176 lines
18 KiB
XML
176 lines
18 KiB
XML
<document id="E59F873454B476F5FBBEF4BF83E35657" ID-DOI="10.26107/RBZ-2020-0052" ID-ISSN="2345-7600" ID-Zenodo-Dep="4576861" ID-ZooBank="C7D4631B-9B02-4E6E-9665-76CCFECA86A0" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1614780803714" checkinUser="carolina" docAuthor="Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Buczkó, Krisztina & Vermeulen, Jaap J." docDate="2020" docId="9565D05FFFC5FFC5FCA05B54FB109D99" docLanguage="en" docName="RafflesBZool.68.350-368.pdf" docOrigin="Raffles Bulletin of Zoology 68" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Clostophis koilobasis Pall-Gergely & Vermeulen 2020, new species" docType="treatment" docVersion="4" lastPageNumber="358" masterDocId="695CA827FFCDFFCDFF9A5A29FFDE9B6C" masterDocTitle="Clostophis Benson, 1860, is not a monotypic diplommatinid but a speciose hypselostomatid (Gastropoda: Eupulmonata), with descriptions of six new species" masterLastPageNumber="368" masterPageNumber="350" pageNumber="358" updateTime="1698878356270" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="6CA4F2B2F1BC1115BBC5D38216AE26A6">Clostophis Benson, 1860, is not a monotypic diplommatinid but a speciose hypselostomatid (Gastropoda: Eupulmonata), with descriptions of six new species</mods:title>
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<mods:name id="4247E44B48D8AD23E23A6C1E3406612D" type="personal">
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<mods:roleTerm id="F26440BDC7A091D3B8461EFA5D4D0EE4">Author</mods:roleTerm>
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<mods:namePart id="E52EA7B8362FD941A2256A9DDA3CBBCE">Páll-Gergely, Barna</mods:namePart>
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<mods:roleTerm id="12EA776618CDDD99A46B455F0564115D">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="3F3690E920A4A3B6C711B671DEF9A991">Hunyadi, András</mods:namePart>
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</mods:name>
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<mods:name id="529C8CE37B94C49FBD18817D294C99C3" type="personal">
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<mods:roleTerm id="632D3DFC195BF0D51F103314AA997338">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="35D33881AA6EF91CAAE7712FDC61E03C">Grego, Jozef</mods:namePart>
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</mods:name>
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<mods:name id="D0B28970DACAE2616B9E08E41CAA0252" type="personal">
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<mods:roleTerm id="E75A4B5A2567F34AB16218585D723527">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="CF2EBD64C5925A41343D23300493F531">Reischütz, Alexander</mods:namePart>
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</mods:name>
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<mods:name id="D4CF87F2640B5741DF845DF0004672A9" type="personal">
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</mods:role>
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<mods:namePart id="A648BA1E849E13EB8C46381A8FD021D5">Buczkó, Krisztina</mods:namePart>
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</mods:name>
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<mods:name id="5BE8EDF6C36A174C33A99A46DB37CD12" type="personal">
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<mods:roleTerm id="7B97D2EDEF151F94D3F982BD2651389F">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="E1CAD4E5378663AB783E4CFF8CF3A387">Vermeulen, Jaap J.</mods:namePart>
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<mods:typeOfResource id="EA57EA1B6ADBCC350BDB64648D19DDF6">text</mods:typeOfResource>
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<mods:title id="57CD789BFD7C6DB1AFD4377D76E8FC8D">Raffles Bulletin of Zoology</mods:title>
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<mods:part id="9EDC8B55C54FF3698E7CF90AD8BE0909">
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<mods:date id="D0E42D96082139603214FD1D4F3427EB">2020</mods:date>
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<mods:detail id="237B60DE56CC624A8F96ECB4B4FB5E95" type="pubDate">
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<mods:number id="5EA236D34BCB839BB09445E70F2D960A">2020-05-08</mods:number>
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<mods:number id="8B9D38FEE70B0636A44493019B53743C">68</mods:number>
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<mods:start id="CA9E52CCE9B13C468E51385FFD4542ED">350</mods:start>
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<mods:end id="067DE50BE199FC6FDE7C8B72BC254742">368</mods:end>
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<mods:classification id="720506915C160D4DA3844FA6287E7D09">journal article</mods:classification>
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<mods:identifier id="27CE854B0C2195627441CB64EDE25646" type="DOI">10.26107/RBZ-2020-0052</mods:identifier>
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<mods:identifier id="98E3F57BE136DCA6408D72FDB33423E4" type="ISSN">2345-7600</mods:identifier>
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<mods:identifier id="DC88EC6B482D385EE2822460303C3B18" type="Zenodo-Dep">4576861</mods:identifier>
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<treatment id="9565D05FFFC5FFC5FCA05B54FB109D99" ID-DOI="http://doi.org/10.5281/zenodo.4576688" ID-Zenodo-Dep="4576688" LSID="urn:lsid:plazi:treatment:9565D05FFFC5FFC5FCA05B54FB109D99" httpUri="http://treatment.plazi.org/id/9565D05FFFC5FFC5FCA05B54FB109D99" lastPageNumber="358" pageId="8" pageNumber="358">
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<subSubSection id="55D632C2FFC5FFC5FCA05B54FB489AD9" pageId="8" pageNumber="358" type="nomenclature">
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<paragraph id="1D736149FFC5FFC5FCA05B54FB489AD9" blockId="8.[826,1439,380,469]" pageId="8" pageNumber="358">
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<heading id="463BD625FFC5FFC5FCA05B54FB489AD9" bold="true" centered="true" fontSize="10" level="2" pageId="8" pageNumber="358" reason="2">
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<emphasis id="2FB8BD5BFFC5FFC5FCA05B54FB489AD9" bold="true" pageId="8" pageNumber="358">
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<taxonomicName id="DACC1ACAFFC5FFC5FCA05B54FABF9AF9" ID-CoL="7QQQ2" authority="Pall-Gergely & Vermeulen" authorityName="Pall-Gergely & Vermeulen" authorityYear="2020" box="[826,1377,380,405]" class="Gastropoda" family="Gastrocoptidae" genus="Clostophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="358" phylum="Mollusca" rank="species" species="koilobasis" status="sp. nov.">
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<emphasis id="2FB8BD5BFFC5FFC5FCA05B54FBFA9AF9" bold="true" box="[826,1060,381,405]" italics="true" pageId="8" pageNumber="358">Clostophis koilobasis</emphasis>
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||
Páll-Gergely & Vermeulen
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</taxonomicName>
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,
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<taxonomicNameLabel id="348B0020FFC5FFC5FAEB5B54FB489AD9" pageId="8" pageNumber="358" rank="species">new species</taxonomicNameLabel>
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||
</emphasis>
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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<subSubSection id="55D632C2FFC5FFC5FBD95B94FB4B9AB9" box="[1091,1173,445,469]" pageId="8" pageNumber="358" type="description">
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<paragraph id="1D736149FFC5FFC5FBD95B94FB4B9AB9" blockId="8.[826,1439,380,469]" box="[1091,1173,445,469]" pageId="8" pageNumber="358">
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(
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<figureCitation id="85F77DCCFFC5FFC5FBD15B94FB539AB9" box="[1099,1165,445,469]" captionStart="Fig" captionStartId="9.[136,171,2035,2057]" captionTargetBox="[172,1438,190,2020]" captionTargetId="figure-8@9.[172,1438,190,2020]" captionTargetPageId="9" captionText="Fig. 6. Clostophis koilobasis Páll-Gergely & Vermeulen, new species (NHMUK 20191126, holotype). A, apertural view; B, dorsal view; C, ventral view; D, lateral view; E–F, apertural dentition; G, protoconch sculpture; H, sculpture of the ventral side; I, sculpture of body whorl (lateral side)." figureDoi="http://doi.org/10.5281/zenodo.4576874" httpUri="https://zenodo.org/record/4576874/files/figure.png" pageId="8" pageNumber="358">Fig. 6</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="55D632C2FFC5FFC5FCB75BD4FC1699D9" pageId="8" pageNumber="358" type="materials_examined">
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<paragraph id="1D736149FFC5FFC5FCB75BD4FC1699D9" blockId="8.[813,1451,509,693]" pageId="8" pageNumber="358">
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<emphasis id="2FB8BD5BFFC5FFC5FCB75BD4FC049979" bold="true" box="[813,986,509,533]" pageId="8" pageNumber="358">Type material.</emphasis>
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<materialsCitation id="ADA46B14FFC5FFC5FC7E5BD4FC1699D9" collectingDate="1998-10-01" collectionCode="NHMUK, JJV" collectorName="Cat Ba Isl., NE & J. J. Vermeulen & A. J. Whitten" country="Vietnam" latitude="20.820833" location="Cat Ba Isl." longLatPrecision="12" longitude="107.0575" pageId="8" pageNumber="358" specimenCode="NHMUK 20191126, JJV 6230" specimenCount="1" stateProvince="Hai Phong" typeStatus="holotype">
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<typeStatus id="C277DFEBFFC5FFC5FC7E5BD4FB979979" box="[996,1097,509,533]" pageId="8" pageNumber="358" type="holotype">Holotype</typeStatus>
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(SH:
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<quantity id="DA34CCACFFC5FFC5FB0A5BD4FB299979" box="[1168,1271,509,533]" metricMagnitude="-3" metricUnit="m" metricValue="1.06" pageId="8" pageNumber="358" unit="mm" value="1.06">1.06 mm</quantity>
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, SW:
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<quantity id="DA34CCACFFC5FFC5FAA55BD4FA7A9979" box="[1343,1444,509,533]" metricMagnitude="-3" metricUnit="m" metricValue="1.22" pageId="8" pageNumber="358" unit="mm" value="1.22">1.22 mm</quantity>
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) (
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<specimenCode id="4D6AC932FFC5FFC5FCAE5834FBC39959" box="[820,1053,541,565]" collectionCode="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="8" pageNumber="358" type="Museum">NHMUK 20191126</specimenCode>
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; original inv. number:
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<specimenCode id="4D6AC932FFC5FFC5FAB45837FA419959" box="[1326,1439,541,565]" collectionCode="JJV" pageId="8" pageNumber="358">JJV 6230</specimenCode>
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),
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<collectingCountry id="65DB21D9FFC5FFC5FCB75814FC529939" box="[813,908,573,597]" name="Vietnam" pageId="8" pageNumber="358">Vietnam</collectingCountry>
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,
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<collectingRegion id="DF08AFABFFC5FFC5FC0F5814FB9E9939" box="[917,1088,573,597]" country="Vietnam" name="Hai Phong" pageId="8" pageNumber="358">Haiphong Prov.</collectingRegion>
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,
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<collectorName id="B039049FFFC5FFC5FBD35814FB379939" box="[1097,1257,573,597]" pageId="8" pageNumber="358">
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<location id="18133792FFC5FFC5FBD35814FB649939" LSID="urn:lsid:plazi:treatment:9565D05FFFC5FFC5FCA05B54FB109D99:18133792FFC5FFC5FBD35814FB649939" box="[1097,1210,573,597]" country="Vietnam" latitude="20.820833" longLatPrecision="12" longitude="107.0575" name="Cat Ba Isl." pageId="8" pageNumber="358" stateProvince="Hai Phong">Cat Ba Isl.</location>
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, NE
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</collectorName>
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coast,
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<geoCoordinate id="78F8078EFFC5FFC5FAAB5814FA759939" box="[1329,1451,572,597]" degrees="20" direction="north" minutes="49.25" orientation="latitude" pageId="8" pageNumber="358" precision="9" value="20.820833">20°49.25′N</geoCoordinate>
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<geoCoordinate id="78F8078EFFC5FFC5FCB75874FC6A9919" box="[813,948,604,629]" degrees="107" direction="east" minutes="03.45" orientation="longitude" pageId="8" pageNumber="358" precision="9" value="107.0575">107°03.45′E</geoCoordinate>
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, dry limestone scree slope with low woodland, in rock crevices, leg.
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<collectorName id="B039049FFFC5FFC5FBB95857FB1699F9" box="[1059,1224,637,661]" pageId="8" pageNumber="358">J.J. Vermeulen</collectorName>
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&
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<collectorName id="B039049FFFC5FFC5FB685857FA5C99F9" box="[1266,1410,637,662]" pageId="8" pageNumber="358">A.J. Whitten</collectorName>
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,
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<date id="69724789FFC5FFC5FA0A5854FC1A99D9" pageId="8" pageNumber="358" value="1998-10-01">
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<collectingDate id="7936BE61FFC5FFC5FA0A5854FC1A99D9" pageId="8" pageNumber="358" value="1998-10-01">01 October 1998</collectingDate>
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</date>
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.
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="55D632C2FFC5FFC5FCB758F4FC049859" pageId="8" pageNumber="358" type="diagnosis">
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<paragraph id="1D736149FFC5FFC5FCB758F4FC049859" blockId="8.[812,1452,733,821]" pageId="8" pageNumber="358">
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<emphasis id="2FB8BD5BFFC5FFC5FCB758F4FC7F9999" bold="true" box="[813,929,733,757]" pageId="8" pageNumber="358">Diagnosis.</emphasis>
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A low conical shell with strongly widened body whorl, very wide umbilicus, and an elevated, distorted, short parietal lamella.
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</paragraph>
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</subSubSection>
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<subSubSection id="55D632C2FFC5FFC5FCB75974FAEF9E79" pageId="8" pageNumber="358" type="description">
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<paragraph id="1D736149FFC5FFC5FCB75974FAEF9E79" blockId="8.[811,1452,861,1301]" pageId="8" pageNumber="358">
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<emphasis id="2FB8BD5BFFC5FFC5FCB75974FC639819" bold="true" box="[813,957,861,885]" pageId="8" pageNumber="358">Description.</emphasis>
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Shell low triangular, wider than tall, with concave sides and strongly widened last whorl. Body whorl rounded, aperture not detached. Protoconch consisting of 1.25 whorls, with very finely raised threads arranged into hardly visible, widely-spaced spiral striae. Entire shell consisting of 4.75 whorls. Teleoconch with irregular, fine growth lines and dense (body whorl) and widely-spaced (previous whorls) spiral striation. Spiral striae ca. 22 on body whorl from lateral view (the
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<typeStatus id="C277DFEBFFC5FFC5FC085E74FC309F19" box="[914,1006,1117,1141]" pageId="8" pageNumber="358" type="holotype">holotype</typeStatus>
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was strongly corroded, therefore the spiral striae were not counted from apertural view). Umbilicus very wide, occupies almost half of the shell width. Aperture subovate, peristome very slightly expanded, not reflected. Parietal tooth elevated, strong, but short, distorted internally (oblique to the growth axis of the body whorl).
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</paragraph>
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</subSubSection>
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<subSubSection id="55D632C2FFC5FFC5FCB75F14FB5C9EF9" pageId="8" pageNumber="358" type="etymology">
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<paragraph id="1D736149FFC5FFC5FCB75F14FB5C9EF9" blockId="8.[811,1452,1341,1429]" pageId="8" pageNumber="358">
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<emphasis id="2FB8BD5BFFC5FFC5FCB75F14FC719E39" bold="true" box="[813,943,1341,1365]" pageId="8" pageNumber="358">Etymology.</emphasis>
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Combination of ‘koilos’ (= hollow in Greek) and ‘basis’ (= base in Greek), referring to the wide umbilicus as well as to the concave spire.
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</paragraph>
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</subSubSection>
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<subSubSection id="55D632C2FFC5FFC5FCB75F94FB109D99" pageId="8" pageNumber="358" type="diagnosis">
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<paragraph id="1D736149FFC5FFC5FCB75F94FB109D99" blockId="8.[811,1454,1469,1781]" pageId="8" pageNumber="358">
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<emphasis id="2FB8BD5BFFC5FFC5FCB75F94FBF29EB9" bold="true" box="[813,1068,1469,1493]" pageId="8" pageNumber="358">Differential diagnosis.</emphasis>
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<taxonomicName id="DACC1ACAFFC5FFC5FBAC5F94FAD19EB9" authority="Páll-Gergely & Hunyadi & Grego & Reischütz & Buczkó & Vermeulen, 2020" authorityName="Páll-Gergely & Hunyadi & Grego & Reischütz & Buczkó & Vermeulen" authorityYear="2020" box="[1078,1295,1469,1493]" class="Gastropoda" family="Gastrocoptidae" genus="Clostophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="358" phylum="Mollusca" rank="species" species="incurvus" status="sp. nov.">
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<emphasis id="2FB8BD5BFFC5FFC5FBAC5F94FAD19EB9" box="[1078,1295,1469,1493]" italics="true" pageId="8" pageNumber="358">Clostophis incurvus</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="348B0020FFC5FFC5FABA5F97FA799EB9" box="[1312,1447,1469,1493]" pageId="8" pageNumber="358" rank="species">new species</taxonomicNameLabel>
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, which is also known from the Halong Bay area, has a taller shell with narrower umbilicus and a strong parietal tooth.
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<taxonomicName id="DACC1ACAFFC5FFC5FCB75C34FBF79D59" authority="Páll-Gergely & Hunyadi & Grego & Reischütz & Buczkó & Vermeulen, 2020" authorityName="Páll-Gergely & Hunyadi & Grego & Reischütz & Buczkó & Vermeulen" authorityYear="2020" box="[813,1065,1565,1589]" class="Gastropoda" family="Gastrocoptidae" genus="Clostophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="358" phylum="Mollusca" rank="species" species="platytrochus" status="sp. nov.">
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<emphasis id="2FB8BD5BFFC5FFC5FCB75C34FBF79D59" box="[813,1065,1565,1589]" italics="true" pageId="8" pageNumber="358">Clostophis platytrochus</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="348B0020FFC5FFC5FBAF5C37FB689D59" box="[1077,1206,1565,1589]" pageId="8" pageNumber="358" rank="species">new species</taxonomicNameLabel>
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, is somewhat similar in shell shape, but has a slightly keeled body whorl, a detached terminal part of the peristome, a strongly oblique aperture, and a strong palatal tooth. Some populations of
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<taxonomicName id="DACC1ACAFFC5FFC5FAA15C54FC769DD9" authority="Páll-Gergely & Hunyadi & Grego & Reischütz & Buczkó & Vermeulen, 2020" authorityName="Páll-Gergely & Hunyadi & Grego & Reischütz & Buczkó & Vermeulen" authorityYear="2020" class="Gastropoda" family="Gastrocoptidae" genus="Clostophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="358" phylum="Mollusca" rank="species" species="multiformis" status="sp. nov.">
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<emphasis id="2FB8BD5BFFC5FFC5FAA15C54FC769DD9" italics="true" pageId="8" pageNumber="358">Clostophis multiformis</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="348B0020FFC5FFC5FC2D5CB7FBE29DD9" box="[951,1084,1693,1717]" pageId="8" pageNumber="358" rank="species">new species</taxonomicNameLabel>
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, are similar to
|
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<taxonomicName id="DACC1ACAFFC5FFC5FB7E5CB7FAB19DD9" authority="Páll-Gergely & Hunyadi & Grego & Reischütz & Buczkó & Vermeulen, 2020" authorityName="Páll-Gergely & Hunyadi & Grego & Reischütz & Buczkó & Vermeulen" authorityYear="2020" box="[1252,1391,1693,1717]" class="Gastropoda" family="Gastrocoptidae" genus="Clostophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="358" phylum="Mollusca" rank="species" species="koilobasis" status="sp. nov.">
|
||
<emphasis id="2FB8BD5BFFC5FFC5FB7E5CB7FAB19DD9" box="[1252,1391,1693,1717]" italics="true" pageId="8" pageNumber="358">C. koilobasis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="348B0020FFC5FFC5FAE55CB7FCA19DB9" pageId="8" pageNumber="358" rank="species">new species</taxonomicNameLabel>
|
||
, in shell shape, but that species always has a low parietal tooth and narrower umbilicus.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |