360 lines
57 KiB
XML
360 lines
57 KiB
XML
<document id="E87C77F24F2586D029E1817A7353F3A4" ID-DOI="10.1649/0010-065X(2001)055[0185:DAROPN]2.0.CO;2" ID-ISSN="1938-4394" ID-Zenodo-Dep="10102726" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630155689397" checkinUser="tatiana" docAuthor="Cicchino, Armando C. & Roig- Juñent, Sergio" docDate="2001" docId="5E5B1142FB7EFFAAE67B5163FE5BF95D" docLanguage="en" docName="0010-065X_2001_055_0185_DAROPN_2.0.CO_2.pdf.imf" docOrigin="The Coleopterists Bulletin 55 (2)" docStyle="DocumentStyle:37D8AFFB4AA6BE9981973FD5310B493E.6:TheColeopBull.2000-2002.journal_article.0cover" docStyleId="37D8AFFB4AA6BE9981973FD5310B493E" docStyleName="TheColeopBull.2000-2002.journal_article.0cover" docStyleVersion="6" docTitle="Paranillopsis Cicchino & Juñent, 2001, new genus" docType="treatment" docVersion="3" lastPageNumber="189" masterDocId="A262693AFB7DFFAFE447506CFFF8FF04" masterDocTitle="Description And Relationships Of Paranillopsis New Genus, Two New Species From Argentina, And A Key To The Neotropical Genera Of The Subtribe Anillina (Coleoptera: Carabidae: Bembidiini)" masterLastPageNumber="193" masterPageNumber="185" pageNumber="188" updateTime="1699622613882" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:mods id="9F3C3BDA77DF8DF87B3EF51534C2C664" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="30E13490596C7DC46885CA9C8594E532">
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<mods:title id="01991AEBC21BFA711E61A1B7BAE97415">Description And Relationships Of Paranillopsis New Genus, Two New Species From Argentina, And A Key To The Neotropical Genera Of The Subtribe Anillina (Coleoptera: Carabidae: Bembidiini)</mods:title>
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<mods:name id="DF01D9A302FF8DBC35DA8E9FEFE1EBD6" type="personal">
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<mods:roleTerm id="5E1B838DD768547368AAB0A23B28A751">Author</mods:roleTerm>
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<mods:namePart id="29D532A4C4D08B7D3AECAF6909017540">Cicchino, Armando C.</mods:namePart>
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</mods:name>
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<mods:name id="8927DE4DD169034ABB707E06315DD41D" type="personal">
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<mods:role id="9C42B8E5268A4580C011952C853914A3">
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<mods:roleTerm id="4255DBB5557F1646A58762FFB38FF88E">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="02BE891EE0FBD708C6AA5167EBB3F464">Roig- Juñent, Sergio</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="21A79BEF1226F77619BD22B28E11D19C">text</mods:typeOfResource>
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<mods:titleInfo id="4BBDC557BEE893D74B479FF6D396D46C">
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<mods:title id="9FF47EA6A255291E39195985AC9F79CA">The Coleopterists Bulletin</mods:title>
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</mods:titleInfo>
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<mods:part id="C1B9C8ADF32726A2DA198398885A5988">
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<mods:date id="5C2EB4E1A30592C1AE826407F7742EA7">2001</mods:date>
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<mods:detail id="BB4ACE948DB2C01907A54D3DA65346C9" type="pubDate">
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<mods:number id="3FD3312FED5C64DE9A43973FE7E79B6F">2001-06-01</mods:number>
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<mods:detail id="DBDC4B6C661C4CBFD4FB9BD9C35A4CF8" type="volume">
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<mods:number id="8F821924DB3159906CBA3F1D0E6D0C24">55</mods:number>
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<mods:detail id="53F1767B40D422082B4BBAB3E2600E24" type="issue">
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<mods:number id="E0F6F0A265A24159AE63F7B2C59FDE8E">2</mods:number>
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<mods:classification id="C0B6E843652443879A862A95D43E262E">journal article</mods:classification>
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<mods:identifier id="262A18DF03D6E9667405283A2F04F7D2" type="DOI">10.1649/0010-065X(2001)055[0185:DAROPN]2.0.CO;2</mods:identifier>
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<mods:identifier id="34E6B694603DD1CC89C5367A6F8D8397" type="ISSN">1938-4394</mods:identifier>
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<mods:identifier id="B11A68BFC4007BE9D10B7BD5F05DDBD1" type="Zenodo-Dep">10102726</mods:identifier>
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</mods:mods>
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<treatment id="5E5B1142FB7EFFAAE67B5163FE5BF95D" LSID="urn:lsid:plazi:treatment:5E5B1142FB7EFFAAE67B5163FE5BF95D" httpUri="http://treatment.plazi.org/id/5E5B1142FB7EFFAAE67B5163FE5BF95D" lastPageId="5" lastPageNumber="189" pageId="3" pageNumber="188">
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<subSubSection id="9EE8F3DFFB7EFFACE67B5163FCB6FE27" box="[572,846,270,292]" pageId="3" pageNumber="188" type="nomenclature">
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<paragraph id="D64DA054FB7EFFACE67B5163FCB6FE27" blockId="3.[572,846,270,292]" box="[572,846,270,292]" pageId="3" pageNumber="188">
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<heading id="8D051738FB7EFFACE67B5163FCB6FE27" box="[572,846,270,292]" centered="true" fontSize="9" level="2" pageId="3" pageNumber="188" reason="2">
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<taxonomicName id="11F2DBD7FB7EFFACE67B5163FD31FE20" ID-CoL="8KY8V" authorityName="Cicchino & Roig-Junient" authorityYear="2001" box="[572,713,271,292]" class="Insecta" family="Carabidae" genus="Paranillopsis" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="188" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="E4867C46FB7EFFACE67B5163FD31FE20" box="[572,713,271,292]" italics="true" pageId="3" pageNumber="188">Paranillopsis</emphasis>
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||
</taxonomicName>
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,
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||
<emphasis id="E4867C46FB7EFFACE69F5162FCB6FE27" bold="true" box="[728,846,270,291]" pageId="3" pageNumber="188">
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||
<taxonomicNameLabel id="FFB5C13DFB7EFFACE69F5162FCB6FE27" box="[728,846,270,291]" pageId="3" pageNumber="188" rank="genus">new genus</taxonomicNameLabel>
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</emphasis>
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||
</heading>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="9EE8F3DFFB7EFFACE5755154FE09FE9A" pageId="3" pageNumber="188" type="etymology">
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<paragraph id="D64DA054FB7EFFACE5755154FE09FE9A" blockId="3.[279,1112,312,547]" pageId="3" pageNumber="188">
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<emphasis id="E4867C46FB7EFFACE5755154FE49FE49" bold="true" box="[306,433,312,333]" pageId="3" pageNumber="188">Etymology.</emphasis>
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The name
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<taxonomicName id="11F2DBD7FB7EFFACE6755155FD47FE4A" authorityName="Cicchino & Roig-Junient" authorityYear="2001" box="[562,703,313,334]" class="Insecta" family="Carabidae" genus="Paranillopsis" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="188" phylum="Arthropoda" rank="genus">
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<emphasis id="E4867C46FB7EFFACE6755155FD47FE4A" box="[562,703,313,334]" italics="true" pageId="3" pageNumber="188">Paranillopsis</emphasis>
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</taxonomicName>
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is based on two classical words, the generic name
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<emphasis id="E4867C46FB7EFFACE5F5513FFDDFFE6C" box="[434,551,339,360]" italics="true" pageId="3" pageNumber="188">
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<taxonomicName id="11F2DBD7FB7EFFACE5F5513FFDDBFE6C" authorityName="Jeannel" authorityYear="1949" box="[434,547,339,360]" class="Insecta" family="Carabidae" genus="Paranillus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="188" phylum="Arthropoda" rank="genus">Paranillus</taxonomicName>
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,
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</emphasis>
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and the suffix
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<emphasis id="E4867C46FB7EFFACE694513FFCF7FE6C" box="[723,783,339,360]" italics="true" pageId="3" pageNumber="188">opsis,</emphasis>
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meaning similar to. Together, these words indicate that the new genus has features similar to, but not the same as,
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<emphasis id="E4867C46FB7EFFACE53C51E5FE09FE9A" box="[379,497,393,414]" italics="true" pageId="3" pageNumber="188">
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<taxonomicName id="11F2DBD7FB7EFFACE53C51E5FE15FE9A" authorityName="Jeannel" authorityYear="1949" box="[379,493,393,414]" class="Insecta" family="Carabidae" genus="Paranillus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="188" phylum="Arthropoda" rank="genus">Paranillus</taxonomicName>
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.
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</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection id="9EE8F3DFFB7EFFACE57551CFFBE8FEBC" box="[306,1040,419,440]" pageId="3" pageNumber="188" type="type_taxon">
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<paragraph id="D64DA054FB7EFFACE57551CFFBE8FEBC" blockId="3.[279,1112,312,547]" box="[306,1040,419,440]" pageId="3" pageNumber="188">
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||
<emphasis id="E4867C46FB7EFFACE57551CFFE3FFEBC" bold="true" box="[306,455,419,440]" pageId="3" pageNumber="188">
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<typeStatus id="09491EF6FB7EFFACE57551CFFE93FEBC" box="[306,363,419,440]" pageId="3" pageNumber="188">Type</typeStatus>
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species.
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</emphasis>
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<taxonomicName id="11F2DBD7FB7EFFACE58851CFFD30FEBC" ID-CoL="4D657" authorityName="Cicchino & Roig-Junent" authorityYear="2001" box="[463,712,419,440]" class="Insecta" family="Carabidae" genus="Paranillopsis" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="188" phylum="Arthropoda" rank="species" species="piguensis" status="sp. nov.">
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<emphasis id="E4867C46FB7EFFACE58851CFFD30FEBC" box="[463,712,419,440]" italics="true" pageId="3" pageNumber="188">Paranillopsis piguensis</emphasis>
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</taxonomicName>
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<emphasis id="E4867C46FB7EFFACE69751CFFCABFEBC" bold="true" box="[720,851,419,440]" pageId="3" pageNumber="188">
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<taxonomicNameLabel id="FFB5C13DFB7EFFACE69751CFFCABFEBC" box="[720,851,419,440]" pageId="3" pageNumber="188" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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, here designated.
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</paragraph>
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</subSubSection>
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<subSubSection id="9EE8F3DFFB7EFFACE57551D1FD09FD27" pageId="3" pageNumber="188" type="diagnosis">
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<paragraph id="D64DA054FB7EFFACE57551D1FD09FD27" blockId="3.[279,1112,312,547]" pageId="3" pageNumber="188">
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<emphasis id="E4867C46FB7EFFACE57551D1FE5BFED6" bold="true" box="[306,419,445,466]" pageId="3" pageNumber="188">Diagnosis.</emphasis>
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Body elongate (
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<figureCitation id="4EC9BCD1FB7EFFACE61051D2FD6EFED7" box="[599,662,446,467]" captionStart="Figs" captionStartId="2.[330,373,1526,1545]" captionTargetBox="[325,1112,263,1502]" captionTargetId="figure-10@2.[281,1156,212,1572]" captionTargetPageId="2" captionText="Figs. 1–10. Paranillopsis piguensis new species. 1) Habitus, dorsal aspect; 2) microsculture mesh pattern and pubescence; 3) antennomere 10 and 11; 4) labrum, ventral aspect, showing epipharynx; 5) labrum and clypeus, dorsal aspect; 6) labium, ventral aspect; 7) labium, apex of glossal sclerite; 8) right mandible, ventral aspect; 9) left mandible, ventral aspect; 10) maxillary palpomeres 3 and 4. Legend: cl, clypeus; e," figureDoi="http://doi.org/10.5281/zenodo.10102728" httpUri="https://zenodo.org/record/10102728/files/figure.png" pageId="3" pageNumber="188">Fig. 1</figureCitation>
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), depigmented, wingless; eye traces lacking; antenna long, extended to base of elytra; mentum tooth present; ligula with rounded paraglossae; elytral chaetotaxy of
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<typeStatus id="09491EF6FB7EFFACE76E519FFCAEFD0C" box="[809,854,499,520]" pageId="3" pageNumber="188">type</typeStatus>
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A (anillienée), without discal fixed setae, except the basal scutellar.
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</paragraph>
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</subSubSection>
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<subSubSection id="9EE8F3DFFB7EFFAAE568525AFE5BF95D" lastPageId="5" lastPageNumber="190" pageId="3" pageNumber="188" type="description">
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||
<paragraph id="D64DA054FB7EFFACE568525AFE0AFA7D" blockId="3.[279,1113,566,1401]" pageId="3" pageNumber="188">
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||
<emphasis id="E4867C46FB7EFFACE568525AFE5FFD4D" bold="true" box="[303,423,566,585]" pageId="3" pageNumber="188">Description.</emphasis>
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Length
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<quantity id="110A0DB1FB7EFFACE5BF525AFD7CFD4D" box="[504,644,566,585]" metricMagnitude="-3" metricUnit="m" metricValue="1.315" metricValueMax="1.57" metricValueMin="1.06" pageId="3" pageNumber="188" unit="mm" value="1.315" valueMax="1.57" valueMin="1.06">1.06–1.57 mm</quantity>
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. Head and elytra integument with microsculpture mesh pattern isodiametric (
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<figureCitation id="4EC9BCD1FB7EFFACE6505222FDA8FD65" box="[535,592,590,609]" captionStart="Figs" captionStartId="2.[330,373,1526,1545]" captionTargetBox="[325,1112,263,1502]" captionTargetId="figure-10@2.[281,1156,212,1572]" captionTargetPageId="2" captionText="Figs. 1–10. Paranillopsis piguensis new species. 1) Habitus, dorsal aspect; 2) microsculture mesh pattern and pubescence; 3) antennomere 10 and 11; 4) labrum, ventral aspect, showing epipharynx; 5) labrum and clypeus, dorsal aspect; 6) labium, ventral aspect; 7) labium, apex of glossal sclerite; 8) right mandible, ventral aspect; 9) left mandible, ventral aspect; 10) maxillary palpomeres 3 and 4. Legend: cl, clypeus; e," figureDoi="http://doi.org/10.5281/zenodo.10102728" httpUri="https://zenodo.org/record/10102728/files/figure.png" pageId="3" pageNumber="188">Fig. 2</figureCitation>
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), pronotum integument smooth; thick short pubescence present over the entire body. Uniform yellow brown color. Head almost same width as pronotum (
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<figureCitation id="4EC9BCD1FB7EFFACE5C75212FE4EFD95" box="[384,438,638,657]" captionStart="Figs" captionStartId="2.[330,373,1526,1545]" captionTargetBox="[325,1112,263,1502]" captionTargetId="figure-10@2.[281,1156,212,1572]" captionTargetPageId="2" captionText="Figs. 1–10. Paranillopsis piguensis new species. 1) Habitus, dorsal aspect; 2) microsculture mesh pattern and pubescence; 3) antennomere 10 and 11; 4) labrum, ventral aspect, showing epipharynx; 5) labrum and clypeus, dorsal aspect; 6) labium, ventral aspect; 7) labium, apex of glossal sclerite; 8) right mandible, ventral aspect; 9) left mandible, ventral aspect; 10) maxillary palpomeres 3 and 4. Legend: cl, clypeus; e," figureDoi="http://doi.org/10.5281/zenodo.10102728" httpUri="https://zenodo.org/record/10102728/files/figure.png" pageId="3" pageNumber="188">Fig. 1</figureCitation>
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), compressed or not in middle. Eyes absent. With four fixed setae, two supraorbital (
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<figureCitation id="4EC9BCD1FB7EFFACE5D252FAFE24FDAD" box="[405,476,662,681]" captionStart="Figs" captionStartId="5.[303,346,1256,1275]" captionTargetBox="[279,1102,265,1232]" captionTargetId="figure-12@5.[231,1159,215,1279]" captionTargetPageId="5" captionText="Figs. 16–18. Paranillopsis piguensis. 16) Head, dorsal aspect; 17) pronotum; 18) elytron, dorsal aspect. Figs. 19–21. Paranillopsis pampensis. 19) Head, dorsal aspect. 20) pronotum; 21) elytron, dorsal aspect. Figs. 22–23. Elytral base, dorsal aspect. 22) Paranillopsis piguensis; 23) P. pampensis. Figs. 24–25. Ovipositor, gonocoxites of Paranillopsis pampensis. 24) Ventral aspect; 25) dorsal aspect. Legend: bs, basal seta; bsp, basal setigerous puncture; cs, clypeal seta; es, ensiform seta; fs, frontal seta; lfc, lateral frontal carina; ns, nematiform seta. os, ocular seta; sos, supraorbital seta; us 1, 4, and 8, umbilical seta 1, 4,and 8; usp 1, umbilical setigerous puncture 1." figureDoi="http://doi.org/10.5281/zenodo.10102732" httpUri="https://zenodo.org/record/10102732/files/figure.png" pageId="3" pageNumber="188">Fig. 16</figureCitation>
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,
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<emphasis id="E4867C46FB7EFFACE5A052FAFDFDFDAD" bold="true" box="[487,517,662,681]" pageId="3" pageNumber="188">sos</emphasis>
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), one frontoparamedial (
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<figureCitation id="4EC9BCD1FB7EFFACE6BD52FAFCB9FDAD" box="[762,833,662,681]" captionStart="Figs" captionStartId="5.[303,346,1256,1275]" captionTargetBox="[279,1102,265,1232]" captionTargetId="figure-12@5.[231,1159,215,1279]" captionTargetPageId="5" captionText="Figs. 16–18. Paranillopsis piguensis. 16) Head, dorsal aspect; 17) pronotum; 18) elytron, dorsal aspect. Figs. 19–21. Paranillopsis pampensis. 19) Head, dorsal aspect. 20) pronotum; 21) elytron, dorsal aspect. Figs. 22–23. Elytral base, dorsal aspect. 22) Paranillopsis piguensis; 23) P. pampensis. Figs. 24–25. Ovipositor, gonocoxites of Paranillopsis pampensis. 24) Ventral aspect; 25) dorsal aspect. Legend: bs, basal seta; bsp, basal setigerous puncture; cs, clypeal seta; es, ensiform seta; fs, frontal seta; lfc, lateral frontal carina; ns, nematiform seta. os, ocular seta; sos, supraorbital seta; us 1, 4, and 8, umbilical seta 1, 4,and 8; usp 1, umbilical setigerous puncture 1." figureDoi="http://doi.org/10.5281/zenodo.10102732" httpUri="https://zenodo.org/record/10102732/files/figure.png" pageId="3" pageNumber="188">Fig. 16</figureCitation>
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,
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<emphasis id="E4867C46FB7EFFACE70A52FAFCA6FDAD" bold="true" box="[845,862,662,681]" pageId="3" pageNumber="188">fs</emphasis>
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), and one ocular (
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<figureCitation id="4EC9BCD1FB7EFFACE04952FAFBACFDAD" box="[1038,1108,662,681]" captionStart="Figs" captionStartId="5.[303,346,1256,1275]" captionTargetBox="[279,1102,265,1232]" captionTargetId="figure-12@5.[231,1159,215,1279]" captionTargetPageId="5" captionText="Figs. 16–18. Paranillopsis piguensis. 16) Head, dorsal aspect; 17) pronotum; 18) elytron, dorsal aspect. Figs. 19–21. Paranillopsis pampensis. 19) Head, dorsal aspect. 20) pronotum; 21) elytron, dorsal aspect. Figs. 22–23. Elytral base, dorsal aspect. 22) Paranillopsis piguensis; 23) P. pampensis. Figs. 24–25. Ovipositor, gonocoxites of Paranillopsis pampensis. 24) Ventral aspect; 25) dorsal aspect. Legend: bs, basal seta; bsp, basal setigerous puncture; cs, clypeal seta; es, ensiform seta; fs, frontal seta; lfc, lateral frontal carina; ns, nematiform seta. os, ocular seta; sos, supraorbital seta; us 1, 4, and 8, umbilical seta 1, 4,and 8; usp 1, umbilical setigerous puncture 1." figureDoi="http://doi.org/10.5281/zenodo.10102732" httpUri="https://zenodo.org/record/10102732/files/figure.png" pageId="3" pageNumber="188">Fig. 16</figureCitation>
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,
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<emphasis id="E4867C46FB7EFFACE55052C2FED4FDC5" bold="true" box="[279,300,686,705]" pageId="3" pageNumber="188">os</emphasis>
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||
). Lateral frontal carina clearly visible (Fig. 19,
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<emphasis id="E4867C46FB7EFFACE74852C2FCDFFDC5" bold="true" box="[783,807,686,705]" pageId="3" pageNumber="188">lfc</emphasis>
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). Antenna moniliform with 11 antennomeres, long, extended to base of elytra, with thick pubescence from the second antennomere, antennomere 11 excavated (
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<figureCitation id="4EC9BCD1FB7EFFACE6EA52B2FD11FDF5" box="[685,745,734,753]" captionStart="Figs" captionStartId="2.[330,373,1526,1545]" captionTargetBox="[325,1112,263,1502]" captionTargetId="figure-10@2.[281,1156,212,1572]" captionTargetPageId="2" captionText="Figs. 1–10. Paranillopsis piguensis new species. 1) Habitus, dorsal aspect; 2) microsculture mesh pattern and pubescence; 3) antennomere 10 and 11; 4) labrum, ventral aspect, showing epipharynx; 5) labrum and clypeus, dorsal aspect; 6) labium, ventral aspect; 7) labium, apex of glossal sclerite; 8) right mandible, ventral aspect; 9) left mandible, ventral aspect; 10) maxillary palpomeres 3 and 4. Legend: cl, clypeus; e," figureDoi="http://doi.org/10.5281/zenodo.10102728" httpUri="https://zenodo.org/record/10102728/files/figure.png" pageId="3" pageNumber="188">Fig. 3</figureCitation>
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,
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<emphasis id="E4867C46FB7EFFACE6B052B2FCF9FDF5" bold="true" box="[759,769,734,753]" pageId="3" pageNumber="188">e</emphasis>
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||
). Labrum (
|
||
<figureCitation id="4EC9BCD1FB7EFFACE73652B2FC55FDF5" box="[881,941,734,753]" captionStart="Figs" captionStartId="2.[330,373,1526,1545]" captionTargetBox="[325,1112,263,1502]" captionTargetId="figure-10@2.[281,1156,212,1572]" captionTargetPageId="2" captionText="Figs. 1–10. Paranillopsis piguensis new species. 1) Habitus, dorsal aspect; 2) microsculture mesh pattern and pubescence; 3) antennomere 10 and 11; 4) labrum, ventral aspect, showing epipharynx; 5) labrum and clypeus, dorsal aspect; 6) labium, ventral aspect; 7) labium, apex of glossal sclerite; 8) right mandible, ventral aspect; 9) left mandible, ventral aspect; 10) maxillary palpomeres 3 and 4. Legend: cl, clypeus; e," figureDoi="http://doi.org/10.5281/zenodo.10102728" httpUri="https://zenodo.org/record/10102728/files/figure.png" pageId="3" pageNumber="188">Fig. 5</figureCitation>
|
||
,
|
||
<emphasis id="E4867C46FB7EFFACE7FC52B2FC39FDF5" bold="true" box="[955,961,734,753]" pageId="3" pageNumber="188">l</emphasis>
|
||
) with six setae dorsally and concave anterior margin (
|
||
<figureCitation id="4EC9BCD1FB7EFFACE6D3529AFD2AFC0D" box="[660,722,758,777]" captionStart="Figs" captionStartId="2.[330,373,1526,1545]" captionTargetBox="[325,1112,263,1502]" captionTargetId="figure-10@2.[281,1156,212,1572]" captionTargetPageId="2" captionText="Figs. 1–10. Paranillopsis piguensis new species. 1) Habitus, dorsal aspect; 2) microsculture mesh pattern and pubescence; 3) antennomere 10 and 11; 4) labrum, ventral aspect, showing epipharynx; 5) labrum and clypeus, dorsal aspect; 6) labium, ventral aspect; 7) labium, apex of glossal sclerite; 8) right mandible, ventral aspect; 9) left mandible, ventral aspect; 10) maxillary palpomeres 3 and 4. Legend: cl, clypeus; e," figureDoi="http://doi.org/10.5281/zenodo.10102728" httpUri="https://zenodo.org/record/10102728/files/figure.png" pageId="3" pageNumber="188">Fig. 4</figureCitation>
|
||
); mandibles short with retinaculum and premolar tooth small and conical (
|
||
<figureCitation id="4EC9BCD1FB7EFFACE6195362FD3FFC25" box="[606,711,782,801]" captionStart="Figs" captionStartId="2.[330,373,1526,1545]" captionTargetBox="[325,1112,263,1502]" captionTargetId="figure-10@2.[281,1156,212,1572]" captionTargetPageId="2" captionText="Figs. 1–10. Paranillopsis piguensis new species. 1) Habitus, dorsal aspect; 2) microsculture mesh pattern and pubescence; 3) antennomere 10 and 11; 4) labrum, ventral aspect, showing epipharynx; 5) labrum and clypeus, dorsal aspect; 6) labium, ventral aspect; 7) labium, apex of glossal sclerite; 8) right mandible, ventral aspect; 9) left mandible, ventral aspect; 10) maxillary palpomeres 3 and 4. Legend: cl, clypeus; e," figureDoi="http://doi.org/10.5281/zenodo.10102728" httpUri="https://zenodo.org/record/10102728/files/figure.png" pageId="3" pageNumber="188">Figs. 9–10</figureCitation>
|
||
); maxillar palpomere 3 large (
|
||
<figureCitation id="4EC9BCD1FB7EFFACE7A05362FBD5FC25" box="[999,1069,782,801]" captionStart="Figs" captionStartId="2.[330,373,1526,1545]" captionTargetBox="[325,1112,263,1502]" captionTargetId="figure-10@2.[281,1156,212,1572]" captionTargetPageId="2" captionText="Figs. 1–10. Paranillopsis piguensis new species. 1) Habitus, dorsal aspect; 2) microsculture mesh pattern and pubescence; 3) antennomere 10 and 11; 4) labrum, ventral aspect, showing epipharynx; 5) labrum and clypeus, dorsal aspect; 6) labium, ventral aspect; 7) labium, apex of glossal sclerite; 8) right mandible, ventral aspect; 9) left mandible, ventral aspect; 10) maxillary palpomeres 3 and 4. Legend: cl, clypeus; e," figureDoi="http://doi.org/10.5281/zenodo.10102728" httpUri="https://zenodo.org/record/10102728/files/figure.png" pageId="3" pageNumber="188">Fig. 10</figureCitation>
|
||
,
|
||
<emphasis id="E4867C46FB7EFFACE0705362FEDBFC3D" bold="true" pageId="3" pageNumber="188">mp 3</emphasis>
|
||
) and palpomere 4 subconical (
|
||
<figureCitation id="4EC9BCD1FB7EFFACE616534AFD61FC3D" box="[593,665,806,825]" captionStart="Figs" captionStartId="2.[330,373,1526,1545]" captionTargetBox="[325,1112,263,1502]" captionTargetId="figure-10@2.[281,1156,212,1572]" captionTargetPageId="2" captionText="Figs. 1–10. Paranillopsis piguensis new species. 1) Habitus, dorsal aspect; 2) microsculture mesh pattern and pubescence; 3) antennomere 10 and 11; 4) labrum, ventral aspect, showing epipharynx; 5) labrum and clypeus, dorsal aspect; 6) labium, ventral aspect; 7) labium, apex of glossal sclerite; 8) right mandible, ventral aspect; 9) left mandible, ventral aspect; 10) maxillary palpomeres 3 and 4. Legend: cl, clypeus; e," figureDoi="http://doi.org/10.5281/zenodo.10102728" httpUri="https://zenodo.org/record/10102728/files/figure.png" pageId="3" pageNumber="188">Fig. 10</figureCitation>
|
||
,
|
||
<emphasis id="E4867C46FB7EFFACE6E1534AFD23FC3D" bold="true" box="[678,731,806,825]" pageId="3" pageNumber="188">mp 4</emphasis>
|
||
); labium with mentum (
|
||
<figureCitation id="4EC9BCD1FB7EFFACE78E534AFBFDFC3D" box="[969,1029,806,825]" captionStart="Figs" captionStartId="2.[330,373,1526,1545]" captionTargetBox="[325,1112,263,1502]" captionTargetId="figure-10@2.[281,1156,212,1572]" captionTargetPageId="2" captionText="Figs. 1–10. Paranillopsis piguensis new species. 1) Habitus, dorsal aspect; 2) microsculture mesh pattern and pubescence; 3) antennomere 10 and 11; 4) labrum, ventral aspect, showing epipharynx; 5) labrum and clypeus, dorsal aspect; 6) labium, ventral aspect; 7) labium, apex of glossal sclerite; 8) right mandible, ventral aspect; 9) left mandible, ventral aspect; 10) maxillary palpomeres 3 and 4. Legend: cl, clypeus; e," figureDoi="http://doi.org/10.5281/zenodo.10102728" httpUri="https://zenodo.org/record/10102728/files/figure.png" pageId="3" pageNumber="188">Fig. 6</figureCitation>
|
||
,
|
||
<emphasis id="E4867C46FB7EFFACE055534AFBDDFC3D" bold="true" box="[1042,1061,806,825]" pageId="3" pageNumber="188">m</emphasis>
|
||
) and submentum not fused; mentum with simple tooth (
|
||
<figureCitation id="4EC9BCD1FB7EFFACE6BE5352FCCCFC55" box="[761,820,830,849]" captionStart="Figs" captionStartId="2.[330,373,1526,1545]" captionTargetBox="[325,1112,263,1502]" captionTargetId="figure-10@2.[281,1156,212,1572]" captionTargetPageId="2" captionText="Figs. 1–10. Paranillopsis piguensis new species. 1) Habitus, dorsal aspect; 2) microsculture mesh pattern and pubescence; 3) antennomere 10 and 11; 4) labrum, ventral aspect, showing epipharynx; 5) labrum and clypeus, dorsal aspect; 6) labium, ventral aspect; 7) labium, apex of glossal sclerite; 8) right mandible, ventral aspect; 9) left mandible, ventral aspect; 10) maxillary palpomeres 3 and 4. Legend: cl, clypeus; e," figureDoi="http://doi.org/10.5281/zenodo.10102728" httpUri="https://zenodo.org/record/10102728/files/figure.png" pageId="3" pageNumber="188">Fig. 6</figureCitation>
|
||
,
|
||
<emphasis id="E4867C46FB7EFFACE7785352FCA2FC55" bold="true" box="[831,858,830,849]" pageId="3" pageNumber="188">mt</emphasis>
|
||
), two setae each in umbilicate punctures at base of median tooth and two lateral setae, epilobes not evident; submentum with four to eight setae; labial palpomere 2 (
|
||
<figureCitation id="4EC9BCD1FB7EFFACE7035302FC87FC85" box="[836,895,878,897]" captionStart="Figs" captionStartId="2.[330,373,1526,1545]" captionTargetBox="[325,1112,263,1502]" captionTargetId="figure-10@2.[281,1156,212,1572]" captionTargetPageId="2" captionText="Figs. 1–10. Paranillopsis piguensis new species. 1) Habitus, dorsal aspect; 2) microsculture mesh pattern and pubescence; 3) antennomere 10 and 11; 4) labrum, ventral aspect, showing epipharynx; 5) labrum and clypeus, dorsal aspect; 6) labium, ventral aspect; 7) labium, apex of glossal sclerite; 8) right mandible, ventral aspect; 9) left mandible, ventral aspect; 10) maxillary palpomeres 3 and 4. Legend: cl, clypeus; e," figureDoi="http://doi.org/10.5281/zenodo.10102728" httpUri="https://zenodo.org/record/10102728/files/figure.png" pageId="3" pageNumber="188">Fig. 6</figureCitation>
|
||
,
|
||
<emphasis id="E4867C46FB7EFFACE7CA5302FC4DFC85" bold="true" box="[909,949,878,897]" pageId="3" pageNumber="188">lp 2</emphasis>
|
||
) with few setae, palpomere 3 slender (
|
||
<figureCitation id="4EC9BCD1FB7EFFACE5B753EAFDD5FC9D" box="[496,557,902,921]" captionStart="Figs" captionStartId="2.[330,373,1526,1545]" captionTargetBox="[325,1112,263,1502]" captionTargetId="figure-10@2.[281,1156,212,1572]" captionTargetPageId="2" captionText="Figs. 1–10. Paranillopsis piguensis new species. 1) Habitus, dorsal aspect; 2) microsculture mesh pattern and pubescence; 3) antennomere 10 and 11; 4) labrum, ventral aspect, showing epipharynx; 5) labrum and clypeus, dorsal aspect; 6) labium, ventral aspect; 7) labium, apex of glossal sclerite; 8) right mandible, ventral aspect; 9) left mandible, ventral aspect; 10) maxillary palpomeres 3 and 4. Legend: cl, clypeus; e," figureDoi="http://doi.org/10.5281/zenodo.10102728" httpUri="https://zenodo.org/record/10102728/files/figure.png" pageId="3" pageNumber="188">Fig. 6</figureCitation>
|
||
,
|
||
<emphasis id="E4867C46FB7EFFACE67B53EAFD9EFC9D" bold="true" box="[572,614,902,921]" pageId="3" pageNumber="188">lp 3</emphasis>
|
||
), as long as palpomere 2; ligula with paraglossae broadly rounded apically (
|
||
<figureCitation id="4EC9BCD1FB7EFFACE65253F2FD6FFCB5" box="[533,663,926,945]" captionStart="Figs" captionStartId="2.[330,373,1526,1545]" captionTargetBox="[325,1112,263,1502]" captionTargetId="figure-10@2.[281,1156,212,1572]" captionTargetPageId="2" captionText="Figs. 1–10. Paranillopsis piguensis new species. 1) Habitus, dorsal aspect; 2) microsculture mesh pattern and pubescence; 3) antennomere 10 and 11; 4) labrum, ventral aspect, showing epipharynx; 5) labrum and clypeus, dorsal aspect; 6) labium, ventral aspect; 7) labium, apex of glossal sclerite; 8) right mandible, ventral aspect; 9) left mandible, ventral aspect; 10) maxillary palpomeres 3 and 4. Legend: cl, clypeus; e," figureDoi="http://doi.org/10.5281/zenodo.10102728" httpUri="https://zenodo.org/record/10102728/files/figure.png" pageId="3" pageNumber="188">Figs. 6 and 7</figureCitation>
|
||
,
|
||
<emphasis id="E4867C46FB7EFFACE6E353F2FD30FCB5" bold="true" box="[676,712,926,945]" pageId="3" pageNumber="188">pag</emphasis>
|
||
); glossal sclerite (
|
||
<figureCitation id="4EC9BCD1FB7EFFACE73C53F2FC05FCB5" box="[891,1021,926,945]" captionStart="Figs" captionStartId="2.[330,373,1526,1545]" captionTargetBox="[325,1112,263,1502]" captionTargetId="figure-10@2.[281,1156,212,1572]" captionTargetPageId="2" captionText="Figs. 1–10. Paranillopsis piguensis new species. 1) Habitus, dorsal aspect; 2) microsculture mesh pattern and pubescence; 3) antennomere 10 and 11; 4) labrum, ventral aspect, showing epipharynx; 5) labrum and clypeus, dorsal aspect; 6) labium, ventral aspect; 7) labium, apex of glossal sclerite; 8) right mandible, ventral aspect; 9) left mandible, ventral aspect; 10) maxillary palpomeres 3 and 4. Legend: cl, clypeus; e," figureDoi="http://doi.org/10.5281/zenodo.10102728" httpUri="https://zenodo.org/record/10102728/files/figure.png" pageId="3" pageNumber="188">Figs. 6 and 7</figureCitation>
|
||
,
|
||
<emphasis id="E4867C46FB7EFFACE04D53F2FBE7FCB5" bold="true" box="[1034,1055,926,945]" pageId="3" pageNumber="188">gs</emphasis>
|
||
) with two central subapical setae. Pronotum transverse, cordiform, maximum width anteriorly and lateral margins posteriorly convergent; with two setae on each side, one anterior and another posterior; posterior margin narrower than anterior (
|
||
<figureCitation id="4EC9BCD1FB7EFFACE70C538AFC7BFCFD" box="[843,899,998,1017]" captionStart="Figs" captionStartId="2.[330,373,1526,1545]" captionTargetBox="[325,1112,263,1502]" captionTargetId="figure-10@2.[281,1156,212,1572]" captionTargetPageId="2" captionText="Figs. 1–10. Paranillopsis piguensis new species. 1) Habitus, dorsal aspect; 2) microsculture mesh pattern and pubescence; 3) antennomere 10 and 11; 4) labrum, ventral aspect, showing epipharynx; 5) labrum and clypeus, dorsal aspect; 6) labium, ventral aspect; 7) labium, apex of glossal sclerite; 8) right mandible, ventral aspect; 9) left mandible, ventral aspect; 10) maxillary palpomeres 3 and 4. Legend: cl, clypeus; e," figureDoi="http://doi.org/10.5281/zenodo.10102728" httpUri="https://zenodo.org/record/10102728/files/figure.png" pageId="3" pageNumber="188">Fig. 1</figureCitation>
|
||
); posterior angles distinct, sharp, and protruded. Elytra flat, long (three ot four times longer than wide), not extended to apex of abdominal tergum VIII, with humeral margin rounded or straight; lateral margins regularly denticulate throughout to insertion of umbilicate seta 8, each denticle at apex with small setula, basal setae present; discal setae absent; umbilical series (
|
||
<figureCitation id="4EC9BCD1FB7EFFACE51A5432FE5AFB75" box="[349,418,1118,1137]" captionStart="Figs" captionStartId="2.[330,373,1526,1545]" captionTargetBox="[325,1112,263,1502]" captionTargetId="figure-10@2.[281,1156,212,1572]" captionTargetPageId="2" captionText="Figs. 1–10. Paranillopsis piguensis new species. 1) Habitus, dorsal aspect; 2) microsculture mesh pattern and pubescence; 3) antennomere 10 and 11; 4) labrum, ventral aspect, showing epipharynx; 5) labrum and clypeus, dorsal aspect; 6) labium, ventral aspect; 7) labium, apex of glossal sclerite; 8) right mandible, ventral aspect; 9) left mandible, ventral aspect; 10) maxillary palpomeres 3 and 4. Legend: cl, clypeus; e," figureDoi="http://doi.org/10.5281/zenodo.10102728" httpUri="https://zenodo.org/record/10102728/files/figure.png" pageId="3" pageNumber="188">Figs. 1</figureCitation>
|
||
,
|
||
<figureCitation id="4EC9BCD1FB7EFFACE5F55432FE33FB75" box="[434,459,1118,1137]" captionStart="Figs" captionStartId="5.[303,346,1256,1275]" captionTargetBox="[279,1102,265,1232]" captionTargetId="figure-12@5.[231,1159,215,1279]" captionTargetPageId="5" captionText="Figs. 16–18. Paranillopsis piguensis. 16) Head, dorsal aspect; 17) pronotum; 18) elytron, dorsal aspect. Figs. 19–21. Paranillopsis pampensis. 19) Head, dorsal aspect. 20) pronotum; 21) elytron, dorsal aspect. Figs. 22–23. Elytral base, dorsal aspect. 22) Paranillopsis piguensis; 23) P. pampensis. Figs. 24–25. Ovipositor, gonocoxites of Paranillopsis pampensis. 24) Ventral aspect; 25) dorsal aspect. Legend: bs, basal seta; bsp, basal setigerous puncture; cs, clypeal seta; es, ensiform seta; fs, frontal seta; lfc, lateral frontal carina; ns, nematiform seta. os, ocular seta; sos, supraorbital seta; us 1, 4, and 8, umbilical seta 1, 4,and 8; usp 1, umbilical setigerous puncture 1." figureDoi="http://doi.org/10.5281/zenodo.10102732" httpUri="https://zenodo.org/record/10102732/files/figure.png" pageId="3" pageNumber="188">18</figureCitation>
|
||
, and 21) of nine setae, each seta inserted in umbilicate setigerous puncture, in three groups: basal, with setae 1–4, medial, with setae 5 and 6; and apical, with setae 7–9; setae 2, 6, and 8 largest; setigerous punctures 1–3 equidistant in sequence; setigerous puncture 4 more isolated, four times distance from 3 as 3 from 2, and similar distance from 5; setigerous punctures 5 and 6 close together; setigerous puncture 7 displaced mediad, 8 and 9 close together in form of pair geminée (series
|
||
<typeStatus id="09491EF6FB7EFFACE7B054BAFBE7FBED" box="[1015,1055,1238,1257]" pageId="3" pageNumber="188">type</typeStatus>
|
||
A or anillienée of
|
||
<bibRefCitation id="B263DDA5FB7EFFACE5D55482FDE9FA05" author="Jeannel" box="[402,529,1262,1281]" pageId="3" pageNumber="188" pagination="33 - 204" refId="ref4069" refString="Jeannel, R. 1963. Monographie des Anillini, Bembidiides endoges. (Col. Trechinae). Memoires du Museum d'Histoire Naturelle 28 (2): 33 - 204." type="journal article" year="1963">Jeannel 1963</bibRefCitation>
|
||
). Recurrent groove present (Fig. 21,
|
||
<emphasis id="E4867C46FB7EFFACE72D5482FC78FA05" bold="true" box="[874,896,1262,1281]" pageId="3" pageNumber="188">rg</emphasis>
|
||
). Protibiae broad, with a large spur on dorsal surface (
|
||
<figureCitation id="4EC9BCD1FB7EFFACE67B556AFD7AFA1D" box="[572,642,1286,1305]" captionStart="Figs" captionStartId="4.[330,373,806,825]" captionTargetBox="[356,1082,264,784]" captionTargetId="figure-12@4.[349,1088,260,788]" captionTargetPageId="4" captionText="Figs. 11–15. Paranillopsis piguensis new species. 11) left protibia, dorsal aspect; 12) right protibia, ventral aspect; 13) sclerites of ovipositor, ventral aspect; 14–15) female genitalia, spermatheca. Legend: as, apical spur; ds, dorsal spur; es, ensiform seta; ns, nematiform seta; pas, preapical spur." figureDoi="http://doi.org/10.5281/zenodo.10102730" httpUri="https://zenodo.org/record/10102730/files/figure.png" pageId="3" pageNumber="188">Fig. 11</figureCitation>
|
||
,
|
||
<emphasis id="E4867C46FB7EFFACE6CA556AFD5BFA1D" bold="true" box="[653,675,1286,1305]" pageId="3" pageNumber="188">ds</emphasis>
|
||
). Apical spur displaced to the middle the tibia (
|
||
<figureCitation id="4EC9BCD1FB7EFFACE5595572FE69FA35" box="[286,401,1310,1329]" captionStart="Figs" captionStartId="4.[330,373,806,825]" captionTargetBox="[356,1082,264,784]" captionTargetId="figure-12@4.[349,1088,260,788]" captionTargetPageId="4" captionText="Figs. 11–15. Paranillopsis piguensis new species. 11) left protibia, dorsal aspect; 12) right protibia, ventral aspect; 13) sclerites of ovipositor, ventral aspect; 14–15) female genitalia, spermatheca. Legend: as, apical spur; ds, dorsal spur; es, ensiform seta; ns, nematiform seta; pas, preapical spur." figureDoi="http://doi.org/10.5281/zenodo.10102730" httpUri="https://zenodo.org/record/10102730/files/figure.png" pageId="3" pageNumber="188">Figs. 11–12</figureCitation>
|
||
,
|
||
<emphasis id="E4867C46FB7EFFACE5DA5572FE4AFA35" bold="true" box="[413,434,1310,1329]" pageId="3" pageNumber="188">as</emphasis>
|
||
). Abdominal sterna IV–VII with two paramedian setae; VIII with four. Gonocoxites wide, stylus arquate (
|
||
<figureCitation id="4EC9BCD1FB7EFFACE626555AFD5EFA4D" box="[609,678,1334,1353]" captionStart="Figs" captionStartId="4.[330,373,806,825]" captionTargetBox="[356,1082,264,784]" captionTargetId="figure-12@4.[349,1088,260,788]" captionTargetPageId="4" captionText="Figs. 11–15. Paranillopsis piguensis new species. 11) left protibia, dorsal aspect; 12) right protibia, ventral aspect; 13) sclerites of ovipositor, ventral aspect; 14–15) female genitalia, spermatheca. Legend: as, apical spur; ds, dorsal spur; es, ensiform seta; ns, nematiform seta; pas, preapical spur." figureDoi="http://doi.org/10.5281/zenodo.10102730" httpUri="https://zenodo.org/record/10102730/files/figure.png" pageId="3" pageNumber="188">Fig. 13</figureCitation>
|
||
) with subapical setose organ with two nematiform setae (
|
||
<figureCitation id="4EC9BCD1FB7EFFACE5D35522FE21FA65" box="[404,473,1358,1377]" captionStart="Figs" captionStartId="4.[330,373,806,825]" captionTargetBox="[356,1082,264,784]" captionTargetId="figure-12@4.[349,1088,260,788]" captionTargetPageId="4" captionText="Figs. 11–15. Paranillopsis piguensis new species. 11) left protibia, dorsal aspect; 12) right protibia, ventral aspect; 13) sclerites of ovipositor, ventral aspect; 14–15) female genitalia, spermatheca. Legend: as, apical spur; ds, dorsal spur; es, ensiform seta; ns, nematiform seta; pas, preapical spur." figureDoi="http://doi.org/10.5281/zenodo.10102730" httpUri="https://zenodo.org/record/10102730/files/figure.png" pageId="3" pageNumber="188">Fig. 13</figureCitation>
|
||
,
|
||
<emphasis id="E4867C46FB7EFFACE5A45522FE01FA65" bold="true" box="[483,505,1358,1377]" pageId="3" pageNumber="188">ns</emphasis>
|
||
) and two lateral strong ensiform setae (
|
||
<figureCitation id="4EC9BCD1FB7EFFACE7295522FC4BFA65" box="[878,947,1358,1377]" captionStart="Figs" captionStartId="4.[330,373,806,825]" captionTargetBox="[356,1082,264,784]" captionTargetId="figure-12@4.[349,1088,260,788]" captionTargetPageId="4" captionText="Figs. 11–15. Paranillopsis piguensis new species. 11) left protibia, dorsal aspect; 12) right protibia, ventral aspect; 13) sclerites of ovipositor, ventral aspect; 14–15) female genitalia, spermatheca. Legend: as, apical spur; ds, dorsal spur; es, ensiform seta; ns, nematiform seta; pas, preapical spur." figureDoi="http://doi.org/10.5281/zenodo.10102730" httpUri="https://zenodo.org/record/10102730/files/figure.png" pageId="3" pageNumber="188">Fig. 13</figureCitation>
|
||
,
|
||
<emphasis id="E4867C46FB7EFFACE7FA5522FC28FA65" bold="true" box="[957,976,1358,1377]" pageId="3" pageNumber="188">es</emphasis>
|
||
); spermatheca rounded (
|
||
<figureCitation id="4EC9BCD1FB7EFFACE535550AFE1FFA7D" box="[370,487,1382,1401]" captionStart="Figs" captionStartId="4.[330,373,806,825]" captionTargetBox="[356,1082,264,784]" captionTargetId="figure-12@4.[349,1088,260,788]" captionTargetPageId="4" captionText="Figs. 11–15. Paranillopsis piguensis new species. 11) left protibia, dorsal aspect; 12) right protibia, ventral aspect; 13) sclerites of ovipositor, ventral aspect; 14–15) female genitalia, spermatheca. Legend: as, apical spur; ds, dorsal spur; es, ensiform seta; ns, nematiform seta; pas, preapical spur." figureDoi="http://doi.org/10.5281/zenodo.10102730" httpUri="https://zenodo.org/record/10102730/files/figure.png" pageId="3" pageNumber="188">Figs. 14–15</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="D64DA054FB7EFFACE57555FDFBA0FAC5" blockId="3.[279,1112,1425,1473]" pageId="3" pageNumber="188">
|
||
<emphasis id="E4867C46FB7EFFACE57555FDFDD4FAA2" bold="true" box="[306,556,1425,1446]" pageId="3" pageNumber="188">Phyletic relationships.</emphasis>
|
||
<taxonomicName id="11F2DBD7FB7EFFACE67155FEFD3BFAA3" authorityName="Cicchino & Roig-Junient" authorityYear="2001" box="[566,707,1426,1447]" class="Insecta" family="Carabidae" genus="Paranillopsis" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="188" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="E4867C46FB7EFFACE67155FEFD3BFAA3" box="[566,707,1426,1447]" italics="true" pageId="3" pageNumber="188">Paranillopsis</emphasis>
|
||
</taxonomicName>
|
||
is included within the subtribe
|
||
<taxonomicName id="11F2DBD7FB7EFFACE06655FDFEC6FAC5" authorityName="Jeannel" authorityYear="1937" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="188" phylum="Arthropoda" rank="subTribe" subTribe="Anillina">Anillina</taxonomicName>
|
||
because it shares with other members of the subtribe the following char
|
||
</paragraph>
|
||
<paragraph id="D64DA054FB7EFFACE550565DFCF0F970" blockId="3.[279,1112,1584,1652]" pageId="3" pageNumber="188">
|
||
excavation;
|
||
<emphasis id="E4867C46FB7EFFACE5CE565CFE55F947" bold="true" box="[393,429,1584,1603]" pageId="3" pageNumber="188">epi,</emphasis>
|
||
epipharynx;
|
||
<emphasis id="E4867C46FB7EFFACE66C565CFDBDF947" bold="true" box="[555,581,1584,1603]" pageId="3" pageNumber="188">gs,</emphasis>
|
||
glossal sclerite;
|
||
<emphasis id="E4867C46FB7EFFACE6A3565CFD08F947" bold="true" box="[740,752,1584,1603]" pageId="3" pageNumber="188">l,</emphasis>
|
||
labrum;
|
||
<emphasis id="E4867C46FB7EFFACE701565CFC71F947" bold="true" box="[838,905,1584,1603]" pageId="3" pageNumber="188">lp 1, 2,</emphasis>
|
||
and
|
||
<emphasis id="E4867C46FB7EFFACE7FF565CFC32F947" bold="true" box="[952,970,1584,1603]" pageId="3" pageNumber="188">3,</emphasis>
|
||
lapial palpomeres 1–3;
|
||
<emphasis id="E4867C46FB7EFFACE5335624FE75F95F" bold="true" box="[372,397,1608,1627]" pageId="3" pageNumber="188">m,</emphasis>
|
||
mentum;
|
||
<emphasis id="E4867C46FB7EFFACE5AA5624FDD8F95F" bold="true" box="[493,544,1608,1627]" pageId="3" pageNumber="188">mp 3</emphasis>
|
||
and
|
||
<emphasis id="E4867C46FB7EFFACE6085624FD99F95F" bold="true" box="[591,609,1608,1627]" pageId="3" pageNumber="188">4,</emphasis>
|
||
maxillary palpomeres 3 and 4;
|
||
<emphasis id="E4867C46FB7EFFACE7C95624FC57F95F" bold="true" box="[910,943,1608,1627]" pageId="3" pageNumber="188">mt,</emphasis>
|
||
tooth of mentum;
|
||
<emphasis id="E4867C46FB7EFFACE550560CFEBAF977" bold="true" box="[279,322,1632,1651]" pageId="3" pageNumber="188">pag,</emphasis>
|
||
paraglossa;
|
||
<emphasis id="E4867C46FB7EFFACE5FE560CFE20F977" bold="true" box="[441,472,1632,1651]" pageId="3" pageNumber="188">pg,</emphasis>
|
||
palpiger;
|
||
<emphasis id="E4867C46FB7EFFACE67E560CFD7AF977" bold="true" box="[569,642,1632,1651]" pageId="3" pageNumber="188">T VIII,</emphasis>
|
||
Tergum VIII.
|
||
</paragraph>
|
||
<caption id="828DF0DCFB79FFABE50D534AFD6AFC85" ID-DOI="http://doi.org/10.5281/zenodo.10102730" ID-Zenodo-Dep="10102730" httpUri="https://zenodo.org/record/10102730/files/figure.png" pageId="4" pageNumber="189" startId="4.[330,373,806,825]" targetBox="[356,1082,264,784]" targetPageId="4">
|
||
<paragraph id="D64DA054FB79FFABE50D534AFD6AFC85" blockId="4.[307,1140,806,897]" pageId="4" pageNumber="189">
|
||
<emphasis id="E4867C46FB79FFABE50D534AFE3BFC3D" bold="true" box="[330,451,806,825]" pageId="4" pageNumber="189">Figs. 11–15.</emphasis>
|
||
<taxonomicName id="11F2DBD7FB79FFABE59D534AFD41FC3D" authorityName="Cicchino & Roig-Junent" authorityYear="2001" box="[474,697,806,825]" class="Insecta" family="Carabidae" genus="Paranillopsis" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="189" phylum="Arthropoda" rank="species" species="piguensis" status="sp. nov.">
|
||
<emphasis id="E4867C46FB79FFABE59D534AFD41FC3D" box="[474,697,806,825]" italics="true" pageId="4" pageNumber="189">Paranillopsis piguensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="FFB5C13DFB79FFABE684534AFCC1FC3D" box="[707,825,806,825]" pageId="4" pageNumber="189" rank="species">new species</taxonomicNameLabel>
|
||
.
|
||
<emphasis id="E4867C46FB79FFABE701534AFC9DFC3D" bold="true" box="[838,869,806,825]" pageId="4" pageNumber="189">11)</emphasis>
|
||
left protibia, dorsal aspect;
|
||
<emphasis id="E4867C46FB79FFABE5745352FEAAFC55" bold="true" box="[307,338,830,849]" pageId="4" pageNumber="189">12)</emphasis>
|
||
right protibia, ventral aspect;
|
||
<emphasis id="E4867C46FB79FFABE62D5352FD71FC55" bold="true" box="[618,649,830,849]" pageId="4" pageNumber="189">13)</emphasis>
|
||
sclerites of ovipositor, ventral aspect;
|
||
<emphasis id="E4867C46FB79FFABE7AA5352FBD7FC55" bold="true" box="[1005,1071,830,849]" pageId="4" pageNumber="189">14–15)</emphasis>
|
||
female genitalia, spermatheca. Legend:
|
||
<emphasis id="E4867C46FB79FFABE62C533AFD7DFC6D" bold="true" box="[619,645,854,873]" pageId="4" pageNumber="189">as,</emphasis>
|
||
apical spur;
|
||
<emphasis id="E4867C46FB79FFABE740533AFCDBFC6D" bold="true" box="[775,803,854,873]" pageId="4" pageNumber="189">ds,</emphasis>
|
||
dorsal spur;
|
||
<emphasis id="E4867C46FB79FFABE7E1533AFC47FC6D" bold="true" box="[934,959,854,873]" pageId="4" pageNumber="189">es,</emphasis>
|
||
ensiform seta;
|
||
<emphasis id="E4867C46FB79FFABE010533AFB8BFC6D" bold="true" box="[1111,1139,854,873]" pageId="4" pageNumber="189">ns,</emphasis>
|
||
nematiform seta;
|
||
<emphasis id="E4867C46FB79FFABE59E5302FDF9FC85" bold="true" box="[473,513,878,897]" pageId="4" pageNumber="189">pas,</emphasis>
|
||
preapical spur.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="D64DA054FB79FFAAE57453A8FE5BF95D" blockId="4.[307,1140,964,1625]" lastBlockId="5.[279,1112,1524,1625]" lastPageId="5" lastPageNumber="190" pageId="4" pageNumber="189">
|
||
acters: Eyes lacking and umbilical series of nine setae, and with the humeral group of four setigerous punctures, 1–3 close together near humerus, 4 distinctly posteriad 3. For drawing phylogenetic inferences within anilline genera there are two different character systems. The first hypothesis proposed by
|
||
<bibRefCitation id="B263DDA5FB79FFABE5745443FE2BFB40" author="Jeannel" box="[307,467,1071,1092]" pageId="4" pageNumber="189" pagination="241 - 396" refId="ref3984" refString="Jeannel, R. 1937. Les Bembidiides endoges (Col. Carabidae). Monographie d'une lignee gondwanienne. Revue Francaise d'Entomologie. 3 (4): 241 - 396." type="journal article" year="1937">Jeannel (1937)</bibRefCitation>
|
||
, and followed by others (
|
||
<bibRefCitation id="B263DDA5FB79FFABE6AD5443FC8BFB40" author="Jeanne" box="[746,883,1071,1092]" pageId="4" pageNumber="189" pagination="83 - 102" refId="ref3946" refString="Jeanne, C. 1973. Sur la signification des Bembidiides endoges de la Region Euromediterraneen (Col. Carabidae, Bembidiinae, Anillini). Nouvelle Revue d'Entomologie 3 (2): 83 - 102." type="journal article" year="1973">Jeanne 1973</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="B263DDA5FB79FFABE7C65443FB9FFB40" author="Vigna Taglianti" box="[897,1127,1071,1092]" pageId="4" pageNumber="189" pagination="163 - 172" refId="ref4392" refString="Vigna Taglianti, A. 1980. Nouvelles domees sur la systematique et la repartition geographique des coleopteres carabiques cavernicoles et endoges du Proche-Orient (Coleoptera, Carabidae). Memoires du Biospelologie 7: 163 - 172." type="journal article" year="1980">Vigna Taglianti 1980</bibRefCitation>
|
||
), is based on the disposition of setae of the umbilical series of elytra. The second one, based on the presence and absence of median mentum tooth, was also proposed by
|
||
<bibRefCitation id="B263DDA5FB79FFABE5FB5413FDA2FB90" author="Jeannel" box="[444,602,1151,1172]" pageId="4" pageNumber="189" pagination="33 - 204" refId="ref4069" refString="Jeannel, R. 1963. Monographie des Anillini, Bembidiides endoges. (Col. Trechinae). Memoires du Museum d'Histoire Naturelle 28 (2): 33 - 204." type="journal article" year="1963">Jeannel (1963)</bibRefCitation>
|
||
, and stated that the arrangement of anilline genera on the basis of
|
||
<specimenCount id="C0F46BDDFB79FFABE59854F5FDB0FBAA" box="[479,584,1177,1198]" pageId="4" pageNumber="189" type="generic" typeStatus="types">two types</specimenCount>
|
||
of umbilical series is inadequate to show relationships. Because the system used by Jeannel in 1963 takes into consideration more features than that used in 1937, we follow the second system to infer the phylogenetic relationships and systematic placement of
|
||
<emphasis id="E4867C46FB79FFABE7A75486FB8CFBFB" box="[992,1140,1258,1279]" italics="true" pageId="4" pageNumber="189">
|
||
<taxonomicName id="11F2DBD7FB79FFABE7A75486FB88FBFB" authorityName="Cicchino & Roig-Junient" authorityYear="2001" box="[992,1136,1258,1279]" class="Insecta" family="Carabidae" genus="Paranillopsis" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="189" phylum="Arthropoda" rank="genus">Paranillopsis</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
Within the subtribe
|
||
<taxonomicName id="11F2DBD7FB79FFABE6575568FD90FA1D" authorityName="Jeannel" authorityYear="1937" box="[528,616,1284,1305]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="189" phylum="Arthropoda" rank="subTribe" subTribe="Anillina">Anillina</taxonomicName>
|
||
,
|
||
<taxonomicName id="11F2DBD7FB79FFABE6305568FCFCFA1D" authorityName="Cicchino & Roig-Junient" authorityYear="2001" box="[631,772,1284,1305]" class="Insecta" family="Carabidae" genus="Paranillopsis" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="189" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="E4867C46FB79FFABE6305568FCFCFA1D" box="[631,772,1284,1305]" italics="true" pageId="4" pageNumber="189">Paranillopsis</emphasis>
|
||
</taxonomicName>
|
||
belongs to the division Phanerodonta (
|
||
<bibRefCitation id="B263DDA5FB79FFABE5395573FDF7FA30" author="Jeannel" box="[382,527,1311,1332]" pageId="4" pageNumber="189" pagination="33 - 204" refId="ref4069" refString="Jeannel, R. 1963. Monographie des Anillini, Bembidiides endoges. (Col. Trechinae). Memoires du Museum d'Histoire Naturelle 28 (2): 33 - 204." type="journal article" year="1963">Jeannel 1963</bibRefCitation>
|
||
), whose members have a mentum tooth, and within this group this genus is included in the phyletic series of
|
||
<emphasis id="E4867C46FB79FFABE73F5556FC15FA4B" box="[888,1005,1338,1359]" italics="true" pageId="4" pageNumber="189">
|
||
<taxonomicName id="11F2DBD7FB79FFABE73F5556FC11FA4B" authorityName="Jeannel" authorityYear="1949" box="[888,1001,1338,1359]" class="Insecta" family="Carabidae" genus="Paranillus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="189" phylum="Arthropoda" rank="genus">Paranillus</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
because the elytra have lost discal setae, and the umbilical series is
|
||
<typeStatus id="09491EF6FB79FFABE7E75538FC35FA6D" box="[928,973,1364,1385]" pageId="4" pageNumber="189">type</typeStatus>
|
||
A (anillienée). This phyletic line comprises only one genus,
|
||
<emphasis id="E4867C46FB79FFABE7695503FC5BFA80" box="[814,931,1391,1412]" italics="true" pageId="4" pageNumber="189">
|
||
<taxonomicName id="11F2DBD7FB79FFABE7695503FC67FA80" authorityName="Jeannel" authorityYear="1949" box="[814,927,1391,1412]" class="Insecta" family="Carabidae" genus="Paranillus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="189" phylum="Arthropoda" rank="genus">Paranillus</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
with eight species restricted to
|
||
<collectingCountry id="AEE5E0C4FB79FFABE5FD55E5FDC2FA9A" box="[442,570,1417,1438]" name="Madagascar" pageId="4" pageNumber="189">Madagascar</collectingCountry>
|
||
. The new genus differs from
|
||
<taxonomicName id="11F2DBD7FB79FFABE73C55E6FC11FA9B" authorityName="Jeannel" authorityYear="1949" box="[891,1001,1418,1439]" class="Insecta" family="Carabidae" genus="Paranillus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="189" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="E4867C46FB79FFABE73C55E6FC11FA9B" box="[891,1001,1418,1439]" italics="true" pageId="4" pageNumber="189">Paranillus</emphasis>
|
||
</taxonomicName>
|
||
in details of the ligula, mentum, and in elytral length. In
|
||
<emphasis id="E4867C46FB79FFABE75D55C8FC56FABD" box="[794,942,1444,1465]" italics="true" pageId="4" pageNumber="189">
|
||
<taxonomicName id="11F2DBD7FB79FFABE75D55C8FC51FABD" authorityName="Cicchino & Roig-Junient" authorityYear="2001" box="[794,937,1444,1465]" class="Insecta" family="Carabidae" genus="Paranillopsis" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="189" phylum="Arthropoda" rank="genus">Paranillopsis</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
with ligula seemingly bilobed apically, the paraglossae are separate from one another and extended beyond the apex of the glossal sclerite; in
|
||
<emphasis id="E4867C46FB79FFABE71155B6FC33FAEB" box="[854,971,1498,1519]" italics="true" pageId="4" pageNumber="189">
|
||
<taxonomicName id="11F2DBD7FB79FFABE71155B6FC3FFAEB" authorityName="Jeannel" authorityYear="1949" box="[854,967,1498,1519]" class="Insecta" family="Carabidae" genus="Paranillus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="189" phylum="Arthropoda" rank="genus">Paranillus</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
with the ligula with apical margin truncate, the paraglossae are absent. In
|
||
<taxonomicName id="11F2DBD7FB79FFABE7F35598FBB9F90D" authorityName="Cicchino & Roig-Junient" authorityYear="2001" box="[948,1089,1524,1545]" class="Insecta" family="Carabidae" genus="Paranillopsis" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="189" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="E4867C46FB79FFABE7F35598FBB9F90D" box="[948,1089,1524,1545]" italics="true" pageId="4" pageNumber="189">Paranillopsis</emphasis>
|
||
</taxonomicName>
|
||
epilobes are not evident; in
|
||
<taxonomicName id="11F2DBD7FB79FFABE6165663FD47F920" authorityName="Jeannel" authorityYear="1949" box="[593,703,1551,1572]" class="Insecta" family="Carabidae" genus="Paranillus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="189" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="E4867C46FB79FFABE6165663FD47F920" box="[593,703,1551,1572]" italics="true" pageId="4" pageNumber="189">Paranillus</emphasis>
|
||
</taxonomicName>
|
||
they are evident. In
|
||
<emphasis id="E4867C46FB79FFABE7F45663FBBFF920" box="[947,1095,1551,1572]" italics="true" pageId="4" pageNumber="189">
|
||
<taxonomicName id="11F2DBD7FB79FFABE7F45663FBBAF920" authorityName="Cicchino & Roig-Junient" authorityYear="2001" box="[947,1090,1551,1572]" class="Insecta" family="Carabidae" genus="Paranillopsis" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="189" phylum="Arthropoda" rank="genus">Paranillopsis</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
the elytra are not extended to abdominal tergum VIII; in
|
||
<emphasis id="E4867C46FB79FFABE7325646FC12F93B" box="[885,1002,1578,1599]" italics="true" pageId="4" pageNumber="189">
|
||
<taxonomicName id="11F2DBD7FB79FFABE7325646FC1EF93B" authorityName="Jeannel" authorityYear="1949" box="[885,998,1578,1599]" class="Insecta" family="Carabidae" genus="Paranillus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="189" phylum="Arthropoda" rank="genus">Paranillus</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
tergum VIII is covered by the elytra. Thus
|
||
<taxonomicName id="11F2DBD7FB79FFABE6D55628FCE7F95D" authorityName="Cicchino & Roig-Junient" authorityYear="2001" box="[658,799,1604,1625]" class="Insecta" family="Carabidae" genus="Paranillopsis" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="189" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="E4867C46FB79FFABE6D55628FCE7F95D" box="[658,799,1604,1625]" italics="true" pageId="4" pageNumber="189">Paranillopsis</emphasis>
|
||
</taxonomicName>
|
||
represents an additional clade (phyletic series of
|
||
<bibRefCitation id="B263DDA5FB78FFAAE5A15598FD8CF90D" author="Jeannel" box="[486,628,1524,1545]" pageId="5" pageNumber="190" pagination="33 - 204" refId="ref4069" refString="Jeannel, R. 1963. Monographie des Anillini, Bembidiides endoges. (Col. Trechinae). Memoires du Museum d'Histoire Naturelle 28 (2): 33 - 204." type="journal article" year="1963">Jeannel 1963</bibRefCitation>
|
||
) in South America, because the other Neotropical genera are grouped in different phyletic series (
|
||
<bibRefCitation id="B263DDA5FB78FFAAE7C05663FBE2F920" author="Jeannel" box="[903,1050,1551,1572]" pageId="5" pageNumber="190" pagination="33 - 204" refId="ref4069" refString="Jeannel, R. 1963. Monographie des Anillini, Bembidiides endoges. (Col. Trechinae). Memoires du Museum d'Histoire Naturelle 28 (2): 33 - 204." type="journal article" year="1963">Jeannel 1963</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="B263DDA5FB78FFAAE06C5663FE61F93A" author="Reichardt" pageId="5" pageNumber="190" refId="ref4186" refString="Reichardt, H. 1970. Un nouveau coleoptere carabique humicole et aveugle des ˆles i Galapagos [Pp. 165 - 169]. In: (Mission Zool. Belge aux ˆles i Galapagos et en Ecuador (N. Leleup and J. Leleup, editors). Volume 2." type="journal volume" year="1970">Reichardt 1970</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="B263DDA5FB78FFAAE5E05645FD74F93A" author="Vigna Taglianti" box="[423,652,1577,1598]" pageId="5" pageNumber="190" pagination="117 - 128" refId="ref4353" refString="Vigna Taglianti, A. 1973. The Anillini of Mexico and Guatemala (Coleoptera, Carabidae). Problemi attuali di Scienza e di Cultura. Quaderno, Accademia Nazionale dei Lincei 171: 117 - 128." type="journal article" year="1973">Vigna Taglianti 1973</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="B263DDA5FB78FFAAE6DD5645FCE5F93A" author="Mateu" box="[666,797,1577,1598]" pageId="5" pageNumber="190" pagination="201 - 207" refId="ref4152" refString="Mateu, J. 1980. Un nouveau Anillini tetramere, Gen. and sp. nov. recolte dans les grottes du Perou. Memoires du Biospelogie 7: 201 - 207." type="journal article" year="1980">Mateu 1980</bibRefCitation>
|
||
; Etonti and Mateu 1992;
|
||
<bibRefCitation id="B263DDA5FB78FFAAE0745645FE6FF95D" author="Zaballos" pageId="5" pageNumber="190" pagination="33 - 43" refId="ref4431" refString="Zaballos, J. P. 1997. Honduranillus balli: un nuevo genero y especie de Anillini (Coleoptera, Caraboidea) de Honduras. Acta Zoologica Mexicana (n. s.) 71: 33 - 43." type="journal article" year="1997">Zaballos 1997</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |