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<mods:title id="C3AA008DCFDD6E77B99028043E1A1241">The South American genus Quemedice Mello-Leitão (Araneae: Sparassidae): familial placement and taxonomic revision</mods:title>
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<mods:namePart id="642159D3D603D78FC8C0188A9D69C98A">Rheims, Cristina A.</mods:namePart>
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<treatment id="03A487D4FFFDFFE1AC90CA586D2AFDB5" ID-DOI="http://doi.org/10.5281/zenodo.6228516" ID-GBIF-Taxon="119370581" ID-Zenodo-Dep="6228516" LSID="urn:lsid:plazi:treatment:03A487D4FFFDFFE1AC90CA586D2AFDB5" httpUri="http://treatment.plazi.org/id/03A487D4FFFDFFE1AC90CA586D2AFDB5" lastPageId="2" lastPageNumber="62" pageId="1" pageNumber="61">
<subSubSection id="C3176549FFFDFFE2AC90CA586E27FCC3" box="[151,528,890,916]" pageId="1" pageNumber="61" type="nomenclature">
<paragraph id="8BB236C2FFFDFFE2AC90CA586E27FCC3" blockId="1.[151,528,890,916]" box="[151,528,890,916]" pageId="1" pageNumber="61">
<heading id="D0FA81AEFFFDFFE2AC90CA586E27FCC3" bold="true" box="[151,528,890,916]" fontSize="11" level="1" pageId="1" pageNumber="61" reason="1">
<taxonomicName id="4C0D4D41FFFDFFE2AC90CA586E27FCC3" ID-CoL="76H7" ID-ENA="1956527" authority="Mello-Leitao, 1942" authorityName="Mello-Leitao" authorityYear="1942" box="[151,528,890,916]" class="Arachnida" family="Sparassidae" genus="Quemedice" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="61" phylum="Arthropoda" rank="genus">
<emphasis id="B979EAD0FFFDFFE2AC90CA586E27FCC3" bold="true" box="[151,528,890,916]" pageId="1" pageNumber="61">
<emphasis id="B979EAD0FFFDFFE2AC90CA586D29FCC3" bold="true" box="[151,286,890,916]" italics="true" pageId="1" pageNumber="61">Quemedice</emphasis>
<bibRefCitation id="EF9C4B33FFFDFFE2AD22CA586E27FCC3" author="Mello-Leitao" box="[293,528,890,916]" lastPageId="1" lastPageNumber="68" pageId="1" pageNumber="67" refString="Mello-Leitao, C. F. de. (1942) Aranas del Chaco y Santiago del Estero. Revista del Museo de La Plata (N. S., Zool.), 2, 381 - 426." type="journal article" year="1942">Mello-Leitão, 1942</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="C3176549FFFDFFE2AC90CAE26EB3FB40" pageId="1" pageNumber="61" type="materials_examined">
<paragraph id="8BB236C2FFFDFFE2AC90CAE26EB3FB40" blockId="1.[151,1436,960,1047]" pageId="1" pageNumber="61">
<typeStatus id="54B68860FFFDFFE2AC90CAE26CFBFC80" box="[151,204,960,983]" pageId="1" pageNumber="61">Type</typeStatus>
species:
<taxonomicName id="4C0D4D41FFFDFFE2AD2ACAE26F31FC80" authority="Mello-Leitao, 1942" authorityName="Mello-Leitao" authorityYear="1942" box="[301,774,960,983]" class="Arachnida" family="Sparassidae" genus="Quemedice" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="61" phylum="Arthropoda" rank="species" species="enigmaticus">
<emphasis id="B979EAD0FFFDFFE2AD2ACAE26E1AFC80" box="[301,557,960,983]" italics="true" pageId="1" pageNumber="61">Quemedice enigmaticus</emphasis>
<bibRefCitation id="EF9C4B33FFFDFFE2AE33CAE26F31FC80" author="Mello-Leitao" box="[564,774,960,983]" lastPageId="1" lastPageNumber="68" pageId="1" pageNumber="67" refString="Mello-Leitao, C. F. de. (1942) Aranas del Chaco y Santiago del Estero. Revista del Museo de La Plata (N. S., Zool.), 2, 381 - 426." type="journal article" year="1942">Mello-Leitão, 1942</bibRefCitation>
</taxonomicName>
, by original designation
<taxonomicName id="4C0D4D41FFFDFFE2AC90CAC26E2AFCA0" authority="Mello-Leitao, 1942: 407" authorityName="Mello-Leitao" authorityPageNumber="407" authorityYear="1942" box="[151,541,992,1015]" class="Arachnida" family="Sparassidae" genus="Quemedice" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="61" phylum="Arthropoda" rank="genus">
<emphasis id="B979EAD0FFFDFFE2AC90CAC26D38FCA0" box="[151,271,992,1015]" italics="true" pageId="1" pageNumber="61">Quemedice</emphasis>
<bibRefCitation id="EF9C4B33FFFDFFE2AD11CAC26DDEFCA0" author="Mello-Leitao" box="[278,489,992,1015]" lastPageId="1" lastPageNumber="68" pageId="1" pageNumber="67" refString="Mello-Leitao, C. F. de. (1942) Aranas del Chaco y Santiago del Estero. Revista del Museo de La Plata (N. S., Zool.), 2, 381 - 426." type="journal article" year="1942">Mello-Leitão, 1942</bibRefCitation>
: 407
</taxonomicName>
408;
<bibRefCitation id="EF9C4B33FFFDFFE2AE58CAC26ECFFCA0" author="Roewer" box="[607,760,992,1015]" pageId="1" pageNumber="68" refString="Roewer, C. F. (1954) Katalog der Araneae von 1758 bis 1940, vol. 1. Institut Royal des Sciences Naturelles de Belgique, Bruxelles, 923 pp." type="book" year="1954">Roewer, 1954</bibRefCitation>
: 790 (
<taxonomicName id="4C0D4D41FFFDFFE2AF3ECAC26F8AFCA0" box="[825,957,992,1015]" class="Arachnida" family="Thomisidae" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="61" phylum="Arthropoda" rank="family">Thomisidae</taxonomicName>
);
