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<document id="8EE7914531B21CBCB5DE82666D5728D2" ID-CLB-Dataset="22330" ID-DOI="10.11646/zootaxa.4772.1.5" ID-GBIF-Dataset="b14b0d0d-cf67-4895-965c-12792c622b21" ID-ISSN="1175-5326" ID-Zenodo-Dep="3814079" ID-ZooBank="B497198A-08CD-4A21-AE04-14390499853B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1588834845818" checkinUser="plazi" docAuthor="Polhemus, Dan A." docDate="2020" docId="03FE00115B71FFDFFF38F91AFDFD096E" docLanguage="en" docName="zootaxa.4772.1.5.pdf" docOrigin="Zootaxa 4772 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Enithares peninsularis Polhemus 2020, new species" docType="treatment" docVersion="7" lastPageNumber="145" masterDocId="FFC778695B7DFFD2FFAFFFC8FFF90E21" masterDocTitle="Nine new species of Enithares (Heteroptera: Notonectidae) from New Guinea, with distributional notes on other species and an updated world checklist" masterLastPageNumber="182" masterPageNumber="132" pageNumber="144" updateTime="1698838866946" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="E36BD8C9FE76DC7FA292A5A9207B356D">
<mods:title id="FC0875C37ABA074B2321504EFCB1F462">Nine new species of Enithares (Heteroptera: Notonectidae) from New Guinea, with distributional notes on other species and an updated world checklist</mods:title>
</mods:titleInfo>
<mods:name id="119B2405F9CB57717F61CB1B9AD00093" type="personal">
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<mods:namePart id="F4D71357FAEAA35ECD425A0078806037">Polhemus, Dan A.</mods:namePart>
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<mods:title id="56B061E3D39921006A9D2A9371EFFEB2">Zootaxa</mods:title>
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<mods:part id="68718756E3D26D16EC0B81CD94275F6F">
<mods:date id="7F6F37F1DE3147C96C967CDC1FB896AE">2020</mods:date>
<mods:detail id="8415F5C355828B4D0FC98DDFAC151AF7" type="pubDate">
<mods:number id="68E1172555EF99F023BD6D4C48B39AA3">2020-05-07</mods:number>
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<mods:number id="71BAE192B398F0C26DE6E53D8AB452D4">4772</mods:number>
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<mods:classification id="339C08C904089F3AF9250149DE333B00">journal article</mods:classification>
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<mods:identifier id="8538514E7EFAAA67378F5FE56C14833C" type="DOI">10.11646/zootaxa.4772.1.5</mods:identifier>
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<treatment id="03FE00115B71FFDFFF38F91AFDFD096E" ID-DOI="http://doi.org/10.5281/zenodo.3815474" ID-GBIF-Taxon="163975550" ID-Zenodo-Dep="3815474" LSID="urn:lsid:plazi:treatment:03FE00115B71FFDFFF38F91AFDFD096E" httpUri="http://treatment.plazi.org/id/03FE00115B71FFDFFF38F91AFDFD096E" lastPageId="13" lastPageNumber="145" pageId="12" pageNumber="144">
<subSubSection id="C34DE28C5B71FFDEFF38F91AFDBA08CD" box="[151,579,1746,1773]" pageId="12" pageNumber="144" type="nomenclature">
<paragraph id="8BE8B1075B71FFDEFF38F91AFDBA08CD" blockId="12.[151,579,1746,1808]" box="[151,579,1746,1773]" pageId="12" pageNumber="144">
<heading id="D0A0066B5B71FFDEFF38F91AFDBA08CD" bold="true" box="[151,579,1746,1773]" fontSize="11" level="1" pageId="12" pageNumber="144" reason="1">
<emphasis id="B9236D155B71FFDEFF38F91AFDBA08CD" bold="true" box="[151,579,1746,1773]" pageId="12" pageNumber="144">
<taxonomicName id="4C57CA845B71FFDEFF38F91AFE5508CC" authority="Polhemus, 2020" authorityName="Polhemus" authorityYear="2020" box="[151,428,1746,1773]" class="Insecta" family="Notonectidae" genus="Enithares" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="144" phylum="Arthropoda" rank="species" species="peninsularis" status="sp. nov.">
<emphasis id="B9236D155B71FFDEFF38F91AFE5508CC" bold="true" box="[151,428,1746,1773]" italics="true" pageId="12" pageNumber="144">Enithares peninsularis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A210D06E5B71FFDEFE1CF91BFDBA08CD" box="[435,579,1746,1773]" pageId="12" pageNumber="144" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34DE28C5B71FFDEFF38F93EFEA10931" box="[151,344,1782,1808]" pageId="12" pageNumber="144" type="description">
<paragraph id="8BE8B1075B71FFDEFF38F93EFEA10931" blockId="12.[151,579,1746,1808]" box="[151,344,1782,1808]" pageId="12" pageNumber="144">
(
<figureCitation id="136CAD825B71FFDEFF30F93EFF040931" box="[159,253,1782,1808]" captionStart="FIGURES 1621" captionStartId="10.[151,264,1690,1715]" captionTargetBox="[151,1436,182,1664]" captionTargetId="figure@10.[151,1436,181,1664]" captionTargetPageId="10" captionText="FIGURES 1621. Enithares species in the E. atra group, male genitalic complex, left lateral view. 16. Enithares orsaki n. sp.; specimen from Papua New Guinea, Madang Prov., Baitabag, CL 7034. 17. Enithares atra Brooks; specimen from Papua New Guinea, Central Prov., Dorobisoro, CL 7264. 18. Enithares insularis n. sp.; specimen from Papua New Guinea, Milne Bay Prov., Fergusson Island, CL 7181. 19. Enithares bosavi n. sp., holotype; specimen from Papua New Guinea, Southern Highlands Prov., Mt. Bosavi, CL 7206. 20. Enithares papua n. sp.; specimen from Papua New Guinea, Central Prov., Tekadu, CL 7157. 21. Enithares peninsularis n. sp.; specimen from Papua New Guinea, Central Prov., Woitape, CL 7248." figureDoi="http://doi.org/10.5281/zenodo.3814089" httpUri="https://zenodo.org/record/3814089/files/figure.png" pageId="12" pageNumber="144">Figs. 21</figureCitation>
