265 lines
40 KiB
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265 lines
40 KiB
XML
<document id="20D3AF24EBB9286A44D85E6B5F543F71" ID-DOI="10.5281/zenodo.4504075" ID-ISSN="2345-7600" ID-Zenodo-Dep="4504075" ID-ZooBank="2300F64E-BD30-4E06-A24F-A226185A4BD3" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1612465590805" checkinUser="carolina" docAuthor="Siriboon, Thanit, Sutcharit, Chirasak, Naggs, Fred, Rowson, Ben & Panha, Somsak" docDate="2014" docId="781A87B5C4469C34FCB53D59FDB29857" docLanguage="en" docName="RafflesBZool.62.161-174.pdf" docOrigin="Raffles Bulletin of Zoology 62" docStyle="DocumentStyle:F2D34FF25DBF9998BB8DF6F24A5B097A.5:RafflesBZool.2014-2018.journal_article" docStyleId="F2D34FF25DBF9998BB8DF6F24A5B097A" docStyleName="RafflesBZool.2014-2018.journal_article" docStyleVersion="5" docTitle="Carinartemis striatus Siriboon & Sutcharit & Naggs & Rowson & Panha 2014, new species" docType="treatment" docUuid="11592BC8-C051-4269-8B1A-FAC73AA8F620" docUuidSource="ZooBank" docVersion="5" lastPageNumber="172" masterDocId="8423FFCDC44C9C3FFFE23E44FFF99F43" masterDocTitle="Revision of the carnivorous snail genus Indoartemon Forcart, 1946 and a new genus Carinartemis from Thailand (Pulmonata: Streptaxidae)" masterLastPageNumber="174" masterPageNumber="161" pageNumber="171" updateTime="1698865986014" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="8EB3772DA592F6E082FEB624E354C85D">Revision of the carnivorous snail genus Indoartemon Forcart, 1946 and a new genus Carinartemis from Thailand (Pulmonata: Streptaxidae)</mods:title>
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<mods:namePart id="12236B4630B58BBF974F065346351527">Siriboon, Thanit</mods:namePart>
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<mods:namePart id="B8B9213941A1E7D3B0E529FF1101ABA0">Sutcharit, Chirasak</mods:namePart>
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<mods:namePart id="7C9CC75DE901FA83EB609E45BC5DC220">Naggs, Fred</mods:namePart>
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<mods:namePart id="B9C2DD38FFBB5EC15BCB3A0CC85AC84F">Rowson, Ben</mods:namePart>
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<mods:namePart id="4D7C9D417A8E4F9F5EC731E76FADD2D9">Panha, Somsak</mods:namePart>
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<mods:title id="43016FA592A64DE043B5B4F87FE4C81D">Raffles Bulletin of Zoology</mods:title>
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<mods:date id="565211DA459BE00C1EC11021D65B3934">2014</mods:date>
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<mods:number id="8CF6E603433AF687D4E2A2D0D6AACDE3">2014-04-04</mods:number>
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<subSubSection id="B8A96528C4469C35FCB53D59FA579C76" box="[855,1454,797,821]" pageId="10" pageNumber="171" type="nomenclature">
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<paragraph id="F00C36A3C4469C35FCB53D59FA579C76" blockId="10.[855,1454,797,853]" box="[855,1454,797,821]" pageId="10" pageNumber="171">
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<heading id="AB4481CFC4469C35FCB53D59FA579C76" bold="true" box="[855,1454,797,821]" centered="true" fontSize="10" level="2" pageId="10" pageNumber="171" reason="2">
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<emphasis id="C2C7EAB1C4469C35FCB53D59FA579C76" bold="true" box="[855,1454,797,821]" pageId="10" pageNumber="171">
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<taxonomicName id="37B34D20C4469C35FCB53D59FAE59C76" ID-CoL="RD94" authority="Siriboon & Panha" authorityName="Siriboon & Sutcharit & Naggs & Rowson & Panha" authorityYear="2014" box="[855,1308,797,821]" class="Gastropoda" family="Streptaxidae" genus="Carinartemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="171" phylum="Mollusca" rank="species" species="striatus" status="sp. nov.">
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<emphasis id="C2C7EAB1C4469C35FCB53D59FBBA9C76" bold="true" box="[855,1091,797,821]" italics="true" pageId="10" pageNumber="171">Carinartemis striatus</emphasis>
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Siriboon & Panha
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</taxonomicName>
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,
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<taxonomicNameLabel id="D9F457CAC4469C35FACB3D5AFA579C76" box="[1321,1454,798,821]" pageId="10" pageNumber="171" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<subSubSection id="B8A96528C4469C35FC7F3D7AFA909C16" box="[925,1385,830,853]" pageId="10" pageNumber="171" type="description">
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<paragraph id="F00C36A3C4469C35FC7F3D7AFA909C16" blockId="10.[855,1454,797,853]" box="[925,1385,830,853]" pageId="10" pageNumber="171">
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(
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<figureCitation id="68882A26C4469C35FC463D7AFC109C16" box="[932,1001,830,853]" captionStart="Fig" captionStartId="2.[136,171,1832,1853]" captionTargetBox="[138,776,989,1810]" captionTargetId="figure-7@2.[137,776,989,1810]" captionTargetPageId="2" captionText="Fig. 1. Distribution map of Indoartemon spp. and Carinartemis spp. Indoartemon spp (white circle): (1) Indoartemon cingalensis (Benson, 1853); (2) Indoartemon layardianus (Benson, 1853); (3) Indoartemon glacilis (Collett, 1898); (4) Indoartemon eburneus (Pfeiffer, 1861); (5) Indoartemon fuchsianus (Gredler, 1881); (6) Indoartemon bidens (Möllendorff, 1883); (7) Indoartemon tridens (Möllendorff, 1898); (8) Indoartemon laevis (Blanford, 1899); (9) Indoartemon prestoni (Gude, 1903); and (10) Indoartemon medius, new species. Carinartemis spp. (black circle): (1) Carinartemis vesperus, new species; and (2) Carinartemis striatus, new species." figureDoi="http://doi.org/10.5281/zenodo.4504077" httpUri="https://zenodo.org/record/4504077/files/figure.png" pageId="10" pageNumber="171">Figs 1</figureCitation>
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,
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<figureCitation id="68882A26C4469C35FC1A3D7AFBE09C16" box="[1016,1049,830,853]" captionStart="Fig" captionStartId="2.[836,872,1975,1996]" captionTargetBox="[836,1473,561,1952]" captionTargetId="figure-361@2.[836,1473,561,1952]" captionTargetPageId="2" captionText="Fig. 2. Living snails of: A, Indoartemon medius, new species, paratype CUMZ 5017 (shell width about 8 mm); B, Carinartemis vesperus, new species, paratype CUMZ 6201 (shell width about 10 mm); C, Carinartemis striatus, new species, paratype CUMZ 6205 (shell width about 12 mm)." figureDoi="http://doi.org/10.5281/zenodo.4504079" httpUri="https://zenodo.org/record/4504079/files/figure.png" pageId="10" pageNumber="171">2C</figureCitation>
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,
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<figureCitation id="68882A26C4469C35FBC43D7AFB9D9C16" box="[1062,1124,830,853]" captionStart="Fig" captionStartId="7.[113,148,2005,2026]" captionTargetBox="[113,1453,825,1982]" captionTargetId="figure-13@7.[113,1453,825,1982]" captionTargetPageId="7" captionText="Fig. 6. A, Shell of Stemmatopsis poirieri lectotype MNHN 26980; B–E, Shells of Carinartemis vesperus, new species, (B) holotype CUMZ 5019 with apertural dentition, (C, D) paratype CUMZ 6200 from Khao Kling, Khao Yoi District, Phetchaburi, and (E) specimen NMW 1955.158.25233 from Khao Yoi, Petchaburi; F, G, Carinartemis striatus, new species, (F) holotype CUMZ 6203 with apertural dentition, and (G) paratype CUMZ 6117 from Tam Kra Sae, Sai Yok, Kanchanaburi." figureDoi="http://doi.org/10.5281/zenodo.4504087" httpUri="https://zenodo.org/record/4504087/files/figure.png" pageId="10" pageNumber="171">6F, G</figureCitation>
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,
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<figureCitation id="68882A26C4469C35FB923D7AFB4A9C16" box="[1136,1203,830,853]" captionStart="Fig" captionStartId="8.[136,171,2035,2056]" captionTargetBox="[146,1462,502,2007]" captionTargetId="figure-66@8.[131,1478,489,2021]" captionTargetPageId="8" captionText="Fig. 7. Genitalia of Carinartemis spp: A, B, Carinartemis vesperus, new species, paratype CUMZ 6201, (A) reproductive system, and (B) insertion of vas deferens into penial sheath; C, D, Carinartemis striatus, new species, paratype CUMZ 6205, (C) reproductive system, and (D) attachment of vas deferens into distal end of the penial sheath." figureDoi="http://doi.org/10.5281/zenodo.4504089" httpUri="https://zenodo.org/record/4504089/files/figure.png" pageId="10" pageNumber="171">7C, D</figureCitation>
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,
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<figureCitation id="68882A26C4469C35FB223D7AFB359C16" box="[1216,1228,830,853]" captionStart="Fig" captionStartId="9.[114,149,1977,1998]" captionTargetBox="[114,1451,379,1954]" captionTargetId="figure-54@9.[108,1458,374,1960]" captionTargetPageId="9" captionText="Fig. 8. Internal sculpture of genitalia of Carinartemis spp: A–F, Carinartemis vesperus, new species, paratype CUMZ 6201, (A) details of atrial pore on the atrium surface, (B) high magnification of penial hooks, (C) top view of penial hook, (D) arrangement of vaginal folds with hooks, (E) high magnification of vaginal hook, and (F) radula morphology. G–K, Carinartemis striatus, new species, paratype CUMZ 6205, (G) details of atrium surface, (H) low magnification shows arrangement of penial hooks, (I) low magnification shows arrangement of vaginal folds, (J) high magnification of penial hooks, and (K) radula morphology." figureDoi="http://doi.org/10.5281/zenodo.4504091" httpUri="https://zenodo.org/record/4504091/files/figure.png" pageId="10" pageNumber="171">8</figureCitation>
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G–K,
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||
<tableCitation id="BD310318C4469C35FAED3D7AFAA79C16" box="[1295,1374,830,853]" captionStart="Table 1" captionStartId="4.[136,189,191,212]" captionText="Table 1. Shell measurements of the three Indoartemon species and two Carinatemis species. Specimen collections and catalogue numbers" pageId="10" pageNumber="171">Table 1</tableCitation>
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)
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="B8A96528C4469C35FCA13D39FB709B3B" pageId="10" pageNumber="171" type="materials_examined">
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<paragraph id="F00C36A3C4469C35FCA13D39FAF09B46" blockId="10.[835,1475,893,1029]" pageId="10" pageNumber="171">
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<emphasis id="C2C7EAB1C4469C35FCA13D39FC2A9CD1" bold="true" box="[835,979,893,914]" pageId="10" pageNumber="171">Type material.</emphasis>
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<materialsCitation id="40DB3CFEC4469C35FC3A3D39FAF19CED" collectionCode="CUMZ" pageId="10" pageNumber="171" specimenCode="CUMZ 6203" specimenCount="1" typeStatus="holotype">
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<typeStatus id="2F088801C4469C35FC3A3D39FBD69CD1" box="[984,1071,893,914]" pageId="10" pageNumber="171" type="holotype">Holotype</typeStatus>
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<specimenCode id="A0159ED8C4469C35FBD13D39FB529CD1" box="[1075,1195,893,914]" collectionCode="CUMZ" country="0" httpUri="http://grbio.org/cool/da58-p0xp" name="Cameroon University, Museum of Zoology" pageId="10" pageNumber="171">CUMZ 6203</specimenCode>
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(
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<figureCitation id="68882A26C4469C35FB543D39FB059CD1" box="[1206,1276,893,914]" captionStart="Fig" captionStartId="7.[113,148,2005,2026]" captionTargetBox="[113,1453,825,1982]" captionTargetId="figure-13@7.[113,1453,825,1982]" captionTargetPageId="7" captionText="Fig. 6. A, Shell of Stemmatopsis poirieri lectotype MNHN 26980; B–E, Shells of Carinartemis vesperus, new species, (B) holotype CUMZ 5019 with apertural dentition, (C, D) paratype CUMZ 6200 from Khao Kling, Khao Yoi District, Phetchaburi, and (E) specimen NMW 1955.158.25233 from Khao Yoi, Petchaburi; F, G, Carinartemis striatus, new species, (F) holotype CUMZ 6203 with apertural dentition, and (G) paratype CUMZ 6117 from Tam Kra Sae, Sai Yok, Kanchanaburi." figureDoi="http://doi.org/10.5281/zenodo.4504087" httpUri="https://zenodo.org/record/4504087/files/figure.png" pageId="10" pageNumber="171">Fig. 6F</figureCitation>
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). Measurement: shell height
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||
<quantity id="374B9B46C4469C35FC643DDDFC2A9CED" box="[902,979,921,942]" metricMagnitude="-3" metricUnit="m" metricValue="9.2" pageId="10" pageNumber="171" unit="mm" value="9.2">9.2 mm</quantity>
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, shell width
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<quantity id="374B9B46C4469C35FBAD3DDDFB5E9CED" box="[1103,1191,921,942]" metricMagnitude="-2" metricUnit="m" metricValue="1.14" pageId="10" pageNumber="171" unit="mm" value="11.4">11.4 mm</quantity>
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, 7 whorls
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||
</materialsCitation>
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.
|
||
<materialsCitation id="40DB3CFEC4469C35FAF03DDDFBC99C88" collectionCode="CUMZ" pageId="10" pageNumber="171" specimenCode="CUMZ 6117" specimenCount="1" typeStatus="paratype">
|
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<typeStatus id="2F088801C4469C35FAF03DDDFA8A9CED" box="[1298,1395,921,942]" pageId="10" pageNumber="171" type="paratype">Paratypes</typeStatus>
|
||
:
|
||
<specimenCode id="A0159ED8C4469C35FA9F3DDDFC8B9C88" collectionCode="CUMZ" country="0" httpUri="http://grbio.org/cool/da58-p0xp" name="Cameroon University, Museum of Zoology" pageId="10" pageNumber="171">CUMZ 6117</specimenCode>
|
||
(5 shells;
|
||
<figureCitation id="68882A26C4469C35FC393DF2FBD19C88" box="[987,1064,950,971]" captionStart="Fig" captionStartId="7.[113,148,2005,2026]" captionTargetBox="[113,1453,825,1982]" captionTargetId="figure-13@7.[113,1453,825,1982]" captionTargetPageId="7" captionText="Fig. 6. A, Shell of Stemmatopsis poirieri lectotype MNHN 26980; B–E, Shells of Carinartemis vesperus, new species, (B) holotype CUMZ 5019 with apertural dentition, (C, D) paratype CUMZ 6200 from Khao Kling, Khao Yoi District, Phetchaburi, and (E) specimen NMW 1955.158.25233 from Khao Yoi, Petchaburi; F, G, Carinartemis striatus, new species, (F) holotype CUMZ 6203 with apertural dentition, and (G) paratype CUMZ 6117 from Tam Kra Sae, Sai Yok, Kanchanaburi." figureDoi="http://doi.org/10.5281/zenodo.4504087" httpUri="https://zenodo.org/record/4504087/files/figure.png" pageId="10" pageNumber="171">Fig. 6G</figureCitation>
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)
|
||
</materialsCitation>
|
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,
|
||
<materialsCitation id="40DB3CFEC4469C35FBDE3DF2FAE79C88" box="[1084,1310,950,971]" collectionCode="CUMZ" pageId="10" pageNumber="171" specimenCode="CUMZ 6204" specimenCount="1" typeStatus="paratype">
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<specimenCode id="A0159ED8C4469C35FBDE3DF2FB409C88" box="[1084,1209,950,971]" collectionCode="CUMZ" country="0" httpUri="http://grbio.org/cool/da58-p0xp" name="Cameroon University, Museum of Zoology" pageId="10" pageNumber="171">CUMZ 6204</specimenCode>
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(3 shells)
|
||
</materialsCitation>
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,
|
||
<materialsCitation id="40DB3CFEC4469C35FACB3DF2FB0C9CAB" collectionCode="CUMZ" pageId="10" pageNumber="171" specimenCode="CUMZ 6205" specimenCount="1" typeStatus="paratype">
|
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<specimenCode id="A0159ED8C4469C35FACB3DF2FA5F9C88" box="[1321,1446,950,971]" collectionCode="CUMZ" country="0" httpUri="http://grbio.org/cool/da58-p0xp" name="Cameroon University, Museum of Zoology" pageId="10" pageNumber="171">CUMZ 6205</specimenCode>
|
||
(
|
||
<specimenCount id="E6B5FD2AC4469C35FA553DF2FC679CAB" pageId="10" pageNumber="171" type="generic">1 specimen</specimenCount>
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in ethanol;
|
||
<figureCitation id="68882A26C4469C35FBF73D97FB9C9CAB" box="[1045,1125,979,1000]" captionStart="Fig" captionStartId="2.[836,872,1975,1996]" captionTargetBox="[836,1473,561,1952]" captionTargetId="figure-361@2.[836,1473,561,1952]" captionTargetPageId="2" captionText="Fig. 2. Living snails of: A, Indoartemon medius, new species, paratype CUMZ 5017 (shell width about 8 mm); B, Carinartemis vesperus, new species, paratype CUMZ 6201 (shell width about 10 mm); C, Carinartemis striatus, new species, paratype CUMZ 6205 (shell width about 12 mm)." figureDoi="http://doi.org/10.5281/zenodo.4504079" httpUri="https://zenodo.org/record/4504079/files/figure.png" pageId="10" pageNumber="171">Figs 2B</figureCitation>
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||
,
|
||
<figureCitation id="68882A26C4469C35FB923D97FB549CAB" box="[1136,1197,979,1000]" captionStart="Fig" captionStartId="8.[136,171,2035,2056]" captionTargetBox="[146,1462,502,2007]" captionTargetId="figure-66@8.[131,1478,489,2021]" captionTargetPageId="8" captionText="Fig. 7. Genitalia of Carinartemis spp: A, B, Carinartemis vesperus, new species, paratype CUMZ 6201, (A) reproductive system, and (B) insertion of vas deferens into penial sheath; C, D, Carinartemis striatus, new species, paratype CUMZ 6205, (C) reproductive system, and (D) attachment of vas deferens into distal end of the penial sheath." figureDoi="http://doi.org/10.5281/zenodo.4504089" httpUri="https://zenodo.org/record/4504089/files/figure.png" pageId="10" pageNumber="171">7A, B</figureCitation>
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,
|
||
<figureCitation id="68882A26C4469C35FB5B3D97FB3D9CAB" box="[1209,1220,979,1000]" captionStart="Fig" captionStartId="9.[114,149,1977,1998]" captionTargetBox="[114,1451,379,1954]" captionTargetId="figure-54@9.[108,1458,374,1960]" captionTargetPageId="9" captionText="Fig. 8. Internal sculpture of genitalia of Carinartemis spp: A–F, Carinartemis vesperus, new species, paratype CUMZ 6201, (A) details of atrial pore on the atrium surface, (B) high magnification of penial hooks, (C) top view of penial hook, (D) arrangement of vaginal folds with hooks, (E) high magnification of vaginal hook, and (F) radula morphology. G–K, Carinartemis striatus, new species, paratype CUMZ 6205, (G) details of atrium surface, (H) low magnification shows arrangement of penial hooks, (I) low magnification shows arrangement of vaginal folds, (J) high magnification of penial hooks, and (K) radula morphology." figureDoi="http://doi.org/10.5281/zenodo.4504091" httpUri="https://zenodo.org/record/4504091/files/figure.png" pageId="10" pageNumber="171">8</figureCitation>
|
||
G–I)
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="40DB3CFEC4469C35FAE23D97FC669B46" collectionCode="NHMUK" pageId="10" pageNumber="171" specimenCode="NHMUK 20130611" specimenCount="1" typeStatus="paratype">
|
||
<specimenCode id="A0159ED8C4469C35FAE23D97FA3B9CAB" box="[1280,1474,979,1000]" collectionCode="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="10" pageNumber="171" type="Museum">NHMUK 20130611</specimenCode>
|
||
(2 shells)
|
||
</materialsCitation>
|
||
,
|
||
<collectionCode id="96A2AE66C4469C35FC483DB4FC239B46" box="[938,986,1008,1029]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="10" pageNumber="171">SMF</collectionCode>
|
||
(2 shells), and
|
||
<collectionCode id="96A2AE66C4469C35FB903DB4FB589B46" box="[1138,1185,1008,1029]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="10" pageNumber="171">ZRC</collectionCode>
|
||
(2 shells).
|
||
</paragraph>
|
||
<paragraph id="F00C36A3C4469C35FCA13A6DFB709B3B" blockId="10.[835,1474,1065,1144]" pageId="10" pageNumber="171">
|
||
<materialsCitation id="40DB3CFEC4469C35FCA13A6DFB7D9B3B" country="Thailand" latitude="14.106528" location="Tam Kra Sae" longLatPrecision="1" longitude="99.166115" municipality="Sai Yok" pageId="10" pageNumber="171" specimenCount="1" stateProvince="Kanchanaburi" typeStatus="holotype">
|
||
<emphasis id="C2C7EAB1C4469C35FCA13A6DFC359B7D" bold="true" box="[835,972,1065,1086]" pageId="10" pageNumber="171">
|
||
<typeStatus id="2F088801C4469C35FCA13A6DFC8C9B7D" box="[835,885,1065,1086]" pageId="10" pageNumber="171">Type</typeStatus>
|
||
locality.
