-content>
-
<specimenCode id="9F974F3103A70CFDB8C5F88006557905">Ar 44997</specimenCode>
</materialsCitation>
.
<materialsCitation id="C34B50D63A3DF20E70543381514904E1" collectingDate="2011-08-07" collectionCode="IZCAS" collectorName="C Wang &amp; Ar" country="China" county="Tianyang County" elevation="467" latitude="23.668716" location="China" longLatPrecision="1" longitude="106.56593" municipality="Baise" specimenCode="Ar 44998" specimenCount="3" specimenCount-male="3" stateProvince="Guangxi" typeStatus="paratype">
<emphasis id="81164AAE724E7D36ACB1C5EA4B247604" bold="true">
<emphasis id="5DC15BD63D301124CDB37C83D82707BC" bold="true" italics="true">
<typeStatus id="6771ED6CFB63F4275E6B91445298C7BD" type="paratype">Paratypes</typeStatus>
</emphasis>
:
<collectingCountry id="105EC176062474CFA67B6E54F5CF7ACF" name="China">China</collectingCountry>
</emphasis>
•
<specimenCount id="C973F21566A7C3DE6934A7E8397E18A8" count="3" type="male">3 ♂</specimenCount>
; same data as for holotype;
<named-content id="983E8E32346E516768BA460CF56AD55E" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/institute-zoology-chinese-academy-sciences" xlink_title="Institute of Zoology, Chinese Academy of Sciences">
<collectionCode id="9977676253471452B899DDCCD67E1A79" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" type="">IZCAS</collectionCode>
</named-content>
-
<specimenCode id="380A8F887295A9852B4B27FE08573EFA">Ar 44998</specimenCode>
–45000
</materialsCitation>
•
<materialsCitation id="80EA74762CC83BC9ACFBE38F063C922D" collectingDate="2011-08-07" collectionCode="IZCAS" collectorName="C Wang &amp; Ar" country="China" county="Tianyang County" elevation="467" latitude="23.668716" location="Dongjing Town" longLatPrecision="1" longitude="106.56593" municipality="Baise" specimenCode="Ar 45001" specimenCount="2" specimenCount-female="2" stateProvince="Guangxi" typeStatus="paratype">
<specimenCount id="99980BF7E0BF4816102093E06B3C337C" count="2" type="female">2 ♀</specimenCount>
; same data as for holotype;
<named-content id="C91E4BB22A501C5FA13F6302B60BB268" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/institute-zoology-chinese-academy-sciences" xlink_title="Institute of Zoology, Chinese Academy of Sciences">
<collectionCode id="096F0808A4A0E6666FA9926175640890" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" type="">IZCAS</collectionCode>
</named-content>
-
<specimenCode id="CE890A0E8A3A46ADA591FF75186459E5">Ar 45001</specimenCode>
–02
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ECSAG" type="etymology">
<paragraph id="9EFF5599F322F31865284EF8B4412629">
<heading id="85917F4885AEEDD543E8862131F56733" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="1DD545B3A7B4495468AD1BDAE4223591">The specific name refers to the type locality; noun in apposition.</paragraph>
</subSubSection>
<subSubSection id="SECID0EHSAG" type="diagnosis">
<paragraph id="3F5B94686F5254DD774D26E7EA65FB94">
<heading id="7F6DDA2B5C4E159A5907A4839B566167" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="9553ED37B6379C43707C5314FF6AF2AD">
The new species resembles
<taxonomicName id="159422C595E7D92C34B00FA0E0CF52E9" authority="Zhang &amp; Peng, 2011" authorityName="Zhang &amp; Peng" authorityYear="2011" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="tianlinensis">
<emphasis id="62DC064C3346FD0457C4D6FB0B737F96" italics="true">B. tianlinensis</emphasis>
<bibRefCitation id="82FD981DEFDA354B43C7201C3D6049D9" DOI="10.11646/zootaxa.2989.1.2" author="Zhang F &amp; Peng Y" issue="1" journalOrPublisher="Zootaxa" pagination="51-68" refId="B15" refString="Zhang F, Peng Y (2011) Eleven new species of the genus Belisana Thorell (Araneae: Pholcidae) from South China. Zootaxa 2989 (1): 51 - 68. https: // doi. org / 10.11646 / zootaxa. 2989.1. 2" title="Eleven new species of the genus Belisana Thorell (Araneae: Pholcidae) from South China." volume="2989" year="2011">Zhang &amp; Peng, 2011</bibRefCitation>
</taxonomicName>
(
<bibRefCitation id="DB8668D07FC8CDE2EDE37E256818F7A6" DOI="10.11646/zootaxa.2989.1.2" author="Zhang F &amp; Peng Y" firstAuthor="Zhang" issue="1" journalOrPublisher="Zootaxa" page="65" pagination="51-68" refId="B15" refString="Zhang F, Peng Y (2011) Eleven new species of the genus Belisana Thorell (Araneae: Pholcidae) from South China. Zootaxa 2989 (1): 51–68. https://doi.org/10.11646/zootaxa.2989.1.2" title="Eleven new species of the genus Belisana Thorell (Araneae: Pholcidae) from South China." volume="2989" year="2011">Zhang and Peng 2011: 65</bibRefCitation>
, fig. 10 A – G) by having similar bulbal apophyses and epigyne (Fig.
<figureCitation id="6E1609E925A720836C3BF4A06DB5CFDA" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Belisana tianyang sp. nov., holotype male (C – F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E – H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 mm (A – D); 0.50 mm (E – H)." figureDoi="10.3897/zookeys.1209.127951.figure7" httpUri="https://binary.pensoft.net/fig/1113531">7 A, C</figureCitation>
), but can be distinguished by retrolateral flap of procursus strongly curved and wide (4 times wider than long, f in Figs
<figureCitation id="EC878E86FFECEE2D9EDB089174BFE069" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Belisana tianyang sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at prolatero-subdistal sclerite, arrow 2 points at prolatero-ventral lamella, arrow 3 points at distal membranous lamella, arrow 4 points at distal spine D retrolateral view, arrow points at dorso-subdistal apophysis). Abbreviations: b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.10 mm (A, B); 0.02 mm (C, D)." figureDoi="10.3897/zookeys.1209.127951.figure6" httpUri="https://binary.pensoft.net/fig/1113530">6 D</figureCitation>
,
<figureCitation id="A986047A2BBE59B6AE27986A6D6ECAE2" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species) A, B Belisana langping sp. nov. C, D B. lingui sp. nov. E, F B. tianyang sp. nov. Abbreviation: f = flap. Scale bars: 0.10 mm." figureDoi="10.3897/zookeys.1209.127951.figure8" httpUri="https://binary.pensoft.net/fig/1113532">8 F</figureCitation>
vs. straight and 2 times wider than long in
<taxonomicName id="8538F28EAE7A2867B4AFEE936266FCA9" authorityName="Zhang &amp; Peng" authorityYear="2011" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="tianlinensis">
<emphasis id="9D80CA94AC67DE8619CCC346A3780281" italics="true">B. tianlinensis</emphasis>
</taxonomicName>
), by male cheliceral distal apophyses long (6 times longer than wide) and tips widely separated (
<abbrev id="ABBRID0ETTAG" xlink_title="distal apophysis">da</abbrev>
in Fig.
<figureCitation id="C9E5F485568F0CE28A31CB05D4C7FAC5" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Belisana tianyang sp. nov., holotype male (C – F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E – H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 mm (A – D); 0.50 mm (E – H)." figureDoi="10.3897/zookeys.1209.127951.figure7" httpUri="https://binary.pensoft.net/fig/1113531">7 D</figureCitation>
vs. 2 times longer than wide and tips closer to each other in
<taxonomicName id="CC7967FC7F07C5AD4E81CA94C6F51441" authorityName="Zhang &amp; Peng" authorityYear="2011" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="tianlinensis">
<emphasis id="C4712005B8CE6FC94D2B64AE1AC4D242" italics="true">B. tianlinensis</emphasis>
</taxonomicName>
), by pore plates curved, anteriorly pointed and posteriorly wide (
<abbrev id="ABBRID0EGUAG" xlink_title="pore plate">pp</abbrev>
in Figs
<figureCitation id="FE86EB9AEB803C5B62FEE7B2E4BE6718" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Belisana tianyang sp. nov., holotype male (C – F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E – H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 mm (A – D); 0.50 mm (E – H)." figureDoi="10.3897/zookeys.1209.127951.figure7" httpUri="https://binary.pensoft.net/fig/1113531">7 B</figureCitation>
,
<figureCitation id="C20FF22BD76E8816D7F478A29E0BC026" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Female genitalia in ventral and dorsal views A, B Belisana langping sp. nov. C, D B. lingui sp. nov. E, F B. tianyang sp. nov. Abbreviations: aa = anterior arch, ep = epigynal pocket, pp = pore plate. Scale bars: 0.10 mm." figureDoi="10.3897/zookeys.1209.127951.figure9" httpUri="https://binary.pensoft.net/fig/1113533">9 F</figureCitation>
vs. nearly triangular in
<taxonomicName id="1DC19CB7677717B9FB08D3BF7D077C72" authorityName="Zhang &amp; Peng" authorityYear="2011" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="tianlinensis">
<emphasis id="79367BC208461D7312BF0FF5D7890D16" italics="true">B. tianlinensis</emphasis>
</taxonomicName>
), and by male clypeus unmodified (Fig.
<figureCitation id="EE4D9D2D8688DF382ECF932BE86E50A1" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Belisana tianyang sp. nov., holotype male (C – F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E – H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 mm (A – D); 0.50 mm (E – H)." figureDoi="10.3897/zookeys.1209.127951.figure7" httpUri="https://binary.pensoft.net/fig/1113531">7 E</figureCitation>
vs. with pointed frontal apophysis in
<taxonomicName id="075A38B18AF7FFFC11C7DF922803D1F7" authorityName="Zhang &amp; Peng" authorityYear="2011" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="tianlinensis">
<emphasis id="50D5E19E42E27A4A28A4BB23167934FF" italics="true">B. tianlinensis</emphasis>
</taxonomicName>
); also distinguished from
<taxonomicName id="5F625FE2EC7E998BA77AFEC71501427A" authority="Chen, Zhang &amp; Zhu, 2009" authorityName="Chen, Zhang &amp; Zhu" authorityYear="2009" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="douqing">
<emphasis id="FF6BAADA72CFCF8289F1A9F33632E653" italics="true">B. douqing</emphasis>
<bibRefCitation id="BB067B48FA2409A9EE91556F95ECE0F6" DOI="10.11646/zootaxa.2092.1.5" author="Chen H &amp; Zhang F &amp; Zhu M" issue="1" journalOrPublisher="Zootaxa" pagination="58-68" refId="B1" refString="Chen H, Zhang F, Zhu M (2009) Four new troglophilous species of the genus Belisana Thorell, 1898 (Araneae, Pholcidae) from Guizhou Province, China. Zootaxa 2092 (1): 58 - 68. https: // doi. org / 10.11646 / zootaxa. 2092.1. 5" title="Four new troglophilous species of the genus Belisana Thorell, 1898 (Araneae, Pholcidae) from Guizhou Province, China." volume="2092" year="2009">Chen, Zhang &amp; Zhu, 2009</bibRefCitation>
</taxonomicName>
(
<bibRefCitation id="1007CDD304061A2C80DB5CDBABF4674D" DOI="10.11646/zootaxa.2092.1.5" author="Chen H &amp; Zhang F &amp; Zhu M" etAl="et al." firstAuthor="Chen" issue="1" journalOrPublisher="Zootaxa" page="59" pagination="58-68" refId="B1" refString="Chen H, Zhang F, Zhu M (2009) Four new troglophilous species of the genus Belisana Thorell, 1898 (Araneae, Pholcidae) from Guizhou Province, China. Zootaxa 2092 (1): 58–68. https://doi.org/10.11646/zootaxa.2092.1.5" title="Four new troglophilous species of the genus Belisana Thorell, 1898 (Araneae, Pholcidae) from Guizhou Province, China." volume="2092" year="2009">Chen et al. 2009: 59</bibRefCitation>
, figs 1–11) by procursus with rectangular distal membranous lamella (arrow 3 in Figs
<figureCitation id="9690FD2438065765B2133DC450D08811" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Belisana tianyang sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at prolatero-subdistal sclerite, arrow 2 points at prolatero-ventral lamella, arrow 3 points at distal membranous lamella, arrow 4 points at distal spine D retrolateral view, arrow points at dorso-subdistal apophysis). Abbreviations: b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.10 mm (A, B); 0.02 mm (C, D)." figureDoi="10.3897/zookeys.1209.127951.figure6" httpUri="https://binary.pensoft.net/fig/1113530">6 C</figureCitation>
,
<figureCitation id="FDB29B9B9E6C9E1E13778C2095EB323F" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species) A, B Belisana langping sp. nov. C, D B. lingui sp. nov. E, F B. tianyang sp. nov. Abbreviation: f = flap. Scale bars: 0.10 mm." figureDoi="10.3897/zookeys.1209.127951.figure8" httpUri="https://binary.pensoft.net/fig/1113532">8 E</figureCitation>
vs. nearly square in
<taxonomicName id="CEB2E5CEB53C1643CBF0F295D060BF23" authorityName="Chen, Zhang &amp; Zhu" authorityYear="2009" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="douqing">
<emphasis id="A86081DAF5E985A88ED228531B229296" italics="true">B. douqing</emphasis>
</taxonomicName>
) and curved retrolateral membranous flap (f in Figs
<figureCitation id="896FEE45312E0C402AE9E66581CFC5C7" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Belisana tianyang sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at prolatero-subdistal sclerite, arrow 2 points at prolatero-ventral lamella, arrow 3 points at distal membranous lamella, arrow 4 points at distal spine D retrolateral view, arrow points at dorso-subdistal apophysis). Abbreviations: b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.10 mm (A, B); 0.02 mm (C, D)." figureDoi="10.3897/zookeys.1209.127951.figure6" httpUri="https://binary.pensoft.net/fig/1113530">6 D</figureCitation>
,
<figureCitation id="0B5C6F5C29DD151CEF3D960C47BB674F" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species) A, B Belisana langping sp. nov. C, D B. lingui sp. nov. E, F B. tianyang sp. nov. Abbreviation: f = flap. Scale bars: 0.10 mm." figureDoi="10.3897/zookeys.1209.127951.figure8" httpUri="https://binary.pensoft.net/fig/1113532">8 F</figureCitation>
vs. angular in
<taxonomicName id="229F1D5182DA010A4C44AFCAD8852BC0" authorityName="Chen, Zhang &amp; Zhu" authorityYear="2009" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="douqing">
<emphasis id="5C42941C5146C043F150464FFB996770" italics="true">B. douqing</emphasis>
</taxonomicName>
) and by pore plates curved, anteriorly pointed and posteriorly wide (
<abbrev id="ABBRID0EBXAG" xlink_title="pore plate">pp</abbrev>
in Figs
<figureCitation id="3F1323BBDDB864428A0BD2A4A901EFDA" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Belisana tianyang sp. nov., holotype male (C – F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E – H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 mm (A – D); 0.50 mm (E – H)." figureDoi="10.3897/zookeys.1209.127951.figure7" httpUri="https://binary.pensoft.net/fig/1113531">7 B</figureCitation>
,
<figureCitation id="C9879D9D5924FE1CA9CD6599DD610AC1" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Female genitalia in ventral and dorsal views A, B Belisana langping sp. nov. C, D B. lingui sp. nov. E, F B. tianyang sp. nov. Abbreviations: aa = anterior arch, ep = epigynal pocket, pp = pore plate. Scale bars: 0.10 mm." figureDoi="10.3897/zookeys.1209.127951.figure9" httpUri="https://binary.pensoft.net/fig/1113533">9 F</figureCitation>
vs. long elliptic in
<taxonomicName id="6F88DC1AC73E916B2656F1D0EE8905C8" authorityName="Chen, Zhang &amp; Zhu" authorityYear="2009" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="douqing">
<emphasis id="D64ABE99BEF971EFABF3D3ED04592177" italics="true">B. douqing</emphasis>
</taxonomicName>
).
</paragraph>
<caption id="A032C0A46EFBB9A7005A4D3D381EE06E" ID-DOI="10.3897/zookeys.1209.127951.figure6" ID-arpha="C8A13633-12CD-5511-AA44-891922113387" httpUri="https://binary.pensoft.net/fig/1113530" startId="F6">
<paragraph id="A0E7A08D5606E5B4453D11C0926DF601">
<label id="BE5374DFA09A5C639820E1484137C069">Figure 6.</label>
</paragraph>
<paragraph id="6586BA2DC4BCEB7FE80A8DFE435081C2">
<taxonomicName id="135D4F4F6D1DA99D78E9E1DF07B4ECC5" authorityName="Zhang &amp; Wang &amp; Li &amp; Zhang &amp; Yao" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="tianyang" status="sp. nov.">
<emphasis id="965B469A79B9D0A2A34251F1A12247EA" italics="true">Belisana tianyang</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9382D0FA67DFF00FA12176B801F84C38" rank="species">sp. nov.</taxonomicNameLabel>
, holotype male
<emphasis id="7582C794878B4DA2758AFA925FF407A0" bold="true">A, B</emphasis>
palp (
<emphasis id="19488AD0CD387E9DF1D78120ECB557A3" bold="true">A</emphasis>
prolateral view
<emphasis id="CD3AE07EF69914D92B8CFC72D46E121B" bold="true">B</emphasis>
retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion)
<emphasis id="60AEAB45505A18D834AE5206F19BE10B" bold="true">C, D</emphasis>
distal part of procursus (
<emphasis id="A2436FFA3B20FA80CAD98852C9C052A3" bold="true">C</emphasis>
prolateral view, arrow 1 points at prolatero-subdistal sclerite, arrow 2 points at prolatero-ventral lamella, arrow 3 points at distal membranous lamella, arrow 4 points at distal spine
<emphasis id="79BE603D1E3E8B5FD3ED2C80E30FF290" bold="true">D</emphasis>
retrolateral view, arrow points at dorso-subdistal apophysis). Abbreviations: b = bulb,
<abbrev id="ABBRID0EXYAG" xlink_title="bulbal apophysis">ba</abbrev>
= bulbal apophysis, e = embolus, f = flap,
<abbrev id="ABBRID0E2YAG" xlink_title="procursus">pr</abbrev>
= procursus. Scale bars: 0.10 mm (
<emphasis id="D01FFE03A8EB8DB105DE86EC7B73589D" bold="true">A, B</emphasis>
); 0.02 mm (
<emphasis id="E000E7A35DEB5C185DA8A354B2F71D89" bold="true">C, D</emphasis>
).
</paragraph>
</caption>
<caption id="5EC147353E8711F85B13AF7A12C69235" ID-DOI="10.3897/zookeys.1209.127951.figure7" ID-arpha="773C46B8-80B1-5CE7-8649-E0A8F3A3F0EA" httpUri="https://binary.pensoft.net/fig/1113531" startId="F7">
<paragraph id="BF35001E1032BF1B7F8FFC0AB3C03D9B">
<label id="D26EEB7EBD1646C43294AEDF8875B8C7">Figure 7.</label>
</paragraph>
<paragraph id="EDEDDB3DD07F26528819E393F8180A78">
<taxonomicName id="E7B1DDB2B1F53BE84D6A6FC89C8429F2" authorityName="Zhang &amp; Wang &amp; Li &amp; Zhang &amp; Yao" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="tianyang" status="sp. nov.">
<emphasis id="9979B17C518007673D459A403F689205" italics="true">Belisana tianyang</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FA1E8F81868CE22C033D1A39A9AB95EE" rank="species">sp. nov.</taxonomicNameLabel>
, holotype male (
<emphasis id="EFEE585CCA933E8B544783CC68A62CBB" bold="true">C – F</emphasis>
) and paratype female (
<emphasis id="7AE2C953B0DA01A52B0A46238B207C12" bold="true">A, B, G, H</emphasis>
)
<emphasis id="C65AE345D65757F5FB2BF5768855D4DD" bold="true">A</emphasis>
epigyne, ventral view
<emphasis id="87FE822FA03EF989871FEB078201CFDB" bold="true">B</emphasis>
vulva, dorsal view
<emphasis id="732BBC6AB0E81186938B57DCC65A90B4" bold="true">C</emphasis>
bulb, prolateral view
<emphasis id="C4945C860B4E804826ECB5815709E600" bold="true">D</emphasis>
chelicerae, frontal view
<emphasis id="9FB26C7CDAA279000588298596F97B4F" bold="true">E – H</emphasis>
habitus (
<emphasis id="04056CCCDCE2679B07B185B4BE39783A" bold="true">E, G</emphasis>
dorsal view
<emphasis id="363A3D290B4CAA408024AD16605640E6" bold="true">F</emphasis>
lateral view
<emphasis id="0B32BCE9E88769042DC625FFB48EC7A5" bold="true">H</emphasis>
ventral view). Abbreviations:
<abbrev id="ABBRID0EP1AG" xlink_title="anterior arch">aa</abbrev>
= anterior arch, b = bulb,
<abbrev id="ABBRID0ET1AG" xlink_title="bulbal apophysis">ba</abbrev>
= bulbal apophysis,
<abbrev id="ABBRID0EX1AG" xlink_title="distal apophysis">da</abbrev>
= distal apophysis, e = embolus,
<abbrev id="ABBRID0E21AG" xlink_title="epigynal pocket">ep</abbrev>
= epigynal pocket,
<abbrev id="ABBRID0E61AG" xlink_title="proximo-lateral apophysis">pa</abbrev>
= proximo-lateral apophysis,
<abbrev id="ABBRID0ED2AG" xlink_title="pore plate">pp</abbrev>
= pore plate. Scale bars: 0.10 mm (
<emphasis id="FB91A633C1E895D27FC1CFB00D936E3F" bold="true">A – D</emphasis>
); 0.50 mm (
<emphasis id="730F747047219D407A540FD23D5A11F1" bold="true">E – H</emphasis>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EQ2AG" type="description">
<paragraph id="9ED203CE80D2BEF13D9988A2879856D4">
<heading id="27B8FE15F1918C01309D723EE2A94102" reason="title">Description.</heading>
</paragraph>
<paragraph id="FAB17CDE7D82A566D115425755F3FAA7">
<emphasis id="A4F20565D00BC6E3321273D48F53AA7B" bold="true">Male</emphasis>
(
<emphasis id="BEA4BE28F9D156C6E7B7E6A9502A5AC6" bold="true" italics="true">
<typeStatus id="90422A61EBDD73A2FB50D81DBAE63CB4" type="holotype">holotype</typeStatus>
</emphasis>
): Total length 1.98 (2.08 with clypeus), prosoma 0.74 long, 0.75 wide, opisthosoma 1.24 long, 0.89 wide. Leg I: 21.95 (5.51, 0.33, 5.44, 9.17, 1.50), leg II missing, leg III: 8.82 (2.44, 0.27, 2.25, 3.13, 0.73), leg IV: 12.12 (3.60, 0.28, 3.04, 4.45, 0.75); tibia I
<abbrev id="ABBRID0E22AG" xlink_title="length/diameter">L / d</abbrev>
: 68. Eye interdistances and diameters:
<abbrev id="ABBRID0E62AG" xlink_title="posterior median eye">PME</abbrev>
–
<abbrev id="ABBRID0ED3AG" xlink_title="posterior median eye">PME</abbrev>
0.10,
<abbrev id="ABBRID0EH3AG" xlink_title="posterior median eye">PME</abbrev>
0.08,
<abbrev id="ABBRID0EL3AG" xlink_title="posterior median eye">PME</abbrev>
–
<abbrev id="ABBRID0EP3AG" xlink_title="anterior lateral eye">ALE</abbrev>
0.02,
<abbrev id="ABBRID0ET3AG" xlink_title="anterior median eye">AME</abbrev>
absent. Sternum width / length: 0.58 / 0.56. Habitus as in Fig.
<figureCitation id="C600DACA2B47A84AEEF1B8F1FC9D14B0" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Belisana tianyang sp. nov., holotype male (C – F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E – H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 mm (A – D); 0.50 mm (E – H)." figureDoi="10.3897/zookeys.1209.127951.figure7" httpUri="https://binary.pensoft.net/fig/1113531">7 E, F</figureCitation>
. Dorsal shield of prosoma yellowish, with indistinct median stripe; sternum yellowish, without marks. Legs whitish, without darker rings. Opisthosoma yellowish, without spots. Thoracic furrow absent. Clypeus unmodified. Chelicerae with pair of proximo-lateral apophyses (
<abbrev id="ABBRID0E23AG" xlink_title="proximo-lateral apophysis">pa</abbrev>
in Fig.
<figureCitation id="6E9527711CA6CD9346DC5A09F877F95C" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Belisana tianyang sp. nov., holotype male (C – F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E – H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 mm (A – D); 0.50 mm (E – H)." figureDoi="10.3897/zookeys.1209.127951.figure7" httpUri="https://binary.pensoft.net/fig/1113531">7 D</figureCitation>
) and pair of distal apophyses (
<abbrev id="ABBRID0ED4AG" xlink_title="distal apophysis">da</abbrev>
in Fig.
<figureCitation id="C3D32A3170BBCA882D834F8A2336E86A" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Belisana tianyang sp. nov., holotype male (C – F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E – H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 mm (A – D); 0.50 mm (E – H)." figureDoi="10.3897/zookeys.1209.127951.figure7" httpUri="https://binary.pensoft.net/fig/1113531">7 D</figureCitation>
; distance between tips: 0.36). Palp as in Fig.
<figureCitation id="F89DF53196091EBC46B6AB024C30873D" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Belisana tianyang sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at prolatero-subdistal sclerite, arrow 2 points at prolatero-ventral lamella, arrow 3 points at distal membranous lamella, arrow 4 points at distal spine D retrolateral view, arrow points at dorso-subdistal apophysis). Abbreviations: b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.10 mm (A, B); 0.02 mm (C, D)." figureDoi="10.3897/zookeys.1209.127951.figure6" httpUri="https://binary.pensoft.net/fig/1113530">6 A, B</figureCitation>
; trochanter with ventral apophysis (arrow 1 in Fig.
<figureCitation id="B700EDB804C4F06FFA8AA0B8CFB543DF" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Belisana tianyang sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at prolatero-subdistal sclerite, arrow 2 points at prolatero-ventral lamella, arrow 3 points at distal membranous lamella, arrow 4 points at distal spine D retrolateral view, arrow points at dorso-subdistal apophysis). Abbreviations: b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.10 mm (A, B); 0.02 mm (C, D)." figureDoi="10.3897/zookeys.1209.127951.figure6" httpUri="https://binary.pensoft.net/fig/1113530">6 B</figureCitation>
); femur with small retrolatero-proximal protrusion (arrow 2 in Fig.
<figureCitation id="E1A4884298C2B3EB99AE85230DD189C7" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Belisana tianyang sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at prolatero-subdistal sclerite, arrow 2 points at prolatero-ventral lamella, arrow 3 points at distal membranous lamella, arrow 4 points at distal spine D retrolateral view, arrow points at dorso-subdistal apophysis). Abbreviations: b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.10 mm (A, B); 0.02 mm (C, D)." figureDoi="10.3897/zookeys.1209.127951.figure6" httpUri="https://binary.pensoft.net/fig/1113530">6 B</figureCitation>
); procursus simple proximally but complex distally, with prolatero-subdistal sclerite (arrow 1 in Figs
<figureCitation id="9F5C2A2F04814CB114CE27CF4D5EA316" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Belisana tianyang sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at prolatero-subdistal sclerite, arrow 2 points at prolatero-ventral lamella, arrow 3 points at distal membranous lamella, arrow 4 points at distal spine D retrolateral view, arrow points at dorso-subdistal apophysis). Abbreviations: b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.10 mm (A, B); 0.02 mm (C, D)." figureDoi="10.3897/zookeys.1209.127951.figure6" httpUri="https://binary.pensoft.net/fig/1113530">6 C</figureCitation>
,
<figureCitation id="2510CE904538AE4034FD12CFCAB648D2" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species) A, B Belisana langping sp. nov. C, D B. lingui sp. nov. E, F B. tianyang sp. nov. Abbreviation: f = flap. Scale bars: 0.10 mm." figureDoi="10.3897/zookeys.1209.127951.figure8" httpUri="https://binary.pensoft.net/fig/1113532">8 E</figureCitation>
), sclerotized prolatero-ventral lamella (arrow 2 in Figs
<figureCitation id="A04693BFE13BEAA8D749EE6BCE501D16" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Belisana tianyang sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at prolatero-subdistal sclerite, arrow 2 points at prolatero-ventral lamella, arrow 3 points at distal membranous lamella, arrow 4 points at distal spine D retrolateral view, arrow points at dorso-subdistal apophysis). Abbreviations: b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.10 mm (A, B); 0.02 mm (C, D)." figureDoi="10.3897/zookeys.1209.127951.figure6" httpUri="https://binary.pensoft.net/fig/1113530">6 C</figureCitation>
,
<figureCitation id="4D90249968937CAEDD016C0A20888EC3" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species) A, B Belisana langping sp. nov. C, D B. lingui sp. nov. E, F B. tianyang sp. nov. Abbreviation: f = flap. Scale bars: 0.10 mm." figureDoi="10.3897/zookeys.1209.127951.figure8" httpUri="https://binary.pensoft.net/fig/1113532">8 E</figureCitation>
), distal membranous lamella (arrow 3 in Figs
<figureCitation id="4E476606EA676B9E6435D75CB0F46CCD" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Belisana tianyang sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at prolatero-subdistal sclerite, arrow 2 points at prolatero-ventral lamella, arrow 3 points at distal membranous lamella, arrow 4 points at distal spine D retrolateral view, arrow points at dorso-subdistal apophysis). Abbreviations: b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.10 mm (A, B); 0.02 mm (C, D)." figureDoi="10.3897/zookeys.1209.127951.figure6" httpUri="https://binary.pensoft.net/fig/1113530">6 C</figureCitation>
,
<figureCitation id="FDA2BC57BA8B77F3A47F066FC1B04ED1" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species) A, B Belisana langping sp. nov. C, D B. lingui sp. nov. E, F B. tianyang sp. nov. Abbreviation: f = flap. Scale bars: 0.10 mm." figureDoi="10.3897/zookeys.1209.127951.figure8" httpUri="https://binary.pensoft.net/fig/1113532">8 E</figureCitation>
), curved distal spine (arrow 4 in Figs
<figureCitation id="34C896587E2550AADE09015E44CC648B" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Belisana tianyang sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at prolatero-subdistal sclerite, arrow 2 points at prolatero-ventral lamella, arrow 3 points at distal membranous lamella, arrow 4 points at distal spine D retrolateral view, arrow points at dorso-subdistal apophysis). Abbreviations: b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.10 mm (A, B); 0.02 mm (C, D)." figureDoi="10.3897/zookeys.1209.127951.figure6" httpUri="https://binary.pensoft.net/fig/1113530">6 C</figureCitation>
,
<figureCitation id="876B854EAD4358BE68808B3AA3B80073" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species) A, B Belisana langping sp. nov. C, D B. lingui sp. nov. E, F B. tianyang sp. nov. Abbreviation: f = flap. Scale bars: 0.10 mm." figureDoi="10.3897/zookeys.1209.127951.figure8" httpUri="https://binary.pensoft.net/fig/1113532">8 E</figureCitation>
), sclerotized dorso-subdistal apophysis (arrow in Figs
<figureCitation id="C78F1DC8EB07F003CCD19EFB49070632" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Belisana tianyang sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at prolatero-subdistal sclerite, arrow 2 points at prolatero-ventral lamella, arrow 3 points at distal membranous lamella, arrow 4 points at distal spine D retrolateral view, arrow points at dorso-subdistal apophysis). Abbreviations: b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.10 mm (A, B); 0.02 mm (C, D)." figureDoi="10.3897/zookeys.1209.127951.figure6" httpUri="https://binary.pensoft.net/fig/1113530">6 D</figureCitation>
,
<figureCitation id="4FE704337388AE492CB2468E4A714928" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species) A, B Belisana langping sp. nov. C, D B. lingui sp. nov. E, F B. tianyang sp. nov. Abbreviation: f = flap. Scale bars: 0.10 mm." figureDoi="10.3897/zookeys.1209.127951.figure8" httpUri="https://binary.pensoft.net/fig/1113532">8 F</figureCitation>
), and retrolateral membranous flap (f in Figs
<figureCitation id="0527E27006AC5E05E02D4F5A71CA429F" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Belisana tianyang sp. nov., holotype male A, B palp (A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus (C prolateral view, arrow 1 points at prolatero-subdistal sclerite, arrow 2 points at prolatero-ventral lamella, arrow 3 points at distal membranous lamella, arrow 4 points at distal spine D retrolateral view, arrow points at dorso-subdistal apophysis). Abbreviations: b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.10 mm (A, B); 0.02 mm (C, D)." figureDoi="10.3897/zookeys.1209.127951.figure6" httpUri="https://binary.pensoft.net/fig/1113530">6 D</figureCitation>
,
<figureCitation id="AC9E5A61F27E1AA41D8597AC9129AE43" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species) A, B Belisana langping sp. nov. C, D B. lingui sp. nov. E, F B. tianyang sp. nov. Abbreviation: f = flap. Scale bars: 0.10 mm." figureDoi="10.3897/zookeys.1209.127951.figure8" httpUri="https://binary.pensoft.net/fig/1113532">8 F</figureCitation>
); bulb with hooked apophysis (
<abbrev id="ABBRID0EH6AG" xlink_title="bulbal apophysis">ba</abbrev>
in Fig.
<figureCitation id="0805C4A7BF39E9FE116029534CFA2585" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Belisana tianyang sp. nov., holotype male (C – F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E – H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 mm (A – D); 0.50 mm (E – H)." figureDoi="10.3897/zookeys.1209.127951.figure7" httpUri="https://binary.pensoft.net/fig/1113531">7 C</figureCitation>
) and simple embolus (e in Fig.
<figureCitation id="9A2D236A923609AA0FE80B4D963AD986" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Belisana tianyang sp. nov., holotype male (C – F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E – H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 mm (A – D); 0.50 mm (E – H)." figureDoi="10.3897/zookeys.1209.127951.figure7" httpUri="https://binary.pensoft.net/fig/1113531">7 C</figureCitation>
). Retrolateral trichobothria on tibia I at 5 % proximally; legs with short vertical setae on metatarsi; tarsus I with 19 distinct pseudosegments.
</paragraph>
<caption id="5B6B9BA7EBAB77C65EB3B5F91E6FEAB4" ID-DOI="10.3897/zookeys.1209.127951.figure8" ID-arpha="E5391B4D-65F8-50C8-ABA4-FA65B1F40FDF" httpUri="https://binary.pensoft.net/fig/1113532" startId="F8">
<paragraph id="79060FC1C90A380D49969C457AC8988F">
<label id="099ACEE7EBDF8E81DCA849059C9D99BB">Figure 8.</label>
</paragraph>
<paragraph id="CDB5432E39D487CBC576AD445C63D848">
Procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species)
<emphasis id="76C77C13B3A6B9C45FDE972D08726E8A" bold="true">A, B</emphasis>
<taxonomicName id="64069888A7B53994B082B4D3A46B9710" authorityName="Zhang &amp; Wang &amp; Li &amp; Zhang &amp; Yao" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="langping" status="sp. nov.">
<emphasis id="BB4E8BE631F1DB7A9A08BAFD99CDE305" italics="true">Belisana langping</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F4E58004C87E582A42899B5288C169EC" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="77C044D8449409F7AE6D3FE8F8C37A38" bold="true">C, D</emphasis>
<taxonomicName id="0D595ACE2C8F2DF96B6C0D227A21FCF6" authorityName="Zhang &amp; Wang &amp; Li &amp; Zhang &amp; Yao" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="lingui" status="sp. nov.">
<emphasis id="339374DB27AF962B5DED5244BF99F5F2" italics="true">B. lingui</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7BC9BA83980743C4DC96BA1C484DC2A0" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="7F7D12A8DE27AD224982FF3F009D4C77" bold="true">E, F</emphasis>
<taxonomicName id="6C064041DCC0E25E12DDED1EEEB43C0F" authorityName="Zhang &amp; Wang &amp; Li &amp; Zhang &amp; Yao" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="tianyang" status="sp. nov.">
<emphasis id="C509B896AE89B805FA8FCB33AE0C451E" italics="true">B. tianyang</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2E877CE1882282C3B655A24C411550F6" rank="species">sp. nov.</taxonomicNameLabel>
Abbreviation: f = flap. Scale bars: 0.10 mm.
</paragraph>
</caption>
<caption id="186D04FFC61A67653CA138D525A5CED0" ID-DOI="10.3897/zookeys.1209.127951.figure9" ID-arpha="402004D2-75A9-5B0C-97EF-28A50679C9EC" httpUri="https://binary.pensoft.net/fig/1113533" startId="F9">
<paragraph id="7A33C4A8C337EB38A434CFF4997C46E9">
<label id="BF8B102EA91002FA498F439AAA4C0188">Figure 9.</label>
</paragraph>
<paragraph id="E77BFCA0110272E1C97056451B34632B">
Female genitalia in ventral and dorsal views
<emphasis id="DAE0B53BA7AF8F3D9CE978F9BE87EA57" bold="true">A, B</emphasis>
<taxonomicName id="B82BCF26CD9574C9F61CF58EC2FA33E0" authorityName="Zhang &amp; Wang &amp; Li &amp; Zhang &amp; Yao" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="langping" status="sp. nov.">
<emphasis id="EA5C6682778E13EAFE649DFEAF768320" italics="true">Belisana langping</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CE9A0826877E2E2C9BFB5F476862BF6A" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="0EF9960DA5535E459BC74413E0E575D8" bold="true">C, D</emphasis>
<taxonomicName id="03D7E15754E65AA0F14DB7E0ECF358AC" authorityName="Zhang &amp; Wang &amp; Li &amp; Zhang &amp; Yao" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="lingui" status="sp. nov.">
<emphasis id="F0DE3DFC3B330612D4FBE7F5C6FBF5F2" italics="true">B. lingui</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D06B7A4954FAFFAE4AACC645320F47AD" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="8E004A57111A7765086AADEDD5402BE3" bold="true">E, F</emphasis>
<taxonomicName id="D1D14C89BAB24573052EF34EBFEFB6F5" authorityName="Zhang &amp; Wang &amp; Li &amp; Zhang &amp; Yao" authorityYear="2024" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="tianyang" status="sp. nov.">
<emphasis id="1951A15FBB7F70EC8521F88CFC13EEFE" italics="true">B. tianyang</emphasis>
</taxonomicName>
<taxonomicNameLabel id="925F0754F68FCA258D16F65A7B109163" rank="species">sp. nov.</taxonomicNameLabel>
Abbreviations:
<abbrev id="ABBRID0EWCBG" xlink_title="anterior arch">aa</abbrev>
= anterior arch,
<abbrev id="ABBRID0E1CBG" xlink_title="epigynal pocket">ep</abbrev>
= epigynal pocket,
<abbrev id="ABBRID0E5CBG" xlink_title="pore plate">pp</abbrev>
= pore plate. Scale bars: 0.10 mm.
</paragraph>
</caption>
<paragraph id="3D3E1DB68A77F7DFBDE42E9F3DC4AE88">
<emphasis id="754AED11E340D99AAD43D34470788A09" bold="true">Female</emphasis>
(
<emphasis id="20A0056BCEE7EE83ABA7788A88CE4F14" bold="true" italics="true">
<typeStatus id="529DCE9C9A6DA53AB02331DD132ECD77" type="paratype">paratype</typeStatus>
</emphasis>
,
<named-content id="681ABCF05FA70081D44DEC8DD9DF9EB8" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/institute-zoology-chinese-academy-sciences" xlink_title="Institute of Zoology, Chinese Academy of Sciences">
<collectionCode id="F3E66F7FA13F41C9BE8601E7257B3C68" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" type="">IZCAS</collectionCode>
</named-content>
-
<specimenCode id="274D151B578BC32A761FB47195AF7B4E">Ar 45001</specimenCode>
): Similar to male, habitus as in Fig.
<figureCitation id="F0EDAF25A9903D79A0DFE66F4F2FD68C" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Belisana tianyang sp. nov., holotype male (C – F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E – H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 mm (A – D); 0.50 mm (E – H)." figureDoi="10.3897/zookeys.1209.127951.figure7" httpUri="https://binary.pensoft.net/fig/1113531">7 G, H</figureCitation>
. Total length 2.14 (2.27 with clypeus), prosoma 0.70 long, 0.79 wide, opisthosoma 1.44 long, 1.32 wide; tibia I: 4.15; tibia I
<abbrev id="ABBRID0EWDBG" xlink_title="length/diameter">L / d</abbrev>
: 52. Eye interdistances and diameters:
<abbrev id="ABBRID0E1DBG" xlink_title="posterior median eye">PME</abbrev>
–
<abbrev id="ABBRID0E5DBG" xlink_title="posterior median eye">PME</abbrev>
0.10,
<abbrev id="ABBRID0ECEBG" xlink_title="posterior median eye">PME</abbrev>
0.08,
<abbrev id="ABBRID0EGEBG" xlink_title="posterior median eye">PME</abbrev>
–
<abbrev id="ABBRID0EKEBG" xlink_title="anterior lateral eye">ALE</abbrev>
0.02,
<abbrev id="ABBRID0EOEBG" xlink_title="anterior median eye">AME</abbrev>
absent. Sternum width / length: 0.55 / 0.54. Epigyne simple and flat, with pair of lateral pockets 0.36 apart (
<abbrev id="ABBRID0ESEBG" xlink_title="epigynal pocket">ep</abbrev>
in Figs
<figureCitation id="6F92B5F3628D7DECB21A18AB78CFD32B" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Belisana tianyang sp. nov., holotype male (C – F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E – H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 mm (A – D); 0.50 mm (E – H)." figureDoi="10.3897/zookeys.1209.127951.figure7" httpUri="https://binary.pensoft.net/fig/1113531">7 A</figureCitation>
,
<figureCitation id="41F03F23A63D0BB126A7EB0F5B4A6297" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Female genitalia in ventral and dorsal views A, B Belisana langping sp. nov. C, D B. lingui sp. nov. E, F B. tianyang sp. nov. Abbreviations: aa = anterior arch, ep = epigynal pocket, pp = pore plate. Scale bars: 0.10 mm." figureDoi="10.3897/zookeys.1209.127951.figure9" httpUri="https://binary.pensoft.net/fig/1113533">9 E</figureCitation>
