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<document id="D5995CAFF152B0AF6BBCB74F62B4995A" ID-DOI="10.11646/zootaxa.2142.1.3" ID-ISSN="1175-5326" ID-Zenodo-Dep="5319920" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629625288872" checkinUser="felipe" docAuthor="Domínguez, Eduardo, Molineri, Carlos &amp; Mariano, Rodolfo" docDate="2009" docId="993987A7FFB2FFDCDAF6FB66325EFC25" docLanguage="en" docName="zt02142p044.pdf" docOrigin="Zootaxa 2142 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.2142.1.3" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Hagenulopsis lipeo Domínguez, Molineri &amp; Mariano, 2009, sp. nov." docType="treatment" docVersion="4" lastPageNumber="40" masterDocId="6500FFDFFFB5FFD7DA61FFFC3700FFFB" masterDocTitle="Revision of the South American species of Hagenulopsis Ulmer and Askola Peters (Ephemeroptera: Leptophlebiidae) with description of six new species" masterLastPageNumber="44" masterPageNumber="29" pageNumber="36" updateTime="1699085826779" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="5FB3103C1FC41BAAF3A0CA0E5E82049C">Revision of the South American species of Hagenulopsis Ulmer and Askola Peters (Ephemeroptera: Leptophlebiidae) with description of six new species</mods:title>
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<mods:namePart id="6FA3A5A5F3E22981BFA22D7B52D21170">Domínguez, Eduardo</mods:namePart>
<mods:affiliation id="7FA2400A72BD9402C6066DA5F64C4D94">CONICET-Facultad de Ciencias Naturales e IML, Universidad Nacional de Tucumán, Argentina</mods:affiliation>
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<mods:namePart id="7F3C04AB88B6B915AE8CB1A7CF4E3688">Molineri, Carlos</mods:namePart>
<mods:affiliation id="A1D42D1668349FB8A808CC759657F1D4">CONICET-Facultad de Ciencias Naturales e IML, Universidad Nacional de Tucumán, Argentina</mods:affiliation>
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<mods:namePart id="4A1E2831BEFD6007200A743BFE8C6E12">Mariano, Rodolfo</mods:namePart>
<mods:affiliation id="9D60CAE604C871181461C1989E9B10DC">Universidade de São Paulo, Faculdade de Filosofia, Ciências e Letras de Ribeirão Preto, Departamento de Biologia, Laboratório de Entomologia Aquática, Ribeirão Preto, Brasil</mods:affiliation>
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<mods:date id="88703EB28712A53EF1DEEB35BFCD45F2">2009</mods:date>
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<treatment id="993987A7FFB2FFDCDAF6FB66325EFC25" ID-DOI="http://doi.org/10.5281/zenodo.5333451" ID-Zenodo-Dep="5333451" LSID="urn:lsid:plazi:treatment:993987A7FFB2FFDCDAF6FB66325EFC25" httpUri="http://treatment.plazi.org/id/993987A7FFB2FFDCDAF6FB66325EFC25" lastPageId="11" lastPageNumber="40" pageId="7" pageNumber="36">
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<paragraph id="112F36B1FFB2FFD0DAF6FB66349FFB4F" blockId="7.[151,927,1178,1204]" box="[151,927,1178,1204]" pageId="7" pageNumber="36">
<heading id="4A6781DDFFB2FFD0DAF6FB66349FFB4F" bold="true" box="[151,927,1178,1204]" fontSize="11" level="1" pageId="7" pageNumber="36" reason="1">
<emphasis id="23E4EAA3FFB2FFD0DAF6FB66349FFB4F" bold="true" box="[151,927,1178,1204]" pageId="7" pageNumber="36">
<taxonomicName id="D6904D32FFB2FFD0DAF6FB66367EFB4F" ID-CoL="3J9HQ" authorityName="Dominguez, Molineri &amp; Mariano" authorityYear="2009" box="[151,382,1178,1204]" class="Insecta" family="Leptophlebiidae" genus="Hagenulopsis" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="36" phylum="Arthropoda" rank="species" species="lipeo" status="sp. nov.">
<emphasis id="23E4EAA3FFB2FFD0DAF6FB66367EFB4F" bold="true" box="[151,382,1178,1204]" italics="true" pageId="7" pageNumber="36">Hagenulopsis lipeo</emphasis>
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<taxonomicNameLabel id="38D757D8FFB2FFD0DBE7FB6636E3FB4F" box="[390,483,1178,1204]" pageId="7" pageNumber="36" rank="species">sp. nov.</taxonomicNameLabel>
Domínguez, Molineri and Mariano
</emphasis>
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</paragraph>
<paragraph id="112F36B1FFB2FFD0DAF6FB18329CF893" blockId="7.[151,1436,1252,2012]" pageId="7" pageNumber="36">
Male imago (
<figureCitation id="89AB2A34FFB2FFD0DB52FB183689FB05" box="[307,393,1252,1278]" captionStart="FIGURES 1924" captionStartId="9.[151,271,1737,1761]" captionTargetBox="[207,1380,194,1713]" captionTargetId="figure-121@9.[207,1381,194,1713]" captionTargetPageId="9" captionText="FIGURES 1924. Hagenulopsis. Figs 1921, H. lipeo. 19, male imago, general aspect, lateral view; 20, head, dorsal view; 21, head, lateral view. Figs 2224, H. diptera. 22, male imago, general aspect, lateral view; 23, head, dorsal view; 24, head, lateral view." figureDoi="http://doi.org/10.5281/zenodo.5319932" httpUri="https://zenodo.org/record/5319932/files/figure.png" pageId="7" pageNumber="36">Fig. 19</figureCitation>
). Length: body, 5.0
<quantity id="D6689B54FFB2FFD0D81AFB1835D7FB05" box="[635,727,1252,1278]" metricMagnitude="-3" metricUnit="m" metricValue="5.8" pageId="7" pageNumber="36" unit="mm" value="5.8">5.8 mm</quantity>
; fore wings, 5.0
<quantity id="D6689B54FFB2FFD0D9C1FB1834FCFB05" box="[928,1020,1252,1278]" metricMagnitude="-3" metricUnit="m" metricValue="5.6" pageId="7" pageNumber="36" unit="mm" value="5.6">5.6 mm</quantity>
. Eyes: upper portion grayish brown meeting dorsally, basally contiguous; facets small and numerous (
<figureCitation id="89AB2A34FFB2FFD0D9F8FAF63329FADF" box="[921,1065,1290,1316]" captionStart="FIGURES 1924" captionStartId="9.[151,271,1737,1761]" captionTargetBox="[207,1380,194,1713]" captionTargetId="figure-121@9.[207,1381,194,1713]" captionTargetPageId="9" captionText="FIGURES 1924. Hagenulopsis. Figs 1921, H. lipeo. 19, male imago, general aspect, lateral view; 20, head, dorsal view; 21, head, lateral view. Figs 2224, H. diptera. 22, male imago, general aspect, lateral view; 23, head, dorsal view; 24, head, lateral view." figureDoi="http://doi.org/10.5281/zenodo.5319932" httpUri="https://zenodo.org/record/5319932/files/figure.png" pageId="7" pageNumber="36">Figs. 2021</figureCitation>
). Facets of lower portion black, small. Head yellowish brown with black markings close to base of eyes. Ocelli whitish, lateral ocelli twice the size of the median one. Antennae: scape yellowish white, pedicel brownish [flagellum broken off and lost]. Thorax. Nota and pleurae reddish brown with carinae darker and whitish membranes; sterna yellowish brown with whitish membranes. Leg I yellowish heavily washed with black specially on femora, tarsi lighter; legs IIIII with coxae and trochanter yellowish white, remainder segments yellowish washed with black specially at apex of femora; longitudinal lighter line on femora IIIII; claws in all legs yellowish washed with black. Wings membrane hyaline uniformly tinted with light brown, veins brownish lighter toward apex, pterostigma clouded white (
<figureCitation id="89AB2A34FFB2FFD0DB2DF9BC36A4F9A1" box="[332,420,1600,1626]" captionStart="FIGURES 3747" captionStartId="13.[151,269,1198,1222]" captionTargetBox="[211,1384,194,1168]" captionTargetId="figure-350@13.[202,1384,194,1174]" captionTargetPageId="13" captionText="FIGURES 3747. Hagenulopsis. Figs 3739, male fore wing: 37, H. zunigae sp. nov.; 38, H. esmeralda sp. nov.; 39, H. lipeo. Fig. 40, H. zunigae sp. nov., genitalia, ventral view. Figs 4142, H. esmeralda sp. nov.: 41, genitalia, ventral view; 42, penes, detail. Figs 4346, H. lipeo: 43, genitalia, ventral view; 44, penes, detail; 45, apex of styliger plate, ventral view; 46, idem, variation. Fig. 47, H. diptera, male head, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5319938" httpUri="https://zenodo.org/record/5319938/files/figure.png" pageId="7" pageNumber="36">Fig. 37</figureCitation>
); no costal cross veins basad to bulla. Abdomen. Terga yellowish heavily washed with black except on two submedial and two sublateral paler areas close to anterior margin on each terga II to VI; sterna yellowish white washed with black except anterior margin and ganglia; posterior margin of sterna VIIVIII translucid. Genitalia (
<figureCitation id="89AB2A34FFB2FFD0D867F9483598F935" box="[518,664,1716,1742]" captionStart="FIGURES 3747" captionStartId="13.[151,269,1198,1222]" captionTargetBox="[211,1384,194,1168]" captionTargetId="figure-350@13.[202,1384,194,1174]" captionTargetPageId="13" captionText="FIGURES 3747. Hagenulopsis. Figs 3739, male fore wing: 37, H. zunigae sp. nov.; 38, H. esmeralda sp. nov.; 39, H. lipeo. Fig. 40, H. zunigae sp. nov., genitalia, ventral view. Figs 4142, H. esmeralda sp. nov.: 41, genitalia, ventral view; 42, penes, detail. Figs 4346, H. lipeo: 43, genitalia, ventral view; 44, penes, detail; 45, apex of styliger plate, ventral view; 46, idem, variation. Fig. 47, H. diptera, male head, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5319938" httpUri="https://zenodo.org/record/5319938/files/figure.png" pageId="7" pageNumber="36">Figs. 4346</figureCitation>
): styliger plate and forceps yellowish brown except inner margins of forceps whitish, penes yellowish white; short styliger plate (ratio maximum width/medial length: 1.952.17) with a similar width along its entire length, hind margin straight to slightly concave; first forceps segment curved medially (ratio length F1/styliger: 2.02.3) (
<figureCitation id="89AB2A34FFB2FFD0D93AF8D434EBF8B9" box="[859,1003,1832,1858]" captionStart="FIGURES 3747" captionStartId="13.[151,269,1198,1222]" captionTargetBox="[211,1384,194,1168]" captionTargetId="figure-350@13.[202,1384,194,1174]" captionTargetPageId="13" captionText="FIGURES 3747. Hagenulopsis. Figs 3739, male fore wing: 37, H. zunigae sp. nov.; 38, H. esmeralda sp. nov.; 39, H. lipeo. Fig. 40, H. zunigae sp. nov., genitalia, ventral view. Figs 4142, H. esmeralda sp. nov.: 41, genitalia, ventral view; 42, penes, detail. Figs 4346, H. lipeo: 43, genitalia, ventral view; 44, penes, detail; 45, apex of styliger plate, ventral view; 46, idem, variation. Fig. 47, H. diptera, male head, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5319938" httpUri="https://zenodo.org/record/5319938/files/figure.png" pageId="7" pageNumber="36">Figs. 4344</figureCitation>
). Caudal filaments, basal half of first segment yellowish brown; rest of filaments yellowish black basally, turning yellowish gradually toward apex.
</paragraph>
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<subSubSection id="598A653AFFB2FFDDDAA7F88935CAFD53" lastPageId="10" lastPageNumber="39" pageId="7" pageNumber="36" type="description">
<paragraph id="112F36B1FFB2FFDFDAA7F88936DFFF23" blockId="7.[151,1436,1252,2012]" lastBlockId="8.[151,1436,152,216]" lastPageId="8" lastPageNumber="37" pageId="7" pageNumber="36">
Female imago. Length: body,
<quantity id="D6689B54FFB2FFD0D845F88935B4F874" box="[548,692,1909,1935]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="5.6" metricValueMin="4.4" pageId="7" pageNumber="36" unit="mm" value="5.0" valueMax="5.6" valueMin="4.4">4.45.6 mm</quantity>
; fore wings, 5.0
<quantity id="D6689B54FFB2FFD0D91FF88934DBF874" box="[894,987,1909,1935]" metricMagnitude="-3" metricUnit="m" metricValue="5.8" pageId="7" pageNumber="36" unit="mm" value="5.8">5.8 mm</quantity>
. Head yellowish brown except base of ocelli blackish, and area between ocelli and eyes whitish. Eyes black. Thorax yellowish brown, sterna yellowish white except darker medially. Some females have lighter wing membranes than the males. Abdomen ventrally with lighter areas covering anterior ¾ of segments IIV; egg guide yellowish black lighter toward apex, apically acute.
</paragraph>
<caption id="45EF6639FFBDFFDFDAF6F86E354BF814" ID-DOI="http://doi.org/10.5281/zenodo.5319928" ID-Zenodo-Dep="5319928" httpUri="https://zenodo.org/record/5319928/files/figure.png" pageId="8" pageNumber="37" startId="8.[151,269,1938,1962]" targetBox="[165,1421,259,1914]" targetPageId="8">
<paragraph id="112F36B1FFBDFFDFDAF6F86E354BF814" blockId="8.[151,1436,1938,2031]" pageId="8" pageNumber="37">
<emphasis id="23E4EAA3FFBDFFDFDAF6F86E365DF851" bold="true" box="[151,349,1938,1962]" pageId="8" pageNumber="37">FIGURES 1318.</emphasis>
<taxonomicName id="D6904D32FFBDFFDFDB02F86F36F6F851" box="[355,502,1939,1962]" class="Insecta" family="Leptophlebiidae" genus="Hagenulopsis" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="37" phylum="Arthropoda" rank="genus">
<emphasis id="23E4EAA3FFBDFFDFDB02F86F36F6F851" box="[355,502,1939,1962]" italics="true" pageId="8" pageNumber="37">Hagenulopsis</emphasis>
</taxonomicName>
. Figs 1315,
<taxonomicName id="D6904D32FFBDFFDFD8E7F86F35F9F851" authorityName="Dominguez, Molineri &amp; Mariano" authorityYear="2009" box="[646,761,1939,1962]" class="Insecta" family="Leptophlebiidae" genus="Hagenulopsis" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="37" phylum="Arthropoda" rank="species" species="zunigae" status="sp. nov.">
<emphasis id="23E4EAA3FFBDFFDFD8E7F86F35F9F851" box="[646,761,1939,1962]" italics="true" pageId="8" pageNumber="37">H. zunigae</emphasis>
</taxonomicName>
<emphasis id="23E4EAA3FFBDFFDFD961F86E3455F851" bold="true" box="[768,853,1938,1962]" pageId="8" pageNumber="37">
<taxonomicNameLabel id="38D757D8FFBDFFDFD961F86E3455F851" box="[768,853,1938,1962]" pageId="8" pageNumber="37" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
13, male imago, general aspect, lateral view; 14, head, dorsal view; 15, head, lateral view. Figs 1618,
<taxonomicName id="D6904D32FFBDFFDFD8FDF84A342DF836" authorityName="Dominguez, Molineri &amp; Bersoza" authorityYear="2009" box="[668,813,1974,1997]" class="Insecta" family="Leptophlebiidae" genus="Hagenulopsis" kingdom="Animalia" order="Ephemeroptera" pageId="8" pageNumber="37" phylum="Arthropoda" rank="species" species="esmeralda" status="sp. nov.">
<emphasis id="23E4EAA3FFBDFFDFD8FDF84A342DF836" box="[668,813,1974,1997]" italics="true" pageId="8" pageNumber="37">H. esmeralda</emphasis>
</taxonomicName>
<emphasis id="23E4EAA3FFBDFFDFD955F849348AF836" bold="true" box="[820,906,1973,1997]" pageId="8" pageNumber="37">
<taxonomicNameLabel id="38D757D8FFBDFFDFD955F849348AF836" box="[820,906,1973,1997]" pageId="8" pageNumber="37" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
16, male imago, general aspect, lateral view; 17, head, dorsal view; 18, head, lateral view.
</paragraph>
</caption>
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<paragraph id="112F36B1FFBCFFDEDAF6F9353681F8DD" blockId="9.[151,1436,1737,1830]" pageId="9" pageNumber="38">
<emphasis id="23E4EAA3FFBCFFDEDAF6F9353661F91A" bold="true" box="[151,353,1737,1761]" pageId="9" pageNumber="38">FIGURES 1924.</emphasis>
<taxonomicName id="D6904D32FFBCFFDEDB08F93636FEF91A" box="[361,510,1738,1761]" class="Insecta" family="Leptophlebiidae" genus="Hagenulopsis" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="38" phylum="Arthropoda" rank="genus">
<emphasis id="23E4EAA3FFBCFFDEDB08F93636FEF91A" box="[361,510,1738,1761]" italics="true" pageId="9" pageNumber="38">Hagenulopsis</emphasis>
</taxonomicName>
. Figs 1921,
<taxonomicName id="D6904D32FFBCFFDED8F4F93635EDF91A" authorityName="Dominguez, Molineri &amp; Mariano" authorityYear="2009" box="[661,749,1738,1761]" class="Insecta" family="Leptophlebiidae" genus="Hagenulopsis" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="38" phylum="Arthropoda" rank="species" species="lipeo">
<emphasis id="23E4EAA3FFBCFFDED8F4F93635EDF91A" box="[661,749,1738,1761]" italics="true" pageId="9" pageNumber="38">H. lipeo</emphasis>
</taxonomicName>
. 19, male imago, general aspect, lateral view; 20, head, dorsal view; 21, head, lateral view. Figs 2224,
<taxonomicName id="D6904D32FFBCFFDED831F91035BEF8F8" authorityName="Ulmer" authorityYear="1920" box="[592,702,1772,1795]" class="Insecta" family="Leptophlebiidae" genus="Hagenulopsis" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="38" phylum="Arthropoda" rank="species" species="diptera">
<emphasis id="23E4EAA3FFBCFFDED831F91035BEF8F8" box="[592,702,1772,1795]" italics="true" pageId="9" pageNumber="38">H. diptera</emphasis>
</taxonomicName>
. 22, male imago, general aspect, lateral view; 23, head, dorsal view; 24, head, lateral view.