|
||
</emphasis>
|
||
<location id="F56C6078C4469C35FC363A6DFBAE9B7D" LSID="urn:lsid:plazi:treatment:781A87B5C4469C34FCB53D59FDB29857:F56C6078C4469C35FC363A6DFBAE9B7D" box="[980,1111,1065,1086]" country="Thailand" latitude="14.106528" longLatPrecision="1" longitude="99.166115" municipality="Sai Yok" name="Tam Kra Sae" pageId="10" pageNumber="171" stateProvince="Kanchanaburi">Tam Kra Sae</location>
|
||
,
|
||
<collectingMunicipality id="1068ACD9C4469C35FB803A6DFB4B9B7D" box="[1122,1202,1065,1086]" pageId="10" pageNumber="171">Sai Yok</collectingMunicipality>
|
||
,
|
||
<collectingRegion id="3277F841C4469C35FB5E3A6DFABC9B7D" box="[1212,1349,1065,1086]" country="Thailand" name="Kanchanaburi" pageId="10" pageNumber="171">Kanchanaburi</collectingRegion>
|
||
,
|
||
<collectingCountry id="88A47633C4469C35FAB23A6DFA519B7D" box="[1360,1448,1065,1086]" name="Thailand" pageId="10" pageNumber="171">Thailand</collectingCountry>
|
||
. A limestone mountain reaching about 200 metres above mean sea level (
|
||
<geoCoordinate id="95875064C4469C35FC623A27FC029B3B" box="[896,1019,1123,1144]" degrees="14" direction="north" minutes="6" orientation="latitude" pageId="10" pageNumber="171" precision="1" seconds="23.5" value="14.106528">14°6'23.5"N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="95875064C4469C35FBE73A27FB849B3B" box="[1029,1149,1123,1144]" degrees="99" direction="east" minutes="9" orientation="longitude" pageId="10" pageNumber="171" precision="1" seconds="58.0" value="99.166115">99°9'58.0"E</geoCoordinate>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B8A96528C4469C35FCA13AE5FB199BBB" pageId="10" pageNumber="171" type="etymology">
|
||
<paragraph id="F00C36A3C4469C35FCA13AE5FB199BBB" blockId="10.[835,1474,1185,1272]" pageId="10" pageNumber="171">
|
||
<emphasis id="C2C7EAB1C4469C35FCA13AE5FC319BFB" bold="true" box="[835,968,1185,1208]" pageId="10" pageNumber="171">Etymology.</emphasis>
|
||
The specific epithet from the Latin
|
||
<emphasis id="C2C7EAB1C4469C35FA923AE5FA3B9BFB" box="[1392,1474,1185,1208]" italics="true" pageId="10" pageNumber="171">“stria”</emphasis>
|
||
meaning “furrow”, in reference to the strong transverse ridges or radial striation on the shell surface.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B8A96528C4469C35FCA13B65FAC2981B" pageId="10" pageNumber="171" type="diagnosis">
|
||
<paragraph id="F00C36A3C4469C35FCA13B65FB239ABB" blockId="10.[835,1474,1313,1528]" pageId="10" pageNumber="171">
|
||
<emphasis id="C2C7EAB1C4469C35FCA13B65FC469A7B" bold="true" box="[835,959,1313,1336]" pageId="10" pageNumber="171">Diagnosis.</emphasis>
|
||
<taxonomicName id="37B34D20C4469C35FC293B65FB3D9A7B" authority="Siriboon & Sutcharit & Naggs & Rowson & Panha, 2014" authorityName="Siriboon & Sutcharit & Naggs & Rowson & Panha" authorityYear="2014" box="[971,1220,1313,1336]" class="Gastropoda" family="Streptaxidae" genus="Carinartemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="171" phylum="Mollusca" rank="species" species="striatus" status="sp. nov.">
|
||
<emphasis id="C2C7EAB1C4469C35FC293B65FB3D9A7B" box="[971,1220,1313,1336]" italics="true" pageId="10" pageNumber="171">Carinartemis striatus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="D9F457CAC4469C35FB353B65FA9F9A7B" box="[1239,1382,1313,1336]" pageId="10" pageNumber="171" rank="species">new species</taxonomicNameLabel>
|
||
, can be distinguished from
|
||
<taxonomicName id="37B34D20C4469C35FBFC3B05FB589A1B" baseAuthorityName="Mollendorff" baseAuthorityYear="1894" box="[1054,1185,1345,1368]" class="Gastropoda" family="Streptaxidae" genus="Haploptychius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="171" phylum="Mollusca" rank="species" species="mirificus">
|
||
<emphasis id="C2C7EAB1C4469C35FBFC3B05FB589A1B" box="[1054,1185,1345,1368]" italics="true" pageId="10" pageNumber="171">H. mirificus</emphasis>
|
||
</taxonomicName>
|
||
in its larger shell, higher spire, strong transverse ridges over the entire shell, subcircular aperture, broadly expanded peristome, and small parietal lamella. Whereas
|
||
<taxonomicName id="37B34D20C4469C35FBED3BE5FB6A9AFB" baseAuthorityName="Mollendorff" baseAuthorityYear="1894" box="[1039,1171,1441,1464]" class="Gastropoda" family="Streptaxidae" genus="Haploptychius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="171" phylum="Mollusca" rank="species" species="mirificus">
|
||
<emphasis id="C2C7EAB1C4469C35FBED3BE5FB6A9AFB" box="[1039,1171,1441,1464]" italics="true" pageId="10" pageNumber="171">H. mirificus</emphasis>
|
||
</taxonomicName>
|
||
has a smaller shell, lower spire, nearly smooth shell surface, triangular aperture and a strong and triangular parietal lamella.
