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<document id="FDC61F455494AE373B84D43872D51917" ID-DOI="10.37828/em.2023.64.1" ID-ISSN="2336-9744" ID-Zenodo-Dep="13248512" ID-ZooBank="urn:lsid:zoobank.org:pub:ED4A9B48-70C2-4B8D-B8F1-513B217A439B" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722974219379" checkinUser="felipe" docAuthor="Volynkin, Anton V." docDate="2023" docId="03A487E9FFE6666CFF6C4D65C303F964" docLanguage="en" docName="EcolMontenegrina.64.1-184.pdf" docOrigin="Ecologica Montenegrina 64" docSource="http://dx.doi.org/10.37828/em.2023.64.1" docStyle="DocumentStyle:3485E46B0EDC64DBF2992EDE30BFB5AE.2:EcolMontenegrina.2017-.journal_article" docStyleId="3485E46B0EDC64DBF2992EDE30BFB5AE" docStyleName="EcolMontenegrina.2017-.journal_article" docStyleVersion="2" docTitle="Siccia nyasa Volynkin, 2023, sp. n." docType="treatment" docUuid="E948DFE2-D6DC-423C-A680-F231E6D24680" docUuidSource="ZooBank" docVersion="2" lastPageNumber="86" masterDocId="FF9DFF91FFB36639FFFB4E67C14AFFAD" masterDocTitle="Contribution to the knowledge of the genus Siccia Walker (= Aemene Walker, syn. n.) in the Afrotropics with descriptions of seventy-three new species, three new subspecies, and a check-list of Asiatic taxa of the genus (Lepidoptera: Erebidae: Arctiinae: Lithosiini)" masterLastPageNumber="184" masterPageNumber="1" pageNumber="86" updateTime="1722983564555" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="01B7D303BCC02CF7129D5C6ED51A12BF">Contribution to the knowledge of the genus Siccia Walker (= Aemene Walker, syn. n.) in the Afrotropics with descriptions of seventy-three new species, three new subspecies, and a check-list of Asiatic taxa of the genus (Lepidoptera: Erebidae: Arctiinae: Lithosiini)</mods:title>
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<mods:namePart id="18E94F535508BBCF6CBE459AF095622C">Volynkin, Anton V.</mods:namePart>
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<mods:affiliation id="73016827DE754936EAD62A9A965AF0CC">The African Natural History Research Trust (ANHRT), Street Court Leominster, Kingsland, HR 6 9 QA, UK. E-mail: anton @ anhrt. org. uk; https: // orcid. org / 0000 - 0001 - 9447 - 4925 &amp; Altai State University, Lenina Avenue 61, RF- 656049, Barnaul, Russia.</mods:affiliation>
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<mods:title id="1C0F79C5ADEFE6F27A7B4EAF41EEFD31">Ecologica Montenegrina</mods:title>
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<mods:date id="6BA0F207E19C82399E651417E6E0E3BD">2023</mods:date>
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<paragraph id="8BB236FFFFE6666CFF6C4D65C03DFCB1" blockId="85.[151,1145,770,860]" box="[151,375,770,796]" pageId="85" pageNumber="86">
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<taxonomicName id="4C0D4D7CFFE6666CFF6C4D65C064FCB1" ID-CoL="BJ4KT" authorityName="Volynkin" authorityYear="2023" box="[151,302,770,796]" class="Insecta" family="Erebidae" genus="Siccia" kingdom="Animalia" order="Lepidoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="nyasa" status="sp. nov.">
<emphasis id="B979EAEDFFE6666CFF6C4D65C064FCB1" bold="true" box="[151,302,770,796]" italics="true" pageId="85" pageNumber="86">Siccia nyasa</emphasis>
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https://zoobank.org/
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urn:lsid:zoobank.org:act:
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<subSubSection id="C3176574FFE6666CFF6C4D25C001FCF1" box="[151,331,834,860]" pageId="85" pageNumber="86" type="description">
<paragraph id="8BB236FFFFE6666CFF6C4D25C001FCF1" blockId="85.[151,1145,770,860]" box="[151,331,834,860]" pageId="85" pageNumber="86">
(