</paragraph>
</caption>
<paragraph id="112F36B1FFBCFFDDDAA7F8A835CAFD53" blockId="9.[151,1436,1876,2018]" lastBlockId="10.[151,1436,152,680]" lastPageId="10" lastPageNumber="39" pageId="9" pageNumber="38">
Mature nymphs. Female length: body,
<quantity id="D6689B54FFBCFFDED8FDF8A83430F895" box="[668,816,1876,1902]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="5.6" metricValueMin="5.4" pageId="9" pageNumber="38" unit="mm" value="5.5" valueMax="5.6" valueMin="5.4">5.45.6 mm</quantity>
, cerci
<quantity id="D6689B54FFBCFFDED9E3F8A83315F895" box="[898,1045,1876,1902]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" metricValueMax="3.9" metricValueMin="3.7" pageId="9" pageNumber="38" unit="mm" value="3.8" valueMax="3.9" valueMin="3.7">3.73.9 mm</quantity>
, terminal filament, 4.85.0 mm. Male length: body
<quantity id="D6689B54FFBCFFDEDB1AF887350EF86E" box="[379,526,1915,1941]" metricMagnitude="-3" metricUnit="m" metricValue="5.449999999999999" metricValueMax="5.6" metricValueMin="5.3" pageId="9" pageNumber="38" unit="mm" value="5.449999999999999" valueMax="5.6" valueMin="5.3">5.35.6 mm</quantity>
, cerci
<quantity id="D6689B54FFBCFFDED83CF88735BCF86E" box="[605,700,1915,1941]" metricMagnitude="-3" metricUnit="m" metricValue="5.3" pageId="9" pageNumber="38" unit="mm" value="5.3">5.3 mm</quantity>
, terminal filament,
<quantity id="D6689B54FFBCFFDED9C6F8873307F86E" box="[935,1031,1915,1941]" metricMagnitude="-3" metricUnit="m" metricValue="3.6" pageId="9" pageNumber="38" unit="mm" value="3.6">3.6 mm</quantity>
. Head. Eyes of male with reddish black upper portion and blackish lower portion; eyes of female black; ocelli whitish with inner half margin blackish. Antennae: scape yellowish, pedicel blackish, flagellum yellowish lighter toward apex. Dorsum of head yellowish brown stained with black in central area. Mouthparts yellowish stained with black at: base of labrum, basal half of mandibles, outer margin and palpi segment I of maxilla, posterior margin of superlinguae, and submentum; molae of mandible and apical row of setae of maxilla orange brown; anteromedian emargination of labrum with 3 denticles, the median smaller than lateral ones. Thorax. nota yellowish white with latero-medial and latero-posterior areas washed with black; pro- and mesosternum yellowish white with ganglia and margins stained with black, metasternum yellowish white almost completely washed with black except inner margin of articulation of coxae. Leg I light orange yellow with coxa and margins of trochanters washed with black, femur almost completely washed with black except lighter oblicuous line. Legs II and III yellowish with coxa and two longitudinal lateromedial areas on femora, and longitudinal stripe on tibiae blackish; tibiae and tarsi darker than femora, claws orange brown. Tarsal claws with 89 denticles. Abdomen dorsally yellowish orange washed heavily with black especially on lateroposterior areas of terga; sterna yellowish with central area tinged with black, lighter in central area. [Posterolateral projection as in
<taxonomicName id="D6904D32FFBFFFDDD876FD94356AFD7A" authorityName="Spieth" authorityYear="1943" box="[535,618,616,641]" class="Insecta" family="Leptophlebiidae" genus="Hagenulopsis" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="39" phylum="Arthropoda" rank="species" species="minuta">
<emphasis id="23E4EAA3FFBFFFDDD876FD94356AFD7A" box="[535,618,616,641]" italics="true" pageId="10" pageNumber="39">minuta</emphasis>
</taxonomicName>
description]. Gills whitish with blackish tracheae. Cerci yellowish orange, few setae on each intersegmental union.
</paragraph>
</subSubSection>
<caption id="45EF6639FFBFFFDDDAF6F8913275F809" ID-DOI="http://doi.org/10.5281/zenodo.5319934" ID-Zenodo-Dep="5319934" httpUri="https://zenodo.org/record/5319934/files/figure.png" pageId="10" pageNumber="39" startId="10.[151,269,1901,1925]" targetBox="[195,1385,727,1874]" targetPageId="10">
<paragraph id="112F36B1FFBFFFDDDAF6F8913275F809" blockId="10.[151,1436,1901,2035]" pageId="10" pageNumber="39">
<emphasis id="23E4EAA3FFBFFFDDDAF6F891365DF87E" bold="true" box="[151,349,1901,1925]" pageId="10" pageNumber="39">FIGURES 2536.</emphasis>
<taxonomicName id="D6904D32FFBFFFDDDB02F89236ABF87E" box="[355,427,1902,1925]" class="Insecta" family="Leptophlebiidae" genus="Askola" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="39" phylum="Arthropoda" rank="genus">
<emphasis id="23E4EAA3FFBFFFDDDB02F89236ABF87E" box="[355,427,1902,1925]" italics="true" pageId="10" pageNumber="39">Askola</emphasis>
</taxonomicName>
. Figs 2527, male fore wing: 25,
<taxonomicName id="D6904D32FFBFFFDDD972F89234D3F87E" authorityName="Dominguez, Molineri &amp; Mariano" authorityYear="2009" box="[787,979,1902,1925]" class="Insecta" family="Leptophlebiidae" genus="Askola" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="39" phylum="Arthropoda" rank="species" species="emmerichi" status="sp. nov.">
<emphasis id="23E4EAA3FFBFFFDDD972F89234D3F87E" box="[787,979,1902,1925]" italics="true" pageId="10" pageNumber="39">Askola emmerichi</emphasis>
</taxonomicName>
<emphasis id="23E4EAA3FFBFFFDDD9B8F8913328F87E" bold="true" box="[985,1064,1901,1925]" pageId="10" pageNumber="39">
<taxonomicNameLabel id="38D757D8FFBFFFDDD9B8F8913328F87E" box="[985,1064,1901,1925]" pageId="10" pageNumber="39" rank="species">sp.nov.</taxonomicNameLabel>
</emphasis>
; 26,
<taxonomicName id="D6904D32FFBFFFDDDE3CF8923211F87E" authorityName="Dominguez, Molineri &amp; Mariano" authorityYear="2009" box="[1117,1297,1902,1925]" class="Insecta" family="Leptophlebiidae" genus="Askola" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="39" phylum="Arthropoda" rank="species" species="paprockii" status="sp. nov.">
<emphasis id="23E4EAA3FFBFFFDDDE3CF8923211F87E" box="[1117,1297,1902,1925]" italics="true" pageId="10" pageNumber="39">Askola paprockii</emphasis>
</taxonomicName>
<emphasis id="23E4EAA3FFBFFFDDDF79F891326DF87E" bold="true" box="[1304,1389,1901,1925]" pageId="10" pageNumber="39">
<taxonomicNameLabel id="38D757D8FFBFFFDDDF79F891326DF87E" box="[1304,1389,1901,1925]" pageId="10" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
; 27,
<taxonomicName id="D6904D32FFBFFFDDDAF6F86E364AF852" authorityName="Dominguez, Molineri &amp; Mariano" authorityYear="2009" box="[151,330,1938,1961]" class="Insecta" family="Leptophlebiidae" genus="Askola" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="39" phylum="Arthropoda" rank="species" species="cipoensis" status="sp. nov.">
<emphasis id="23E4EAA3FFBFFFDDDAF6F86E364AF852" box="[151,330,1938,1961]" italics="true" pageId="10" pageNumber="39">Askola cipoensis</emphasis>
</taxonomicName>
<emphasis id="23E4EAA3FFBFFFDDDB33F86D36A8F852" bold="true" box="[338,424,1937,1961]" pageId="10" pageNumber="39">
<taxonomicNameLabel id="38D757D8FFBFFFDDDB33F86D36A8F852" box="[338,424,1937,1961]" pageId="10" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Figs 2830,
<taxonomicName id="D6904D32FFBFFFDDD857F86E35F7F852" authorityName="Dominguez, Molineri &amp; Mariano" authorityYear="2009" box="[566,759,1938,1961]" class="Insecta" family="Leptophlebiidae" genus="Askola" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="39" phylum="Arthropoda" rank="species" species="emmerichi" status="sp. nov.">
<emphasis id="23E4EAA3FFBFFFDDD857F86E35F7F852" box="[566,759,1938,1961]" italics="true" pageId="10" pageNumber="39">Askola emmerichi</emphasis>
</taxonomicName>
<emphasis id="23E4EAA3FFBFFFDDD89FF86D344DF852" bold="true" box="[766,845,1937,1961]" pageId="10" pageNumber="39">
<taxonomicNameLabel id="38D757D8FFBFFFDDD89FF86D344DF852" box="[766,845,1937,1961]" pageId="10" pageNumber="39" rank="species">sp.nov.</taxonomicNameLabel>
</emphasis>
: 28, genitalia, ventral view; 29, genitalia, lateral view; 30, penes, detail. Figs 3133,
<taxonomicName id="D6904D32FFBFFFDDDBB4F84A3589F836" authorityName="Dominguez, Molineri &amp; Mariano" authorityYear="2009" box="[469,649,1974,1997]" class="Insecta" family="Leptophlebiidae" genus="Askola" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="39" phylum="Arthropoda" rank="species" species="paprockii" status="sp. nov.">
<emphasis id="23E4EAA3FFBFFFDDDBB4F84A3589F836" box="[469,649,1974,1997]" italics="true" pageId="10" pageNumber="39">Askola paprockii</emphasis>
</taxonomicName>
<emphasis id="23E4EAA3FFBFFFDDD8EEF84935E4F836" bold="true" box="[655,740,1973,1997]" pageId="10" pageNumber="39">
<taxonomicNameLabel id="38D757D8FFBFFFDDD8EEF84935E4F836" box="[655,740,1973,1997]" pageId="10" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
: 31, genitalia, ventral view; 32, genitalia, lateral view; 33, penes, detail. Figs 3436,
<taxonomicName id="D6904D32FFBFFFDDDB05F8273516F809" authorityName="Dominguez, Molineri &amp; Mariano" authorityYear="2009" box="[356,534,2011,2034]" class="Insecta" family="Leptophlebiidae" genus="Askola" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="39" phylum="Arthropoda" rank="species" species="cipoensis" status="sp. nov.">
<emphasis id="23E4EAA3FFBFFFDDDB05F8273516F809" box="[356,534,2011,2034]" italics="true" pageId="10" pageNumber="39">Askola cipoensis</emphasis>
</taxonomicName>
<emphasis id="23E4EAA3FFBFFFDDD87CF8273572F808" bold="true" box="[541,626,2011,2035]" pageId="10" pageNumber="39">
<taxonomicNameLabel id="38D757D8FFBFFFDDD87CF8273572F808" box="[541,626,2011,2035]" pageId="10" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
: 34, genitalia, ventral view; 35, genitalia, lateral view; 36, penes, detail.
</paragraph>
</caption>
<subSubSection id="598A653AFFBEFFDCDAA7FF6B35F8FE61" pageId="11" pageNumber="40" type="materials_examined">
<paragraph id="112F36B1FFBEFFDCDAA7FF6B35F8FE61" blockId="11.[151,1437,151,990]" pageId="11" pageNumber="40">
<emphasis id="23E4EAA3FFBEFFDCDAA7FF6B363DFF4A" bold="true" box="[198,317,151,177]" pageId="11" pageNumber="40">Material:</emphasis>
<materialsCitation id="A1F83CECFFBEFFDCDB25FF6435E5FF23" collectingDate="2004-11-13" collectorName="PN Baritu" country="Argentina" pageId="11" pageNumber="40" specimenCount="106" specimenCount-female="8" specimenCount-male="98" stateProvince="Salta" typeStatus="holotype">
<typeStatus id="CE2B8813FFBEFFDCDB25FF6436B1FF49" box="[324,433,152,178]" pageId="11" pageNumber="40" type="holotype">Holotype</typeStatus>
male imago from
<collectingCountry id="69877621FFBEFFDCD8E9FF643438FF49" box="[648,824,152,178]" name="Argentina" pageId="11" pageNumber="40">ARGENTINA</collectingCountry>
:
<collectingRegion id="D354F853FFBEFFDCD923FF64347FFF49" box="[834,895,152,178]" country="Argentina" name="Salta" pageId="11" pageNumber="40">Salta</collectingRegion>
,
<collectorName id="BC655367FFBEFFDCD9EAFF643303FF49" box="[907,1027,152,178]" pageId="11" pageNumber="40">PN Baritu</collectorName>
, Lipeo, estación 5,
<date id="652E1071FFBEFFDCDE84FF643274FF49" box="[1253,1396,152,178]" pageId="11" pageNumber="40" value="2004-11-13">
<collectingDate id="756AE999FFBEFFDCDE84FF643274FF49" box="[1253,1396,152,178]" pageId="11" pageNumber="40" value="2004-11-13">13-XI-2004</collectingDate>
</date>
;
<specimenCount id="0796FD38FFBEFFDCDF1EFF6437D0FF23" pageId="11" pageNumber="40" type="male">98 male</specimenCount>
and
<specimenCount id="0796FD38FFBEFFDCDB6DFF423673FF23" box="[268,371,190,216]" pageId="11" pageNumber="40" type="female">8 female</specimenCount>
imagos same data as holotype
</materialsCitation>
.
<materialsCitation id="A1F83CECFFBEFFDCD890FF423301FF04" collectingDate="2004-11-13" collectorName="E. Dominguez" country="Argentina" county="Parque Nacional Baritu" location="Los Naranjos" municipality="Lipeo" pageId="11" pageNumber="40" specimenCount="1" stateProvince="Salta" typeStatus="paratype">
<typeStatus id="CE2B8813FFBEFFDCD890FF423469FF23" box="[753,873,190,216]" pageId="11" pageNumber="40" type="paratype">Paratypes</typeStatus>
: 18 nymphs from
<collectingCountry id="69877621FFBEFFDCDE22FF4233F4FF23" box="[1091,1268,190,216]" name="Argentina" pageId="11" pageNumber="40">ARGENTINA</collectingCountry>
:
<collectingRegion id="D354F853FFBEFFDCDF61FF42323EFF23" box="[1280,1342,190,216]" country="Argentina" name="Salta" pageId="11" pageNumber="40">Salta</collectingRegion>
,
<collectingCounty id="F84E4E3DFFBEFFDCDF2AFF423655FF04" pageId="11" pageNumber="40">Parque Nacional Baritu</collectingCounty>
,
<collectingMunicipality id="F14BACCBFFBEFFDCDB00FF1936A8FF04" box="[353,424,229,255]" pageId="11" pageNumber="40">Lipeo</collectingMunicipality>
, Aº
<location id="144F606AFFBEFFDCDBBAFF19357CFF04" LSID="urn:lsid:plazi:treatment:993987A7FFB2FFDCDAF6FB66325EFC25:144F606AFFBEFFDCDBBAFF19357CFF04" box="[475,636,229,255]" country="Argentina" county="Parque Nacional Baritu" municipality="Lipeo" name="Los Naranjos" pageId="11" pageNumber="40" stateProvince="Salta">Los Naranjos</location>
,
<date id="652E1071FFBEFFDCD8E9FF193414FF04" box="[648,788,229,255]" pageId="11" pageNumber="40" value="2004-11-13">
<collectingDate id="756AE999FFBEFFDCD8E9FF193414FF04" box="[648,788,229,255]" pageId="11" pageNumber="40" value="2004-11-13">13-XI-2004</collectingDate>
</date>
,
<collectorName id="BC655367FFBEFFDCD940FF1934CBFF04" box="[801,971,229,255]" pageId="11" pageNumber="40">E. Domínguez</collectorName>
col.
</materialsCitation>
;
<materialsCitation id="A1F83CECFFBEFFDCDE6FFF1936FEFEDD" collectingDate="2004-11-17" collectorName="E. Dominguez" country="Argentina" county="Parque Nacional Baritu" location="Baritu" municipality="Lipeo" pageId="11" pageNumber="40" specimenCount="1" stateProvince="Salta" typeStatus="paratype">
2 nymphs, same data as holotype, except Aº
<location id="144F606AFFBEFFDCDB75FEF03661FEDD" LSID="urn:lsid:plazi:treatment:993987A7FFB2FFDCDAF6FB66325EFC25:144F606AFFBEFFDCDB75FEF03661FEDD" box="[276,353,268,294]" country="Argentina" county="Parque Nacional Baritu" municipality="Lipeo" name="Baritu" pageId="11" pageNumber="40" stateProvince="Salta">Baritu</location>
,
<date id="652E1071FFBEFFDCDB0FFEF036FEFEDD" box="[366,510,268,294]" pageId="11" pageNumber="40" value="2004-11-17">
<collectingDate id="756AE999FFBEFFDCDB0FFEF036FEFEDD" box="[366,510,268,294]" pageId="11" pageNumber="40" value="2004-11-17">17-XI-2004</collectingDate>
</date>
</materialsCitation>
.
<materialsCitation id="A1F83CECFFBEFFDCD86AFEF035F4FE61" collectingDate="1993-04-19" collectingDateMax="2000-03-03" collectingDateMin="1993-04-19" collectionCode="IFML, MZSP" collectorName="El Carmen &amp; Dominguez &amp; Molineri &amp; E. Dominguez &amp; M. Paco &amp; All" country="Bolivia" county="Other" elevation="1100" location="Las Lanzas" municipality="El Carmen" pageId="11" pageNumber="40" specimenCount="10" specimenCount-male="10" stateProvince="Santa Cruz">
<collectingCounty id="F84E4E3DFFBEFFDCD86AFEF0354FFEDD" box="[523,591,268,294]" pageId="11" pageNumber="40">Other</collectingCounty>
material: Jujuy,
<collectorName id="BC655367FFBEFFDCD979FEF0349CFEDD" box="[792,924,268,294]" pageId="11" pageNumber="40">
<collectingMunicipality id="F14BACCBFFBEFFDCD979FEF0349CFEDD" box="[792,924,268,294]" pageId="11" pageNumber="40">El Carmen</collectingMunicipality>
</collectorName>
, Aº
<location id="144F606AFFBEFFDCD9B1FEF0335AFEDD" LSID="urn:lsid:plazi:treatment:993987A7FFB2FFDCDAF6FB66325EFC25:144F606AFFBEFFDCD9B1FEF0335AFEDD" box="[976,1114,268,294]" country="Bolivia" county="Other" municipality="El Carmen" name="Las Lanzas" pageId="11" pageNumber="40" stateProvince="Santa Cruz">Las Lanzas</location>
,
<date id="652E1071FFBEFFDCDE06FEF033E6FEDD" box="[1127,1254,268,294]" pageId="11" pageNumber="40" value="2000-03-03">
<collectingDate id="756AE999FFBEFFDCDE06FEF033E6FEDD" box="[1127,1254,268,294]" pageId="11" pageNumber="40" value="2000-03-03">3-III-2000</collectingDate>
</date>
,
<collectorName id="BC655367FFBEFFDCDE92FEF0327DFEDD" box="[1267,1405,268,294]" pageId="11" pageNumber="40">Domínguez</collectorName>
&amp;
<collectorName id="BC655367FFBEFFDCDAF6FECE3602FEB7" box="[151,258,306,332]" pageId="11" pageNumber="40">Molineri</collectorName>
cols.
<collectingCountry id="69877621FFBEFFDCDB2CFECE36CEFEB7" box="[333,462,306,332]" name="Bolivia" pageId="11" pageNumber="40">BOLIVIA</collectingCountry>
: Dpto.
<collectingRegion id="D354F853FFBEFFDCD848FECE35B4FEB7" box="[553,692,306,332]" country="Bolivia" name="Santa Cruz" pageId="11" pageNumber="40">Santa Cruz</collectingRegion>
,
<location id="144F606AFFBEFFDCD8A0FECE3455FEB7" LSID="urn:lsid:plazi:treatment:993987A7FFB2FFDCDAF6FB66325EFC25:144F606AFFBEFFDCD8A0FECE3455FEB7" box="[705,853,306,332]" country="Bolivia" county="Other" municipality="El Carmen" name="Ichilo Prov." pageId="11" pageNumber="40" stateProvince="Santa Cruz">Ichilo Prov.</location>
,
<locationDeviation id="009D2F93FFBEFFDCD903FECE33F4FEB7" box="[866,1268,306,332]" location="Hierba Buena Militar" pageId="11" pageNumber="40">
<quantity id="D6689B54FFBEFFDCD903FECE34B0FEB7" box="[866,944,306,332]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="11" pageNumber="40" unit="km" value="10.0">10 km</quantity>
NE Hierba Buena Militar
</locationDeviation>
,
<location id="144F606AFFBEFFDCDF63FECE37EFFE88" LSID="urn:lsid:plazi:treatment:993987A7FFB2FFDCDAF6FB66325EFC25:144F606AFFBEFFDCDF63FECE37EFFE88" country="Bolivia" county="Other" municipality="El Carmen" name="Quebrada El Bronce" pageId="11" pageNumber="40" stateProvince="Santa Cruz">Quebrada El Bronce</location>
,
<quantity id="D6689B54FFBEFFDCDA9AFEA53656FE88" box="[251,342,345,371]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="11" pageNumber="40" unit="m" value="1100.0">
<elevation id="9ABDD182FFBEFFDCDA9AFEA53656FE88" box="[251,342,345,371]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="11" pageNumber="40" unit="m" value="1100.0">1100 m</elevation>
</quantity>
,
<date id="652E1071FFBEFFDCDB02FEA536EEFE88" box="[355,494,345,371]" pageId="11" pageNumber="40" value="1993-04-19">
<collectingDate id="756AE999FFBEFFDCDB02FEA536EEFE88" box="[355,494,345,371]" pageId="11" pageNumber="40" value="1993-04-19">19-IV-1993</collectingDate>
</date>
,
<collectorName id="BC655367FFBEFFDCDB9AFEA535A5FE88" box="[507,677,345,371]" pageId="11" pageNumber="40">E. Domínguez</collectorName>
&amp;
<collectorName id="BC655367FFBEFFDCD8ADFEA5342FFE88" box="[716,815,345,371]" pageId="11" pageNumber="40">M. Paco</collectorName>
cols.
<collectorName id="BC655367FFBEFFDCD917FEA5349CFE88" box="[886,924,345,371]" pageId="11" pageNumber="40">All</collectorName>
material deposited at
<collectionCode id="7781AE74FFBEFFDCDEC3FEA533EBFE88" box="[1186,1259,345,371]" pageId="11" pageNumber="40">IFML</collectionCode>
, except
<specimenCount id="0796FD38FFBEFFDCDF2DFEA5329CFE88" box="[1356,1436,345,371]" pageId="11" pageNumber="40" type="male">5 male</specimenCount>
imagos at
<collectionCode id="7781AE74FFBEFFDCDB6EFE7C365BFE61" box="[271,347,384,410]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="11" pageNumber="40" type="Museum">MZSP</collectionCode>
and
<specimenCount id="0796FD38FFBEFFDCDBF4FE7C36E3FE61" box="[405,483,384,410]" pageId="11" pageNumber="40" type="male">5 male</specimenCount>
imagos at MUSENUV
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="598A653AFFBEFFDCDAA7FE5A35DAFE3B" box="[198,730,422,448]" pageId="11" pageNumber="40" type="etymology">
<paragraph id="112F36B1FFBEFFDCDAA7FE5A35DAFE3B" blockId="11.[151,1437,151,990]" box="[198,730,422,448]" pageId="11" pageNumber="40">
<emphasis id="23E4EAA3FFBEFFDCDAA7FE5A3657FE3B" bold="true" box="[198,343,422,448]" pageId="11" pageNumber="40">Etymology:</emphasis>
Lipeo, name of the
<typeStatus id="CE2B8813FFBEFFDCD822FE5A3575FE3B" box="[579,629,422,448]" pageId="11" pageNumber="40">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="598A653AFFBEFFDCDAA7FE3134D2FD34" pageId="11" pageNumber="40" type="biology_ecology">
<paragraph id="112F36B1FFBEFFDCDAA7FE3134D2FD34" blockId="11.[151,1437,151,990]" pageId="11" pageNumber="40">
<emphasis id="23E4EAA3FFBEFFDCDAA7FE31362EFE1C" bold="true" box="[198,302,461,487]" pageId="11" pageNumber="40">Biology:</emphasis>
<taxonomicName id="D6904D32FFBEFFDCDB5DFE313525FE1D" authorityName="Dominguez, Molineri &amp; Mariano" authorityYear="2009" box="[316,549,461,486]" class="Insecta" family="Leptophlebiidae" genus="Hagenulopsis" kingdom="Animalia" order="Ephemeroptera" pageId="11" pageNumber="40" phylum="Arthropoda" rank="species" species="lipeo" status="sp. nov.">
<emphasis id="23E4EAA3FFBEFFDCDB5DFE313525FE1D" box="[316,549,461,486]" italics="true" pageId="11" pageNumber="40">Hagenulopsis lipeo</emphasis>
</taxonomicName>
<emphasis id="23E4EAA3FFBEFFDCD853FE313596FE1C" bold="true" box="[562,662,461,487]" pageId="11" pageNumber="40">
<taxonomicNameLabel id="38D757D8FFBEFFDCD853FE313596FE1C" box="[562,662,461,487]" pageId="11" pageNumber="40" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
was collected in a small effluent of the Lipeo river, in
<collectingRegion id="D354F853FFBEFFDCDF00FE313602FDF5" country="Argentina" name="Salta" pageId="11" pageNumber="40">Salta Province</collectingRegion>
, NW
<collectingCountry id="69877621FFBEFFDCDB24FE0836BDFDF5" box="[325,445,500,526]" name="Argentina" pageId="11" pageNumber="40">Argentina</collectingCountry>
. The stream was about
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wide and between
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to
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deep. The bottom was covered with rocks, with some sandy areas. A mating flight was observed during the morning hours,
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above the water course, performing the typical up and down pattern of most leptophlebiid species. It is interesting to note that the pattern does not seem, at least on the main features, to be affected by the dipterous conditions of the specimens. The size of the swarm was approximately
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long and
<quantity id="D6689B54FFBEFFDCDE1AFD7233A8FD53" box="[1147,1192,654,680]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="11" pageNumber="40" unit="m" value="1.0">1 m</quantity>
width, and extended for about an hour. A few females were caught among the flying males.
</paragraph>
</subSubSection>
<subSubSection id="598A653AFFBEFFDCDAA7FD27325EFC25" pageId="11" pageNumber="40" type="discussion">
<paragraph id="112F36B1FFBEFFDCDAA7FD27325EFC25" blockId="11.[151,1437,151,990]" pageId="11" pageNumber="40">
<emphasis id="23E4EAA3FFBEFFDCDAA7FD273654FD0E" bold="true" box="[198,340,731,757]" pageId="11" pageNumber="40">Discussion:</emphasis>
This species seems to have a rather extended distribution from NW
<collectingCountry id="69877621FFBEFFDCDECBFD20321FFD0D" box="[1194,1311,732,758]" name="Argentina" pageId="11" pageNumber="40">Argentina</collectingCountry>
to central
<collectingCountry id="69877621FFBEFFDCDAF6FCFE37F1FCE7" box="[151,241,770,796]" name="Bolivia" pageId="11" pageNumber="40">Bolivia</collectingCountry>
. It has been observed certain variation in the shape of the apex of the male styliger plate (
<figureCitation id="89AB2A34FFBEFFDCDF61FCFE328FFCE7" box="[1280,1423,770,796]" captionStart="FIGURES 3747" captionStartId="13.[151,269,1198,1222]" captionTargetBox="[211,1384,194,1168]" captionTargetId="figure-350@13.[202,1384,194,1174]" captionTargetPageId="13" captionText="FIGURES 3747. Hagenulopsis. Figs 3739, male fore wing: 37, H. zunigae sp. nov.; 38, H. esmeralda sp. nov.; 39, H. lipeo. Fig. 40, H. zunigae sp. nov., genitalia, ventral view. Figs 4142, H. esmeralda sp. nov.: 41, genitalia, ventral view; 42, penes, detail. Figs 4346, H. lipeo: 43, genitalia, ventral view; 44, penes, detail; 45, apex of styliger plate, ventral view; 46, idem, variation. Fig. 47, H. diptera, male head, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5319938" httpUri="https://zenodo.org/record/5319938/files/figure.png" pageId="11" pageNumber="40">Figs. 4546</figureCitation>
), that is due to the angle of observation. It can be separated from the other- species of the genus by the following combination of characters. Imago: 1) general coloration dark (yellowish, heavily washed with black), pattern as in
<figureCitation id="89AB2A34FFBEFFDCDBF9FC8A36F3FC6B" box="[408,499,886,912]" captionStart="FIGURES 1924" captionStartId="9.[151,271,1737,1761]" captionTargetBox="[207,1380,194,1713]" captionTargetId="figure-121@9.[207,1381,194,1713]" captionTargetPageId="9" captionText="FIGURES 1924. Hagenulopsis. Figs 1921, H. lipeo. 19, male imago, general aspect, lateral view; 20, head, dorsal view; 21, head, lateral view. Figs 2224, H. diptera. 22, male imago, general aspect, lateral view; 23, head, dorsal view; 24, head, lateral view." figureDoi="http://doi.org/10.5281/zenodo.5319932" httpUri="https://zenodo.org/record/5319932/files/figure.png" pageId="11" pageNumber="40">Fig. 19</figureCitation>
;
<figureCitation id="89AB2A34FFBEFFDCD863FC8A3512FC6B" box="[514,530,886,912]" captionStart="FIGURES 16" captionStartId="4.[151,272,1935,1959]" captionTargetBox="[252,1334,187,1911]" captionTargetId="figure-65@4.[252,1335,187,1911]" captionTargetPageId="4" captionText="FIGURES 16. Askola. Figs 13, Askola emmerichi, sp. nov., male imago. 1, general aspect, lateral view; 2, head, dorsal view; 3, head, lateral view. Figs 46, Askola paprockii sp. nov., male imago. 4, body, dorsal view; 5, idem, ventral view; 6, head, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5319924" httpUri="https://zenodo.org/record/5319924/files/figure.png" pageId="11" pageNumber="40">2</figureCitation>
) eyes of male meeting on -meson of head (
<figureCitation id="89AB2A34FFBEFFDCDE53FC8A33CBFC6B" box="[1074,1227,886,912]" captionStart="FIGURES 1924" captionStartId="9.[151,271,1737,1761]" captionTargetBox="[207,1380,194,1713]" captionTargetId="figure-121@9.[207,1381,194,1713]" captionTargetPageId="9" captionText="FIGURES 1924. Hagenulopsis. Figs 1921, H. lipeo. 19, male imago, general aspect, lateral view; 20, head, dorsal view; 21, head, lateral view. Figs 2224, H. diptera. 22, male imago, general aspect, lateral view; 23, head, dorsal view; 24, head, lateral view." figureDoi="http://doi.org/10.5281/zenodo.5319932" httpUri="https://zenodo.org/record/5319932/files/figure.png" pageId="11" pageNumber="40">Figs. 2021</figureCitation>
); 3) first forceps segment not more than 2.3 times length of styliger plate, and curved medially (
<figureCitation id="89AB2A34FFBEFFDCDE0EFC6133CCFC4C" box="[1135,1228,925,951]" captionStart="FIGURES 3747" captionStartId="13.[151,269,1198,1222]" captionTargetBox="[211,1384,194,1168]" captionTargetId="figure-350@13.[202,1384,194,1174]" captionTargetPageId="13" captionText="FIGURES 3747. Hagenulopsis. Figs 3739, male fore wing: 37, H. zunigae sp. nov.; 38, H. esmeralda sp. nov.; 39, H. lipeo. Fig. 40, H. zunigae sp. nov., genitalia, ventral view. Figs 4142, H. esmeralda sp. nov.: 41, genitalia, ventral view; 42, penes, detail. Figs 4346, H. lipeo: 43, genitalia, ventral view; 44, penes, detail; 45, apex of styliger plate, ventral view; 46, idem, variation. Fig. 47, H. diptera, male head, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5319938" httpUri="https://zenodo.org/record/5319938/files/figure.png" pageId="11" pageNumber="40">Fig. 43</figureCitation>
); 4) hind femora yellowish washed with black, heavier on apical 1/4, with a longitudinal lighter line; 5) cerci dark brown.
</paragraph>
</subSubSection>
</treatment>
</document>