|
||
</paragraph>
|
||
<paragraph id="F00C36A3C4469C35FCA13865FAC2981B" blockId="10.[835,1475,1569,1880]" pageId="10" pageNumber="171">
|
||
Compared with
|
||
<taxonomicName id="37B34D20C4469C35FC0F3865FB90997B" baseAuthorityName="Gould" baseAuthorityYear="1856" box="[1005,1129,1569,1592]" class="Gastropoda" family="Streptaxidae" genus="Haploptychius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="171" phylum="Mollusca" rank="species" species="exacutus">
|
||
<emphasis id="C2C7EAB1C4469C35FC0F3865FB90997B" box="[1005,1129,1569,1592]" italics="true" pageId="10" pageNumber="171">H. exacutus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="37B34D20C4469C35FB793865FAF2997B" baseAuthorityName="Benson" baseAuthorityYear="1859" box="[1179,1291,1569,1592]" class="Gastropoda" family="Streptaxidae" genus="Haploptychius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="171" phylum="Mollusca" rank="species" species="sankeyi">
|
||
<emphasis id="C2C7EAB1C4469C35FB793865FAF2997B" box="[1179,1291,1569,1592]" italics="true" pageId="10" pageNumber="171">H. sankeyi</emphasis>
|
||
</taxonomicName>
|
||
, this new species differs by having a strong peripheral keel around nearly the entire penultimate whorl, the last whorl less being inflated and much deviated from the vertical axis, and with strong transverse ridges over the entire shell. In comparison, these two species have a strong peripheral keel over less than half of penultimate whorl, and have the last whorl inflated and less deviated from vertical axis. The transverse ridges of
|
||
<taxonomicName id="37B34D20C4469C35FA4A3945FC59987B" baseAuthorityName="Gould" baseAuthorityYear="1856" class="Gastropoda" family="Streptaxidae" genus="Haploptychius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="171" phylum="Mollusca" rank="species" species="exacutus">
|
||
<emphasis id="C2C7EAB1C4469C35FA4A3945FC59987B" italics="true" pageId="10" pageNumber="171">H. exacutus</emphasis>
|
||
</taxonomicName>
|
||
disappear below the periphery of the penultimate whorl and it has two distinct parietal lamellae.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B8A96528C4469C34FCA139C5FE9F9917" lastPageId="11" lastPageNumber="172" pageId="10" pageNumber="171" type="description">
|
||
<paragraph id="F00C36A3C4469C34FCA139C5FDC39E17" blockId="10.[835,1475,1921,2072]" lastBlockId="11.[113,753,189,340]" lastPageId="11" lastPageNumber="172" pageId="10" pageNumber="171">
|
||
<emphasis id="C2C7EAB1C4469C35FCA139C5FC2C98DB" bold="true" box="[835,981,1921,1944]" pageId="10" pageNumber="171">Description.</emphasis>
|
||
Shell oblique-heliciform, white, dull, and translucent; whorls 7, spire low conical with distinct suture. Shell surface glossy, with strong transverse ridges and varices present. Embryonic shell with about 2½ whorls and smooth surface; following whorls regularly expanding. Shell periphery wide and sharply keeled around nearly the entire penultimate whorl; last whorl axially deflected. Umbilicus open and shallow. Aperture semi-ovate, peristome discontinuous, thickened, expanded and reflected. Apertural dentition of one parietal lamella (
|
||
<figureCitation id="68882A26C4479C34FE393F79FDD79E17" box="[475,558,317,340]" captionStart="Fig" captionStartId="7.[113,148,2005,2026]" captionTargetBox="[113,1453,825,1982]" captionTargetId="figure-13@7.[113,1453,825,1982]" captionTargetPageId="7" captionText="Fig. 6. A, Shell of Stemmatopsis poirieri lectotype MNHN 26980; B–E, Shells of Carinartemis vesperus, new species, (B) holotype CUMZ 5019 with apertural dentition, (C, D) paratype CUMZ 6200 from Khao Kling, Khao Yoi District, Phetchaburi, and (E) specimen NMW 1955.158.25233 from Khao Yoi, Petchaburi; F, G, Carinartemis striatus, new species, (F) holotype CUMZ 6203 with apertural dentition, and (G) paratype CUMZ 6117 from Tam Kra Sae, Sai Yok, Kanchanaburi." figureDoi="http://doi.org/10.5281/zenodo.4504087" httpUri="https://zenodo.org/record/4504087/files/figure.png" pageId="11" pageNumber="172">Fig. 6F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="F00C36A3C4479C34FF933F39FF249D17" blockId="11.[113,753,381,596]" pageId="11" pageNumber="172">
|
||
<emphasis id="C2C7EAB1C4479C34FF933F39FF329ED7" bold="true" box="[113,203,381,404]" pageId="11" pageNumber="172">Radula.</emphasis>
|
||
Teeth arranged in anteriorly V-shaped rows, each row containing 45–49 teeth with the formula (22-24)-1-(22- 24); central tooth small, short, triangular with pointed cusp. Lateral and marginal teeth undifferentiated, unicuspid and lanceolate; lateral teeth gradually reduced in length and size with outer teeth much smaller and shorter than inner teeth (
|
||
<figureCitation id="68882A26C4479C34FF9B3C79FF299D17" box="[121,208,573,596]" captionStart="Fig" captionStartId="9.[114,149,1977,1998]" captionTargetBox="[114,1451,379,1954]" captionTargetId="figure-54@9.[108,1458,374,1960]" captionTargetPageId="9" captionText="Fig. 8. Internal sculpture of genitalia of Carinartemis spp: A–F, Carinartemis vesperus, new species, paratype CUMZ 6201, (A) details of atrial pore on the atrium surface, (B) high magnification of penial hooks, (C) top view of penial hook, (D) arrangement of vaginal folds with hooks, (E) high magnification of vaginal hook, and (F) radula morphology. G–K, Carinartemis striatus, new species, paratype CUMZ 6205, (G) details of atrium surface, (H) low magnification shows arrangement of penial hooks, (I) low magnification shows arrangement of vaginal folds, (J) high magnification of penial hooks, and (K) radula morphology." figureDoi="http://doi.org/10.5281/zenodo.4504091" httpUri="https://zenodo.org/record/4504091/files/figure.png" pageId="11" pageNumber="172">Fig. 8K</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="F00C36A3C4479C34FF933C39FEA79CF7" blockId="11.[113,753,637,948]" pageId="11" pageNumber="172">
|
||
<emphasis id="C2C7EAB1C4479C34FF933C39FEDA9DD7" bold="true" box="[113,291,637,660]" pageId="11" pageNumber="172">Genital organs.</emphasis>
|
||
Atrium (at) short. Penis (p) long, slender, becoming slightly broad in middle and then tapering distally. Penial sheath (ps) thin, extending about quarter of total penis length; penial sheath retractor muscle (psr) very thin, originating at atrium and body wall, inserting distally on penial sheath (
|
||
<figureCitation id="68882A26C4479C34FEFB3D59FE8D9C77" box="[281,372,797,820]" captionStart="Fig" captionStartId="8.[136,171,2035,2056]" captionTargetBox="[146,1462,502,2007]" captionTargetId="figure-66@8.[131,1478,489,2021]" captionTargetPageId="8" captionText="Fig. 7. Genitalia of Carinartemis spp: A, B, Carinartemis vesperus, new species, paratype CUMZ 6201, (A) reproductive system, and (B) insertion of vas deferens into penial sheath; C, D, Carinartemis striatus, new species, paratype CUMZ 6205, (C) reproductive system, and (D) attachment of vas deferens into distal end of the penial sheath." figureDoi="http://doi.org/10.5281/zenodo.4504089" httpUri="https://zenodo.org/record/4504089/files/figure.png" pageId="11" pageNumber="172">Fig. 7C</figureCitation>
|
||
). Vas deferens (vd) slender, very long, attached to distal end of penial sheath with very thin connective tissue (
|
||
<figureCitation id="68882A26C4479C34FED43D19FE729C37" box="[310,395,861,884]" captionStart="Fig" captionStartId="8.[136,171,2035,2056]" captionTargetBox="[146,1462,502,2007]" captionTargetId="figure-66@8.[131,1478,489,2021]" captionTargetPageId="8" captionText="Fig. 7. Genitalia of Carinartemis spp: A, B, Carinartemis vesperus, new species, paratype CUMZ 6201, (A) reproductive system, and (B) insertion of vas deferens into penial sheath; C, D, Carinartemis striatus, new species, paratype CUMZ 6205, (C) reproductive system, and (D) attachment of vas deferens into distal end of the penial sheath." figureDoi="http://doi.org/10.5281/zenodo.4504089" httpUri="https://zenodo.org/record/4504089/files/figure.png" pageId="11" pageNumber="172">Fig. 7D</figureCitation>
|
||
). Penial retractor muscle (pr) thin and very long, inserting distally on penis at the penis and vas deferens junction.
|
||
</paragraph>
|
||
<paragraph id="F00C36A3C4479C34FF933D99FE169BD7" blockId="11.[113,752,989,1172]" pageId="11" pageNumber="172">
|
||
Internal wall of atrium generally corrugated with sparse atrial pores (
|
||
<figureCitation id="68882A26C4479C34FF1C3DB9FEAE9B57" box="[254,343,1021,1044]" captionStart="Fig" captionStartId="9.[114,149,1977,1998]" captionTargetBox="[114,1451,379,1954]" captionTargetId="figure-54@9.[108,1458,374,1960]" captionTargetPageId="9" captionText="Fig. 8. Internal sculpture of genitalia of Carinartemis spp: A–F, Carinartemis vesperus, new species, paratype CUMZ 6201, (A) details of atrial pore on the atrium surface, (B) high magnification of penial hooks, (C) top view of penial hook, (D) arrangement of vaginal folds with hooks, (E) high magnification of vaginal hook, and (F) radula morphology. G–K, Carinartemis striatus, new species, paratype CUMZ 6205, (G) details of atrium surface, (H) low magnification shows arrangement of penial hooks, (I) low magnification shows arrangement of vaginal folds, (J) high magnification of penial hooks, and (K) radula morphology." figureDoi="http://doi.org/10.5281/zenodo.4504091" httpUri="https://zenodo.org/record/4504091/files/figure.png" pageId="11" pageNumber="172">Fig. 8G</figureCitation>
|
||
). Penial wall with pale brown penial hooks, about 1 hook/200 μm
|
||
<superScript id="07C69BEBC4479C34FE513A58FE429B69" attach="left" box="[435,443,1052,1066]" fontSize="6" pageId="11" pageNumber="172">2</superScript>
|
||
; hooks located on irregular trapezoidal papillae (pl). Penial hooks short (<
|
||
<quantity id="374B9B46C4479C34FD913A79FD2A9B17" box="[627,723,1085,1108]" metricMagnitude="-5" metricUnit="m" metricValue="4.0" pageId="11" pageNumber="172" unit="mm" value="0.04">0.04 mm</quantity>