<bibRefCitation id="EF9C4B33FFFDFFE2AFD7CAC26845FCA0" author="Homann" box="[976,1138,992,1015]" pageId="1" pageNumber="67" refString="Homann, H. (1975) Die Stellung der Thomisidae und der Philodromidae im System der Araneae (Chelicerata, Arachnida). Zeitschrift fur Morphologie und Okologie der Tiere, 80, 181 - 202." type="journal article" year="1975">Homann, 1975</bibRefCitation>
: 183 (transf. to
<taxonomicName id="4C0D4D41FFFDFFE2A91CCAC26CC7FB40" class="Arachnida" family="Philodromidae" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="61" phylum="Arthropoda" rank="family">Philodromidae</taxonomicName>
);
<bibRefCitation id="EF9C4B33FFFDFFE2AD04CD226D95FB40" author="Brignoli" box="[259,418,1024,1047]" pageId="1" pageNumber="67" refString="Brignoli, P. M. (1983) A catalogue of the Araneae described between 1940 and 1981, Manchester University Press, Manchester, 755 pp." type="book" year="1983">Brignoli, 1983</bibRefCitation>
: 596;
<bibRefCitation id="EF9C4B33FFFDFFE2ADE5CD226EB7FB40" author="Platnick" box="[482,640,1024,1047]" pageId="1" pageNumber="68" refString="Platnick, N. I. (2008) The world spider catalog, version 8.5, American Museum of Natural History. Available from: http: / / research. amnh. org / entomology / spiders / catalog / index. html (accessed 30 March 2008)." type="journal article" year="2008">Platnick, 2008</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3176549FFFDFFE2AC90CD606871FBAB" pageId="1" pageNumber="61" type="diagnosis">
<paragraph id="8BB236C2FFFDFFE2AC90CD606871FBAB" blockId="1.[151,1437,1090,2036]" pageId="1" pageNumber="61">
<emphasis id="B979EAD0FFFDFFE2AC90CD606D27FB0B" bold="true" box="[151,272,1090,1116]" pageId="1" pageNumber="61">Diagnosis</emphasis>
. Distinguished from remaining sparassid genera by the combination of the following characters: elongate opisthosoma (
<figureCitation id="13362A47FFFDFFE2ADA3CD486E7EFBD3" box="[420,585,1130,1156]" captionStart="FIGURES 1 7" captionStartId="3.[151,269,1780,1804]" captionTargetBox="[264,1323,263,1753]" captionTargetId="figure@3.[264,1323,263,1754]" captionTargetPageId="3" captionText="FIGURES 1 7. Quemedice enigmaticus Mello Leitão, preserved specimens. 1 4 Female (holotype, MLP 15450), 1 3 habitus (1 dorsal; 2 ventral; 3 lateral), 4 Epigynum, ventral; 5 7. Male (MACN Ar 10758), habitus (5 dorsal; 6 lateral, 7 ventral). Scale bars: 1 2, 4 7, 2 mm; 3, 0.25 mm." httpUri="https://zenodo.org/record/274360/files/figure.png" pageId="1" pageNumber="61">Figs 12, 47</figureCitation>
), male palp with massive embolus retrolaterally bent with median region curved dorsad and behind the base and emerging dorsally to rest behind an apophysis at the base of the embolus (
<figureCitation id="13362A47FFFDFFE2ACCCCD986D93FB83" box="[203,420,1210,1236]" captionStart="FIGURES 8 16. 8 12" captionStartId="4.[151,269,1715,1739]" captionTargetBox="[231,1365,476,1684]" captionTargetId="figure@4.[231,1366,474,1687]" captionTargetPageId="4" captionText="FIGURES 8 16. 8 12. Quemedice enigmaticus Mello Leitão (8 9 IBSP 67822; 10 MACN Ar 10758; 11 12 IBSP 1899). 8 10 Left male palp (8 ventral, 9 retrolateral, 10 ditto, detail of expanded embolus); 11 12. Epigynum (11 ventral, 12 dorsal). 13 16. Quemedice piracuruca n. sp. (13 14, paratype, IBSP 28565; 15 16, paratype, MPEG 6402). 13 14 Left male palp (arrows indicate small teeth) (13 ventral, 14 dorsal); 15 16. Epigynum (15 ventral, 16 dorsal). (C = cymbium; CD = copulatory duct; CO = copulatory opening; db = dorsal branch of RTA; EA = embolus apophysis; EB = embolus base; E = embolus; FD = fertilization duct; HR = hyaline receptacle; MS = median septum; LL = lateral lobes; ST = subtegulum; T = tegulum; vb = ventral branch of RTA). Scale bars: 8 9, 11 16, 0.5 mm; 10, 0.1 mm." httpUri="https://zenodo.org/record/274361/files/figure.png" pageId="1" pageNumber="61">Figs 810, 1314</figureCitation>
). Females are distinguished by having the copulatory ducts distally convoluted with anterior membranous receptacles and inconspicuous spermathecae (
<figureCitation id="13362A47FFFDFFE2AFB6CDC0680FFBAB" box="[945,1080,1250,1276]" captionStart="FIGURES 8 16. 8 12" captionStartId="4.[151,269,1715,1739]" captionTargetBox="[231,1365,476,1684]" captionTargetId="figure@4.[231,1366,474,1687]" captionTargetPageId="4" captionText="FIGURES 8 16. 8 12. Quemedice enigmaticus Mello Leitão (8 9 IBSP 67822; 10 MACN Ar 10758; 11 12 IBSP 1899). 8 10 Left male palp (8 ventral, 9 retrolateral, 10 ditto, detail of expanded embolus); 11 12. Epigynum (11 ventral, 12 dorsal). 13 16. Quemedice piracuruca n. sp. (13 14, paratype, IBSP 28565; 15 16, paratype, MPEG 6402). 13 14 Left male palp (arrows indicate small teeth) (13 ventral, 14 dorsal); 15 16. Epigynum (15 ventral, 16 dorsal). (C = cymbium; CD = copulatory duct; CO = copulatory opening; db = dorsal branch of RTA; EA = embolus apophysis; EB = embolus base; E = embolus; FD = fertilization duct; HR = hyaline receptacle; MS = median septum; LL = lateral lobes; ST = subtegulum; T = tegulum; vb = ventral branch of RTA). Scale bars: 8 9, 11 16, 0.5 mm; 10, 0.1 mm." httpUri="https://zenodo.org/record/274361/files/figure.png" pageId="1" pageNumber="61">Figs 12, 16</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3176549FFFDFFE2ACC2CC286DD9F8A3" pageId="1" pageNumber="61" type="description">
<paragraph id="8BB236C2FFFDFFE2ACC2CC286883F853" blockId="1.[151,1437,1090,2036]" pageId="1" pageNumber="61">
<emphasis id="B979EAD0FFFDFFE2ACC2CC286D62FA73" bold="true" box="[197,341,1290,1316]" pageId="1" pageNumber="61">Description</emphasis>
. Prosoma longer than wide (
<figureCitation id="13362A47FFFDFFE2AEA8CC286F2BFA73" box="[687,796,1290,1316]" captionStart="FIGURES 1 7" captionStartId="3.[151,269,1780,1804]" captionTargetBox="[264,1323,263,1753]" captionTargetId="figure@3.[264,1323,263,1754]" captionTargetPageId="3" captionText="FIGURES 1 7. Quemedice enigmaticus Mello Leitão, preserved specimens. 1 4 Female (holotype, MLP 15450), 1 3 habitus (1 dorsal; 2 ventral; 3 lateral), 4 Epigynum, ventral; 5 7. Male (MACN Ar 10758), habitus (5 dorsal; 6 lateral, 7 ventral). Scale bars: 1 2, 4 7, 2 mm; 3, 0.25 mm." httpUri="https://zenodo.org/record/274360/files/figure.png" pageId="1" pageNumber="61">Figs 1, 5</figureCitation>
); cephalic region slightly higher than thoracic region. Eyes arranged in two recurved rows; AME slightly larger than ALE and farther apart from each other than from laterals; PME slightly smaller than PLE, equidistant (
<figureCitation id="13362A47FFFDFFE2AF5DCC786FF0FA23" box="[858,967,1370,1396]" captionStart="FIGURES 1 7" captionStartId="3.[151,269,1780,1804]" captionTargetBox="[264,1323,263,1753]" captionTargetId="figure@3.[264,1323,263,1754]" captionTargetPageId="3" captionText="FIGURES 1 7. Quemedice enigmaticus Mello Leitão, preserved specimens. 1 4 Female (holotype, MLP 15450), 1 3 habitus (1 dorsal; 2 ventral; 3 lateral), 4 Epigynum, ventral; 5 7. Male (MACN Ar 10758), habitus (5 dorsal; 6 lateral, 7 ventral). Scale bars: 1 2, 4 7, 2 mm; 3, 0.25 mm." httpUri="https://zenodo.org/record/274360/files/figure.png" pageId="1" pageNumber="61">Figs 1, 5</figureCitation>
). Chelicerae with two promarginal and two retromarginal teeth; intermarginal denticles absent (
<figureCitation id="13362A47FFFDFFE2AF36CCA06F8BFACB" box="[817,956,1410,1436]" captionStart="FIGURES 17 22" captionStartId="5.[151,269,1693,1717]" captionTargetBox="[264,1322,389,1669]" captionTargetId="figure@5.[264,1323,389,1670]" captionTargetPageId="5" captionText="FIGURES 17 22. Quemedice enigmaticus Mello Leitão (MACN Ar 10758). 17 22 Left chelicera (17 posterior, 18 retromargin, 19 ditto, detail of cheliceral gland area, 20 ectal, 21 frontal, 22 promargin) (CB = cheliceral boss; CGA = cheliceral gland area; M = macrosetae). Scale bars: 17, 20 21, 200 µm; 18, 22, 100 µm; 19, 50 µm." httpUri="https://zenodo.org/record/274362/files/figure.png" pageId="1" pageNumber="61">Figs 1722</figureCitation>
). Internal keel at base of fang with 8-10 strong setae in one or two rows (
<figureCitation id="13362A47FFFDFFE2AE24CC886F3EFA93" box="[547,777,1450,1476]" captionStart="FIGURES 17 22" captionStartId="5.[151,269,1693,1717]" captionTargetBox="[264,1322,389,1669]" captionTargetId="figure@5.[264,1323,389,1670]" captionTargetPageId="5" captionText="FIGURES 17 22. Quemedice enigmaticus Mello Leitão (MACN Ar 10758). 17 22 Left chelicera (17 posterior, 18 retromargin, 19 ditto, detail of cheliceral gland area, 20 ectal, 21 frontal, 22 promargin) (CB = cheliceral boss; CGA = cheliceral gland area; M = macrosetae). Scale bars: 17, 20 21, 200 µm; 18, 22, 100 µm; 19, 50 µm." httpUri="https://zenodo.org/record/274362/files/figure.png" pageId="1" pageNumber="61">Figs 1718, 2122</figureCitation>
). Cheliceral boss on ectal margin (
<figureCitation id="13362A47FFFDFFE2A8AECC886957FA93" box="[1193,1376,1450,1476]" captionStart="FIGURES 17 22" captionStartId="5.[151,269,1693,1717]" captionTargetBox="[264,1322,389,1669]" captionTargetId="figure@5.[264,1323,389,1670]" captionTargetPageId="5" captionText="FIGURES 17 22. Quemedice enigmaticus Mello Leitão (MACN Ar 10758). 17 22 Left chelicera (17 posterior, 18 retromargin, 19 ditto, detail of cheliceral gland area, 20 ectal, 21 frontal, 22 promargin) (CB = cheliceral boss; CGA = cheliceral gland area; M = macrosetae). Scale bars: 17, 20 21, 200 µm; 18, 22, 100 µm; 19, 50 µm." httpUri="https://zenodo.org/record/274362/files/figure.png" pageId="1" pageNumber="61">Figs 17, 2021</figureCitation>
) and gland area on retromargin (
<figureCitation id="13362A47FFFDFFE2ADDACCF06E02FABB" box="[477,565,1490,1516]" captionStart="FIGURES 17 22" captionStartId="5.[151,269,1693,1717]" captionTargetBox="[264,1322,389,1669]" captionTargetId="figure@5.[264,1323,389,1670]" captionTargetPageId="5" captionText="FIGURES 17 22. Quemedice enigmaticus Mello Leitão (MACN Ar 10758). 17 22 Left chelicera (17 posterior, 18 retromargin, 19 ditto, detail of cheliceral gland area, 20 ectal, 21 frontal, 22 promargin) (CB = cheliceral boss; CGA = cheliceral gland area; M = macrosetae). Scale bars: 17, 20 21, 200 µm; 18, 22, 100 µm; 19, 50 µm." httpUri="https://zenodo.org/record/274362/files/figure.png" pageId="1" pageNumber="61">Fig. 19</figureCitation>
). Abdomen elongate, three times longer than wide (
<figureCitation id="13362A47FFFDFFE2A8A4CCF0697AFABB" box="[1187,1357,1490,1516]" captionStart="FIGURES 1 7" captionStartId="3.[151,269,1780,1804]" captionTargetBox="[264,1323,263,1753]" captionTargetId="figure@3.[264,1323,263,1754]" captionTargetPageId="3" captionText="FIGURES 1 7. Quemedice enigmaticus Mello Leitão, preserved specimens. 1 4 Female (holotype, MLP 15450), 1 3 habitus (1 dorsal; 2 ventral; 3 lateral), 4 Epigynum, ventral; 5 7. Male (MACN Ar 10758), habitus (5 dorsal; 6 lateral, 7 ventral). Scale bars: 1 2, 4 7, 2 mm; 3, 0.25 mm." httpUri="https://zenodo.org/record/274360/files/figure.png" pageId="1" pageNumber="61">Figs 12, 47</figureCitation>
). Legs laterigrade (2413). Leg spination in males: femora IIV p1-1-1; d0-1-1; r0-1-1; patellae III p1; IIIIV d1; tibiae IIV p1-0-1; d1-0-1; r1-0-1;
<date id="FFB31002FFFDFFE2AE3DCF006E44F96B" box="[570,627,1570,1596]" pageId="1" pageNumber="61">v2-2</date>
-0; metatarsi IIII p1-1-0; r1-1-0;
<date id="FFB31002FFFDFFE2A801CF006808F96B" box="[1030,1087,1570,1596]" pageId="1" pageNumber="61">v2-2</date>
-0; IV p1-1-2; r1-1-2;
<date id="FFB31002FFFDFFE2A941CF006948F96B" box="[1350,1407,1570,1596]" pageId="1" pageNumber="61">v2-2</date>
-0. Leg spination in females as in males, except patellae IIV 0; tibiae IIV d0. Metatarsi IIV with dorsal trilobate membrane at distal end, with median lobe slightly more developed than laterals (
<figureCitation id="13362A47FFFDFFE2A897CF50692DF9DB" box="[1168,1306,1650,1676]" captionStart="FIGURES 23 28" captionStartId="6.[151,269,1943,1967]" captionTargetBox="[233,1367,545,1917]" captionTargetId="figure@6.[233,1368,545,1917]" captionTargetPageId="6" captionText="FIGURES 23 28. Quemedice enigmaticus Mello Leitão (MACN Ar 10758). 23 27 Male, left leg IV (23 claw, retrolateral apical, 24 tarsal trichobothria, dorsal, 25 tarsal organ, dorsal, 26 trilobate membrane at distal end of metatarsus, prolateral, 27 ditto, dorsal; 28 Epiandrium, ventral view. Scale bars: 23, 26 27, 100 µm; 28, 50 µm; 25, 20 µm; 24, 10 µm." httpUri="https://zenodo.org/record/274363/files/figure.png" pageId="1" pageNumber="61">Figs 2627</figureCitation>
). Tarsi and anterior half of metatarsi densely scopulated. Tarsal organ capsulated, with rounded opening, located dorsally at the distal end of tarsi (
<figureCitation id="13362A47FFFDFFE2ADB1CFE06E39F98B" box="[438,526,1730,1756]" captionStart="FIGURES 23 28" captionStartId="6.[151,269,1943,1967]" captionTargetBox="[233,1367,545,1917]" captionTargetId="figure@6.[233,1368,545,1917]" captionTargetPageId="6" captionText="FIGURES 23 28. Quemedice enigmaticus Mello Leitão (MACN Ar 10758). 23 27 Male, left leg IV (23 claw, retrolateral apical, 24 tarsal trichobothria, dorsal, 25 tarsal organ, dorsal, 26 trilobate membrane at distal end of metatarsus, prolateral, 27 ditto, dorsal; 28 Epiandrium, ventral view. Scale bars: 23, 26 27, 100 µm; 28, 50 µm; 25, 20 µm; 24, 10 µm." httpUri="https://zenodo.org/record/274363/files/figure.png" pageId="1" pageNumber="61">Fig. 25</figureCitation>
). Trichobothrium with basal crescent plate projecting over smooth distal plate (
<figureCitation id="13362A47FFFDFFE2AC98CFC86CC0F853" box="[159,247,1770,1796]" captionStart="FIGURES 23 28" captionStartId="6.[151,269,1943,1967]" captionTargetBox="[233,1367,545,1917]" captionTargetId="figure@6.[233,1368,545,1917]" captionTargetPageId="6" captionText="FIGURES 23 28. Quemedice enigmaticus Mello Leitão (MACN Ar 10758). 23 27 Male, left leg IV (23 claw, retrolateral apical, 24 tarsal trichobothria, dorsal, 25 tarsal organ, dorsal, 26 trilobate membrane at distal end of metatarsus, prolateral, 27 ditto, dorsal; 28 Epiandrium, ventral view. Scale bars: 23, 26 27, 100 µm; 28, 50 µm; 25, 20 µm; 24, 10 µm." httpUri="https://zenodo.org/record/274363/files/figure.png" pageId="1" pageNumber="61">Fig. 24</figureCitation>
). Tarsi with two pectinated claws, with ca. 25 teeth each, and claw tufts (
<figureCitation id="13362A47FFFDFFE2A848CFC86890F853" box="[1103,1191,1770,1796]" captionStart="FIGURES 23 28" captionStartId="6.[151,269,1943,1967]" captionTargetBox="[233,1367,545,1917]" captionTargetId="figure@6.[233,1368,545,1917]" captionTargetPageId="6" captionText="FIGURES 23 28. Quemedice enigmaticus Mello Leitão (MACN Ar 10758). 23 27 Male, left leg IV (23 claw, retrolateral apical, 24 tarsal trichobothria, dorsal, 25 tarsal organ, dorsal, 26 trilobate membrane at distal end of metatarsus, prolateral, 27 ditto, dorsal; 28 Epiandrium, ventral view. Scale bars: 23, 26 27, 100 µm; 28, 50 µm; 25, 20 µm; 24, 10 µm." httpUri="https://zenodo.org/record/274363/files/figure.png" pageId="1" pageNumber="61">Fig. 23</figureCitation>
).
</paragraph>
<paragraph id="8BB236C2FFFDFFE2ACC2CE316DD9F8A3" blockId="1.[151,1437,1090,2036]" pageId="1" pageNumber="61">
<emphasis id="B979EAD0FFFDFFE2ACC2CE316D09F87B" box="[197,318,1811,1836]" italics="true" pageId="1" pageNumber="61">Male palp</emphasis>
. Tibia short with one strong basal spine prolaterally and one dorsally (
<figureCitation id="13362A47FFFDFFE2A88ECE306967F87B" box="[1161,1360,1810,1836]" captionStart="FIGURES 8 16. 8 12" captionStartId="4.[151,269,1715,1739]" captionTargetBox="[231,1365,476,1684]" captionTargetId="figure@4.[231,1366,474,1687]" captionTargetPageId="4" captionText="FIGURES 8 16. 8 12. Quemedice enigmaticus Mello Leitão (8 9 IBSP 67822; 10 MACN Ar 10758; 11 12 IBSP 1899). 8 10 Left male palp (8 ventral, 9 retrolateral, 10 ditto, detail of expanded embolus); 11 12. Epigynum (11 ventral, 12 dorsal). 13 16. Quemedice piracuruca n. sp. (13 14, paratype, IBSP 28565; 15 16, paratype, MPEG 6402). 13 14 Left male palp (arrows indicate small teeth) (13 ventral, 14 dorsal); 15 16. Epigynum (15 ventral, 16 dorsal). (C = cymbium; CD = copulatory duct; CO = copulatory opening; db = dorsal branch of RTA; EA = embolus apophysis; EB = embolus base; E = embolus; FD = fertilization duct; HR = hyaline receptacle; MS = median septum; LL = lateral lobes; ST = subtegulum; T = tegulum; vb = ventral branch of RTA). Scale bars: 8 9, 11 16, 0.5 mm; 10, 0.1 mm." httpUri="https://zenodo.org/record/274361/files/figure.png" pageId="1" pageNumber="61">Figs 89, 1314</figureCitation>
). RTA distally bifid (
<figureCitation id="13362A47FFFDFFE2AD3ACE186E34F803" box="[317,515,1850,1876]" captionStart="FIGURES 8 16. 8 12" captionStartId="4.[151,269,1715,1739]" captionTargetBox="[231,1365,476,1684]" captionTargetId="figure@4.[231,1366,474,1687]" captionTargetPageId="4" captionText="FIGURES 8 16. 8 12. Quemedice enigmaticus Mello Leitão (8 9 IBSP 67822; 10 MACN Ar 10758; 11 12 IBSP 1899). 8 10 Left male palp (8 ventral, 9 retrolateral, 10 ditto, detail of expanded embolus); 11 12. Epigynum (11 ventral, 12 dorsal). 13 16. Quemedice piracuruca n. sp. (13 14, paratype, IBSP 28565; 15 16, paratype, MPEG 6402). 13 14 Left male palp (arrows indicate small teeth) (13 ventral, 14 dorsal); 15 16. Epigynum (15 ventral, 16 dorsal). (C = cymbium; CD = copulatory duct; CO = copulatory opening; db = dorsal branch of RTA; EA = embolus apophysis; EB = embolus base; E = embolus; FD = fertilization duct; HR = hyaline receptacle; MS = median septum; LL = lateral lobes; ST = subtegulum; T = tegulum; vb = ventral branch of RTA). Scale bars: 8 9, 11 16, 0.5 mm; 10, 0.1 mm." httpUri="https://zenodo.org/record/274361/files/figure.png" pageId="1" pageNumber="61">Figs 89, 1314</figureCitation>
). Cymbium with oval, median alveolus and two prolateral spines at base, one of them stronger. Subtegulum retrolaterally displaced and visible in ventral and retrolateral views. Tegulum Sshaped, slightly spiraled perpendicularly to the main palpal axis, towards the tip of the cymbium (
<figureCitation id="13362A47FFFDFFE2A92CCEA86CD3F89B" captionStart="FIGURES 8 16. 8 12" captionStartId="4.[151,269,1715,1739]" captionTargetBox="[231,1365,476,1684]" captionTargetId="figure@4.[231,1366,474,1687]" captionTargetPageId="4" captionText="FIGURES 8 16. 8 12. Quemedice enigmaticus Mello Leitão (8 9 IBSP 67822; 10 MACN Ar 10758; 11 12 IBSP 1899). 8 10 Left male palp (8 ventral, 9 retrolateral, 10 ditto, detail of expanded embolus); 11 12. Epigynum (11 ventral, 12 dorsal). 13 16. Quemedice piracuruca n. sp. (13 14, paratype, IBSP 28565; 15 16, paratype, MPEG 6402). 13 14 Left male palp (arrows indicate small teeth) (13 ventral, 14 dorsal); 15 16. Epigynum (15 ventral, 16 dorsal). (C = cymbium; CD = copulatory duct; CO = copulatory opening; db = dorsal branch of RTA; EA = embolus apophysis; EB = embolus base; E = embolus; FD = fertilization duct; HR = hyaline receptacle; MS = median septum; LL = lateral lobes; ST = subtegulum; T = tegulum; vb = ventral branch of RTA). Scale bars: 8 9, 11 16, 0.5 mm; 10, 0.1 mm." httpUri="https://zenodo.org/record/274361/files/figure.png" pageId="1" pageNumber="61">Figs 89, 1314</figureCitation>
). Conductor absent. Embolus massive, retrolaterally directed and medially curving behind base and emerging at the top (
