treatments-xml/data/44/38/87/443887FA661CFFADFF6FFD72FBB84A16.xml

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<mods:title id="4871E1273FEE57BCEE8A75E94CE88993">Phylogenetic relationships of Tectoribates: nymphal characters of new North American species place the genus in Tegoribatidae (Acari, Oribatida)</mods:title>
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<mods:namePart id="30CE843052D16EE27E81FB19D7AA103A">Behan-Pelletier, Valerie M.</mods:namePart>
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<mods:namePart id="23339F641C05E0A688E1B2F166AAEB48">Walter, David E.</mods:namePart>
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<mods:date id="05C7CE7A50769164CC5DAB86D2E5191F">2013</mods:date>
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<treatment id="443887FA661CFFADFF6FFD72FBB84A16" ID-DOI="http://doi.org/10.5281/zenodo.5671481" ID-GBIF-Taxon="119583772" ID-Zenodo-Dep="5671481" LSID="urn:lsid:plazi:treatment:443887FA661CFFADFF6FFD72FBB84A16" httpUri="http://treatment.plazi.org/id/443887FA661CFFADFF6FFD72FBB84A16" lastPageId="10" lastPageNumber="469" pageId="2" pageNumber="461">
<subSubSection id="848B6567661CFFA5FF6FFD72FE1849D5" box="[151,461,653,679]" pageId="2" pageNumber="461" type="nomenclature">
<paragraph id="CC2E36EC661CFFA5FF6FFD72FE1849D5" blockId="2.[151,461,653,679]" box="[151,461,653,679]" pageId="2" pageNumber="461">
<heading id="97668180661CFFA5FF6FFD72FE1849D5" bold="true" box="[151,461,653,679]" fontSize="11" level="1" pageId="2" pageNumber="461" reason="1">
<taxonomicName id="0B914D6F661CFFA5FF6FFD72FE1849D5" authority="Berlese 1910" authorityName="Berlese" authorityYear="1910" box="[151,461,653,679]" class="Arachnida" family="Oribatellidae" genus="Tectoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="2" pageNumber="461" phylum="Arthropoda" rank="genus">
<emphasis id="FEE5EAFE661CFFA5FF6FFD72FE1849D5" bold="true" box="[151,461,653,679]" pageId="2" pageNumber="461">
<emphasis id="FEE5EAFE661CFFA5FF6FFD72FEFC49D5" bold="true" box="[151,297,653,679]" italics="true" pageId="2" pageNumber="461">Tectoribates</emphasis>
Berlese 1910
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="848B6567661CFFA5FF6FFD2BFBE548AC" pageId="2" pageNumber="461" type="materials_examined">
<paragraph id="CC2E36EC661CFFA5FF6FFD2BFC514998" blockId="2.[151,1337,724,776]" box="[151,900,724,746]" pageId="2" pageNumber="461">
<emphasis id="FEE5EAFE661CFFA5FF6FFD2BFEF34998" bold="true" box="[151,294,724,746]" pageId="2" pageNumber="461">
<typeStatus id="132A884E661CFFA5FF6FFD2BFF194998" box="[151,204,724,746]" pageId="2" pageNumber="461">Type</typeStatus>
species:
</emphasis>
<taxonomicName id="0B914D6F661CFFA5FED3FD2BFCEE4998" authority="Berlese, 1910" authorityName="Berlese" authorityYear="1910" box="[299,827,724,746]" class="Arachnida" family="Ceratozetidae" genus="Sphaerozetes" kingdom="Animalia" order="Sarcoptiformes" pageId="2" pageNumber="461" phylum="Arthropoda" rank="species" species="proximus" subGenus="Tectoribates">
<emphasis id="FEE5EAFE661CFFA5FED3FD2BFD724998" box="[299,679,724,746]" italics="true" pageId="2" pageNumber="461">Sphaerozetes (Tectoribates) proximus</emphasis>
Berlese, 1910
</taxonomicName>
, p. 264
</paragraph>
<paragraph id="CC2E36EC661CFFA5FF6FFD0DFAEC487A" blockId="2.[151,1337,724,776]" box="[151,1337,754,776]" pageId="2" pageNumber="461">
<emphasis id="FEE5EAFE661CFFA5FF6FFD0DFF29487A" bold="true" box="[151,252,754,776]" pageId="2" pageNumber="461">Synonym</emphasis>
: 
<taxonomicName id="0B914D6F661CFFA5FEF1FD0DFDDB487A" authority="Kunst, 1962" authorityName="Kunst" authorityYear="1962" box="[265,526,754,776]" class="Arachnida" family="Oribatellidae" genus="Anoribatella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="2" pageNumber="461" phylum="Arthropoda" rank="genus">
<emphasis id="FEE5EAFE661CFFA5FEF1FD0DFE5C487A" box="[265,393,754,776]" italics="true" pageId="2" pageNumber="461">Anoribatella</emphasis>
Kunst, 1962
</taxonomicName>
, 
<typeStatus id="132A884E661CFFA5FDE0FD0CFD9F487A" box="[536,586,755,776]" pageId="2" pageNumber="461">Type</typeStatus>
species: 
<taxonomicName id="0B914D6F661CFFA5FD5FFD0DFBC6487A" authority="Schuster, 1958" authorityName="Schuster" authorityYear="1958" box="[679,1043,754,776]" class="Arachnida" family="Achipteriidae" genus="Anachipteria" kingdom="Animalia" order="Sarcoptiformes" pageId="2" pageNumber="461" phylum="Arthropoda" rank="species" species="ornata">
<emphasis id="FEE5EAFE661CFFA5FD5FFD0DFCA3487A" box="[679,886,754,776]" italics="true" pageId="2" pageNumber="461">Anachipteria ornata</emphasis>
Schuster, 1958
</taxonomicName>
, p. 90; Bernini (1973) p. 541
</paragraph>
<paragraph id="CC2E36EC661CFFA5FF6FFCCAFE71483C" blockId="2.[151,1086,821,990]" box="[151,420,821,846]" pageId="2" pageNumber="461">Included extant species:</paragraph>
<paragraph id="CC2E36EC661CFFA5FF6FFCA6FBE548AC" blockId="2.[151,1086,821,990]" pageId="2" pageNumber="461">
<taxonomicName id="0B914D6F661CFFA5FF6FFCA6FD914800" authority="Berlese, 1910" authorityName="Berlese" authorityYear="1910" box="[151,580,857,882]" class="Arachnida" family="Oribatellidae" genus="Tectoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="2" pageNumber="461" phylum="Arthropoda" rank="species" species="proximus">
<emphasis id="FEE5EAFE661CFFA5FF6FFCA6FE5B4803" box="[151,398,857,882]" italics="true" pageId="2" pageNumber="461">Tectoribates proximus</emphasis>
(Berlese, 1910)
</taxonomicName>
, known from the Palaearctic and Neotropics; 
<taxonomicName id="0B914D6F661CFFA5FF6FFC82FD3148E4" authority="Balogh &amp; Mahunka, 1965" authorityName="Balogh &amp; Mahunka" authorityYear="1965" box="[151,740,893,918]" class="Arachnida" family="Oribatellidae" genus="Tectoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="2" pageNumber="461" phylum="Arthropoda" rank="species" species="deserticola">
<emphasis id="FEE5EAFE661CFFA5FF6FFC82FE7648E4" box="[151,419,893,918]" italics="true" pageId="2" pageNumber="461">Tectoribates deserticola</emphasis>
(Balogh &amp; Mahunka, 1965)
</taxonomicName>
, known from 
<collectingCountry id="B486767C661CFFA5FC78FC82FC2548E4" box="[896,1008,893,918]" name="Mongolia" pageId="2" pageNumber="461">Mongolia</collectingCountry>
; 
<taxonomicName id="0B914D6F661CFFA5FF6FFC5EFD4448C8" authority="Bayartogtokh, 1997" authorityName="Bayartogtokh" authorityYear="1997" box="[151,657,929,954]" class="Arachnida" family="Oribatellidae" genus="Tectoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="2" pageNumber="461" phylum="Arthropoda" rank="species" species="mongolicus">
<emphasis id="FEE5EAFE661CFFA5FF6FFC5EFE7D48C8" box="[151,424,929,954]" italics="true" pageId="2" pageNumber="461">Tectoribates mongolicus</emphasis>
Bayartogtokh, 1997
</taxonomicName>
, known from 
<collectingCountry id="B486767C661CFFA5FCD4FC5EFC4848C8" box="[812,925,929,954]" name="Mongolia" pageId="2" pageNumber="461">Mongolia</collectingCountry>
; 
<taxonomicName id="0B914D6F661CFFA5FF6FFC3AFDE948AC" authority="Schuster, 1958" authorityName="Schuster" authorityYear="1958" box="[151,572,965,990]" class="Arachnida" family="Oribatellidae" genus="Tectoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="2" pageNumber="461" phylum="Arthropoda" rank="species" species="ornatus">
<emphasis id="FEE5EAFE661CFFA5FF6FFC3AFEA848AF" box="[151,381,965,990]" italics="true" pageId="2" pageNumber="461">Tectoribates ornatus</emphasis>
(Schuster, 1958)
</taxonomicName>
known from the Palaearctic and Neotropics.
</paragraph>
</subSubSection>
<subSubSection id="848B6567661CFFA5FF6FFBF2FEE44C37" pageId="2" pageNumber="461" type="diagnosis">
<paragraph id="CC2E36EC661CFFA5FF6FFBF2FC0F4D52" blockId="2.[151,1437,1037,2006]" pageId="2" pageNumber="461">
<emphasis id="FEE5EAFE661CFFA5FF6FFBF2FEC44F54" bold="true" box="[151,273,1037,1062]" pageId="2" pageNumber="461">Diagnosis.</emphasis>
<emphasis id="FEE5EAFE661CFFA5FEE1FBF2FE834F54" box="[281,342,1037,1062]" italics="true" pageId="2" pageNumber="461">Adult</emphasis>
. Species comprising this genus are unique among poronotic 
<taxonomicName id="0B914D6F661CFFA5FBF2FBF2FAAB4F54" authority="(Grandjean 1953b)" baseAuthorityName="Grandjean" baseAuthorityYear="1953" box="[1034,1406,1037,1062]" class="Arachnida" infraOrder="Brachypylina" kingdom="Animalia" order="Sarcoptiformes" pageId="2" pageNumber="461" phylum="Arthropoda" rank="subTribe" subTribe="Brachypylina">Brachypylina (Grandjean 1953b)</taxonomicName>
in having the following combination of character states. Integument with micropuncta throughout. Rostrum with medial keel, or ridge extending from medial margin posteriorly to junction of lamella and prodorsum; rostrum with or without medial incision. Lamellae long, broad, following contour of prodorsum, with large cusps. Lamellar cusps effacing on prodorsum medially posterior to area of attachment of lamellae to prodorsum. Translamella absent. Bothridial wall distally expanded, with scales. Notogaster with posterior notogastral tectum, with immovable pteromorphs lacking anteroventral dens. Octotaxic system as 4 pairs of porose areas or saccules. Notogaster with 10 pairs of setae. Notogaster usually with, occasionally without, subsurface hexagonal pattern anteriorly. Lenticulus absent. Genal tooth present. Tutorium broad, lamelliform, cusp triangular. Discidium strongly curved lamina interlocking with proximal edge of trochanter IV. Epimere I with ’necklace’ of small tubercles anteriorly, extending between setae 
<emphasis id="FEE5EAFE661CFFA5FD4AFA89FD194EFC" box="[690,716,1398,1422]" italics="true" pageId="2" pageNumber="461">1c</emphasis>
. Epimeral seta 
<emphasis id="FEE5EAFE661CFFA5FC74FA89FC734EFC" box="[908,934,1398,1422]" italics="true" pageId="2" pageNumber="461">1c</emphasis>
thickest seta. Genital plates with 6 setae. Postanal porose area present. Axillary saccule present. Palp tarsal eupathidium 
<emphasis id="FEE5EAFE661CFFA5FBC3FA65FBBF4EC3" box="[1083,1130,1434,1457]" italics="true" pageId="2" pageNumber="461">acm</emphasis>
fused to solenidion. Legs heterotridactylous. Genua I and II with large ventral tooth. Seta 
<emphasis id="FEE5EAFE661CFFA5FC6EFA41FC744EA7" box="[918,929,1470,1493]" italics="true" pageId="2" pageNumber="461">s</emphasis>
on tarsus I not eupathidial. Tarsus II with solenidion 
<emphasis id="FEE5EAFE661CFFA5FEE0FA1DFEE64E8C" box="[280,307,1506,1534]" italics="true" pageId="2" pageNumber="461">ω1</emphasis>
most proximal setiform structure; solenidia shorter than setae on tarsus II. One or two minute anterodorsal knobs or spines present on tibia I, close to solenidia 
<emphasis id="FEE5EAFE661CFFA5FC8AF9F6FC544D52" box="[882,897,1545,1568]" italics="true" pageId="2" pageNumber="461">φ</emphasis>
1 and 
<emphasis id="FEE5EAFE661CFFA5FC44F9F6FC1E4D52" box="[956,971,1545,1568]" italics="true" pageId="2" pageNumber="461">φ</emphasis>
2.
</paragraph>
<paragraph id="CC2E36EC661CFFA5FF3FF9CFFEE44C37" blockId="2.[151,1437,1037,2006]" pageId="2" pageNumber="461">
<emphasis id="FEE5EAFE661CFFA5FF3FF9CFFEEB4D35" box="[199,318,1584,1607]" italics="true" pageId="2" pageNumber="461">Immatures</emphasis>
: Deutonymph, Tritonymph. Apheredermous, plicate, without hysterosomal macrosclerites or excentrosclerites, cerotegument as tightly packed platelets, 
<quantity id="0B699B09661CFFA5FCD7F9ACFCA34D1E" box="[815,886,1619,1644]" metricMagnitude="-2" metricUnit="m" metricValue="6.35" metricValueMax="10.16" metricValueMin="2.54" pageId="2" pageNumber="461" unit="in" value="2.5" valueMax="4.0" valueMin="1.0">1–4 in</quantity>
diameter, covering all of body and leg segments. Prodorsal porose regions present. Gastronotal setation unideficient; nymphs with 15 pairs of setae (
<emphasis id="FEE5EAFE661CFFA5FADBF988FAE74DE6" box="[1315,1330,1655,1684]" italics="true" pageId="2" pageNumber="461">f1</emphasis>
, absent). Gastronotal setae of 
<emphasis id="FEE5EAFE661CFFA5FE84F961FE5D4DC7" box="[380,392,1694,1717]" italics="true" pageId="2" pageNumber="461">c</emphasis>
, 
<emphasis id="FEE5EAFE661CFFA5FE6EF962FE714DC4" box="[406,420,1693,1718]" italics="true" pageId="2" pageNumber="461">d</emphasis>
, and 
<emphasis id="FEE5EAFE661CFFA5FE18F962FE3D4DC4" box="[480,488,1693,1718]" italics="true" pageId="2" pageNumber="461">l</emphasis>
series monomorphic, phylliform; setae of 
<emphasis id="FEE5EAFE661CFFA5FC3AF961FC054DC7" box="[962,976,1694,1717]" italics="true" pageId="2" pageNumber="461">p</emphasis>
series phylliform as for other gastronotal setae, or setose. Lamellar, interlamellar setae and setae 
<emphasis id="FEE5EAFE661CFFA5FCFDF93EFCC64DA8" box="[773,787,1729,1754]" italics="true" pageId="2" pageNumber="461">d</emphasis>
of at least femora I and II phylliform. Opisthonotal gland present. Apodemato-acetabular tracheal system or porose homologues absent. Paraprocts atrichous in deutonymph. Cupule development normal. Bothridium cup-shaped. Humeral organ absent. Seta 
<emphasis id="FEE5EAFE661CFFA5FBBAF8F6FB854C50" box="[1090,1104,1801,1826]" italics="true" pageId="2" pageNumber="461">d</emphasis>
present on tibiae I to IV and genua I to III.
</paragraph>
</subSubSection>
<subSubSection id="848B6567661CFFA4FF3FF8AEFC424EB4" lastPageId="3" lastPageNumber="462" pageId="2" pageNumber="461" type="description">
<paragraph id="CC2E36EC661CFFA5FF3FF8AEFAF24CFC" blockId="2.[151,1437,1037,2006]" pageId="2" pageNumber="461">
<emphasis id="FEE5EAFE661CFFA5FF3FF8AEFE844C18" bold="true" box="[199,337,1873,1898]" pageId="2" pageNumber="461">Description</emphasis>
. 
<emphasis id="FEE5EAFE661CFFA5FE9AF8AEFDF84C18" box="[354,557,1873,1898]" italics="true" pageId="2" pageNumber="461">Adult. Integument</emphasis>
. Cerotegument present laterally on podosoma, finely and densely granulate. Integument pitted with micropuncta throughout, puncta or foveae in area between genital and anal plate.
</paragraph>
<paragraph id="CC2E36EC661CFFA4FF3FF866FDA14ADE" blockId="2.[151,1437,1037,2006]" lastBlockId="3.[151,1437,151,2021]" lastPageId="3" lastPageNumber="462" pageId="2" pageNumber="461">
<emphasis id="FEE5EAFE661CFFA5FF3FF866FE914CC0" box="[199,324,1945,1970]" italics="true" pageId="2" pageNumber="461">Prodorsum</emphasis>
. Rostrum with dentate to almost smooth margin, with or without medial incision; medial keel extending from medial margin posteriorly to junction of lamella and prodorsum (
<figureCitation id="54AA2A69661CFFA5FBB7F842FB794CA4" box="[1103,1196,1981,2006]" captionStart="FIGURE 2 A – E" captionStartId="7.[151,250,1866,1888]" captionTargetBox="[229,1346,193,1835]" captionTargetId="figure@7.[229,1346,193,1837]" captionTargetPageId="7" captionText="FIGURE 2 A – E. Tectoribates alcescampestris sp. nov., scanning electron microscope images of adult female from Alberta, Canada, A. habitus dorsal; B, habitus ventral, with arrow to custodium; C, dorsofrontal aspect with arrow to medial ridge on rostrum and arrow to proximal solenidion on tarsus II; D, gnathosoma, with arrow to small lateral tectum on mentum, and arrow to ‘ necklace’ of tubercles on epimere 1; E, anterolateral of prodorsum, with arrow to genal tooth. Scale bar: A, B, C = 20 µm, D, E = 10 µm." httpUri="https://zenodo.org/record/247814/files/figure.png" pageId="2" pageNumber="461">Fig. 2C</figureCitation>
). Rostral setae long, heavily barbed laterally, directed anteromedially. Lamellae long, broad, following contour of prodorsum, with large, cusps; cusps parallel and closely adjacent medially, or diverging anteriorly; cusps with lateral dens, with or without medial dens, shorter or longer than lateral dens (
<figureCitation id="54AA2A69661DFFA4FCDDFF20FCA14B8A" box="[805,884,223,248]" captionStart="FIGURE 1" captionStartId="6.[151,250,1962,1984]" captionTargetBox="[215,1363,200,1920]" captionTargetId="figure@6.[212,1368,195,1923]" captionTargetPageId="6" captionText="FIGURE 1. Tectoribates alcescampestris sp. nov., adult female, dorsal aspect, legs not illustrated, except for proximal segments of leg I. Scale bar = 50 µm." httpUri="https://zenodo.org/record/247813/files/figure.png" pageId="3" pageNumber="462">Figs. 1</figureCitation>
, 
<figureCitation id="54AA2A69661DFFA4FC7CFF1FFC7C4B8A" box="[900,937,224,248]" captionStart="FIGURE 2 A – E" captionStartId="7.[151,250,1866,1888]" captionTargetBox="[229,1346,193,1835]" captionTargetId="figure@7.[229,1346,193,1837]" captionTargetPageId="7" captionText="FIGURE 2 A – E. Tectoribates alcescampestris sp. nov., scanning electron microscope images of adult female from Alberta, Canada, A. habitus dorsal; B, habitus ventral, with arrow to custodium; C, dorsofrontal aspect with arrow to medial ridge on rostrum and arrow to proximal solenidion on tarsus II; D, gnathosoma, with arrow to small lateral tectum on mentum, and arrow to ‘ necklace’ of tubercles on epimere 1; E, anterolateral of prodorsum, with arrow to genal tooth. Scale bar: A, B, C = 20 µm, D, E = 10 µm." httpUri="https://zenodo.org/record/247814/files/figure.png" pageId="3" pageNumber="462">2A</figureCitation>
, 
<figureCitation id="54AA2A69661DFFA4FC4FFF1FFC124B8A" box="[951,967,224,248]" captionStart="FIGURE 4" captionStartId="11.[151,250,1950,1972]" captionTargetBox="[286,1291,266,1939]" captionTargetId="figure@11.[285,1303,265,1940]" captionTargetPageId="11" captionText="FIGURE 4. Tectoribates borealis sp. nov., adult female, dorsal aspect, legs not illustrated, except for proximal segments of leg I. Scale bar = 50 µm." httpUri="https://zenodo.org/record/247816/files/figure.png" pageId="3" pageNumber="462">4</figureCitation>
, 
<figureCitation id="54AA2A69661DFFA4FC2DFF1FFC3A4B8A" box="[981,1007,224,248]" captionStart="FIGURES 11 A – F" captionStartId="18.[151,264,1843,1865]" captionTargetBox="[163,1406,193,1798]" captionTargetId="figure@18.[163,1407,193,1800]" captionTargetPageId="18" captionText="FIGURES 11 A – F. Differential interference contrast microscope images of adults: A, B: Tectoribates alcescampestris sp. nov., adult female; A, prodorsum, dorsal aspect, with arrow to lamellae effacing on prodorsum medially (4 layers combined); B, gnathosoma with arrow to axillary saccule; C – E: Tectoribates borealis sp. nov., C, notogaster, anterior, showing pattern in lenticular region; D, prodorsum, with arrow to lamellae effacing on prodorsum medially (3 layers combined); E, pedotectum I, with arrow to posteroventral region; F, Tectoribates campestris sp. nov., notogaster, anterior, showing pattern in lenticular region; insert showing notogastral saccule Sa. Scale bars: A, D = 50 µm, B = 10 µm, C, E, F = 15 µm." httpUri="https://zenodo.org/record/247822/files/figure.png" pageId="3" pageNumber="462">11</figureCitation>
A, D, 
<figureCitation id="54AA2A69661DFFA4FBC1FF1FFB8D4B8A" box="[1081,1112,224,248]" captionStart="FIGURE 12" captionStartId="21.[151,250,1967,1989]" captionTargetBox="[232,1336,196,1910]" captionTargetId="figure@21.[224,1346,193,1915]" captionTargetPageId="21" captionText="FIGURE 12. Tectoribates campestris sp. nov., adult female, A, dorsal aspect, legs not illustrated, except for proximal segments of leg I. Scale bar = 50 µm." httpUri="https://zenodo.org/record/247823/files/figure.png" pageId="3" pageNumber="462">12</figureCitation>
); lamellar cusps effacing on prodorsum medially, posterior to area of attachment of lamellae to prodorsum (
<figureCitation id="54AA2A69661DFFA4FBE8FEFBFB404A6E" box="[1040,1173,260,285]" captionStart="FIGURE 2 A – E" captionStartId="7.[151,250,1866,1888]" captionTargetBox="[229,1346,193,1835]" captionTargetId="figure@7.[229,1346,193,1837]" captionTargetPageId="7" captionText="FIGURE 2 A – E. Tectoribates alcescampestris sp. nov., scanning electron microscope images of adult female from Alberta, Canada, A. habitus dorsal; B, habitus ventral, with arrow to custodium; C, dorsofrontal aspect with arrow to medial ridge on rostrum and arrow to proximal solenidion on tarsus II; D, gnathosoma, with arrow to small lateral tectum on mentum, and arrow to ‘ necklace’ of tubercles on epimere 1; E, anterolateral of prodorsum, with arrow to genal tooth. Scale bar: A, B, C = 20 µm, D, E = 10 µm." httpUri="https://zenodo.org/record/247814/files/figure.png" pageId="3" pageNumber="462">Figs. 2A, C</figureCitation>
, 
<figureCitation id="54AA2A69661DFFA4FB59FEFBFB224A6E" box="[1185,1271,260,284]" captionStart="FIGURES 11 A – F" captionStartId="18.[151,264,1843,1865]" captionTargetBox="[163,1406,193,1798]" captionTargetId="figure@18.[163,1407,193,1800]" captionTargetPageId="18" captionText="FIGURES 11 A – F. Differential interference contrast microscope images of adults: A, B: Tectoribates alcescampestris sp. nov., adult female; A, prodorsum, dorsal aspect, with arrow to lamellae effacing on prodorsum medially (4 layers combined); B, gnathosoma with arrow to axillary saccule; C – E: Tectoribates borealis sp. nov., C, notogaster, anterior, showing pattern in lenticular region; D, prodorsum, with arrow to lamellae effacing on prodorsum medially (3 layers combined); E, pedotectum I, with arrow to posteroventral region; F, Tectoribates campestris sp. nov., notogaster, anterior, showing pattern in lenticular region; insert showing notogastral saccule Sa. Scale bars: A, D = 50 µm, B = 10 µm, C, E, F = 15 µm." httpUri="https://zenodo.org/record/247822/files/figure.png" pageId="3" pageNumber="462">11A, D</figureCitation>
). Translamella absent. Longitudinal thickening present medially on lamella with nerve canal of seta 
<emphasis id="FEE5EAFE661DFFA4FBBFFED8FB8E4A32" box="[1095,1115,295,320]" italics="true" pageId="3" pageNumber="462">le</emphasis>
extending through thickness (
<figureCitation id="54AA2A69661DFFA4FF67FEB3FEDE4A16" box="[159,267,332,357]" captionStart="FIGURES 11 A – F" captionStartId="18.[151,264,1843,1865]" captionTargetBox="[163,1406,193,1798]" captionTargetId="figure@18.[163,1407,193,1800]" captionTargetPageId="18" captionText="FIGURES 11 A – F. Differential interference contrast microscope images of adults: A, B: Tectoribates alcescampestris sp. nov., adult female; A, prodorsum, dorsal aspect, with arrow to lamellae effacing on prodorsum medially (4 layers combined); B, gnathosoma with arrow to axillary saccule; C – E: Tectoribates borealis sp. nov., C, notogaster, anterior, showing pattern in lenticular region; D, prodorsum, with arrow to lamellae effacing on prodorsum medially (3 layers combined); E, pedotectum I, with arrow to posteroventral region; F, Tectoribates campestris sp. nov., notogaster, anterior, showing pattern in lenticular region; insert showing notogastral saccule Sa. Scale bars: A, D = 50 µm, B = 10 µm, C, E, F = 15 µm." httpUri="https://zenodo.org/record/247822/files/figure.png" pageId="3" pageNumber="462">Fig. 11D</figureCitation>
). Interlamellar setae subequal in thickness to lamellar setae. Exobothridial seta thin, finely barbed. Bothridial wall distally expanded, with scales, without lateral indentation (
<figureCitation id="54AA2A69661DFFA4FC2DFE90FBF74AFA" box="[981,1058,367,392]" captionStart="FIGURE 1" captionStartId="6.[151,250,1962,1984]" captionTargetBox="[215,1363,200,1920]" captionTargetId="figure@6.[212,1368,195,1923]" captionTargetPageId="6" captionText="FIGURE 1. Tectoribates alcescampestris sp. nov., adult female, dorsal aspect, legs not illustrated, except for proximal segments of leg I. Scale bar = 50 µm." httpUri="https://zenodo.org/record/247813/files/figure.png" pageId="3" pageNumber="462">Figs. 1</figureCitation>
, 
<figureCitation id="54AA2A69661DFFA4FBD7FE8FFB804AFA" box="[1071,1109,368,392]" captionStart="FIGURE 2 A – E" captionStartId="7.[151,250,1866,1888]" captionTargetBox="[229,1346,193,1835]" captionTargetId="figure@7.[229,1346,193,1837]" captionTargetPageId="7" captionText="FIGURE 2 A – E. Tectoribates alcescampestris sp. nov., scanning electron microscope images of adult female from Alberta, Canada, A. habitus dorsal; B, habitus ventral, with arrow to custodium; C, dorsofrontal aspect with arrow to medial ridge on rostrum and arrow to proximal solenidion on tarsus II; D, gnathosoma, with arrow to small lateral tectum on mentum, and arrow to ‘ necklace’ of tubercles on epimere 1; E, anterolateral of prodorsum, with arrow to genal tooth. Scale bar: A, B, C = 20 µm, D, E = 10 µm." httpUri="https://zenodo.org/record/247814/files/figure.png" pageId="3" pageNumber="462">2A</figureCitation>
). Porose area Ad not evident. Dorsophragmata fused basally or separate.
</paragraph>
<paragraph id="CC2E36EC661DFFA4FF3FFE46FA59492D" blockId="3.[151,1437,151,2021]" pageId="3" pageNumber="462">
<emphasis id="FEE5EAFE661DFFA4FF3FFE46FE9D4AA2" box="[199,328,441,464]" italics="true" pageId="3" pageNumber="462">Notogaster.</emphasis>
Longer than wide, with immovable pteromorphs having smooth margins without dens anteroventrally (
<figureCitation id="54AA2A69661DFFA4FEA8FE23FE7D4A86" box="[336,424,476,501]" captionStart="FIGURE 2 A – E" captionStartId="7.[151,250,1866,1888]" captionTargetBox="[229,1346,193,1835]" captionTargetId="figure@7.[229,1346,193,1837]" captionTargetPageId="7" captionText="FIGURE 2 A – E. Tectoribates alcescampestris sp. nov., scanning electron microscope images of adult female from Alberta, Canada, A. habitus dorsal; B, habitus ventral, with arrow to custodium; C, dorsofrontal aspect with arrow to medial ridge on rostrum and arrow to proximal solenidion on tarsus II; D, gnathosoma, with arrow to small lateral tectum on mentum, and arrow to ‘ necklace’ of tubercles on epimere 1; E, anterolateral of prodorsum, with arrow to genal tooth. Scale bar: A, B, C = 20 µm, D, E = 10 µm." httpUri="https://zenodo.org/record/247814/files/figure.png" pageId="3" pageNumber="462">Fig. 2B</figureCitation>
). Anterior margin of notogaster undulating, without ridges in region lateral to bothridium. Anterior of notogaster with or without distinct hexagonal pattern in transmitted light (
<figureCitation id="54AA2A69661DFFA4FB8AFE00FB14496A" box="[1138,1217,511,536]" captionStart="FIGURE 1" captionStartId="6.[151,250,1962,1984]" captionTargetBox="[215,1363,200,1920]" captionTargetId="figure@6.[212,1368,195,1923]" captionTargetPageId="6" captionText="FIGURE 1. Tectoribates alcescampestris sp. nov., adult female, dorsal aspect, legs not illustrated, except for proximal segments of leg I. Scale bar = 50 µm." httpUri="https://zenodo.org/record/247813/files/figure.png" pageId="3" pageNumber="462">Figs. 1</figureCitation>
, 
<figureCitation id="54AA2A69661DFFA4FB2AFDFFFB37496A" box="[1234,1250,512,536]" captionStart="FIGURE 4" captionStartId="11.[151,250,1950,1972]" captionTargetBox="[286,1291,266,1939]" captionTargetId="figure@11.[285,1303,265,1940]" captionTargetPageId="11" captionText="FIGURE 4. Tectoribates borealis sp. nov., adult female, dorsal aspect, legs not illustrated, except for proximal segments of leg I. Scale bar = 50 µm." httpUri="https://zenodo.org/record/247816/files/figure.png" pageId="3" pageNumber="462">4</figureCitation>
, 
<figureCitation id="54AA2A69661DFFA4FB08FDFFFAB74965" box="[1264,1378,512,536]" captionStart="FIGURES 11 A – F" captionStartId="18.[151,264,1843,1865]" captionTargetBox="[163,1406,193,1798]" captionTargetId="figure@18.[163,1407,193,1800]" captionTargetPageId="18" captionText="FIGURES 11 A – F. Differential interference contrast microscope images of adults: A, B: Tectoribates alcescampestris sp. nov., adult female; A, prodorsum, dorsal aspect, with arrow to lamellae effacing on prodorsum medially (4 layers combined); B, gnathosoma with arrow to axillary saccule; C – E: Tectoribates borealis sp. nov., C, notogaster, anterior, showing pattern in lenticular region; D, prodorsum, with arrow to lamellae effacing on prodorsum medially (3 layers combined); E, pedotectum I, with arrow to posteroventral region; F, Tectoribates campestris sp. nov., notogaster, anterior, showing pattern in lenticular region; insert showing notogastral saccule Sa. Scale bars: A, D = 50 µm, B = 10 µm, C, E, F = 15 µm." httpUri="https://zenodo.org/record/247822/files/figure.png" pageId="3" pageNumber="462">11A, C, F</figureCitation>
, 
<figureCitation id="54AA2A69661DFFA4FA88FDFFFA5A496A" box="[1392,1423,512,536]" captionStart="FIGURE 12" captionStartId="21.[151,250,1967,1989]" captionTargetBox="[232,1336,196,1910]" captionTargetId="figure@21.[224,1346,193,1915]" captionTargetPageId="21" captionText="FIGURE 12. Tectoribates campestris sp. nov., adult female, A, dorsal aspect, legs not illustrated, except for proximal segments of leg I. Scale bar = 50 µm." httpUri="https://zenodo.org/record/247823/files/figure.png" pageId="3" pageNumber="462">12</figureCitation>
). Octotaxic system expressed as 4 pairs of porose areas or saccules (
<figureCitation id="54AA2A69661DFFA4FC55FDDBFC2A494E" box="[941,1023,548,573]" captionStart="FIGURE 1" captionStartId="6.[151,250,1962,1984]" captionTargetBox="[215,1363,200,1920]" captionTargetId="figure@6.[212,1368,195,1923]" captionTargetPageId="6" captionText="FIGURE 1. Tectoribates alcescampestris sp. nov., adult female, dorsal aspect, legs not illustrated, except for proximal segments of leg I. Scale bar = 50 µm." httpUri="https://zenodo.org/record/247813/files/figure.png" pageId="3" pageNumber="462">Figs. 1</figureCitation>
, 
<figureCitation id="54AA2A69661DFFA4FBE9FDDBFBF4494E" box="[1041,1057,548,572]" captionStart="FIGURE 4" captionStartId="11.[151,250,1950,1972]" captionTargetBox="[286,1291,266,1939]" captionTargetId="figure@11.[285,1303,265,1940]" captionTargetPageId="11" captionText="FIGURE 4. Tectoribates borealis sp. nov., adult female, dorsal aspect, legs not illustrated, except for proximal segments of leg I. Scale bar = 50 µm." httpUri="https://zenodo.org/record/247816/files/figure.png" pageId="3" pageNumber="462">4</figureCitation>
, 
<figureCitation id="54AA2A69661DFFA4FBC9FDDBFB8E494E" box="[1073,1115,548,572]" captionStart="FIGURES 11 A – F" captionStartId="18.[151,264,1843,1865]" captionTargetBox="[163,1406,193,1798]" captionTargetId="figure@18.[163,1407,193,1800]" captionTargetPageId="18" captionText="FIGURES 11 A – F. Differential interference contrast microscope images of adults: A, B: Tectoribates alcescampestris sp. nov., adult female; A, prodorsum, dorsal aspect, with arrow to lamellae effacing on prodorsum medially (4 layers combined); B, gnathosoma with arrow to axillary saccule; C – E: Tectoribates borealis sp. nov., C, notogaster, anterior, showing pattern in lenticular region; D, prodorsum, with arrow to lamellae effacing on prodorsum medially (3 layers combined); E, pedotectum I, with arrow to posteroventral region; F, Tectoribates campestris sp. nov., notogaster, anterior, showing pattern in lenticular region; insert showing notogastral saccule Sa. Scale bars: A, D = 50 µm, B = 10 µm, C, E, F = 15 µm." httpUri="https://zenodo.org/record/247822/files/figure.png" pageId="3" pageNumber="462">11F</figureCitation>
, insert, 12); without sexual dimorphism. With 10 pairs of short setae; distance 
<emphasis id="FEE5EAFE661DFFA4FD29FDB8FCD34916" box="[721,774,583,612]" italics="true" pageId="3" pageNumber="462">h1-h1</emphasis>
shorter than distance 
<emphasis id="FEE5EAFE661DFFA4FC04FDB6FBE44916" box="[1020,1073,585,612]" italics="true" pageId="3" pageNumber="462">p1-p1</emphasis>
; 5 pairs of lyrifissures present.
</paragraph>
<paragraph id="CC2E36EC661DFFA4FF3FFD91FC1E48F0" blockId="3.[151,1437,151,2021]" pageId="3" pageNumber="462">
<emphasis id="FEE5EAFE661DFFA4FF3FFD91FDDE49F5" box="[199,523,622,647]" italics="true" pageId="3" pageNumber="462">Lateral Region of Podosoma</emphasis>
. Genal tooth short, blunt triangular, with longitudinal ridge ventrolaterally (
<figureCitation id="54AA2A69661DFFA4FA9CFD91FF6D49D9" captionStart="FIGURE 2 A – E" captionStartId="7.[151,250,1866,1888]" captionTargetBox="[229,1346,193,1835]" captionTargetId="figure@7.[229,1346,193,1837]" captionTargetPageId="7" captionText="FIGURE 2 A – E. Tectoribates alcescampestris sp. nov., scanning electron microscope images of adult female from Alberta, Canada, A. habitus dorsal; B, habitus ventral, with arrow to custodium; C, dorsofrontal aspect with arrow to medial ridge on rostrum and arrow to proximal solenidion on tarsus II; D, gnathosoma, with arrow to small lateral tectum on mentum, and arrow to ‘ necklace’ of tubercles on epimere 1; E, anterolateral of prodorsum, with arrow to genal tooth. Scale bar: A, B, C = 20 µm, D, E = 10 µm." httpUri="https://zenodo.org/record/247814/files/figure.png" pageId="3" pageNumber="462">Figs. 2E</figureCitation>
, arrow, 8D, 9D). Tutorium broad, lamelliform, cusp triangular, lying parallel to dorsal contour of prodorsum in lateral aspect, extending anterior of insertion of rostral seta (
<figureCitation id="54AA2A69661DFFA4FCBBFD49FC7049BD" box="[835,933,694,719]" captionStart="FIGURE 2 A – E" captionStartId="7.[151,250,1866,1888]" captionTargetBox="[229,1346,193,1835]" captionTargetId="figure@7.[229,1346,193,1837]" captionTargetPageId="7" captionText="FIGURE 2 A – E. Tectoribates alcescampestris sp. nov., scanning electron microscope images of adult female from Alberta, Canada, A. habitus dorsal; B, habitus ventral, with arrow to custodium; C, dorsofrontal aspect with arrow to medial ridge on rostrum and arrow to proximal solenidion on tarsus II; D, gnathosoma, with arrow to small lateral tectum on mentum, and arrow to ‘ necklace’ of tubercles on epimere 1; E, anterolateral of prodorsum, with arrow to genal tooth. Scale bar: A, B, C = 20 µm, D, E = 10 µm." httpUri="https://zenodo.org/record/247814/files/figure.png" pageId="3" pageNumber="462">Figs. 2E</figureCitation>
, 
<figureCitation id="54AA2A69661DFFA4FC4AFD48FC0249BD" box="[946,983,695,719]" captionStart="FIGURES 9 A – D" captionStartId="16.[151,264,1636,1658]" captionTargetBox="[151,1436,193,1614]" captionTargetId="figure@16.[151,1436,193,1615]" captionTargetPageId="16" captionText="FIGURES 9 A – D. Tectoribates borealis sp. nov., scanning electron microscope images of adult female from Alberta, Canada; A, ventral region showing discidium, custodium and foveae on lateral of ventral plate; B, detail of foveae on notogaster, showing foveae mostly plugged with cerotegument; C, prodorsum, anterolateral aspect, with arrow to small teeth lateral to solenidia on tibia I; D, rostrum, laterofrontal aspect, with arrow to proximal solenidion on tarsus II. Scale bar: A, B, C = 10 µm, D = 1 µm." httpUri="https://zenodo.org/record/247820/files/figure.png" pageId="3" pageNumber="462">9D</figureCitation>
). Ventrally directed rostral ridge ventral to rostral seta present or absent. Pedotectum I convex dorsally, without ventral depression (
<figureCitation id="54AA2A69661DFFA4FB2AFD25FAEF4981" box="[1234,1338,730,755]" captionStart="FIGURES 11 A – F" captionStartId="18.[151,264,1843,1865]" captionTargetBox="[163,1406,193,1798]" captionTargetId="figure@18.[163,1407,193,1800]" captionTargetPageId="18" captionText="FIGURES 11 A – F. Differential interference contrast microscope images of adults: A, B: Tectoribates alcescampestris sp. nov., adult female; A, prodorsum, dorsal aspect, with arrow to lamellae effacing on prodorsum medially (4 layers combined); B, gnathosoma with arrow to axillary saccule; C – E: Tectoribates borealis sp. nov., C, notogaster, anterior, showing pattern in lenticular region; D, prodorsum, with arrow to lamellae effacing on prodorsum medially (3 layers combined); E, pedotectum I, with arrow to posteroventral region; F, Tectoribates campestris sp. nov., notogaster, anterior, showing pattern in lenticular region; insert showing notogastral saccule Sa. Scale bars: A, D = 50 µm, B = 10 µm, C, E, F = 15 µm." httpUri="https://zenodo.org/record/247822/files/figure.png" pageId="3" pageNumber="462">Fig. 11E</figureCitation>
, arrow). Pedotectum II present, without tubercle close to body wall. Custodium present, with short to long, free distal point (
<figureCitation id="54AA2A69661DFFA4FF67FCDDFED44849" box="[159,257,802,827]" captionStart="FIGURE 2 A – E" captionStartId="7.[151,250,1866,1888]" captionTargetBox="[229,1346,193,1835]" captionTargetId="figure@7.[229,1346,193,1837]" captionTargetPageId="7" captionText="FIGURE 2 A – E. Tectoribates alcescampestris sp. nov., scanning electron microscope images of adult female from Alberta, Canada, A. habitus dorsal; B, habitus ventral, with arrow to custodium; C, dorsofrontal aspect with arrow to medial ridge on rostrum and arrow to proximal solenidion on tarsus II; D, gnathosoma, with arrow to small lateral tectum on mentum, and arrow to ‘ necklace’ of tubercles on epimere 1; E, anterolateral of prodorsum, with arrow to genal tooth. Scale bar: A, B, C = 20 µm, D, E = 10 µm." httpUri="https://zenodo.org/record/247814/files/figure.png" pageId="3" pageNumber="462">Figs. 2B</figureCitation>
, arrow, 5). Discidium strongly curved lamina interlocking with proximal edge of trochanter IV (
<figureCitation id="54AA2A69661DFFA4FACDFCDDFA424849" box="[1333,1431,802,827]" captionStart="FIGURE 2 A – E" captionStartId="7.[151,250,1866,1888]" captionTargetBox="[229,1346,193,1835]" captionTargetId="figure@7.[229,1346,193,1837]" captionTargetPageId="7" captionText="FIGURE 2 A – E. Tectoribates alcescampestris sp. nov., scanning electron microscope images of adult female from Alberta, Canada, A. habitus dorsal; B, habitus ventral, with arrow to custodium; C, dorsofrontal aspect with arrow to medial ridge on rostrum and arrow to proximal solenidion on tarsus II; D, gnathosoma, with arrow to small lateral tectum on mentum, and arrow to ‘ necklace’ of tubercles on epimere 1; E, anterolateral of prodorsum, with arrow to genal tooth. Scale bar: A, B, C = 20 µm, D, E = 10 µm." httpUri="https://zenodo.org/record/247814/files/figure.png" pageId="3" pageNumber="462">Figs. 2B</figureCitation>
, 
<figureCitation id="54AA2A69661DFFA4FF6FFCB8FF69482D" box="[151,188,839,863]" captionStart="FIGURES 9 A – D" captionStartId="16.[151,264,1636,1658]" captionTargetBox="[151,1436,193,1614]" captionTargetId="figure@16.[151,1436,193,1615]" captionTargetPageId="16" captionText="FIGURES 9 A – D. Tectoribates borealis sp. nov., scanning electron microscope images of adult female from Alberta, Canada; A, ventral region showing discidium, custodium and foveae on lateral of ventral plate; B, detail of foveae on notogaster, showing foveae mostly plugged with cerotegument; C, prodorsum, anterolateral aspect, with arrow to small teeth lateral to solenidia on tibia I; D, rostrum, laterofrontal aspect, with arrow to proximal solenidion on tarsus II. Scale bar: A, B, C = 10 µm, D = 1 µm." httpUri="https://zenodo.org/record/247820/files/figure.png" pageId="3" pageNumber="462">9A</figureCitation>
). Porose area Al present, proximal in position, difficult to see, because overlain by base of lamella and tutorium. Humerosejugal porose organs Am and Ah each expressed as porose area.