,
<figureCitation id="136CAD825B71FFDEFEA6F93EFEA90931" box="[265,336,1782,1808]" captionStart-0="FIGURE 30" captionStart-1="FIGURES 3134" captionStartId-0="17.[151,250,1174,1199]" captionStartId-1="20.[151,264,1869,1894]" captionTargetBox-0="[158,1429,354,1139]" captionTargetBox-1="[151,1436,181,1844]" captionTargetId-0="figure@17.[151,1436,345,1149]" captionTargetId-1="figure@20.[151,1436,181,1844]" captionTargetPageId-0="17" captionTargetPageId-1="20" captionText-0="FIGURE 30. Distribution of species in the E. atra group on New Guinea. Square = E. ziwa n. sp.; triangles = E. stylata Lansbury; pentagon = E. peninsularis n. sp." captionText-1="FIGURES 3134. Enithares species in the E. atra group, dorsal habitus photographs. 3132. Enithares peninsularis n. sp., male and female; specimens from Papua New Guinea, Central Prov., CL 7248. 3334. Enithares tagula n. sp., male and female; specimens from Papua New Guinea, Milne Bay Prov., CL 7190." figureDoi-0="http://doi.org/10.5281/zenodo.3814095" figureDoi-1="http://doi.org/10.5281/zenodo.3814097" httpUri-0="https://zenodo.org/record/3814095/files/figure.png" httpUri-1="https://zenodo.org/record/3814097/files/figure.png" pageId="12" pageNumber="144">3032</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C34DE28C5B71FFDFFF38F8F6FEA70EF6" lastPageId="13" lastPageNumber="145" pageId="12" pageNumber="144" type="materials_examined">
<paragraph id="8BE8B1075B71FFDEFF38F8F6FA6509C9" blockId="12.[151,1437,1853,2025]" pageId="12" pageNumber="144">
<emphasis id="B9236D155B71FFDEFF38F8F6FE460979" bold="true" box="[151,447,1853,1880]" pageId="12" pageNumber="144">Type material examined</emphasis>
.
<materialsCitation id="3B3FBB5A5B71FFDEFE7EF8F6FB4F0981" ID-GBIF-Occurrence="2610672370" collectingDate="2003-10-01" collectionCode="A, BPBM" country="Papua New Guinea" county="Udabe River" elevation="1630" latitude="-8.53389" location="Central Prov." longLatPrecision="21" longitude="147.23778" municipality="Owen Stanley Range" pageId="12" pageNumber="144" specimenCount="1" typeStatus="holotype">
<typeStatus id="54EC0FA55B71FFDEFE7EF8F6FDC50979" box="[465,572,1854,1880]" pageId="12" pageNumber="144" type="holotype">Holotype</typeStatus>
, male (dissected):
<collectingCountry id="F340F1975B71FFDEFCB7F8F6FBCA0979" box="[792,1075,1854,1881]" name="Papua New Guinea" pageId="12" pageNumber="144">PAPUA NEW GUINEA</collectingCountry>
,
<location id="8E88E7DC5B71FFDEFBEDF8F5FB110978" LSID="urn:lsid:plazi:treatment:03FE00115B71FFDFFF38F91AFDFD096E:8E88E7DC5B71FFDEFBEDF8F5FB110978" box="[1090,1256,1853,1881]" country="Papua New Guinea" county="Udabe River" latitude="-8.53389" longLatPrecision="21" longitude="147.23778" municipality="Owen Stanley Range" name="Central Prov." pageId="12" pageNumber="144">
<emphasis id="B9236D155B71FFDEFBEDF8F5FB1B0979" bold="true" box="[1090,1250,1853,1880]" pageId="12" pageNumber="144">Central Prov</emphasis>
.
</location>
,
<collectingMunicipality id="6B8C2B7D5B71FFDEFB54F8F6FF18095C" pageId="12" pageNumber="144">Owen Stanley Range</collectingMunicipality>
,
<collectingCounty id="6289C98B5B71FFDEFF5DF8AAFE7C095D" box="[242,389,1890,1916]" pageId="12" pageNumber="144">Udabe River</collectingCounty>
headwaters,
<quantity id="4CAF1CE25B71FFDEFD8EF8A9FD7C095D" box="[545,645,1889,1916]" metricMagnitude="3" metricUnit="m" metricValue="1.75" pageId="12" pageNumber="144" unit="km" value="1.75">1.75 km</quantity>
.
<locationDeviation id="9A5AA8255B71FFDEFD3BF8ABFC95095D" box="[660,876,1890,1917]" location="Woitape" pageId="12" pageNumber="144">WNW of Woitape</locationDeviation>
,
<quantity id="4CAF1CE25B71FFDEFCD4F8A9FBD9095C" box="[891,1056,1889,1917]" metricMagnitude="3" metricUnit="m" metricValue="1.63" metricValueMax="1.645" metricValueMin="1.6149999999999998" pageId="12" pageNumber="144" unit="m" value="1630.0" valueMax="1645.0" valueMin="1615.0">
<elevation id="007A56345B71FFDEFCD4F8A9FBD9095C" box="[891,1056,1889,1917]" metricMagnitude="3" metricUnit="m" metricValue="1.63" metricValueMax="1.645" metricValueMin="1.6149999999999998" pageId="12" pageNumber="144" unit="m" value="1630.0" valueMax="1645.0" valueMin="1615.0">16151645 m</elevation>
</quantity>
.,
<geoCoordinate id="EE63D7C05B71FFDEFB9AF8A9FB53095D" box="[1077,1194,1889,1916]" degrees="8" direction="south" minutes="32" orientation="latitude" pageId="12" pageNumber="144" precision="15" seconds="02" value="-8.53389">8°3202S</geoCoordinate>
,
<geoCoordinate id="EE63D7C05B71FFDEFB14F8A9FAB7095D" box="[1211,1358,1889,1916]" degrees="147" direction="east" minutes="14" orientation="longitude" pageId="12" pageNumber="144" precision="15" seconds="16" value="147.23778">147°1416E</geoCoordinate>
, water temp. 16.518.5 °C.,
<date id="FFE997C75B71FFDEFE22F84EFDBB0981" box="[397,578,1926,1952]" pageId="12" pageNumber="144" value="2003-10-01">
<collectingDate id="EFAD6E2F5B71FFDEFE22F84EFDBB0981" box="[397,578,1926,1952]" pageId="12" pageNumber="144" value="2003-10-01">1 October 2003</collectingDate>
</date>
, 09:0014:00 hrs., CL 7247, D.
<collectionCode id="ED4629C25B71FFDEFC19F84EFC340981" box="[950,973,1926,1952]" pageId="12" pageNumber="144">A</collectionCode>
. Polhemus (
<collectionCode id="ED4629C25B71FFDEFBF5F84EFB570981" box="[1114,1198,1926,1952]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" lsid="urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="12" pageNumber="144" type="Museum">BPBM</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B3FBB5A5B71FFDEFB6DF84FFF0B09C8" ID-GBIF-Occurrence="2610672585" collectingDate="2003-10-01" collectionCode="A, USNM, BPBM" country="Papua New Guinea" county="Udabe River" elevation="1630" latitude="-8.53389" location="Polhemus" longLatPrecision="21" longitude="147.23778" municipality="Owen Stanley Range" pageId="12" pageNumber="144" specimenCount="9" specimenCount-female="7" specimenCount-male="2" stateProvince="Central Prov" typeStatus="paratype">
<emphasis id="B9236D155B71FFDEFB6DF84FFACD0981" box="[1218,1332,1927,1952]" italics="true" pageId="12" pageNumber="144">
<typeStatus id="54EC0FA55B71FFDEFB6DF84FFACD0981" box="[1218,1332,1927,1952]" pageId="12" pageNumber="144" type="paratype">Paratypes</typeStatus>
</emphasis>
:
<collectingCountry id="F340F1975B71FFDEFAEAF84EFEB709E5" name="Papua New Guinea" pageId="12" pageNumber="144">PAPUA NEW GUINEA</collectingCountry>
,
<emphasis id="B9236D155B71FFDEFEF3F861FE0309E5" bold="true" box="[348,506,1961,1988]" pageId="12" pageNumber="144">
<collectingRegion id="49937FE55B71FFDEFEF3F861FE0309E5" box="[348,506,1961,1988]" pageId="12" pageNumber="144">Central Prov</collectingRegion>
</emphasis>
.,
<specimenCount id="9D517A8E5B71FFDEFDBEF862FD9509E5" box="[529,620,1962,1988]" pageId="12" pageNumber="144" type="male">2 males</specimenCount>
,
<specimenCount id="9D517A8E5B71FFDEFDD5F863FD1509E5" box="[634,748,1962,1989]" pageId="12" pageNumber="144" type="female">7 females</specimenCount>
, same data as holotype, CL 7247, D.
<collectionCode id="ED4629C25B71FFDEFB30F862FB4F09E5" box="[1183,1206,1962,1988]" pageId="12" pageNumber="144">A</collectionCode>
.
<location id="8E88E7DC5B71FFDEFB6CF862FAC809E5" LSID="urn:lsid:plazi:treatment:03FE00115B71FFDFFF38F91AFDFD096E:8E88E7DC5B71FFDEFB6CF862FAC809E5" box="[1219,1329,1962,1988]" country="Papua New Guinea" name="Polhemus" pageId="12" pageNumber="144" stateProvince="Central Prov">Polhemus</location>
(
<collectionCode id="ED4629C25B71FFDEFAECF862FA6109E5" box="[1347,1432,1962,1988]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="12" pageNumber="144" type="Museum">USNM</collectionCode>
,
<collectionCode id="ED4629C25B71FFDEFF38F807FF1309C8" box="[151,234,1999,2025]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" lsid="urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="12" pageNumber="144" type="Museum">BPBM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B3FBB5A5B71FFDEFF52F806FA6509C9" ID-GBIF-Occurrence="2610672542" box="[253,1436,1998,2025]" country="Papua New Guinea" location="Woitape" pageId="12" pageNumber="144" specimenCount="6" specimenCount-female="2" specimenCount-immature="1" specimenCount-male="3" stateProvince="Central Prov" typeStatus="paratype">
<specimenCount id="9D517A8E5B71FFDEFF52F806FEAF09C9" box="[253,342,1998,2024]" pageId="12" pageNumber="144" type="male">3 males</specimenCount>
,
<specimenCount id="9D517A8E5B71FFDEFECEF806FE2909C9" box="[353,464,1998,2024]" pageId="12" pageNumber="144" type="female">2 females</specimenCount>
,
<specimenCount id="9D517A8E5B71FFDEFE74F806FDA709C9" box="[475,606,1998,2024]" pageId="12" pageNumber="144" type="immature">1 immature</specimenCount>
, Owen Stanley Range, small hill streamlet,
<quantity id="4CAF1CE25B71FFDEFBE6F806FB6309C9" box="[1097,1178,1998,2024]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="12" pageNumber="144" unit="km" value="1.3">1.3 km</quantity>
. SSE of
<location id="8E88E7DC5B71FFDEFB55F806FAA309C9" LSID="urn:lsid:plazi:treatment:03FE00115B71FFDFFF38F91AFDFD096E:8E88E7DC5B71FFDEFB55F806FAA309C9" box="[1274,1370,1998,2024]" country="Papua New Guinea" name="Woitape" pageId="12" pageNumber="144" stateProvince="Central Prov">Woitape</location>
, 1600
</materialsCitation>
</paragraph>
<paragraph id="8BE8B1075B70FFDFFF38FF51FEA70EF6" blockId="13.[151,1437,152,1871]" pageId="13" pageNumber="145">
m.,
<materialsCitation id="3B3FBB5A5B70FFDFFF6BFF50FAD80E92" ID-GBIF-Occurrence="2610672385" box="[196,1313,152,179]" collectingDate="2003-10-02" collectionCode="A" latitude="-8.55" longLatPrecision="21" longitude="147.25945" pageId="13" pageNumber="145" specimenCount="1">
<geoCoordinate id="EE63D7C05B70FFDFFF6BFF50FEC30E92" box="[196,314,152,179]" degrees="8" direction="south" minutes="33" orientation="latitude" pageId="13" pageNumber="145" precision="15" seconds="00" value="-8.55">8°3300S</geoCoordinate>
,
<geoCoordinate id="EE63D7C05B70FFDFFEE8FF50FE230E92" box="[327,474,152,179]" degrees="147" direction="east" minutes="15" orientation="longitude" pageId="13" pageNumber="145" precision="15" seconds="34" value="147.25945">147°1534E</geoCoordinate>
, water temp. 17.5 °C.,
<date id="FFE997C75B70FFDFFD4BFF51FC610E92" box="[740,920,153,179]" pageId="13" pageNumber="145" value="2003-10-02">
<collectingDate id="EFAD6E2F5B70FFDFFD4BFF51FC610E92" box="[740,920,153,179]" pageId="13" pageNumber="145" value="2003-10-02">2 October 2003</collectingDate>
</date>
, 08:3009:15 hrs., CL 7248, D.
<collectionCode id="ED4629C25B70FFDFFAA5FF51FAD80E92" box="[1290,1313,153,179]" pageId="13" pageNumber="145">A</collectionCode>
</materialsCitation>
. Polhemus (
<collectionCode id="ED4629C25B70FFDFFF30FF75FF0D0EF6" box="[159,244,189,215]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="13" pageNumber="145" type="Museum">USNM</collectionCode>
,
<collectionCode id="ED4629C25B70FFDFFF50FF75FEAB0EF6" box="[255,338,189,215]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" lsid="urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="13" pageNumber="145" type="Museum">BPBM</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C34DE28C5B70FFDFFF68FF28FC1E0AE6" pageId="13" pageNumber="145" type="description">
<paragraph id="8BE8B1075B70FFDFFF68FF28FC0D0EDA" blockId="13.[151,1437,152,1871]" box="[199,1012,224,251]" pageId="13" pageNumber="145">
<emphasis id="B9236D155B70FFDFFF68FF28FEA80EDA" bold="true" box="[199,337,224,251]" pageId="13" pageNumber="145">Description</emphasis>
.
<emphasis id="B9236D155B70FFDFFECFFF29FE640EDA" bold="true" box="[352,413,225,251]" pageId="13" pageNumber="145">Male</emphasis>
: Length
<quantity id="4CAF1CE25B70FFDFFDAEFF29FD8C0EDA" box="[513,629,225,251]" metricMagnitude="-2" metricUnit="m" metricValue="1.22" pageId="13" pageNumber="145" unit="mm" value="12.2">12.20 mm</quantity>
, width across pronotum
<quantity id="4CAF1CE25B70FFDFFC26FF29FC090EDA" box="[905,1008,225,251]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" pageId="13" pageNumber="145" unit="mm" value="4.6">4.60 mm</quantity>
.
</paragraph>
<paragraph id="8BE8B1075B70FFDFFF68FECDFD890C1E" blockId="13.[151,1437,152,1871]" pageId="13" pageNumber="145">
<emphasis id="B9236D155B70FFDFFF68FECDFEBB0F3E" box="[199,322,261,287]" italics="true" pageId="13" pageNumber="145">Coloration</emphasis>
: Ground color dark brown (
<figureCitation id="136CAD825B70FFDFFD2FFECDFD2A0F3E" box="[640,723,261,287]" captionStart="FIGURES 3134" captionStartId="20.[151,264,1869,1894]" captionTargetBox="[151,1436,181,1844]" captionTargetId="figure@20.[151,1436,181,1844]" captionTargetPageId="20" captionText="FIGURES 3134. Enithares species in the E. atra group, dorsal habitus photographs. 3132. Enithares peninsularis n. sp., male and female; specimens from Papua New Guinea, Central Prov., CL 7248. 3334. Enithares tagula n. sp., male and female; specimens from Papua New Guinea, Milne Bay Prov., CL 7190." figureDoi="http://doi.org/10.5281/zenodo.3814097" httpUri="https://zenodo.org/record/3814097/files/figure.png" pageId="13" pageNumber="145">Fig. 31</figureCitation>
). Vertex and frons yellowish-orange; frons bearing a pair (1+1) of small irregular brown patches near anterior apex to either side of midline, and another roughly triangular brown patch along midline above base of labrum. Pronotum with anterior third lying between lateral foveae brown, bordered posteriorly by a creamy white transverse fascia, posterior half of pronotum dark brown. Scutellum dark brownish-black, posterior apex dark yellow. Hemelytra medium brown hyaline, certain areas appearing darker due to dorsal color showing through; wing membrane dark fumate brown. Legs yellowish brown, central portions of trochanters and femora, and anterior faces of fore tibia suffused with darker reddish-brown, hind femur narrowly margined with dark brown, a second similar narrow longitudinal stripe running parallel to this on ventral face. Venter brown, coxal plates margined with tan.
</paragraph>
<paragraph id="8BE8B1075B70FFDFFF68FD81FDA60D36" blockId="13.[151,1437,152,1871]" pageId="13" pageNumber="145">
<emphasis id="B9236D155B70FFDFFF68FD81FE4E0C42" box="[199,439,585,611]" italics="true" pageId="13" pageNumber="145">Structural characters</emphasis>
: Head broadly rounded anteriorly when viewed dorsally. Head length 1.00; greatest width 3.60, equal to 0.78 pronotal width; anterior width of vertex 1.40, equal to 1.4× head length. Synthlipsis 0.98, about 0.70 anterior width of vertex and clearly shorter than pronotum. Pronotal length along midline 1.40, humeral width 4.60, lateral margins convex, posterior margin weakly sinuate. Dorsal margin of pronotal fovea directed caudad behind eyes. Nodal furrow curved weakly cephalad, removed by over 2.0× its length from membranal suture, length 0.60, distance to membranal suture 1.40.
</paragraph>
<paragraph id="8BE8B1075B70FFDFFF68FCE9FB350D86" blockId="13.[151,1437,152,1871]" pageId="13" pageNumber="145">Front and hind legs typical for genus, lacking unusual modifications. Middle trochanter rounded. Middle femur with single large subapical tooth, bordered basally by about 8 small black pegs, distally by about 4 similar pegs. Lengths of leg segments as follows: fore femurtibiatarsal 1tarsal 2 = 2.00/2.00/0.80/0.40; middle femurtibiatarsal 1tarsal 2 = 2.30/2.10/0.90/0.60; hind femurtibiatarsal 1tarsal 2 = 4.80/4.00/1.80/1.10.</paragraph>
<paragraph id="8BE8B1075B70FFDFFF68FC79FB560DEA" blockId="13.[151,1437,152,1871]" box="[199,1199,945,971]" pageId="13" pageNumber="145">Ventral abdomen with metaxyphus elongate triangular, slightly concave, apex acuminate.</paragraph>
<paragraph id="8BE8B1075B70FFDFFF68FC1DFA9C0A5E" blockId="13.[151,1437,152,1871]" pageId="13" pageNumber="145">
<emphasis id="B9236D155B70FFDFFF68FC1DFE970DCE" box="[199,366,981,1007]" italics="true" pageId="13" pageNumber="145">Male genitalia</emphasis>
when viewed laterally (
<figureCitation id="136CAD825B70FFDFFD2AFC1DFD210DCE" box="[645,728,981,1007]" captionStart="FIGURES 1621" captionStartId="10.[151,264,1690,1715]" captionTargetBox="[151,1436,182,1664]" captionTargetId="figure@10.[151,1436,181,1664]" captionTargetPageId="10" captionText="FIGURES 1621. Enithares species in the E. atra group, male genitalic complex, left lateral view. 16. Enithares orsaki n. sp.; specimen from Papua New Guinea, Madang Prov., Baitabag, CL 7034. 17. Enithares atra Brooks; specimen from Papua New Guinea, Central Prov., Dorobisoro, CL 7264. 18. Enithares insularis n. sp.; specimen from Papua New Guinea, Milne Bay Prov., Fergusson Island, CL 7181. 19. Enithares bosavi n. sp., holotype; specimen from Papua New Guinea, Southern Highlands Prov., Mt. Bosavi, CL 7206. 20. Enithares papua n. sp.; specimen from Papua New Guinea, Central Prov., Tekadu, CL 7157. 21. Enithares peninsularis n. sp.; specimen from Papua New Guinea, Central Prov., Woitape, CL 7248." figureDoi="http://doi.org/10.5281/zenodo.3814089" httpUri="https://zenodo.org/record/3814089/files/figure.png" pageId="13" pageNumber="145">Fig. 21</figureCitation>
) with proctiger roughly rectangular, apex angular; pygophore with posteroventral angle forming an obtuse angle bearing a prominent setal tuft, posterior lobe of pygophore well produced, dorsal margin truncate, slightly notched anterodorsally; paramere large, tapering, apex blunt, reaching to apex of posterior lobe; lateral arm of basal plate large, basal section gently curving, bearing an anteroventral process, distal section gradually tapering, apex blunt, rounded; aedeagus semicircular, with numerous transverse folds.
</paragraph>
<paragraph id="8BE8B1075B70FFDFFF68FB41FC1E0AE6" blockId="13.[151,1437,152,1871]" pageId="13" pageNumber="145">
<emphasis id="B9236D155B70FFDFFF68FB41FEE70A82" bold="true" box="[199,286,1161,1187]" pageId="13" pageNumber="145">Female</emphasis>
: Length
<quantity id="4CAF1CE25B70FFDFFE28FB41FE040A82" box="[391,509,1161,1187]" metricMagnitude="-2" metricUnit="m" metricValue="1.27" pageId="13" pageNumber="145" unit="mm" value="12.7">12.70 mm</quantity>
, width across pronotum
<quantity id="4CAF1CE25B70FFDFFCB8FB41FC860A82" box="[791,895,1161,1187]" metricMagnitude="-3" metricUnit="m" metricValue="4.9" pageId="13" pageNumber="145" unit="mm" value="4.9">4.90 mm</quantity>
. Similar to male in general structure and coloration, but with pronotum and scutellum entirely yellowish-orange (
<figureCitation id="136CAD825B70FFDFFC29FB65FC220AE6" box="[902,987,1197,1223]" captionStart="FIGURES 3134" captionStartId="20.[151,264,1869,1894]" captionTargetBox="[151,1436,181,1844]" captionTargetId="figure@20.[151,1436,181,1844]" captionTargetPageId="20" captionText="FIGURES 3134. Enithares species in the E. atra group, dorsal habitus photographs. 3132. Enithares peninsularis n. sp., male and female; specimens from Papua New Guinea, Central Prov., CL 7248. 3334. Enithares tagula n. sp., male and female; specimens from Papua New Guinea, Milne Bay Prov., CL 7190." figureDoi="http://doi.org/10.5281/zenodo.3814097" httpUri="https://zenodo.org/record/3814097/files/figure.png" pageId="13" pageNumber="145">Fig. 32</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C34DE28C5B70FFDFFF68FB19FE9C0B2E" pageId="13" pageNumber="145" type="etymology">
<paragraph id="8BE8B1075B70FFDFFF68FB19FE9C0B2E" blockId="13.[151,1437,152,1871]" pageId="13" pageNumber="145">
<emphasis id="B9236D155B70FFDFFF68FB19FEB10ACA" bold="true" box="[199,328,1233,1259]" pageId="13" pageNumber="145">Etymology</emphasis>
. The name “peninsularis” refers to the geographic range of this species, which is confined to the Papuan Peninsula.
</paragraph>
</subSubSection>
<subSubSection id="C34DE28C5B70FFDFFF68FAD0FDFD096E" pageId="13" pageNumber="145" type="discussion">
<paragraph id="8BE8B1075B70FFDFFF68FAD0FF170872" blockId="13.[151,1437,152,1871]" pageId="13" pageNumber="145">
<emphasis id="B9236D155B70FFDFFF68FAD0FEBC0B12" bold="true" box="[199,325,1304,1331]" pageId="13" pageNumber="145">Discussion</emphasis>
. In terms of size,
<taxonomicName id="4C57CA845B70FFDFFDBEFAD2FD430B12" authorityName="Polhemus" authorityYear="2020" box="[529,698,1305,1331]" class="Insecta" family="Notonectidae" genus="Enithares" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="145" phylum="Arthropoda" rank="species" species="peninsularis">
<emphasis id="B9236D155B70FFDFFDBEFAD2FD430B12" box="[529,698,1305,1331]" italics="true" pageId="13" pageNumber="145">E. peninsularis</emphasis>
</taxonomicName>
is one of the largest
<taxonomicName id="4C57CA845B70FFDFFC09FAD1FBEB0B12" authorityName="Spinola" authorityYear="1837" box="[934,1042,1305,1331]" class="Insecta" family="Notonectidae" genus="Enithares" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis id="B9236D155B70FFDFFC09FAD1FBEB0B12" box="[934,1042,1305,1331]" italics="true" pageId="13" pageNumber="145">Enithares</emphasis>
</taxonomicName>
species occurring on New
<collectingCountry id="F340F1975B70FFDFFAEBFAD1FA610B12" box="[1348,1432,1305,1331]" name="Guinea-Bissau" pageId="13" pageNumber="145">Guinea</collectingCountry>
, with body lengths in both sexes exceeding
<quantity id="4CAF1CE25B70FFDFFDD4FAF5FD340B76" box="[635,717,1341,1367]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="13" pageNumber="145" unit="mm" value="12.0">12 mm</quantity>
. The genitalic morphology of
<taxonomicName id="4C57CA845B70FFDFFB8EFAF6FB320B76" authorityName="Polhemus" authorityYear="2020" box="[1057,1227,1341,1367]" class="Insecta" family="Notonectidae" genus="Enithares" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="145" phylum="Arthropoda" rank="species" species="peninsularis">
<emphasis id="B9236D155B70FFDFFB8EFAF6FB320B76" box="[1057,1227,1341,1367]" italics="true" pageId="13" pageNumber="145">E. peninsularis</emphasis>
</taxonomicName>
shares similarities to that of
<taxonomicName id="4C57CA845B70FFDFFEADFAAAFE920B5A" authorityName="Lansbury" authorityYear="1968" box="[258,363,1377,1403]" class="Insecta" family="Notonectidae" genus="Enithares" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="145" phylum="Arthropoda" rank="species" species="stylata">
<emphasis id="B9236D155B70FFDFFEADFAAAFE920B5A" box="[258,363,1377,1403]" italics="true" pageId="13" pageNumber="145">E. stylata</emphasis>
</taxonomicName>
, another high altitude species, particularly in regard to the blunt, dorsally notched posterior lobe on the pygophore (PL), and the elongate paramere with a rounded apex. However, the shape of the proctiger when viewed laterally is roughly rectangular in
<taxonomicName id="4C57CA845B70FFDFFDC6FA62FCE80BE2" authorityName="Polhemus" authorityYear="2020" box="[617,785,1449,1475]" class="Insecta" family="Notonectidae" genus="Enithares" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="145" phylum="Arthropoda" rank="species" species="peninsularis">
<emphasis id="B9236D155B70FFDFFDC6FA62FCE80BE2" box="[617,785,1449,1475]" italics="true" pageId="13" pageNumber="145">E. peninsularis</emphasis>
</taxonomicName>
, versus roughly triangular in
<taxonomicName id="4C57CA845B70FFDFFBF4FA62FB3C0BE2" authorityName="Lansbury" authorityYear="1968" box="[1115,1221,1449,1475]" class="Insecta" family="Notonectidae" genus="Enithares" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="145" phylum="Arthropoda" rank="species" species="stylata">
<emphasis id="B9236D155B70FFDFFBF4FA62FB3C0BE2" box="[1115,1221,1449,1475]" italics="true" pageId="13" pageNumber="145">E. stylata</emphasis>
</taxonomicName>
(compare
<figureCitation id="136CAD825B70FFDFFA94FA61FA6E0BE2" box="[1339,1431,1449,1475]" captionStart="FIGURES 1621" captionStartId="10.[151,264,1690,1715]" captionTargetBox="[151,1436,182,1664]" captionTargetId="figure@10.[151,1436,181,1664]" captionTargetPageId="10" captionText="FIGURES 1621. Enithares species in the E. atra group, male genitalic complex, left lateral view. 16. Enithares orsaki n. sp.; specimen from Papua New Guinea, Madang Prov., Baitabag, CL 7034. 17. Enithares atra Brooks; specimen from Papua New Guinea, Central Prov., Dorobisoro, CL 7264. 18. Enithares insularis n. sp.; specimen from Papua New Guinea, Milne Bay Prov., Fergusson Island, CL 7181. 19. Enithares bosavi n. sp., holotype; specimen from Papua New Guinea, Southern Highlands Prov., Mt. Bosavi, CL 7206. 20. Enithares papua n. sp.; specimen from Papua New Guinea, Central Prov., Tekadu, CL 7157. 21. Enithares peninsularis n. sp.; specimen from Papua New Guinea, Central Prov., Woitape, CL 7248." figureDoi="http://doi.org/10.5281/zenodo.3814089" httpUri="https://zenodo.org/record/3814089/files/figure.png" pageId="13" pageNumber="145">Figs. 21</figureCitation>
,
<figureCitation id="136CAD825B70FFDFFF38FA05FF4E0BC6" box="[151,183,1485,1511]" captionStart="FIGURES 3540" captionStartId="23.[151,264,1623,1648]" captionTargetBox="[170,1386,169,1547]" captionTargetId="figure@23.[151,1400,151,1568]" captionTargetPageId="23" captionText="FIGURES 3540. Enithares species in the E. atra and E. bakeri groups, male genitalic complex, left lateral view. 35. Enithares elongata Lansbury; specimen from Papua New Guinea, Morobe Prov., Wareo. 36. Enithares ziwa n. sp.; specimen from Indonesia, Papua Prov., Ziwa River, CL 7100. 37. Enithares stylata Lansbury; specimen from Papua New Guinea, Morobe Prov., Wau, CL 1817. 38. Enithares tagula n. sp.; specimen from Papua New Guinea, Milne Bay Prov. Tagula Is., CL 7190. 39. Enithares bakeri Brooks, specimen from Indonesia, West Papua Prov., Aimasi River, CL 2651. 40. Enithares loria Brooks; specimen from Indonesia, Papua Prov., Siewa, CL 7085." figureDoi="http://doi.org/10.5281/zenodo.3814099" httpUri="https://zenodo.org/record/3814099/files/figure.png" pageId="13" pageNumber="145">37</figureCitation>
). In addition, the shape of the LABP will immediately separate the two species, with that of
<taxonomicName id="4C57CA845B70FFDFFB68FA06FA880BC6" authorityName="Polhemus" authorityYear="2020" box="[1223,1393,1485,1511]" class="Insecta" family="Notonectidae" genus="Enithares" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="145" phylum="Arthropoda" rank="species" species="peninsularis">
<emphasis id="B9236D155B70FFDFFB68FA06FA880BC6" box="[1223,1393,1485,1511]" italics="true" pageId="13" pageNumber="145">E. peninsularis</emphasis>
</taxonomicName>
being stout and gently curving, with a blunt apex and a prominent ventral process, whereas that of
<taxonomicName id="4C57CA845B70FFDFFB78FA3AFAB8082A" authorityName="Lansbury" authorityYear="1968" box="[1239,1345,1521,1547]" class="Insecta" family="Notonectidae" genus="Enithares" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="145" phylum="Arthropoda" rank="species" species="stylata">
<emphasis id="B9236D155B70FFDFFB78FA3AFAB8082A" box="[1239,1345,1521,1547]" italics="true" pageId="13" pageNumber="145">E. stylata</emphasis>
</taxonomicName>
is more strongly curved and tapering, coming to an acute, slightly hooked apex, and lacks a ventral process (compare
<figureCitation id="136CAD825B70FFDFFACAF9DDFF4C0872" captionStart="FIGURES 1621" captionStartId="10.[151,264,1690,1715]" captionTargetBox="[151,1436,182,1664]" captionTargetId="figure@10.[151,1436,181,1664]" captionTargetPageId="10" captionText="FIGURES 1621. Enithares species in the E. atra group, male genitalic complex, left lateral view. 16. Enithares orsaki n. sp.; specimen from Papua New Guinea, Madang Prov., Baitabag, CL 7034. 17. Enithares atra Brooks; specimen from Papua New Guinea, Central Prov., Dorobisoro, CL 7264. 18. Enithares insularis n. sp.; specimen from Papua New Guinea, Milne Bay Prov., Fergusson Island, CL 7181. 19. Enithares bosavi n. sp., holotype; specimen from Papua New Guinea, Southern Highlands Prov., Mt. Bosavi, CL 7206. 20. Enithares papua n. sp.; specimen from Papua New Guinea, Central Prov., Tekadu, CL 7157. 21. Enithares peninsularis n. sp.; specimen from Papua New Guinea, Central Prov., Woitape, CL 7248." figureDoi="http://doi.org/10.5281/zenodo.3814089" httpUri="https://zenodo.org/record/3814089/files/figure.png" pageId="13" pageNumber="145">Figs. 21</figureCitation>
,
<figureCitation id="136CAD825B70FFDFFF6EF9F1FF180872" box="[193,225,1593,1619]" captionStart="FIGURES 3540" captionStartId="23.[151,264,1623,1648]" captionTargetBox="[170,1386,169,1547]" captionTargetId="figure@23.[151,1400,151,1568]" captionTargetPageId="23" captionText="FIGURES 3540. Enithares species in the E. atra and E. bakeri groups, male genitalic complex, left lateral view. 35. Enithares elongata Lansbury; specimen from Papua New Guinea, Morobe Prov., Wareo. 36. Enithares ziwa n. sp.; specimen from Indonesia, Papua Prov., Ziwa River, CL 7100. 37. Enithares stylata Lansbury; specimen from Papua New Guinea, Morobe Prov., Wau, CL 1817. 38. Enithares tagula n. sp.; specimen from Papua New Guinea, Milne Bay Prov. Tagula Is., CL 7190. 39. Enithares bakeri Brooks, specimen from Indonesia, West Papua Prov., Aimasi River, CL 2651. 40. Enithares loria Brooks; specimen from Indonesia, Papua Prov., Siewa, CL 7085." figureDoi="http://doi.org/10.5281/zenodo.3814099" httpUri="https://zenodo.org/record/3814099/files/figure.png" pageId="13" pageNumber="145">37</figureCitation>
).
</paragraph>
<paragraph id="8BE8B1075B70FFDFFF68F994FCA808BA" blockId="13.[151,1437,152,1871]" pageId="13" pageNumber="145">
<emphasis id="B9236D155B70FFDFFF68F994FE7E0856" bold="true" box="[199,391,1628,1655]" pageId="13" pageNumber="145">Ecological notes</emphasis>
. At the
<typeStatus id="54EC0FA55B70FFDFFE4EF995FDE80856" box="[481,529,1629,1655]" pageId="13" pageNumber="145">type</typeStatus>
locality near Woitape (CL 7247),
<taxonomicName id="4C57CA845B70FFDFFC3DF996FBC50856" authorityName="Polhemus" authorityYear="2020" box="[914,1084,1629,1655]" class="Insecta" family="Notonectidae" genus="Enithares" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="145" phylum="Arthropoda" rank="species" species="peninsularis">
<emphasis id="B9236D155B70FFDFFC3DF996FBC50856" box="[914,1084,1629,1655]" italics="true" pageId="13" pageNumber="145">E. peninsularis</emphasis>
</taxonomicName>
was taken from the still waters of a deep, algae-filled side pool formed behind large boulders.
</paragraph>
<paragraph id="8BE8B1075B70FFDFFF68F96DFDFD096E" blockId="13.[151,1437,152,1871]" pageId="13" pageNumber="145">
<taxonomicName id="4C57CA845B70FFDFFF68F96DFE3D089E" authorityName="Polhemus" authorityYear="2020" box="[199,452,1701,1727]" class="Insecta" family="Notonectidae" genus="Enithares" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="145" phylum="Arthropoda" rank="species" species="peninsularis">
<emphasis id="B9236D155B70FFDFFF68F96DFE3D089E" box="[199,452,1701,1727]" italics="true" pageId="13" pageNumber="145">Enithares peninsularis</emphasis>
</taxonomicName>
appears to be restricted to the higher elevations in the Owen Stanley Range of the Papuan Peninsula (
<figureCitation id="136CAD825B70FFDFFEEEF901FE6C08C2" box="[321,405,1737,1763]" captionStart="FIGURE 30" captionStartId="17.[151,250,1174,1199]" captionTargetBox="[158,1429,354,1139]" captionTargetId="figure@17.[151,1436,345,1149]" captionTargetPageId="17" captionText="FIGURE 30. Distribution of species in the E. atra group on New Guinea. Square = E. ziwa n. sp.; triangles = E. stylata Lansbury; pentagon = E. peninsularis n. sp." figureDoi="http://doi.org/10.5281/zenodo.3814095" httpUri="https://zenodo.org/record/3814095/files/figure.png" pageId="13" pageNumber="145">Fig. 30</figureCitation>
). The only other
<taxonomicName id="4C57CA845B70FFDFFDFEF901FD4408C2" authorityName="Spinola" authorityYear="1837" box="[593,701,1737,1763]" class="Insecta" family="Notonectidae" genus="Enithares" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis id="B9236D155B70FFDFFDFEF901FD4408C2" box="[593,701,1737,1763]" italics="true" pageId="13" pageNumber="145">Enithares</emphasis>
</taxonomicName>
species on New
<collectingCountry id="F340F1975B70FFDFFCD8F901FC3108C2" box="[887,968,1737,1763]" name="Guinea-Bissau" pageId="13" pageNumber="145">Guinea</collectingCountry>
of similar large size is
<taxonomicName id="4C57CA845B70FFDFFB66F902FACB08C2" authorityName="Lansbury" authorityYear="1968" box="[1225,1330,1737,1763]" class="Insecta" family="Notonectidae" genus="Enithares" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="145" phylum="Arthropoda" rank="species" species="stylata">
<emphasis id="B9236D155B70FFDFFB66F902FACB08C2" box="[1225,1330,1737,1763]" italics="true" pageId="13" pageNumber="145">E. stylata</emphasis>
</taxonomicName>
, which is widespread through the central highlands of
<collectingCountry id="F340F1975B70FFDFFD24F925FC9D0926" box="[651,868,1773,1799]" name="Papua New Guinea" pageId="13" pageNumber="145">Papua New Guinea</collectingCountry>
west of the Aure Scarp (
<figureCitation id="136CAD825B70FFDFFBD6F925FB370926" box="[1145,1230,1773,1799]" captionStart="FIGURE 30" captionStartId="17.[151,250,1174,1199]" captionTargetBox="[158,1429,354,1139]" captionTargetId="figure@17.[151,1436,345,1149]" captionTargetPageId="17" captionText="FIGURE 30. Distribution of species in the E. atra group on New Guinea. Square = E. ziwa n. sp.; triangles = E. stylata Lansbury; pentagon = E. peninsularis n. sp." figureDoi="http://doi.org/10.5281/zenodo.3814095" httpUri="https://zenodo.org/record/3814095/files/figure.png" pageId="13" pageNumber="145">Fig. 30</figureCitation>
). Based on known records,
<taxonomicName id="4C57CA845B70FFDFFF57F8DAFE5A090A" authorityName="Polhemus" authorityYear="2020" box="[248,419,1809,1835]" class="Insecta" family="Notonectidae" genus="Enithares" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="145" phylum="Arthropoda" rank="species" species="peninsularis">
<emphasis id="B9236D155B70FFDFFF57F8DAFE5A090A" box="[248,419,1809,1835]" italics="true" pageId="13" pageNumber="145">E. peninsularis</emphasis>
</taxonomicName>
occupies the Owen Stanley Mountains area of freshwater endemism (Area 31) as defined by D.
<bibRefCitation id="EFC6CCF65B70FFDFFF73F8FDFDF9096E" author="Polhemus, D. A. &amp; Allen, G. A." box="[220,512,1845,1871]" pageId="13" pageNumber="145" pagination="207 - 245" refId="ref39028" refString="Polhemus, D. A. &amp; Allen, G. A. (2007) Freshwater biogeography of Papua. In: Marshall, A. J. &amp; Beehler, B. M. (Eds.), Ecology of Papua. Part 1. Periplus Editions (HK) Ltd., Singapore, pp. 207 - 245." type="book chapter" year="2007">Polhemus &amp; Allen (2007)</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>