355 lines
48 KiB
XML
355 lines
48 KiB
XML
<document id="A4FEAD7A65114B91C975E8AFC331FEAE" ID-CLB-Dataset="22330" ID-DOI="10.11646/zootaxa.4772.1.5" ID-GBIF-Dataset="b14b0d0d-cf67-4895-965c-12792c622b21" ID-ISSN="1175-5326" ID-Zenodo-Dep="3814079" ID-ZooBank="B497198A-08CD-4A21-AE04-14390499853B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1588834845818" checkinUser="plazi" docAuthor="Polhemus, Dan A." docDate="2020" docId="03FE00115B73FFC3FF38F886FE6C0F62" docLanguage="en" docName="zootaxa.4772.1.5.pdf" docOrigin="Zootaxa 4772 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Enithares ziwa Polhemus 2020, new species" docType="treatment" docVersion="7" lastPageNumber="149" masterDocId="FFC778695B7DFFD2FFAFFFC8FFF90E21" masterDocTitle="Nine new species of Enithares (Heteroptera: Notonectidae) from New Guinea, with distributional notes on other species and an updated world checklist" masterLastPageNumber="182" masterPageNumber="132" pageNumber="146" updateTime="1698838866946" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="D4EBE547039B22E5109FBAA508A383A9">Nine new species of Enithares (Heteroptera: Notonectidae) from New Guinea, with distributional notes on other species and an updated world checklist</mods:title>
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<mods:name id="74A431A7D6B63547BEA82DD2210093F8" type="personal">
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<mods:namePart id="C1633BF4DD8A2DCC9C00B70B265A77E1">Polhemus, Dan A.</mods:namePart>
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<mods:title id="A206850D1F7E86BB0E5BBF83BAD769A0">Zootaxa</mods:title>
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<mods:part id="B33F1994205423B9FEC7872E1F39A517">
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<mods:date id="FF8C0C8F8BF83DB838D6AD5A4790CB49">2020</mods:date>
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<mods:number id="1FF5CCBA6F61E7A8EC055467CB4E4C09">2020-05-07</mods:number>
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<mods:number id="4EEA0748F37E7F638D941D7AE86D783C">4772</mods:number>
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<mods:identifier id="6AB41A22F510BE8150CC742B66D7A385" type="DOI">10.11646/zootaxa.4772.1.5</mods:identifier>
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<treatment id="03FE00115B73FFC3FF38F886FE6C0F62" ID-DOI="http://doi.org/10.5281/zenodo.3815470" ID-GBIF-Taxon="163975549" ID-Zenodo-Dep="3815470" LSID="urn:lsid:plazi:treatment:03FE00115B73FFC3FF38F886FE6C0F62" httpUri="http://treatment.plazi.org/id/03FE00115B73FFC3FF38F886FE6C0F62" lastPageId="17" lastPageNumber="149" pageId="14" pageNumber="146">
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<subSubSection id="C34DE28C5B73FFDCFF38F886FE1B0948" box="[151,482,1870,1897]" pageId="14" pageNumber="146" type="nomenclature">
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<paragraph id="8BE8B1075B73FFDCFF38F886FE1B0948" blockId="14.[151,482,1870,1934]" box="[151,482,1870,1897]" pageId="14" pageNumber="146">
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<heading id="D0A0066B5B73FFDCFF38F886FE1B0948" bold="true" box="[151,482,1870,1897]" fontSize="11" level="1" pageId="14" pageNumber="146" reason="1">
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<emphasis id="B9236D155B73FFDCFF38F886FE1B0948" bold="true" box="[151,482,1870,1897]" pageId="14" pageNumber="146">
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<taxonomicName id="4C57CA845B73FFDCFF38F886FEB50948" authority="Polhemus, 2020" authorityName="Polhemus" authorityYear="2020" box="[151,332,1870,1897]" class="Insecta" family="Notonectidae" genus="Enithares" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="146" phylum="Arthropoda" rank="species" species="ziwa" status="sp. nov.">
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<emphasis id="B9236D155B73FFDCFF38F886FEB50948" bold="true" box="[151,332,1870,1897]" italics="true" pageId="14" pageNumber="146">Enithares ziwa</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A210D06E5B73FFDCFEFCF887FE1B0948" box="[339,482,1871,1897]" pageId="14" pageNumber="146" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C34DE28C5B73FFDCFF38F8BBFEA109AF" box="[151,344,1907,1934]" pageId="14" pageNumber="146" type="description">
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<paragraph id="8BE8B1075B73FFDCFF38F8BBFEA109AF" blockId="14.[151,482,1870,1934]" box="[151,344,1907,1934]" pageId="14" pageNumber="146">
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(
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<figureCitation id="136CAD825B73FFDCFF30F8BBFF0409AC" box="[159,253,1907,1933]" captionStart="FIGURES 23–29" captionStartId="16.[151,264,1746,1771]" captionTargetBox="[199,1389,181,1721]" captionTargetId="figure@16.[199,1389,181,1721]" captionTargetPageId="16" captionText="FIGURES 23–29. Enithares species in the E. atra group, dorsal habitus photographs. 23–24. Enithares elongata Lansbury, male and female; specimens from Indonesia, Papua Province, Cyclops Mountains, CL 7145. 25–26. Enithares papua n. sp., male and female; specimens from Papua New Guinea, Southern Highlands Prov., male specimen from Swinging Bridge, CL 7020, female specimen from Tubage, CL 7022. 27. Enithares ziwa n. sp., male holotype, specimen from Indonesia, Papua Province, upper Ziwa River, CL 7100. 28–29. Enithares stylata Lansbury, male and female; specimens from Papua New Guinea, Western Highlands Prov., near Mt. Hagen, CL 1780." figureDoi="http://doi.org/10.5281/zenodo.3814093" httpUri="https://zenodo.org/record/3814093/files/figure.png" pageId="14" pageNumber="146">Figs. 27</figureCitation>
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,
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<figureCitation id="136CAD825B73FFDCFEA6F8BBFEDE09AC" box="[265,295,1907,1933]" captionStart="FIGURE 30" captionStartId="17.[151,250,1174,1199]" captionTargetBox="[158,1429,354,1139]" captionTargetId="figure@17.[151,1436,345,1149]" captionTargetPageId="17" captionText="FIGURE 30. Distribution of species in the E. atra group on New Guinea. Square = E. ziwa n. sp.; triangles = E. stylata Lansbury; pentagon = E. peninsularis n. sp." figureDoi="http://doi.org/10.5281/zenodo.3814095" httpUri="https://zenodo.org/record/3814095/files/figure.png" pageId="14" pageNumber="146">30</figureCitation>
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,
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<figureCitation id="136CAD825B73FFDCFE9CF8BBFEA909AF" box="[307,336,1907,1934]" captionStart="FIGURES 35–40" captionStartId="23.[151,264,1623,1648]" captionTargetBox="[170,1386,169,1547]" captionTargetId="figure@23.[151,1400,151,1568]" captionTargetPageId="23" captionText="FIGURES 35–40. Enithares species in the E. atra and E. bakeri groups, male genitalic complex, left lateral view. 35. Enithares elongata Lansbury; specimen from Papua New Guinea, Morobe Prov., Wareo. 36. Enithares ziwa n. sp.; specimen from Indonesia, Papua Prov., Ziwa River, CL 7100. 37. Enithares stylata Lansbury; specimen from Papua New Guinea, Morobe Prov., Wau, CL 1817. 38. Enithares tagula n. sp.; specimen from Papua New Guinea, Milne Bay Prov. Tagula Is., CL 7190. 39. Enithares bakeri Brooks, specimen from Indonesia, West Papua Prov., Aimasi River, CL 2651. 40. Enithares loria Brooks; specimen from Indonesia, Papua Prov., Siewa, CL 7085." figureDoi="http://doi.org/10.5281/zenodo.3814099" httpUri="https://zenodo.org/record/3814099/files/figure.png" pageId="14" pageNumber="146">36</figureCitation>
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)
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C34DE28C5B73FFDDFF38F87EFEA70EDA" lastPageId="15" lastPageNumber="147" pageId="14" pageNumber="146" type="materials_examined">
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||
<paragraph id="8BE8B1075B73FFDCFF38F87EFA6509D5" blockId="14.[151,1436,1973,2036]" pageId="14" pageNumber="146">
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||
<emphasis id="B9236D155B73FFDCFF38F87EFE4009F1" bold="true" box="[151,441,1973,2000]" pageId="14" pageNumber="146">Type material examined</emphasis>
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||
.
