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<document id="16EF4785DFE2A2D119834268DE31B7E2" ID-CLB-Dataset="5025" ID-DOI="10.1649/0010-065X-72.3.503" ID-GBIF-Dataset="a3f528b0-9dd1-4059-9e1f-0a88c23f1e46" ID-ISSN="1938-4394" ID-Zenodo-Dep="4789344" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1621946641600" checkinUser="carolina" docAuthor="Etzler, Frank E. &amp; Johnson, Paul J." docDate="2018" docId="2E1004476013540BFD4D06788E3BFA4E" docLanguage="en" docName="TheColeopBull.72.3.503-521.pdf" docOrigin="The Coleopterists Bulletin 72 (3)" docSource="http://dx.doi.org/10.1649/0010-065x-72.3.503" docStyle="DocumentStyle:6D94AA43905A5ECAD3A4A280834A9134.5:TheColeopBull.2017-.journal_article.0cover" docStyleId="6D94AA43905A5ECAD3A4A280834A9134" docStyleName="TheColeopBull.2017-.journal_article.0cover" docStyleVersion="5" docTitle="Athoplastus rainieri Etzler &amp; Johnson 2018, new combination" docType="treatment" docVersion="8" lastPageNumber="518" masterDocId="D2297C3F601D5404FFBE057D8C77FFA2" masterDocTitle="Athoplastus Johnson and Etzler (Coleoptera: Elateridae: Dendrometrinae), a New Genus of Click Beetle from the Northwestern Continental USA" masterLastPageNumber="521" masterPageNumber="503" pageNumber="517" updateTime="1698990595291" updateUser="ExternalLinkService">
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<mods:title id="219BBC5A68FBF07FBAA73877EF6A801C">Athoplastus Johnson and Etzler (Coleoptera: Elateridae: Dendrometrinae), a New Genus of Click Beetle from the Northwestern Continental USA</mods:title>
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<mods:namePart id="649DBFBDF15D8FC2BA0FBE49D85E3E89">Etzler, Frank E.</mods:namePart>
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<mods:title id="47FB1417AB0C8839D8D05E7E8B7507AD">The Coleopterists Bulletin</mods:title>
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<mods:date id="F193375C865E5863A8012545A62E298B">2018</mods:date>
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<treatment id="2E1004476013540BFD4D06788E3BFA4E" ID-DOI="http://doi.org/10.5281/zenodo.5189242" ID-GBIF-Taxon="184010383" ID-Zenodo-Dep="5189242" LSID="urn:lsid:plazi:treatment:2E1004476013540BFD4D06788E3BFA4E" httpUri="http://treatment.plazi.org/id/2E1004476013540BFD4D06788E3BFA4E" lastPageId="15" lastPageNumber="518" pageId="14" pageNumber="517">
<subSubSection id="EEA3E6DA6013540AFD4D06788832FCF2" pageId="14" pageNumber="517" type="nomenclature">
<paragraph id="A606B5516013540AFD4D0678887AFC97" blockId="14.[755,1145,773,848]" pageId="14" pageNumber="517">
<heading id="FD4E023D6013540AFD4D0678887AFC97" centered="true" fontSize="9" level="2" pageId="14" pageNumber="517" reason="2">
<emphasis id="94CD69436013540AFD4D0678887AFC97" bold="true" pageId="14" pageNumber="517">
<taxonomicName id="61B9CED26013540AFD4D06788803FCB8" authority="(Van Dyke, 1932)" authorityName="Etzler &amp; Johnson" authorityYear="2018" baseAuthorityName="Van Dyke" baseAuthorityYear="1932" box="[755,1140,773,794]" class="Insecta" family="Elateridae" genus="Athoplastus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="517" phylum="Arthropoda" rank="species" species="rainieri" status="comb. nov.">
<emphasis id="94CD69436013540AFD4D06788FCEFCB8" bold="true" box="[755,953,773,794]" italics="true" pageId="14" pageNumber="517">Athoplastus rainieri</emphasis>
(
<bibRefCitation id="C228C8A06013540AFC760678881AFCB8" author="Van Dyke, E. C." box="[968,1133,773,794]" pageId="14" pageNumber="517" pagination="291 - 465" refId="ref12676" refString="Van Dyke, E. C. 1932. Miscellaneous studies in the Elateridae and related families of Coleoptera. Proceedings of the California Academy of Science 20: 291 - 465." type="journal article" year="1932">Van Dyke, 1932</bibRefCitation>
)
</taxonomicName>
,
<taxonomicNameLabel id="8FFED4386013540AFCE0065D887AFC97" box="[862,1037,800,821]" pageId="14" pageNumber="517" rank="species">new combination</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="A606B5516013540AFC9906468832FCF2" blockId="14.[755,1145,773,848]" box="[807,1093,826,848]" pageId="14" pageNumber="517">
(Adult:
<figureCitation id="3E82A9D46013540AFCCC06468FC0FCF2" box="[882,951,827,848]" captionStart="Figs" captionStartId="4.[197,236,1615,1634]" captionTargetBox="[245,1120,196,1597]" captionTargetId="figure-0@4.[243,1122,194,1599]" captionTargetPageId="4" captionText="Figs. 14. Athoplastus species, dorsal habitus. 1) A. acutus; 2) A. palouse; 3) A. lochsa; 4) A. rainieri." figureDoi="http://doi.org/10.5281/zenodo.4789354" httpUri="https://zenodo.org/record/4789354/files/figure.png" pageId="14" pageNumber="517">Figs. 4</figureCitation>
,
<figureCitation id="3E82A9D46013540AFC7C06468FB8FCF2" box="[962,975,827,848]" captionStart="Figs" captionStartId="5.[175,214,1615,1634]" captionTargetBox="[206,1118,195,1597]" captionTargetId="figure-0@5.[204,1120,194,1599]" captionTargetPageId="5" captionText="Figs. 59. Athoplastus species. 5) A. acutus, female dorsal habitus; 6) A. acutus, male dorsal habitus, darker coloration; 7) A. acutus, male ventral habitus; 8) A. rainier, prothorax, lateral view (arrow indicates start of area between procoxae); 9) A. acutus, prothorax, lateral view (arrow indicates start of expanded area between procoxae)." figureDoi="http://doi.org/10.5281/zenodo.4789370" httpUri="https://zenodo.org/record/4789370/files/figure.png" pageId="14" pageNumber="517">8</figureCitation>
,
<figureCitation id="3E82A9D46013540AFC620646886DFCF2" box="[988,1050,826,848]" captionStart="Figs" captionStartId="7.[175,214,1615,1634]" captionTargetBox="[246,1077,201,1597]" captionTargetId="figure-1@7.[244,1080,194,1599]" captionTargetPageId="7" captionText="Figs. 1627. Athoplastus species, adeagi, dorsal view, ventral view, and close up of paramere apex, respectively." figureDoi="http://doi.org/10.5281/zenodo.4789394" httpUri="https://zenodo.org/record/4789394/files/figure.png" pageId="14" pageNumber="517">2224</figureCitation>
,
<figureCitation id="3E82A9D46013540AFB9B06468849FCF2" box="[1061,1086,827,848]" captionStart="Figs" captionStartId="9.[175,214,1615,1634]" captionTargetBox="[219,1104,195,1597]" captionTargetId="figure-1@9.[217,1106,194,1599]" captionTargetPageId="9" captionText="Figs. 2830. Athoplastus species. 28) A. rainier, female, internal genitalia, v = vagina, u = uterus, acgld = accessory gland, bc = bursa copulatrix (after Han et al. 2012); 29) Distribution map of examined specimens, with rivers; 30) Neighbor-joining tree of Montana populations of A. acutus, COI mtDNA, 500 runs, scale bar = 1% difference." figureDoi="http://doi.org/10.5281/zenodo.4789442" httpUri="https://zenodo.org/record/4789442/files/figure.png" pageId="14" pageNumber="517">28</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="EEA3E6DA6013540AFD7D06198F4DFC0D" pageId="14" pageNumber="517" type="reference_group">
<paragraph id="A606B5516013540AFD7D0619882AFCD8" blockId="14.[707,1193,868,943]" box="[707,1117,868,890]" pageId="14" pageNumber="517">
<treatmentCitationGroup id="86A9927F6013540AFD7D0619882AFCD8" box="[707,1117,868,890]" pageId="14" pageNumber="517">
<treatmentCitation id="271893406013540AFD7D06198850FCD8" author="Van Dyke, E. C." box="[707,1063,868,890]" page="397" pageId="14" pageNumber="517" year="1932">
<taxonomicName id="61B9CED26013540AFD7D06198850FCD8" authority="Van Dyke 1932: 397" authorityName="Van Dyke" authorityPageNumber="397" authorityYear="1932" box="[707,1063,868,890]" class="Insecta" family="Elateridae" genus="Ludius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="517" phylum="Arthropoda" rank="species" species="rainieri">
<emphasis id="94CD69436013540AFD7D06198F24FCD8" box="[707,851,868,890]" italics="true" pageId="14" pageNumber="517">Ludius rainieri</emphasis>
<bibRefCitation id="C228C8A06013540AFCE506188850FCD8" author="Van Dyke, E. C." box="[859,1063,869,890]" pageId="14" pageNumber="517" pagination="291 - 465" refId="ref12676" refString="Van Dyke, E. C. 1932. Miscellaneous studies in the Elateridae and related families of Coleoptera. Proceedings of the California Academy of Science 20: 291 - 465." type="journal article" year="1932">Van Dyke 1932: 397</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, 421.
</treatmentCitationGroup>
</paragraph>
<paragraph id="A606B5516013540AFD7D06028F4DFC0D" blockId="14.[707,1193,868,943]" pageId="14" pageNumber="517">
<taxonomicName id="61B9CED26013540AFD7D06028817FC37" authority="(Van Dyke, 1932)" baseAuthorityName="Van Dyke" baseAuthorityYear="1932" box="[707,1120,895,917]" class="Insecta" family="Elateridae" genus="Ctenicera" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="517" phylum="Arthropoda" rank="species" species="rainieri">
<emphasis id="94CD69436013540AFD7D06028F09FC37" box="[707,894,895,917]" italics="true" pageId="14" pageNumber="517">Ctenicera rainieri</emphasis>
(
<bibRefCitation id="C228C8A06013540AFC2406FD882EFC37" author="Van Dyke, E. C." box="[922,1113,896,917]" pageId="14" pageNumber="517" pagination="291 - 465" refId="ref12676" refString="Van Dyke, E. C. 1932. Miscellaneous studies in the Elateridae and related families of Coleoptera. Proceedings of the California Academy of Science 20: 291 - 465." type="journal article" year="1932">Van Dyke, 1932</bibRefCitation>
)
</taxonomicName>
;
<bibRefCitation id="C228C8A06013540AFBC406FD88DEFC37" author="Lane, M. C." box="[1146,1193,896,917]" pageId="14" pageNumber="517" pagination="6 - 34" refId="ref12170" refString="Lane, M. C. 1971. Family Elateridae [pp. 6 - 34, excluding Cardiophorinae]. In: The Beetles of the Pacific Northwest, Part V: Rhipiceroida, Sternoxi, Phytophaga, Rynchophora, and Lamellicornia (M. H. Hatch, editor). University of Washington Press, Seattle, WA." type="book chapter" year="1971">Lane</bibRefCitation>
1971: 32.
</paragraph>
</subSubSection>
<subSubSection id="EEA3E6DA6013540AFD6506B88F97FB6B" pageId="14" pageNumber="517" type="diagnosis">
<paragraph id="A606B5516013540AFD6506B88F97FB6B" blockId="14.[707,1194,965,1730]" pageId="14" pageNumber="517">
<emphasis id="94CD69436013540AFD6506B88F35FC78" bold="true" box="[731,834,965,986]" pageId="14" pageNumber="517">Diagnosis.</emphasis>
This species is distinguished from all other species of the genus by the combination of close pronotal punctures (separated by less than one diameter of a puncture) (
<figureCitation id="3E82A9D46013540AFC1801688F9CFB88" box="[934,1003,1045,1066]" captionStart="Figs" captionStartId="6.[197,236,1734,1753]" captionTargetBox="[177,1188,196,1716]" captionTargetId="figure-0@6.[175,1190,194,1718]" captionTargetPageId="6" captionText="Figs. 1015. Athoplastus species. 10) A. acutus, prothorax, dorsal view; 11) A. lochsa, prothorax, dorsal view; 12) A. acutus, left antenna; 13) A. lochsa, left antenna; 14) A. palouse, left antenna; 15) A. acutus, female, left wing. Scale bars for antennae = 1 mm; for wing = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4789388" httpUri="https://zenodo.org/record/4789388/files/figure.png" pageId="14" pageNumber="517">Fig. 10</figureCitation>
), sloped prosternum (
<figureCitation id="3E82A9D46013540AFD7401528F73FBE6" box="[714,772,1071,1092]" captionStart="Figs" captionStartId="5.[175,214,1615,1634]" captionTargetBox="[206,1118,195,1597]" captionTargetId="figure-0@5.[204,1120,194,1599]" captionTargetPageId="5" captionText="Figs. 59. Athoplastus species. 5) A. acutus, female dorsal habitus; 6) A. acutus, male dorsal habitus, darker coloration; 7) A. acutus, male ventral habitus; 8) A. rainier, prothorax, lateral view (arrow indicates start of area between procoxae); 9) A. acutus, prothorax, lateral view (arrow indicates start of expanded area between procoxae)." figureDoi="http://doi.org/10.5281/zenodo.4789370" httpUri="https://zenodo.org/record/4789370/files/figure.png" pageId="14" pageNumber="517">Fig. 8</figureCitation>
), and dark markings at each puncture along the striae of the elytra (
<figureCitation id="3E82A9D46013540AFC2201378FA1FBFD" box="[924,982,1098,1119]" captionStart="Figs" captionStartId="4.[197,236,1615,1634]" captionTargetBox="[245,1120,196,1597]" captionTargetId="figure-0@4.[243,1122,194,1599]" captionTargetPageId="4" captionText="Figs. 14. Athoplastus species, dorsal habitus. 1) A. acutus; 2) A. palouse; 3) A. lochsa; 4) A. rainieri." figureDoi="http://doi.org/10.5281/zenodo.4789354" httpUri="https://zenodo.org/record/4789354/files/figure.png" pageId="14" pageNumber="517">Fig. 4</figureCitation>
). This last character is likely not a good species-level character and may be related to specimen development.
<taxonomicName id="61B9CED26013540AFB8701038F7CFB0C" authorityName="Etzler &amp; Johnson" authorityYear="2018" baseAuthorityName="Van Dyke" baseAuthorityYear="1932" class="Insecta" family="Elateridae" genus="Athoplastus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="517" phylum="Arthropoda" rank="species" species="rainieri">
<emphasis id="94CD69436013540AFB8701038F7CFB0C" italics="true" pageId="14" pageNumber="517">Athoplastus rainieri</emphasis>
</taxonomicName>
is quite similar to
<taxonomicName id="61B9CED26013540AFC0501E48857FB0D" authorityName="Etzler and Johnson" authorityYear="2018" box="[955,1056,1177,1199]" class="Insecta" family="Elateridae" genus="Athoplastus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="517" phylum="Arthropoda" rank="species" species="palouse">
<emphasis id="94CD69436013540AFC0501E48FBEFB0D" box="[955,969,1177,1199]" italics="true" pageId="14" pageNumber="517">A</emphasis>
.
<emphasis id="94CD69436013540AFC6801E48857FB0D" box="[982,1056,1177,1199]" italics="true" pageId="14" pageNumber="517">palouse</emphasis>
</taxonomicName>
, which can be distinguished as noted above.
</paragraph>
</subSubSection>
<subSubSection id="EEA3E6DA6013540BFD6501B38D7BFB5F" lastPageId="15" lastPageNumber="518" pageId="14" pageNumber="517" type="description">
<paragraph id="A606B5516013540BFD6501B38DAAFCA7" blockId="14.[707,1194,965,1730]" lastBlockId="15.[151,639,194,1516]" lastPageId="15" lastPageNumber="518" pageId="14" pageNumber="517">
<emphasis id="94CD69436013540AFD6501B38FE2FB41" bold="true" box="[731,917,1230,1251]" pageId="14" pageNumber="517">Description. Male.</emphasis>
Length
<quantity id="614118B46013540AFC5B01B3885CFB41" box="[997,1067,1230,1251]" metricMagnitude="-2" metricUnit="m" metricValue="1.6" pageId="14" pageNumber="517" unit="mm" value="16.0">16 mm</quantity>
, width
<quantity id="614118B46013540AFBCC01B388DEFB41" box="[1138,1193,1230,1251]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="14" pageNumber="517" unit="mm" value="4.0">4 mm</quantity>
(measured across bases of elytra). Red-brown, carina and tips of mandibles darker; antennae, palps, coxae, and tarsi lighter, almost tawny yellow (
<figureCitation id="3E82A9D46013540AFB3D00638EA6FAEC" captionStart="Figs" captionStartId="4.[197,236,1615,1634]" captionTargetBox="[245,1120,196,1597]" captionTargetId="figure-0@4.[243,1122,194,1599]" captionTargetPageId="4" captionText="Figs. 14. Athoplastus species, dorsal habitus. 1) A. acutus; 2) A. palouse; 3) A. lochsa; 4) A. rainieri." figureDoi="http://doi.org/10.5281/zenodo.4789354" httpUri="https://zenodo.org/record/4789354/files/figure.png" pageId="14" pageNumber="517">Fig. 4</figureCitation>
).
<emphasis id="94CD69436013540AFD5F00448F69FAEC" bold="true" box="[737,798,1337,1358]" pageId="14" pageNumber="517">Head:</emphasis>
Ocular index = 61.1.
<emphasis id="94CD69436013540AFC510044882FFAEC" bold="true" box="[1007,1112,1337,1358]" pageId="14" pageNumber="517">Antennae:</emphasis>
Ratio of antennomeres (211): 1: 1.25: 3: 3.25: 3.5: 3.25: 3.75: 2(?): X: X. [Antennomere 9 appears to be broken, and 10 and 11 are missing on the specimen measured.]
<emphasis id="94CD69436013540AFC8500DE8FDEFA1A" bold="true" box="[827,937,1443,1464]" pageId="14" pageNumber="517">Pronotum:</emphasis>
1.1X wider than long. Lateral margin completely carinate, slightly sinuate, almost subparallel anterad hind angles; convergent and slightly dorsoventrally flattened along lateral edges on anterior fourth. Punctures distinct, subumbilicate and dense on disc, separated by less than 0.2X their own diameter, with 2 “dimples” on either side of midline just anterad midlength, with less close punctures; punctures becoming clearly umbilicate and denser along anterior and lateral margins.
<emphasis id="94CD69436013540AFD4003D08F09F960" bold="true" box="[766,894,1709,1730]" pageId="14" pageNumber="517">Prosternum:</emphasis>
Slightly convex with dense, shallow, umbilicate punctures; anterior lobe short, broadly rounded, shallowly deflexed. Posterior process short and arcuate, gradually raised ventrally between procoxae, then sharply deflexed posteriorly; distinct ridge present immediately before procoxal cavity and along ventral surface of prosternal process.
<emphasis id="94CD69436012540BFE93041C8D13FED4" bold="true" box="[301,356,353,374]" pageId="15" pageNumber="518">Legs:</emphasis>
Tarsomere 5 equal to tarsomere 1 on pro- and mesotarsi, tarsomere 5 missing on both metatarsi.
<emphasis id="94CD69436012540BFEFD04EB8DFCFE09" bold="true" box="[323,395,406,427]" pageId="15" pageNumber="518">Elytra:</emphasis>
Elongate, subparallel, 4X pronotal length; striae shallowly impressed, deeper apically, punctures marked by darkening of the integument; intervals shallowly convex, punctures fine, shallow, and irregularly distributed.
<emphasis id="94CD69436012540BFDA2077D8E09FDB7" bold="true" box="[540,638,512,533]" pageId="15" pageNumber="518">Aedeagus</emphasis>
(
<figureCitation id="3E82A9D46012540BFF2007668D62FD92" box="[158,277,538,560]" captionStart="Figs" captionStartId="7.[175,214,1615,1634]" captionTargetBox="[246,1077,201,1597]" captionTargetId="figure-1@7.[244,1080,194,1599]" captionTargetPageId="7" captionText="Figs. 1627. Athoplastus species, adeagi, dorsal view, ventral view, and close up of paramere apex, respectively." figureDoi="http://doi.org/10.5281/zenodo.4789394" httpUri="https://zenodo.org/record/4789394/files/figure.png" pageId="15" pageNumber="518">Figs. 2224</figureCitation>
): Basal piece approximately 1/3 total length, with deep, narrowly U-shaped, membranous section on ventral surface (
<figureCitation id="3E82A9D46012540BFE18072D8D87FDC7" box="[422,496,592,613]" captionStart="Figs" captionStartId="7.[175,214,1615,1634]" captionTargetBox="[246,1077,201,1597]" captionTargetId="figure-1@7.[244,1080,194,1599]" captionTargetPageId="7" captionText="Figs. 1627. Athoplastus species, adeagi, dorsal view, ventral view, and close up of paramere apex, respectively." figureDoi="http://doi.org/10.5281/zenodo.4789394" httpUri="https://zenodo.org/record/4789394/files/figure.png" pageId="15" pageNumber="518">Fig. 23</figureCitation>
); median lobe 1.2X length of parameres, slightly deflexed ventrally at tip, median carina on ventral surface, basal struts following edges of median lobe; parameres wide basally, edges slightly sinuate and gradually narrowing toward tip, tip with subacute lateral expansion bearing 2 setae (
<figureCitation id="3E82A9D46012540BFE37078D8DA5FCA7" box="[393,466,752,773]" captionStart="Figs" captionStartId="7.[175,214,1615,1634]" captionTargetBox="[246,1077,201,1597]" captionTargetId="figure-1@7.[244,1080,194,1599]" captionTargetPageId="7" captionText="Figs. 1627. Athoplastus species, adeagi, dorsal view, ventral view, and close up of paramere apex, respectively." figureDoi="http://doi.org/10.5281/zenodo.4789394" httpUri="https://zenodo.org/record/4789394/files/figure.png" pageId="15" pageNumber="518">Fig. 24</figureCitation>
).
</paragraph>
<paragraph id="A606B5516012540BFF1106778C90FBE1" blockId="15.[151,639,194,1516]" pageId="15" pageNumber="518">
<emphasis id="94CD69436012540BFF1106778C88FCBD" bold="true" box="[175,255,778,799]" pageId="15" pageNumber="518">Female.</emphasis>
Length
<quantity id="614118B46012540BFEEE06778DE2FCBD" box="[336,405,778,799]" metricMagnitude="-2" metricUnit="m" metricValue="2.1" pageId="15" pageNumber="518" unit="mm" value="21.0">21 mm</quantity>
, width
<quantity id="614118B46012540BFE6306778E62FCBD" box="[477,533,778,799]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="15" pageNumber="518" unit="mm" value="5.0">5 mm</quantity>
(measured across bases of elytra). Yellow-brown.
<emphasis id="94CD69436012540BFDAA06588E25FC98" bold="true" box="[532,594,805,826]" pageId="15" pageNumber="518">Head:</emphasis>
Ocular index = 72.7.
<emphasis id="94CD69436012540BFEF906428DD2FCF6" bold="true" box="[327,421,831,852]" pageId="15" pageNumber="518">Antenna:</emphasis>
Ratio of antennomeres 211: 1: 1.6: 2.2: 2: 1.8: 1.6: 1.8: 1.6: X: X: X. [Note: Antennae are broken on specimen examined, so the last three antennomeres of longest intact antenna are missing.]
<emphasis id="94CD69436012540BFED806D78DA4FC1D" bold="true" box="[358,467,938,959]" pageId="15" pageNumber="518">Pronotum:</emphasis>
1.25X wider than long, hind angles divergent, sides broadly sinuate.
<emphasis id="94CD69436012540BFF2906A28CB9FC56" bold="true" box="[151,206,991,1012]" pageId="15" pageNumber="518">Legs:</emphasis>
Tarsomere 5 intermediate in size between tarsomeres 1 and 2 on metatarsi; pro- and mesotarsi as in male.
<emphasis id="94CD69436012540BFEB401698DD8FB8B" bold="true" box="[266,431,1044,1065]" pageId="15" pageNumber="518">Scutellar shield:</emphasis>
Setae radiating from midline.
</paragraph>
<paragraph id="A606B5516012540BFF1101348D7BFB5F" blockId="15.[151,639,194,1516]" pageId="15" pageNumber="518">
<emphasis id="94CD69436012540BFF1101348E7CFBFC" bold="true" box="[175,523,1097,1118]" pageId="15" pageNumber="518">Specimens Examined (3). Male.</emphasis>
<materialsCitation id="16D1BF0C6012540BFDA201348D58FB0C" ID-GBIF-Occurrence="3333036335" collectingDate="1936-05-13" collectingDateMax="1937-05-26" collectingDateMin="1936-05-13" collectionCode="CASC, USNM" collectorName="E. C. Van Dyke" country="United States of America" county="Pierce Co." location="Mt. Rainier" pageId="15" pageNumber="518" specimenCount="1" stateProvince="Washington" typeStatus="holotype">
<emphasis id="94CD69436012540BFDA201348DF3FBDA" bold="true" pageId="15" pageNumber="518">
<typeStatus id="79020BF36012540BFDA201348E0DFBFC" box="[540,634,1097,1118]" pageId="15" pageNumber="518" type="holotype">Holotype</typeStatus>
.
<collectingCountry id="DEAEF5C16012540BFF29011E8CBDFBDA" box="[151,202,1123,1144]" name="United States of America" pageId="15" pageNumber="518">USA</collectingCountry>
:
<collectingRegion id="647D7BB36012540BFF69011E8DF6FBDA" box="[215,385,1123,1144]" country="United States of America" name="Washington" pageId="15" pageNumber="518">WASHINGTON</collectingRegion>
:
</emphasis>
[
<collectingCounty id="4F67CDDD6012540BFE2C011E8D8AFBDA" box="[402,509,1123,1144]" pageId="15" pageNumber="518">Pierce Co.</collectingCounty>
];
<location id="A366E38A6012540BFDB1011E8E0AFBDA" LSID="urn:lsid:plazi:treatment:2E1004476013540BFD4D06788E3BFA4E:A366E38A6012540BFDB1011E8E0AFBDA" box="[527,637,1123,1144]" country="United States of America" county="Pierce Co." name="Mt. Rainier" pageId="15" pageNumber="518" stateProvince="Washington">Mt. Rainier</location>
[
<collectorName id="0B4CD0876012540BFF2001038D44FB31" box="[158,307,1150,1171]" pageId="15" pageNumber="518">E.C. Van Dyke</collectorName>
, date unknown] (
<collectionCode id="C0A82D946012540BFE6401038E6BFB31" box="[474,540,1150,1171]" pageId="15" pageNumber="518">CASC</collectionCode>
, 1,
<collectionCode id="C0A82D946012540BFD8101038E0AFB31" box="[575,637,1150,1171]" pageId="15" pageNumber="518">CASC</collectionCode>
type no. 3181).
</materialsCitation>
<materialsCitation id="16D1BF0C6012540BFE8901E48D7BFB5F" ID-GBIF-Occurrence="3333036337" collectingDate="1936-05-13" collectingDateMax="1937-05-26" collectingDateMin="1936-05-13" collectionCode="CASC, USNM" collectorName="M. C. Lane" country="United States of America" county="Pierce Co." location="Mt. Rainier" pageId="15" pageNumber="518" specimenCount="1" stateProvince="Washington" typeStatus="holotype">
<emphasis id="94CD69436012540BFE8901E48E0AFB0C" bold="true" box="[311,637,1177,1198]" pageId="15" pageNumber="518">
Female.
<collectingCountry id="DEAEF5C16012540BFE3101E48DB5FB0C" box="[399,450,1177,1198]" name="United States of America" pageId="15" pageNumber="518">USA</collectingCountry>
:
<collectingRegion id="647D7BB36012540BFE6E01E48E0DFB0C" box="[464,634,1177,1198]" country="United States of America" name="Washington" pageId="15" pageNumber="518">WASHINGTON</collectingRegion>
:
</emphasis>
[Kittitas Co.]
<collectorName id="0B4CD0876012540BFEA201CE8DD7FB6A" box="[284,416,1203,1224]" pageId="15" pageNumber="518">Lk. Cle Elum</collectorName>
;
<date id="D20793916012540BFE1501CE8E7CFB6A" box="[427,523,1203,1224]" pageId="15" pageNumber="518" value="1936-05-13">
<collectingDate id="C2436A796012540BFE1501CE8E7CFB6A" box="[427,523,1203,1224]" pageId="15" pageNumber="518" value="1936-05-13">13.v.1936</collectingDate>
</date>
;
<collectorName id="0B4CD0876012540BFDA801CE8E0AFB6A" box="[534,637,1203,1224]" pageId="15" pageNumber="518">M.C. Lane</collectorName>
(1,
<collectionCode id="C0A82D946012540BFF0B01B38C89FB41" box="[181,254,1230,1251]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="15" pageNumber="518" type="Museum">USNM</collectionCode>
);
<collectorName id="0B4CD0876012540BFEB201B38DFEFB41" box="[268,393,1230,1251]" pageId="15" pageNumber="518">Lk. Cle Elum</collectorName>
[?],
<date id="D20793916012540BFE0F01B38E66FB41" box="[433,529,1230,1251]" pageId="15" pageNumber="518" value="1937-05-26">
<collectingDate id="C2436A796012540BFE0F01B38E66FB41" box="[433,529,1230,1251]" pageId="15" pageNumber="518" value="1937-05-26">26.v.1937</collectingDate>
</date>
,
<collectorName id="0B4CD0876012540BFDA701B38E0AFB41" box="[537,637,1230,1251]" pageId="15" pageNumber="518">M.C. Lane</collectorName>
(1,
<collectionCode id="C0A82D946012540BFF0701958D75FB5F" box="[185,258,1256,1277]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="15" pageNumber="518" type="Museum">USNM</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="EEA3E6DA6012540BFF11007E8E3BFA4E" pageId="15" pageNumber="518" type="distribution">
<paragraph id="A606B5516012540BFF11007E8E18FA90" blockId="15.[151,639,194,1516]" pageId="15" pageNumber="518">
<emphasis id="94CD69436012540BFF11007E8D47FABA" bold="true" box="[175,304,1283,1304]" pageId="15" pageNumber="518">Distribution.</emphasis>
This species is only known from the Cascade Range of mountains in
<collectingRegion id="647D7BB36012540BFE4900608E1CFA90" box="[503,619,1309,1330]" country="United States of America" name="Washington" pageId="15" pageNumber="518">Washington</collectingRegion>
.
</paragraph>
<paragraph id="A606B5516012540BFF1100458E3BFA4E" blockId="15.[151,639,194,1516]" pageId="15" pageNumber="518">
<emphasis id="94CD69436012540BFF1100458DF5FAEF" bold="true" box="[175,386,1336,1357]" pageId="15" pageNumber="518">Taxonomic Remarks.</emphasis>
<bibRefCitation id="C228C8A06012540BFE3800458E53FAEF" author="Van Dyke, E. C." box="[390,548,1336,1357]" pageId="15" pageNumber="518" pagination="291 - 465" refId="ref12676" refString="Van Dyke, E. C. 1932. Miscellaneous studies in the Elateridae and related families of Coleoptera. Proceedings of the California Academy of Science 20: 291 - 465." type="journal article" year="1932">Van Dyke (1932)</bibRefCitation>
mentions a specimen seen “in the collection of Prof. W. J. Chamberlin at Corvallis, Oregon.” Chamberlain s collection was deposited at the Oregon State Arthropod Collection (OSAC) at Oregon State University (
<bibRefCitation id="C228C8A06012540BFF7800C08D4EFA70" author="Oman, P." box="[198,313,1469,1490]" pageId="15" pageNumber="518" pagination="165" refId="ref12277" refString="Oman, P. 1970. The Oregon State University insect collection. Bulletin of the Entomological Society of America 16: 165." type="journal article" year="1970">
<collectingCountry id="DEAEF5C16012540BFF7800C08D77FA70" box="[198,256,1469,1490]" name="Oman" pageId="15" pageNumber="518">Oman</collectingCountry>
1970
</bibRefCitation>
). However, a search of the OSAC yielded no specimen matching this description.
</paragraph>
</subSubSection>
</treatment>
</document>