). Vulva with ridge-shaped anterior arch (
<abbrev id="ABBRID0E5EBG" xlink_title="anterior arch">aa</abbrev>
in Figs
<figureCitation id="2AF02AEF9A72914B3E64D232D1E47A56" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Belisana tianyang sp. nov., holotype male (C – F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E – H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 mm (A – D); 0.50 mm (E – H)." figureDoi="10.3897/zookeys.1209.127951.figure7" httpUri="https://binary.pensoft.net/fig/1113531">7 B</figureCitation>
,
<figureCitation id="5B6BC8F6B47C47769C3E02D36A623DD9" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Female genitalia in ventral and dorsal views A, B Belisana langping sp. nov. C, D B. lingui sp. nov. E, F B. tianyang sp. nov. Abbreviations: aa = anterior arch, ep = epigynal pocket, pp = pore plate. Scale bars: 0.10 mm." figureDoi="10.3897/zookeys.1209.127951.figure9" httpUri="https://binary.pensoft.net/fig/1113533">9 F</figureCitation>
) and pair of curved, anteriorly pointed and posteriorly wide pore plates (
<abbrev id="ABBRID0EKFBG" xlink_title="pore plate">pp</abbrev>
in Figs
<figureCitation id="7E49C8C609AC919A9020C4CB13B9B798" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Belisana tianyang sp. nov., holotype male (C – F) and paratype female (A, B, G, H) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E – H habitus (E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 mm (A – D); 0.50 mm (E – H)." figureDoi="10.3897/zookeys.1209.127951.figure7" httpUri="https://binary.pensoft.net/fig/1113531">7 B</figureCitation>
,
<figureCitation id="3600AD2AF0C83148044AEC4816B80C1B" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Female genitalia in ventral and dorsal views A, B Belisana langping sp. nov. C, D B. lingui sp. nov. E, F B. tianyang sp. nov. Abbreviations: aa = anterior arch, ep = epigynal pocket, pp = pore plate. Scale bars: 0.10 mm." figureDoi="10.3897/zookeys.1209.127951.figure9" httpUri="https://binary.pensoft.net/fig/1113533">9 F</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EWFBG" type="variation">
<paragraph id="4282E7285C0FA832B71A13C026734A45">
<heading id="DF809F4BDC09B3A27415BD19C013E851" reason="title">Variation.</heading>
</paragraph>
<paragraph id="9983AF8ABDAC0221C53C99E31A4382AC">
Tibia I in
<specimenCount id="07063DEF81CB6A5B30DA5D1C19584D8E" count="3" type="male">three male</specimenCount>
<typeStatus id="7616D6FB40B796A8155628E98B63F30A" type="paratype">paratypes</typeStatus>
(
<named-content id="F50BE15E798A949EC770566E4D04D0B7" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/institute-zoology-chinese-academy-sciences" xlink_title="Institute of Zoology, Chinese Academy of Sciences">
<collectionCode id="2D3D58D2AE80A13F20982B684CCF4344" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" type="">IZCAS</collectionCode>
</named-content>
-
<specimenCode id="35B69D1F233D709D95214BC2F4686F43">Ar 44998</specimenCode>
–45000): 5.38, 5.64, 5.77. Tibia I in another female
<typeStatus id="1C031DEC96686B06571015B061FDC11C" type="paratype">paratype</typeStatus>
(
<named-content id="F2C2E56B4F965F0C590CBF94DE8FF631" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/institute-zoology-chinese-academy-sciences" xlink_title="Institute of Zoology, Chinese Academy of Sciences">
<collectionCode id="72792A41AE80CA4966C9E06A737B92D9" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" type="">IZCAS</collectionCode>
</named-content>
-
<specimenCode id="9A1E7028B30288E85E58D450917C26F0">Ar 45002</specimenCode>
) missing.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EEGBG" type="habitat">
<paragraph id="65E245E1E349CC35AB5EC84B9CAD4E1E">
<heading id="18A837813BA5335114A86DA27736BA5D" reason="title">Habitat.</heading>
</paragraph>
<paragraph id="F787627BFEB1048D907CD8DDB47A482F">The species was found in the dark zone inside the cave.</paragraph>
</subSubSection>
<subSubSection id="SECID0EJGBG" type="distribution">
<paragraph id="26CD00CA8D7CD808BB53ECE710E96869">
<heading id="EE42DDE8D0FBD070918C3D98AE814C3E" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="B0835EC80C6981D069E689E1BAE1F592">
<collectingCountry id="7FE79A7506F17005E64B1D10125BE74A" name="China">China</collectingCountry>
(
<collectingRegion id="E63C715156F726CD97166B55658033CD" country="China" name="Guangxi">Guangxi</collectingRegion>
, type locality; Fig.
<figureCitation id="6421B14868F281EF8B9F3F1D7F1D9527" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Distribution records of the Belisana species from Guangxi, China 1 Belisana colubrina Zhang &amp; Peng, 2011 2 B. guilin Yao &amp; Li, 2020 3 B. langping sp. nov. 4 B. lingui sp. nov. 5 B. naling Yao &amp; Li, 2020 6 B. parallelica Zhang &amp; Peng, 2011 7 B. tianlinensis Zhang &amp; Peng, 2011 8 B. tianyang sp. nov. 9 B. tongle Zhang, Chen &amp; Zhu, 2008 10 B. xuanguan Yao &amp; Li, 2020 11 B. zhangi Tong &amp; Li, 2007." figureDoi="10.3897/zookeys.1209.127951.figure1" httpUri="https://binary.pensoft.net/fig/1113525">1</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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