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<materialsCitation id="3B3FBB5A5B73FFDCFE67F87DFA6509D5" ID-GBIF-Occurrence="2610672615" country="Indonesia" county="New Guinea" elevation="1050" latitude="-3.1447222" location="upper Ziwa River" longLatPrecision="21" longitude="136.57361" municipality="Wapoga Alpha" pageId="14" pageNumber="146" specimenCount="1" stateProvince="Papua Barat" typeStatus="holotype">
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<typeStatus id="54EC0FA55B73FFDCFE67F87DFDCA09EE" box="[456,563,1973,1999]" pageId="14" pageNumber="146" type="holotype">Holotype</typeStatus>
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, male:
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<collectingCountry id="F340F1975B73FFDCFD2AF87DFCD909EE" box="[645,800,1973,1999]" name="Indonesia" pageId="14" pageNumber="146">INDONESIA</collectingCountry>
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,
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<collectingRegion id="49937FE55B73FFDCFC83F87DFC0909F1" box="[812,1008,1973,2000]" country="Indonesia" name="Papua Barat" pageId="14" pageNumber="146">
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<emphasis id="B9236D155B73FFDCFC83F87DFC1009F1" bold="true" box="[812,1001,1973,2000]" pageId="14" pageNumber="146">Irian Jaya Prov</emphasis>
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||
.
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||
</collectingRegion>
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||
[
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<collectingRegion id="49937FE55B73FFDCFC51F87EFB7409F1" box="[1022,1165,1973,2000]" country="Indonesia" name="Papua Barat" pageId="14" pageNumber="146">Papua Prov.</collectingRegion>
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],
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<collectingCounty id="6289C98B5B73FFDCFB0FF87EFAC909EE" box="[1184,1328,1973,2000]" pageId="14" pageNumber="146">New Guinea</collectingCounty>
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, pools on rainforest floor near PTFI
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<collectingMunicipality id="6B8C2B7D5B73FFDCFE16F812FDA409D2" box="[441,605,2009,2036]" pageId="14" pageNumber="146">Wapoga Alpha</collectingMunicipality>
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||
drilling camp,
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||
<location id="8E88E7DC5B73FFDCFCAAF812FC3109D2" LSID="urn:lsid:plazi:treatment:03FE00115B73FFC3FF38F886FE6C0F62:8E88E7DC5B73FFDCFCAAF812FC3109D2" box="[773,968,2009,2036]" country="Indonesia" county="New Guinea" latitude="-3.1447222" longLatPrecision="21" longitude="136.57361" municipality="Wapoga Alpha" name="upper Ziwa River" pageId="14" pageNumber="146" stateProvince="Papua Barat">
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||
upper
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||
<location id="8E88E7DC5B73FFDCFCE5F811FC3109D2" LSID="urn:lsid:plazi:treatment:03FE00115B73FFC3FF38F886FE6C0F62:8E88E7DC5B73FFDCFCE5F811FC3109D2" box="[842,968,2009,2035]" country="Indonesia" county="New Guinea" latitude="-3.1447222" longLatPrecision="21" longitude="136.57361" municipality="Wapoga Alpha" name="Ziwa River valley" pageId="14" pageNumber="146" stateProvince="Papua Barat">Ziwa River</location>
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</location>
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valley,
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<quantity id="4CAF1CE25B73FFDCFBB2F811FB8F09D5" box="[1053,1142,2009,2036]" metricMagnitude="3" metricUnit="m" metricValue="1.05" pageId="14" pageNumber="146" unit="m" value="1050.0">
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<elevation id="007A56345B73FFDCFBB2F811FB8F09D5" box="[1053,1142,2009,2036]" metricMagnitude="3" metricUnit="m" metricValue="1.05" pageId="14" pageNumber="146" unit="m" value="1050.0">1050 m</elevation>
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||
</quantity>
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||
.,
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||
<geoCoordinate id="EE63D7C05B73FFDCFB2AF811FB0309D5" box="[1157,1274,2009,2036]" degrees="3" direction="south" minutes="08" orientation="latitude" pageId="14" pageNumber="146" precision="15" seconds="41" value="-3.1447222">3°08′41′′S</geoCoordinate>
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,
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||
<geoCoordinate id="EE63D7C05B73FFDCFAAAF811FA6109D5" box="[1285,1432,2009,2036]" degrees="136" direction="east" minutes="34" orientation="longitude" pageId="14" pageNumber="146" precision="15" seconds="25" value="136.57361">136°34′25′′E</geoCoordinate>
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,
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</materialsCitation>
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||
</paragraph>
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||
<paragraph id="8BE8B1075B72FFDDFF38FF51FEA70EDA" blockId="15.[151,1437,153,2046]" pageId="15" pageNumber="147">
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<materialsCitation id="3B3FBB5A5B72FFDDFF38FF51FAD80E92" ID-GBIF-Occurrence="2610672571" box="[151,1313,153,179]" collectingDate="1998-04-18" collectingDateMax="1998-04-19" collectingDateMin="1998-04-18" collectionCode="A" pageId="15" pageNumber="147" specimenCount="1">
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water temp. 20° C.,
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<date id="FFE997C75B72FFDDFED1FF51FDE60E92" box="[382,543,153,179]" pageId="15" pageNumber="147" value="1998-04-18">
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||
<collectingDate id="EFAD6E2F5B72FFDDFED1FF51FDE60E92" box="[382,543,153,179]" pageId="15" pageNumber="147" value="1998-04-18">18 April 1998</collectingDate>
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</date>
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, 10:00–17:00 hrs.,
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<date id="FFE997C75B72FFDDFD55FF51FC620E92" box="[762,923,153,179]" pageId="15" pageNumber="147" value="1998-04-19">
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||
<collectingDate id="EFAD6E2F5B72FFDDFD55FF51FC620E92" box="[762,923,153,179]" pageId="15" pageNumber="147" value="1998-04-19">19 April 1998</collectingDate>
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</date>
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, 14:00–18:00 hrs., CL 7100, D.
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<collectionCode id="ED4629C25B72FFDDFAA5FF51FAD80E92" box="[1290,1313,153,179]" pageId="15" pageNumber="147">A</collectionCode>
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||
</materialsCitation>
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||
.
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<materialsCitation id="3B3FBB5A5B72FFDDFA81FF51FE820EF6" ID-GBIF-Occurrence="2610672466" collectionCode="USNM" location="Polhemus" pageId="15" pageNumber="147" specimenCount="1" typeStatus="paratype">
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||
<location id="8E88E7DC5B72FFDDFA81FF51FA650E92" LSID="urn:lsid:plazi:treatment:03FE00115B73FFC3FF38F886FE6C0F62:8E88E7DC5B72FFDDFA81FF51FA650E92" box="[1326,1436,153,179]" name="Polhemus" pageId="15" pageNumber="147">Polhemus</location>
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||
(
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||
<collectionCode id="ED4629C25B72FFDDFF30FF75FF0F0EF6" box="[159,246,189,215]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="15" pageNumber="147" type="Museum">USNM</collectionCode>
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||
).
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||
<emphasis id="B9236D155B72FFDDFEA6FF76FE820EF6" box="[265,379,190,215]" italics="true" pageId="15" pageNumber="147">
|
||
<typeStatus id="54EC0FA55B72FFDDFEA6FF76FE820EF6" box="[265,379,190,215]" pageId="15" pageNumber="147" type="paratype">Paratypes</typeStatus>
|
||
</emphasis>
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||
</materialsCitation>
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||
:
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||
<materialsCitation id="3B3FBB5A5B72FFDDFE26FF75FADA0EF6" ID-GBIF-Occurrence="2610672354" box="[393,1315,188,215]" collectionCode="A" country="Indonesia" county="New Guinea" elevation="1050" latitude="-3.1447222" location="upper Ziwa River" longLatPrecision="21" longitude="136.57361" municipality="Wapoga Alpha" pageId="15" pageNumber="147" specimenCount="4" specimenCount-female="4" stateProvince="Papua Barat" typeStatus="paratype">
|
||
<collectingCountry id="F340F1975B72FFDDFE26FF75FDDD0EF6" box="[393,548,189,215]" name="Indonesia" pageId="15" pageNumber="147">INDONESIA</collectingCountry>
|
||
,
|
||
<emphasis id="B9236D155B72FFDDFD80FF74FD090EF6" bold="true" box="[559,752,188,215]" pageId="15" pageNumber="147">
|
||
<collectingRegion id="49937FE55B72FFDDFD80FF74FD090EF6" box="[559,752,188,215]" country="Indonesia" name="Papua Barat" pageId="15" pageNumber="147">Irian Jaya Prov.</collectingRegion>
|
||
</emphasis>
|
||
:
|
||
<specimenCount id="9D517A8E5B72FFDDFD51FF75FC940EF6" box="[766,877,189,215]" pageId="15" pageNumber="147" type="female">4 females</specimenCount>
|
||
, same data as holotype, CL 7100, D.
|
||
<collectionCode id="ED4629C25B72FFDDFAA3FF75FADA0EF6" box="[1292,1315,189,215]" pageId="15" pageNumber="147">A</collectionCode>
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B3FBB5A5B72FFDDFA81FF75FEA30EDA" ID-GBIF-Occurrence="2610672407" collectionCode="USNM, BPBM" country="Indonesia" location="Polhemus" pageId="15" pageNumber="147" specimenCount="1" stateProvince="Papua Barat" typeStatus="paratype">
|
||
<location id="8E88E7DC5B72FFDDFA81FF75FA650EF6" LSID="urn:lsid:plazi:treatment:03FE00115B73FFC3FF38F886FE6C0F62:8E88E7DC5B72FFDDFA81FF75FA650EF6" box="[1326,1436,189,215]" country="Indonesia" name="Polhemus" pageId="15" pageNumber="147" stateProvince="Papua Barat">Polhemus</location>
|
||
(
|
||
<collectionCode id="ED4629C25B72FFDDFF30FF29FF0D0EDA" box="[159,244,225,251]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="15" pageNumber="147" type="Museum">USNM</collectionCode>
|
||
,
|
||
<collectionCode id="ED4629C25B72FFDDFF50FF29FEAB0EDA" box="[255,338,225,251]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" lsid="urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="15" pageNumber="147" type="Museum">BPBM</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34DE28C5B72FFDDFF68FECCFB3B0AC9" pageId="15" pageNumber="147" type="description">
|
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<paragraph id="8BE8B1075B72FFDDFF68FECCFC0D0F3E" blockId="15.[151,1437,153,2046]" box="[199,1012,260,287]" pageId="15" pageNumber="147">
|
||
<emphasis id="B9236D155B72FFDDFF68FECCFEA80F3E" bold="true" box="[199,337,260,287]" pageId="15" pageNumber="147">Description</emphasis>
|
||
.
|
||
<emphasis id="B9236D155B72FFDDFECFFECDFE640F3E" bold="true" box="[352,413,261,287]" pageId="15" pageNumber="147">Male</emphasis>
|
||
: Length
|
||
<quantity id="4CAF1CE25B72FFDDFDAEFECDFD8C0F3E" box="[513,629,261,287]" metricMagnitude="-2" metricUnit="m" metricValue="1.09" pageId="15" pageNumber="147" unit="mm" value="10.9">10.90 mm</quantity>
|
||
, width across pronotum
|
||
<quantity id="4CAF1CE25B72FFDDFC26FECDFC090F3E" box="[905,1008,261,287]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="15" pageNumber="147" unit="mm" value="4.2">4.20 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BE8B1075B72FFDDFF68FEE1FC9A0C1F" blockId="15.[151,1437,153,2046]" pageId="15" pageNumber="147">
|
||
<emphasis id="B9236D155B72FFDDFF68FEE1FEBB0F62" box="[199,322,297,323]" italics="true" pageId="15" pageNumber="147">Coloration</emphasis>
|
||
: Ground color black, with frons and vertex pale, limited pale markings on pronotum and hemelytra (
|
||
<figureCitation id="136CAD825B72FFDDFF0FFE85FF0D0F46" box="[160,244,333,359]" captionStart="FIGURES 23–29" captionStartId="16.[151,264,1746,1771]" captionTargetBox="[199,1389,181,1721]" captionTargetId="figure@16.[199,1389,181,1721]" captionTargetPageId="16" captionText="FIGURES 23–29. Enithares species in the E. atra group, dorsal habitus photographs. 23–24. Enithares elongata Lansbury, male and female; specimens from Indonesia, Papua Province, Cyclops Mountains, CL 7145. 25–26. Enithares papua n. sp., male and female; specimens from Papua New Guinea, Southern Highlands Prov., male specimen from Swinging Bridge, CL 7020, female specimen from Tubage, CL 7022. 27. Enithares ziwa n. sp., male holotype, specimen from Indonesia, Papua Province, upper Ziwa River, CL 7100. 28–29. Enithares stylata Lansbury, male and female; specimens from Papua New Guinea, Western Highlands Prov., near Mt. Hagen, CL 1780." figureDoi="http://doi.org/10.5281/zenodo.3814093" httpUri="https://zenodo.org/record/3814093/files/figure.png" pageId="15" pageNumber="147">Fig. 27</figureCitation>
|
||
). Vertex and frons dark yellow, with a pair (1+1) of irregular dark brown patches on frons at apex of head; 3 irregular dark brown patches present on lower frons, running transversely above base of labrum. Pronotum black, with incipient pale creamy white fascia across transverse midline. Scutellum uniformly shining black. Hemelytra black, with limited creamy white markings along embolar suture; wing membrane dark fumate brown. Legs yellowish-brown dorsally, medium brown ventrally, with anterior faces of fore and middle femora weakly suffused with reddish brown; posterior femur narrowly margined with black along anterior face; ventral surfaces of all trochanters and femora sparingly marked with darker brown. Venter brown.
|
||
</paragraph>
|
||
<paragraph id="8BE8B1075B72FFDDFF68FD80FDD90D37" blockId="15.[151,1437,153,2046]" pageId="15" pageNumber="147">
|
||
<emphasis id="B9236D155B72FFDDFF68FD80FE4E0C43" box="[199,439,584,610]" italics="true" pageId="15" pageNumber="147">Structural characters</emphasis>
|
||
: Head broadly rounded anteriorly when viewed dorsally. Head length 1.10; greatest width 3.40, equal to 0.81 pronotal width; anterior width of vertex 1.30, equal to 1.18 head length. Synthlipsis 0.75, about 0.58 anterior width of vertex and clearly shorter than pronotum. Pronotal length along midline 1.30, humeral width 4.20, lateral margins convex, posterior margin weakly sinuate. Dorsal margin of pronotal fovea directed caudad behind eyes (
|
||
<figureCitation id="136CAD825B72FFDDFEBDFD10FE9E0CD3" box="[274,359,728,754]" captionStart="FIGURES 23–29" captionStartId="16.[151,264,1746,1771]" captionTargetBox="[199,1389,181,1721]" captionTargetId="figure@16.[199,1389,181,1721]" captionTargetPageId="16" captionText="FIGURES 23–29. Enithares species in the E. atra group, dorsal habitus photographs. 23–24. Enithares elongata Lansbury, male and female; specimens from Indonesia, Papua Province, Cyclops Mountains, CL 7145. 25–26. Enithares papua n. sp., male and female; specimens from Papua New Guinea, Southern Highlands Prov., male specimen from Swinging Bridge, CL 7020, female specimen from Tubage, CL 7022. 27. Enithares ziwa n. sp., male holotype, specimen from Indonesia, Papua Province, upper Ziwa River, CL 7100. 28–29. Enithares stylata Lansbury, male and female; specimens from Papua New Guinea, Western Highlands Prov., near Mt. Hagen, CL 1780." figureDoi="http://doi.org/10.5281/zenodo.3814093" httpUri="https://zenodo.org/record/3814093/files/figure.png" pageId="15" pageNumber="147">Fig. 27</figureCitation>
|
||
). Nodal furrow nearly straight, removed by 2.3× its length from membranal suture, length 0.60, distance to membranal suture 1.40.
|
||
</paragraph>
|
||
<paragraph id="8BE8B1075B72FFDDFF68FCE8FE680DE8" blockId="15.[151,1437,153,2046]" pageId="15" pageNumber="147">Fore and hind legs typical for genus, lacking unusual modifications. Middle trochanter rounded. Middle femur with single moderately large, elongate, sharp subapical tooth, bordered basally by about 16 small black pegs, distally by about 3 similar pegs. Lengths of leg segments as follows: fore femur–tibia–tarsal 1–tarsal 2 = 1.70/1.70/0.70/0.45; middle femur–tibia–tarsal 1–tarsal 2 = 2.50/2.20/0.90/0.50; hind femur–tibia–tarsal 1–tarsal 2 = 4.30/3.60/1.80/0.90.</paragraph>
|
||
<paragraph id="8BE8B1075B72FFDDFF68FC1BFBCB0DCC" blockId="15.[151,1437,153,2046]" box="[199,1074,979,1005]" pageId="15" pageNumber="147">Ventral abdomen with metaxyphus triangular, slightly concave, tip acuminate.</paragraph>
|
||
<paragraph id="8BE8B1075B72FFDDFF68FC3FFBD30A81" blockId="15.[151,1437,153,2046]" pageId="15" pageNumber="147">
|
||
<emphasis id="B9236D155B72FFDDFF68FC3FFE960A30" box="[199,367,1015,1041]" italics="true" pageId="15" pageNumber="147">Male genitalia</emphasis>
|
||
when viewed laterally (
|
||
<figureCitation id="136CAD825B72FFDDFD26FC3FFD270A33" box="[649,734,1015,1042]" captionStart="FIGURES 35–40" captionStartId="23.[151,264,1623,1648]" captionTargetBox="[170,1386,169,1547]" captionTargetId="figure@23.[151,1400,151,1568]" captionTargetPageId="23" captionText="FIGURES 35–40. Enithares species in the E. atra and E. bakeri groups, male genitalic complex, left lateral view. 35. Enithares elongata Lansbury; specimen from Papua New Guinea, Morobe Prov., Wareo. 36. Enithares ziwa n. sp.; specimen from Indonesia, Papua Prov., Ziwa River, CL 7100. 37. Enithares stylata Lansbury; specimen from Papua New Guinea, Morobe Prov., Wau, CL 1817. 38. Enithares tagula n. sp.; specimen from Papua New Guinea, Milne Bay Prov. Tagula Is., CL 7190. 39. Enithares bakeri Brooks, specimen from Indonesia, West Papua Prov., Aimasi River, CL 2651. 40. Enithares loria Brooks; specimen from Indonesia, Papua Prov., Siewa, CL 7085." figureDoi="http://doi.org/10.5281/zenodo.3814099" httpUri="https://zenodo.org/record/3814099/files/figure.png" pageId="15" pageNumber="147">Fig. 36</figureCitation>
|
||
) with posterior lobe of pygophore with posteroventral angle sharp, bearing a prominent tapering setal tuft, posterior margin strongly bowed posteriorly and bearing long setae, apex truncate, slightly notched anteriorly on dorsal margin; paramere elongate, slightly tapering, apex rounded, exceeding dorsal margin of posterior lobe; lateral arm of basal plate sinuate, basal half stout, distal half strongly tapering, apex acuminate; aedeagus semicircular, with numerous transverse folds.
|
||
</paragraph>
|
||
<paragraph id="8BE8B1075B72FFDDFF68FB62FB3B0AC9" blockId="15.[151,1437,153,2046]" pageId="15" pageNumber="147">
|
||
<emphasis id="B9236D155B72FFDDFF68FB62FEE70AE5" bold="true" box="[199,286,1194,1220]" pageId="15" pageNumber="147">Female</emphasis>
|
||
: Length 10.50, width across pronotum
|
||
<quantity id="4CAF1CE25B72FFDDFD7EFB62FCCE0AE4" box="[721,823,1194,1221]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" pageId="15" pageNumber="147" unit="mm" value="4.1">4.10 mm</quantity>
|
||
. Similar to male in general structure and coloration, but with coloration paler, ground color medium brown, scutellum entirely creamy yellowish-white.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34DE28C5B72FFDDFF68FB3AFB010B2D" box="[199,1272,1266,1292]" pageId="15" pageNumber="147" type="etymology">
|
||
<paragraph id="8BE8B1075B72FFDDFF68FB3AFB010B2D" blockId="15.[151,1437,153,2046]" box="[199,1272,1266,1292]" pageId="15" pageNumber="147">
|
||
<emphasis id="B9236D155B72FFDDFF68FB3AFEB10B2D" bold="true" box="[199,328,1266,1292]" pageId="15" pageNumber="147">Etymology</emphasis>
|
||
. The name “ziwa” is a noun in apposition and refers to the Ziwa River
|
||
<typeStatus id="54EC0FA55B72FFDDFBC8FB3AFB6E0B2D" box="[1127,1175,1266,1292]" pageId="15" pageNumber="147">type</typeStatus>
|
||
locality.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34DE28C5B72FFC3FF68FADDFE6C0F62" lastPageId="17" lastPageNumber="149" pageId="15" pageNumber="147" type="discussion">
|
||
<paragraph id="8BE8B1075B72FFDDFF68FADDFEA6080A" blockId="15.[151,1437,153,2046]" pageId="15" pageNumber="147">
|
||
<emphasis id="B9236D155B72FFDDFF68FADDFEBC0B11" bold="true" box="[199,325,1301,1328]" pageId="15" pageNumber="147">Discussion</emphasis>
|
||
. In regard to its male genital structures,
|
||
<taxonomicName id="4C57CA845B72FFDDFC8CFADEFC350B11" authorityName="Polhemus" authorityYear="2020" box="[803,972,1302,1328]" class="Insecta" family="Notonectidae" genus="Enithares" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="147" phylum="Arthropoda" rank="species" species="ziwa">
|
||
<emphasis id="B9236D155B72FFDDFC8CFADEFC350B11" box="[803,972,1302,1328]" italics="true" pageId="15" pageNumber="147">Enithares ziwa</emphasis>
|
||
</taxonomicName>
|
||
is similar to
|
||
<taxonomicName id="4C57CA845B72FFDDFBC3FADFFB200B11" authorityName="Lansbury" authorityYear="1968" box="[1132,1241,1302,1328]" class="Insecta" family="Notonectidae" genus="Enithares" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="147" phylum="Arthropoda" rank="species" species="stylata">
|
||
<emphasis id="B9236D155B72FFDDFBC3FADFFB200B11" box="[1132,1241,1302,1328]" italics="true" pageId="15" pageNumber="147">E. stylata</emphasis>
|
||
</taxonomicName>
|
||
from the central mountains of
|
||
<collectingCountry id="F340F1975B72FFDDFE9BFAF2FDEB0B72" box="[308,530,1337,1364]" name="Papua New Guinea" pageId="15" pageNumber="147">Papua New Guinea</collectingCountry>
|
||
, and the paramere shapes are similar in the two species. However, the posterior lobe of the pygophore in
|
||
<taxonomicName id="4C57CA845B72FFDDFE12FA96FDE80B56" authorityName="Polhemus" authorityYear="2020" box="[445,529,1374,1399]" class="Insecta" family="Notonectidae" genus="Enithares" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="147" phylum="Arthropoda" rank="species" species="ziwa">
|
||
<emphasis id="B9236D155B72FFDDFE12FA96FDE80B56" box="[445,529,1374,1399]" italics="true" pageId="15" pageNumber="147">E. ziwa</emphasis>
|
||
</taxonomicName>
|
||
has the posteroventral angle far more sharp and pronounced, and the posterior margin of the PL more prominently bowed (compare
|
||
<figureCitation id="136CAD825B72FFDDFD5BFA49FC870BBA" box="[756,894,1409,1435]" captionStart="FIGURES 35–40" captionStartId="23.[151,264,1623,1648]" captionTargetBox="[170,1386,169,1547]" captionTargetId="figure@23.[151,1400,151,1568]" captionTargetPageId="23" captionText="FIGURES 35–40. Enithares species in the E. atra and E. bakeri groups, male genitalic complex, left lateral view. 35. Enithares elongata Lansbury; specimen from Papua New Guinea, Morobe Prov., Wareo. 36. Enithares ziwa n. sp.; specimen from Indonesia, Papua Prov., Ziwa River, CL 7100. 37. Enithares stylata Lansbury; specimen from Papua New Guinea, Morobe Prov., Wau, CL 1817. 38. Enithares tagula n. sp.; specimen from Papua New Guinea, Milne Bay Prov. Tagula Is., CL 7190. 39. Enithares bakeri Brooks, specimen from Indonesia, West Papua Prov., Aimasi River, CL 2651. 40. Enithares loria Brooks; specimen from Indonesia, Papua Prov., Siewa, CL 7085." figureDoi="http://doi.org/10.5281/zenodo.3814099" httpUri="https://zenodo.org/record/3814099/files/figure.png" pageId="15" pageNumber="147">Figs. 36, 37</figureCitation>
|
||
). The LABP in both species is strongly tapering on the distal half, but in
|
||
<taxonomicName id="4C57CA845B72FFDDFE08FA6EFDE80B9E" authorityName="Lansbury" authorityYear="1968" box="[423,529,1445,1471]" class="Insecta" family="Notonectidae" genus="Enithares" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="147" phylum="Arthropoda" rank="species" species="stylata">
|
||
<emphasis id="B9236D155B72FFDDFE08FA6EFDE80B9E" box="[423,529,1445,1471]" italics="true" pageId="15" pageNumber="147">E. stylata</emphasis>
|
||
</taxonomicName>
|
||
it comes to a slightly hooked apex, whereas in
|
||
<taxonomicName id="4C57CA845B72FFDDFB8DFA6EFB8D0B9E" authorityName="Polhemus" authorityYear="2020" box="[1058,1140,1446,1471]" class="Insecta" family="Notonectidae" genus="Enithares" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="147" phylum="Arthropoda" rank="species" species="ziwa">
|
||
<emphasis id="B9236D155B72FFDDFB8DFA6EFB8D0B9E" box="[1058,1140,1446,1471]" italics="true" pageId="15" pageNumber="147">E. ziwa</emphasis>
|
||
</taxonomicName>
|
||
the apex is sharp and acuminate (compare
|
||
<figureCitation id="136CAD825B72FFDDFEF3FA01FE1C0BC2" box="[348,485,1481,1507]" captionStart="FIGURES 35–40" captionStartId="23.[151,264,1623,1648]" captionTargetBox="[170,1386,169,1547]" captionTargetId="figure@23.[151,1400,151,1568]" captionTargetPageId="23" captionText="FIGURES 35–40. Enithares species in the E. atra and E. bakeri groups, male genitalic complex, left lateral view. 35. Enithares elongata Lansbury; specimen from Papua New Guinea, Morobe Prov., Wareo. 36. Enithares ziwa n. sp.; specimen from Indonesia, Papua Prov., Ziwa River, CL 7100. 37. Enithares stylata Lansbury; specimen from Papua New Guinea, Morobe Prov., Wau, CL 1817. 38. Enithares tagula n. sp.; specimen from Papua New Guinea, Milne Bay Prov. Tagula Is., CL 7190. 39. Enithares bakeri Brooks, specimen from Indonesia, West Papua Prov., Aimasi River, CL 2651. 40. Enithares loria Brooks; specimen from Indonesia, Papua Prov., Siewa, CL 7085." figureDoi="http://doi.org/10.5281/zenodo.3814099" httpUri="https://zenodo.org/record/3814099/files/figure.png" pageId="15" pageNumber="147">Figs. 36, 37</figureCitation>
|
||
). The two species may also be segregated on the basis of size, with
|
||
<taxonomicName id="4C57CA845B72FFDDFB70FA02FAB00BC2" authorityName="Lansbury" authorityYear="1968" box="[1247,1353,1481,1507]" class="Insecta" family="Notonectidae" genus="Enithares" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="147" phylum="Arthropoda" rank="species" species="stylata">
|
||
<emphasis id="B9236D155B72FFDDFB70FA02FAB00BC2" box="[1247,1353,1481,1507]" italics="true" pageId="15" pageNumber="147">E. stylata</emphasis>
|
||
</taxonomicName>
|
||
having body lengths ranging from 11.75–13.00 mm, whereas all examples of
|
||
<taxonomicName id="4C57CA845B72FFDDFC1AFA26FBF00826" authorityName="Polhemus" authorityYear="2020" box="[949,1033,1518,1543]" class="Insecta" family="Notonectidae" genus="Enithares" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="147" phylum="Arthropoda" rank="species" species="ziwa">
|
||
<emphasis id="B9236D155B72FFDDFC1AFA26FBF00826" box="[949,1033,1518,1543]" italics="true" pageId="15" pageNumber="147">E. ziwa</emphasis>
|
||
</taxonomicName>
|
||
so far known have body lengths of
|
||
<quantity id="4CAF1CE25B72FFDDFF38F9D9FEF0080A" box="[151,265,1553,1579]" metricMagnitude="-2" metricUnit="m" metricValue="1.09" pageId="15" pageNumber="147" unit="mm" value="10.9">10.90 mm</quantity>
|
||
or less.
|
||
</paragraph>
|
||
<paragraph id="8BE8B1075B72FFDDFF68F9FDFDFD096B" blockId="15.[151,1437,153,2046]" pageId="15" pageNumber="147">
|
||
<taxonomicName id="4C57CA845B72FFDDFF68F9FDFE96086F" authorityName="Polhemus" authorityYear="2020" box="[199,367,1589,1615]" class="Insecta" family="Notonectidae" genus="Enithares" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="147" phylum="Arthropoda" rank="species" species="ziwa">
|
||
<emphasis id="B9236D155B72FFDDFF68F9FDFE96086F" box="[199,367,1589,1615]" italics="true" pageId="15" pageNumber="147">Enithares ziwa</emphasis>
|
||
</taxonomicName>
|
||
is currently known from a single series taken in the upper Wapoga River basin, draining the north slopes of the central mountains in western New
|
||
<collectingCountry id="F340F1975B72FFDDFCA3F990FCA40853" box="[780,861,1624,1650]" name="Guinea-Bissau" pageId="15" pageNumber="147">Guinea</collectingCountry>
|
||
(
|
||
<figureCitation id="136CAD825B72FFDDFCC0F990FC3F0852" box="[879,966,1624,1651]" captionStart="FIGURE 30" captionStartId="17.[151,250,1174,1199]" captionTargetBox="[158,1429,354,1139]" captionTargetId="figure@17.[151,1436,345,1149]" captionTargetPageId="17" captionText="FIGURE 30. Distribution of species in the E. atra group on New Guinea. Square = E. ziwa n. sp.; triangles = E. stylata Lansbury; pentagon = E. peninsularis n. sp." figureDoi="http://doi.org/10.5281/zenodo.3814095" httpUri="https://zenodo.org/record/3814095/files/figure.png" pageId="15" pageNumber="147">Fig. 30</figureCitation>
|
||
). It will likely prove to be more broadly distributed when further collections are obtained from the remote regions lying between the
|
||
<collectingCountry id="F340F1975B72FFDDFB10F9B5FA6508B7" box="[1215,1436,1660,1687]" name="Papua New Guinea" pageId="15" pageNumber="147">Papua New Guinea</collectingCountry>
|
||
border and the Paniai Lakes. Based on apparent homologies in the male gentalic characters,
|
||
<taxonomicName id="4C57CA845B72FFDDFB0DF969FB0D089B" authorityName="Polhemus" authorityYear="2020" box="[1186,1268,1697,1722]" class="Insecta" family="Notonectidae" genus="Enithares" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="147" phylum="Arthropoda" rank="species" species="ziwa">
|
||
<emphasis id="B9236D155B72FFDDFB0DF969FB0D089B" box="[1186,1268,1697,1722]" italics="true" pageId="15" pageNumber="147">E. ziwa</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C57CA845B72FFDDFAACF969FA94089B" authorityName="Lansbury" authorityYear="1968" box="[1283,1389,1696,1722]" class="Insecta" family="Notonectidae" genus="Enithares" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="147" phylum="Arthropoda" rank="species" species="stylata">
|
||
<emphasis id="B9236D155B72FFDDFAACF969FA94089B" box="[1283,1389,1696,1722]" italics="true" pageId="15" pageNumber="147">E. stylata</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C57CA845B72FFDDFF38F90DFEB908FF" authorityName="Polhemus" authorityYear="2020" box="[151,320,1732,1758]" class="Insecta" family="Notonectidae" genus="Enithares" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="147" phylum="Arthropoda" rank="species" species="peninsularis">
|
||
<emphasis id="B9236D155B72FFDDFF38F90DFEB908FF" box="[151,320,1732,1758]" italics="true" pageId="15" pageNumber="147">E. peninsularis</emphasis>
|
||
</taxonomicName>
|
||
appear to be related, large-sized species occurring at elevations above
|
||
<quantity id="4CAF1CE25B72FFDDFBF2F90CFB4B08FE" box="[1117,1202,1732,1759]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="15" pageNumber="147" unit="m" value="1000.0">1000 m</quantity>
|
||
in the central mountains of New
|
||
<collectingCountry id="F340F1975B72FFDDFE81F920FE7B0923" box="[302,386,1768,1794]" name="Guinea-Bissau" pageId="15" pageNumber="147">Guinea</collectingCountry>
|
||
, which sequentially replace each other as one moves west-to-east along the uplift. Within this assemblage,
|
||
<taxonomicName id="4C57CA845B72FFDDFE8AF8C5FE8F0907" authorityName="Polhemus" authorityYear="2020" box="[293,374,1805,1830]" class="Insecta" family="Notonectidae" genus="Enithares" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="147" phylum="Arthropoda" rank="species" species="ziwa">
|
||
<emphasis id="B9236D155B72FFDDFE8AF8C5FE8F0907" box="[293,374,1805,1830]" italics="true" pageId="15" pageNumber="147">E. ziwa</emphasis>
|
||
</taxonomicName>
|
||
occupies the West Papuan Central Highlands area of freshwater endemism (Area 20) as defined by D.
|
||
<bibRefCitation id="EFC6CCF65B72FFDDFF73F8F8FDF9096B" author="Polhemus, D. A. & Allen, G. A." box="[220,512,1840,1866]" pageId="15" pageNumber="147" pagination="207 - 245" refId="ref39028" refString="Polhemus, D. A. & Allen, G. A. (2007) Freshwater biogeography of Papua. In: Marshall, A. J. & Beehler, B. M. (Eds.), Ecology of Papua. Part 1. Periplus Editions (HK) Ltd., Singapore, pp. 207 - 245." type="book chapter" year="2007">Polhemus & Allen (2007)</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BE8B1075B72FFDDFF68F89BFEA609DF" blockId="15.[151,1437,153,2046]" pageId="15" pageNumber="147">
|
||
This is the species treated in part under the concept of “
|
||
<taxonomicName id="4C57CA845B72FFDDFCEDF89CFC57094F" authorityName="Spinola" authorityYear="1837" box="[834,942,1876,1902]" class="Insecta" family="Notonectidae" genus="Enithares" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="147" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9236D155B72FFDDFCEDF89CFC57094F" box="[834,942,1876,1902]" italics="true" pageId="15" pageNumber="147">Enithares</emphasis>
|
||
</taxonomicName>
|
||
sp. undet #2” by D.
|
||
<bibRefCitation id="EFC6CCF65B72FFDDFB33F89BFAA7094F" author="Polhemus, D. A. & Polhemus, J. T." box="[1180,1374,1875,1902]" pageId="15" pageNumber="147" pagination="197 - 216" refId="ref39092" refString="Polhemus, D. A. & Polhemus, J. T. (2000) A biodiversity survey of aquatic insects in Ajkwa River Basin and adjacent areas, Irian Jaya, Indonesia. Tropical Biodiversity, 5 (3), 197 - 216." type="journal article" year="2000">Polhemus (2000)</bibRefCitation>
|
||
from sampling station
|
||
<quantity id="4CAF1CE25B72FFDDFEFAF8BFFE7409B0" box="[341,397,1911,1938]" metricMagnitude="0" metricUnit="m" metricValue="1.4224" pageId="15" pageNumber="147" unit="in" value="56.0">56 in</quantity>
|
||
the Conservation International rapid assessment report covering surveys in the Wapoga River Basin of north Indonesian New
|
||
<collectingCountry id="F340F1975B72FFDDFE55F853FDB60994" box="[506,591,1947,1973]" name="Guinea-Bissau" pageId="15" pageNumber="147">Guinea</collectingCountry>
|
||
. The collections from this site were initially thought to contain only a single
|
||
<taxonomicName id="4C57CA845B72FFDDFF38F877FEFA09F8" authorityName="Spinola" authorityYear="1837" box="[151,259,1983,2009]" class="Insecta" family="Notonectidae" genus="Enithares" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="147" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9236D155B72FFDDFF38F877FEFA09F8" box="[151,259,1983,2009]" italics="true" pageId="15" pageNumber="147">Enithares</emphasis>
|
||
</taxonomicName>
|
||
species, but upon closer examination proved to be a mixed series containing both
|
||
<taxonomicName id="4C57CA845B72FFDDFB06F808FA6509F8" authority="Lansbury" authorityName="Lansbury" authorityYear="1974" box="[1193,1436,1983,2009]" class="Insecta" family="Notonectidae" genus="Enithares" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="147" phylum="Arthropoda" rank="species" species="elongata">
|
||
<emphasis id="B9236D155B72FFDDFB06F808FAD209F8" box="[1193,1323,1983,2009]" italics="true" pageId="15" pageNumber="147">E. elongata</emphasis>
|
||
Lansbury
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C57CA845B72FFDDFF68F82CFEE009DC" authority="Polhemus, 2020" authorityName="Polhemus" authorityYear="2020" box="[199,281,2020,2045]" class="Insecta" family="Notonectidae" genus="Enithares" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="147" phylum="Arthropoda" rank="species" species="ziwa" status="sp. nov.">
|
||
<emphasis id="B9236D155B72FFDDFF68F82CFEE009DC" box="[199,281,2020,2045]" italics="true" pageId="15" pageNumber="147">E. ziwa</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9236D155B72FFDDFE8FF82CFEA609DF" bold="true" box="[288,351,2020,2046]" pageId="15" pageNumber="147">
|
||
<taxonomicNameLabel id="A210D06E5B72FFDDFE8FF82CFEA609DF" box="[288,351,2020,2046]" pageId="15" pageNumber="147" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</paragraph>
|
||
<caption id="DF28E18F5B6DFFC2FF38F91AFD5309BE" ID-DOI="http://doi.org/10.5281/zenodo.3814093" ID-Zenodo-Dep="3814093" httpUri="https://zenodo.org/record/3814093/files/figure.png" pageId="16" pageNumber="148" startId="16.[151,264,1746,1771]" targetBox="[199,1389,181,1721]" targetPageId="16">
|
||
<paragraph id="8BE8B1075B6DFFC2FF38F91AFD5309BE" blockId="16.[151,1437,1746,1951]" pageId="16" pageNumber="148">
|
||
<emphasis id="B9236D155B6DFFC2FF38F91AFEA908CA" bold="true" box="[151,336,1746,1771]" pageId="16" pageNumber="148">FIGURES 23–29</emphasis>
|
||
.
|
||
<taxonomicName id="4C57CA845B6DFFC2FEF0F91BFE3808CA" authorityName="Spinola" authorityYear="1837" box="[351,449,1747,1771]" class="Insecta" family="Notonectidae" genus="Enithares" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="148" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9236D155B6DFFC2FEF0F91BFE3808CA" box="[351,449,1747,1771]" italics="true" pageId="16" pageNumber="148">Enithares</emphasis>
|
||
</taxonomicName>
|
||
species in the
|
||
<taxonomicName id="4C57CA845B6DFFC2FDF1F91CFD5E08CA" box="[606,679,1748,1771]" class="Insecta" family="Notonectidae" genus="Enithares" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="148" phylum="Arthropoda" rank="species" species="atra">
|
||
<emphasis id="B9236D155B6DFFC2FDF1F91CFD5E08CA" box="[606,679,1748,1771]" italics="true" pageId="16" pageNumber="148">E. atra</emphasis>
|
||
</taxonomicName>
|
||
group, dorsal habitus photographs. 23–24.
|
||
<taxonomicName id="4C57CA845B6DFFC2FBC3F91BFA6008CA" authority="Lansbury" authorityName="Lansbury" authorityYear="1974" box="[1132,1433,1746,1771]" class="Insecta" family="Notonectidae" genus="Enithares" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="148" phylum="Arthropoda" rank="species" species="elongata">
|
||
<emphasis id="B9236D155B6DFFC2FBC3F91BFAC908CA" box="[1132,1328,1747,1771]" italics="true" pageId="16" pageNumber="148">Enithares elongata</emphasis>
|
||
Lansbury
|
||
</taxonomicName>
|
||
, male and female; specimens from Indonesia, Papua Province, Cyclops Mountains, CL 7145. 25–26.
|
||
<taxonomicName id="4C57CA845B6DFFC2FB05F93FFAAB092E" authority="Polhemus, 2020" authorityName="Polhemus" authorityYear="2020" box="[1194,1362,1783,1807]" class="Insecta" family="Notonectidae" genus="Enithares" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="148" phylum="Arthropoda" rank="species" species="papua" status="sp. nov.">
|
||
<emphasis id="B9236D155B6DFFC2FB05F93FFAAB092E" box="[1194,1362,1783,1807]" italics="true" pageId="16" pageNumber="148">Enithares papua</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9236D155B6DFFC2FAF4F93FFA6F092E" bold="true" box="[1371,1430,1783,1807]" pageId="16" pageNumber="148">
|
||
<taxonomicNameLabel id="A210D06E5B6DFFC2FAF4F93FFA6F092E" box="[1371,1430,1783,1807]" pageId="16" pageNumber="148" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, male and female; specimens from Papua New Guinea, Southern Highlands Prov., male specimen from Swinging Bridge, CL 7020, female specimen from Tubage, CL 7022. 27.
|
||
<taxonomicName id="4C57CA845B6DFFC2FD34F8F7FCC80977" authority="Polhemus, 2020" authorityName="Polhemus" authorityYear="2020" box="[667,817,1855,1879]" class="Insecta" family="Notonectidae" genus="Enithares" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="148" phylum="Arthropoda" rank="species" species="ziwa" status="sp. nov.">
|
||
<emphasis id="B9236D155B6DFFC2FD34F8F7FCC80977" box="[667,817,1855,1879]" italics="true" pageId="16" pageNumber="148">Enithares ziwa</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9236D155B6DFFC2FC98F8F7FC960976" bold="true" box="[823,879,1855,1879]" pageId="16" pageNumber="148">
|
||
<taxonomicNameLabel id="A210D06E5B6DFFC2FC98F8F7FC960976" box="[823,879,1855,1879]" pageId="16" pageNumber="148" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, male holotype, specimen from Indonesia, Papua Province, upper Ziwa River, CL 7100. 28–29.
|
||
<taxonomicName id="4C57CA845B6DFFC2FDE6F8ABFCA6095A" authority="Lansbury" authorityName="Lansbury" authorityYear="1968" box="[585,863,1890,1915]" class="Insecta" family="Notonectidae" genus="Enithares" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="148" phylum="Arthropoda" rank="species" species="stylata">
|
||
<emphasis id="B9236D155B6DFFC2FDE6F8ABFD0E095A" box="[585,759,1891,1915]" italics="true" pageId="16" pageNumber="148">Enithares stylata</emphasis>
|
||
Lansbury
|
||
</taxonomicName>
|
||
, male and female; specimens from Papua New Guinea, Western Highlands Prov., near Mt. Hagen, CL 1780.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BE8B1075B6CFFC3FF68FF50FE6C0F62" blockId="17.[151,1436,152,323]" pageId="17" pageNumber="149">
|
||
<emphasis id="B9236D155B6CFFC3FF68FF50FE7E0E92" bold="true" box="[199,391,152,179]" pageId="17" pageNumber="149">Ecological notes</emphasis>
|
||
. The
|
||
<typeStatus id="54EC0FA55B6CFFC3FE68FF51FE0E0E92" box="[455,503,153,179]" pageId="17" pageNumber="149">type</typeStatus>
|
||
series of
|
||
<taxonomicName id="4C57CA845B6CFFC3FDCCFF52FD4C0E92" authorityName="Polhemus" authorityYear="2020" box="[611,693,154,179]" class="Insecta" family="Notonectidae" genus="Enithares" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="149" phylum="Arthropoda" rank="species" species="ziwa">
|
||
<emphasis id="B9236D155B6CFFC3FDCCFF52FD4C0E92" box="[611,693,154,179]" italics="true" pageId="17" pageNumber="149">E. ziwa</emphasis>
|
||
</taxonomicName>
|
||
was taken from a pooled area along the margin of the Ziwa River, a swift mountain torrent draining from the northern slope of the New
|
||
<collectingCountry id="F340F1975B6CFFC3FC0FFF75FC080EF6" box="[928,1009,189,215]" name="Guinea-Bissau" pageId="17" pageNumber="149">Guinea</collectingCountry>
|
||
central ranges. The pool was adjacent to a set of cobble riffles, which graded into swift rapids in the main channel. The habitat preference of this species is similar to that of
|
||
<taxonomicName id="4C57CA845B6CFFC3FEDAFECEFE190F3E" authorityName="Lansbury" authorityYear="1968" box="[373,480,261,287]" class="Insecta" family="Notonectidae" genus="Enithares" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="149" phylum="Arthropoda" rank="species" species="stylata">
|
||
<emphasis id="B9236D155B6CFFC3FEDAFECEFE190F3E" box="[373,480,261,287]" italics="true" pageId="17" pageNumber="149">E. stylata</emphasis>
|
||
</taxonomicName>
|
||
, which also occurs in slower areas along the margins of swift rivers in the highlands of
|
||
<collectingCountry id="F340F1975B6CFFC3FF19FEE1FE6B0F62" box="[182,402,297,323]" name="Papua New Guinea" pageId="17" pageNumber="149">Papua New Guinea</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |