218 lines
25 KiB
XML
218 lines
25 KiB
XML
<document id="60318174053DCF27AB71BD411AAE23F1" ID-DOI="10.5281/zenodo.156112" ID-GBIF-Dataset="f86b2225-fd69-4d14-888f-1656863a3244" ID-ISSN="11755326" ID-Zenodo-Dep="156112" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459440574016" checkinUser="plazi" docAuthor="Béthoux, Olivier & Nel, André" docDate="2002" docId="03A487F05712FB72FECA8BFF9F20A3FD" docLanguage="en" docName="zt00096.pdf" docOrigin="Zootaxa 96" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Permoraphidia magnifica Béthoux & Nel, 2002, sp. nov." docType="treatment" docVersion="15" lastPageNumber="41" masterDocId="FF9DFF885734FB5AFFC288139B43A04B" masterDocTitle="Venation pattern and revision of Orthoptera sensu nov. and sister groups. Phylogeny of Palaeozoic and Mesozoic Orthoptera sensu nov." masterLastPageNumber="88" masterPageNumber="1" pageNumber="39" updateTime="1698189919819" updateUser="plazi">
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<mods:mods id="6F483953B060862AAB19CB9152807AB0" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="2A1A0C471094100E60EF25AAB978E404">
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<mods:title id="10AEA81DC8317935F5DC2388D795ED35">Venation pattern and revision of Orthoptera sensu nov. and sister groups. Phylogeny of Palaeozoic and Mesozoic Orthoptera sensu nov.</mods:title>
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</mods:titleInfo>
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<mods:name id="F7BFC7BB6A2438ECFDFE68BC24B6C443" type="personal">
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<mods:role id="B14EC81D710C2400350186C17859A3FA">
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<mods:roleTerm id="2BBF3A86D868BB00BF67ADCDA7A68682">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="A371ABA0C822ED4130130F2B1ECE6794">Béthoux, Olivier</mods:namePart>
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</mods:name>
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<mods:name id="63E43FABA63EAC0946F940361A1985EA" type="personal">
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<mods:role id="5EC6F40AED7BF38F7AD3E179A214C19D">
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<mods:roleTerm id="907A4EBF5B2B41A1E515099B95BF6B7D">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="C30704A36194CC1A14B3780BDCB497A4">Nel, André</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="C16EA77A4F972B8BA14E5FA952A34BBD">text</mods:typeOfResource>
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<mods:titleInfo id="C649B98501FA4DD9EF210E68DFE1BE7C">
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<mods:title id="203535041E3E167EF6EC922EC1ADD011">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="FDFD619051FB9BFC3BBEC8022D87C1C7">
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<mods:date id="72BA309F8D0F85753FE74F0520576914">2002</mods:date>
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<mods:detail id="F385D4AA068815BBC1E563141CB378FD" type="volume">
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<mods:number id="51DD48FD8C8D8B09AC45A98AFC73FDB7">96</mods:number>
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</mods:detail>
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<mods:extent id="A45B52DE43DC270550DA908C8AE31908" unit="page">
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<mods:start id="8301FF7BECB96F9D102AE74D3226B83E">1</mods:start>
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<mods:end id="8DC1FA11D681EBAFF368F1F80D7292B4">88</mods:end>
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</mods:extent>
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</mods:part>
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<mods:classification id="AD18362FE64B589A8C5C2032615CDE1F">journal article</mods:classification>
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<mods:identifier id="DDDB522A783D1812A3FFB96BD253CC38" type="DOI">10.5281/zenodo.156112</mods:identifier>
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<mods:identifier id="A283C6086E7F5FE5DDC8466DB0F51372" type="GBIF-Dataset">f86b2225-fd69-4d14-888f-1656863a3244</mods:identifier>
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<mods:identifier id="65A26B7A73FA878D7ADB3BDA5AF77ACF" type="ISSN">11755326</mods:identifier>
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<mods:identifier id="32502BBD544FA9AFD59A6D8BFEAB9F06" type="Zenodo-Dep">156112</mods:identifier>
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</mods:mods>