|
||
in length), slightly expanding at base, tips sharp and curved towards genital orifice (
|
||
<figureCitation id="68882A26C4479C34FE903A39FE1B9BD7" box="[370,482,1149,1172]" captionStart="Fig" captionStartId="9.[114,149,1977,1998]" captionTargetBox="[114,1451,379,1954]" captionTargetId="figure-54@9.[108,1458,374,1960]" captionTargetPageId="9" captionText="Fig. 8. Internal sculpture of genitalia of Carinartemis spp: A–F, Carinartemis vesperus, new species, paratype CUMZ 6201, (A) details of atrial pore on the atrium surface, (B) high magnification of penial hooks, (C) top view of penial hook, (D) arrangement of vaginal folds with hooks, (E) high magnification of vaginal hook, and (F) radula morphology. G–K, Carinartemis striatus, new species, paratype CUMZ 6205, (G) details of atrium surface, (H) low magnification shows arrangement of penial hooks, (I) low magnification shows arrangement of vaginal folds, (J) high magnification of penial hooks, and (K) radula morphology." figureDoi="http://doi.org/10.5281/zenodo.4504091" httpUri="https://zenodo.org/record/4504091/files/figure.png" pageId="11" pageNumber="172">Fig. 8H, J</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="F00C36A3C4479C34FF933AF9FD8C9A97" blockId="11.[113,753,1213,1492]" pageId="11" pageNumber="172">
|
||
Vagina (v) short and stout. Gametolytic duct (gd) long, extending as far as albumin gland; gametolytic sac (gs) ovate. Free oviduct (fo) long, proximally large with equivalent diameter with vagina, and tapering to a smaller tube distally. Oviduct (ov) enlarged and folded; prostate gland inconspicuous and bound to oviduct. Talon (ta) small, very short and club shaped. Hermaphroditic duct (hd) bearing seminal vesicle (sv) about as long as the length from talon to branching point of seminal vesicle (
|
||
<figureCitation id="68882A26C4479C34FDF13BF9FD919A97" box="[531,616,1469,1492]" captionStart="Fig" captionStartId="8.[136,171,2035,2056]" captionTargetBox="[146,1462,502,2007]" captionTargetId="figure-66@8.[131,1478,489,2021]" captionTargetPageId="8" captionText="Fig. 7. Genitalia of Carinartemis spp: A, B, Carinartemis vesperus, new species, paratype CUMZ 6201, (A) reproductive system, and (B) insertion of vas deferens into penial sheath; C, D, Carinartemis striatus, new species, paratype CUMZ 6205, (C) reproductive system, and (D) attachment of vas deferens into distal end of the penial sheath." figureDoi="http://doi.org/10.5281/zenodo.4504089" httpUri="https://zenodo.org/record/4504089/files/figure.png" pageId="11" pageNumber="172">Fig. 7C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="F00C36A3C4479C34FF933BB9FE9F9917" blockId="11.[113,752,1533,1620]" pageId="11" pageNumber="172">
|
||
Proximal vaginal wall with undulated parallel vaginal folds, arched distally; transverse parallel cavity present; vaginal hooks absent (
|
||
<figureCitation id="68882A26C4479C34FEEF3879FEA09917" box="[269,345,1597,1620]" captionStart="Fig" captionStartId="9.[114,149,1977,1998]" captionTargetBox="[114,1451,379,1954]" captionTargetId="figure-54@9.[108,1458,374,1960]" captionTargetPageId="9" captionText="Fig. 8. Internal sculpture of genitalia of Carinartemis spp: A–F, Carinartemis vesperus, new species, paratype CUMZ 6201, (A) details of atrial pore on the atrium surface, (B) high magnification of penial hooks, (C) top view of penial hook, (D) arrangement of vaginal folds with hooks, (E) high magnification of vaginal hook, and (F) radula morphology. G–K, Carinartemis striatus, new species, paratype CUMZ 6205, (G) details of atrium surface, (H) low magnification shows arrangement of penial hooks, (I) low magnification shows arrangement of vaginal folds, (J) high magnification of penial hooks, and (K) radula morphology." figureDoi="http://doi.org/10.5281/zenodo.4504091" httpUri="https://zenodo.org/record/4504091/files/figure.png" pageId="11" pageNumber="172">Fig. 8I</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B8A96528C4479C34FF933839FDE799F7" pageId="11" pageNumber="172" type="distribution">
|
||
<paragraph id="F00C36A3C4479C34FF933839FDE799F7" blockId="11.[113,752,1661,1716]" pageId="11" pageNumber="172">
|
||
<emphasis id="C2C7EAB1C4479C34FF933839FEF899D7" bold="true" box="[113,257,1661,1684]" pageId="11" pageNumber="172">Distribution.</emphasis>
|
||
This new species was found from the limestone karst near the very famous historic site.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B8A96528C4479C34FF933899FDB29857" pageId="11" pageNumber="172" type="discussion">
|
||
<paragraph id="F00C36A3C4479C34FF933899FDB29857" blockId="11.[113,751,1757,1812]" pageId="11" pageNumber="172">
|
||
<emphasis id="C2C7EAB1C4479C34FF933899FF1C99B7" bold="true" box="[113,229,1757,1780]" pageId="11" pageNumber="172">Remarks.</emphasis>
|
||
Apparently rare. Recent extensive searching revealed the living animal for the first time.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |