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<document id="3ADD8B63B66825CF9DCB6B72A98BE0D9" ID-CLB-Dataset="57608" ID-DOI="10.11646/zootaxa.5276.1.1" ID-GBIF-Dataset="590e0e3a-f3ea-4420-b976-b668987049e6" ID-ISSN="1175­5334" ID-Zenodo-Dep="7907294" ID-ZooBank="92300500-BB24-45B0-8ADD-977C3220A069" IM.bibliography_approvedBy="tatiana" IM.illustrations_approvedBy="tatiana" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="tatiana" IM.treatments_approvedBy="tatiana" checkinTime="1682929863379" checkinUser="plazi" docAuthor="Camargo, Alexssandro, Vieira, Rodrigo &amp; Rafael, José Albertino" docDate="2023" docId="03A4878EEE593857BEE1FA57F3E9FD7B" docLanguage="en" docName="zootaxa.5276.1.1.pdf" docOrigin="Zootaxa 5276" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Catostola complicata, comb. nov." docType="treatment" docVersion="5" lastPageNumber="32" masterDocId="FF9DFFF6EE423848BE76FFD5F253FFD2" masterDocTitle="Taxonomic review of Ctenodontina Enderlein, 1914 with the revalidation of Catostola Hull, 1958 (Diptera: Asilidae: Asilinae) and description of a new species" masterLastPageNumber="71" masterPageNumber="1" pageNumber="28" updateTime="1699515144633" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="3F891C746FE2672B683DB8B9E81C590F">Taxonomic review of Ctenodontina Enderlein, 1914 with the revalidation of Catostola Hull, 1958 (Diptera: Asilidae: Asilinae) and description of a new species</mods:title>
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<mods:namePart id="992ED828E1A56EA63A17FE65FF3D9E4E">Camargo, Alexssandro</mods:namePart>
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<mods:namePart id="3B34726587859DE7FAEB1E6675109051">Vieira, Rodrigo</mods:namePart>
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<mods:namePart id="FD5A93963233BD24CD0D28AFBFC315A1">Rafael, José Albertino</mods:namePart>
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<treatment id="03A4878EEE593857BEE1FA57F3E9FD7B" ID-DOI="http://doi.org/10.5281/zenodo.7907326" ID-GBIF-Taxon="209588243" ID-Zenodo-Dep="7907326" LSID="urn:lsid:plazi:treatment:03A4878EEE593857BEE1FA57F3E9FD7B" httpUri="http://treatment.plazi.org/id/03A4878EEE593857BEE1FA57F3E9FD7B" lastPageId="31" lastPageNumber="32" pageId="27" pageNumber="28">
<subSubSection id="C3176513EE593853BEE1FA57F086FA4F" box="[151,725,1410,1437]" pageId="27" pageNumber="28" type="nomenclature">
<paragraph id="8BB23698EE593853BEE1FA57F086FA4F" blockId="27.[151,725,1410,1473]" box="[151,725,1410,1437]" pageId="27" pageNumber="28">
<heading id="D0FA81F4EE593853BEE1FA57F086FA4F" bold="true" box="[151,725,1410,1437]" fontSize="11" level="1" pageId="27" pageNumber="28" reason="1">
<emphasis id="B979EA8AEE593853BEE1FA57F086FA4F" bold="true" box="[151,725,1410,1437]" pageId="27" pageNumber="28">
<taxonomicName id="4C0D4D1BEE593853BEE1FA57F014FA4E" ID-CoL="BDNLX" authority="(James, 1953)" baseAuthorityName="James" baseAuthorityYear="1953" box="[151,583,1410,1437]" class="Insecta" family="Asilidae" genus="Catostola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="complicata" status="comb. nov.">
<emphasis id="B979EA8AEE593853BEE1FA57F3C6FA4F" bold="true" box="[151,405,1410,1437]" italics="true" pageId="27" pageNumber="28">Catostola complicata</emphasis>
(
<bibRefCitation id="EF9C4B69EE593853BFD2FA57F06CFA4E" author="James, M. T." box="[420,575,1410,1436]" pageId="27" pageNumber="28" pagination="46 - 57" refId="ref35216" refString="James, M. T. (1953) The Diptera collected on the Cockerell and Hubbell Expedition to Honduras: Part II. Asilidae. Journal of the Washington Academy of Sciences, 43 (2), 46 - 57." type="journal article" year="1953">James, 1953</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A24A57F1EE593853BC38FA56F086FA4F" box="[590,725,1411,1437]" pageId="27" pageNumber="28" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3176513EE593853BEE1FA72F302FA13" box="[151,337,1447,1473]" pageId="27" pageNumber="28" type="description">
<paragraph id="8BB23698EE593853BEE1FA72F302FA13" blockId="27.[151,725,1410,1473]" box="[151,337,1447,1473]" pageId="27" pageNumber="28">
(
<figureCitation id="13362A1DEE593853BEE9FA72F372FA13" box="[159,289,1447,1473]" captionStart-0="FIGURE 16" captionStart-1="FIGURE 17" captionStartId-0="29.[151,250,1943,1967]" captionStartId-1="30.[151,250,1897,1921]" captionTargetBox-0="[203,1385,181,1919]" captionTargetBox-1="[218,1368,181,1873]" captionTargetId-0="figure-17@29.[203,1385,181,1919]" captionTargetId-1="figure-21@30.[218,1368,181,1873]" captionTargetPageId-0="29" captionTargetPageId-1="30" captionText-0="FIGURE 16. Catostola complicata (James, 1953) comb. nov., holotype male. A. Habitus, lateral (insert: labels, not to scale); B. Same, dorsal; C. Head, frontal; D. Same, lateral. Scale bar 1 mm." captionText-1="FIGURE 17. Catostola complicata (James, 1953) comb. nov., holotype male. A. Wing, dorsal; B. Scutellum, dorsal; C. Hind femur, anterior; D. Terminalia, lateral; E. Same, dorsolateral; F. Same, dorsal; G. Same, ventral. Abbreviations: cerc: cercus; epand: epandrium; goncx: gonocoxite; gonst: gonostylus; hypd: hypandrium; st: sternite; subepand scl: subepandrial sclerite tg: tergite. Scale bar 1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.7907356" figureDoi-1="http://doi.org/10.5281/zenodo.7907358" httpUri-0="https://zenodo.org/record/7907356/files/figure.png" httpUri-1="https://zenodo.org/record/7907358/files/figure.png" pageId="27" pageNumber="28">Figs 1617</figureCitation>
,
<figureCitation id="13362A1DEE593853BF5AFA72F31AFA13" box="[300,329,1447,1473]" captionStart="FIGURE 44" captionStartId="66.[151,250,1682,1706]" captionTargetBox="[151,1435,525,1658]" captionTargetId="figure-172@66.[151,1435,525,1658]" captionTargetPageId="66" captionText="FIGURE 44. Distribution map of Catostola species: Catostola baleta (Walker, 1849) comb. nov., Catostola complicata (James, 1953) comb. nov., Catostola indecisa sp. nov., and Catostola martini (Fisher, 1992) comb. nov." figureDoi="http://doi.org/10.5281/zenodo.7907427" httpUri="https://zenodo.org/record/7907427/files/figure.png" pageId="27" pageNumber="28">44</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3176513EE593853BEE1FA3EF0E3F94E" pageId="27" pageNumber="28" type="reference_group">
<paragraph id="8BB23698EE593853BEE1FA3EF36CF9F0" blockId="27.[151,1436,1515,1693]" pageId="27" pageNumber="28">
<treatmentCitationGroup id="AB1D11B6EE593853BEE1FA3EF36CF9F0" pageId="27" pageNumber="28">
<treatmentCitation id="0AAC1089EE593853BEE1FA3EF000F9D1" author="James, M. T." box="[151,595,1515,1539]" page="52" pageId="27" pageNumber="28" year="1953">
<taxonomicName id="4C0D4D1BEE593853BEE1FA3EF000F9D1" ID-CoL="4BQQ3" authority="James, 1953: 52" authorityName="James" authorityPageNumber="52" authorityYear="1953" box="[151,595,1515,1539]" class="Insecta" family="Asilidae" genus="Pachychoeta" kingdom="Animalia" order="Diptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="complicata">
<emphasis id="B979EA8AEE593853BEE1FA3EF3C4F9D1" box="[151,407,1515,1539]" italics="true" pageId="27" pageNumber="28">Pachychoeta complicata</emphasis>
<bibRefCitation id="EF9C4B69EE593853BFD4FA39F000F9D1" author="James, M. T." box="[418,595,1515,1539]" pageId="27" pageNumber="28" pagination="46 - 57" refId="ref35216" refString="James, M. T. (1953) The Diptera collected on the Cockerell and Hubbell Expedition to Honduras: Part II. Asilidae. Journal of the Washington Academy of Sciences, 43 (2), 46 - 57." type="journal article" year="1953">James, 1953: 52</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, fig. 6;
<treatmentCitation id="0AAC1089EE593853BCD9FA3EF1BDF9D1" author="Martin, C. H. &amp; Papavero, N." box="[687,1006,1515,1539]" page="71" pageId="27" pageNumber="28" year="1970">
<bibRefCitation id="EF9C4B69EE593853BCD9FA3EF1BDF9D1" author="Martin, C. H. &amp; Papavero, N." box="[687,1006,1515,1539]" pageId="27" pageNumber="28" pagination="1 - 139" refId="ref35563" refString="Martin, C. H. &amp; Papavero, N. (1970) Family Asilidae. In: Papavero, N. (Ed.), A Catalogue of the Diptera of the Americas South of the United States 35 b. Museu de Zoologia, Universidade de S &quot; o Paulo, S &quot; o Paulo, pp 1 - 139." type="book chapter" year="1970">Martin &amp; Papavero, 1970: 71</bibRefCitation>
</treatmentCitation>
(catalogue);
<treatmentCitation id="0AAC1089EE593853BA08FA3EF766F9D1" author="Martin, C. H." box="[1150,1333,1515,1539]" page="84" pageId="27" pageNumber="28" year="1975">
<bibRefCitation id="EF9C4B69EE593853BA08FA3EF766F9D1" author="Martin, C. H." box="[1150,1333,1515,1539]" pageId="27" pageNumber="28" pagination="1 - 107" refId="ref35510" refString="Martin, C. H. (1975) The generic and specific characters of four old and six new Asilini genera in the western United States, Mexico and Central America (Diptera: Asilidae). Occasional papers of the California Academy of Sciences, 119, 1 - 107." type="journal article" year="1975">Martin, 1975: 84</bibRefCitation>
</treatmentCitation>
, 86 (key, comments).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BB23698EE593853BEE1F9FCF313F98D" blockId="27.[151,1436,1515,1693]" pageId="27" pageNumber="28">
<treatmentCitationGroup id="AB1D11B6EE593853BEE1F9FCF313F98D" pageId="27" pageNumber="28">
<taxonomicName id="4C0D4D1BEE593853BEE1F9FCF335F992" authorityName="Strecker" authorityYear="1898" box="[151,358,1576,1600]" class="Insecta" family="Asilidae" genus="Lecania" kingdom="Animalia" order="Diptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="complicata">
<emphasis id="B979EA8AEE593853BEE1F9FCF335F992" box="[151,358,1576,1600]" italics="true" pageId="27" pageNumber="28">Lecania complicata</emphasis>
</taxonomicName>
;
<bibRefCitation id="EF9C4B69EE593853BF0FF9FDF3BCF992" author="Hull, F. M." box="[377,495,1576,1600]" pageId="27" pageNumber="28" pagination="1 - 432" refId="ref35163" refString="Hull, F. M. (1962) Robber flies of the world. The genera of the family Asilidae. Smithsonian Institution Bulletin, 294 (1 &amp; 2), 1 - 432 &amp; 433 - 907. https: // doi. org / 10.5479 / si. 03629236.224" type="journal article" year="1962">Hull, 1962</bibRefCitation>
(2): 481;
<treatmentCitation id="0AAC1089EE593853BC25F9FDF1CAF993" author="Artigas, J. N. &amp; Papavero, N." box="[595,921,1576,1601]" page="37" pageId="27" pageNumber="28" year="1995">
<bibRefCitation id="EF9C4B69EE593853BC25F9FDF1CAF993" author="Artigas, J. N. &amp; Papavero, N." box="[595,921,1576,1601]" pageId="27" pageNumber="28" pagination="33 - 56" refId="ref34602" refString="Artigas, J. N. &amp; Papavero, N. (1995) The American genera of Asilidae (Diptera): Keys for identification with an atlas of female spermathecae and other morphological details. IX. 10. Subfamily Asilinae Leach - Lecania - Group, with a catalogue of Neotropical species. Theoria, 4, 33 - 56." type="journal article" year="1995">Artigas &amp; Papavero, 1995: 37</bibRefCitation>
</treatmentCitation>
(
<taxonomicName id="4C0D4D1BEE593853BDD8F9FCF653F992" ID-CoL="9CJMC" authorityName="Macquart" authorityYear="1838" box="[942,1024,1577,1600]" class="Insecta" family="Asilidae" genus="Lecania" kingdom="Animalia" order="Diptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="genus">
<emphasis id="B979EA8AEE593853BDD8F9FCF653F992" box="[942,1024,1577,1600]" italics="true" pageId="27" pageNumber="28">Lecania</emphasis>
</taxonomicName>
-group catalogue);
<treatmentCitation id="0AAC1089EE593853BABAF9FCF7CFF993" author="Papavero, N." box="[1228,1436,1577,1601]" page="31" pageId="27" pageNumber="28" year="2009">
<bibRefCitation id="EF9C4B69EE593853BABAF9FCF7CFF993" author="Papavero, N." box="[1228,1436,1577,1601]" pageId="27" pageNumber="28" pagination="1 - 178" refId="ref35724" refString="Papavero, N. (2009) Catalogue of Neotropical Diptera. Asilidae. Neotropical Diptera, 17, 1 - 178." type="journal article" year="2009">Papavero, 2009: 31</bibRefCitation>
</treatmentCitation>
(catalogue).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BB23698EE593853BEE1F9B3F0E3F94E" blockId="27.[151,1436,1515,1693]" pageId="27" pageNumber="28">
<treatmentCitationGroup id="AB1D11B6EE593853BEE1F9B3F0E3F94E" pageId="27" pageNumber="28">
<taxonomicName id="4C0D4D1BEE593853BEE1F9B3F3C8F9AC" baseAuthorityName="James" baseAuthorityYear="1953" box="[151,411,1638,1662]" class="Insecta" family="Asilidae" genus="Ctenodontina" kingdom="Animalia" order="Diptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="complicata">
<emphasis id="B979EA8AEE593853BEE1F9B3F3C8F9AC" box="[151,411,1638,1662]" italics="true" pageId="27" pageNumber="28">Ctenodontina complicata</emphasis>
</taxonomicName>
;
<treatmentCitation id="0AAC1089EE593853BFDCF9B3F032F9AC" author="Fisher, E. M." box="[426,609,1638,1662]" page="619" pageId="27" pageNumber="28" year="2009">
<bibRefCitation id="EF9C4B69EE593853BFDCF9B3F032F9AC" author="Fisher, E. M." box="[426,609,1638,1662]" pageId="27" pageNumber="28" refId="ref34954" refString="Fisher, E. M. (2009) Asilidae (robber flies, assassin flies, moscas cazadoras, moscas ladronas). In: Brown, B. V., Borkent, A., Cumming, J. M., Wood, D. M., Woodley, N. E. &amp; Zumbado, M. A. (Eds.), Manual of Central American Diptera. Vol. 1. National Research Council Research Press, Ottawa, Ontario, pp. 585 - 632." type="book" year="2009">Fisher, 2009: 619</bibRefCitation>
</treatmentCitation>
(Manual of Central American
<taxonomicName id="4C0D4D1BEE593853BDD3F9B3F1A2F9AC" ID-CoL="D2P" box="[933,1009,1638,1662]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="order">Diptera</taxonomicName>
[Fisher confused
<collectingCountry id="F31A7608EE593853BAD9F9B3F750F9AC" box="[1199,1283,1638,1662]" name="Mauritius" pageId="27" pageNumber="28">Maurice</collectingCountry>
T. James with James S. Hine in the authorship of the species]).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3176513EE593853BEE1F91FF1FEF86F" pageId="27" pageNumber="28" type="diagnosis">
<paragraph id="8BB23698EE593853BEE1F91FF1FEF86F" blockId="27.[151,1437,1738,2017]" pageId="27" pageNumber="28">
<emphasis id="B979EA8AEE593853BEE1F91FF341F936" bold="true" box="[151,274,1738,1764]" pageId="27" pageNumber="28">Diagnosis.</emphasis>
Face white pruinose, frons and vertex brown pruinose; mystacal macrosetae white with a row of dorsolateral black macrosetae (
<figureCitation id="13362A1DEE593853BF82F93AF02DF8DB" box="[500,638,1775,1801]" captionStart="FIGURE 16" captionStartId="29.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1919]" captionTargetId="figure-17@29.[203,1385,181,1919]" captionTargetPageId="29" captionText="FIGURE 16. Catostola complicata (James, 1953) comb. nov., holotype male. A. Habitus, lateral (insert: labels, not to scale); B. Same, dorsal; C. Head, frontal; D. Same, lateral. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7907356" httpUri="https://zenodo.org/record/7907356/files/figure.png" pageId="27" pageNumber="28">Fig. 16CD</figureCitation>
); thorax brownish grey pruinose with dark brown pruinose paramedian stripes and spots (
<figureCitation id="13362A1DEE593853BF17F8C6F394F8FF" box="[353,455,1811,1837]" captionStart="FIGURE 16" captionStartId="29.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1919]" captionTargetId="figure-17@29.[203,1385,181,1919]" captionTargetPageId="29" captionText="FIGURE 16. Catostola complicata (James, 1953) comb. nov., holotype male. A. Habitus, lateral (insert: labels, not to scale); B. Same, dorsal; C. Head, frontal; D. Same, lateral. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7907356" httpUri="https://zenodo.org/record/7907356/files/figure.png" pageId="27" pageNumber="28">Fig. 16B</figureCitation>
); legs yellow with apical third of mid and hind femora dark brown (
<figureCitation id="13362A1DEE593853BAB2F8C6F703F8FF" box="[1220,1360,1811,1837]" captionStart="FIGURE 16" captionStartId="29.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1919]" captionTargetId="figure-17@29.[203,1385,181,1919]" captionTargetPageId="29" captionText="FIGURE 16. Catostola complicata (James, 1953) comb. nov., holotype male. A. Habitus, lateral (insert: labels, not to scale); B. Same, dorsal; C. Head, frontal; D. Same, lateral. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7907356" httpUri="https://zenodo.org/record/7907356/files/figure.png" pageId="27" pageNumber="28">Figs 16AB</figureCitation>
,
<figureCitation id="13362A1DEE593853BB2AF8C6F7DCF8FF" box="[1372,1423,1811,1837]" captionStart="FIGURE 17" captionStartId="30.[151,250,1897,1921]" captionTargetBox="[218,1368,181,1873]" captionTargetId="figure-21@30.[218,1368,181,1873]" captionTargetPageId="30" captionText="FIGURE 17. Catostola complicata (James, 1953) comb. nov., holotype male. A. Wing, dorsal; B. Scutellum, dorsal; C. Hind femur, anterior; D. Terminalia, lateral; E. Same, dorsolateral; F. Same, dorsal; G. Same, ventral. Abbreviations: cerc: cercus; epand: epandrium; goncx: gonocoxite; gonst: gonostylus; hypd: hypandrium; st: sternite; subepand scl: subepandrial sclerite tg: tergite. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7907358" httpUri="https://zenodo.org/record/7907358/files/figure.png" pageId="27" pageNumber="28">17C</figureCitation>
); abdomen dark brown with brownish grey lateral margins (
<figureCitation id="13362A1DEE593853BD68F8E2F1F4F883" box="[798,935,1847,1873]" captionStart="FIGURE 16" captionStartId="29.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1919]" captionTargetId="figure-17@29.[203,1385,181,1919]" captionTargetPageId="29" captionText="FIGURE 16. Catostola complicata (James, 1953) comb. nov., holotype male. A. Habitus, lateral (insert: labels, not to scale); B. Same, dorsal; C. Head, frontal; D. Same, lateral. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7907356" httpUri="https://zenodo.org/record/7907356/files/figure.png" pageId="27" pageNumber="28">Fig. 16AB</figureCitation>
); terminalia dark reddish to black (
<figureCitation id="13362A1DEE593853BB5CF8E2F2FCF8A7" captionStart="FIGURE 17" captionStartId="30.[151,250,1897,1921]" captionTargetBox="[218,1368,181,1873]" captionTargetId="figure-21@30.[218,1368,181,1873]" captionTargetPageId="30" captionText="FIGURE 17. Catostola complicata (James, 1953) comb. nov., holotype male. A. Wing, dorsal; B. Scutellum, dorsal; C. Hind femur, anterior; D. Terminalia, lateral; E. Same, dorsolateral; F. Same, dorsal; G. Same, ventral. Abbreviations: cerc: cercus; epand: epandrium; goncx: gonocoxite; gonst: gonostylus; hypd: hypandrium; st: sternite; subepand scl: subepandrial sclerite tg: tergite. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7907358" httpUri="https://zenodo.org/record/7907358/files/figure.png" pageId="27" pageNumber="28">Fig. 17D G</figureCitation>
); S8 with a few, sparse white setae and with a mid-posterior spatulate projection slightly constricted medially with long dark brown to black macrosetae and setae laterally and apically, these macrosetae with white tips (
<figureCitation id="13362A1DEE593853BB19F8AAF2BDF86F" captionStart="FIGURE 17" captionStartId="30.[151,250,1897,1921]" captionTargetBox="[218,1368,181,1873]" captionTargetId="figure-21@30.[218,1368,181,1873]" captionTargetPageId="30" captionText="FIGURE 17. Catostola complicata (James, 1953) comb. nov., holotype male. A. Wing, dorsal; B. Scutellum, dorsal; C. Hind femur, anterior; D. Terminalia, lateral; E. Same, dorsolateral; F. Same, dorsal; G. Same, ventral. Abbreviations: cerc: cercus; epand: epandrium; goncx: gonocoxite; gonst: gonostylus; hypd: hypandrium; st: sternite; subepand scl: subepandrial sclerite tg: tergite. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7907358" httpUri="https://zenodo.org/record/7907358/files/figure.png" pageId="27" pageNumber="28">Fig. 17DG</figureCitation>
); epandrium with a finger-like dorsal projection (
<figureCitation id="13362A1DEE593853BD60F876F1F2F86F" box="[790,929,1955,1981]" captionStart="FIGURE 17" captionStartId="30.[151,250,1897,1921]" captionTargetBox="[218,1368,181,1873]" captionTargetId="figure-21@30.[218,1368,181,1873]" captionTargetPageId="30" captionText="FIGURE 17. Catostola complicata (James, 1953) comb. nov., holotype male. A. Wing, dorsal; B. Scutellum, dorsal; C. Hind femur, anterior; D. Terminalia, lateral; E. Same, dorsolateral; F. Same, dorsal; G. Same, ventral. Abbreviations: cerc: cercus; epand: epandrium; goncx: gonocoxite; gonst: gonostylus; hypd: hypandrium; st: sternite; subepand scl: subepandrial sclerite tg: tergite. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7907358" httpUri="https://zenodo.org/record/7907358/files/figure.png" pageId="27" pageNumber="28">Fig. 17DG</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3176513EE593854BEB1F813F0E6F80F" lastPageId="28" lastPageNumber="29" pageId="27" pageNumber="28" type="description">
<paragraph id="8BB23698EE593854BEB1F813F15CFE27" blockId="27.[151,1437,1738,2017]" lastBlockId="28.[151,1437,151,2013]" lastPageId="28" lastPageNumber="29" pageId="27" pageNumber="28">
<emphasis id="B979EA8AEE593853BEB1F813F09DF832" bold="true" box="[199,718,1990,2017]" pageId="27" pageNumber="28">
Redescription.
<typeStatus id="54B6883AEE593853BF0BF812F3B9F833" box="[381,490,1991,2017]" pageId="27" pageNumber="28" type="holotype">Holotype</typeStatus>
male (
<figureCitation id="13362A1DEE593853BC4AF813F092F832" box="[572,705,1990,2016]" captionStart-0="FIGURE 16" captionStart-1="FIGURE 17" captionStartId-0="29.[151,250,1943,1967]" captionStartId-1="30.[151,250,1897,1921]" captionTargetBox-0="[203,1385,181,1919]" captionTargetBox-1="[218,1368,181,1873]" captionTargetId-0="figure-17@29.[203,1385,181,1919]" captionTargetId-1="figure-21@30.[218,1368,181,1873]" captionTargetPageId-0="29" captionTargetPageId-1="30" captionText-0="FIGURE 16. Catostola complicata (James, 1953) comb. nov., holotype male. A. Habitus, lateral (insert: labels, not to scale); B. Same, dorsal; C. Head, frontal; D. Same, lateral. Scale bar 1 mm." captionText-1="FIGURE 17. Catostola complicata (James, 1953) comb. nov., holotype male. A. Wing, dorsal; B. Scutellum, dorsal; C. Hind femur, anterior; D. Terminalia, lateral; E. Same, dorsolateral; F. Same, dorsal; G. Same, ventral. Abbreviations: cerc: cercus; epand: epandrium; goncx: gonocoxite; gonst: gonostylus; hypd: hypandrium; st: sternite; subepand scl: subepandrial sclerite tg: tergite. Scale bar 1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.7907356" figureDoi-1="http://doi.org/10.5281/zenodo.7907358" httpUri-0="https://zenodo.org/record/7907356/files/figure.png" httpUri-1="https://zenodo.org/record/7907358/files/figure.png" pageId="27" pageNumber="28">Figs 1617</figureCitation>
).
</emphasis>
Head (
<figureCitation id="13362A1DEE593853BD55F812F1FCF833" box="[803,943,1991,2017]" captionStart="FIGURE 16" captionStartId="29.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1919]" captionTargetId="figure-17@29.[203,1385,181,1919]" captionTargetPageId="29" captionText="FIGURE 16. Catostola complicata (James, 1953) comb. nov., holotype male. A. Habitus, lateral (insert: labels, not to scale); B. Same, dorsal; C. Head, frontal; D. Same, lateral. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7907356" httpUri="https://zenodo.org/record/7907356/files/figure.png" pageId="27" pageNumber="28">Fig. 16CD</figureCitation>
): eyes black; face white pruinose; mystacal macrosetae white, except by a dorsolateral row of dark brown to black macrosetae (
<figureCitation id="13362A1DEE5E3854BA07FF42F6ACFF63" box="[1137,1279,151,177]" captionStart="FIGURE 16" captionStartId="29.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1919]" captionTargetId="figure-17@29.[203,1385,181,1919]" captionTargetPageId="29" captionText="FIGURE 16. Catostola complicata (James, 1953) comb. nov., holotype male. A. Habitus, lateral (insert: labels, not to scale); B. Same, dorsal; C. Head, frontal; D. Same, lateral. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7907356" httpUri="https://zenodo.org/record/7907356/files/figure.png" pageId="28" pageNumber="29">Fig. 16CD</figureCitation>
); subvibrissal macrosetae dark brown to black with a few white setae mixed; gena white pruinose; palpus dark brown with dark brown macrosetae and setae, except ventrally with white setae; proboscis black, labial setae yellowish, proboscial setae white; antennal socket black, white pruinose; frons brown pruinose with a triangular reddish apruinose central spot; orbital setae mixed dark brown, yellow and white; ocellar tubercle black, brown pruinose, with six pairs of dark brown proclinate setae; vertex brown pruinose; occiput greyish pruinose becoming white pruinose laterally and on lower occipital margin; 56 black postocular macrosetae (distal one pale), other dorsal occipital setae black and yellowish; lateral occipital and lower occipital setae white. Antenna: scape black with a reddish ring apically, white pruinose and black setose; pedicel reddish brown with yellowish pruinosity, white lateroventrally, black setose with a few very short yellowish setae ventrally (
<figureCitation id="13362A1DEE5E3854BC0FFE0EF150FE27" box="[633,771,475,501]" captionStart="FIGURE 16" captionStartId="29.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1919]" captionTargetId="figure-17@29.[203,1385,181,1919]" captionTargetPageId="29" captionText="FIGURE 16. Catostola complicata (James, 1953) comb. nov., holotype male. A. Habitus, lateral (insert: labels, not to scale); B. Same, dorsal; C. Head, frontal; D. Same, lateral. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7907356" httpUri="https://zenodo.org/record/7907356/files/figure.png" pageId="28" pageNumber="29">Fig. 16CD</figureCitation>
).
</paragraph>
<paragraph id="8BB23698EE5E3854BEB1FE2AF779FC53" blockId="28.[151,1437,151,2013]" pageId="28" pageNumber="29">
Thorax (
<figureCitation id="13362A1DEE5E3854BF5BFE2AF3EDFDCB" box="[301,446,511,537]" captionStart="FIGURE 16" captionStartId="29.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1919]" captionTargetId="figure-17@29.[203,1385,181,1919]" captionTargetPageId="29" captionText="FIGURE 16. Catostola complicata (James, 1953) comb. nov., holotype male. A. Habitus, lateral (insert: labels, not to scale); B. Same, dorsal; C. Head, frontal; D. Same, lateral. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7907356" httpUri="https://zenodo.org/record/7907356/files/figure.png" pageId="28" pageNumber="29">Fig. 16AB</figureCitation>
): brown pruinose; antepronotum and postpronotum brownish grey laterally; scutum with a pair of dark brown pruinose paramedian stripes rounded anteriorly, laterally and fading after postsutural spot; pre and postsutural spots and prescutellar spots dark brown pruinose; scutum laterally and posteriorly and transverse suture brownish grey pruinose; a triangular apruinose black spot at posterior scutal margin; postalar wall and scutellum brownish grey pruinose (
<figureCitation id="13362A1DEE5E3854BC17FD5AF099FD7B" box="[609,714,655,681]" captionStart="FIGURE 17" captionStartId="30.[151,250,1897,1921]" captionTargetBox="[218,1368,181,1873]" captionTargetId="figure-21@30.[218,1368,181,1873]" captionTargetPageId="30" captionText="FIGURE 17. Catostola complicata (James, 1953) comb. nov., holotype male. A. Wing, dorsal; B. Scutellum, dorsal; C. Hind femur, anterior; D. Terminalia, lateral; E. Same, dorsolateral; F. Same, dorsal; G. Same, ventral. Abbreviations: cerc: cercus; epand: epandrium; goncx: gonocoxite; gonst: gonostylus; hypd: hypandrium; st: sternite; subepand scl: subepandrial sclerite tg: tergite. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7907358" httpUri="https://zenodo.org/record/7907358/files/figure.png" pageId="28" pageNumber="29">Fig. 17B</figureCitation>
); pleura greyish pruinose, except proepisternum, proepimeron, anepisternum, dorsal part of katepisternum, katepimeron and posterior part of metanepisternum brownish grey pruinose. Chaetotaxy: antepronotal macrosetae and setae pale yellow with a few, very short black setae posteriorly; 2 notopleural; 1 supra alar; 1 postalar and 3 pairs of dorsocentral black postsutural macrosetae; scutum with short black acrostichal setae and thin yellowish setae on postpronotal lobe, anteriorly; scutellum with short sparse black discal and marginal scutellar setae (
<figureCitation id="13362A1DEE5E3854BC44FC96F0C9FC8F" box="[562,666,835,861]" captionStart="FIGURE 17" captionStartId="30.[151,250,1897,1921]" captionTargetBox="[218,1368,181,1873]" captionTargetId="figure-21@30.[218,1368,181,1873]" captionTargetPageId="30" captionText="FIGURE 17. Catostola complicata (James, 1953) comb. nov., holotype male. A. Wing, dorsal; B. Scutellum, dorsal; C. Hind femur, anterior; D. Terminalia, lateral; E. Same, dorsolateral; F. Same, dorsal; G. Same, ventral. Abbreviations: cerc: cercus; epand: epandrium; goncx: gonocoxite; gonst: gonostylus; hypd: hypandrium; st: sternite; subepand scl: subepandrial sclerite tg: tergite. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7907358" httpUri="https://zenodo.org/record/7907358/files/figure.png" pageId="28" pageNumber="29">Fig. 17B</figureCitation>
); anepisternum with short, thin yellowish and black setae dorsally; katatergite with white macrosetae and setae; remainder of pleura with thin and very sparse whitish setae.
</paragraph>
<paragraph id="8BB23698EE5E3854BEB1FC5EF69FFBC3" blockId="28.[151,1437,151,2013]" pageId="28" pageNumber="29">
Wings (
<figureCitation id="13362A1DEE5E3854BF6BFC5EF3D5FC77" box="[285,390,907,933]" captionStart="FIGURE 17" captionStartId="30.[151,250,1897,1921]" captionTargetBox="[218,1368,181,1873]" captionTargetId="figure-21@30.[218,1368,181,1873]" captionTargetPageId="30" captionText="FIGURE 17. Catostola complicata (James, 1953) comb. nov., holotype male. A. Wing, dorsal; B. Scutellum, dorsal; C. Hind femur, anterior; D. Terminalia, lateral; E. Same, dorsolateral; F. Same, dorsal; G. Same, ventral. Abbreviations: cerc: cercus; epand: epandrium; goncx: gonocoxite; gonst: gonostylus; hypd: hypandrium; st: sternite; subepand scl: subepandrial sclerite tg: tergite. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7907358" httpUri="https://zenodo.org/record/7907358/files/figure.png" pageId="28" pageNumber="29">Fig. 17A</figureCitation>
): translucent, apical third slightly yellowish translucent; veins dark brown;
<emphasis id="B979EA8AEE5E3854BAB1FC5EF6B3FC79" box="[1223,1248,907,939]" italics="true" pageId="28" pageNumber="29">
R
<subScript id="178934DDEE5E3854BAAEFC48F6B3FC79" attach="left" box="[1240,1248,925,939]" fontSize="6" pageId="28" pageNumber="29">4</subScript>
</emphasis>
slightly rounded in the junction with
<emphasis id="B979EA8AEE5E3854BFF4FC7AF3C8FC1D" box="[386,411,943,975]" italics="true" pageId="28" pageNumber="29">
R
<subScript id="178934DDEE5E3854BFE5FC14F3C8FC1D" attach="left" box="[403,411,961,975]" fontSize="6" pageId="28" pageNumber="29">5</subScript>
</emphasis>
; bifurcation of
<emphasis id="B979EA8AEE5E3854BC24FC7AF038FC1D" box="[594,619,943,975]" italics="true" pageId="28" pageNumber="29">
R
<subScript id="178934DDEE5E3854BC15FC14F038FC1D" attach="left" box="[611,619,961,975]" fontSize="6" pageId="28" pageNumber="29">4</subScript>
</emphasis>
and
<emphasis id="B979EA8AEE5E3854BCD0FC7AF0ECFC1D" box="[678,703,943,975]" italics="true" pageId="28" pageNumber="29">
R
<subScript id="178934DDEE5E3854BCC1FC14F0ECFC1D" attach="left" box="[695,703,961,975]" fontSize="6" pageId="28" pageNumber="29">5</subScript>
</emphasis>
beyond discal cell at a distance equivalent to the length of two crossvein
<emphasis id="B979EA8AEE5E3854BF7DFC06F360FC3E" box="[267,307,979,1004]" italics="true" pageId="28" pageNumber="29">r-m</emphasis>
; crossvein
<emphasis id="B979EA8AEE5E3854BFCEFC06F3B3FC3E" box="[440,480,979,1004]" italics="true" pageId="28" pageNumber="29">r-m</emphasis>
shortly after the middle of discal cell at a distance equivalent to the length of one crossvein
<emphasis id="B979EA8AEE5E3854BF7EFC22F369FBC2" box="[264,314,1015,1040]" italics="true" pageId="28" pageNumber="29">m-m</emphasis>
; cells
<emphasis id="B979EA8AEE5E3854BFF5FC22F3CCFBC5" box="[387,415,1015,1047]" italics="true" pageId="28" pageNumber="29">
m
<subScript id="178934DDEE5E3854BFE1FBDCF3CCFBC5" attach="left" box="[407,415,1033,1047]" fontSize="6" pageId="28" pageNumber="29">3</subScript>
</emphasis>
and
<emphasis id="B979EA8AEE5E3854BFA0FC22F3ADFBC2" box="[470,510,1015,1040]" italics="true" pageId="28" pageNumber="29">cua</emphasis>
closed before wing margin. Halteres: yellow, knob dark yellow.
</paragraph>
<paragraph id="8BB23698EE5E3854BEB1FBCEF637FAAB" blockId="28.[151,1437,151,2013]" pageId="28" pageNumber="29">
Legs (
<figureCitation id="13362A1DEE5E3854BF7BFBCEF3CAFBE7" box="[269,409,1051,1077]" captionStart="FIGURE 16" captionStartId="29.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1919]" captionTargetId="figure-17@29.[203,1385,181,1919]" captionTargetPageId="29" captionText="FIGURE 16. Catostola complicata (James, 1953) comb. nov., holotype male. A. Habitus, lateral (insert: labels, not to scale); B. Same, dorsal; C. Head, frontal; D. Same, lateral. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7907356" httpUri="https://zenodo.org/record/7907356/files/figure.png" pageId="28" pageNumber="29">Figs 16AB</figureCitation>
,
<figureCitation id="13362A1DEE5E3854BFD3FBCEF38BFBE7" box="[421,472,1051,1077]" captionStart="FIGURE 17" captionStartId="30.[151,250,1897,1921]" captionTargetBox="[218,1368,181,1873]" captionTargetId="figure-21@30.[218,1368,181,1873]" captionTargetPageId="30" captionText="FIGURE 17. Catostola complicata (James, 1953) comb. nov., holotype male. A. Wing, dorsal; B. Scutellum, dorsal; C. Hind femur, anterior; D. Terminalia, lateral; E. Same, dorsolateral; F. Same, dorsal; G. Same, ventral. Abbreviations: cerc: cercus; epand: epandrium; goncx: gonocoxite; gonst: gonostylus; hypd: hypandrium; st: sternite; subepand scl: subepandrial sclerite tg: tergite. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7907358" httpUri="https://zenodo.org/record/7907358/files/figure.png" pageId="28" pageNumber="29">17C</figureCitation>
): yellow, except coxae and trochanters black and brownish grey pruinose, apical third of mid and hind femora dark brown (hind femora with a brown anterodorsal stripe fading proximally), apex of fore and mid tibiae, anteroventral stripe and apical half of hind tibia and all tarsi dark brown (darker on hind leg). Chaetotaxy: coxae and trochanters with white macrosetae and setae; hind trochanter with 3 posterior white macrosetae; fore femur with 56 ventral yellow macrosetae; mid femur with 45 ventral (whitish), 7 anteroventral (proximal whitish), 3 anterior, 1 posterodorsal preapical and 3 posterior (2 proximal yellow) black macrosetae; hind femur with 1 anteroventral preapical, 1 anterior, 1 anterodorsal preapical, and 1 posterodorsal preapical black macrosetae; other short femoral setae black anterodorsally and yellowish posteroventrally; tibiae and tarsi with black macrosetae and short black setae (tibiae with a few short yellowish setae anterodorsally); brush setae golden reddish yellow; pulvilli and empodium dark reddish; claws black, dark reddish basally.
</paragraph>
<paragraph id="8BB23698EE5E3854BEB1FA56F0B2F9FF" blockId="28.[151,1437,151,2013]" pageId="28" pageNumber="29">
Abdomen (
<figureCitation id="13362A1DEE5E3854BF3FFA56F38BFA4F" box="[329,472,1411,1437]" captionStart="FIGURE 16" captionStartId="29.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1919]" captionTargetId="figure-17@29.[203,1385,181,1919]" captionTargetPageId="29" captionText="FIGURE 16. Catostola complicata (James, 1953) comb. nov., holotype male. A. Habitus, lateral (insert: labels, not to scale); B. Same, dorsal; C. Head, frontal; D. Same, lateral. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7907356" httpUri="https://zenodo.org/record/7907356/files/figure.png" pageId="28" pageNumber="29">Fig. 16AB</figureCitation>
): dark brown pruinose, tergites laterally brownish grey pruinose, T6 and 7 silvery pruinose; T1 with many lateral marginal whitish macrosetae and setae; T2 with 4, T3 with 8, T4 with 7, T57 only with 34 thin posterior marginal whitish setae; short sparse setae on tergites whitish on T1 and 2 and dark brown on other tergites becoming whitish at lateral margins; sternites brownish grey pruinose and with sparse, thin whitish setae; anterior margin of S7 with an apruinose band.
</paragraph>
<paragraph id="8BB23698EE5E3854BEB1F9E2F1A1F86B" blockId="28.[151,1437,151,2013]" pageId="28" pageNumber="29">
Terminalia (
<figureCitation id="13362A1DEE5E3854BF27F9E2F38CF983" box="[337,479,1591,1617]" captionStart="FIGURE 17" captionStartId="30.[151,250,1897,1921]" captionTargetBox="[218,1368,181,1873]" captionTargetId="figure-21@30.[218,1368,181,1873]" captionTargetPageId="30" captionText="FIGURE 17. Catostola complicata (James, 1953) comb. nov., holotype male. A. Wing, dorsal; B. Scutellum, dorsal; C. Hind femur, anterior; D. Terminalia, lateral; E. Same, dorsolateral; F. Same, dorsal; G. Same, ventral. Abbreviations: cerc: cercus; epand: epandrium; goncx: gonocoxite; gonst: gonostylus; hypd: hypandrium; st: sternite; subepand scl: subepandrial sclerite tg: tergite. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7907358" httpUri="https://zenodo.org/record/7907358/files/figure.png" pageId="28" pageNumber="29">Fig. 17DG</figureCitation>
): shining dark reddish to black; T8 yellowish pruinose at posterior margin and with brown to black macrosetae and setae (
<figureCitation id="13362A1DEE5E3854BC25F98EF08FF9A7" box="[595,732,1627,1653]" captionStart="FIGURE 17" captionStartId="30.[151,250,1897,1921]" captionTargetBox="[218,1368,181,1873]" captionTargetId="figure-21@30.[218,1368,181,1873]" captionTargetPageId="30" captionText="FIGURE 17. Catostola complicata (James, 1953) comb. nov., holotype male. A. Wing, dorsal; B. Scutellum, dorsal; C. Hind femur, anterior; D. Terminalia, lateral; E. Same, dorsolateral; F. Same, dorsal; G. Same, ventral. Abbreviations: cerc: cercus; epand: epandrium; goncx: gonocoxite; gonst: gonostylus; hypd: hypandrium; st: sternite; subepand scl: subepandrial sclerite tg: tergite. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7907358" httpUri="https://zenodo.org/record/7907358/files/figure.png" pageId="28" pageNumber="29">Fig. 17DF</figureCitation>
); S8 with a few, sparse white setae and with a mid-posterior spatulate projection slightly constricted medially with long dark brown to black macrosetae and setae laterally and apically, these macrosetae with white tips (
<figureCitation id="13362A1DEE5E3854BCF2F976F141F96F" box="[644,786,1699,1725]" captionStart="FIGURE 17" captionStartId="30.[151,250,1897,1921]" captionTargetBox="[218,1368,181,1873]" captionTargetId="figure-21@30.[218,1368,181,1873]" captionTargetPageId="30" captionText="FIGURE 17. Catostola complicata (James, 1953) comb. nov., holotype male. A. Wing, dorsal; B. Scutellum, dorsal; C. Hind femur, anterior; D. Terminalia, lateral; E. Same, dorsolateral; F. Same, dorsal; G. Same, ventral. Abbreviations: cerc: cercus; epand: epandrium; goncx: gonocoxite; gonst: gonostylus; hypd: hypandrium; st: sternite; subepand scl: subepandrial sclerite tg: tergite. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7907358" httpUri="https://zenodo.org/record/7907358/files/figure.png" pageId="28" pageNumber="29">Fig. 17DG</figureCitation>
); epandrium black with a group of dark reddish to black macrosetae proximally, ventrally and dorsally, very constricted medially forming a dorsal finger-like projection with reddish setae dorsally and ventrally, apex of projection more or less triangular with rounded tip, epandrial projection dark reddish internally (
<figureCitation id="13362A1DEE5E3854BC56F8DAF0FBF8FB" box="[544,680,1807,1833]" captionStart="FIGURE 17" captionStartId="30.[151,250,1897,1921]" captionTargetBox="[218,1368,181,1873]" captionTargetId="figure-21@30.[218,1368,181,1873]" captionTargetPageId="30" captionText="FIGURE 17. Catostola complicata (James, 1953) comb. nov., holotype male. A. Wing, dorsal; B. Scutellum, dorsal; C. Hind femur, anterior; D. Terminalia, lateral; E. Same, dorsolateral; F. Same, dorsal; G. Same, ventral. Abbreviations: cerc: cercus; epand: epandrium; goncx: gonocoxite; gonst: gonostylus; hypd: hypandrium; st: sternite; subepand scl: subepandrial sclerite tg: tergite. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7907358" httpUri="https://zenodo.org/record/7907358/files/figure.png" pageId="28" pageNumber="29">Fig. 17DF</figureCitation>
); hypandrium black at posterior corners with setae of same colour; gonocoxite black proximally becoming dark reddish distally, without setae and with a more or less triangular shape apically, rounded on tip; gonostylus dark reddish with short reddish setae; cercus reddish dark to black with short dark reddish to black setae and yellowish setae apically (
<figureCitation id="13362A1DEE5E3854BD63F8AEF1F3F847" box="[789,928,1915,1941]" captionStart="FIGURE 17" captionStartId="30.[151,250,1897,1921]" captionTargetBox="[218,1368,181,1873]" captionTargetId="figure-21@30.[218,1368,181,1873]" captionTargetPageId="30" captionText="FIGURE 17. Catostola complicata (James, 1953) comb. nov., holotype male. A. Wing, dorsal; B. Scutellum, dorsal; C. Hind femur, anterior; D. Terminalia, lateral; E. Same, dorsolateral; F. Same, dorsal; G. Same, ventral. Abbreviations: cerc: cercus; epand: epandrium; goncx: gonocoxite; gonst: gonostylus; hypd: hypandrium; st: sternite; subepand scl: subepandrial sclerite tg: tergite. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7907358" httpUri="https://zenodo.org/record/7907358/files/figure.png" pageId="28" pageNumber="29">Fig. 17DG</figureCitation>
); subepandrial sclerite with a pair of rounded lobes ventrally without setae, dorsal half black with yellowish setae apically.
</paragraph>
<paragraph id="8BB23698EE5E3854BEB1F816F0E6F80F" blockId="28.[151,1437,151,2013]" box="[199,693,1986,2013]" pageId="28" pageNumber="29">
<emphasis id="B979EA8AEE5E3854BEB1F816F377F80F" bold="true" box="[199,292,1987,2013]" pageId="28" pageNumber="29">Length.</emphasis>
Body: 17.85 mm; wing: 11.81 mm.
</paragraph>
</subSubSection>
<caption id="DF726610EE5F3855BEE1F842F11FF801" ID-DOI="http://doi.org/10.5281/zenodo.7907356" ID-Zenodo-Dep="7907356" httpUri="https://zenodo.org/record/7907356/files/figure.png" pageId="29" pageNumber="30" startId="29.[151,250,1943,1967]" targetBox="[203,1385,181,1919]" targetPageId="29" targetType="figure">
<paragraph id="8BB23698EE5F3855BEE1F842F11FF801" blockId="29.[151,1436,1943,2004]" pageId="29" pageNumber="30">
<emphasis id="B979EA8AEE5F3855BEE1F842F373F87D" bold="true" box="[151,288,1943,1967]" pageId="29" pageNumber="30">FIGURE 16.</emphasis>
<taxonomicName id="4C0D4D1BEE5F3855BF51F84DF0C4F87D" authority="(James, 1953)" baseAuthorityName="James" baseAuthorityYear="1953" box="[295,663,1943,1968]" class="Insecta" family="Asilidae" genus="Catostola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="complicata" status="comb. nov.">
<emphasis id="B979EA8AEE5F3855BF51F84DF051F862" box="[295,514,1944,1968]" italics="true" pageId="29" pageNumber="30">Catostola complicata</emphasis>
(
<bibRefCitation id="EF9C4B69EE5F3855BC66F84DF0C3F87D" author="James, M. T." box="[528,656,1943,1968]" pageId="29" pageNumber="30" pagination="46 - 57" refId="ref35216" refString="James, M. T. (1953) The Diptera collected on the Cockerell and Hubbell Expedition to Honduras: Part II. Asilidae. Journal of the Washington Academy of Sciences, 43 (2), 46 - 57." type="journal article" year="1953">James, 1953</bibRefCitation>
)
</taxonomicName>
<emphasis id="B979EA8AEE5F3855BCE8F84DF140F862" bold="true" box="[670,787,1944,1968]" pageId="29" pageNumber="30">
<taxonomicNameLabel id="A24A57F1EE5F3855BCE8F84DF140F862" box="[670,787,1944,1968]" pageId="29" pageNumber="30" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
, holotype male.
<emphasis id="B979EA8AEE5F3855BDC9F842F184F87D" bold="true" box="[959,983,1943,1967]" pageId="29" pageNumber="30">A.</emphasis>
Habitus, lateral (insert: labels, not to scale);
<emphasis id="B979EA8AEE5F3855BEE1F869F2FDF806" bold="true" box="[151,174,1980,2004]" pageId="29" pageNumber="30">B.</emphasis>
Same, dorsal;
<emphasis id="B979EA8AEE5F3855BF33F86EF30EF801" bold="true" box="[325,349,1979,2003]" pageId="29" pageNumber="30">C.</emphasis>
Head, frontal;
<emphasis id="B979EA8AEE5F3855BF81F869F043F806" bold="true" box="[503,528,1980,2004]" pageId="29" pageNumber="30">D.</emphasis>
Same, lateral. Scale bar 1 mm.
</paragraph>
</caption>
<caption id="DF726610EE5C3856BEE1F8BCF3D9F83E" ID-DOI="http://doi.org/10.5281/zenodo.7907358" ID-Zenodo-Dep="7907358" httpUri="https://zenodo.org/record/7907358/files/figure.png" pageId="30" pageNumber="31" startId="30.[151,250,1897,1921]" targetBox="[218,1368,181,1873]" targetPageId="30" targetType="figure">
<paragraph id="8BB23698EE5C3856BEE1F8BCF3D9F83E" blockId="30.[151,1437,1897,2029]" pageId="30" pageNumber="31">
<emphasis id="B979EA8AEE5C3856BEE1F8BCF373F853" bold="true" box="[151,288,1897,1921]" pageId="30" pageNumber="31">FIGURE 17.</emphasis>
<taxonomicName id="4C0D4D1BEE5C3856BF51F8BCF0C5F853" authority="(James, 1953)" baseAuthorityName="James" baseAuthorityYear="1953" box="[295,662,1897,1921]" class="Insecta" family="Asilidae" genus="Catostola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="complicata" status="comb. nov.">
<emphasis id="B979EA8AEE5C3856BF51F8BCF052F853" box="[295,513,1897,1921]" italics="true" pageId="30" pageNumber="31">Catostola complicata</emphasis>
(
<bibRefCitation id="EF9C4B69EE5C3856BC79F8BCF0DCF853" author="James, M. T." box="[527,655,1897,1921]" pageId="30" pageNumber="31" pagination="46 - 57" refId="ref35216" refString="James, M. T. (1953) The Diptera collected on the Cockerell and Hubbell Expedition to Honduras: Part II. Asilidae. Journal of the Washington Academy of Sciences, 43 (2), 46 - 57." type="journal article" year="1953">James, 1953</bibRefCitation>
)
</taxonomicName>
<emphasis id="B979EA8AEE5C3856BCEBF8BCF141F853" bold="true" box="[669,786,1897,1921]" pageId="30" pageNumber="31">
<taxonomicNameLabel id="A24A57F1EE5C3856BCEBF8BCF141F853" box="[669,786,1897,1921]" pageId="30" pageNumber="31" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
, holotype male.
<emphasis id="B979EA8AEE5C3856BDCBF8BCF185F853" bold="true" box="[957,982,1897,1921]" pageId="30" pageNumber="31">A.</emphasis>
Wing, dorsal;
<emphasis id="B979EA8AEE5C3856BA1AF8BCF6D0F853" bold="true" box="[1132,1155,1897,1921]" pageId="30" pageNumber="31">B.</emphasis>
Scutellum, dorsal;
<emphasis id="B979EA8AEE5C3856BB3CF8BCF730F853" bold="true" box="[1354,1379,1897,1921]" pageId="30" pageNumber="31">C.</emphasis>
Hind femur, anterior;
<emphasis id="B979EA8AEE5C3856BF4BF858F305F877" bold="true" box="[317,342,1933,1957]" pageId="30" pageNumber="31">D.</emphasis>
Terminalia, lateral;
<emphasis id="B979EA8AEE5C3856BC52F858F068F877" bold="true" box="[548,571,1933,1957]" pageId="30" pageNumber="31">E.</emphasis>
Same, dorsolateral;
<emphasis id="B979EA8AEE5C3856BD7BF858F173F877" bold="true" box="[781,800,1933,1957]" pageId="30" pageNumber="31">F.</emphasis>
Same, dorsal;
<emphasis id="B979EA8AEE5C3856BDCCF858F187F877" bold="true" box="[954,980,1933,1957]" pageId="30" pageNumber="31">G.</emphasis>
Same, ventral.
<emphasis id="B979EA8AEE5C3856BA02F858F746F877" bold="true" box="[1140,1301,1933,1957]" pageId="30" pageNumber="31">Abbreviations:</emphasis>
cerc: cercus; epand: epandrium; goncx: gonocoxite; gonst: gonostylus; hypd: hypandrium; st: sternite; subepand scl: subepandrial sclerite tg: tergite. Scale bar 1 mm.
</paragraph>
</caption>
<subSubSection id="C3176513EE5D3857BEB1FF42F03CFE7F" pageId="31" pageNumber="32" type="materials_examined">
<paragraph id="8BB23698EE5D3857BEB1FF42F14EFF63" blockId="31.[151,1437,150,684]" box="[199,797,150,177]" pageId="31" pageNumber="32">
<emphasis id="B979EA8AEE5D3857BEB1FF42F3E1FF62" bold="true" box="[199,434,150,177]" pageId="31" pageNumber="32">
<typeStatus id="54B6883AEE5D3857BEB1FF42F367FF63" box="[199,308,151,177]" pageId="31" pageNumber="32" type="holotype">Holotype</typeStatus>
condition.
</emphasis>
Postpedicel missing (
<figureCitation id="13362A1DEE5D3857BCDEFF42F143FF63" box="[680,784,151,177]" captionStart="FIGURE 16" captionStartId="29.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1919]" captionTargetId="figure-17@29.[203,1385,181,1919]" captionTargetPageId="29" captionText="FIGURE 16. Catostola complicata (James, 1953) comb. nov., holotype male. A. Habitus, lateral (insert: labels, not to scale); B. Same, dorsal; C. Head, frontal; D. Same, lateral. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7907356" httpUri="https://zenodo.org/record/7907356/files/figure.png" pageId="31" pageNumber="32">Fig. 16D</figureCitation>
).
</paragraph>
<paragraph id="8BB23698EE5D3857BEB1FF6EF3F1FF07" blockId="31.[151,1437,150,684]" box="[199,418,187,213]" pageId="31" pageNumber="32">
<emphasis id="B979EA8AEE5D3857BEB1FF6EF376FF07" bold="true" box="[199,293,187,213]" pageId="31" pageNumber="32">Female.</emphasis>
Unknown.
</paragraph>
<paragraph id="8BB23698EE5D3857BEB1FF0BF03CFE7F" blockId="31.[151,1437,150,684]" pageId="31" pageNumber="32">
<emphasis id="B979EA8AEE5D3857BEB1FF0BF383FF2A" bold="true" box="[199,464,222,248]" pageId="31" pageNumber="32">Taxonomic discussion.</emphasis>
This species can be differentiated from other
<taxonomicName id="4C0D4D1BEE5D3857BDC8FF0AF6D3FF2B" box="[958,1152,223,249]" pageId="31" pageNumber="32">
<emphasis id="B979EA8AEE5D3857BDC8FF0AF678FF2B" box="[958,1067,223,249]" italics="true" pageId="31" pageNumber="32">Catostola</emphasis>
species
</taxonomicName>
mainly by the yellow legs with apical third of mid and hind femora dark brown (
<figureCitation id="13362A1DEE5D3857BC86FED6F12FFECF" box="[752,892,259,285]" captionStart="FIGURE 16" captionStartId="29.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1919]" captionTargetId="figure-17@29.[203,1385,181,1919]" captionTargetPageId="29" captionText="FIGURE 16. Catostola complicata (James, 1953) comb. nov., holotype male. A. Habitus, lateral (insert: labels, not to scale); B. Same, dorsal; C. Head, frontal; D. Same, lateral. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7907356" httpUri="https://zenodo.org/record/7907356/files/figure.png" pageId="31" pageNumber="32">Figs 16AB</figureCitation>
,
<figureCitation id="13362A1DEE5D3857BDF1FED6F1E4FECF" box="[903,951,259,285]" captionStart="FIGURE 17" captionStartId="30.[151,250,1897,1921]" captionTargetBox="[218,1368,181,1873]" captionTargetId="figure-21@30.[218,1368,181,1873]" captionTargetPageId="30" captionText="FIGURE 17. Catostola complicata (James, 1953) comb. nov., holotype male. A. Wing, dorsal; B. Scutellum, dorsal; C. Hind femur, anterior; D. Terminalia, lateral; E. Same, dorsolateral; F. Same, dorsal; G. Same, ventral. Abbreviations: cerc: cercus; epand: epandrium; goncx: gonocoxite; gonst: gonostylus; hypd: hypandrium; st: sternite; subepand scl: subepandrial sclerite tg: tergite. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7907358" httpUri="https://zenodo.org/record/7907358/files/figure.png" pageId="31" pageNumber="32">17C</figureCitation>
) and the epandrium with a long, finger-like dorsal projection (
<figureCitation id="13362A1DEE5D3857BF12FEF2F3A3FE93" box="[356,496,295,321]" captionStart="FIGURE 17" captionStartId="30.[151,250,1897,1921]" captionTargetBox="[218,1368,181,1873]" captionTargetId="figure-21@30.[218,1368,181,1873]" captionTargetPageId="30" captionText="FIGURE 17. Catostola complicata (James, 1953) comb. nov., holotype male. A. Wing, dorsal; B. Scutellum, dorsal; C. Hind femur, anterior; D. Terminalia, lateral; E. Same, dorsolateral; F. Same, dorsal; G. Same, ventral. Abbreviations: cerc: cercus; epand: epandrium; goncx: gonocoxite; gonst: gonostylus; hypd: hypandrium; st: sternite; subepand scl: subepandrial sclerite tg: tergite. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7907358" httpUri="https://zenodo.org/record/7907358/files/figure.png" pageId="31" pageNumber="32">Fig. 17DG</figureCitation>
). The most similar species is
<taxonomicName id="4C0D4D1BEE5D3857BD4DFEF2F1A5FE93" baseAuthorityName="Walker" baseAuthorityYear="1849" box="[827,1014,295,321]" class="Insecta" family="Asilidae" genus="Catostola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="baleta" status="comb. nov.">
<emphasis id="B979EA8AEE5D3857BD4DFEF2F1A5FE93" box="[827,1014,295,321]" italics="true" pageId="31" pageNumber="32">Catostola baleta</emphasis>
</taxonomicName>
<emphasis id="B979EA8AEE5D3857BD8BFEF2F62DFE93" bold="true" box="[1021,1150,295,321]" pageId="31" pageNumber="32">
<taxonomicNameLabel id="A24A57F1EE5D3857BD8BFEF2F62DFE93" box="[1021,1150,295,321]" pageId="31" pageNumber="32" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
that can be distinguished through epandrial finger-like projection which is constricted short before the mid-length of the epandrium with the projection slightly longer than half of epandrial length (
<figureCitation id="13362A1DEE5D3857BD7BFEBAF1C0FE5B" box="[781,915,367,393]" captionStart="FIGURE 17" captionStartId="30.[151,250,1897,1921]" captionTargetBox="[218,1368,181,1873]" captionTargetId="figure-21@30.[218,1368,181,1873]" captionTargetPageId="30" captionText="FIGURE 17. Catostola complicata (James, 1953) comb. nov., holotype male. A. Wing, dorsal; B. Scutellum, dorsal; C. Hind femur, anterior; D. Terminalia, lateral; E. Same, dorsolateral; F. Same, dorsal; G. Same, ventral. Abbreviations: cerc: cercus; epand: epandrium; goncx: gonocoxite; gonst: gonostylus; hypd: hypandrium; st: sternite; subepand scl: subepandrial sclerite tg: tergite. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7907358" httpUri="https://zenodo.org/record/7907358/files/figure.png" pageId="31" pageNumber="32">Fig. 17DE</figureCitation>
) and the S8 mid-posterior projection which is slightly constricted at the base (
<figureCitation id="13362A1DEE5D3857BF8CFE46F031FE7F" box="[506,610,403,429]" captionStart="FIGURE 17" captionStartId="30.[151,250,1897,1921]" captionTargetBox="[218,1368,181,1873]" captionTargetId="figure-21@30.[218,1368,181,1873]" captionTargetPageId="30" captionText="FIGURE 17. Catostola complicata (James, 1953) comb. nov., holotype male. A. Wing, dorsal; B. Scutellum, dorsal; C. Hind femur, anterior; D. Terminalia, lateral; E. Same, dorsolateral; F. Same, dorsal; G. Same, ventral. Abbreviations: cerc: cercus; epand: epandrium; goncx: gonocoxite; gonst: gonostylus; hypd: hypandrium; st: sternite; subepand scl: subepandrial sclerite tg: tergite. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7907358" httpUri="https://zenodo.org/record/7907358/files/figure.png" pageId="31" pageNumber="32">Fig. 17G</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3176513EE5D3857BEB1FE63F7D0FDEF" pageId="31" pageNumber="32" type="distribution">
<paragraph id="8BB23698EE5D3857BEB1FE63F7D0FDEF" blockId="31.[151,1437,150,684]" pageId="31" pageNumber="32">
<emphasis id="B979EA8AEE5D3857BEB1FE63F382FE02" bold="true" box="[199,465,438,464]" pageId="31" pageNumber="32">
Distribution (
<figureCitation id="13362A1DEE5D3857BF1AFE63F397FE02" box="[364,452,438,464]" captionStart="FIGURE 44" captionStartId="66.[151,250,1682,1706]" captionTargetBox="[151,1435,525,1658]" captionTargetId="figure-172@66.[151,1435,525,1658]" captionTargetPageId="66" captionText="FIGURE 44. Distribution map of Catostola species: Catostola baleta (Walker, 1849) comb. nov., Catostola complicata (James, 1953) comb. nov., Catostola indecisa sp. nov., and Catostola martini (Fisher, 1992) comb. nov." figureDoi="http://doi.org/10.5281/zenodo.7907427" httpUri="https://zenodo.org/record/7907427/files/figure.png" pageId="31" pageNumber="32">Fig. 44</figureCitation>
).
</emphasis>
<collectingCountry id="F31A7608EE5D3857BFADFE62F01BFE03" box="[475,584,439,465]" name="Honduras" pageId="31" pageNumber="32">Honduras</collectingCountry>
(
<collectingRegion id="49C9F87AEE5D3857BC2CFE62F09AFE03" box="[602,713,439,465]" country="Honduras" name="Atlantida" pageId="31" pageNumber="32">Atlántida</collectingRegion>
). The original description was based on
<specimenCount id="9D0BFD11EE5D3857BAD7FE62F765FE03" box="[1185,1334,439,465]" pageId="31" pageNumber="32" type="generic" typeStatus="holotype">one holotype</specimenCount>
and
<specimenCount id="9D0BFD11EE5D3857BB04FE62F351FE27" pageId="31" pageNumber="32" type="generic" typeStatus="paratypes">two paratypes</specimenCount>
from the same locality as the
<typeStatus id="54B6883AEE5D3857BC39FE0EF0E2FE27" box="[591,689,475,501]" pageId="31" pageNumber="32" type="holotype">holotype</typeStatus>
but collected on
<date id="FFB31058EE5D3857BD06FE0EF1AEFE27" box="[880,1021,475,501]" pageId="31" pageNumber="32" value="1923-05-02">May 2, 1923</date>
(
<bibRefCitation id="EF9C4B69EE5D3857BA7DFE0EF6C9FE27" author="James, M. T." box="[1035,1178,474,501]" pageId="31" pageNumber="32" pagination="46 - 57" refId="ref35216" refString="James, M. T. (1953) The Diptera collected on the Cockerell and Hubbell Expedition to Honduras: Part II. Asilidae. Journal of the Washington Academy of Sciences, 43 (2), 46 - 57." type="journal article" year="1953">James, 1953</bibRefCitation>
). In this study, only the
<typeStatus id="54B6883AEE5D3857BEE1FE2AF2AAFDCB" box="[151,249,511,537]" pageId="31" pageNumber="32" type="holotype">holotype</typeStatus>
was examined.According to Erika Tucker (
<emphasis id="B979EA8AEE5D3857BCA9FE2AF149FDCB" box="[735,794,511,537]" italics="true" pageId="31" pageNumber="32">in litt</emphasis>
., 2019), curator at the University of Michigan Museum of Zoology (UMMZ), the
<typeStatus id="54B6883AEE5D3857BFE8FDF6F05AFDEF" box="[414,521,547,573]" pageId="31" pageNumber="32" type="paratype">paratypes</typeStatus>
were not located in the respective collection and its whereabouts are unknown.
</paragraph>
</subSubSection>
<subSubSection id="C3176513EE5D3857BEB1FD93F3E9FD7B" pageId="31" pageNumber="32" type="materials_examined">
<paragraph id="8BB23698EE5D3857BEB1FD93F3E9FD7B" blockId="31.[151,1437,150,684]" pageId="31" pageNumber="32">
<emphasis id="B979EA8AEE5D3857BEB1FD93F3FDFDB2" bold="true" box="[199,430,582,608]" pageId="31" pageNumber="32">Material examined.</emphasis>
<materialsCitation id="3B653CC5EE5D3857BFC5FD92F124FD57" collectionCode="T" collectorName="H. Hubbell &amp; James" country="Honduras" latitude="15.773334" location="Guaimas District" longLatPrecision="21" longitude="-87.46527" municipality="Tela" pageId="31" pageNumber="32" specimenCount="1" stateProvince="Atlantida" typeStatus="holotype">
<emphasis id="B979EA8AEE5D3857BFC5FD92F098FDB2" bold="true" box="[435,715,582,609]" pageId="31" pageNumber="32">
<typeStatus id="54B6883AEE5D3857BFC5FD92F071FDB3" box="[435,546,583,609]" pageId="31" pageNumber="32" type="holotype">Holotype</typeStatus>
.
<collectingCountry id="F31A7608EE5D3857BC5DFD93F098FDB2" box="[555,715,582,608]" name="Honduras" pageId="31" pageNumber="32">HONDURAS</collectingCountry>
</emphasis>
1923 [
<emphasis id="B979EA8AEE5D3857BD60FD93F1DBFDB3" bold="true" box="[790,904,582,609]" pageId="31" pageNumber="32">
<collectingRegion id="49C9F87AEE5D3857BD60FD93F1DBFDB3" box="[790,904,582,609]" country="Honduras" name="Atlantida" pageId="31" pageNumber="32">Atlántida</collectingRegion>
</emphasis>
]
<collectingMunicipality id="6BD6ACE2EE5D3857BDE3FD92F19AFDB3" box="[917,969,583,609]" pageId="31" pageNumber="32">Tela</collectingMunicipality>
: [
<geoCoordinate id="EE39505FEE5D3857BDAEFD9DF633FDB1" box="[984,1120,584,611]" degrees="15" direction="north" minutes="46" orientation="latitude" pageId="31" pageNumber="32" precision="15" seconds="24" value="15.773334">15°4624″N</geoCoordinate>
<geoCoordinate id="EE39505FEE5D3857BA12FD9DF6A6FDB1" box="[1124,1269,584,611]" degrees="87" direction="west" minutes="27" orientation="longitude" pageId="31" pageNumber="32" precision="15" seconds="55" value="-87.46527">87°2755″W</geoCoordinate>
], v.10
<location id="8ED26043EE5D3857BB4FFD92F2B8FD57" LSID="urn:lsid:plazi:treatment:03A4878EEE593857BEE1FA57F3E9FD7B:8ED26043EE5D3857BB4FFD92F2B8FD57" country="Honduras" latitude="15.773334" longLatPrecision="21" longitude="-87.46527" municipality="Tela" name="Guaimas District" pageId="31" pageNumber="32" stateProvince="Atlantida">Guaimas District</location>
508
<collectionCode id="ED1CAE5DEE5D3857BF5FFDBEF36FFD57" box="[297,316,619,645]" name="Tavera, Department of Geology and Geophysics" pageId="31" pageNumber="32">T</collectionCode>
.
<collectorName id="26F8534EEE5D3857BF36FDBEF3EDFD57" box="[320,446,619,645]" pageId="31" pageNumber="32">H. Hubbell</collectorName>
/
<taxonomicName id="4C0D4D1BEE5D3857BFADFDBEF0A2FD57" authorityName="James" authorityYear="1953" box="[475,753,619,645]" class="Insecta" family="Asilidae" genus="Pachychoeta" kingdom="Animalia" order="Diptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="complicata">
<emphasis id="B979EA8AEE5D3857BFADFDBEF0A2FD57" box="[475,753,619,645]" italics="true" pageId="31" pageNumber="32">Pachychoeta complicata</emphasis>
</taxonomicName>
1 ♁
<collectorName id="26F8534EEE5D3857BD45FDBEF124FD57" box="[819,887,619,645]" pageId="31" pageNumber="32">James</collectorName>
</materialsCitation>
<materialsCitation id="3B653CC5EE5D3857BDF4FDBEF3E0FD7A" collectionCode="UMMZ" country="Honduras" pageId="31" pageNumber="32" specimenCount="1" stateProvince="Atlantida" typeStatus="holotype">
<typeStatus id="54B6883AEE5D3857BDF4FDBEF645FD57" box="[898,1046,619,645]" pageId="31" pageNumber="32" type="holotype">HOLOTYPE</typeStatus>
/ bar code UMMZI-163425 (
<figureCitation id="13362A1DEE5D3857BB19FDBEF29BFD7B" captionStart="FIGURE 16" captionStartId="29.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1919]" captionTargetId="figure-17@29.[203,1385,181,1919]" captionTargetPageId="29" captionText="FIGURE 16. Catostola complicata (James, 1953) comb. nov., holotype male. A. Habitus, lateral (insert: labels, not to scale); B. Same, dorsal; C. Head, frontal; D. Same, lateral. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7907356" httpUri="https://zenodo.org/record/7907356/files/figure.png" pageId="31" pageNumber="32">Fig. 16A</figureCitation>
[insert])
<emphasis id="B979EA8AEE5D3857BF5FFD45F3E0FD7A" bold="true" box="[297,435,654,684]" pageId="31" pageNumber="32">
(♁
<collectionCode id="ED1CAE5DEE5D3857BF38FD5BF3FFFD7A" box="[334,428,654,680]" country="USA" httpUri="http://grbio.org/cool/ra64-3eva" name="University of Michigan, Museum of Zoology" pageId="31" pageNumber="32" type="University or college">UMMZ</collectionCode>
)
</emphasis>
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>