</paragraph>
<paragraph id="CC2E36EC661DFFA4FF3FFC71FA494F61" blockId="3.[151,1437,151,2021]" pageId="3" pageNumber="462">
<emphasis id="FEE5EAFE661DFFA4FF3FFC71FEC048D5" box="[199,277,910,935]" italics="true" pageId="3" pageNumber="462">Ventral</emphasis>
. Epimere I with single ‘necklace’ of small tubercles anteriorly, extending between setae 
<emphasis id="FEE5EAFE661DFFA4FB0DFC70FAC548D5" box="[1269,1296,911,935]" italics="true" pageId="3" pageNumber="462">1c</emphasis>
(
<figureCitation id="54AA2A69661DFFA4FAE7FC71FABB48D5" box="[1311,1390,910,935]" captionStart="FIGURE 6" captionStartId="13.[151,250,1919,1941]" captionTargetBox="[267,1320,196,1897]" captionTargetId="figure@13.[260,1326,193,1898]" captionTargetPageId="13" captionText="FIGURE 6. Tectoribates borealis sp. nov., adult female, ventral aspect, legs not illustrated. Scale bar = 50 µm." httpUri="https://zenodo.org/record/247818/files/figure.png" pageId="3" pageNumber="462">Figs. 6</figureCitation>
, 
<figureCitation id="54AA2A69661DFFA4FA82FC70FA4948D5" box="[1402,1436,911,935]" captionStart="FIGURES 8 A – D" captionStartId="15.[151,264,1836,1858]" captionTargetBox="[151,1436,193,1814]" captionTargetId="figure@15.[151,1436,193,1815]" captionTargetPageId="15" captionText="FIGURES 8 A – D. Tectoribates borealis sp. nov., scanning electron microscope images of adult female from Alberta, Canada, A. habitus dorsal; B, prodorsum, with arrow to small teeth lateral to solenidia on tarsus I, and arrow to exobothridial seta; C, habitus ventral, with arrow to foveae between genital and anal plates; D, gnathosoma, with arrow to band of tubercles on anterior of coxisternal region. Scale bar: A, B, C = 20 µm, D = 10 µm." httpUri="https://zenodo.org/record/247819/files/figure.png" pageId="3" pageNumber="462">8D</figureCitation>
arrow). Epimeral setal formula 3-1-3-3 or 3-1-2-3; seta 
<emphasis id="FEE5EAFE661DFFA4FCFBFC4CFCCB48B9" box="[771,798,947,971]" italics="true" pageId="3" pageNumber="462">4c</emphasis>
short and thin, seta 
<emphasis id="FEE5EAFE661DFFA4FC05FC4CFBCD48B9" box="[1021,1048,947,971]" italics="true" pageId="3" pageNumber="462">1c</emphasis>
thickest seta (
<figureCitation id="54AA2A69661DFFA4FB4FFC4DFACC48B9" box="[1207,1305,946,971]" captionStart="FIGURE 2 A – E" captionStartId="7.[151,250,1866,1888]" captionTargetBox="[229,1346,193,1835]" captionTargetId="figure@7.[229,1346,193,1837]" captionTargetPageId="7" captionText="FIGURE 2 A – E. Tectoribates alcescampestris sp. nov., scanning electron microscope images of adult female from Alberta, Canada, A. habitus dorsal; B, habitus ventral, with arrow to custodium; C, dorsofrontal aspect with arrow to medial ridge on rostrum and arrow to proximal solenidion on tarsus II; D, gnathosoma, with arrow to small lateral tectum on mentum, and arrow to ‘ necklace’ of tubercles on epimere 1; E, anterolateral of prodorsum, with arrow to genal tooth. Scale bar: A, B, C = 20 µm, D, E = 10 µm." httpUri="https://zenodo.org/record/247814/files/figure.png" pageId="3" pageNumber="462">Figs. 2B</figureCitation>
, 
<figureCitation id="54AA2A69661DFFA4FADDFC4DFAE048B9" box="[1317,1333,946,971]" captionStart="FIGURE 6" captionStartId="13.[151,250,1919,1941]" captionTargetBox="[267,1320,196,1897]" captionTargetId="figure@13.[260,1326,193,1898]" captionTargetPageId="13" captionText="FIGURE 6. Tectoribates borealis sp. nov., adult female, ventral aspect, legs not illustrated. Scale bar = 50 µm." httpUri="https://zenodo.org/record/247818/files/figure.png" pageId="3" pageNumber="462">6</figureCitation>
). Genital plates with 6 setae; 1 pair aggenital setae; 3 pairs adanal setae and 2 pairs anal setae (
<figureCitation id="54AA2A69661DFFA4FB6AFC29FB0E489D" box="[1170,1243,982,1007]" captionStart="FIGURE 6" captionStartId="13.[151,250,1919,1941]" captionTargetBox="[267,1320,196,1897]" captionTargetId="figure@13.[260,1326,193,1898]" captionTargetPageId="13" captionText="FIGURE 6. Tectoribates borealis sp. nov., adult female, ventral aspect, legs not illustrated. Scale bar = 50 µm." httpUri="https://zenodo.org/record/247818/files/figure.png" pageId="3" pageNumber="462">Fig. 6</figureCitation>
). Lyrifissure 
<emphasis id="FEE5EAFE661DFFA4FA80FC29FA49489D" box="[1400,1436,982,1007]" italics="true" pageId="3" pageNumber="462">iad</emphasis>
anterolateral to anal plate. Postanal porose area present (not evident on 
<figureCitation id="54AA2A69661DFFA4FC57FC05FC254F61" box="[943,1008,1018,1043]" captionStart="FIGURE 6" captionStartId="13.[151,250,1919,1941]" captionTargetBox="[267,1320,196,1897]" captionTargetId="figure@13.[260,1326,193,1898]" captionTargetPageId="13" captionText="FIGURE 6. Tectoribates borealis sp. nov., adult female, ventral aspect, legs not illustrated. Scale bar = 50 µm." httpUri="https://zenodo.org/record/247818/files/figure.png" pageId="3" pageNumber="462">Fig. 6</figureCitation>
because of curvature of ventral plate).
</paragraph>
<paragraph id="CC2E36EC661DFFA4FF3FFBE1FE854FD1" blockId="3.[151,1437,151,2021]" pageId="3" pageNumber="462">
<taxonomicName id="0B914D6F661DFFA4FF3FFBE1FE834F45" box="[199,342,1054,1079]" class="Secernentea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="3" pageNumber="462" phylum="Nematoda" rank="genus">
<emphasis id="FEE5EAFE661DFFA4FF3FFBE1FE834F45" box="[199,342,1054,1079]" italics="true" pageId="3" pageNumber="462">Gnathosoma</emphasis>
</taxonomicName>
. Axillary saccule present at base of palp (
<figureCitation id="54AA2A69661DFFA4FCC7FBE1FC704F45" box="[831,933,1054,1079]" captionStart="FIGURES 11 A – F" captionStartId="18.[151,264,1843,1865]" captionTargetBox="[163,1406,193,1798]" captionTargetId="figure@18.[163,1407,193,1800]" captionTargetPageId="18" captionText="FIGURES 11 A – F. Differential interference contrast microscope images of adults: A, B: Tectoribates alcescampestris sp. nov., adult female; A, prodorsum, dorsal aspect, with arrow to lamellae effacing on prodorsum medially (4 layers combined); B, gnathosoma with arrow to axillary saccule; C – E: Tectoribates borealis sp. nov., C, notogaster, anterior, showing pattern in lenticular region; D, prodorsum, with arrow to lamellae effacing on prodorsum medially (3 layers combined); E, pedotectum I, with arrow to posteroventral region; F, Tectoribates campestris sp. nov., notogaster, anterior, showing pattern in lenticular region; insert showing notogastral saccule Sa. Scale bars: A, D = 50 µm, B = 10 µm, C, E, F = 15 µm." httpUri="https://zenodo.org/record/247822/files/figure.png" pageId="3" pageNumber="462">Fig. 11B</figureCitation>
, arrow). Chelicera chelate-dentate. Mentum with short 
<taxonomicName id="0B914D6F661DFFA4FEE0FBBCFE984F28" box="[280,333,1091,1114]" class="Arachnida" family="Oribatellidae" genus="Oribata" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="3" pageNumber="462" phylum="Arthropoda" rank="species" species="tecta">tecta</taxonomicName>
laterally, anterolateral of setae 
<emphasis id="FEE5EAFE661DFFA4FD3CFBBDFD074F29" box="[708,722,1090,1115]" italics="true" pageId="3" pageNumber="462">h</emphasis>
, without recurved ridge distally (
<figureCitation id="54AA2A69661DFFA4FB98FBBDFB684F29" box="[1120,1213,1090,1115]" captionStart="FIGURE 2 A – E" captionStartId="7.[151,250,1866,1888]" captionTargetBox="[229,1346,193,1835]" captionTargetId="figure@7.[229,1346,193,1837]" captionTargetPageId="7" captionText="FIGURE 2 A – E. Tectoribates alcescampestris sp. nov., scanning electron microscope images of adult female from Alberta, Canada, A. habitus dorsal; B, habitus ventral, with arrow to custodium; C, dorsofrontal aspect with arrow to medial ridge on rostrum and arrow to proximal solenidion on tarsus II; D, gnathosoma, with arrow to small lateral tectum on mentum, and arrow to ‘ necklace’ of tubercles on epimere 1; E, anterolateral of prodorsum, with arrow to genal tooth. Scale bar: A, B, C = 20 µm, D, E = 10 µm." httpUri="https://zenodo.org/record/247814/files/figure.png" pageId="3" pageNumber="462">Fig. 2D</figureCitation>
, arrow). Palp setal formula 0–2–1–3–9(1); eupathidium 
<emphasis id="FEE5EAFE661DFFA4FDBFFB97FDA34F0D" box="[583,630,1128,1151]" italics="true" pageId="3" pageNumber="462">acm</emphasis>
shorter than solenidion, forming double horn with solenidion along length (Fig. 7E).
</paragraph>
<paragraph id="CC2E36EC661DFFA4FF3FFB4FFC424EB4" blockId="3.[151,1437,151,2021]" pageId="3" pageNumber="462">
<emphasis id="FEE5EAFE661DFFA4FF3FFB4FFED64FB5" box="[199,259,1200,1223]" italics="true" pageId="3" pageNumber="462">Legs.</emphasis>
Heterotridactylous (Figs. 7A–D). Leg setal formulas I–IV, solenidia in parentheses (based on species described herein and 
<taxonomicName id="0B914D6F661DFFA4FE7EFB2BFD6F4F99" authority="Bernini 1973" authorityName="Bernini" authorityYear="1973" box="[390,698,1234,1259]" class="Arachnida" family="Oribatellidae" genus="Tectoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="3" pageNumber="462" phylum="Arthropoda" rank="species" species="proximus">
<emphasis id="FEE5EAFE661DFFA4FE7EFB2BFDDC4F99" box="[390,521,1236,1259]" italics="true" pageId="3" pageNumber="462">T. proximus</emphasis>
(Bernini 1973)
</taxonomicName>
: I: 1-5-3(1)-4(2)-19(2); II: 1-5-3(1)-4(1)-15(2); III: 2-3-1(1)-3(1)- 15; IV: 1-2-2-3(1)-12. Genua I and II with large distally directed tooth ventrally (Figs. 7A, B, left arrow). Porose areas on femora I to IV and trochanters III and IV. Seta 
<emphasis id="FEE5EAFE661DFFA4FCF6FAE3FCCC4E41" box="[782,793,1308,1331]" italics="true" pageId="3" pageNumber="462">s</emphasis>
on tarsus I not eupathidial, positioned distal of setae (
<emphasis id="FEE5EAFE661DFFA4FA85FAE3FA5E4E41" box="[1405,1419,1308,1331]" italics="true" pageId="3" pageNumber="462">a</emphasis>
). Tarsus II with proximal solenidion 
<emphasis id="FEE5EAFE661DFFA4FDDFFABFFD964E2E" box="[551,579,1344,1372]" italics="true" pageId="3" pageNumber="462">ω1</emphasis>
most proximal setiform structure on segment; solenidia on tarsus II shorter than setae on segment (Figs. 7B right arrow, 2C). Femur III with seta 
<emphasis id="FEE5EAFE661DFFA4FC64FA9AFC784E0C" box="[924,941,1381,1406]" italics="true" pageId="3" pageNumber="462">l’</emphasis>
present; seta 
<emphasis id="FEE5EAFE661DFFA4FBBCFA99FB824E0F" box="[1092,1111,1382,1405]" italics="true" pageId="3" pageNumber="462">v'</emphasis>
of genua I and II present. On genua and tibiae I and II setae 
<emphasis id="FEE5EAFE661DFFA4FE03FA76FDC74ED0" box="[507,530,1417,1442]" italics="true" pageId="3" pageNumber="462">l”</emphasis>
thicker, more heavily barbed than 
<emphasis id="FEE5EAFE661DFFA4FC5BFA76FC614ED0" box="[931,948,1417,1442]" italics="true" pageId="3" pageNumber="462">l’</emphasis>
. One or two minute anterodorsal knobs or spines present on tibia I close to solenidia 
<emphasis id="FEE5EAFE661DFFA4FD89FA51FD554EB7" box="[625,640,1454,1477]" italics="true" pageId="3" pageNumber="462">φ</emphasis>
1 and 
<emphasis id="FEE5EAFE661DFFA4FD44FA51FD1E4EB7" box="[700,715,1454,1477]" italics="true" pageId="3" pageNumber="462">φ</emphasis>
2 (
<figureCitation id="54AA2A69661DFFA4FD1AFA52FCEC4EB4" box="[738,825,1453,1478]" captionStart="FIGURES 9 A – D" captionStartId="16.[151,264,1636,1658]" captionTargetBox="[151,1436,193,1614]" captionTargetId="figure@16.[151,1436,193,1615]" captionTargetPageId="16" captionText="FIGURES 9 A – D. Tectoribates borealis sp. nov., scanning electron microscope images of adult female from Alberta, Canada; A, ventral region showing discidium, custodium and foveae on lateral of ventral plate; B, detail of foveae on notogaster, showing foveae mostly plugged with cerotegument; C, prodorsum, anterolateral aspect, with arrow to small teeth lateral to solenidia on tibia I; D, rostrum, laterofrontal aspect, with arrow to proximal solenidion on tarsus II. Scale bar: A, B, C = 10 µm, D = 1 µm." httpUri="https://zenodo.org/record/247820/files/figure.png" pageId="3" pageNumber="462">Fig. 9C</figureCitation>
, arrow).
</paragraph>
</subSubSection>
<subSubSection id="848B6567661DFFA3FF3FFA2BFEF94DCD" lastPageId="4" lastPageNumber="463" pageId="3" pageNumber="462" type="discussion">
<paragraph id="CC2E36EC661DFFA4FF3FFA2BFCA34D0F" blockId="3.[151,1437,151,2021]" pageId="3" pageNumber="462">
<emphasis id="FEE5EAFE661DFFA4FF3FFA2BFEE14E9F" bold="true" box="[199,308,1492,1517]" pageId="3" pageNumber="462">Remarks</emphasis>
. 1. The hexagonal pattern present anteriorly on the adult notogaster in most (but not all) species of 
<taxonomicName id="0B914D6F661DFFA4FF6FFA07FEF44D63" box="[151,289,1528,1553]" class="Arachnida" family="Oribatellidae" genus="Tectoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="3" pageNumber="462" phylum="Arthropoda" rank="genus">
<emphasis id="FEE5EAFE661DFFA4FF6FFA07FEF44D63" box="[151,289,1528,1553]" italics="true" pageId="3" pageNumber="462">Tectoribates</emphasis>
</taxonomicName>
is a cuticular structure not visible on the surface, i.e., not visible with SEM (
<figureCitation id="54AA2A69661DFFA4FB6CFA07FACE4D63" box="[1172,1307,1528,1553]" captionStart="FIGURES 8 A – D" captionStartId="15.[151,264,1836,1858]" captionTargetBox="[151,1436,193,1814]" captionTargetId="figure@15.[151,1436,193,1815]" captionTargetPageId="15" captionText="FIGURES 8 A – D. Tectoribates borealis sp. nov., scanning electron microscope images of adult female from Alberta, Canada, A. habitus dorsal; B, prodorsum, with arrow to small teeth lateral to solenidia on tarsus I, and arrow to exobothridial seta; C, habitus ventral, with arrow to foveae between genital and anal plates; D, gnathosoma, with arrow to band of tubercles on anterior of coxisternal region. Scale bar: A, B, C = 20 µm, D = 10 µm." httpUri="https://zenodo.org/record/247819/files/figure.png" pageId="3" pageNumber="462">Figs. 8A, B</figureCitation>
). It has the same position as the lenticulus or “clear-spot” of many taxa, however, there is no difference in pigmentation of the cuticle associated with the hexagonal pattern. Whether this is a light sensory structure, as with the lenticulus of 
<taxonomicName id="0B914D6F661DFFA4FF6FF99BFE744D0F" authority="Banks, 1895" authorityName="Banks" authorityYear="1895" box="[151,417,1636,1661]" class="Arachnida" family="Oribatellidae" genus="Oribatella" kingdom="Animalia" order="Sarcoptiformes" pageId="3" pageNumber="462" phylum="Arthropoda" rank="genus">
<emphasis id="FEE5EAFE661DFFA4FF6FF99BFED94D0F" box="[151,268,1636,1661]" italics="true" pageId="3" pageNumber="462">Oribatella</emphasis>
Banks, 1895
</taxonomicName>
, (Alberti &amp; Fernandez 1990) is unknown.
</paragraph>
<paragraph id="CC2E36EC661DFFA4FF3FF976FEAA4CB3" blockId="3.[151,1437,151,2021]" pageId="3" pageNumber="462">
2. In 
<taxonomicName id="0B914D6F661DFFA4FEFBF977FE594DD3" box="[259,396,1672,1697]" class="Arachnida" family="Oribatellidae" genus="Tectoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="3" pageNumber="462" phylum="Arthropoda" rank="genus">
<emphasis id="FEE5EAFE661DFFA4FEFBF977FE594DD3" box="[259,396,1672,1697]" italics="true" pageId="3" pageNumber="462">Tectoribates</emphasis>
</taxonomicName>
the lamellar cusps do not fuse, rather they efface (merge) on the prodorsum posterior to the region of attachment of the lamella to the prodorsum. In many poronotic 
<taxonomicName id="0B914D6F661DFFA4FC21F953FBA64DB7" box="[985,1139,1708,1733]" class="Arachnida" infraOrder="Brachypylina" kingdom="Animalia" order="Sarcoptiformes" pageId="3" pageNumber="462" phylum="Arthropoda" rank="infraOrder">Brachypylina</taxonomicName>
, lamellar cusp fusion is at the region of attachment of the lamella to the prodorsum, e.g., the mycobatid 
<taxonomicName id="0B914D6F661DFFA4FBDFF92FFA4D4D9B" authority="Behan-Pelletier 1998" authorityName="Behan-Pelletier" authorityYear="1998" box="[1063,1432,1744,1769]" class="Arachnida" family="Mycobatidae" genus="Selvazetes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="3" pageNumber="462" phylum="Arthropoda" rank="genus">
<emphasis id="FEE5EAFE661DFFA4FBDFF92FFB4F4D9B" box="[1063,1178,1744,1769]" italics="true" pageId="3" pageNumber="462">Selvazetes</emphasis>
Behan-Pelletier 1998
</taxonomicName>
. Alternatively, fusion of lamellar cusps can be anterior to the area of attachment of the lamella to the prodorsum, and a ‘pocket’ is formed, e.g., the oribatellid 
<taxonomicName id="0B914D6F661DFFA4FD59F8E7FC9F4C43" box="[673,842,1816,1841]" class="Arachnida" family="Oribatellidae" genus="Ophidiotrichus" kingdom="Animalia" order="Sarcoptiformes" pageId="3" pageNumber="462" phylum="Arthropoda" rank="genus">
<emphasis id="FEE5EAFE661DFFA4FD59F8E7FC9F4C43" box="[673,842,1816,1841]" italics="true" pageId="3" pageNumber="462">Ophidiotrichus</emphasis>
</taxonomicName>
and many species of 
<taxonomicName id="0B914D6F661DFFA4FBB0F8E7FEE74C27" authority="Behan-Pelletier &amp; Walter 2012" authorityName="Behan-Pelletier &amp; Walter" authorityYear="2012" class="Arachnida" family="Oribatellidae" genus="Oribatella" kingdom="Animalia" order="Sarcoptiformes" pageId="3" pageNumber="462" phylum="Arthropoda" rank="genus">
<emphasis id="FEE5EAFE661DFFA4FBB0F8E7FB684C43" box="[1096,1213,1816,1841]" italics="true" pageId="3" pageNumber="462">Oribatella</emphasis>
(Behan-Pelletier &amp; Walter 2012)
</taxonomicName>
. In 
<taxonomicName id="0B914D6F661DFFA4FE9DF8C3FE3A4C27" box="[357,495,1852,1877]" class="Arachnida" family="Oribatellidae" genus="Tectoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="3" pageNumber="462" phylum="Arthropoda" rank="genus">
<emphasis id="FEE5EAFE661DFFA4FE9DF8C3FE3A4C27" box="[357,495,1852,1877]" italics="true" pageId="3" pageNumber="462">Tectoribates</emphasis>
</taxonomicName>
, the area of attachment of the lamella to the prodorsum is outlined by thicker integument on 
<figureCitation id="54AA2A69661DFFA4FEC6F89FFE034C0A" box="[318,470,1888,1913]" captionStart="FIGURES 11 A – F" captionStartId="18.[151,264,1843,1865]" captionTargetBox="[163,1406,193,1798]" captionTargetId="figure@18.[163,1407,193,1800]" captionTargetPageId="18" captionText="FIGURES 11 A – F. Differential interference contrast microscope images of adults: A, B: Tectoribates alcescampestris sp. nov., adult female; A, prodorsum, dorsal aspect, with arrow to lamellae effacing on prodorsum medially (4 layers combined); B, gnathosoma with arrow to axillary saccule; C – E: Tectoribates borealis sp. nov., C, notogaster, anterior, showing pattern in lenticular region; D, prodorsum, with arrow to lamellae effacing on prodorsum medially (3 layers combined); E, pedotectum I, with arrow to posteroventral region; F, Tectoribates campestris sp. nov., notogaster, anterior, showing pattern in lenticular region; insert showing notogastral saccule Sa. Scale bars: A, D = 50 µm, B = 10 µm, C, E, F = 15 µm." httpUri="https://zenodo.org/record/247822/files/figure.png" pageId="3" pageNumber="462">Figs. 11 A, D</figureCitation>
; but dorsomedially the lamellar cusps efface on the prodorsum posterior to this region. This gives the appearance of a narrow cleft extending posteriorly from the region of lamellar attachment to the prodorsum medially.
</paragraph>
<paragraph id="CC2E36EC661DFFA3FF3FF832FB854AFA" blockId="3.[151,1437,151,2021]" lastBlockId="4.[151,1436,151,1619]" lastPageId="4" lastPageNumber="463" pageId="3" pageNumber="462">
3. Coxisternal setation. Bernini (1973) gave an epimeral setation of 3-1-3-3 for both 
<taxonomicName id="0B914D6F661DFFA4FB8DF833FAB94C96" box="[1141,1388,1996,2021]" class="Arachnida" family="Oribatellidae" genus="Tectoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="3" pageNumber="462" phylum="Arthropoda" rank="species" species="proximus">
<emphasis id="FEE5EAFE661DFFA4FB8DF833FAB94C96" box="[1141,1388,1996,2021]" italics="true" pageId="3" pageNumber="462">Tectoribates proximus</emphasis>
</taxonomicName>
and 
<taxonomicName id="0B914D6F661AFFA3FF6FFF66FEDD4BC2" box="[151,264,153,176]" class="Arachnida" family="Oribatellidae" genus="Tectoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="463" phylum="Arthropoda" rank="species" species="ornatus">
<emphasis id="FEE5EAFE661AFFA3FF6FFF66FEDD4BC2" box="[151,264,153,176]" italics="true" pageId="4" pageNumber="463">T. ornatus</emphasis>
</taxonomicName>
, though seta 
<emphasis id="FEE5EAFE661AFFA3FE59FF67FE6E4BC2" box="[417,443,152,176]" italics="true" pageId="4" pageNumber="463">3c</emphasis>
does not appear in his illustration of the venter of 
<taxonomicName id="0B914D6F661AFFA3FC04FF66FBAA4BC2" box="[1020,1151,153,176]" class="Arachnida" family="Oribatellidae" genus="Tectoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="463" phylum="Arthropoda" rank="species" species="proximus">
<emphasis id="FEE5EAFE661AFFA3FC04FF66FBAA4BC2" box="[1020,1151,153,176]" italics="true" pageId="4" pageNumber="463">T. proximus</emphasis>
</taxonomicName>
. Similarly, Bayartogtokh (1997) did not show seta 
<emphasis id="FEE5EAFE661AFFA3FE4EFF43FE044BA6" box="[438,465,188,212]" italics="true" pageId="4" pageNumber="463">3c</emphasis>
in his illustration of the venter of 
<taxonomicName id="0B914D6F661AFFA3FCAEFF42FC264BA7" box="[854,1011,188,213]" class="Arachnida" family="Oribatellidae" genus="Tectoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="463" phylum="Arthropoda" rank="species" species="mongolicus">
<emphasis id="FEE5EAFE661AFFA3FCAEFF42FC264BA7" box="[854,1011,188,213]" italics="true" pageId="4" pageNumber="463">T. mongolicus</emphasis>
</taxonomicName>
and Balogh and Mahunka (1965) did not describe or illustrate the epimeral setation of 
<taxonomicName id="0B914D6F661AFFA3FD46FF1EFC834B8A" box="[702,854,223,248]" class="Arachnida" family="Oribatellidae" genus="Tectoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="463" phylum="Arthropoda" rank="species" species="deserticola">
<emphasis id="FEE5EAFE661AFFA3FD46FF1EFC834B8A" box="[702,854,223,248]" italics="true" pageId="4" pageNumber="463">T. deserticola</emphasis>
</taxonomicName>
. The three North American species of 
<taxonomicName id="0B914D6F661AFFA3FAF4FF20FA404B8A" box="[1292,1429,223,248]" class="Arachnida" family="Oribatellidae" genus="Tectoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="463" phylum="Arthropoda" rank="genus">
<emphasis id="FEE5EAFE661AFFA3FAF4FF20FA404B8A" box="[1292,1429,223,248]" italics="true" pageId="4" pageNumber="463">Tectoribates</emphasis>
</taxonomicName>
, described below, have an epimeral setation of 3-1-2-3. It is clearly seta 
<emphasis id="FEE5EAFE661AFFA3FC44FEFBFC034A6E" box="[956,982,260,284]" italics="true" pageId="4" pageNumber="463">3c</emphasis>
that is absent, as seen by comparing the region lateral of the custodium in 
<figureCitation id="54AA2A69661AFFA3FDEDFED8FDBE4A32" box="[533,619,295,320]" captionStart="FIGURES 9 A – D" captionStartId="16.[151,264,1636,1658]" captionTargetBox="[151,1436,193,1614]" captionTargetId="figure@16.[151,1436,193,1615]" captionTargetPageId="16" captionText="FIGURES 9 A – D. Tectoribates borealis sp. nov., scanning electron microscope images of adult female from Alberta, Canada; A, ventral region showing discidium, custodium and foveae on lateral of ventral plate; B, detail of foveae on notogaster, showing foveae mostly plugged with cerotegument; C, prodorsum, anterolateral aspect, with arrow to small teeth lateral to solenidia on tibia I; D, rostrum, laterofrontal aspect, with arrow to proximal solenidion on tarsus II. Scale bar: A, B, C = 10 µm, D = 1 µm." httpUri="https://zenodo.org/record/247820/files/figure.png" pageId="4" pageNumber="463">Fig. 9A</figureCitation>
with that in 
<taxonomicName id="0B914D6F661AFFA3FD02FED8FCBA4A32" box="[762,879,295,320]" class="Arachnida" family="Oribatellidae" genus="Oribatella" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="463" phylum="Arthropoda" rank="genus">
<emphasis id="FEE5EAFE661AFFA3FD02FED8FCBA4A32" box="[762,879,295,320]" italics="true" pageId="4" pageNumber="463">Oribatella</emphasis>
</taxonomicName>
species (e.g. Figs. 29A–D in Behan-Pelletier and Walter 2012). The ease of misinterpretation of setation on epimeres III and IV in 
<taxonomicName id="0B914D6F661AFFA3FBC6FEB3FB004A17" box="[1086,1237,332,357]" class="Arachnida" infraOrder="Brachypylina" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="463" phylum="Arthropoda" rank="infraOrder">Brachypylina</taxonomicName>
was discussed in detail by Sidorchuk &amp; Norton (2010; their Miscellaneous morphological remarks, 4).
</paragraph>
<paragraph id="CC2E36EC661AFFA3FF3FFE6BFC564981" blockId="4.[151,1436,151,1619]" pageId="4" pageNumber="463">
4. We attributed immatures to 
<taxonomicName id="0B914D6F661AFFA3FDD5FE6BFCBC4ADF" box="[557,873,404,429]" class="Arachnida" family="Oribatellidae" genus="Tectoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="463" phylum="Arthropoda" rank="species" species="alcecampestris">
<emphasis id="FEE5EAFE661AFFA3FDD5FE6BFCBC4ADF" box="[557,873,404,429]" italics="true" pageId="4" pageNumber="463">Tectoribates alcecampestris</emphasis>
</taxonomicName>
and 
<taxonomicName id="0B914D6F661AFFA3FC5FFE6AFBF44ADF" box="[935,1057,404,429]" class="Arachnida" family="Oribatellidae" genus="Tectoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="463" phylum="Arthropoda" rank="species" species="borealis">
<emphasis id="FEE5EAFE661AFFA3FC5FFE6AFBF44ADF" box="[935,1057,404,429]" italics="true" pageId="4" pageNumber="463">T. borealis</emphasis>
</taxonomicName>
(see below) for several reasons. First, they co-ocurred in samples with adult 
<taxonomicName id="0B914D6F661AFFA3FD7BFE48FCD84AA2" box="[643,781,439,464]" class="Arachnida" family="Oribatellidae" genus="Tectoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="463" phylum="Arthropoda" rank="genus">
<emphasis id="FEE5EAFE661AFFA3FD7BFE48FCD84AA2" box="[643,781,439,464]" italics="true" pageId="4" pageNumber="463">Tectoribates</emphasis>
</taxonomicName>
and were of appropriate size. Other brachypyline adults of similar size that were present in these samples included species of 
<taxonomicName id="0B914D6F661AFFA3FC71FE23FBA94A87" authority="Hull, 1916" authorityName="Hull" authorityYear="1916" box="[905,1148,476,501]" class="Arachnida" family="Autognetidae" genus="Autogneta" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="463" phylum="Arthropoda" rank="genus">
<emphasis id="FEE5EAFE661AFFA3FC71FE23FC294A81" box="[905,1020,476,499]" italics="true" pageId="4" pageNumber="463">Autogneta</emphasis>
Hull, 1916
</taxonomicName>
, 
<taxonomicName id="0B914D6F661AFFA3FB73FE23FF1A496A" authority="Oudemans, 1930" authorityName="Oudemans" authorityYear="1930" class="Arachnida" family="Thyrisomidae" genus="Banksinoma" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="463" phylum="Arthropoda" rank="genus">
<emphasis id="FEE5EAFE661AFFA3FB73FE23FAC34A87" box="[1163,1302,476,501]" italics="true" pageId="4" pageNumber="463">Banksinoma</emphasis>
Oudemans, 1930
</taxonomicName>
, and 
<taxonomicName id="0B914D6F661AFFA3FEF3FE00FD96496A" authority="Berlese, 1896" authorityName="Berlese" authorityYear="1896" box="[267,579,511,536]" class="Arachnida" family="Tectocepheidae" genus="Tectocepheus" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="463" phylum="Arthropoda" rank="genus">
<emphasis id="FEE5EAFE661AFFA3FEF3FE00FE74496A" box="[267,417,511,536]" italics="true" pageId="4" pageNumber="463">Tectocepheus</emphasis>
Berlese, 1896
</taxonomicName>
, but immatures of these genera are already known and are distinctly different from our suspected 
<taxonomicName id="0B914D6F661AFFA3FE78FDDBFDDC494F" box="[384,521,548,573]" class="Arachnida" family="Oribatellidae" genus="Tectoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="463" phylum="Arthropoda" rank="genus">
<emphasis id="FEE5EAFE661AFFA3FE78FDDBFDDC494F" box="[384,521,548,573]" italics="true" pageId="4" pageNumber="463">Tectoribates</emphasis>
</taxonomicName>
nymphs: representative works include Grandjean (1960) (
<taxonomicName id="0B914D6F661AFFA3FB4AFDDBFAF04949" box="[1202,1317,548,571]" class="Arachnida" family="Autognetidae" genus="Autogneta" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="463" phylum="Arthropoda" rank="genus">
<emphasis id="FEE5EAFE661AFFA3FB4AFDDBFAF04949" box="[1202,1317,548,571]" italics="true" pageId="4" pageNumber="463">Autogneta</emphasis>
</taxonomicName>
), Ermilov (2010) (
<taxonomicName id="0B914D6F661AFFA3FF0BFDB8FEAB4912" box="[243,382,583,608]" class="Arachnida" family="Thyrisomidae" genus="Banksinoma" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="463" phylum="Arthropoda" rank="genus">
<emphasis id="FEE5EAFE661AFFA3FF0BFDB8FEAB4912" box="[243,382,583,608]" italics="true" pageId="4" pageNumber="463">Banksinoma</emphasis>
</taxonomicName>
) and Pfingstl and Krisper (2011) (
<taxonomicName id="0B914D6F661AFFA3FCFCFDB8FC4F4912" box="[772,922,583,608]" class="Arachnida" family="Tectocepheidae" genus="Tectocepheus" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="463" phylum="Arthropoda" rank="genus">
<emphasis id="FEE5EAFE661AFFA3FCFCFDB8FC4F4912" box="[772,922,583,608]" italics="true" pageId="4" pageNumber="463">Tectocepheus</emphasis>
</taxonomicName>
). In addition, the immatures described below share two important traits with 
<taxonomicName id="0B914D6F661AFFA3FE00FD93FD5449F7" box="[504,641,620,645]" class="Arachnida" family="Oribatellidae" genus="Tectoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="463" phylum="Arthropoda" rank="genus">
<emphasis id="FEE5EAFE661AFFA3FE00FD93FD5449F7" box="[504,641,620,645]" italics="true" pageId="4" pageNumber="463">Tectoribates</emphasis>
</taxonomicName>
adults that are not found in adults or immatures of these other genera: they have an axillary saccule at the base of the palp, and the palp solenidion is fused along the distal half of its length with euapthidium 
<emphasis id="FEE5EAFE661AFFA3FE4FFD4AFE3049BE" box="[439,485,693,716]" italics="true" pageId="4" pageNumber="463">acm</emphasis>
. However, proximal solenidion 
<emphasis id="FEE5EAFE661AFFA3FCA1FD4AFCA149A3" box="[857,884,693,721]" italics="true" pageId="4" pageNumber="463">ω1</emphasis>
on tritonymphal tarsus II is in line with seta 
<emphasis id="FEE5EAFE661AFFA3FA7BFD4BFA4949BE" box="[1411,1436,692,716]" italics="true" pageId="4" pageNumber="463">ft’</emphasis>
rather than being the most proximal setiform structure on segment.
</paragraph>
<paragraph id="CC2E36EC661AFFA3FF3FFD01FF3A48B9" blockId="4.[151,1436,151,1619]" pageId="4" pageNumber="463">
5. Bernini (1973) did not consider 
<taxonomicName id="0B914D6F661AFFA3FDB1FD01FC314865" authority="Balogh, 1959" authorityName="Balogh" authorityYear="1959" box="[585,996,766,791]" class="Arachnida" family="Oribatellidae" genus="Tectoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="463" phylum="Arthropoda" rank="species" species="africanus">
<emphasis id="FEE5EAFE661AFFA3FDB1FD01FC964865" box="[585,835,766,791]" italics="true" pageId="4" pageNumber="463">Tectoribates africanus</emphasis>
Balogh, 1959
</taxonomicName>
, a member of the genus, on the basis of the morphology of the rostrum and the lamellar cusps. He did not record seeing specimens, nor suggest another placement for this species. Balogh and Balogh (2002) placed this species in the oribatellid genus 
<taxonomicName id="0B914D6F661AFFA3FAF5FCB9FECC48F1" authority="Popp, 1960" authorityName="Popp" authorityYear="1960" class="Arachnida" family="Achipteriidae" genus="Plakoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="463" phylum="Arthropoda" rank="genus">
<emphasis id="FEE5EAFE661AFFA3FAF5FCB9FA49482D" box="[1293,1436,838,863]" italics="true" pageId="4" pageNumber="463">Plakoribates</emphasis>
Popp, 1960
</taxonomicName>
, however, Subías (2004, 2013) retained this species in 
<taxonomicName id="0B914D6F661AFFA3FC79FC95FBDF48F1" box="[897,1034,874,899]" class="Arachnida" family="Oribatellidae" genus="Tectoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="463" phylum="Arthropoda" rank="genus">
<emphasis id="FEE5EAFE661AFFA3FC79FC95FBDF48F1" box="[897,1034,874,899]" italics="true" pageId="4" pageNumber="463">Tectoribates</emphasis>
</taxonomicName>
. We do not consider it a member of 
<taxonomicName id="0B914D6F661AFFA3FF6FFC71FEF548D5" box="[151,288,910,935]" class="Arachnida" family="Oribatellidae" genus="Tectoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="463" phylum="Arthropoda" rank="genus">
<emphasis id="FEE5EAFE661AFFA3FF6FFC71FEF548D5" box="[151,288,910,935]" italics="true" pageId="4" pageNumber="463">Tectoribates</emphasis>
</taxonomicName>
, because of the cup-shape morphology of the bothridium, but cannot suggest another placement for the species.
</paragraph>
<paragraph id="CC2E36EC661AFFA3FF3FFC29FCC04E09" blockId="4.[151,1436,151,1619]" pageId="4" pageNumber="463">
6. The genus 
<taxonomicName id="0B914D6F661AFFA3FE97FC29FD3E489D" authority="Subias, 1977" authorityName="Subias" authorityYear="1977" box="[367,747,982,1007]" class="Arachnida" family="Oribatellidae" genus="Pseudotectoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="463" phylum="Arthropoda" rank="genus">
<emphasis id="FEE5EAFE661AFFA3FE97FC29FD93489D" box="[367,582,982,1007]" italics="true" pageId="4" pageNumber="463">Pseudotectoribates</emphasis>
Subías, 1977
</taxonomicName>
, with 
<taxonomicName id="0B914D6F661AFFA3FCC3FC29FAD0489D" authority="Mihelcic, 1956" authorityName="Mihelcic" authorityYear="1956" box="[827,1285,982,1007]" class="Arachnida" family="Achipteriidae" genus="Anachipteria" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="463" phylum="Arthropoda" rank="species" species="subsimilis">
<emphasis id="FEE5EAFE661AFFA3FCC3FC29FB9C489D" box="[827,1097,982,1007]" italics="true" pageId="4" pageNumber="463">Anachipteria subsimilis</emphasis>
Mihelčič, 1956
</taxonomicName>
(= 
<taxonomicName id="0B914D6F661AFFA3FAC0FC27FEF94F61" authority="Subias, 1977" authorityName="Subias" authorityYear="1977" class="Arachnida" family="Oribatellidae" genus="Pseudotectoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="463" phylum="Arthropoda" rank="species" species="bellus">
<emphasis id="FEE5EAFE661AFFA3FAC0FC27FA49489D" box="[1336,1436,982,1007]" italics="true" pageId="4" pageNumber="463">P. bellus</emphasis>
Subías, 1977
</taxonomicName>
). as 
<typeStatus id="132A884E661AFFA3FEA6FC04FE5B4F61" box="[350,398,1019,1043]" pageId="4" pageNumber="463">type</typeStatus>
species, is similar in appearance to 
<taxonomicName id="0B914D6F661AFFA3FCDCFC05FC784F61" box="[804,941,1018,1043]" class="Arachnida" family="Oribatellidae" genus="Tectoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="463" phylum="Arthropoda" rank="genus">
<emphasis id="FEE5EAFE661AFFA3FCDCFC05FC784F61" box="[804,941,1018,1043]" italics="true" pageId="4" pageNumber="463">Tectoribates</emphasis>
</taxonomicName>
, a fact remarked on by Schuster (1958) and Bernini (1973). The genus was defined by Subías (1977) as having lamellae independent along their length, a bothridial seta (sensillus) that is truncated distally, a notogaster with 4 pairs of porose areas and 13 pairs of setae. The three included species are heterotridactylous, as are 
<taxonomicName id="0B914D6F661AFFA3FCD8FB99FC7C4F0D" box="[800,937,1126,1151]" class="Arachnida" family="Oribatellidae" genus="Tectoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="463" phylum="Arthropoda" rank="genus">
<emphasis id="FEE5EAFE661AFFA3FCD8FB99FC7C4F0D" box="[800,937,1126,1151]" italics="true" pageId="4" pageNumber="463">Tectoribates</emphasis>
</taxonomicName>
spp. The illustration of 
<taxonomicName id="0B914D6F661AFFA3FB38FB97FA984F0D" box="[1216,1357,1126,1151]" class="Arachnida" family="Achipteriidae" genus="Anachipteria" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="463" phylum="Arthropoda" rank="species" species="subsimilis">
<emphasis id="FEE5EAFE661AFFA3FB38FB97FA984F0D" box="[1216,1357,1126,1151]" italics="true" pageId="4" pageNumber="463">P. subsimilis</emphasis>
</taxonomicName>
shows the distinctive hexagonal pattern that is found anteriorly on the notogaster of many species of 
<taxonomicName id="0B914D6F661AFFA3FB4FFB75FA944FD1" box="[1207,1345,1162,1187]" class="Arachnida" family="Oribatellidae" genus="Tectoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="463" phylum="Arthropoda" rank="genus">
<emphasis id="FEE5EAFE661AFFA3FB4FFB75FA944FD1" box="[1207,1345,1162,1187]" italics="true" pageId="4" pageNumber="463">Tectoribates</emphasis>
</taxonomicName>
(Subías 1977, Fig. 200). Pérez-Iñigo (1993) and Subías (2004, 2013) considered this genus a member of 
<taxonomicName id="0B914D6F661AFFA3FB2FFB51FAB84FB5" box="[1239,1389,1198,1223]" class="Arachnida" family="Oribatellidae" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="463" phylum="Arthropoda" rank="family">Oribatellidae</taxonomicName>
. We have examined specimens of the 
<typeStatus id="132A884E661AFFA3FDE5FB2CFD984F99" box="[541,589,1235,1259]" pageId="4" pageNumber="463">type</typeStatus>
species 
<taxonomicName id="0B914D6F661AFFA3FD4CFB2DFBD34F99" box="[692,1030,1234,1259]" class="Arachnida" family="Oribatellidae" genus="Pseudotectoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="463" phylum="Arthropoda" rank="species" species="subsimilis">
<emphasis id="FEE5EAFE661AFFA3FD4CFB2DFBD34F99" box="[692,1030,1234,1259]" italics="true" pageId="4" pageNumber="463">Pseudotectoribates subsimilis</emphasis>
</taxonomicName>
from 
<collectingCountry id="B486767C661AFFA3FBAAFB2DFB474F99" box="[1106,1170,1234,1259]" name="Spain" pageId="4" pageNumber="463">Spain</collectingCountry>
(Madrid, Valdelaguna, collector, L. S. Subías) and specimens of 
<taxonomicName id="0B914D6F661AFFA3FD8CFB09FC9F4E7D" box="[628,842,1270,1295]" class="Arachnida" family="Oribatellidae" genus="Pseudotectoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="463" phylum="Arthropoda" rank="genus">
<emphasis id="FEE5EAFE661AFFA3FD8CFB09FC9F4E7D" box="[628,842,1270,1295]" italics="true" pageId="4" pageNumber="463">Pseudotectoribates</emphasis>
</taxonomicName>
sp. from Montpellier, 
<collectingCountry id="B486767C661AFFA3FBB7FB07FB4E4E7D" box="[1103,1179,1272,1295]" name="France" pageId="4" pageNumber="463">France</collectingCountry>
(collector, W. Knülle) and they differ from the diagnosis of 
<taxonomicName id="0B914D6F661AFFA3FDC6FAE5FD104E41" box="[574,709,1306,1331]" class="Arachnida" family="Oribatellidae" genus="Tectoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="463" phylum="Arthropoda" rank="genus">
<emphasis id="FEE5EAFE661AFFA3FDC6FAE5FD104E41" box="[574,709,1306,1331]" italics="true" pageId="4" pageNumber="463">Tectoribates</emphasis>
</taxonomicName>
only in having 13 pairs of notogastral setae. The lamellar cusps are independent and efface on the surface of the prodorsum, as in 
<taxonomicName id="0B914D6F661AFFA3FC53FAC1FBE14E25" box="[939,1076,1342,1367]" class="Arachnida" family="Oribatellidae" genus="Tectoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="463" phylum="Arthropoda" rank="genus">
<emphasis id="FEE5EAFE661AFFA3FC53FAC1FBE14E25" box="[939,1076,1342,1367]" italics="true" pageId="4" pageNumber="463">Tectoribates</emphasis>
</taxonomicName>
, and the necklace of tubercles extending between epimeral setae 
<emphasis id="FEE5EAFE661AFFA3FDE0FA9CFDE64E09" box="[536,563,1379,1403]" italics="true" pageId="4" pageNumber="463">1c</emphasis>
is present ventrally.
</paragraph>
<paragraph id="CC2E36EC661AFFA3FF3FFA77FA8B4D21" blockId="4.[151,1436,151,1619]" pageId="4" pageNumber="463">
7. Sidorchuk and Norton (2010) established a 
<typeStatus id="132A884E661AFFA3FD34FA77FCF04ED2" box="[716,805,1416,1440]" pageId="4" pageNumber="463" type="neotype">neotype</typeStatus>
and provided a photograph for the Eocene Baltic amber species 
<taxonomicName id="0B914D6F661AFFA3FF0DFA55FD444EB1" authority="Sellnick, 1931" authorityName="Sellnick" authorityYear="1931" box="[245,657,1450,1475]" class="Arachnida" family="Oribatellidae" genus="Tectoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="463" phylum="Arthropoda" rank="species" species="parvus">
<emphasis id="FEE5EAFE661AFFA3FF0DFA55FE0D4EB1" box="[245,472,1450,1475]" italics="true" pageId="4" pageNumber="463">Tectoribates parvus</emphasis>
Sellnick, 1931
</taxonomicName>
. 
<collectingCountry id="B486767C661AFFA3FD5AFA54FD144EB1" box="[674,705,1451,1475]" name="American Samoa" pageId="4" pageNumber="463">As</collectingCountry>
they noted this species was transferred from 
<taxonomicName id="0B914D6F661AFFA3FB17FA55FAAD4EB1" box="[1263,1400,1450,1475]" class="Arachnida" family="Oribatellidae" genus="Tectoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="463" phylum="Arthropoda" rank="genus">
<emphasis id="FEE5EAFE661AFFA3FB17FA55FAAD4EB1" box="[1263,1400,1450,1475]" italics="true" pageId="4" pageNumber="463">Tectoribates</emphasis>
</taxonomicName>
to 
<taxonomicName id="0B914D6F661AFFA3FF6FFA31FE094E95" authority="Balogh, 1943" authorityName="Balogh" authorityYear="1943" box="[151,476,1486,1511]" class="Arachnida" family="Unduloribatidae" genus="Unduloribates" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="463" phylum="Arthropoda" rank="genus">
<emphasis id="FEE5EAFE661AFFA3FF6FFA31FEEC4E95" box="[151,313,1486,1511]" italics="true" pageId="4" pageNumber="463">Unduloribates</emphasis>
Balogh, 1943
</taxonomicName>
, by Krivolutsky 
<emphasis id="FEE5EAFE661AFFA3FD63FA2FFD014E95" box="[667,724,1486,1511]" italics="true" pageId="4" pageNumber="463">et al.</emphasis>
(1990) and listed among 
<taxonomicName id="0B914D6F661AFFA3FC02FA31FB594E95" box="[1018,1164,1486,1511]" class="Arachnida" family="Oribatellidae" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="463" phylum="Arthropoda" rank="family">Oribatellidae</taxonomicName>
by these authors, based only on the original description. They noted the resemblance to 
<taxonomicName id="0B914D6F661AFFA3FC95FA0DFBDA4D79" box="[877,1039,1522,1547]" class="Arachnida" family="Unduloribatidae" genus="Unduloribates" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="463" phylum="Arthropoda" rank="genus">
<emphasis id="FEE5EAFE661AFFA3FC95FA0DFBDA4D79" box="[877,1039,1522,1547]" italics="true" pageId="4" pageNumber="463">Unduloribates</emphasis>
</taxonomicName>
in general appearance, but that the length of 264 Μm, is “less than half the length of the smallest reported extant 
<taxonomicName id="0B914D6F661AFFA3FC0CF9E9FB434D5D" box="[1012,1174,1558,1583]" class="Arachnida" family="Unduloribatidae" genus="Unduloribates" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="463" phylum="Arthropoda" rank="genus">
<emphasis id="FEE5EAFE661AFFA3FC0CF9E9FB434D5D" box="[1012,1174,1558,1583]" italics="true" pageId="4" pageNumber="463">Unduloribates</emphasis>
</taxonomicName>
specimen”. Because of the poor condition of the specimen they could not classify this species and its placement awaits further study.
</paragraph>
<paragraph id="CC2E36EC661AFFA3FF6FF959FEF94DCD" blockId="4.[151,300,1702,1727]" box="[151,300,1702,1727]" pageId="4" pageNumber="463">
<emphasis id="FEE5EAFE661AFFA3FF6FF959FEF94DCD" bold="true" box="[151,300,1702,1727]" pageId="4" pageNumber="463">Descriptions</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="848B6567661AFFA3FF6FF911FE6A4C59" pageId="4" pageNumber="463" type="description">
<paragraph id="CC2E36EC661AFFA3FF6FF911FE6A4C59" blockId="4.[151,574,1774,1835]" pageId="4" pageNumber="463">
<emphasis id="FEE5EAFE661AFFA3FF6FF911FDEB4C75" bold="true" box="[151,574,1774,1799]" pageId="4" pageNumber="463">
<taxonomicName id="0B914D6F661AFFA3FF6FF911FE0B4C75" box="[151,478,1774,1799]" class="Arachnida" family="Oribatellidae" genus="Tectoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="463" phylum="Arthropoda" rank="species" species="alcescampestris" status="sp. nov.">
<emphasis id="FEE5EAFE661AFFA3FF6FF911FE0B4C75" bold="true" box="[151,478,1774,1799]" italics="true" pageId="4" pageNumber="463">Tectoribates alcescampestris</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E5D65785661AFFA3FE1CF911FDEB4C75" box="[484,574,1774,1799]" pageId="4" pageNumber="463" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="54AA2A69661AFFA3FF67F8EDFF394C59" box="[159,236,1810,1835]" captionStart="FIGURE 1" captionStartId="6.[151,250,1962,1984]" captionTargetBox="[215,1363,200,1920]" captionTargetId="figure@6.[212,1368,195,1923]" captionTargetPageId="6" captionText="FIGURE 1. Tectoribates alcescampestris sp. nov., adult female, dorsal aspect, legs not illustrated, except for proximal segments of leg I. Scale bar = 50 µm." httpUri="https://zenodo.org/record/247813/files/figure.png" pageId="4" pageNumber="463">Figs. 1</figureCitation>
, 
<figureCitation id="54AA2A69661AFFA3FF02F8ECFEEB4C59" box="[250,318,1811,1835]" captionStart="FIGURE 2 A – E" captionStartId="7.[151,250,1866,1888]" captionTargetBox="[229,1346,193,1835]" captionTargetId="figure@7.[229,1346,193,1837]" captionTargetPageId="7" captionText="FIGURE 2 A – E. Tectoribates alcescampestris sp. nov., scanning electron microscope images of adult female from Alberta, Canada, A. habitus dorsal; B, habitus ventral, with arrow to custodium; C, dorsofrontal aspect with arrow to medial ridge on rostrum and arrow to proximal solenidion on tarsus II; D, gnathosoma, with arrow to small lateral tectum on mentum, and arrow to ‘ necklace’ of tubercles on epimere 1; E, anterolateral of prodorsum, with arrow to genal tooth. Scale bar: A, B, C = 20 µm, D, E = 10 µm." httpUri="https://zenodo.org/record/247814/files/figure.png" pageId="4" pageNumber="463">2A–E</figureCitation>
, 
<figureCitation id="54AA2A69661AFFA3FEB2F8ECFE8F4C59" box="[330,346,1811,1835]" captionStart="FIGURE 3" captionStartId="8.[151,250,1964,1986]" captionTargetBox="[301,1289,199,1923]" captionTargetId="figure@8.[292,1289,193,1930]" captionTargetPageId="8" captionText="FIGURE 3. Tectoribates alcescampestris sp. nov., tritonymph, dorsal aspect, legs not illustrated, except for proximal segments of leg I. Scale bar = 50 µm." httpUri="https://zenodo.org/record/247815/files/figure.png" pageId="4" pageNumber="463">3</figureCitation>
, 
<figureCitation id="54AA2A69661AFFA3FE9EF8ECFE554C59" box="[358,384,1811,1835]" captionStart="FIGURES 11 A – F" captionStartId="18.[151,264,1843,1865]" captionTargetBox="[163,1406,193,1798]" captionTargetId="figure@18.[163,1407,193,1800]" captionTargetPageId="18" captionText="FIGURES 11 A – F. Differential interference contrast microscope images of adults: A, B: Tectoribates alcescampestris sp. nov., adult female; A, prodorsum, dorsal aspect, with arrow to lamellae effacing on prodorsum medially (4 layers combined); B, gnathosoma with arrow to axillary saccule; C – E: Tectoribates borealis sp. nov., C, notogaster, anterior, showing pattern in lenticular region; D, prodorsum, with arrow to lamellae effacing on prodorsum medially (3 layers combined); E, pedotectum I, with arrow to posteroventral region; F, Tectoribates campestris sp. nov., notogaster, anterior, showing pattern in lenticular region; insert showing notogastral saccule Sa. Scale bars: A, D = 50 µm, B = 10 µm, C, E, F = 15 µm." httpUri="https://zenodo.org/record/247822/files/figure.png" pageId="4" pageNumber="463">11</figureCitation>
A, B)
</paragraph>
</subSubSection>
<subSubSection id="848B6567661AFFA2FF6FF8A5FE544ADF" lastPageId="5" lastPageNumber="464" pageId="4" pageNumber="463" type="materials_examined">
<paragraph id="CC2E36EC661AFFA2FF6FF8A5FDFE4A16" blockId="4.[151,1436,1882,2015]" lastBlockId="5.[151,1437,151,2021]" lastPageId="5" lastPageNumber="464" pageId="4" pageNumber="463">
<emphasis id="FEE5EAFE661AFFA3FF6FF8A5FE514C01" bold="true" box="[151,388,1882,1907]" pageId="4" pageNumber="463">Material examined.</emphasis>
<typeStatus id="132A884E661AFFA3FE77F8A5FE294C01" box="[399,508,1882,1907]" pageId="4" pageNumber="463" type="holotype">Holotype</typeStatus>
: adult female, 
<collectingCountry id="B486767C661AFFA3FD53F8A5FCD74C01" box="[683,770,1882,1907]" name="Canada" pageId="4" pageNumber="463">Canada</collectingCountry>
, Alberta, Lamont Co., Moose Pasture, 
<quantity id="0B699B09661AFFA3FB36F8A4FACC4C01" box="[1230,1305,1882,1907]" metricMagnitude="4" metricUnit="m" metricValue="1.1" pageId="4" pageNumber="463" unit="km" value="11.0">11 km</quantity>
S Lamont, aspen litter under snow (53.654167, -112.762778), 
<date id="B82F102C661AFFA3FD11F880FC7C4CE5" box="[745,937,1919,1944]" pageId="4" pageNumber="463" value="2012-01-22">22 January 2012</date>
(D.E. Walter), deposited in the CNC, 
<typeStatus id="132A884E661AFFA3FA94F87FFA494CEA" box="[1388,1436,1920,1944]" pageId="4" pageNumber="463">type</typeStatus>
number 24148. 
<typeStatus id="132A884E661AFFA3FEACF85BFE104CC9" box="[340,453,1956,1979]" pageId="4" pageNumber="463" type="paratype">
<emphasis id="FEE5EAFE661AFFA3FEACF85BFE104CC9" box="[340,453,1956,1979]" italics="true" pageId="4" pageNumber="463">Paratypes</emphasis>
</typeStatus>
<specimenCount id="DA97FD65661AFFA3FE2BF85CFD9D4CC9" box="[467,584,1954,1979]" pageId="4" pageNumber="463" type="female">4 females</specimenCount>
, 
<specimenCount id="DA97FD65661AFFA3FDA1F85CFD604CC9" box="[601,693,1954,1979]" pageId="4" pageNumber="463" type="male">2 males</specimenCount>
with same data as 
<typeStatus id="132A884E661AFFA3FC51F85DFBDA4CC9" box="[937,1039,1954,1979]" pageId="4" pageNumber="463" type="holotype">holotype</typeStatus>
; same location, all (53.656667, - 112.759444), (D.E. Walter), 
<specimenCount id="DA97FD65661AFFA3FE1AF838FD924CAD" box="[482,583,1990,2015]" pageId="4" pageNumber="463" type="female">1 female</specimenCount>
ex littoral moss on margin of slough, 
<date id="B82F102C661AFFA3FBF6F838FB754CAD" box="[1038,1184,1991,2015]" pageId="4" pageNumber="463" value="2008-06-01">1 June 2008</date>
; 
<specimenCount id="DA97FD65661AFFA3FB48F838FAD54CAD" box="[1200,1280,1990,2015]" pageId="4" pageNumber="463" type="adult">1 adult</specimenCount>
ex 
<taxonomicName id="0B914D6F661AFFA2FAC8F839FF284BC2" class="Agaricomycetes" family="Pleurotaceae" genus="Pleurotus" kingdom="Fungi" lastPageId="5" lastPageNumber="464" order="Agaricales" pageId="4" pageNumber="463" phylum="Basidiomycota" rank="species" species="ostreatus">
<emphasis id="FEE5EAFE661AFFA2FAC8F839FF284BC2" italics="true" lastPageId="5" lastPageNumber="464" pageId="4" pageNumber="463">Pleurotus ostreatus</emphasis>
</taxonomicName>
on aspen log, 
<date id="B82F102C661BFFA2FE50FF68FDEA4BC2" box="[424,575,151,176]" pageId="5" pageNumber="464" value="2011-07-16">16 July 2011</date>
; 
<specimenCount id="DA97FD65661BFFA2FDB5FF67FD6E4BC2" box="[589,699,151,176]" pageId="5" pageNumber="464" type="female">2 females</specimenCount>
ex lichen, moss and polypore fungus from aspen log, 
<date id="B82F102C661BFFA2FAD7FF67FF074BA6" pageId="5" pageNumber="464" value="2012-03-11">11 March 2012</date>
, 
<specimenCount id="DA97FD65661BFFA2FF26FF43FEEC4BA7" box="[222,313,188,213]" pageId="5" pageNumber="464" type="male">3 males</specimenCount>
, 
<specimenCount id="DA97FD65661BFFA2FEBEFF43FE794BA7" box="[326,428,188,213]" pageId="5" pageNumber="464" type="female">1 female</specimenCount>
, 1 deutonymph, 1 tritonymph from lichen, moss and polypore on wood; Alberta: ABMI site 1635 NW, (49.41763, -113.99814), 
<quantity id="0B699B09661BFFA2FDAEFF20FD5A4B8A" box="[598,655,223,248]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="464" unit="km" value="5.0">5 km</quantity>
south-west Pincher Creek ex grass, sedge sod, 
<date id="B82F102C661BFFA2FB5BFF20FAC84B8A" box="[1187,1309,223,248]" pageId="5" pageNumber="464" value="2008-06-12">12.vi.2008</date>
; 
<specimenCount id="DA97FD65661BFFA2FAD1FF1FFA564B8A" box="[1321,1411,223,248]" pageId="5" pageNumber="464" type="male">4 males</specimenCount>
, 
<specimenCount id="DA97FD65661BFFA2FA76FF1FFF3B4A6F" pageId="5" pageNumber="464" type="female">2 females</specimenCount>
ABMI site 1412, NE, SE (51.39874, -112.25328) ex prairie sod, 
<date id="B82F102C661BFFA2FC31FEFBFBB44A6E" box="[969,1121,260,285]" pageId="5" pageNumber="464" value="2012-05-31">31 May 2012</date>
; 
<specimenCount id="DA97FD65661BFFA2FB95FEFBFB094A6F" box="[1133,1244,260,285]" pageId="5" pageNumber="464" type="female">2 females</specimenCount>
, ABMI site 1192 NE (52.66613, -110.69303), ex litter in upland aspen mixed woodland, 
<date id="B82F102C661BFFA2FC31FED7FBB34A32" box="[969,1126,296,320]" pageId="5" pageNumber="464" value="2008-06-19">19 June 2008</date>
. 
<typeStatus id="132A884E661BFFA2FB8AFED7FB0A4A32" box="[1138,1247,296,320]" pageId="5" pageNumber="464" type="paratype">Paratypes</typeStatus>
deposited in the PMAE.IZ, CNC, USNM, and RNC.
</paragraph>
<paragraph id="CC2E36EC661BFFA2FF3FFE90FE544ADF" blockId="5.[151,1437,151,2021]" pageId="5" pageNumber="464">
<emphasis id="FEE5EAFE661BFFA2FF3FFE90FDD14AFA" bold="true" box="[199,516,367,392]" pageId="5" pageNumber="464">Other material examined.</emphasis>
Additional individuals from the 
<typeStatus id="132A884E661BFFA2FC70FE8FFC6D4AFA" box="[904,952,368,392]" pageId="5" pageNumber="464">type</typeStatus>
locality were studied under the scanning electron microscope.
</paragraph>
</subSubSection>
<subSubSection id="848B6567661BFFA2FF3FFE48FB904865" pageId="5" pageNumber="464" type="diagnosis">
<paragraph id="CC2E36EC661BFFA2FF3FFE48FB904865" blockId="5.[151,1437,151,2021]" pageId="5" pageNumber="464">
<emphasis id="FEE5EAFE661BFFA2FF3FFE48FE944AA2" bold="true" box="[199,321,439,464]" pageId="5" pageNumber="464">Diagnosis.</emphasis>
Adult. Total length 275–305. Micropuncta between genital and anal plates subequal in size, though more spaced than micropuncta on rest of body. Rostral margin smooth, with medial V or U-shaped indentation. Notogaster anteriorly without distinct hexagonal pattern. Notogaster with porose areas, 
<quantity id="0B699B09661BFFA2FB2EFE00FAF6496A" box="[1238,1315,511,536]" metricMagnitude="-1" metricUnit="m" metricValue="1.651" metricValueMax="2.032" metricValueMin="1.27" pageId="5" pageNumber="464" unit="in" value="6.5" valueMax="8.0" valueMin="5.0">5-8 in</quantity>
diameter. Notogastral setae smooth, 15–27, with seta 
<emphasis id="FEE5EAFE661BFFA2FD78FDDAFD59494E" box="[640,652,549,572]" italics="true" pageId="5" pageNumber="464">c</emphasis>
longest; 
<emphasis id="FEE5EAFE661BFFA2FD0CFDDBFCC5494F" box="[756,784,548,573]" italics="true" pageId="5" pageNumber="464">lm</emphasis>
posterior or medial to Aa and 
<emphasis id="FEE5EAFE661BFFA2FB90FDDBFBAB494F" box="[1128,1150,548,573]" italics="true" pageId="5" pageNumber="464">lp</emphasis>
anterior to medial of A1; mutual distance of 
<emphasis id="FEE5EAFE661BFFA2FE97FDB8FE7C4916" box="[367,425,583,612]" italics="true" pageId="5" pageNumber="464">h1–h1</emphasis>
about 28, of 
<emphasis id="FEE5EAFE661BFFA2FDB8FDB6FDAF4916" box="[576,634,585,612]" italics="true" pageId="5" pageNumber="464">p1–p1</emphasis>
about 44. Lamella about 79 long, of which cusp 37–41; 11–14 wide at level of insertion of seta 
<emphasis id="FEE5EAFE661BFFA2FE38FD91FE0149F5" box="[448,468,622,647]" italics="true" pageId="5" pageNumber="464">le</emphasis>
. Lamellar cusps parallel medially or diverging anteriorly. Lamellar cleft extending posteriorly about 5–9 from area of attachment of lamella to prodorsum; distance between lamellae in this region (width of cleft) about 4. Lamellar cusp with short pointed lateral dens, 8–12 long, medial dens absent. Lamellar seta arising anteriorly on cusp, about 40 long. Bothridial seta club-shaped, 59–64. Seta 
<emphasis id="FEE5EAFE661BFFA2FB88FD23FB5D4981" box="[1136,1160,732,755]" italics="true" pageId="5" pageNumber="464">ex</emphasis>
about 23. Tutorium 69– 72 long, of which cusp about 32. Custodium 10–12 long. Epimeral seta 
<emphasis id="FEE5EAFE661BFFA2FC44FD00FC034865" box="[956,982,767,791]" italics="true" pageId="5" pageNumber="464">1c</emphasis>
about 30.
</paragraph>
</subSubSection>
<subSubSection id="848B6567661BFFAEFF3FFCDDFC964F4F" lastPageId="9" lastPageNumber="468" pageId="5" pageNumber="464" type="description">
<paragraph id="CC2E36EC661BFFA2FF3FFCDDFB71482D" blockId="5.[151,1437,151,2021]" pageId="5" pageNumber="464">
<emphasis id="FEE5EAFE661BFFA2FF3FFCDDFE8C4849" bold="true" box="[199,345,802,827]" pageId="5" pageNumber="464">Description.</emphasis>
Adults. 
<emphasis id="FEE5EAFE661BFFA2FE40FCDBFD904849" box="[440,581,804,827]" italics="true" pageId="5" pageNumber="464">Dimensions:</emphasis>
Total length: females (
<emphasis id="FEE5EAFE661BFFA2FCB1FCDBFC824849" box="[841,855,804,827]" italics="true" pageId="5" pageNumber="464">n</emphasis>
= 2) 303 (300, 305); males (
<emphasis id="FEE5EAFE661BFFA2FB62FCDBFB7D4849" box="[1178,1192,804,827]" italics="true" pageId="5" pageNumber="464">n</emphasis>
= 4) 289 (range 275– 300). Notogastral width: females (
<emphasis id="FEE5EAFE661BFFA2FDE0FCB7FDF3482D" box="[536,550,840,863]" italics="true" pageId="5" pageNumber="464">n</emphasis>
= 2) 200 (200, 200); males (
<emphasis id="FEE5EAFE661BFFA2FC90FCB7FCA3482D" box="[872,886,840,863]" italics="true" pageId="5" pageNumber="464">n</emphasis>
= 4) 198 (range 190–200).
</paragraph>
<paragraph id="CC2E36EC661BFFA2FF3FFC93FE75489D" blockId="5.[151,1437,151,2021]" pageId="5" pageNumber="464">
<emphasis id="FEE5EAFE661BFFA2FF3FFC93FE9B48F1" box="[199,334,876,899]" italics="true" pageId="5" pageNumber="464">Integument:</emphasis>
Prodorsum, notogaster, pedotecta, anal plates, genital plates, venter and mentum micropunctate; puncta between genital and anal plates subequal in size, more widely spaced than micropuncta elsewhere on body (
<figureCitation id="54AA2A69661BFFA2FF67FC4DFF2C48B9" box="[159,249,946,971]" captionStart="FIGURE 2 A – E" captionStartId="7.[151,250,1866,1888]" captionTargetBox="[229,1346,193,1835]" captionTargetId="figure@7.[229,1346,193,1837]" captionTargetPageId="7" captionText="FIGURE 2 A – E. Tectoribates alcescampestris sp. nov., scanning electron microscope images of adult female from Alberta, Canada, A. habitus dorsal; B, habitus ventral, with arrow to custodium; C, dorsofrontal aspect with arrow to medial ridge on rostrum and arrow to proximal solenidion on tarsus II; D, gnathosoma, with arrow to small lateral tectum on mentum, and arrow to ‘ necklace’ of tubercles on epimere 1; E, anterolateral of prodorsum, with arrow to genal tooth. Scale bar: A, B, C = 20 µm, D, E = 10 µm." httpUri="https://zenodo.org/record/247814/files/figure.png" pageId="5" pageNumber="464">Fig. 2B</figureCitation>
). Lamellae, pedotectum, mentum, coxisternal region with fine irregular striae (
<figureCitation id="54AA2A69661BFFA2FB80FC4DFB6B48B9" box="[1144,1214,946,971]" captionStart="FIGURE 2 A – E" captionStartId="7.[151,250,1866,1888]" captionTargetBox="[229,1346,193,1835]" captionTargetId="figure@7.[229,1346,193,1837]" captionTargetPageId="7" captionText="FIGURE 2 A – E. Tectoribates alcescampestris sp. nov., scanning electron microscope images of adult female from Alberta, Canada, A. habitus dorsal; B, habitus ventral, with arrow to custodium; C, dorsofrontal aspect with arrow to medial ridge on rostrum and arrow to proximal solenidion on tarsus II; D, gnathosoma, with arrow to small lateral tectum on mentum, and arrow to ‘ necklace’ of tubercles on epimere 1; E, anterolateral of prodorsum, with arrow to genal tooth. Scale bar: A, B, C = 20 µm, D, E = 10 µm." httpUri="https://zenodo.org/record/247814/files/figure.png" pageId="5" pageNumber="464">Fig. 2</figureCitation>
); tutorial cusp with distinct striae (
<figureCitation id="54AA2A69661BFFA2FEC4FC29FE41489D" box="[316,404,982,1007]" captionStart="FIGURE 2 A – E" captionStartId="7.[151,250,1866,1888]" captionTargetBox="[229,1346,193,1835]" captionTargetId="figure@7.[229,1346,193,1837]" captionTargetPageId="7" captionText="FIGURE 2 A – E. Tectoribates alcescampestris sp. nov., scanning electron microscope images of adult female from Alberta, Canada, A. habitus dorsal; B, habitus ventral, with arrow to custodium; C, dorsofrontal aspect with arrow to medial ridge on rostrum and arrow to proximal solenidion on tarsus II; D, gnathosoma, with arrow to small lateral tectum on mentum, and arrow to ‘ necklace’ of tubercles on epimere 1; E, anterolateral of prodorsum, with arrow to genal tooth. Scale bar: A, B, C = 20 µm, D, E = 10 µm." httpUri="https://zenodo.org/record/247814/files/figure.png" pageId="5" pageNumber="464">Fig. 2E</figureCitation>
).
</paragraph>
<paragraph id="CC2E36EC661BFFA2FF3FFC05FDC54E09" blockId="5.[151,1437,151,2021]" pageId="5" pageNumber="464">
<emphasis id="FEE5EAFE661BFFA2FF3FFC05FE984F61" box="[199,333,1018,1043]" italics="true" pageId="5" pageNumber="464">Prodorsum:</emphasis>
Rostrum with distinct, strongly convex medial crest, rostral margin smooth, with medial U to Vshaped indentation (
<figureCitation id="54AA2A69661BFFA2FE83FBE1FE2B4F45" box="[379,510,1054,1079]" captionStart="FIGURE 2 A – E" captionStartId="7.[151,250,1866,1888]" captionTargetBox="[229,1346,193,1835]" captionTargetId="figure@7.[229,1346,193,1837]" captionTargetPageId="7" captionText="FIGURE 2 A – E. Tectoribates alcescampestris sp. nov., scanning electron microscope images of adult female from Alberta, Canada, A. habitus dorsal; B, habitus ventral, with arrow to custodium; C, dorsofrontal aspect with arrow to medial ridge on rostrum and arrow to proximal solenidion on tarsus II; D, gnathosoma, with arrow to small lateral tectum on mentum, and arrow to ‘ necklace’ of tubercles on epimere 1; E, anterolateral of prodorsum, with arrow to genal tooth. Scale bar: A, B, C = 20 µm, D, E = 10 µm." httpUri="https://zenodo.org/record/247814/files/figure.png" pageId="5" pageNumber="464">Figs. 2C, E</figureCitation>
). Rostrum with deep, cerotegument-filled groove, ventral to seta 
<emphasis id="FEE5EAFE661BFFA2FB19FBDFFB2D4F45" box="[1249,1272,1056,1079]" italics="true" pageId="5" pageNumber="464">ro</emphasis>
(
<figureCitation id="54AA2A69661BFFA2FAF1FBE1FA5A4F45" box="[1289,1423,1054,1079]" captionStart="FIGURE 2 A – E" captionStartId="7.[151,250,1866,1888]" captionTargetBox="[229,1346,193,1835]" captionTargetId="figure@7.[229,1346,193,1837]" captionTargetPageId="7" captionText="FIGURE 2 A – E. Tectoribates alcescampestris sp. nov., scanning electron microscope images of adult female from Alberta, Canada, A. habitus dorsal; B, habitus ventral, with arrow to custodium; C, dorsofrontal aspect with arrow to medial ridge on rostrum and arrow to proximal solenidion on tarsus II; D, gnathosoma, with arrow to small lateral tectum on mentum, and arrow to ‘ necklace’ of tubercles on epimere 1; E, anterolateral of prodorsum, with arrow to genal tooth. Scale bar: A, B, C = 20 µm, D, E = 10 µm." httpUri="https://zenodo.org/record/247814/files/figure.png" pageId="5" pageNumber="464">Figs. 2D, E</figureCitation>
). Cerotegument thickly developed in area defined by this groove and genal tooth. Lamella about 79 long, of which cusp 37–41; 11–14 wide at level of insertion of seta 
<emphasis id="FEE5EAFE661BFFA2FD0CFB99FCDD4F0D" box="[756,776,1126,1151]" italics="true" pageId="5" pageNumber="464">le</emphasis>
. Lamellar cusps parallel medially or diverging anteriorly. Lamellar cleft extending posteriorly about 5–9 from area of attachment of lamella to prodorsum; distance between lamellae in this region (width of cleft) about 4. Lamellar cusp with short, pointed lateral dens, 8–12 long, medial dens absent. Seta 
<emphasis id="FEE5EAFE661BFFA2FE9DFB2BFEA94F99" box="[357,380,1236,1259]" italics="true" pageId="5" pageNumber="464">ro</emphasis>
48–54 long, strongly barbed along length, acuminate, curved anteromedially. Lamellar seta arising anteriorly on cusp, about 40 long, heavily barbed, acuminate distally. Seta 
<emphasis id="FEE5EAFE661BFFA2FB8CFB07FB5F4E7D" box="[1140,1162,1272,1295]" italics="true" pageId="5" pageNumber="464">in</emphasis>
about 72 long, barbed (subequally to 
<emphasis id="FEE5EAFE661BFFA2FEBCFAE5FE8D4E41" box="[324,344,1306,1331]" italics="true" pageId="5" pageNumber="464">le</emphasis>
), acuminate; mutual distance about 33 (
<figureCitation id="54AA2A69661BFFA2FCD5FAE5FCA44E41" box="[813,881,1306,1331]" captionStart="FIGURE 1" captionStartId="6.[151,250,1962,1984]" captionTargetBox="[215,1363,200,1920]" captionTargetId="figure@6.[212,1368,195,1923]" captionTargetPageId="6" captionText="FIGURE 1. Tectoribates alcescampestris sp. nov., adult female, dorsal aspect, legs not illustrated, except for proximal segments of leg I. Scale bar = 50 µm." httpUri="https://zenodo.org/record/247813/files/figure.png" pageId="5" pageNumber="464">Fig. 1</figureCitation>
). Bothridial seta club-shaped, 59–64 long, with barbed, rounded head directed slightly anteromedially (
<figureCitation id="54AA2A69661BFFA2FCD2FAC1FCAD4E25" box="[810,888,1342,1367]" captionStart="FIGURE 1" captionStartId="6.[151,250,1962,1984]" captionTargetBox="[215,1363,200,1920]" captionTargetId="figure@6.[212,1368,195,1923]" captionTargetPageId="6" captionText="FIGURE 1. Tectoribates alcescampestris sp. nov., adult female, dorsal aspect, legs not illustrated, except for proximal segments of leg I. Scale bar = 50 µm." httpUri="https://zenodo.org/record/247813/files/figure.png" pageId="5" pageNumber="464">Figs 1</figureCitation>
, 
<figureCitation id="54AA2A69661BFFA2FC74FAC0FC7A4E25" box="[908,943,1343,1367]" captionStart="FIGURE 2 A – E" captionStartId="7.[151,250,1866,1888]" captionTargetBox="[229,1346,193,1835]" captionTargetId="figure@7.[229,1346,193,1837]" captionTargetPageId="7" captionText="FIGURE 2 A – E. Tectoribates alcescampestris sp. nov., scanning electron microscope images of adult female from Alberta, Canada, A. habitus dorsal; B, habitus ventral, with arrow to custodium; C, dorsofrontal aspect with arrow to medial ridge on rostrum and arrow to proximal solenidion on tarsus II; D, gnathosoma, with arrow to small lateral tectum on mentum, and arrow to ‘ necklace’ of tubercles on epimere 1; E, anterolateral of prodorsum, with arrow to genal tooth. Scale bar: A, B, C = 20 µm, D, E = 10 µm." httpUri="https://zenodo.org/record/247814/files/figure.png" pageId="5" pageNumber="464">2E</figureCitation>
). Seta 
<emphasis id="FEE5EAFE661BFFA2FBFFFABFFBF54E25" box="[1031,1056,1344,1367]" italics="true" pageId="5" pageNumber="464">ex</emphasis>
thin, weakly barbed, about 23. Dorsophragmata separate at base.
</paragraph>
<paragraph id="CC2E36EC661BFFA2FF3FFA79FBFC4EB1" blockId="5.[151,1437,151,2021]" pageId="5" pageNumber="464">
<emphasis id="FEE5EAFE661BFFA2FF3FFA79FDDF4EED" box="[199,522,1414,1439]" italics="true" pageId="5" pageNumber="464">Lateral region of podosoma:</emphasis>
Tutorium 69–72 long, of which cusp about 32. Custodium 10–12 long (
<figureCitation id="54AA2A69661BFFA2FACEFA79FA5B4EED" box="[1334,1422,1414,1439]" captionStart="FIGURE 2 A – E" captionStartId="7.[151,250,1866,1888]" captionTargetBox="[229,1346,193,1835]" captionTargetId="figure@7.[229,1346,193,1837]" captionTargetPageId="7" captionText="FIGURE 2 A – E. Tectoribates alcescampestris sp. nov., scanning electron microscope images of adult female from Alberta, Canada, A. habitus dorsal; B, habitus ventral, with arrow to custodium; C, dorsofrontal aspect with arrow to medial ridge on rostrum and arrow to proximal solenidion on tarsus II; D, gnathosoma, with arrow to small lateral tectum on mentum, and arrow to ‘ necklace’ of tubercles on epimere 1; E, anterolateral of prodorsum, with arrow to genal tooth. Scale bar: A, B, C = 20 µm, D, E = 10 µm." httpUri="https://zenodo.org/record/247814/files/figure.png" pageId="5" pageNumber="464">Fig. 2B</figureCitation>
). Porose area Al about 
<quantity id="0B699B09661BFFA2FE70FA54FE664EB1" box="[392,435,1450,1475]" metricMagnitude="-1" metricUnit="m" metricValue="2.032" pageId="5" pageNumber="464" unit="in" value="8.0">8 in</quantity>
diameter (difficult to see because of proximal position).
</paragraph>
<paragraph id="CC2E36EC661BFFA2FF3FFA2FFE184D24" blockId="5.[151,1437,151,2021]" pageId="5" pageNumber="464">
<emphasis id="FEE5EAFE661BFFA2FF3FFA2FFE994E95" box="[199,332,1488,1511]" italics="true" pageId="5" pageNumber="464">Notogaster:</emphasis>
Slightly longer than wide: 1.1:1. Anteriorly without hexagonal pattern. Octotaxic system expressed as porose areas, 
<quantity id="0B699B09661BFFA2FE37FA0DFDCC4D79" box="[463,537,1522,1547]" metricMagnitude="-1" metricUnit="m" metricValue="1.651" metricValueMax="2.032" metricValueMin="1.27" pageId="5" pageNumber="464" unit="in" value="6.5" valueMax="8.0" valueMin="5.0">5–8 in</quantity>
diameter. Setae smooth, 15–27, with seta 
<emphasis id="FEE5EAFE661BFFA2FBFDFA0BFBC44D79" box="[1029,1041,1524,1547]" italics="true" pageId="5" pageNumber="464">c</emphasis>
longest; 
<emphasis id="FEE5EAFE661BFFA2FB86FA0DFB4F4D79" box="[1150,1178,1522,1547]" italics="true" pageId="5" pageNumber="464">lm</emphasis>
usually medial to Aa, sometimes posterior to (
<figureCitation id="54AA2A69661BFFA2FE50F9E9FE3F4D5D" box="[424,490,1558,1583]" captionStart="FIGURE 1" captionStartId="6.[151,250,1962,1984]" captionTargetBox="[215,1363,200,1920]" captionTargetId="figure@6.[212,1368,195,1923]" captionTargetPageId="6" captionText="FIGURE 1. Tectoribates alcescampestris sp. nov., adult female, dorsal aspect, legs not illustrated, except for proximal segments of leg I. Scale bar = 50 µm." httpUri="https://zenodo.org/record/247813/files/figure.png" pageId="5" pageNumber="464">Fig. 1</figureCitation>
); 
<emphasis id="FEE5EAFE661BFFA2FDFCF9E9FDCF4D5D" box="[516,538,1558,1583]" italics="true" pageId="5" pageNumber="464">lp</emphasis>
usually anterior or medial to and closely adjacent A1; mutual distance of 
<emphasis id="FEE5EAFE661BFFA2FA9BF9E9FA494D46" box="[1379,1436,1558,1588]" italics="true" pageId="5" pageNumber="464">h1–h1</emphasis>
about 28, of 
<emphasis id="FEE5EAFE661BFFA2FEDCF9C1FE8B4D28" box="[292,350,1598,1626]" italics="true" pageId="5" pageNumber="464">p1–p1</emphasis>
about 44.
</paragraph>
<paragraph id="CC2E36EC661BFFA2FF3FF99BFCE14D9B" blockId="5.[151,1437,151,2021]" pageId="5" pageNumber="464">
<emphasis id="FEE5EAFE661BFFA2FF3FF99BFEA24D0E" box="[199,375,1636,1661]" italics="true" pageId="5" pageNumber="464">Ventral Region:</emphasis>
Epimeral setae mostly about 7–30 long, barbed, acuminate, with 
<emphasis id="FEE5EAFE661BFFA2FBA4F99AFBA24D0F" box="[1116,1143,1637,1661]" italics="true" pageId="5" pageNumber="464">1c</emphasis>
more heavily barbed, and thicker than other epimeral setae, about 30; 
<emphasis id="FEE5EAFE661BFFA2FD7FF976FD764DD3" box="[647,675,1673,1697]" italics="true" pageId="5" pageNumber="464">1a</emphasis>
, 
<emphasis id="FEE5EAFE661BFFA2FD4AF976FC994DD3" box="[690,844,1673,1697]" italics="true" pageId="5" pageNumber="464">2a, 3a, 4a, 4c</emphasis>
about 7, 
<emphasis id="FEE5EAFE661BFFA2FC4FF977FC114DD3" box="[951,964,1672,1697]" italics="true" pageId="5" pageNumber="464">1</emphasis>
b 
<date id="B82F102C661BFFA2FC22F977FBE34DD3" box="[986,1078,1672,1697]" pageId="5" pageNumber="464" value="2011-03-16">
11–16 
<emphasis id="FEE5EAFE661BFFA2FBDFF977FBE34DD3" box="[1063,1078,1672,1697]" italics="true" pageId="5" pageNumber="464">3</emphasis>
</date>
b, 4b 14–18, 
<emphasis id="FEE5EAFE661BFFA2FB32F976FB0D4DD3" box="[1226,1240,1673,1697]" italics="true" pageId="5" pageNumber="464">3</emphasis>
c absent. Genital, aggenital, anal and adanal setae smooth; genital setae about 10, others about 6. Genital setae 3+3. Lyrifissure 
<emphasis id="FEE5EAFE661BFFA2FA80F953FA494DB7" box="[1400,1436,1708,1733]" italics="true" pageId="5" pageNumber="464">iad</emphasis>
anterolateral of anal plate. Postanal porose area about 
<date id="B82F102C661BFFA2FD0CF92EFCC24D9A" box="[756,791,1745,1769]" pageId="5" pageNumber="464">9 x</date>
5.
</paragraph>
<paragraph id="CC2E36EC661BFFA2FF3FF90BFA424C42" blockId="5.[151,1437,151,2021]" pageId="5" pageNumber="464">
<taxonomicName id="0B914D6F661BFFA2FF3FF90BFE734C7F" authority="Small" authorityName="Small" box="[199,422,1780,1805]" class="Secernentea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="5" pageNumber="464" phylum="Nematoda" rank="genus">
<emphasis id="FEE5EAFE661BFFA2FF3FF90BFE8A4C7F" box="[199,351,1780,1805]" italics="true" pageId="5" pageNumber="464">Gnathosoma:</emphasis>
Small
</taxonomicName>
axillary saccule present at base of palp, 
<date id="B82F102C661BFFA2FC9FF90BFC5C4C7E" box="[871,905,1780,1805]" pageId="5" pageNumber="464">6 x</date>
2 (
<figureCitation id="54AA2A69661BFFA2FC54F90BFBDB4C7F" box="[940,1038,1780,1805]" captionStart="FIGURES 11 A – F" captionStartId="18.[151,264,1843,1865]" captionTargetBox="[163,1406,193,1798]" captionTargetId="figure@18.[163,1407,193,1800]" captionTargetPageId="18" captionText="FIGURES 11 A – F. Differential interference contrast microscope images of adults: A, B: Tectoribates alcescampestris sp. nov., adult female; A, prodorsum, dorsal aspect, with arrow to lamellae effacing on prodorsum medially (4 layers combined); B, gnathosoma with arrow to axillary saccule; C – E: Tectoribates borealis sp. nov., C, notogaster, anterior, showing pattern in lenticular region; D, prodorsum, with arrow to lamellae effacing on prodorsum medially (3 layers combined); E, pedotectum I, with arrow to posteroventral region; F, Tectoribates campestris sp. nov., notogaster, anterior, showing pattern in lenticular region; insert showing notogastral saccule Sa. Scale bars: A, D = 50 µm, B = 10 µm, C, E, F = 15 µm." httpUri="https://zenodo.org/record/247822/files/figure.png" pageId="5" pageNumber="464">Fig. 11B</figureCitation>
). Cheliceral length 60–70. Mentum with small tectum, anterolateral of seta 
<emphasis id="FEE5EAFE661BFFA2FDA9F8E7FD8A4C43" box="[593,607,1816,1841]" italics="true" pageId="5" pageNumber="464">h</emphasis>
; without recurved ridge distally (
<figureCitation id="54AA2A69661BFFA2FC2DF8E7FBFB4C43" box="[981,1070,1816,1841]" captionStart="FIGURE 2 A – E" captionStartId="7.[151,250,1866,1888]" captionTargetBox="[229,1346,193,1835]" captionTargetId="figure@7.[229,1346,193,1837]" captionTargetPageId="7" captionText="FIGURE 2 A – E. Tectoribates alcescampestris sp. nov., scanning electron microscope images of adult female from Alberta, Canada, A. habitus dorsal; B, habitus ventral, with arrow to custodium; C, dorsofrontal aspect with arrow to medial ridge on rostrum and arrow to proximal solenidion on tarsus II; D, gnathosoma, with arrow to small lateral tectum on mentum, and arrow to ‘ necklace’ of tubercles on epimere 1; E, anterolateral of prodorsum, with arrow to genal tooth. Scale bar: A, B, C = 20 µm, D, E = 10 µm." httpUri="https://zenodo.org/record/247814/files/figure.png" pageId="5" pageNumber="464">Fig. 2D</figureCitation>
, arrow). Length of seta 
<emphasis id="FEE5EAFE661BFFA2FAC2F8E7FA454C43" box="[1338,1424,1816,1841]" italics="true" pageId="5" pageNumber="464">m&gt;a&gt;h</emphasis>
.
</paragraph>
<paragraph id="CC2E36EC661BFFA2FF3FF8C2FB8C4C0B" blockId="5.[151,1437,151,2021]" pageId="5" pageNumber="464">
<emphasis id="FEE5EAFE661BFFA2FF3FF8C2FED34C26" box="[199,262,1853,1876]" italics="true" pageId="5" pageNumber="464">Legs:</emphasis>
Setae 
<emphasis id="FEE5EAFE661BFFA2FEA8F8C3FEB24C27" box="[336,359,1852,1877]" italics="true" pageId="5" pageNumber="464">l”</emphasis>
of genua and tibiae I and II more barbed than other setae on these segments, 
<emphasis id="FEE5EAFE661BFFA2FB32F8C3FB374C27" box="[1226,1250,1852,1877]" italics="true" pageId="5" pageNumber="464">l”</emphasis>
genu II spinose; 
<emphasis id="FEE5EAFE661BFFA2FF6FF89FFF7B4C0B" box="[151,174,1888,1913]" italics="true" pageId="5" pageNumber="464">l”</emphasis>
genu and tibia I about 14 and 17, 
<emphasis id="FEE5EAFE661BFFA2FDD5F89FFD914C0B" box="[557,580,1888,1913]" italics="true" pageId="5" pageNumber="464">l”</emphasis>
genu and tibia II about 14 and 20, respectively.
</paragraph>
<paragraph id="CC2E36EC661BFFA2FF3FF87BFAFB4CEF" blockId="5.[151,1437,151,2021]" box="[199,1326,1924,1949]" pageId="5" pageNumber="464">
<emphasis id="FEE5EAFE661BFFA2FF3FF87BFE8C4CEF" bold="true" box="[199,345,1924,1949]" pageId="5" pageNumber="464">Description.</emphasis>
<emphasis id="FEE5EAFE661BFFA2FEA7F87AFDBF4CEE" box="[351,618,1925,1948]" italics="true" pageId="5" pageNumber="464">Immatures Dimensions:</emphasis>
Total length: deutonymph (
<emphasis id="FEE5EAFE661BFFA2FC5BF87AFC644CEE" box="[931,945,1925,1948]" italics="true" pageId="5" pageNumber="464">n</emphasis>
= 1) 288; tritonymph (
<emphasis id="FEE5EAFE661BFFA2FB4AF87AFB154CEE" box="[1202,1216,1925,1948]" italics="true" pageId="5" pageNumber="464">n</emphasis>
= 1) 312.
</paragraph>
<paragraph id="CC2E36EC661BFFA2FF3FF857FCD34C97" blockId="5.[151,1437,151,2021]" pageId="5" pageNumber="464">
Tritonymph. 
<emphasis id="FEE5EAFE661BFFA2FEA4F856FE0F4CB2" box="[348,474,1961,1984]" italics="true" pageId="5" pageNumber="464">Integument</emphasis>
: Sclerotized with light brown coloration. Cerotegument of tightly packed platelets, about 
<quantity id="0B699B09661BFFA2FF24F832FED24C97" box="[220,263,1996,2021]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="5" pageNumber="464" unit="in" value="1.0">1 in</quantity>
width covering all of body and leg segments.
</paragraph>
<caption id="98EE66646618FFA1FF6FF855FDC74CAC" httpUri="https://zenodo.org/record/247813/files/figure.png" pageId="6" pageNumber="465" targetBox="[215,1363,200,1920]" targetPageId="6">
<paragraph id="CC2E36EC6618FFA1FF6FF855FDC74CAC" blockId="6.[151,1435,1962,2014]" pageId="6" pageNumber="465">
<emphasis id="FEE5EAFE6618FFA1FF6FF855FECD4CB2" bold="true" box="[151,280,1962,1984]" pageId="6" pageNumber="465">FIGURE 1.</emphasis>
<taxonomicName id="0B914D6F6618FFA1FEDCF855FD984CB2" box="[292,589,1962,1984]" class="Arachnida" family="Oribatellidae" genus="Tectoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="6" pageNumber="465" phylum="Arthropoda" rank="species" species="alcescampestris" status="sp. nov.">
<emphasis id="FEE5EAFE6618FFA1FEDCF855FD984CB2" box="[292,589,1962,1984]" italics="true" pageId="6" pageNumber="465">Tectoribates alcescampestris</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E5D657856618FFA1FDA1F855FD7A4CB2" box="[601,687,1962,1984]" pageId="6" pageNumber="465" rank="species">
<emphasis id="FEE5EAFE6618FFA1FDA1F855FD7A4CB2" bold="true" box="[601,687,1962,1984]" pageId="6" pageNumber="465">sp. nov.</emphasis>
</taxonomicNameLabel>
, adult female, dorsal aspect, legs not illustrated, except for proximal segments of leg I. Scale bar = 50 µm.
</paragraph>
</caption>
<caption id="98EE66646619FFA0FF6FF8B5FE8D4CA8" httpUri="https://zenodo.org/record/247814/files/figure.png" pageId="7" pageNumber="466" targetBox="[229,1346,193,1835]" targetPageId="7">
<paragraph id="CC2E36EC6619FFA0FF6FF8B5FE8D4CA8" blockId="7.[151,1436,1866,2010]" pageId="7" pageNumber="466">
<emphasis id="FEE5EAFE6619FFA0FF6FF8B5FE904C12" bold="true" box="[151,325,1866,1888]" pageId="7" pageNumber="466">FIGURE 2A–E.</emphasis>
<taxonomicName id="0B914D6F6619FFA0FEB5F8B5FDA14C12" box="[333,628,1866,1888]" class="Arachnida" family="Oribatellidae" genus="Tectoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="7" pageNumber="466" phylum="Arthropoda" rank="species" species="alcescampestris" status="sp. nov.">
<emphasis id="FEE5EAFE6619FFA0FEB5F8B5FDA14C12" box="[333,628,1866,1888]" italics="true" pageId="7" pageNumber="466">Tectoribates alcescampestris</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E5D657856619FFA0FD84F8B5FD1A4C12" box="[636,719,1866,1888]" pageId="7" pageNumber="466" rank="species">
<emphasis id="FEE5EAFE6619FFA0FD84F8B5FD1A4C12" bold="true" box="[636,719,1866,1888]" pageId="7" pageNumber="466">sp. nov.</emphasis>
</taxonomicNameLabel>
, scanning electron microscope images of adult female from Alberta, Canada, A. habitus dorsal; B, habitus ventral, with arrow to custodium; C, dorsofrontal aspect with arrow to medial ridge on rostrum and arrow to proximal solenidion on tarsus II; D, 
<taxonomicName id="0B914D6F6619FFA0FD04F878FCAC4CEF" box="[764,889,1927,1949]" class="Secernentea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="7" pageNumber="479" phylum="Nematoda" rank="genus">gnathosoma</taxonomicName>
, with arrow to small lateral tectum on mentum, and arrow to ‘necklace’ of tubercles on epimere 1; E, anterolateral of prodorsum, with arrow to genal tooth. Scale bar: A, B, C = 20 µm, D, E = 10 µm.
</paragraph>
</caption>
<caption id="98EE66646616FFAFFF6FF853FE7B4C93" httpUri="https://zenodo.org/record/247815/files/figure.png" pageId="8" pageNumber="467" targetBox="[301,1289,199,1923]" targetPageId="8">
<paragraph id="CC2E36EC6616FFAFFF6FF853FE7B4C93" blockId="8.[151,1435,1964,2017]" pageId="8" pageNumber="467">
<emphasis id="FEE5EAFE6616FFAFFF6FF853FEC74CB0" bold="true" box="[151,274,1964,1986]" pageId="8" pageNumber="467">FIGURE 3.</emphasis>
<taxonomicName id="0B914D6F6616FFAFFEE0F853FDE94CB0" box="[280,572,1964,1986]" class="Arachnida" family="Oribatellidae" genus="Tectoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="8" pageNumber="467" phylum="Arthropoda" rank="species" species="alcescampestris" status="sp. nov.">
<emphasis id="FEE5EAFE6616FFAFFEE0F853FDE94CB0" box="[280,572,1964,1986]" italics="true" pageId="8" pageNumber="467">Tectoribates alcescampestris</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E5D657856616FFAFFDBAF853FD444CB0" box="[578,657,1964,1986]" pageId="8" pageNumber="467" rank="species">
<emphasis id="FEE5EAFE6616FFAFFDBAF853FD444CB0" bold="true" box="[578,657,1964,1986]" pageId="8" pageNumber="467">sp. nov.</emphasis>
</taxonomicNameLabel>
, tritonymph, dorsal aspect, legs not illustrated, except for proximal segments of leg I. Scale bar = 50 µm.
</paragraph>
</caption>
<paragraph id="CC2E36EC6617FFAEFF3FFF68FCB24A17" blockId="9.[151,1437,151,1085]" pageId="9" pageNumber="468">
<emphasis id="FEE5EAFE6617FFAEFF3FFF68FE984BC2" box="[199,333,151,176]" italics="true" pageId="9" pageNumber="468">Prodorsum:</emphasis>
Aspis with transverse porose area posterior to setae 
<emphasis id="FEE5EAFE6617FFAEFC67FF66FC604BC2" box="[927,949,153,176]" italics="true" pageId="9" pageNumber="468">in</emphasis>
and bothridia. Lateral porose areas present anterior of bothridia (porose areas not illustrated in 
<figureCitation id="54AA2A696617FFAEFD21FF43FCCE4BA6" box="[729,795,188,213]" captionStart="FIGURE 3" captionStartId="8.[151,250,1964,1986]" captionTargetBox="[301,1289,199,1923]" captionTargetId="figure@8.[292,1289,193,1930]" captionTargetPageId="8" captionText="FIGURE 3. Tectoribates alcescampestris sp. nov., tritonymph, dorsal aspect, legs not illustrated, except for proximal segments of leg I. Scale bar = 50 µm." httpUri="https://zenodo.org/record/247815/files/figure.png" pageId="9" pageNumber="468">Fig. 3</figureCitation>
because of difficulty in removing cerotegument). Seta 
<emphasis id="FEE5EAFE6617FFAEFA7CFF42FA494BA6" box="[1412,1436,189,212]" italics="true" pageId="9" pageNumber="468">ro</emphasis>
finely barbed along length, tapered, about 27 long. Seta 
<emphasis id="FEE5EAFE6617FFAEFCD4FF20FC954B8A" box="[812,832,223,248]" italics="true" pageId="9" pageNumber="468">le</emphasis>
barbed, phylliform, about 20 long, 10 at maximum width. Seta 
<emphasis id="FEE5EAFE6617FFAEFEE3FEFAFEE44A6E" box="[283,305,261,284]" italics="true" pageId="9" pageNumber="468">in</emphasis>
barbed, phylliform, about 26 long. Mutual distance of setae 
<emphasis id="FEE5EAFE6617FFAEFC29FEFAFB9C4A6F" box="[977,1097,260,285]" italics="true" pageId="9" pageNumber="468">ro-ro, le-le</emphasis>
and 
<emphasis id="FEE5EAFE6617FFAEFB86FEFAFB664A6E" box="[1150,1203,261,284]" italics="true" pageId="9" pageNumber="468">in-in</emphasis>
about 18, 22 and 52, respectively. Seta 
<emphasis id="FEE5EAFE6617FFAEFE9DFED6FEAB4A32" box="[357,382,297,320]" italics="true" pageId="9" pageNumber="468">ex</emphasis>
about 7 long, thick, heavily barbed, penicillate. Bothridial seta 60–66 long, barbed, clavate, head subequal in length to stalk, more tapered than that of adult.
</paragraph>
<paragraph id="CC2E36EC6617FFAEFF3FFE8FFE9B4953" blockId="9.[151,1437,151,1085]" pageId="9" pageNumber="468">
<emphasis id="FEE5EAFE6617FFAEFF3FFE8FFE764AFA" box="[199,419,368,392]" italics="true" pageId="9" pageNumber="468">Gastronotic Region</emphasis>
: Integument without sclerotization; orientation of plicae more transverse anteriorly, more longitudinal posteriorly (
<figureCitation id="54AA2A696617FFAEFE48FE6BFE204ADE" box="[432,501,404,429]" captionStart="FIGURE 3" captionStartId="8.[151,250,1964,1986]" captionTargetBox="[301,1289,199,1923]" captionTargetId="figure@8.[292,1289,193,1930]" captionTargetPageId="8" captionText="FIGURE 3. Tectoribates alcescampestris sp. nov., tritonymph, dorsal aspect, legs not illustrated, except for proximal segments of leg I. Scale bar = 50 µm." httpUri="https://zenodo.org/record/247815/files/figure.png" pageId="9" pageNumber="468">Fig. 3</figureCitation>
). Setae of 
<emphasis id="FEE5EAFE6617FFAEFD88FE6AFD624ADF" box="[624,695,404,429]" italics="true" pageId="9" pageNumber="468">c, d, l,</emphasis>
and 
<emphasis id="FEE5EAFE6617FFAEFD08FE6BFD2B4ADF" box="[752,766,404,429]" italics="true" pageId="9" pageNumber="468">h</emphasis>
series about 17–26 long, with seta 
<emphasis id="FEE5EAFE6617FFAEFB69FE6AFB704AC3" box="[1169,1189,405,433]" italics="true" pageId="9" pageNumber="468">c3</emphasis>
shortest seta (
<figureCitation id="54AA2A696617FFAEFAB0FE6BFA584ADE" box="[1352,1421,404,429]" captionStart="FIGURE 3" captionStartId="8.[151,250,1964,1986]" captionTargetBox="[301,1289,199,1923]" captionTargetId="figure@8.[292,1289,193,1930]" captionTargetPageId="8" captionText="FIGURE 3. Tectoribates alcescampestris sp. nov., tritonymph, dorsal aspect, legs not illustrated, except for proximal segments of leg I. Scale bar = 50 µm." httpUri="https://zenodo.org/record/247815/files/figure.png" pageId="9" pageNumber="468">Fig. 3</figureCitation>
). Setae of 
<emphasis id="FEE5EAFE6617FFAEFEFBFE43FEC44AA1" box="[259,273,444,467]" italics="true" pageId="9" pageNumber="468">p</emphasis>
series (visible only in ventral aspect); 
<emphasis id="FEE5EAFE6617FFAEFD1CFE43FD2F4AA5" box="[740,762,444,471]" italics="true" pageId="9" pageNumber="468">p1</emphasis>
narrow, barbed, phylliform, about 14, 
<emphasis id="FEE5EAFE6617FFAEFB31FE43FB0A4AA5" box="[1225,1247,444,471]" italics="true" pageId="9" pageNumber="468">p2</emphasis>
heavily barbed, tapered, about 14, 
<emphasis id="FEE5EAFE6617FFAEFE9EFE1DFEA94A8C" box="[358,380,482,510]" italics="true" pageId="9" pageNumber="468">p3</emphasis>
barbed, tapered, about 14. Mutual distance of setae 
<emphasis id="FEE5EAFE6617FFAEFC3DFE1EFBB74A88" box="[965,1122,481,506]" italics="true" pageId="9" pageNumber="468">da-da, dm-dm</emphasis>
and 
<emphasis id="FEE5EAFE6617FFAEFB61FE1EFB0E4A88" box="[1177,1243,481,506]" italics="true" pageId="9" pageNumber="468">dp-dp</emphasis>
about 51, 33 and 23, respectively.
</paragraph>
<paragraph id="CC2E36EC6617FFAEFF3FFDD3FE9549C3" blockId="9.[151,1437,151,1085]" pageId="9" pageNumber="468">
<emphasis id="FEE5EAFE6617FFAEFF3FFDD3FEA14936" box="[199,372,556,581]" italics="true" pageId="9" pageNumber="468">Ventral Region</emphasis>
: Medial margin of epimeral plates weakly defined. Epimere I with narrow mentotectum developed laterally. Epimeral, genital, aggenital, anal and adanal setae smooth, acuminate, about 10 long. Porose integument surrounding opening of opisthonotal gland not evident (possibly because area covered by cerotegument).
</paragraph>
<paragraph id="CC2E36EC6617FFAEFF3FFD43FDBA498B" blockId="9.[151,1437,151,1085]" pageId="9" pageNumber="468">
<taxonomicName id="0B914D6F6617FFAEFF3FFD43FE1249A7" authority="Axillary" authorityName="Axillary" box="[199,455,700,725]" class="Secernentea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="9" pageNumber="468" phylum="Nematoda" rank="genus">
<emphasis id="FEE5EAFE6617FFAEFF3FFD43FE8349A7" box="[199,342,700,725]" italics="true" pageId="9" pageNumber="468">Gnathosoma</emphasis>
: Axillary
</taxonomicName>
saccule present. Palpal eupathidium 
<emphasis id="FEE5EAFE6617FFAEFC78FD42FC7B49A6" box="[896,942,701,724]" italics="true" pageId="9" pageNumber="468">acm</emphasis>
attached to solenidion along distal threequarters of solenidial length (as for adult).
</paragraph>
<paragraph id="CC2E36EC6617FFAEFF3FFCFAFAAD48DE" blockId="9.[151,1437,151,1085]" pageId="9" pageNumber="468">
<emphasis id="FEE5EAFE6617FFAEFF3FFCFAFF29486E" box="[199,252,773,796]" italics="true" pageId="9" pageNumber="468">Legs</emphasis>
: Proral setae of tarsus I appear eupathidial. Subunguinal seta of tarsus I of normal form and inserted proximal to antelateral pair as in adult. Porose areas present on femora I to IV. Distal setae on tarsi I–IV blunt. Setae 
<emphasis id="FEE5EAFE6617FFAEFF24FCB3FF3F4817" box="[220,234,844,869]" italics="true" pageId="9" pageNumber="468">d</emphasis>
of femora I and II barbed, phylliform; seta 
<emphasis id="FEE5EAFE6617FFAEFD19FCB3FD3A4817" box="[737,751,844,869]" italics="true" pageId="9" pageNumber="468">d</emphasis>
of femur III narrow, phylliform; seta 
<emphasis id="FEE5EAFE6617FFAEFB59FCB3FB7A4817" box="[1185,1199,844,869]" italics="true" pageId="9" pageNumber="468">d</emphasis>
of femur IV barbed, tapered. Solenidion and coupled seta 
<emphasis id="FEE5EAFE6617FFAEFDC2FC8FFD9D48FB" box="[570,584,880,905]" italics="true" pageId="9" pageNumber="468">d</emphasis>
on tibia I on anterodorsal tubercle. Solenidion 
<emphasis id="FEE5EAFE6617FFAEFBA1FC8EFBBD48FA" box="[1113,1128,881,904]" italics="true" pageId="9" pageNumber="468">φ</emphasis>
on tibia II and III subequal in length to seta 
<emphasis id="FEE5EAFE6617FFAEFEA8FC6BFE8B48DF" box="[336,350,916,941]" italics="true" pageId="9" pageNumber="468">d</emphasis>
; solenidion 
<emphasis id="FEE5EAFE6617FFAEFE10FC6AFE2248DE" box="[488,503,917,940]" italics="true" pageId="9" pageNumber="468">φ</emphasis>
on tibia 
<date id="B82F102C6617FFAEFDA3FC6AFD5E48DF" box="[603,651,916,941]" pageId="9" pageNumber="468">III 4</date>
/5 length seta 
<emphasis id="FEE5EAFE6617FFAEFCD0FC6BFCE348DF" box="[808,822,916,941]" italics="true" pageId="9" pageNumber="468">d</emphasis>
. Solenidia 
<emphasis id="FEE5EAFE6617FFAEFC4CFC6AFC1448DF" box="[948,961,917,941]" italics="true" pageId="9" pageNumber="468">δ</emphasis>
on genua I, 
<date id="B82F102C6617FFAEFBB5FC6AFB1248DF" box="[1101,1223,916,941]" pageId="9" pageNumber="468">II and III 4</date>
/5 length seta 
<emphasis id="FEE5EAFE6617FFAEFA9BFC6BFAA448DF" box="[1379,1393,916,941]" italics="true" pageId="9" pageNumber="468">d</emphasis>
.
</paragraph>
<paragraph id="CC2E36EC6617FFAEFF3FFC47FB2548A3" blockId="9.[151,1437,151,1085]" box="[199,1264,952,977]" pageId="9" pageNumber="468">
Deutonymph. 
<collectingCountry id="B486767C6617FFAEFE9FFC46FE5348A3" box="[359,390,953,977]" name="American Samoa" pageId="9" pageNumber="468">As</collectingCountry>
for tritonymph, except overall size and length of setae proportionally smaller.
</paragraph>
<paragraph id="CC2E36EC6617FFAEFF3FFC23FB154F6B" blockId="9.[151,1437,151,1085]" pageId="9" pageNumber="468">
<emphasis id="FEE5EAFE6617FFAEFF3FFC23FEBE4887" bold="true" box="[199,363,988,1013]" pageId="9" pageNumber="468">Development.</emphasis>
Development of epimeral setae (deutonymph to adult): 3-1-2-2, 3-1-2-3, 3-1-2-3. Genital, aggenital, anal and adanal setae (deutonymph to adult): 3-5-6, 1-1-1, 0-2-2, 2-3-3, respectively.
</paragraph>
<paragraph id="CC2E36EC6617FFAEFF3FFBDBFC964F4F" blockId="9.[151,1437,151,1085]" box="[199,835,1060,1085]" pageId="9" pageNumber="468">
Development of leg setae and solenidia given in 
<tableCitation id="811303576617FFAEFD11FBDBFCE94F4F" box="[745,828,1060,1085]" captionStart="TABLE 1" captionStartId="9.[151,239,1132,1155]" captionTargetBox="[159,1399,1215,1903]" captionTargetPageId="9" captionText="TABLE 1. Setiform organs in deutonymph, tritonymph and adult Tectoribates alcescampestris sp. nov. Setae (Roman) and solenidia (Greek) are indicated; parentheses indicate pairs of setae." httpUri="http://table.plazi.org/id/98EE66646617FFAEFF6FFB93FC584FD1" pageId="9" pageNumber="468" tableUuid="98EE66646617FFAEFF6FFB93FC584FD1">Table 1</tableCitation>
.
</paragraph>
</subSubSection>
<caption id="98EE66646617FFAEFF6FFB93FC584FD1" ID-Table-UUID="98EE66646617FFAEFF6FFB93FC584FD1" httpUri="http://table.plazi.org/id/98EE66646617FFAEFF6FFB93FC584FD1" pageId="9" pageNumber="468" targetBox="[159,1399,1215,1903]" targetIsTable="true" targetPageId="9">
<paragraph id="CC2E36EC6617FFAEFF6FFB93FC584FD1" blockId="9.[151,1436,1132,1187]" pageId="9" pageNumber="468">
<emphasis id="FEE5EAFE6617FFAEFF6FFB93FEDF4FF1" bold="true" box="[151,266,1132,1155]" pageId="9" pageNumber="468">TABLE 1.</emphasis>
Setiform organs in deutonymph, tritonymph and adult 
<taxonomicName id="0B914D6F6617FFAEFC99FB93FB404FF1" box="[865,1173,1132,1155]" class="Arachnida" family="Oribatellidae" genus="Tectoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="9" pageNumber="468" phylum="Arthropoda" rank="species" species="alcescampestris" status="sp. nov.">
<emphasis id="FEE5EAFE6617FFAEFC99FB93FB404FF1" box="[865,1173,1132,1155]" italics="true" pageId="9" pageNumber="468">Tectoribates alcescampestris</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E5D657856617FFAEFB65FB93FB214FF1" box="[1181,1268,1132,1155]" pageId="9" pageNumber="468" rank="species">
<emphasis id="FEE5EAFE6617FFAEFB65FB93FB214FF1" bold="true" box="[1181,1268,1132,1155]" pageId="9" pageNumber="468">sp. nov.</emphasis>
</taxonomicNameLabel>
Setae (Roman) and solenidia (Greek) are indicated; parentheses indicate pairs of setae.
</paragraph>
</caption>
<paragraph id="CC2E36EC6617FFAEFE68FB40FBC44C1D" pageId="9" pageNumber="468">
<table id="BE91C44C66170058FF67FB40FAA24C1D" box="[159,1399,1215,1903]" gridcols="6" gridrows="17" pageId="9" pageNumber="468">
<tr id="72A134AE66170058FF67FB40FAA24FA7" box="[159,1399,1215,1237]" gridrow="0" pageId="9" pageNumber="468" rowspan-0="1">
<th id="31705DD266170058FE68FB40FDD54FA7" box="[400,512,1215,1237]" gridcol="1" gridrow="0" pageId="9" pageNumber="468">
<emphasis id="FEE5EAFE6617FFAEFE68FB40FDD54FA7" box="[400,512,1215,1237]" italics="true" pageId="9" pageNumber="468">Trochanter</emphasis>
</th>
<th id="31705DD266170058FDD5FB40FDAB4FA7" box="[557,638,1215,1237]" gridcol="2" gridrow="0" pageId="9" pageNumber="468">
<emphasis id="FEE5EAFE6617FFAEFDD5FB3FFDA44FA7" box="[557,625,1216,1237]" italics="true" pageId="9" pageNumber="468">Femur</emphasis>
</th>
<th id="31705DD266170058FD55FB40FD244FA7" box="[685,753,1215,1237]" gridcol="3" gridrow="0" pageId="9" pageNumber="468">
<emphasis id="FEE5EAFE6617FFAEFD55FB3FFD314FA7" box="[685,740,1216,1237]" italics="true" pageId="9" pageNumber="468">Genu</emphasis>
</th>
<th id="31705DD266170058FCA3FB40FC034FA7" box="[859,982,1215,1237]" gridcol="4" gridrow="0" pageId="9" pageNumber="468">
<emphasis id="FEE5EAFE6617FFAEFCA3FB40FC5A4FA7" box="[859,911,1215,1237]" italics="true" pageId="9" pageNumber="468">Tibia</emphasis>
</th>
<th id="31705DD266170058FBF0FB40FAA24FA7" box="[1032,1399,1215,1237]" gridcol="5" gridrow="0" pageId="9" pageNumber="468">
<emphasis id="FEE5EAFE6617FFAEFBF1FB3FFB994FA7" box="[1033,1100,1216,1237]" italics="true" pageId="9" pageNumber="468">Tarsus</emphasis>
</th>
</tr>
<tr id="72A134AE66170058FF67FB16FAA24F8C" box="[159,1399,1257,1278]" gridrow="1" pageId="9" pageNumber="468" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1">
<th id="31705DD266170058FF67FB16FEF24F8C" box="[159,295,1257,1278]" gridcol="0" gridrow="1" pageId="9" pageNumber="468">
<emphasis id="FEE5EAFE6617FFAEFF67FB16FF064F8C" box="[159,211,1257,1278]" italics="true" pageId="9" pageNumber="468">Leg I</emphasis>
</th>
</tr>
<tr id="72A134AE66170058FF67FAEEFAA24E59" box="[159,1399,1297,1323]" gridrow="2" pageId="9" pageNumber="468">
<th id="31705DD266170058FF67FAEEFEF24E59" box="[159,295,1297,1323]" gridcol="0" gridrow="2" pageId="9" pageNumber="468">
<emphasis id="FEE5EAFE6617FFAEFF5DFAEEFEF24E55" box="[165,295,1297,1319]" italics="true" pageId="9" pageNumber="468">Deutonymph</emphasis>
</th>
<td id="31705DD266170058FE68FAEEFDD54E59" box="[400,512,1297,1323]" gridcol="1" gridrow="2" pageId="9" pageNumber="468">
<emphasis id="FEE5EAFE6617FFAEFE68FAEDFE4D4E55" box="[400,408,1298,1319]" italics="true" pageId="9" pageNumber="468">-</emphasis>
</td>
<td id="31705DD266170058FDD5FAEEFDAB4E59" box="[557,638,1297,1323]" gridcol="2" gridrow="2" pageId="9" pageNumber="468">
<emphasis id="FEE5EAFE6617FFAEFDD5FAEEFDAB4E55" box="[557,638,1297,1319]" italics="true" pageId="9" pageNumber="468">d bv”(l)</emphasis>
</td>
<td id="31705DD266170058FD55FAEEFD244E59" box="[685,753,1297,1323]" gridcol="3" gridrow="2" pageId="9" pageNumber="468">
<emphasis id="FEE5EAFE6617FFAEFD55FAEEFD3F4E55" box="[685,746,1297,1319]" italics="true" pageId="9" pageNumber="468">d σ (l)</emphasis>
</td>
<td id="31705DD266170058FCA3FAEEFC034E59" box="[859,982,1297,1323]" gridcol="4" gridrow="2" pageId="9" pageNumber="468">
<emphasis id="FEE5EAFE6617FFAEFCA3FAEEFCA94E55" box="[859,892,1297,1319]" italics="true" pageId="9" pageNumber="468">d φ</emphasis>
<emphasis id="FEE5EAFE6617FFAEFC84FAE0FC564E59" box="[892,899,1311,1323]" italics="true" pageId="9" pageNumber="468">1</emphasis>
<emphasis id="FEE5EAFE6617FFAEFC71FAEEFC054E55" box="[905,976,1297,1319]" italics="true" pageId="9" pageNumber="468">(l) v’ φ</emphasis>
<emphasis id="FEE5EAFE6617FFAEFC37FAE0FC034E59" box="[975,982,1311,1323]" italics="true" pageId="9" pageNumber="468">2</emphasis>
</td>
<td id="31705DD266170058FBF0FAEEFAA24E59" box="[1032,1399,1297,1323]" gridcol="5" gridrow="2" pageId="9" pageNumber="468">
<emphasis id="FEE5EAFE6617FFAEFBF1FAEEFA854E55" box="[1033,1360,1297,1319]" italics="true" pageId="9" pageNumber="468">(ft) (tc) (p) (u) s (a) (pv) (pl) e ω</emphasis>
<emphasis id="FEE5EAFE6617FFAEFAA9FAE0FA8D4E59" box="[1361,1368,1311,1323]" italics="true" pageId="9" pageNumber="468">1</emphasis>
<emphasis id="FEE5EAFE6617FFAEFAA6FAEDFAA54E55" box="[1374,1392,1298,1319]" italics="true" pageId="9" pageNumber="468">ω</emphasis>
<emphasis id="FEE5EAFE6617FFAEFA88FAE0FAA24E59" box="[1392,1399,1311,1323]" italics="true" pageId="9" pageNumber="468">2</emphasis>
</td>
</tr>
<tr id="72A134AE66170058FF67FAC2FAA24E21" box="[159,1399,1341,1363]" gridrow="3" pageId="9" pageNumber="468">
<th id="31705DD266170058FF67FAC2FEF24E21" box="[159,295,1341,1363]" gridcol="0" gridrow="3" pageId="9" pageNumber="468">
<emphasis id="FEE5EAFE6617FFAEFF5DFAC2FECF4E21" box="[165,282,1341,1363]" italics="true" pageId="9" pageNumber="468">Tritonymph</emphasis>
</th>
<td id="31705DD266170058FE68FAC2FDD54E21" box="[400,512,1341,1363]" gridcol="1" gridrow="3" pageId="9" pageNumber="468">
<emphasis id="FEE5EAFE6617FFAEFE68FAC1FE714E21" box="[400,420,1342,1363]" italics="true" pageId="9" pageNumber="468">v’</emphasis>
</td>
<td id="31705DD266170058FDD5FAC2FDAB4E21" box="[557,638,1341,1363]" gridcol="2" gridrow="3" pageId="9" pageNumber="468">
<emphasis id="FEE5EAFE6617FFAEFDD5FAC1FD944E21" box="[557,577,1342,1363]" italics="true" pageId="9" pageNumber="468">v’</emphasis>
</td>
<td id="31705DD266170058FD55FAC2FD244E21" box="[685,753,1341,1363]" gridcol="3" gridrow="3" pageId="9" pageNumber="468">
<emphasis id="FEE5EAFE6617FFAEFD55FAC1FD144E21" box="[685,705,1342,1363]" italics="true" pageId="9" pageNumber="468">v’</emphasis>
</td>
<td id="31705DD266170058FCA3FAC2FC034E21" box="[859,982,1341,1363]" gridcol="4" gridrow="3" pageId="9" pageNumber="468">
<emphasis id="FEE5EAFE6617FFAEFCA3FAC1FCA14E21" box="[859,884,1342,1363]" italics="true" pageId="9" pageNumber="468">v”</emphasis>
</td>
<td id="31705DD266170058FBF0FAC2FAA24E21" box="[1032,1399,1341,1363]" gridcol="5" gridrow="3" pageId="9" pageNumber="468">
<emphasis id="FEE5EAFE6617FFAEFBF1FAC1FBF24E21" box="[1033,1063,1342,1363]" italics="true" pageId="9" pageNumber="468">(it)</emphasis>
</td>
</tr>
<tr id="72A134AE66170058FF67FA98FAA24E0F" box="[159,1399,1383,1405]" gridrow="4" pageId="9" pageNumber="468">
<th id="31705DD266170058FF67FA98FEF24E0F" box="[159,295,1383,1405]" gridcol="0" gridrow="4" pageId="9" pageNumber="468">
<emphasis id="FEE5EAFE6617FFAEFF5DFA98FF094E0F" box="[165,220,1383,1405]" italics="true" pageId="9" pageNumber="468">Adult</emphasis>
</th>
<td id="31705DD266170058FE68FA98FDD54E0F" box="[400,512,1383,1405]" gridcol="1" gridrow="4" pageId="9" pageNumber="468">
<emphasis id="FEE5EAFE6617FFAEFE68FA97FE4D4E0F" box="[400,408,1384,1405]" italics="true" pageId="9" pageNumber="468">-</emphasis>
</td>
<td id="31705DD266170058FDD5FA98FDAB4E0F" box="[557,638,1383,1405]" gridcol="2" gridrow="4" pageId="9" pageNumber="468">
<emphasis id="FEE5EAFE6617FFAEFDD5FA97FD944E0F" box="[557,577,1384,1405]" italics="true" pageId="9" pageNumber="468">v’</emphasis>
</td>
<td id="31705DD266170058FD55FA98FD244E0F" box="[685,753,1383,1405]" gridcol="3" gridrow="4" pageId="9" pageNumber="468">
<emphasis id="FEE5EAFE6617FFAEFD55FA98FD1A4E0F" box="[685,719,1383,1405]" italics="true" pageId="9" pageNumber="468">[d]</emphasis>
</td>
<td id="31705DD266170058FCA3FA98FC034E0F" box="[859,982,1383,1405]" gridcol="4" gridrow="4" pageId="9" pageNumber="468">
<emphasis id="FEE5EAFE6617FFAEFCA3FA98FCA94E0F" box="[859,892,1383,1405]" italics="true" pageId="9" pageNumber="468">[d]</emphasis>
</td>
<td id="31705DD266170058FBF0FA98FAA24E0F" box="[1032,1399,1383,1405]" gridcol="5" gridrow="4" pageId="9" pageNumber="468">
<emphasis id="FEE5EAFE6617FFAEFBF1FA97FBC84E0F" box="[1033,1053,1384,1405]" italics="true" pageId="9" pageNumber="468">v’</emphasis>
</td>
</tr>
<tr id="72A134AE66170058FF67FA6EFAA24ED4" box="[159,1399,1425,1446]" gridrow="5" pageId="9" pageNumber="468" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1">
<th id="31705DD266170058FF67FA6EFEF24ED4" box="[159,295,1425,1446]" gridcol="0" gridrow="5" pageId="9" pageNumber="468">
<emphasis id="FEE5EAFE6617FFAEFF67FA6EFF094ED4" box="[159,220,1425,1446]" italics="true" pageId="9" pageNumber="468">Leg II</emphasis>
</th>
</tr>
<tr id="72A134AE66170058FF67FA46FAA24EA1" box="[159,1399,1465,1491]" gridrow="6" pageId="9" pageNumber="468">
<th id="31705DD266170058FF67FA46FEF24EA1" box="[159,295,1465,1491]" gridcol="0" gridrow="6" pageId="9" pageNumber="468">
<emphasis id="FEE5EAFE6617FFAEFF5DFA46FEF24EBD" box="[165,295,1465,1487]" italics="true" pageId="9" pageNumber="468">Deutonymph</emphasis>
</th>
<td id="31705DD266170058FE68FA46FDD54EA1" box="[400,512,1465,1491]" gridcol="1" gridrow="6" pageId="9" pageNumber="468">
<emphasis id="FEE5EAFE6617FFAEFE68FA45FE4D4EBD" box="[400,408,1466,1487]" italics="true" pageId="9" pageNumber="468">-</emphasis>
</td>
<td id="31705DD266170058FDD5FA46FDAB4EA1" box="[557,638,1465,1491]" gridcol="2" gridrow="6" pageId="9" pageNumber="468">
<emphasis id="FEE5EAFE6617FFAEFDD5FA46FDAE4EBD" box="[557,635,1465,1487]" italics="true" pageId="9" pageNumber="468">d bv” l’</emphasis>
</td>
<td id="31705DD266170058FD55FA46FD244EA1" box="[685,753,1465,1491]" gridcol="3" gridrow="6" pageId="9" pageNumber="468">
<emphasis id="FEE5EAFE6617FFAEFD55FA46FD3F4EBD" box="[685,746,1465,1487]" italics="true" pageId="9" pageNumber="468">d σ (l)</emphasis>
</td>
<td id="31705DD266170058FCA3FA46FC034EA1" box="[859,982,1465,1491]" gridcol="4" gridrow="6" pageId="9" pageNumber="468">
<emphasis id="FEE5EAFE6617FFAEFCA3FA46FC784EBD" box="[859,941,1465,1487]" italics="true" pageId="9" pageNumber="468">d φ (l)v’</emphasis>
</td>
<td id="31705DD266170058FBF0FA46FAA24EA1" box="[1032,1399,1465,1491]" gridcol="5" gridrow="6" pageId="9" pageNumber="468">
<emphasis id="FEE5EAFE6617FFAEFBF0FA45FAC14EBD" box="[1032,1300,1465,1487]" italics="true" pageId="9" pageNumber="468">(p) (tc) (ft) (u) s (a) (pv) ω</emphasis>
<emphasis id="FEE5EAFE6617FFAEFAECFA38FACE4EA1" box="[1300,1307,1479,1491]" italics="true" pageId="9" pageNumber="468">1</emphasis>
<emphasis id="FEE5EAFE6617FFAEFAE6FA45FAE54EBD" box="[1310,1328,1466,1487]" italics="true" pageId="9" pageNumber="468">ω</emphasis>
<emphasis id="FEE5EAFE6617FFAEFAC8FA38FAE24EA1" box="[1328,1335,1479,1491]" italics="true" pageId="9" pageNumber="468">2</emphasis>
</td>
</tr>
<tr id="72A134AE66170058FF67FA1AFAA24E89" box="[159,1399,1509,1531]" gridrow="7" pageId="9" pageNumber="468">
<th id="31705DD266170058FF67FA1AFEF24E89" box="[159,295,1509,1531]" gridcol="0" gridrow="7" pageId="9" pageNumber="468">Tritonymph</th>
<td id="31705DD266170058FE68FA1AFDD54E89" box="[400,512,1509,1531]" gridcol="1" gridrow="7" pageId="9" pageNumber="468">v’</td>
<td id="31705DD266170058FDD5FA1AFDAB4E89" box="[557,638,1509,1531]" gridcol="2" gridrow="7" pageId="9" pageNumber="468">l”</td>
<td id="31705DD266170058FD55FA1AFD244E89" box="[685,753,1509,1531]" gridcol="3" gridrow="7" pageId="9" pageNumber="468">v’</td>
<td id="31705DD266170058FCA3FA1AFC034E89" box="[859,982,1509,1531]" gridcol="4" gridrow="7" pageId="9" pageNumber="468">v”</td>
<td id="31705DD266170058FBF0FA1AFAA24E89" box="[1032,1399,1509,1531]" gridcol="5" gridrow="7" pageId="9" pageNumber="468">(it)</td>
</tr>
<tr id="72A134AE66170058FF67F9F1FAA24D57" box="[159,1399,1550,1573]" gridrow="8" pageId="9" pageNumber="468" rowspan-5="1">
<th id="31705DD266170058FF67F9F1FEF24D57" box="[159,295,1550,1573]" gridcol="0" gridrow="8" pageId="9" pageNumber="468">Adult</th>
<td id="31705DD266170058FE68F9F1FDD54D57" box="[400,512,1550,1573]" gridcol="1" gridrow="8" pageId="9" pageNumber="468">-</td>
<td id="31705DD266170058FDD5F9F1FDAB4D57" box="[557,638,1550,1573]" gridcol="2" gridrow="8" pageId="9" pageNumber="468">v’</td>
<td id="31705DD266170058FD55F9F1FD244D57" box="[685,753,1550,1573]" gridcol="3" gridrow="8" pageId="9" pageNumber="468">[d]</td>
<td id="31705DD266170058FCA3F9F1FC034D57" box="[859,982,1550,1573]" gridcol="4" gridrow="8" pageId="9" pageNumber="468">[d]</td>
</tr>
<tr id="72A134AE66170058FF67F9C6FAA24D3C" box="[159,1399,1593,1614]" gridrow="9" pageId="9" pageNumber="468" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1">
<th id="31705DD266170058FF67F9C6FEF24D3C" box="[159,295,1593,1614]" gridcol="0" gridrow="9" pageId="9" pageNumber="468">Leg III</th>
</tr>
<tr id="72A134AE66170058FF67F99EFAA24D05" box="[159,1399,1633,1655]" gridrow="10" pageId="9" pageNumber="468">
<th id="31705DD266170058FF67F99EFEF24D05" box="[159,295,1633,1655]" gridcol="0" gridrow="10" pageId="9" pageNumber="468">Deutonymph</th>
<td id="31705DD266170058FE68F99EFDD54D05" box="[400,512,1633,1655]" gridcol="1" gridrow="10" pageId="9" pageNumber="468">v’</td>
<td id="31705DD266170058FDD5F99EFDAB4D05" box="[557,638,1633,1655]" gridcol="2" gridrow="10" pageId="9" pageNumber="468">d ev”l”</td>
<td id="31705DD266170058FD55F99EFD244D05" box="[685,753,1633,1655]" gridcol="3" gridrow="10" pageId="9" pageNumber="468">d σ l’ -</td>
<td id="31705DD266170058FCA3F99EFC034D05" box="[859,982,1633,1655]" gridcol="4" gridrow="10" pageId="9" pageNumber="468">d φ v’</td>
<td id="31705DD266170058FBF0F99EFAA24D05" box="[1032,1399,1633,1655]" gridcol="5" gridrow="10" pageId="9" pageNumber="468">(p) (tc) (ft) (u) s (a) (pv)</td>
</tr>
<tr id="72A134AE66170058FF67F974FAA24DD3" box="[159,1399,1675,1697]" gridrow="11" pageId="9" pageNumber="468">
<th id="31705DD266170058FF67F974FEF24DD3" box="[159,295,1675,1697]" gridcol="0" gridrow="11" pageId="9" pageNumber="468">Tritonymph</th>
<td id="31705DD266170058FE68F974FDD54DD3" box="[400,512,1675,1697]" gridcol="1" gridrow="11" pageId="9" pageNumber="468">l’</td>
<td id="31705DD266170058FDD5F974FDAB4DD3" box="[557,638,1675,1697]" gridcol="2" gridrow="11" pageId="9" pageNumber="468">-</td>
<td id="31705DD266170058FD55F974FD244DD3" box="[685,753,1675,1697]" gridcol="3" gridrow="11" pageId="9" pageNumber="468">-</td>
<td id="31705DD266170058FCA3F974FC034DD3" box="[859,982,1675,1697]" gridcol="4" gridrow="11" pageId="9" pageNumber="468">(l)</td>
<td id="31705DD266170058FBF0F974FAA24DD3" box="[1032,1399,1675,1697]" gridcol="5" gridrow="11" pageId="9" pageNumber="468">(it)</td>
</tr>
<tr id="72A134AE66170058FF67F94BFAA24DB8" box="[159,1399,1716,1738]" gridrow="12" pageId="9" pageNumber="468">
<th id="31705DD266170058FF67F94BFEF24DB8" box="[159,295,1716,1738]" gridcol="0" gridrow="12" pageId="9" pageNumber="468">Adult</th>
<td id="31705DD266170058FE68F94BFDD54DB8" box="[400,512,1716,1738]" gridcol="1" gridrow="12" pageId="9" pageNumber="468">-</td>
<td id="31705DD266170058FDD5F94BFDAB4DB8" box="[557,638,1716,1738]" gridcol="2" gridrow="12" pageId="9" pageNumber="468">-</td>
<td id="31705DD266170058FD55F94BFD244DB8" box="[685,753,1716,1738]" gridcol="3" gridrow="12" pageId="9" pageNumber="468">[d]</td>
<td id="31705DD266170058FCA3F94BFC034DB8" box="[859,982,1716,1738]" gridcol="4" gridrow="12" pageId="9" pageNumber="468">[d]</td>
<td id="31705DD266170058FBF0F94BFAA24DB8" box="[1032,1399,1716,1738]" gridcol="5" gridrow="12" pageId="9" pageNumber="468">-</td>
</tr>
<tr id="72A134AE66170058FF67F921FAA24D81" box="[159,1399,1758,1779]" gridrow="13" pageId="9" pageNumber="468" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1">
<th id="31705DD266170058FF67F921FEF24D81" box="[159,295,1758,1779]" gridcol="0" gridrow="13" pageId="9" pageNumber="468">Leg IV</th>
</tr>
<tr id="72A134AE66170058FF67F8F9FAA24C6F" box="[159,1399,1798,1821]" gridrow="14" pageId="9" pageNumber="468">
<th id="31705DD266170058FF67F8F9FEF24C6F" box="[159,295,1798,1821]" gridcol="0" gridrow="14" pageId="9" pageNumber="468">Deutonymph</th>
<td id="31705DD266170058FE68F8F9FDD54C6F" box="[400,512,1798,1821]" gridcol="1" gridrow="14" pageId="9" pageNumber="468">-</td>
<td id="31705DD266170058FDD5F8F9FDAB4C6F" box="[557,638,1798,1821]" gridcol="2" gridrow="14" pageId="9" pageNumber="468">d ev”</td>
<td id="31705DD266170058FD55F8F9FD244C6F" box="[685,753,1798,1821]" gridcol="3" gridrow="14" pageId="9" pageNumber="468">d’ l”</td>
<td id="31705DD266170058FCA3F8F9FC034C6F" box="[859,982,1798,1821]" gridcol="4" gridrow="14" pageId="9" pageNumber="468">d φ v’</td>
<td id="31705DD266170058FBF0F8F9FAA24C6F" box="[1032,1399,1798,1821]" gridcol="5" gridrow="14" pageId="9" pageNumber="468">(p) ft” (u) (pv) (tc) (a) s</td>
</tr>
<tr id="72A134AE66170058FF67F8CFFAA24C34" box="[159,1399,1840,1862]" gridrow="15" pageId="9" pageNumber="468" rowspan-3="1">
<th id="31705DD266170058FF67F8CFFEF24C34" box="[159,295,1840,1862]" gridcol="0" gridrow="15" pageId="9" pageNumber="468">Tritonymph</th>
<td id="31705DD266170058FE68F8CFFDD54C34" box="[400,512,1840,1862]" gridcol="1" gridrow="15" pageId="9" pageNumber="468">v’</td>
<td id="31705DD266170058FDD5F8CFFDAB4C34" box="[557,638,1840,1862]" gridcol="2" gridrow="15" pageId="9" pageNumber="468">-</td>
<td id="31705DD266170058FCA3F8CFFC034C34" box="[859,982,1840,1862]" gridcol="4" gridrow="15" pageId="9" pageNumber="468">l”</td>
<td id="31705DD266170058FBF0F8CFFAA24C34" box="[1032,1399,1840,1862]" gridcol="5" gridrow="15" pageId="9" pageNumber="468">-</td>
</tr>
<tr id="72A134AE66170058FF67F8A6FAA24C1D" box="[159,1399,1881,1903]" gridrow="16" pageId="9" pageNumber="468">
<th id="31705DD266170058FF67F8A6FEF24C1D" box="[159,295,1881,1903]" gridcol="0" gridrow="16" pageId="9" pageNumber="468">Adult</th>
<td id="31705DD266170058FE68F8A6FDD54C1D" box="[400,512,1881,1903]" gridcol="1" gridrow="16" pageId="9" pageNumber="468">-</td>
<td id="31705DD266170058FDD5F8A6FDAB4C1D" box="[557,638,1881,1903]" gridcol="2" gridrow="16" pageId="9" pageNumber="468">-</td>
<td id="31705DD266170058FD55F8A6FD244C1D" box="[685,753,1881,1903]" gridcol="3" gridrow="16" pageId="9" pageNumber="468">-</td>
<td id="31705DD266170058FCA3F8A6FC034C1D" box="[859,982,1881,1903]" gridcol="4" gridrow="16" pageId="9" pageNumber="468">l’ [d]</td>
<td id="31705DD266170058FBF0F8A6FAA24C1D" box="[1032,1399,1881,1903]" gridcol="5" gridrow="16" pageId="9" pageNumber="468">-</td>
</tr>
</table>
</paragraph>
<subSubSection id="848B65676617FFAEFF3FF84BFAFF4CBE" box="[199,1322,1972,1997]" pageId="9" pageNumber="468" type="etymology">
<paragraph id="CC2E36EC6617FFAEFF3FF84BFAFF4CBE" blockId="9.[199,1435,1972,2033]" box="[199,1322,1972,1997]" pageId="9" pageNumber="468">
<emphasis id="FEE5EAFE6617FFAEFF3FF84BFE9B4CBF" bold="true" box="[199,334,1972,1997]" pageId="9" pageNumber="468">Etymology.</emphasis>
The specific epithet, 
<taxonomicName id="0B914D6F6617FFAEFDC7F84BFD274CBF" box="[575,754,1972,1997]" class="Arachnida" family="Oribatellidae" genus="Tectoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="9" pageNumber="468" phylum="Arthropoda" rank="species" species="alcescampestris">
<emphasis id="FEE5EAFE6617FFAEFDC7F84BFD274CBF" box="[575,754,1972,1997]" italics="true" pageId="9" pageNumber="468">alcescampestris</emphasis>
</taxonomicName>
, is the Latin for the 
<typeStatus id="132A884E6617FFAEFC2EF84AFBD34CBF" box="[982,1030,1973,1997]" pageId="9" pageNumber="468">type</typeStatus>
locality – Moose Pasture.
</paragraph>
</subSubSection>
<subSubSection id="848B65676617FFADFF3FF827FBB84A16" lastPageId="10" lastPageNumber="469" pageId="9" pageNumber="468" type="discussion">
<paragraph id="CC2E36EC6617FFADFF3FF827FD4B4BA7" blockId="9.[199,1435,1972,2033]" lastBlockId="10.[151,1436,151,357]" lastPageId="10" lastPageNumber="469" pageId="9" pageNumber="468">
<emphasis id="FEE5EAFE6617FFAEFF3FF827FEE14C83" bold="true" box="[199,308,2008,2033]" pageId="9" pageNumber="468">Remarks</emphasis>
. This species is unique among known 
<taxonomicName id="0B914D6F6617FFAEFD19F827FCBF4C83" box="[737,874,2008,2033]" class="Arachnida" family="Oribatellidae" genus="Tectoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="9" pageNumber="468" phylum="Arthropoda" rank="genus">
<emphasis id="FEE5EAFE6617FFAEFD19F827FCBF4C83" box="[737,874,2008,2033]" italics="true" pageId="9" pageNumber="468">Tectoribates</emphasis>
</taxonomicName>
in lacking the hexagonal pattern on the anterior of the notogaster in the region commonly occupied by a lenticulus in brachypyline taxa. Whether this absence is a plesiomorphy or a secondary loss is unknown.
</paragraph>
<paragraph id="CC2E36EC6614FFADFF3FFF20FBB84A16" blockId="10.[151,1436,151,357]" pageId="10" pageNumber="469">
<taxonomicName id="0B914D6F6614FFADFF3FFF20FDC74B8A" box="[199,530,223,248]" class="Arachnida" family="Oribatellidae" genus="Tectoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="10" pageNumber="469" phylum="Arthropoda" rank="species" species="alcescampestris">
<emphasis id="FEE5EAFE6614FFADFF3FFF20FDC74B8A" box="[199,530,223,248]" italics="true" pageId="10" pageNumber="469">Tectoribates alcescampestris</emphasis>
</taxonomicName>
has been collected primarily in aspen parkland in forest litter and fungal sporocarps and in other open sites covered by grasses and sedges. Gravid females carry up to 2 large eggs. Gut contents are primarily lightly pigmented fungal hyphae and spores. The association of immatures with adults was addressed under the Discussion following the generic diagnosis (See above, Remark 4).
</paragraph>
</subSubSection>
</treatment>
</document>