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<treatment id="03A487F05712FB72FECA8BFF9F20A3FD" ID-DOI="http://doi.org/10.5281/zenodo.6278056" ID-GBIF-Taxon="119330387" ID-Zenodo-Dep="6278056" LSID="urn:lsid:plazi:treatment:03A487F05712FB72FECA8BFF9F20A3FD" httpUri="http://treatment.plazi.org/id/03A487F05712FB72FECA8BFF9F20A3FD" lastPageId="40" lastPageNumber="41" pageId="38" pageNumber="39">
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<subSubSection id="C317656D5712FB7CFECA8BFF99E6A44E" box="[264,677,1003,1030]" pageId="38" pageNumber="39" type="nomenclature">
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<paragraph id="8BB236E65712FB7CFECA8BFF99E6A44E" blockId="38.[264,677,1003,1030]" box="[264,677,1003,1030]" pageId="38" pageNumber="39">
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<heading id="D0FA818A5712FB7CFECA8BFF99E6A44E" bold="true" box="[264,677,1003,1030]" fontSize="11" level="1" pageId="38" pageNumber="39" reason="1">
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<emphasis id="B979EAF45712FB7CFECA8BFF99E6A44E" bold="true" box="[264,677,1003,1030]" pageId="38" pageNumber="39">
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<taxonomicName id="4C0D4D655712FB7CFECA8BFF9902A44D" ID-CoL="4F6S2" box="[264,577,1004,1030]" class="Insecta" family="Permoraphidiidae" genus="Permoraphidia" kingdom="Animalia" order="Orthoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="magnifica" status="sp. nov.">
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<emphasis id="B979EAF45712FB7CFECA8BFF9902A44D" bold="true" box="[264,577,1004,1030]" italics="true" pageId="38" pageNumber="39">Permoraphidia magnifica</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A24A578F5712FB7CFD858BF899E6A44E" box="[583,677,1003,1029]" pageId="38" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C317656D5712FB72FECA8C2F9F20A3FD" lastPageId="40" lastPageNumber="41" pageId="38" pageNumber="39" type="materials_examined">
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<paragraph id="8BB236E65712FB7CFECA8C2F9FADA41D" blockId="38.[264,1325,1084,1870]" box="[264,1262,1084,1110]" pageId="38" pageNumber="39">
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Material.
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<typeStatus id="54B688445712FB7CFEB88C2F9AA4A41D" box="[378,487,1084,1110]" pageId="38" pageNumber="39" type="holotype">Holotype</typeStatus>
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specimen MNHNLPB.7519 (Inv. number of collection 19072).
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</paragraph>
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<paragraph id="8BB236E65712FB7CFEFA8C779868A435" blockId="38.[264,1325,1084,1870]" box="[312,811,1124,1150]" pageId="38" pageNumber="39">
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Geological settings. Permian,
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<collectingCountry id="F31A76765712FB7CFD5A8C77986BA435" box="[664,808,1124,1150]" name="Madagascar" pageId="38" pageNumber="39">Madagascar</collectingCountry>
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.
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</paragraph>
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<paragraph id="8BB236E65712FB7CFEFA8C9F98BBA4ED" blockId="38.[264,1325,1084,1870]" box="[312,1016,1164,1190]" pageId="38" pageNumber="39">
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Etymology. After the excellent preservation of the
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<typeStatus id="54B688445712FB7CFC488C9F98B6A4ED" box="[906,1013,1164,1190]" pageId="38" pageNumber="39" type="holotype">holotype</typeStatus>
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.
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</paragraph>
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<paragraph id="8BB236E65712FB7CFEFA8CA79939A50D" blockId="38.[264,1325,1084,1870]" pageId="38" pageNumber="39">
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Note. This specimen is in a flat and elliptic nodule. No further indication about the exact origin and age of this fossil is known. There is no available record of a Permian nodule outcrop in
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<collectingCountry id="F31A76765712FB7CFE1C8D17992DA555" box="[478,622,1284,1310]" name="Madagascar" pageId="38" pageNumber="39">Madagascar</collectingCountry>
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.
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<collectingCountry id="F31A76765712FB7CFDBA8D1799DBA555" box="[632,664,1284,1310]" name="American Samoa" pageId="38" pageNumber="39">As</collectingCountry>
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this fossil is very well preserved, new investigations on its origin should be undertaken.
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</paragraph>
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<paragraph id="8BB236E65712FB7CFEFA8D479F02A67D" blockId="38.[264,1325,1084,1870]" pageId="38" pageNumber="39">
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Diagnosis. Differences with
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<taxonomicName id="4C0D4D655712FB7CFD538D47986DA526" box="[657,814,1364,1389]" class="Insecta" family="Permoraphidiidae" genus="Permoraphidia" kingdom="Animalia" order="Orthoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="americana">
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<emphasis id="B979EAF45712FB7CFD538D47986DA526" box="[657,814,1364,1389]" italics="true" pageId="38" pageNumber="39">P. americana</emphasis>
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</taxonomicName>
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are as follows: CuA + CuPa with more than two branches, forewing nearly twice as long as that of
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<emphasis id="B979EAF45712FB7CFC028D6F9F14A5DE" box="[960,1111,1404,1429]" italics="true" pageId="38" pageNumber="39">P. a m e r i c a n a</emphasis>
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. It differs from
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<taxonomicName id="4C0D4D655712FB7CFAD78D6F9A21A5F6" class="Insecta" family="Permoraphidiidae" genus="Permoraphidia" kingdom="Animalia" order="Orthoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="grandis">
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<emphasis id="B979EAF45712FB7CFAD78D6F9A21A5F6" italics="true" pageId="38" pageNumber="39">P. grandis</emphasis>
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</taxonomicName>
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in its non sclerotized pterostigma, resembling that of
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<taxonomicName id="4C0D4D655712FB7CFBCA8DB79FE4A5F6" box="[1032,1191,1444,1469]" class="Insecta" family="Permoraphidiidae" genus="Permoraphidia" kingdom="Animalia" order="Orthoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="americana">
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<emphasis id="B979EAF45712FB7CFBCA8DB79FE4A5F6" box="[1032,1191,1444,1469]" italics="true" pageId="38" pageNumber="39">P. americana</emphasis>
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</taxonomicName>
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. Also, the fusion of
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<collectingCountry id="F31A76765712FB7CFEB88DDF9ADDA5AD" box="[378,414,1484,1510]" name="Poland" pageId="38" pageNumber="39">RP</collectingCountry>
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with MA is long, distinctly longer than in
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<taxonomicName id="4C0D4D655712FB7CFC5E8DDF9F57A5AE" box="[924,1044,1484,1509]" class="Insecta" family="Permoraphidiidae" genus="Permoraphidia" kingdom="Animalia" order="Orthoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="grandis">
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<emphasis id="B979EAF45712FB7CFC5E8DDF9F57A5AE" box="[924,1044,1484,1509]" italics="true" pageId="38" pageNumber="39">P. grandis</emphasis>
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</taxonomicName>
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and comparable to that of
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<taxonomicName id="4C0D4D655712FB7CFEE98DE799EAA645" authority="Carpenter, 1943" authorityName="Carpenter" authorityYear="1943" box="[299,681,1524,1550]" class="Insecta" family="Permoraphidiidae" genus="Permoraphidia" kingdom="Animalia" order="Orthoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="americana">
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<emphasis id="B979EAF45712FB7CFEE98DE79A8AA646" box="[299,457,1524,1549]" italics="true" pageId="38" pageNumber="39">P. americana</emphasis>
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(
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<bibRefCitation id="EF9C4B175712FB7CFE198DE799E3A645" author="Carpenter" box="[475,672,1524,1550]" pageId="38" pageNumber="67" refString="Carpenter, F. M. (1943) The Lower Permian insects of Kansas. Part 9: the orders Neuroptera, Raphidiodea, Caloneurodea and Protorthoptera (Probnisidae), with additional Protodonata and Megasecoptera. Proceedings of the American Academy of Arts and Sciences, 75, 55 - 84." type="journal article" year="1943">Carpenter, 1943</bibRefCitation>
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)
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</taxonomicName>
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. Unfortunately,
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<taxonomicName id="4C0D4D655712FB7CFCA98DE798A6A646" box="[875,997,1524,1549]" class="Insecta" family="Permoraphidiidae" genus="Permoraphidia" kingdom="Animalia" order="Orthoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="grandis">
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<emphasis id="B979EAF45712FB7CFCA98DE798A6A646" box="[875,997,1524,1549]" italics="true" pageId="38" pageNumber="39">P. grandis</emphasis>
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</taxonomicName>
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is based on an incomplete forewing with ramifications of CuA + CuPa incompletely preserved.
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</paragraph>
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<paragraph id="8BB236E65712FB7CFEFA8E57998AA6CD" blockId="38.[264,1325,1084,1870]" pageId="38" pageNumber="39">Description. Print and counter print of a nearly complete body with the four wings, three legs and antennae in connection.</paragraph>
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<paragraph id="8BB236E65712FB7CFEFA8E879A38A705" blockId="38.[264,1325,1084,1870]" pageId="38" pageNumber="39">
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Body length about 13.0 mm. Antenna short
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<quantity id="4CF59B035712FB7CFCFC8E8798DAA6E5" box="[830,921,1684,1710]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="38" pageNumber="39" unit="mm" value="1.9">1.9 mm</quantity>
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long. Metathoracic leg saltatorial, with the femora broadened
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<quantity id="4CF59B035712FB7CFD898EAF99E6A69D" box="[587,677,1724,1750]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="38" pageNumber="39" unit="mm" value="1.3">1.3 mm</quantity>
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wide and
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<quantity id="4CF59B035712FB7CFCDC8EAF983BA69D" box="[798,888,1724,1750]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" pageId="38" pageNumber="39" unit="mm" value="5.1">5.1 mm</quantity>
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long, tibia
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<quantity id="4CF59B035712FB7CFC3F8EAF9F14A69D" box="[1021,1111,1724,1750]" metricMagnitude="-3" metricUnit="m" metricValue="6.3" pageId="38" pageNumber="39" unit="mm" value="6.3">6.3 mm</quantity>
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long, two rows of strong and stout outer tibial spines as in modern
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<taxonomicName id="4C0D4D655712FB7CFC8A8EF79889A6B5" box="[840,970,1764,1790]" class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Orthoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="order">Orthoptera</taxonomicName>
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, tarsi
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<quantity id="4CF59B035712FB7CFBD28EF79F28A6B5" box="[1040,1131,1764,1790]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="38" pageNumber="39" unit="mm" value="0.8">0.8 mm</quantity>
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long; other legs more poorly preserved; two cerci probably visible but no ovipositor at the apex of the abdomen.
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</paragraph>
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<paragraph id="8BB236E65713FB7DFEFA890F981EA41D" blockId="39.[264,1324,284,1110]" pageId="39" pageNumber="40">
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Forewing (figs 15ab, 16)
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<quantity id="4CF59B035713FB7DFDAE890F9997A17D" box="[620,724,284,310]" metricMagnitude="-2" metricUnit="m" metricValue="1.68" pageId="39" pageNumber="40" unit="mm" value="16.8">16.8 mm</quantity>
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long,
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<quantity id="4CF59B035713FB7DFCDC890F983BA17D" box="[798,888,284,310]" metricMagnitude="-3" metricUnit="m" metricValue="3.6" pageId="39" pageNumber="40" unit="mm" value="3.6">3.6 mm</quantity>
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wide, with relatively few and spaced crossveins; ScA short, poorly preserved; ScP slightly zigzagged, with simple, slightly curved and regularly spaced anterior branches; area between ScP and costal margin
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<quantity id="4CF59B035713FB7DFAC4897F9A75A1E5" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="39" pageNumber="40" unit="mm" value="0.7">0.7 mm</quantity>
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wide; costal margin with a posterior concavity at the level of the distal end of ScP; area between R and ScP narrow, dark with eight or nine short crossveins between them; pterostigma dark, but not sclerotized, with four to five faint crossveins in it; base of
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<collectingCountry id="F31A76765713FB7DFACA89F79E6FA1B5" box="[1288,1324,484,510]" name="Poland" pageId="39" pageNumber="40">RP</collectingCountry>
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||
just distal to mid length of wing, with a short but strong crossvein between R and MA just basal of it; basal free part of
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<collectingCountry id="F31A76765713FB7DFDB18A2799D4A205" box="[627,663,564,590]" name="Poland" pageId="39" pageNumber="40">RP</collectingCountry>
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||
relatively short,
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<quantity id="4CF59B035713FB7DFCAB8A279884A205" box="[873,967,564,590]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="39" pageNumber="40" unit="mm" value="1.3">1.3 mm</quantity>
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long, without any crossvein reaching it; fusion of
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<collectingCountry id="F31A76765713FB7DFDC08A4F9965A23D" box="[514,550,604,630]" name="Poland" pageId="39" pageNumber="40">RP</collectingCountry>
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||
with MA
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<quantity id="4CF59B035713FB7DFD5D8A4F99BBA23D" box="[671,760,604,630]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="39" pageNumber="40" unit="mm" value="0.8">0.8 mm</quantity>
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long;
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<collectingCountry id="F31A76765713FB7DFC808A4F9825A23D" box="[834,870,604,630]" name="Poland" pageId="39" pageNumber="40">RP</collectingCountry>
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with two or three distal main branches distally ramified; wing apex darkly pigmented; basal stem M + CuA diverges from R opposite to end of ScA on anterior margin; CuPa reaching M + CuA basal of origin of distal free part of CuA + CuPa, just distal to the first third of the wing; CuA + CuPa separates distally, posteriorly irregularly pectinate, with five to six branches reaching posterior wing margin; MP separating from MA close to base of CuA + CuPa; MP very distally and weakly ramified; MA with a distinct angle at the level of the crossvein between R and MA; distal free part of MA simple (right wing) or ramified (left wing); MA not divided into MA1 and MA2; CuPa simple, separated from CuPa at about the basal third of the wing; CuPa and CuPb separated near the wing base; CuPb is a weak simple vein, very near and markedly parallel with 1A; 1A strong and simple; two other longitudinal anal veins; ends of first posterior branch of CuA + CuPa, CuPa, CuPb and 1A strongly approximate at posterior wing margin.
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</paragraph>
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<caption id="DF72666E5713FB7DFECA8E6C9A37A6BD" httpUri="https://zenodo.org/record/156126/files/figure.png" pageId="39" pageNumber="40" targetBox="[265,1309,1178,1646]" targetPageId="39">
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<paragraph id="8BB236E65713FB7DFECA8E6C9A37A6BD" blockId="39.[264,1323,1663,1782]" pageId="39" pageNumber="40">
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<emphasis id="B979EAF45713FB7DFECA8E6C9ADBA6DC" bold="true" box="[264,408,1663,1687]" pageId="39" pageNumber="40">FIGURE 15.</emphasis>
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<taxonomicName id="4C0D4D655713FB7DFE5D8E6C99F5A6DD" box="[415,694,1663,1686]" class="Insecta" family="Permoraphidiidae" genus="Permoraphidia" kingdom="Animalia" order="Orthoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="magnifica" status="sp. nov.">
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<emphasis id="B979EAF45713FB7DFE5D8E6C99F5A6DD" box="[415,694,1663,1686]" italics="true" pageId="39" pageNumber="40">Permoraphidia magnifica</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A24A578F5713FB7DFD028E6C985BA6DD" box="[704,792,1663,1686]" pageId="39" pageNumber="40" rank="species">sp. nov.</taxonomicNameLabel>
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, holotype specimen MNHNLPB.7519ab (print and counterpart). (scale bar represents 5 mm). 15a: reconstruction of left forewing; 15b: reconstruction of right forewing; 15c: reconstruction of left hindwing; 15d: reconstruction of right hindwing.
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</paragraph>
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</caption>
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<paragraph id="8BB236E65713FB72FEFA8F349E5BA1CD" blockId="39.[264,1324,1831,1897]" lastBlockId="40.[264,1324,284,950]" lastPageId="40" lastPageNumber="41" pageId="39" pageNumber="40">
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Hindwing (figs 15.c,d), slightly shorter than forewing, about
|
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<quantity id="4CF59B035713FB7DFBCE8F349F36A70A" box="[1036,1141,1831,1857]" metricMagnitude="-2" metricUnit="m" metricValue="1.0699999999999998" pageId="39" pageNumber="40" unit="mm" value="10.7">10.7 mm</quantity>
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long; anal area only partly preserved; CuA + CuPa with two apical branches; MA very briefly fused with
|
||
<collectingCountry id="F31A7676571CFB72FE85890F9A28A17D" box="[327,363,284,310]" name="Poland" pageId="40" pageNumber="41">RP</collectingCountry>
|
||
in left wing but separated in right wing (with short crossvein between them); pterostigma and wing apex darkly pigmented; presence of a short secondary longitudinal vein between CuPa and CuPb; posterior wing margin angular at the distal end of CuPb.
|
||
</paragraph>
|
||
<paragraph id="8BB236E6571CFB72FEFA8987981CA1B5" blockId="40.[264,1324,284,950]" pageId="40" pageNumber="41">
|
||
Discussion. The presence of saltatorial metathoracic legs confirms an accurate assignment of
|
||
<taxonomicName id="4C0D4D65571CFB72FE0389AF99B4A19E" box="[449,759,444,469]" class="Insecta" family="Permoraphidiidae" genus="Permoraphidia" kingdom="Animalia" order="Orthoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="magnifica">
|
||
<emphasis id="B979EAF4571CFB72FE0389AF99B4A19E" box="[449,759,444,469]" italics="true" pageId="40" pageNumber="41">Permoraphidia magnifica</emphasis>
|
||
</taxonomicName>
|
||
and the
|
||
<taxonomicName id="4C0D4D65571CFB72FCAD89AF9F03A19D" box="[879,1088,444,470]" class="Insecta" family="Permoraphidiidae" kingdom="Animalia" order="Orthoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="family">Permoraphidiidae</taxonomicName>
|
||
to the
|
||
<taxonomicName id="4C0D4D65571CFB72FB6789AF9E6BA19D" box="[1189,1320,444,470]" class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Orthoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="order">Orthoptera</taxonomicName>
|
||
,
|
||
<emphasis id="B979EAF4571CFB72FECA89F79A16A1B6" box="[264,341,484,509]" italics="true" pageId="40" pageNumber="41">contra</emphasis>
|
||
<bibRefCitation id="EF9C4B17571CFB72FE9E89F7994DA1B5" author="Tillyard" box="[348,526,484,510]" pageId="40" pageNumber="71" refString="Tillyard, R. J. (1932) Kansas Permian Insects. Part 14. The order Neuroptera. American Journal of Sciences, (5), 23, 1 - 30." type="journal article" year="1932">Tillyard (1932)</bibRefCitation>
|
||
(assignment in
|
||
<taxonomicName id="4C0D4D65571CFB72FD0B89F79810A1B5" box="[713,851,484,510]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="order">Neuroptera</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BB236E6571CFB72FEFA8A1F9F20A3FD" blockId="40.[264,1324,284,950]" pageId="40" pageNumber="41">
|
||
Following the key of families of ‘Orthoptera’ proposed by
|
||
<bibRefCitation id="EF9C4B17571CFB72FC3F8A1F9FE9A26D" author="Sharov" box="[1021,1194,524,550]" pageId="40" pageNumber="71" refString="Sharov, A. G. (1968) Filogeniya ortopteroidnykh nasekomykh. Trudy Paleontologicheskovo Instituta Akademii nauk SSSR, 118, 1 - 216. [in Russian, Translated in English in 1971: Phylogeny of the Orthopteroidea. Israel program for scientific translations, Jerusalem, 118: 1 - 251.]" type="journal article" year="1968">Sharov (1968)</bibRefCitation>
|
||
, this fossil falls into the family
|
||
<taxonomicName id="4C0D4D65571CFB72FDC38A279992A205" box="[513,721,564,590]" class="Insecta" family="Permoraphidiidae" kingdom="Animalia" order="Orthoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="family">Permoraphidiidae</taxonomicName>
|
||
rather than
|
||
<taxonomicName id="4C0D4D65571CFB72FCA58A279898A205" box="[871,987,564,590]" class="Insecta" family="Elcanidae" kingdom="Animalia" order="Orthoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="family">Elcanidae</taxonomicName>
|
||
because its CuPa is fused with M + CuA instead of being fused with the distal free part of CuA and this fusion is well distal of the level of distal end of ScA.
|
||
<bibRefCitation id="EF9C4B17571CFB72FCFE8A979F49A2D5" author="Carpenter" box="[828,1034,644,670]" pageId="40" pageNumber="67" refString="Carpenter, F. M. (1992). Superclass Hexapoda. In: Moore, R. C. & Kaesler, R. L. (Eds) Treatise on Invertebrate Paleontology. The Geological Society of America and the University of Kansas, Boulder, Colorado, (R), Arthropoda 4 (3 / 4), xxi + 655 pp." type="book" year="1992">Carpenter (1992)</bibRefCitation>
|
||
indicated in his family diagnosis of the
|
||
<taxonomicName id="4C0D4D65571CFB72FE058ABF99D4A28D" box="[455,663,684,710]" class="Insecta" family="Permoraphidiidae" kingdom="Animalia" order="Orthoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="family">Permoraphidiidae</taxonomicName>
|
||
that the hindwing MA is anastomosed with
|
||
<collectingCountry id="F31A7676571CFB72FB588ABF9FFDA28D" box="[1178,1214,684,710]" name="Poland" pageId="40" pageNumber="41">RP</collectingCountry>
|
||
, which is apparently not a stable character as this is not the case on the right hindwing in our fossil.
|
||
<taxonomicName id="4C0D4D65571CFB72FECA8AEF997DA35E" box="[264,574,764,789]" class="Insecta" family="Permoraphidiidae" genus="Permoraphidia" kingdom="Animalia" order="Orthoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="magnifica" status="sp. nov.">
|
||
<emphasis id="B979EAF4571CFB72FECA8AEF997DA35E" box="[264,574,764,789]" italics="true" pageId="40" pageNumber="41">Permoraphidia magnifica</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24A578F571CFB72FD8E8AEF99EEA35D" box="[588,685,764,790]" pageId="40" pageNumber="41" rank="species">sp. nov.</taxonomicNameLabel>
|
||
shares with the two other
|
||
<taxonomicName id="4C0D4D65571CFB72FBCA8AEF9FF8A35E" box="[1032,1211,764,789]" class="Insecta" family="Permoraphidiidae" genus="Permoraphidia" kingdom="Animalia" order="Orthoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B979EAF4571CFB72FBCA8AEF9FF8A35E" box="[1032,1211,764,789]" italics="true" pageId="40" pageNumber="41">Permoraphidia</emphasis>
|
||
</taxonomicName>
|
||
spp. the presence of a strong crossvein between R and MA just basal of the origin of
|
||
<collectingCountry id="F31A7676571CFB72FB648B379F89A375" box="[1190,1226,804,830]" name="Poland" pageId="40" pageNumber="41">RP</collectingCountry>
|
||
(unique character which is a synapomorphy of the family). The character ‘base of MP very close to that of CuA + CuPa ’ is also probably derived, but maybe more homoplasic as it is also present in
|
||
<taxonomicName id="4C0D4D65571CFB72FE408B8F9941A3FE" box="[386,514,924,949]" class="Insecta" family="Bouretidae" genus="Petrelcana" kingdom="Animalia" order="Orthoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B979EAF4571CFB72FE408B8F9941A3FE" box="[386,514,924,949]" italics="true" pageId="40" pageNumber="41">Petrelcana</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C0D4D65571CFB72FDF98B8F99C0A3FE" box="[571,643,924,949]" class="Insecta" family="Oedischiidae" genus="Iasvia" kingdom="Animalia" order="Orthoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B979EAF4571CFB72FDF98B8F99C0A3FE" box="[571,643,924,949]" italics="true" pageId="40" pageNumber="41">Iasvia</emphasis>
|
||
</taxonomicName>
|
||
(herein assigned to
|
||
<taxonomicName id="4C0D4D65571CFB72FCAD8B8F9F1BA3FD" box="[879,1112,924,950]" class="Insecta" family="Oedischiidae" kingdom="Animalia" order="Orthoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="subFamily" subFamily="Mezenoedischiinae">Mezenoedischiinae</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |