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<document id="3ADC94E0599400DB18938C00F6A4C7B3" ID-CLB-Dataset="111181" ID-DOI="10.11646/zootaxa.5169.5.1" ID-GBIF-Dataset="d8cabaec-c613-4882-8a1c-e08d69a084e7" ID-ISSN="1175-5326" ID-Zenodo-Dep="6952712" ID-ZooBank="EB5A2C38-2D31-4F02-BEC1-2AA31761EFEA" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1659441294752" checkinUser="plazi" docAuthor="Atherton, Sarah &amp; Jondelius, Ulf" docDate="2022" docId="CF6B87F3E62F8503FF6FF934FC2DFB87" docLanguage="en" docName="zootaxa.5169.5.1.pdf" docOrigin="Zootaxa 5169 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pseudoposthia WESTbLaD 1946" docType="treatment" docVersion="8" lastPageNumber="422" masterDocId="3352FF8BE63B8516FFF8FFEEFF82FFC9" masterDocTitle="Diversity in the family Isodiametridae (Acoela): New species bring back old problems" masterLastPageNumber="424" masterPageNumber="401" pageNumber="421" updateTime="1699363415466" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="58786988293ECAF2B4CB87B35426D11F">Diversity in the family Isodiametridae (Acoela): New species bring back old problems</mods:title>
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<mods:namePart id="1096DACEE446E0881ED4DB23E7370E99">Atherton, Sarah</mods:namePart>
<mods:nameIdentifier id="51136928C40619C9829D0F7780CE3E6F" type="ORCID">0000-0003-3024-472X</mods:nameIdentifier>
<mods:affiliation id="3ADD2A930468673050B6FF84050AACD1">Department of Zoology, Naturhistoriska riksmuseet, Stockholm, Sweden &amp; sarah. atherton @ nrm. se; https: // orcid. org / 0000 - 0003 - 3024 - 472 X</mods:affiliation>
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<mods:namePart id="B1776032E4EAB3170E7AC033CBC10E42">Jondelius, Ulf</mods:namePart>
<mods:affiliation id="E2B42FFED332EE258D00899277CDEDE9">Department of Zoology, Naturhistoriska riksmuseet, Stockholm, Sweden &amp; Department of Zoology, Systematics and Evolution, Stockholm Universitet, Stockholm Sweden &amp; ulf. jondelius @ nrm. se; https: // orcid. ord / 0000 - 0003 - 2847 - 2192</mods:affiliation>
<mods:nameIdentifier id="4554627C527A764B29FC3D9FA741965F" type="email">ulf.jondelius@nrm.se</mods:nameIdentifier>
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<mods:title id="FBD3B0B1456B6E6E89A05F7CD7F930A5">Zootaxa</mods:title>
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<mods:date id="0B811931B5F428CDE44C698E8FD1D552">2022</mods:date>
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<mods:number id="CAB0E30B43864F16EEAC18F5497E757C">2022-08-02</mods:number>
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<treatment id="CF6B87F3E62F8503FF6FF934FC2DFB87" ID-DOI="http://doi.org/10.5281/zenodo.6953871" ID-GBIF-Taxon="198491535" ID-Zenodo-Dep="6953871" LSID="urn:lsid:plazi:treatment:CF6B87F3E62F8503FF6FF934FC2DFB87" httpUri="http://treatment.plazi.org/id/CF6B87F3E62F8503FF6FF934FC2DFB87" lastPageId="21" lastPageNumber="422" pageId="20" pageNumber="421">
<subSubSection id="0FD8656EE62F8502FF6FF934FEBDF93C" box="[151,319,1754,1781]" pageId="20" pageNumber="421" type="nomenclature">
<paragraph id="477D36E5E62F8502FF6FF934FEBDF93C" blockId="20.[151,1436,1754,2033]" box="[151,319,1754,1781]" pageId="20" pageNumber="421">
<heading id="1C358189E62F8502FF6FF934FEBDF93C" bold="true" box="[151,319,1754,1781]" fontSize="11" level="1" pageId="20" pageNumber="421" reason="1">
<taxonomicName id="80C24D66E62F8502FF6FF934FEBDF93C" ID-CoL="7PWNB" authorityName="WESTbLaD" authorityYear="1946" box="[151,319,1754,1781]" family="Isodiametridae" genus="Pseudoposthia" kingdom="Animalia" order="Acoela" pageId="20" pageNumber="421" phylum="Xenacoelomorpha" rank="genus">
<emphasis id="75B6EAF7E62F8502FF6FF934FEBDF93C" bold="true" box="[151,319,1754,1781]" italics="true" pageId="20" pageNumber="421">Pseudoposthia</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="0FD8656EE62F8503FF3FF911FDA2FE64" lastPageId="21" lastPageNumber="422" pageId="20" pageNumber="421" type="description">
<paragraph id="477D36E5E62F8502FF3FF911FC89F838" blockId="20.[151,1436,1754,2033]" pageId="20" pageNumber="421">
Only a single species of
<taxonomicName id="80C24D66E62F8502FE2EF911FDF9F8D0" authorityName="WESTbLaD" authorityYear="1946" box="[470,635,1791,1817]" family="Isodiametridae" genus="Pseudoposthia" kingdom="Animalia" order="Acoela" pageId="20" pageNumber="421" phylum="Xenacoelomorpha" rank="genus">
<emphasis id="75B6EAF7E62F8502FE2EF911FDF9F8D0" box="[470,635,1791,1817]" italics="true" pageId="20" pageNumber="421">Pseudoposthia</emphasis>
</taxonomicName>
,
<taxonomicName id="80C24D66E62F8502FD70F911FCD4F8D1" authorityName="Westblad" authorityYear="1946" box="[648,854,1791,1816]" family="Isodiametridae" genus="Pseudoposthia" kingdom="Animalia" order="Acoela" pageId="20" pageNumber="421" phylum="Xenacoelomorpha" rank="species" species="macrogonopora">
<emphasis id="75B6EAF7E62F8502FD70F911FCD4F8D1" box="[648,854,1791,1816]" italics="true" pageId="20" pageNumber="421">P. macrogonopora</emphasis>
</taxonomicName>
, was previously known, and its reproductive system notably also does not follow that which is typical of
<taxonomicName id="80C24D66E62F8502FD2FF8CDFC03F8F4" authorityName="HOOGE &amp; TYLER" authorityYear="2005" box="[727,897,1827,1853]" family="Isodiametridae" kingdom="Animalia" order="Acoela" pageId="20" pageNumber="421" phylum="Xenacoelomorpha" rank="family">Isodiametridae</taxonomicName>
. In this case,
<taxonomicName id="80C24D66E62F8502FBEAF8CDFB5DF8F5" authorityName="Westblad" authorityYear="1946" box="[1042,1247,1827,1852]" family="Isodiametridae" genus="Pseudoposthia" kingdom="Animalia" order="Acoela" pageId="20" pageNumber="421" phylum="Xenacoelomorpha" rank="species" species="macrogonopora">
<emphasis id="75B6EAF7E62F8502FBEAF8CDFB5DF8F5" box="[1042,1247,1827,1852]" italics="true" pageId="20" pageNumber="421">P. macrogonopora</emphasis>
</taxonomicName>
does not possess a muscular seminal vesicle with an invaginated isodiametric penis but rather a false seminal vesicle unconnected to the muscular penis (
<bibRefCitation id="23534B14E62F8502FE81F885FDA4F84C" author="Westblad, E." box="[377,550,1899,1925]" pageId="20" pageNumber="421" pagination="1 - 56" refId="ref15876" refString="Westblad, E. (1946) Studien uber Skandinavische Turbellaria Acoela IV. Arkiv for Zoologi, 38, 1 - 56." type="journal article" year="1946">Westblad 1946</bibRefCitation>
). Its reproductive system is additionally unusual for
<taxonomicName id="80C24D66E62F8502FB8DF885FB46F84C" box="[1141,1220,1899,1925]" kingdom="Animalia" order="Acoela" pageId="20" pageNumber="421" phylum="Xenacoelomorpha" rank="order">Acoela</taxonomicName>
in that the seminal vesicle is not fully posterior to the ovaries. Sequences of
<taxonomicName id="80C24D66E62F8502FCE3F861FC68F861" authorityName="Westblad" authorityYear="1946" box="[795,1002,1935,1960]" family="Isodiametridae" genus="Pseudoposthia" kingdom="Animalia" order="Acoela" pageId="20" pageNumber="421" phylum="Xenacoelomorpha" rank="species" species="macrogonopora">
<emphasis id="75B6EAF7E62F8502FCE3F861FC68F861" box="[795,1002,1935,1960]" italics="true" pageId="20" pageNumber="421">P. macrogonopora</emphasis>
</taxonomicName>
are presented here for the first time as well as two additional new species of
<taxonomicName id="80C24D66E62F8502FDB2F85DFD6DF804" authorityName="WESTbLaD" authorityYear="1946" box="[586,751,1971,1997]" family="Isodiametridae" genus="Pseudoposthia" kingdom="Animalia" order="Acoela" pageId="20" pageNumber="421" phylum="Xenacoelomorpha" rank="genus">
<emphasis id="75B6EAF7E62F8502FDB2F85DFD6DF804" box="[586,751,1971,1997]" italics="true" pageId="20" pageNumber="421">Pseudoposthia</emphasis>
</taxonomicName>
. Results from our phylogenetic analyses find the clade does nest within
<taxonomicName id="80C24D66E62F8502FEE2F839FE42F838" authorityName="HOOGE &amp; TYLER" authorityYear="2005" box="[282,448,2007,2033]" family="Isodiametridae" kingdom="Animalia" order="Acoela" pageId="20" pageNumber="421" phylum="Xenacoelomorpha" rank="family">Isodiametridae</taxonomicName>
with high support (
<figureCitation id="DFF92A60E62F8502FD63F839FD78F838" box="[667,762,2007,2033]" captionStart="FIGURE 1" captionStartId="4.[151,250,1943,1967]" captionTargetBox="[232,1353,189,1906]" captionTargetId="figure-23@4.[226,1362,179,1918]" captionTargetPageId="4" captionText="FIGURE 1. Concatenated 18S and COI gene tree summary. Percent bootstrap values are given at each node. New species are highlighted in red." figureDoi="http://doi.org/10.5281/zenodo.6952714" httpUri="https://zenodo.org/record/6952714/files/figure.png" pageId="20" pageNumber="421">Figure 1</figureCitation>
).
</paragraph>
<paragraph id="477D36E5E62E8503FF3FFF79FDA2FE64" blockId="21.[151,1437,151,429]" pageId="21" pageNumber="422">
The reproductive systems of both new species are similar to that of
<taxonomicName id="80C24D66E62E8503FC49FF79FBFCFF79" authorityName="Westblad" authorityYear="1946" box="[945,1150,151,176]" family="Isodiametridae" genus="Pseudoposthia" kingdom="Animalia" order="Acoela" pageId="21" pageNumber="422" phylum="Xenacoelomorpha" rank="species" species="macrogonopora">
<emphasis id="75B6EAF7E62E8503FC49FF79FBFCFF79" box="[945,1150,151,176]" italics="true" pageId="21" pageNumber="422">P. macrogonopora</emphasis>
</taxonomicName>
, both in the anterior position of the male system, the false seminal vesicle unconnected to a penis and the lack of female accessory organs. Additionally, all three species are unpigmented and possess numerous rhabdoid glands scattered across the body.
<taxonomicName id="80C24D66E62E8503FA70FF31FF79FED4" authority="Atherton &amp; Jondelius, 2022" authorityName="Atherton &amp; Jondelius" authorityYear="2022" family="Isodiametridae" genus="Pseudoposthia" kingdom="Animalia" order="Acoela" pageId="21" pageNumber="422" phylum="Xenacoelomorpha" rank="species" species="hanssoni" status="sp. nov.">
<emphasis id="75B6EAF7E62E8503FA70FF31FF79FED4" italics="true" pageId="21" pageNumber="422">P. hanssoni</emphasis>
</taxonomicName>
<emphasis id="75B6EAF7E62E8503FEFAFEEDFEDEFED4" bold="true" box="[258,348,259,285]" pageId="21" pageNumber="422">
<taxonomicNameLabel id="6E85578CE62E8503FEFAFEEDFEDEFED4" box="[258,348,259,285]" pageId="21" pageNumber="422" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be differentiated from
<taxonomicName id="80C24D66E62E8503FD68FEEDFCDDFED5" authorityName="Westblad" authorityYear="1946" box="[656,863,259,284]" family="Isodiametridae" genus="Pseudoposthia" kingdom="Animalia" order="Acoela" pageId="21" pageNumber="422" phylum="Xenacoelomorpha" rank="species" species="macrogonopora">
<emphasis id="75B6EAF7E62E8503FD68FEEDFCDDFED5" box="[656,863,259,284]" italics="true" pageId="21" pageNumber="422">P. macrogonopora</emphasis>
</taxonomicName>
by its smaller body size, a smaller sized penis that is also straight, and by the large vacuoles that are restricted to the posterior body half only.
<taxonomicName id="80C24D66E62E8503FB59FEC9FB68FE88" authority="Atherton &amp; Jondelius, 2022" authorityName="Atherton &amp; Jondelius" authorityYear="2022" box="[1185,1258,295,321]" family="Isodiametridae" genus="Pseudoposthia" kingdom="Animalia" order="Acoela" pageId="21" pageNumber="422" phylum="Xenacoelomorpha" rank="species" species="lata" status="sp. nov.">
<emphasis id="75B6EAF7E62E8503FB59FEC9FB68FE88" box="[1185,1258,295,321]" italics="true" pageId="21" pageNumber="422">P. lata</emphasis>
</taxonomicName>
<emphasis id="75B6EAF7E62E8503FB09FEC9FAC9FE88" bold="true" box="[1265,1355,295,321]" pageId="21" pageNumber="422">
<taxonomicNameLabel id="6E85578CE62E8503FB09FEC9FAC9FE88" box="[1265,1355,295,321]" pageId="21" pageNumber="422" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be distinguished from
<taxonomicName id="80C24D66E62E8503FE8CFEA5FDC6FEAD" authorityName="Westblad" authorityYear="1946" box="[372,580,331,356]" family="Isodiametridae" genus="Pseudoposthia" kingdom="Animalia" order="Acoela" pageId="21" pageNumber="422" phylum="Xenacoelomorpha" rank="species" species="macrogonopora">
<emphasis id="75B6EAF7E62E8503FE8CFEA5FDC6FEAD" box="[372,580,331,356]" italics="true" pageId="21" pageNumber="422">P. macrogonopora</emphasis>
</taxonomicName>
and
<taxonomicName id="80C24D66E62E8503FD86FEA5FD7DFEAC" authority="Atherton &amp; Jondelius, 2022" authorityName="Atherton &amp; Jondelius" authorityYear="2022" box="[638,767,331,357]" family="Isodiametridae" genus="Pseudoposthia" kingdom="Animalia" order="Acoela" pageId="21" pageNumber="422" phylum="Xenacoelomorpha" rank="species" species="hanssoni" status="sp. nov.">
<emphasis id="75B6EAF7E62E8503FD86FEA5FD7DFEAC" box="[638,767,331,357]" italics="true" pageId="21" pageNumber="422">P. hanssoni</emphasis>
</taxonomicName>
<emphasis id="75B6EAF7E62E8503FCFFFEA5FCE0FEAC" bold="true" box="[775,866,331,357]" pageId="21" pageNumber="422">
<taxonomicNameLabel id="6E85578CE62E8503FCFFFEA5FCE0FEAC" box="[775,866,331,357]" pageId="21" pageNumber="422" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in its general body shape, the medium size of the frontal organs and by the fact that the muscular penis is absent.
<tableCitation id="0A40035EE62E8503FCA4FE81FC2CFE40" box="[860,942,367,393]" captionStart="TABLE 3" captionStartId="21.[151,239,476,500]" captionTargetPageId="21" captionText="TABLE 3. Selected morphological characters for species of Pseudoposthia." httpUri="http://table.plazi.org/id/13BD666DE62E8503FF6FFE32FC43FE3D" pageId="21" pageNumber="422" tableUuid="13BD666DE62E8503FF6FFE32FC43FE3D">Table 3</tableCitation>
summarizes the primary morphological differences between the three species.
</paragraph>
</subSubSection>
<caption id="13BD666DE62E8503FF6FFE32FC43FE3D" ID-Table-UUID="13BD666DE62E8503FF6FFE32FC43FE3D" box="[151,961,476,500]" httpUri="http://table.plazi.org/id/13BD666DE62E8503FF6FFE32FC43FE3D" pageId="21" pageNumber="422" startId="21.[151,239,476,500]" targetBox="[162,1401,516,812]" targetIsTable="true" targetPageId="21">
<paragraph id="477D36E5E62E8503FF6FFE32FC43FE3D" blockId="21.[151,961,476,500]" box="[151,961,476,500]" pageId="21" pageNumber="422">
<emphasis id="75B6EAF7E62E8503FF6FFE32FE88FE3D" bold="true" box="[151,266,476,500]" pageId="21" pageNumber="422">TABLE 3.</emphasis>
Selected morphological characters for species of
<taxonomicName id="80C24D66E62E8503FCE5FE32FC38FE3D" authorityName="WESTbLaD" authorityYear="1946" box="[797,954,476,500]" family="Isodiametridae" genus="Pseudoposthia" kingdom="Animalia" order="Acoela" pageId="21" pageNumber="422" phylum="Xenacoelomorpha" rank="genus">
<emphasis id="75B6EAF7E62E8503FCE5FE32FC38FE3D" box="[797,954,476,500]" italics="true" pageId="21" pageNumber="422">Pseudoposthia</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="477D36E5E62E8503FE1CFDEBFB2AFCE5" pageId="21" pageNumber="422">
<table id="35C2C445E62E7AE9FF5AFDEAFAFBFCE5" box="[162,1401,516,812]" gridcols="4" gridrows="8" pageId="21" pageNumber="422">
<tr id="F9F234A7E62E7AE9FF5AFDEAFAFBFDD5" box="[162,1401,516,540]" gridrow="0" pageId="21" pageNumber="422" rowspan-0="1">
<th id="BA235DDBE62E7AE9FE1CFDEAFC87FDD5" box="[484,773,516,540]" gridcol="1" gridrow="0" pageId="21" pageNumber="422">
<taxonomicName id="80C24D66E62E8503FE1CFDEBFD1DFDD5" authorityName="Westblad" authorityYear="1946" box="[484,671,517,540]" family="Isodiametridae" genus="Pseudoposthia" kingdom="Animalia" order="Acoela" pageId="21" pageNumber="422" phylum="Xenacoelomorpha" rank="species" species="macrogonopora">
<emphasis id="75B6EAF7E62E8503FE1CFDEBFD1DFDD5" box="[484,671,517,540]" italics="true" pageId="21" pageNumber="422">P. macrogonopora</emphasis>
</taxonomicName>
</th>
<th id="BA235DDBE62E7AE9FCDEFDEAFC5DFDD5" box="[806,991,516,540]" gridcol="2" gridrow="0" pageId="21" pageNumber="422">
<taxonomicName id="80C24D66E62E8503FCDEFDEBFCE4FDD5" authority="Atherton &amp; Jondelius, 2022" authorityName="Atherton &amp; Jondelius" authorityYear="2022" box="[806,870,516,540]" family="Isodiametridae" genus="Pseudoposthia" kingdom="Animalia" order="Acoela" pageId="21" pageNumber="422" phylum="Xenacoelomorpha" rank="species" species="lata" status="sp. nov.">
<emphasis id="75B6EAF7E62E8503FCDEFDEBFCE4FDD5" box="[806,870,516,540]" italics="true" pageId="21" pageNumber="422">P. lata</emphasis>
</taxonomicName>
<emphasis id="75B6EAF7E62E8503FC94FDEAFC3FFDD5" bold="true" box="[876,957,516,540]" pageId="21" pageNumber="422">
<taxonomicNameLabel id="6E85578CE62E8503FC94FDEAFC3FFDD5" box="[876,957,516,540]" pageId="21" pageNumber="422" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</th>
<th id="BA235DDBE62E7AE9FB9FFDEAFAFBFDD5" box="[1127,1401,516,540]" gridcol="3" gridrow="0" pageId="21" pageNumber="422">
<taxonomicName id="80C24D66E62E8503FB9FFDEBFB58FDD5" authority="Atherton &amp; Jondelius, 2022" authorityName="Atherton &amp; Jondelius" authorityYear="2022" box="[1127,1242,516,540]" family="Isodiametridae" genus="Pseudoposthia" kingdom="Animalia" order="Acoela" pageId="21" pageNumber="422" phylum="Xenacoelomorpha" rank="species" species="hanssoni" status="sp. nov.">
<emphasis id="75B6EAF7E62E8503FB9FFDEBFB58FDD5" box="[1127,1242,516,540]" italics="true" pageId="21" pageNumber="422">P. hanssoni</emphasis>
</taxonomicName>
<emphasis id="75B6EAF7E62E8503FB19FDEAFAB3FDD5" bold="true" box="[1249,1329,516,540]" pageId="21" pageNumber="422">
<taxonomicNameLabel id="6E85578CE62E8503FB19FDEAFAB3FDD5" box="[1249,1329,516,540]" pageId="21" pageNumber="422" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</th>
</tr>
<tr id="F9F234A7E62E7AE9FF5AFDC5FAFBFD8A" box="[162,1401,555,579]" gridrow="1" pageId="21" pageNumber="422">
<th id="BA235DDBE62E7AE9FF5AFDC5FEDFFD8A" box="[162,349,555,579]" gridcol="0" gridrow="1" pageId="21" pageNumber="422">Body length (mm)</th>
<td id="BA235DDBE62E7AE9FE1CFDC5FC87FD8A" box="[484,773,555,579]" gridcol="1" gridrow="1" pageId="21" pageNumber="422">0.70</td>
<td id="BA235DDBE62E7AE9FCDEFDC5FC5DFD8A" box="[806,991,555,579]" gridcol="2" gridrow="1" pageId="21" pageNumber="422">0.35–0.50</td>
<td id="BA235DDBE62E7AE9FB9FFDC5FAFBFD8A" box="[1127,1401,555,579]" gridcol="3" gridrow="1" pageId="21" pageNumber="422">0.45</td>
</tr>
<tr id="F9F234A7E62E7AE9FF5AFDBCFAFBFDA3" box="[162,1401,594,618]" gridrow="2" pageId="21" pageNumber="422">
<th id="BA235DDBE62E7AE9FF5AFDBCFEDFFDA3" box="[162,349,594,618]" gridcol="0" gridrow="2" pageId="21" pageNumber="422">Body width (mm)</th>
<td id="BA235DDBE62E7AE9FE1CFDBCFC87FDA3" box="[484,773,594,618]" gridcol="1" gridrow="2" pageId="21" pageNumber="422">0.20</td>
<td id="BA235DDBE62E7AE9FCDEFDBCFC5DFDA3" box="[806,991,594,618]" gridcol="2" gridrow="2" pageId="21" pageNumber="422">0.25</td>
<td id="BA235DDBE62E7AE9FB9FFDBCFAFBFDA3" box="[1127,1401,594,618]" gridcol="3" gridrow="2" pageId="21" pageNumber="422">0.15</td>
</tr>
<tr id="F9F234A7E62E7AE9FF5AFD97FAFBFD58" box="[162,1401,633,657]" gridrow="3" pageId="21" pageNumber="422">
<th id="BA235DDBE62E7AE9FF5AFD97FEDFFD58" box="[162,349,633,657]" gridcol="0" gridrow="3" pageId="21" pageNumber="422">Frontal organ</th>
<td id="BA235DDBE62E7AE9FE1CFD97FC87FD58" box="[484,773,633,657]" gridcol="1" gridrow="3" pageId="21" pageNumber="422">Well-developed</td>
<td id="BA235DDBE62E7AE9FCDEFD97FC5DFD58" box="[806,991,633,657]" gridcol="2" gridrow="3" pageId="21" pageNumber="422">Medium</td>
<td id="BA235DDBE62E7AE9FB9FFD97FAFBFD58" box="[1127,1401,633,657]" gridcol="3" gridrow="3" pageId="21" pageNumber="422">Well-developed</td>
</tr>
<tr id="F9F234A7E62E7AE9FF5AFD4EFAFBFD71" box="[162,1401,672,696]" gridrow="4" pageId="21" pageNumber="422">
<th id="BA235DDBE62E7AE9FF5AFD4EFEDFFD71" box="[162,349,672,696]" gridcol="0" gridrow="4" pageId="21" pageNumber="422">Vacuoles</th>
<td id="BA235DDBE62E7AE9FE1CFD4EFC87FD71" box="[484,773,672,696]" gridcol="1" gridrow="4" pageId="21" pageNumber="422">Numerous along entire body</td>
<td id="BA235DDBE62E7AE9FCDEFD4EFC5DFD71" box="[806,991,672,696]" gridcol="2" gridrow="4" pageId="21" pageNumber="422">Posterior end only</td>
<td id="BA235DDBE62E7AE9FB9FFD4EFAFBFD71" box="[1127,1401,672,696]" gridcol="3" gridrow="4" pageId="21" pageNumber="422">Numerous in posterior half</td>
</tr>
<tr id="F9F234A7E62E7AE9FF5AFD28FAFBFD16" box="[162,1401,710,735]" gridrow="5" pageId="21" pageNumber="422">
<th id="BA235DDBE62E7AE9FF5AFD28FEDFFD16" box="[162,349,710,735]" gridcol="0" gridrow="5" pageId="21" pageNumber="422">Penis length (µm)</th>
<td id="BA235DDBE62E7AE9FE1CFD28FC87FD16" box="[484,773,710,735]" gridcol="1" gridrow="5" pageId="21" pageNumber="422">30</td>
<td id="BA235DDBE62E7AE9FCDEFD28FC5DFD16" box="[806,991,710,735]" gridcol="2" gridrow="5" pageId="21" pageNumber="422">none</td>
<td id="BA235DDBE62E7AE9FB9FFD28FAFBFD16" box="[1127,1401,710,735]" gridcol="3" gridrow="5" pageId="21" pageNumber="422">9–10</td>
</tr>
<tr id="F9F234A7E62E7AE9FF5AFD03FAFBFCCC" box="[162,1401,749,773]" gridrow="6" pageId="21" pageNumber="422">
<th id="BA235DDBE62E7AE9FF5AFD03FEDFFCCC" box="[162,349,749,773]" gridcol="0" gridrow="6" pageId="21" pageNumber="422">Penis shape</th>
<td id="BA235DDBE62E7AE9FE1CFD03FC87FCCC" box="[484,773,749,773]" gridcol="1" gridrow="6" pageId="21" pageNumber="422">curved</td>
<td id="BA235DDBE62E7AE9FCDEFD03FC5DFCCC" box="[806,991,749,773]" gridcol="2" gridrow="6" pageId="21" pageNumber="422">none</td>
<td id="BA235DDBE62E7AE9FB9FFD03FAFBFCCC" box="[1127,1401,749,773]" gridcol="3" gridrow="6" pageId="21" pageNumber="422">straight</td>
</tr>
<tr id="F9F234A7E62E7AE9FF5AFCFAFAFBFCE5" box="[162,1401,788,812]" gridrow="7" pageId="21" pageNumber="422">
<th id="BA235DDBE62E7AE9FF5AFCFAFEDFFCE5" box="[162,349,788,812]" gridcol="0" gridrow="7" pageId="21" pageNumber="422">Depth</th>
<td id="BA235DDBE62E7AE9FE1CFCFAFC87FCE5" box="[484,773,788,812]" gridcol="1" gridrow="7" pageId="21" pageNumber="422">15–20 m</td>
<td id="BA235DDBE62E7AE9FCDEFCFAFC5DFCE5" box="[806,991,788,812]" gridcol="2" gridrow="7" pageId="21" pageNumber="422">6–10 m</td>
<td id="BA235DDBE62E7AE9FB9FFCFAFAFBFCE5" box="[1127,1401,788,812]" gridcol="3" gridrow="7" pageId="21" pageNumber="422">4–6 m</td>
</tr>
</table>
</paragraph>
<subSubSection id="0FD8656EE62E8503FF3FFCB5FC2DFB87" pageId="21" pageNumber="422" type="diagnosis">
<paragraph id="477D36E5E62E8503FF3FFCB5FC2DFB87" blockId="21.[151,1437,859,1102]" pageId="21" pageNumber="422">
<emphasis id="75B6EAF7E62E8503FF3FFCB5FE2AFCBC" box="[199,424,859,885]" italics="true" pageId="21" pageNumber="422">Emended diagnosis.</emphasis>
Body without pigmentation. Frontal organ present, often well-developed. Rhabdoid glands numerous, scattered. Mouth ventral, located approximately at the body’s midpoint and slightly anterior to the male gonopore. Ovaries and testes paired. Seminal bursa, vagina and female gonopore absent. Male reproductive system and gonopore present anterior to the posteriormost portion of the ovary. Male gonopore ventral, surrounded by a heavy muscular sphincter. Penis, when present, without ejactulatory duct and unconnected to a seminal vesicle. False seminal vesicle without walls or musculature present, connecting directly to the gonopore posterior to the penis. Numerous gland cells associated with penis and male gonopore.
</paragraph>
</subSubSection>
</treatment>
</document>
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