<figureCitation id="13362A7AFFE6666CFF644D25C040FCF1" box="[159,266,834,860]" captionStart="Figures 223239" captionStartId="121.[151,237,1965,1988]" captionTargetBox="[154,1433,190,1914]" captionTargetId="figure-6@121.[151,1436,189,1928]" captionTargetPageId="121" captionText="Figures 223239. Siccia spp., adults. Depositories of the specimens: 223 in MHNG (photo by B. Landry); 224, 225, 227, 229231, and 238, 239 in ANHRT (227: ex AKW); 226 and 233 in NHMUK (©The Trustees of NHMUK); 228, 236 and 237 in ALW; 232 in MWM/ZSM; 234 and 235 in MfN." figureDoi="http://doi.org/10.5281/zenodo.13248557" httpUri="https://zenodo.org/record/13248557/files/figure.png" pageId="85" pageNumber="86">Figs 233</figureCitation>
,
<figureCitation id="13362A7AFFE6666CFEED4D25C009FCF1" box="[278,323,834,860]" captionStart="Figures 423426" captionStartId="159.[151,237,1996,2019]" captionTargetBox="[189,1430,190,1906]" captionTargetId="figure-6@159.[151,1436,189,1928]" captionTargetPageId="159" captionText="Figures 423426. Siccia spp., male genitalia. Depositories of the specimens dissected: 423 and 424 in ANHRT; 425 in MWM/ZSM; 426 in NHMUK (©The Trustees of NHMUK)." figureDoi="http://doi.org/10.5281/zenodo.13248661" httpUri="https://zenodo.org/record/13248661/files/figure.png" pageId="85" pageNumber="86">426</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3176574FFE6666CFF6C4DE2C443FC4E" pageId="85" pageNumber="86" type="materials_examined">
<paragraph id="8BB236FFFFE6666CFF6C4DE2C443FC4E" blockId="85.[151,1437,901,1737]" pageId="85" pageNumber="86">
<emphasis id="B979EAEDFFE6666CFF6C4DE2C01BFC32" bold="true" box="[151,337,901,927]" pageId="85" pageNumber="86">Type material</emphasis>
.
<materialsCitation id="3B653CA2FFE6666CFE924DE2C443FC4E" collectingDate="1913-04-04" collectionCode="NHMUK" country="Malawi" location="Volynkin" pageId="85" pageNumber="86" specimenCount="1" stateProvince="Mulanje" typeStatus="holotype">
<emphasis id="B979EAEDFFE6666CFE924DE2C091FC32" bold="true" box="[361,475,901,927]" pageId="85" pageNumber="86">
<typeStatus id="54B6885DFFE6666CFE924DE2C091FC32" box="[361,475,901,927]" pageId="85" pageNumber="86" type="holotype">Holotype</typeStatus>
</emphasis>
(
<figureCitation id="13362A7AFFE6666CFE0E4DE2C322FC32" box="[501,616,901,927]" captionStart="Figures 223239" captionStartId="121.[151,237,1965,1988]" captionTargetBox="[154,1433,190,1914]" captionTargetId="figure-6@121.[151,1436,189,1928]" captionTargetPageId="121" captionText="Figures 223239. Siccia spp., adults. Depositories of the specimens: 223 in MHNG (photo by B. Landry); 224, 225, 227, 229231, and 238, 239 in ANHRT (227: ex AKW); 226 and 233 in NHMUK (©The Trustees of NHMUK); 228, 236 and 237 in ALW; 232 in MWM/ZSM; 234 and 235 in MfN." figureDoi="http://doi.org/10.5281/zenodo.13248557" httpUri="https://zenodo.org/record/13248557/files/figure.png" pageId="85" pageNumber="86">Figs 233</figureCitation>
,
<figureCitation id="13362A7AFFE6666CFD854DE2C3FAFC32" box="[638,688,901,927]" captionStart="Figures 423426" captionStartId="159.[151,237,1996,2019]" captionTargetBox="[189,1430,190,1906]" captionTargetId="figure-6@159.[151,1436,189,1928]" captionTargetPageId="159" captionText="Figures 423426. Siccia spp., male genitalia. Depositories of the specimens dissected: 423 and 424 in ANHRT; 425 in MWM/ZSM; 426 in NHMUK (©The Trustees of NHMUK)." figureDoi="http://doi.org/10.5281/zenodo.13248661" httpUri="https://zenodo.org/record/13248661/files/figure.png" pageId="85" pageNumber="86">426</figureCitation>
): male, [
<collectingCountry id="F31A766FFFE6666CFCDF4DE2C2CEFC32" box="[804,900,901,927]" name="Malawi" pageId="85" pageNumber="86">Malawi</collectingCountry>
,
<collectingRegion id="49C9F81DFFE6666CFC624DE2C2B5FC32" box="[921,1023,901,927]" country="Malawi" name="Mulanje" pageId="85" pageNumber="86">Mulanje</collectingRegion>
] “Mlanje, | Nyasaland. | 4·4·13 [
<date id="FFB3103FFFE6666CFF644DC0C059FC6C" box="[159,275,935,961]" pageId="85" pageNumber="86" value="1913-04-04">
<collectingDate id="EFF7E9D7FFE6666CFF644DC0C059FC6C" box="[159,275,935,961]" pageId="85" pageNumber="86" value="1913-04-04">4.iv.1913</collectingDate>
</date>
]” / “Rothschild | Bequest | B.M.1939-1.” / QR-code label with unique number “NHMUK010292545”, gen. prep. No.: NHMUK014331251 (prepared by
<location id="8ED26024FFE6666CFC044DAEC53AFC4E" LSID="urn:lsid:plazi:treatment:03A487E9FFE6666CFF6C4D65C303F964:8ED26024FFE6666CFC044DAEC53AFC4E" box="[1023,1136,969,995]" country="Malawi" name="Volynkin" pageId="85" pageNumber="86" stateProvince="Mulanje">Volynkin</location>
) (
<collectionCode id="ED1CAE3AFFE6666CFB7C4DAEC5B7FC4E" box="[1159,1277,969,995]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="85" pageNumber="86" type="Museum">NHMUK</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3176574FFE6666CFF0E4D8CC2CBF9EF" pageId="85" pageNumber="86" type="diagnosis">
<paragraph id="8BB236FFFFE6666CFF0E4D8CC570FA52" blockId="85.[151,1437,901,1737]" pageId="85" pageNumber="86">
<emphasis id="B979EAEDFFE6666CFF0E4D8CC024FBA8" bold="true" box="[245,366,1003,1029]" pageId="85" pageNumber="86">Diagnosis</emphasis>
. The forewing length is 8.0 mm in the male
<typeStatus id="54B6885DFFE6666CFC2C4D8CC508FBA8" box="[983,1090,1003,1029]" pageId="85" pageNumber="86" type="holotype">holotype</typeStatus>
.
<taxonomicName id="4C0D4D7CFFE6666CFBAD4D8CC5B9FBA8" authorityName="Volynkin" authorityYear="2023" box="[1110,1267,1003,1029]" class="Insecta" family="Erebidae" genus="Siccia" kingdom="Animalia" order="Lepidoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="nyasa">
<emphasis id="B979EAEDFFE6666CFBAD4D8CC5B9FBA8" box="[1110,1267,1003,1029]" italics="true" pageId="85" pageNumber="86">Siccia nyasa</emphasis>
</taxonomicName>
is externally reminiscent of
<taxonomicName id="4C0D4D7CFFE6666CFEB64A6BC08FFB8B" authorityName="Volynkin" authorityYear="2023" box="[333,453,1036,1062]" class="Insecta" family="Erebidae" genus="Siccia" kingdom="Animalia" order="Lepidoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="hackeri">
<emphasis id="B979EAEDFFE6666CFEB64A6BC08FFB8B" box="[333,453,1036,1062]" italics="true" pageId="85" pageNumber="86">S. hackeri</emphasis>
</taxonomicName>
but can be distinguished from it by the broader forewing with a somewhat more convex costal margin, the paler, creamy forewing ground colour (it is pale ochreous in the congener), the larger spots of the antemedial line, the larger discal spot, and the considerably paler, greyish hindwing whereas it is brownish-grey in
<taxonomicName id="4C0D4D7CFFE6666CFDF64A15C3CFFB21" authorityName="Volynkin" authorityYear="2023" box="[525,645,1138,1164]" class="Insecta" family="Erebidae" genus="Siccia" kingdom="Animalia" order="Lepidoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="hackeri">
<emphasis id="B979EAEDFFE6666CFDF64A15C3CFFB21" box="[525,645,1138,1164]" italics="true" pageId="85" pageNumber="86">S. hackeri</emphasis>
</taxonomicName>
. The male genital capsule of the new species is reminiscent of
<taxonomicName id="4C0D4D7CFFE6666CFA7D4A15C1A5FB00" authorityName="Volynkin" authorityYear="2023" class="Insecta" family="Erebidae" genus="Siccia" kingdom="Animalia" order="Lepidoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="hackeri">
<emphasis id="B979EAEDFFE6666CFA7D4A15C1A5FB00" italics="true" pageId="85" pageNumber="86">S. hackeri</emphasis>
</taxonomicName>
due to the lack of the penicular setae and small subuncal lobes, but differs clearly from it in the medially concave costa lacking the medial processes but bearing a cluster of spinules proximally. The sacculur structure of
<taxonomicName id="4C0D4D7CFFE6666CFE6F4AB0C0B2FB5C" authorityName="Volynkin" authorityYear="2023" box="[404,504,1239,1265]" class="Insecta" family="Erebidae" genus="Siccia" kingdom="Animalia" order="Lepidoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="nyasa">
<emphasis id="B979EAEDFFE6666CFE6F4AB0C0B2FB5C" box="[404,504,1239,1265]" italics="true" pageId="85" pageNumber="86">S. nyasa</emphasis>
</taxonomicName>
is strongly different from
<taxonomicName id="4C0D4D7CFFE6666CFCC34AB0C2FAFB5C" authorityName="Volynkin" authorityYear="2023" box="[824,944,1239,1265]" class="Insecta" family="Erebidae" genus="Siccia" kingdom="Animalia" order="Lepidoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="hackeri">
<emphasis id="B979EAEDFFE6666CFCC34AB0C2FAFB5C" box="[824,944,1239,1265]" italics="true" pageId="85" pageNumber="86">S. hackeri</emphasis>
</taxonomicName>
and is similar to that of
<taxonomicName id="4C0D4D7CFFE6666CFB274AB0C43CFB5C" authorityName="Romieux" authorityYear="1937" box="[1244,1398,1239,1265]" class="Insecta" family="Erebidae" genus="Siccia" kingdom="Animalia" order="Lepidoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="bicolorata">
<emphasis id="B979EAEDFFE6666CFB274AB0C43CFB5C" box="[1244,1398,1239,1265]" italics="true" pageId="85" pageNumber="86">S. bicolorata</emphasis>
</taxonomicName>
,
<taxonomicName id="4C0D4D7CFFE6666CFA7D4AB0C1B9FABF" class="Insecta" family="Erebidae" genus="Siccia" kingdom="Animalia" order="Lepidoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="mukuyu">
<emphasis id="B979EAEDFFE6666CFA7D4AB0C1B9FABF" italics="true" pageId="85" pageNumber="86">S. mukuyu</emphasis>
</taxonomicName>
,
<taxonomicName id="4C0D4D7CFFE6666CFEF94A9FC0C5FABF" box="[258,399,1272,1298]" class="Insecta" family="Erebidae" genus="Siccia" kingdom="Animalia" order="Lepidoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="kundalila">
<emphasis id="B979EAEDFFE6666CFEF94A9FC0C5FABF" box="[258,399,1272,1298]" italics="true" pageId="85" pageNumber="86">S. kundalila</emphasis>
</taxonomicName>
and
<taxonomicName id="4C0D4D7CFFE6666CFE314A9FC37AFABF" authorityName="Volynkin" authorityYear="2023" box="[458,560,1272,1298]" class="Insecta" family="Erebidae" genus="Siccia" kingdom="Animalia" order="Lepidoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="lydiae">
<emphasis id="B979EAEDFFE6666CFE314A9FC37AFABF" box="[458,560,1272,1298]" italics="true" pageId="85" pageNumber="86">S. lydiae</emphasis>
</taxonomicName>
due to the presence of the proximal process and the narrow distal process but the latter one is more elongate and directed distally in the current new species whereas those processes are directed ventrally in the aforementioned congeners. Unlike
<taxonomicName id="4C0D4D7CFFE6666CFC794B5BC2B2FAFB" authorityName="Volynkin" authorityYear="2023" box="[898,1016,1340,1366]" class="Insecta" family="Erebidae" genus="Siccia" kingdom="Animalia" order="Lepidoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="hackeri">
<emphasis id="B979EAEDFFE6666CFC794B5BC2B2FAFB" box="[898,1016,1340,1366]" italics="true" pageId="85" pageNumber="86">S. hackeri</emphasis>
</taxonomicName>
, the distal section of the costa of
<taxonomicName id="4C0D4D7CFFE6666CFA7C4B5BC196FAD5" authorityName="Volynkin" authorityYear="2023" class="Insecta" family="Erebidae" genus="Siccia" kingdom="Animalia" order="Lepidoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="nyasa">
<emphasis id="B979EAEDFFE6666CFA7C4B5BC196FAD5" italics="true" pageId="85" pageNumber="86">S. nyasa</emphasis>
</taxonomicName>
is considerably narrower bearing a cluster of markedly smaller spines apically. Additionally, compared to
<taxonomicName id="4C0D4D7CFFE6666CFF4D4B18C061FA34" authorityName="Volynkin" authorityYear="2023" box="[182,299,1407,1433]" class="Insecta" family="Erebidae" genus="Siccia" kingdom="Animalia" order="Lepidoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="hackeri">
<emphasis id="B979EAEDFFE6666CFF4D4B18C061FA34" box="[182,299,1407,1433]" italics="true" pageId="85" pageNumber="86">S. hackeri</emphasis>
</taxonomicName>
, the uncus of
<taxonomicName id="4C0D4D7CFFE6666CFE294B18C37EFA34" authorityName="Volynkin" authorityYear="2023" box="[466,564,1407,1433]" class="Insecta" family="Erebidae" genus="Siccia" kingdom="Animalia" order="Lepidoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="nyasa">
<emphasis id="B979EAEDFFE6666CFE294B18C37EFA34" box="[466,564,1407,1433]" italics="true" pageId="85" pageNumber="86">S. nyasa</emphasis>
</taxonomicName>
is shorter, and the anellus lacks processes. The phallus of the new species is shorter and broader than in
<taxonomicName id="4C0D4D7CFFE6666CFDE74BC6C3DDFA16" authorityName="Volynkin" authorityYear="2023" box="[540,663,1441,1467]" class="Insecta" family="Erebidae" genus="Siccia" kingdom="Animalia" order="Lepidoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="hackeri">
<emphasis id="B979EAEDFFE6666CFDE74BC6C3DDFA16" box="[540,663,1441,1467]" italics="true" pageId="85" pageNumber="86">S. hackeri</emphasis>
</taxonomicName>
with a considerably longer carina. The vesica of
<taxonomicName id="4C0D4D7CFFE6666CFAE84BC6C431FA16" authorityName="Volynkin" authorityYear="2023" box="[1299,1403,1441,1467]" class="Insecta" family="Erebidae" genus="Siccia" kingdom="Animalia" order="Lepidoptera" pageId="85" pageNumber="86" phylum="Arthropoda" rank="species" species="nyasa">
<emphasis id="B979EAEDFFE6666CFAE84BC6C431FA16" box="[1299,1403,1441,1467]" italics="true" pageId="85" pageNumber="86">S. nyasa</emphasis>
</taxonomicName>
is significantly broader than in the similar congener and has a large, trilobate dorsal diverticulum, a longer and narrower distal diverticulum, and bears a shorter and thinner terminal cornutus.
</paragraph>
<paragraph id="8BB236FFFFE6666CFF0E4861C358F98D" blockId="85.[151,1437,901,1737]" box="[245,530,1542,1568]" pageId="85" pageNumber="86">The female is unknown.</paragraph>
<paragraph id="8BB236FFFFE6666CFF0E484FC2CBF9EF" blockId="85.[151,1437,901,1737]" box="[245,897,1576,1602]" pageId="85" pageNumber="86">
<emphasis id="B979EAEDFFE6666CFF0E484FC0FFF9EF" bold="true" box="[245,437,1576,1602]" pageId="85" pageNumber="86">Molecular data</emphasis>
. Currently unavailable for this species.
</paragraph>
</subSubSection>
<subSubSection id="C3176574FFE6666CFF0E482DC052F92B" pageId="85" pageNumber="86" type="distribution">
<paragraph id="8BB236FFFFE6666CFF0E482DC052F92B" blockId="85.[151,1437,901,1737]" pageId="85" pageNumber="86">
<emphasis id="B979EAEDFFE6666CFF0E482DC0C5F9C9" bold="true" box="[245,399,1610,1636]" pageId="85" pageNumber="86">Distribution</emphasis>
. The new species is currently known only from its
<typeStatus id="54B6885DFFE6666CFC0B482DC569F9C9" box="[1008,1059,1610,1636]" pageId="85" pageNumber="86">type</typeStatus>
locality in the Southern Region of
<collectingCountry id="F31A766FFFE6666CFF4C480BC05EF92B" box="[183,276,1644,1670]" name="Malawi" pageId="85" pageNumber="86">Malawi</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C3176574FFE6666CFF0E48EAC303F964" pageId="85" pageNumber="86" type="etymology">
<paragraph id="8BB236FFFFE6666CFF0E48EAC303F964" blockId="85.[151,1437,901,1737]" pageId="85" pageNumber="86">
<emphasis id="B979EAEDFFE6666CFF0E48EAC037F90A" bold="true" box="[245,381,1677,1703]" pageId="85" pageNumber="86">Etymology</emphasis>
. The new species is named after the Nyasa people inhabiting
<collectingCountry id="F31A766FFFE6666CFB7148EAC5ADF90A" box="[1162,1255,1677,1703]" name="Malawi" pageId="85" pageNumber="86">Malawi</collectingCountry>
, south-western
<collectingCountry id="F31A766FFFE6666CFF6C48C8C049F964" box="[151,259,1711,1737]" name="Tanzania" pageId="85" pageNumber="86">Tanzania</collectingCountry>
and northern
<collectingCountry id="F31A766FFFE6666CFE5C48C8C30CF964" box="[423,582,1711,1737]" name="Mozambique" pageId="85" pageNumber="86">Mozambique</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>