<figureCitation id="13362A47FFFDFFE2AD8ECEF86DD6F8A3" box="[393,481,2010,2036]" captionStart="FIGURES 8 16. 8 12" captionStartId="4.[151,269,1715,1739]" captionTargetBox="[231,1365,476,1684]" captionTargetId="figure@4.[231,1366,474,1687]" captionTargetPageId="4" captionText="FIGURES 8 16. 8 12. Quemedice enigmaticus Mello Leitão (8 9 IBSP 67822; 10 MACN Ar 10758; 11 12 IBSP 1899). 8 10 Left male palp (8 ventral, 9 retrolateral, 10 ditto, detail of expanded embolus); 11 12. Epigynum (11 ventral, 12 dorsal). 13 16. Quemedice piracuruca n. sp. (13 14, paratype, IBSP 28565; 15 16, paratype, MPEG 6402). 13 14 Left male palp (arrows indicate small teeth) (13 ventral, 14 dorsal); 15 16. Epigynum (15 ventral, 16 dorsal). (C = cymbium; CD = copulatory duct; CO = copulatory opening; db = dorsal branch of RTA; EA = embolus apophysis; EB = embolus base; E = embolus; FD = fertilization duct; HR = hyaline receptacle; MS = median septum; LL = lateral lobes; ST = subtegulum; T = tegulum; vb = ventral branch of RTA). Scale bars: 8 9, 11 16, 0.5 mm; 10, 0.1 mm." httpUri="https://zenodo.org/record/274361/files/figure.png" pageId="1" pageNumber="61">Fig. 10</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3176549FFFEFFE1ACC2C9BA6E16FE2D" pageId="2" pageNumber="62" type="reference_group">
<paragraph id="8BB236C2FFFEFFE1ACC2C9BA6E16FE2D" blockId="2.[151,1436,152,738]" pageId="2" pageNumber="62">
<taxonomicName id="4C0D4D41FFFEFFE1ACC2C9BA6D0BFFE6" ID-CoL="4D55" box="[197,316,152,177]" class="Magnoliopsida" family="Apocynaceae" genus="Epigynum" kingdom="Plantae" order="Gentianales" pageId="2" pageNumber="62" phylum="Tracheophyta" rank="genus">
<emphasis id="B979EAD0FFFEFFE1ACC2C9BA6D0BFFE6" box="[197,316,152,177]" italics="true" pageId="2" pageNumber="62">Epigynum</emphasis>
</taxonomicName>
. Epigynal field divided into median septum and lateral lobes; lateral borders slightly rounded covering longer than wide median septum with pair of anterior copulatory openings (
<figureCitation id="13362A47FFFEFFE1A893C9E268D4FF8D" box="[1172,1251,192,218]" captionStart="FIGURES 1 7" captionStartId="3.[151,269,1780,1804]" captionTargetBox="[264,1323,263,1753]" captionTargetId="figure@3.[264,1323,263,1754]" captionTargetPageId="3" captionText="FIGURES 1 7. Quemedice enigmaticus Mello Leitão, preserved specimens. 1 4 Female (holotype, MLP 15450), 1 3 habitus (1 dorsal; 2 ventral; 3 lateral), 4 Epigynum, ventral; 5 7. Male (MACN Ar 10758), habitus (5 dorsal; 6 lateral, 7 ventral). Scale bars: 1 2, 4 7, 2 mm; 3, 0.25 mm." httpUri="https://zenodo.org/record/274360/files/figure.png" pageId="2" pageNumber="62">Figs 4</figureCitation>
,
<figureCitation id="13362A47FFFEFFE1A8F6C9E26909FF8D" box="[1265,1342,192,218]" captionStart="FIGURES 8 16. 8 12" captionStartId="4.[151,269,1715,1739]" captionTargetBox="[231,1365,476,1684]" captionTargetId="figure@4.[231,1366,474,1687]" captionTargetPageId="4" captionText="FIGURES 8 16. 8 12. Quemedice enigmaticus Mello Leitão (8 9 IBSP 67822; 10 MACN Ar 10758; 11 12 IBSP 1899). 8 10 Left male palp (8 ventral, 9 retrolateral, 10 ditto, detail of expanded embolus); 11 12. Epigynum (11 ventral, 12 dorsal). 13 16. Quemedice piracuruca n. sp. (13 14, paratype, IBSP 28565; 15 16, paratype, MPEG 6402). 13 14 Left male palp (arrows indicate small teeth) (13 ventral, 14 dorsal); 15 16. Epigynum (15 ventral, 16 dorsal). (C = cymbium; CD = copulatory duct; CO = copulatory opening; db = dorsal branch of RTA; EA = embolus apophysis; EB = embolus base; E = embolus; FD = fertilization duct; HR = hyaline receptacle; MS = median septum; LL = lateral lobes; ST = subtegulum; T = tegulum; vb = ventral branch of RTA). Scale bars: 8 9, 11 16, 0.5 mm; 10, 0.1 mm." httpUri="https://zenodo.org/record/274361/files/figure.png" pageId="2" pageNumber="62">11, 15</figureCitation>
). Vulva with copulatory ducts posteriorly convoluted, with large anterior membranous receptacles (“Subepigynealer Sack”
<emphasis id="B979EAD0FFFEFFE1ACECC8326D1BFE7E" box="[235,300,272,297]" italics="true" pageId="2" pageNumber="62">sensu</emphasis>
<bibRefCitation id="EF9C4B33FFFEFFE1AD3DC8326D89FE7D" author="Jarvi" box="[314,446,272,298]" pageId="2" pageNumber="67" refString="Jarvi, T. H. (1912) Das Vaginalsystem der Sparassiden. I. Allgemeiner Teil. Annales Academiae scientiarum Fennicae, 4, 1 - 131." type="journal article" year="1912">Järvi 1912</bibRefCitation>
, considered to be non-homologous to the &quot;spermathecae&quot;
<emphasis id="B979EAD0FFFEFFE1A89BC83268EAFE7E" box="[1180,1245,272,297]" italics="true" pageId="2" pageNumber="62">sensu</emphasis>
<bibRefCitation id="EF9C4B33FFFEFFE1A8ECC83269ABFE7D" author="Sierwald" box="[1259,1436,272,298]" pageId="2" pageNumber="68" refString="Sierwald, P. (1989). Morphology and ontogeny of female copulatory organs in American Pisauridae, with special reference to homologous features (Arachnida: Araneae). Smithsonian Contributions to Zoology, 484, 1 - 24." type="journal article" year="1989">Sierwald 1989</bibRefCitation>
because they are not associated to the fertilization ducts); fertilization ducts short, hook shaped, directed anteriorly and outwards (
<figureCitation id="13362A47FFFEFFE1AD8BC8426E23FE2D" box="[396,532,352,378]" captionStart="FIGURES 8 16. 8 12" captionStartId="4.[151,269,1715,1739]" captionTargetBox="[231,1365,476,1684]" captionTargetId="figure@4.[231,1366,474,1687]" captionTargetPageId="4" captionText="FIGURES 8 16. 8 12. Quemedice enigmaticus Mello Leitão (8 9 IBSP 67822; 10 MACN Ar 10758; 11 12 IBSP 1899). 8 10 Left male palp (8 ventral, 9 retrolateral, 10 ditto, detail of expanded embolus); 11 12. Epigynum (11 ventral, 12 dorsal). 13 16. Quemedice piracuruca n. sp. (13 14, paratype, IBSP 28565; 15 16, paratype, MPEG 6402). 13 14 Left male palp (arrows indicate small teeth) (13 ventral, 14 dorsal); 15 16. Epigynum (15 ventral, 16 dorsal). (C = cymbium; CD = copulatory duct; CO = copulatory opening; db = dorsal branch of RTA; EA = embolus apophysis; EB = embolus base; E = embolus; FD = fertilization duct; HR = hyaline receptacle; MS = median septum; LL = lateral lobes; ST = subtegulum; T = tegulum; vb = ventral branch of RTA). Scale bars: 8 9, 11 16, 0.5 mm; 10, 0.1 mm." httpUri="https://zenodo.org/record/274361/files/figure.png" pageId="2" pageNumber="62">Figs 12, 16</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3176549FFFEFFE1ACC2C8A56D2AFDB5" pageId="2" pageNumber="62" type="discussion">
<paragraph id="8BB236C2FFFEFFE1ACC2C8A56D2AFDB5" blockId="2.[151,1436,152,738]" pageId="2" pageNumber="62">
<emphasis id="B979EAD0FFFEFFE1ACC2C8A56D09FEF6" bold="true" box="[197,318,391,417]" pageId="2" pageNumber="62">Remarks.</emphasis>
Within
<taxonomicName id="4C0D4D41FFFEFFE1AD9BC8AA6E1EFEF5" box="[412,553,392,418]" class="Arachnida" family="Sparassidae" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="62" phylum="Arthropoda" rank="family">Sparassidae</taxonomicName>
,
<taxonomicName id="4C0D4D41FFFEFFE1AE32C8AA6E8EFEF6" box="[565,697,392,417]" class="Arachnida" family="Sparassidae" genus="Quemedice" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="62" phylum="Arthropoda" rank="genus">
<emphasis id="B979EAD0FFFEFFE1AE32C8AA6E8EFEF6" box="[565,697,392,417]" italics="true" pageId="2" pageNumber="62">Quemedice</emphasis>
</taxonomicName>
seems to belong to Sparassinae due to the presence of only two promarginal teeth in the chelicerae, absence of intermarginal denticles, median hook of trilobate membrane as large as or slightly larger than lateral projections and a short-toothed female palpal claw (
<bibRefCitation id="EF9C4B33FFFEFFE1A8C8C8FA6960FEA5" author="Jager" box="[1231,1367,472,498]" pageId="2" pageNumber="67" refString="Jager, P. (1998) First results of a taxonomic revision of the SE Asian Sparassidae (Araneae). In: P. A. Selden (Ed.), Proceedings of the 17 th European Colloquium of Arachnology, Edimburgh 1997, pp. 53 - 59." type="journal article" year="1998">Jäger 1998</bibRefCitation>
). The genus seems to be closely related to
<taxonomicName id="4C0D4D41FFFEFFE1AE45CB226F3EFD4D" authority="Simon" authorityName="Simon" box="[578,777,512,538]" class="Arachnida" family="Sparassidae" genus="Macrinus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="62" phylum="Arthropoda" rank="genus">
<emphasis id="B979EAD0FFFEFFE1AE45CB226E85FD4E" box="[578,690,512,537]" italics="true" pageId="2" pageNumber="62">Macrinus</emphasis>
Simon
</taxonomicName>
,
<taxonomicName id="4C0D4D41FFFEFFE1AF12CB226FE0FD4D" authority="Simon" authorityName="Simon" box="[789,983,512,538]" class="Arachnida" family="Sparassidae" genus="Vindullus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="62" phylum="Arthropoda" rank="genus">
<emphasis id="B979EAD0FFFEFFE1AF12CB226FB4FD4E" box="[789,899,512,537]" italics="true" pageId="2" pageNumber="62">Vindullus</emphasis>
Simon
</taxonomicName>
and
<taxonomicName id="4C0D4D41FFFEFFE1A817CB2268C3FD4D" authority="Kammerer" authorityName="Kammerer" box="[1040,1268,512,538]" class="Arachnida" family="Sparassidae" genus="Nolavia" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="62" phylum="Arthropoda" rank="genus">
<emphasis id="B979EAD0FFFEFFE1A817CB22685AFD4E" box="[1040,1133,512,537]" italics="true" pageId="2" pageNumber="62">Nolavia</emphasis>
Kammerer
</taxonomicName>
, with whom it shares the presence of only two pairs of ventral spines on tibiae I-IV, the tegulum slightly spiraled perpendicularly to the main palpal axis, towards the tip of the cymbium (
<figureCitation id="13362A47FFFEFFE1AF7ACB726FCAFD3D" box="[893,1021,592,618]" captionStart="FIGURES 8 16. 8 12" captionStartId="4.[151,269,1715,1739]" captionTargetBox="[231,1365,476,1684]" captionTargetId="figure@4.[231,1366,474,1687]" captionTargetPageId="4" captionText="FIGURES 8 16. 8 12. Quemedice enigmaticus Mello Leitão (8 9 IBSP 67822; 10 MACN Ar 10758; 11 12 IBSP 1899). 8 10 Left male palp (8 ventral, 9 retrolateral, 10 ditto, detail of expanded embolus); 11 12. Epigynum (11 ventral, 12 dorsal). 13 16. Quemedice piracuruca n. sp. (13 14, paratype, IBSP 28565; 15 16, paratype, MPEG 6402). 13 14 Left male palp (arrows indicate small teeth) (13 ventral, 14 dorsal); 15 16. Epigynum (15 ventral, 16 dorsal). (C = cymbium; CD = copulatory duct; CO = copulatory opening; db = dorsal branch of RTA; EA = embolus apophysis; EB = embolus base; E = embolus; FD = fertilization duct; HR = hyaline receptacle; MS = median septum; LL = lateral lobes; ST = subtegulum; T = tegulum; vb = ventral branch of RTA). Scale bars: 8 9, 11 16, 0.5 mm; 10, 0.1 mm." httpUri="https://zenodo.org/record/274361/files/figure.png" pageId="2" pageNumber="62">Figs. 8, 13</figureCitation>
;
<bibRefCitation id="EF9C4B33FFFEFFE1A80DCB72689DFD3D" author="Rheims" box="[1034,1194,592,618]" pageId="2" pageNumber="68" refString="Rheims, C. A. (2007) Revision of the Neotropical spider genus Macrinus (Araneae, Sparassidae). Journal of Arachnology, 35, 159 - 170." type="journal article" year="2007">Rheims 2007</bibRefCitation>
: figs. 22, 28, 34) the female
<taxonomicName id="4C0D4D41FFFEFFE1ACE8CB5A6D53FDC5" box="[239,356,632,658]" class="Magnoliopsida" family="Apocynaceae" genus="Epigynum" kingdom="Plantae" order="Gentianales" pageId="2" pageNumber="62" phylum="Tracheophyta" rank="genus">epigynum</taxonomicName>
with simple, rounded lateral lobes partially covering the median septum (
<figureCitation id="13362A47FFFEFFE1A8CACB5A6962FDC5" box="[1229,1365,632,658]" captionStart="FIGURES 8 16. 8 12" captionStartId="4.[151,269,1715,1739]" captionTargetBox="[231,1365,476,1684]" captionTargetId="figure@4.[231,1366,474,1687]" captionTargetPageId="4" captionText="FIGURES 8 16. 8 12. Quemedice enigmaticus Mello Leitão (8 9 IBSP 67822; 10 MACN Ar 10758; 11 12 IBSP 1899). 8 10 Left male palp (8 ventral, 9 retrolateral, 10 ditto, detail of expanded embolus); 11 12. Epigynum (11 ventral, 12 dorsal). 13 16. Quemedice piracuruca n. sp. (13 14, paratype, IBSP 28565; 15 16, paratype, MPEG 6402). 13 14 Left male palp (arrows indicate small teeth) (13 ventral, 14 dorsal); 15 16. Epigynum (15 ventral, 16 dorsal). (C = cymbium; CD = copulatory duct; CO = copulatory opening; db = dorsal branch of RTA; EA = embolus apophysis; EB = embolus base; E = embolus; FD = fertilization duct; HR = hyaline receptacle; MS = median septum; LL = lateral lobes; ST = subtegulum; T = tegulum; vb = ventral branch of RTA). Scale bars: 8 9, 11 16, 0.5 mm; 10, 0.1 mm." httpUri="https://zenodo.org/record/274361/files/figure.png" pageId="2" pageNumber="62">Figs 11, 15</figureCitation>
;
<bibRefCitation id="EF9C4B33FFFEFFE1A964CB5A6D3EFDED" author="Rheims" pageId="2" pageNumber="68" refString="Rheims, C. A. (2007) Revision of the Neotropical spider genus Macrinus (Araneae, Sparassidae). Journal of Arachnology, 35, 159 - 170." type="journal article" year="2007">Rheims 2007</bibRefCitation>
: figs 24, 30, 36) and vulva with large anterior hyaline receptacles (
<figureCitation id="13362A47FFFEFFE1A82ECB826884FDED" box="[1065,1203,672,698]" captionStart="FIGURES 8 16. 8 12" captionStartId="4.[151,269,1715,1739]" captionTargetBox="[231,1365,476,1684]" captionTargetId="figure@4.[231,1366,474,1687]" captionTargetPageId="4" captionText="FIGURES 8 16. 8 12. Quemedice enigmaticus Mello Leitão (8 9 IBSP 67822; 10 MACN Ar 10758; 11 12 IBSP 1899). 8 10 Left male palp (8 ventral, 9 retrolateral, 10 ditto, detail of expanded embolus); 11 12. Epigynum (11 ventral, 12 dorsal). 13 16. Quemedice piracuruca n. sp. (13 14, paratype, IBSP 28565; 15 16, paratype, MPEG 6402). 13 14 Left male palp (arrows indicate small teeth) (13 ventral, 14 dorsal); 15 16. Epigynum (15 ventral, 16 dorsal). (C = cymbium; CD = copulatory duct; CO = copulatory opening; db = dorsal branch of RTA; EA = embolus apophysis; EB = embolus base; E = embolus; FD = fertilization duct; HR = hyaline receptacle; MS = median septum; LL = lateral lobes; ST = subtegulum; T = tegulum; vb = ventral branch of RTA). Scale bars: 8 9, 11 16, 0.5 mm; 10, 0.1 mm." httpUri="https://zenodo.org/record/274361/files/figure.png" pageId="2" pageNumber="62">Figs 12, 16</figureCitation>
;
<bibRefCitation id="EF9C4B33FFFEFFE1A8C5CB826954FDED" author="Rheims" box="[1218,1379,672,698]" pageId="2" pageNumber="68" refString="Rheims, C. A. (2007) Revision of the Neotropical spider genus Macrinus (Araneae, Sparassidae). Journal of Arachnology, 35, 159 - 170." type="journal article" year="2007">Rheims 2007</bibRefCitation>
: figs 25, 31, 37).
</paragraph>
</subSubSection>
</treatment>
</document>