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<document id="392BA334E8DC70E081C2177CEF49DD79" ID-DOI="10.11646/zootaxa.1957.1.1" ID-ISSN="1175­5334" ID-Zenodo-Dep="5241137" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629590285433" checkinUser="felipe" docAuthor="Sartori, Michel, Peters, Janice G. &amp; Hubbard, Michael D." docDate="2008" docId="039AE86B4403FFFECE8E7285FE14F94C" docLanguage="en" docName="zt01957p051.pdf" docOrigin="Zootaxa 1957 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.1957.1.1" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Teloganodes tuberculatus Sartori &amp; Peters &amp; Hubbard 2008, sp. nov." docType="treatment" docVersion="4" lastPageNumber="15" masterDocId="FFA390134408FFF0CE197632FFA3FFCD" masterDocTitle="A revision of Oriental Teloganodidae (Insecta, Ephemeroptera, Ephemerelloidea)" masterLastPageNumber="51" masterPageNumber="1" pageNumber="12" updateTime="1699082080968" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="1AE653CF52281A6B06BE83ABB3847F13">A revision of Oriental Teloganodidae (Insecta, Ephemeroptera, Ephemerelloidea)</mods:title>
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<mods:namePart id="C7512A5E3A9098B34EBC24F942806AD3">Sartori, Michel</mods:namePart>
<mods:affiliation id="4A68E266DC33FE9183E90DA1EB5A3F0F">Museum of zoology, Palais de Rumine, Place Riponne 6, CH- 1014 Lausanne, Switzerland. E-mail: michel. sartori @ vd. ch Laboratory of Aquatic Entomology, Florida A &amp; M University, Tallahassee, FL 32307, USA</mods:affiliation>
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<mods:namePart id="4A05D78C66BE3907ADF992059D5CB46A">Peters, Janice G.</mods:namePart>
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<mods:namePart id="F8996E59BC06C8CA56818B9F3BAF0A1C">Hubbard, Michael D.</mods:namePart>
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<mods:title id="AE6E4DDCB0E825F1C84F3D9D44CE19F7">Zootaxa</mods:title>
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<mods:date id="8FFA8B4FC82CF39E5B1A0B920BEC6955">2008</mods:date>
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<mods:number id="9A9ECCB76314684A1A74713D4C8615B6">1957</mods:number>
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<treatment id="039AE86B4403FFFECE8E7285FE14F94C" ID-DOI="http://doi.org/10.5281/zenodo.5242472" ID-Zenodo-Dep="5242472" LSID="urn:lsid:plazi:treatment:039AE86B4403FFFECE8E7285FE14F94C" httpUri="http://treatment.plazi.org/id/039AE86B4403FFFECE8E7285FE14F94C" lastPageId="14" lastPageNumber="15" pageId="11" pageNumber="12">
<subSubSection id="C3290AF64403FFFBCE8E7285FD32FB1C" box="[151,657,1207,1233]" pageId="11" pageNumber="12" type="nomenclature">
<paragraph id="8B8C597D4403FFFBCE8E7285FD32FB1C" blockId="11.[151,733,1207,1268]" box="[151,657,1207,1233]" pageId="11" pageNumber="12">
<heading id="D0C4EE114403FFFBCE8E7285FD32FB1C" bold="true" box="[151,657,1207,1233]" fontSize="11" level="1" pageId="11" pageNumber="12" reason="1">
<emphasis id="B947856F4403FFFBCE8E7285FD32FB1C" bold="true" box="[151,657,1207,1233]" pageId="11" pageNumber="12">
<taxonomicName id="4C3322FE4403FFFBCE8E7285FD8FFB1C" ID-CoL="7BPTG" authority="Sartori" authorityName="Sartori &amp; Peters &amp; Hubbard" authorityYear="2008" box="[151,556,1207,1233]" class="Insecta" family="Teloganodidae" genus="Teloganodes" kingdom="Animalia" order="Ephemeroptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="tuberculatus" status="sp. nov.">
<emphasis id="B947856F4403FFFBCE8E7285FE68FB1C" bold="true" box="[151,459,1207,1233]" italics="true" pageId="11" pageNumber="12">Teloganodes tuberculatus</emphasis>
Sartori
</taxonomicName>
<taxonomicNameLabel id="A27438144403FFFBCC2D7285FD32FB1C" box="[564,657,1207,1233]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3290AF64403FFFBCE8E72E8FD7EFB39" box="[151,733,1242,1268]" pageId="11" pageNumber="12" type="description">
<paragraph id="8B8C597D4403FFFBCE8E72E8FD7EFB39" blockId="11.[151,733,1207,1268]" box="[151,733,1242,1268]" pageId="11" pageNumber="12">
(
<figureCitation id="130845F84403FFFBCE8672E8FE8EFB39" box="[159,301,1242,1268]" captionStart="FIGURES 1625" captionStartId="15.[151,269,1790,1814]" captionTargetBox="[180,1416,190,1708]" captionTargetId="figure-93@15.[151,1436,150,1738]" captionTargetPageId="15" captionText="FIGURES 1625. Labrum (16, 18, 20, 22, 24), with details of the dorsal setae (17, 19, 21, 23, 25), of Oriental Teloganodidae. Insertion of the setae indicated by dots on the labrum. Figs. 1617: Teloganodes tuberculatus; Figs. 1819: T. hubbardi; Figs. 2021: T. jacobusi; Figs. 2223: Dudgeodes hutanis; Figs. 2425: Derlethina eloisae. All labra at same scale and all setae at same scale, two times enlarged compared to the labra." figureDoi="http://doi.org/10.5281/zenodo.5241153" httpUri="https://zenodo.org/record/5241153/files/figure.png" pageId="11" pageNumber="12">Figs. 16, 17</figureCitation>
,
<figureCitation id="130845F84403FFFBCF2372E8FEF9FB39" box="[314,346,1242,1268]" captionStart="FIGURES 2632" captionStartId="16.[151,269,1832,1856]" captionTargetBox="[206,1387,216,1762]" captionTargetId="figure-68@16.[159,1444,194,1782]" captionTargetPageId="16" captionText="FIGURES 2632. Right mandible of Oriental Teloganodidae. Fig. 26: Teloganodes tuberculatus; Fig. 27: T. jacobusi; Fig. 28: Dudgeodes hutanis; Fig. 29: D. pescadori (shape of the new mandible in dotted line); Fig. 30: D. celebensis; Fig. 31: D. ulmeri; Fig. 32: Derlethina eloisae. All same scale." figureDoi="http://doi.org/10.5281/zenodo.5241155" httpUri="https://zenodo.org/record/5241155/files/figure.png" pageId="11" pageNumber="12">26</figureCitation>
,
<figureCitation id="130845F84403FFFBCF7C72E8FE26FB39" box="[357,389,1242,1268]" captionStart="FIGURES 3339" captionStartId="17.[151,269,1831,1855]" captionTargetBox="[200,1380,212,1766]" captionTargetId="figure-69@17.[171,1456,197,1785]" captionTargetPageId="17" captionText="FIGURES 3339. Left mandible of Oriental Teloganodidae. Fig. 33: Teloganodes tuberculatus; Fig. 34: T. jacobusi; Fig. 35: Dudgeodes hutanis; Fig. 36: D. pescadori (shape of the new mandible in dotted line); Fig. 37: D. celebensis; Fig. 38: D. ulmeri; Fig. 39: Derlethina eloisae. All same scale." figureDoi="http://doi.org/10.5281/zenodo.5241159" httpUri="https://zenodo.org/record/5241159/files/figure.png" pageId="11" pageNumber="12">33</figureCitation>
,
<figureCitation id="130845F84403FFFBCF8B72E8FE11FB39" box="[402,434,1242,1268]" captionStart="FIGURES 4046" captionStartId="18.[151,269,1781,1805]" captionTargetBox="[260,1412,191,1697]" captionTargetId="figure-70@18.[151,1436,156,1744]" captionTargetPageId="18" captionText="FIGURES 4046. Maxillae (40, 41) and detail of the setae on the apico-ventral part (4246) of Oriental Teloganodidae. Fig. 40: Teloganodes tuberculatus; Fig. 41: Dudgeodes hutanis; Fig. 42: T. jacobusi; Fig. 43: T. kodai; Fig. 44: D. pescadori; Fig. 45: D. ulmeri; Fig. 46: Derlethina eloisae." figureDoi="http://doi.org/10.5281/zenodo.5241161" httpUri="https://zenodo.org/record/5241161/files/figure.png" pageId="11" pageNumber="12">40</figureCitation>
,
<figureCitation id="130845F84403FFFBCFA472E8FE7FFB39" box="[445,476,1242,1268]" captionStart="FIGURES 4758" captionStartId="21.[151,269,1874,1898]" captionTargetBox="[186,1404,554,1862]" captionTargetId="figure-209@21.[151,1436,542,1868]" captionTargetPageId="21" captionText="FIGURES 4758. Hypopharynx (4749), labium (50) and labial palp (5158) of Oriental Teloganodidae. Fig. 47: Teloganodes jacobusi (setae on the left superlingua not drawn); Fig. 48: T. hubbardi (half right); Fig. 49: Dudgeodes hutanis (half left); Fig. 50: T. jacobusi; Fig. 51: T. tuberculatus; Fig. 52: T. hubbardi; Fig. 53: D. hutanis; Fig. 54: D. stephani; Fig. 55: D. celebensis; Fig. 56: D. ulmeri; Fig. 57: D. pescadori; Fig. 58; Derlethina eloisae. Hypopharynx at same scale; labium and labial palps at same scale." figureDoi="http://doi.org/10.5281/zenodo.5241167" httpUri="https://zenodo.org/record/5241167/files/figure.png" pageId="11" pageNumber="12">51</figureCitation>
,
<figureCitation id="130845F84403FFFBCFF072E8FDAAFB39" box="[489,521,1242,1268]" captionStart="FIGURES 5964" captionStartId="23.[151,269,1824,1848]" captionTargetBox="[179,1393,223,1780]" captionTargetId="figure-78@23.[151,1436,197,1785]" captionTargetPageId="23" captionText="FIGURES 5964. Forefemur (59, 60, 61, 63) and setae on the transverse row (62, 64) of Teloganodes spp. Insertion of the setae on the margins indicated by dots; only some setae drawn. Fig. 59: T. tuberculatus; Fig. 60: T. kodai; Figs. 61 62: T. jacobusi; Figs. 6364: T. hubbardi. Femora at same scale; setae at same scale." figureDoi="http://doi.org/10.5281/zenodo.5241169" httpUri="https://zenodo.org/record/5241169/files/figure.png" pageId="11" pageNumber="12">59</figureCitation>
, 8286, 120122)
</paragraph>
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<subSubSection id="C3290AF64403FFFBCE8E7318FF45FA71" pageId="11" pageNumber="12" type="materials_examined">
<paragraph id="8B8C597D4403FFFBCE8E7318FD8CFAA1" blockId="11.[151,1436,1322,2028]" pageId="11" pageNumber="12">
<emphasis id="B947856F4403FFFBCE8E7318FE2DFA89" bold="true" box="[151,398,1322,1348]" pageId="11" pageNumber="12">Material examined.</emphasis>
<materialsCitation id="3B5B53204403FFFBCF807318FD8CFAA1" collectingDate="1978-03-17" collectionCode="USNM" collectorName="M. Hubbard" country="Sri Lanka" county="Khone Palama" location="Kandy District" municipality="Haloya" pageId="11" pageNumber="12" specimenCount="1" typeStatus="holotype">
<typeStatus id="5488E7DF4403FFFBCF807318FDA8FA89" box="[409,523,1322,1348]" pageId="11" pageNumber="12" type="holotype">Holotype</typeStatus>
: 1 nymph (not mature),
<collectingCountry id="F32419ED4403FFFBCD2D7318FC0EFA89" box="[820,941,1322,1348]" name="Sri Lanka" pageId="11" pageNumber="12">Sri Lanka</collectingCountry>
,
<location id="8EEC0FA64403FFFBCDA57318FBD3FA89" LSID="urn:lsid:plazi:treatment:039AE86B4403FFFECE8E7285FE14F94C:8EEC0FA64403FFFBCDA57318FBD3FA89" box="[956,1136,1322,1348]" country="Sri Lanka" county="Khone Palama" municipality="Haloya" name="Kandy District" pageId="11" pageNumber="12">Kandy District</location>
,
<collectingMunicipality id="6BE8C3074403FFFBCA667318FB75FA89" box="[1151,1238,1322,1348]" pageId="11" pageNumber="12">Haloya</collectingMunicipality>
,
<collectingCounty id="62ED21F14403FFFBCAFF7318FA34FA89" box="[1254,1431,1322,1348]" pageId="11" pageNumber="12">Khone Palama</collectingCounty>
,
<date id="FF8D7FBD4403FFFBCE8E7360FEBEFAA1" box="[151,285,1362,1388]" pageId="11" pageNumber="12" value="1978-03-17">
<collectingDate id="EFC986554403FFFBCE8E7360FEBEFAA1" box="[151,285,1362,1388]" pageId="11" pageNumber="12" value="1978-03-17">17.III.1978</collectingDate>
</date>
,
<collectorName id="26C63CAB4403FFFBCF307360FE1BFAA1" box="[297,440,1362,1388]" pageId="11" pageNumber="12">M. Hubbard</collectorName>
[
<collectionCode id="ED22C1B84403FFFBCFDE7360FD86FAA1" box="[455,549,1362,1388]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="11" pageNumber="12" type="Museum">USNM</collectionCode>
].
</materialsCitation>
</paragraph>
<paragraph id="8B8C597D4403FFFBCEDF7348FF45FA71" blockId="11.[151,1436,1322,2028]" pageId="11" pageNumber="12">
<materialsCitation id="3B5B53204403FFFBCEDF7348FF41FA71" collectingDate="1978-03-17" collectionCode="FAMU, MZL" collectorName="M. Hubbard" country="Sri Lanka" county="Khone Palama" location="Kandy District" municipality="Haloya" pageId="11" pageNumber="12" specimenCount="1" typeStatus="paratype">
<typeStatus id="5488E7DF4403FFFBCEDF7348FE9EFA59" box="[198,317,1402,1428]" pageId="11" pageNumber="12" type="paratype">Paratypes</typeStatus>
: 3 nymphs, same data as holotype [
<quantity id="4CCBF4984403FFFBCCEB7348FC87FA59" box="[754,804,1402,1428]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="11" pageNumber="12" unit="in" value="2.0">2 in</quantity>
<collectionCode id="ED22C1B84403FFFBCD367348FC26FA59" box="[815,901,1402,1428]" pageId="11" pageNumber="12">FAMU</collectionCode>
, 1 partially mounted on slide preparation in
<collectionCode id="ED22C1B84403FFFBCE8E7390FF78FA71" box="[151,219,1442,1468]" country="Switzerland" httpUri="http://grbio.org/cool/pm5v-0p6q" name="Musee Zoologique" pageId="11" pageNumber="12">MZL</collectionCode>
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.
</paragraph>
</subSubSection>
<subSubSection id="C3290AF64403FFFECEDF73F8FE46FA74" lastPageId="14" lastPageNumber="15" pageId="11" pageNumber="12" type="description">
<paragraph id="8B8C597D4403FFFBCEDF73F8FE1DFA29" blockId="11.[151,1436,1322,2028]" box="[198,446,1482,1508]" pageId="11" pageNumber="12">
<emphasis id="B947856F4403FFFBCEDF73F8FEFDFA29" bold="true" box="[198,350,1482,1508]" pageId="11" pageNumber="12">Description.</emphasis>
Nymph
</paragraph>
<paragraph id="8B8C597D4403FFFBCEDC73C0FB26F9C1" blockId="11.[151,1436,1322,2028]" box="[197,1157,1522,1548]" pageId="11" pageNumber="12">
Body length at least
<quantity id="4CCBF4984403FFFBCFAF73C0FDACF9C1" box="[438,527,1522,1548]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="11" pageNumber="12" unit="mm" value="4.5">4.5 mm</quantity>
without cerci. Cerci length ca. 1.6 times body length.
</paragraph>
<paragraph id="8B8C597D4403FFFBCEDC7028FE0CF991" blockId="11.[151,1436,1322,2028]" pageId="11" pageNumber="12">
General colouration of head, thorax and abdomen medium brown dorsally; sterna and legs uniformly yellowish (
<figureCitation id="130845F84403FFFBCEEC7070FE01F991" box="[245,418,1602,1628]" captionStart="FIGURES 120125" captionStartId="33.[151,269,1811,1835]" captionTargetBox="[151,1436,197,1784]" captionTargetId="figure-49@33.[151,1436,197,1784]" captionTargetPageId="33" captionText="FIGURES 120125. Holotypes of Teloganodes spp. in dorsal view (120, 123), abdomen in ventral view (121, 124) and in lateral view (122, 125). Figs. 120122: T. tuberculatus; Figs. 123125: T. kodai." figureDoi="http://doi.org/10.5281/zenodo.5241188" httpUri="https://zenodo.org/record/5241188/files/figure.png" pageId="11" pageNumber="12">Figs. 120121</figureCitation>
).
</paragraph>
<paragraph id="8B8C597D4403FFFCCE8E7058FD63FECF" blockId="11.[151,1436,1322,2028]" lastBlockId="12.[151,1436,152,258]" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12">
Outer margin of head fringed with a row of setae from behind eyes to labrum insertion (
<figureCitation id="130845F84403FFFBCAB77058FAB5F949" box="[1198,1302,1642,1668]" captionStart="FIGURES 120125" captionStartId="33.[151,269,1811,1835]" captionTargetBox="[151,1436,197,1784]" captionTargetId="figure-49@33.[151,1436,197,1784]" captionTargetPageId="33" captionText="FIGURES 120125. Holotypes of Teloganodes spp. in dorsal view (120, 123), abdomen in ventral view (121, 124) and in lateral view (122, 125). Figs. 120122: T. tuberculatus; Figs. 123125: T. kodai." figureDoi="http://doi.org/10.5281/zenodo.5241188" httpUri="https://zenodo.org/record/5241188/files/figure.png" pageId="11" pageNumber="12">Fig. 120</figureCitation>
). Antennae short, 0.7 times head width, flagellum with 1011 segments. Labrum compact, ca. 2 times wider than long, with smooth anterior emargination (
<figureCitation id="130845F84403FFFBCC237088FD32F919" box="[570,657,1722,1748]" captionStart="FIGURES 1625" captionStartId="15.[151,269,1790,1814]" captionTargetBox="[180,1416,190,1708]" captionTargetId="figure-93@15.[151,1436,150,1738]" captionTargetPageId="15" captionText="FIGURES 1625. Labrum (16, 18, 20, 22, 24), with details of the dorsal setae (17, 19, 21, 23, 25), of Oriental Teloganodidae. Insertion of the setae indicated by dots on the labrum. Figs. 1617: Teloganodes tuberculatus; Figs. 1819: T. hubbardi; Figs. 2021: T. jacobusi; Figs. 2223: Dudgeodes hutanis; Figs. 2425: Derlethina eloisae. All labra at same scale and all setae at same scale, two times enlarged compared to the labra." figureDoi="http://doi.org/10.5281/zenodo.5241153" httpUri="https://zenodo.org/record/5241153/files/figure.png" pageId="11" pageNumber="12">Fig. 16</figureCitation>
); dorsal face covered medially by 24 rows of long feathered setae (
<figureCitation id="130845F84403FFFBCE8670D0FF54F931" box="[159,247,1762,1788]" captionStart="FIGURES 1625" captionStartId="15.[151,269,1790,1814]" captionTargetBox="[180,1416,190,1708]" captionTargetId="figure-93@15.[151,1436,150,1738]" captionTargetPageId="15" captionText="FIGURES 1625. Labrum (16, 18, 20, 22, 24), with details of the dorsal setae (17, 19, 21, 23, 25), of Oriental Teloganodidae. Insertion of the setae indicated by dots on the labrum. Figs. 1617: Teloganodes tuberculatus; Figs. 1819: T. hubbardi; Figs. 2021: T. jacobusi; Figs. 2223: Dudgeodes hutanis; Figs. 2425: Derlethina eloisae. All labra at same scale and all setae at same scale, two times enlarged compared to the labra." figureDoi="http://doi.org/10.5281/zenodo.5241153" httpUri="https://zenodo.org/record/5241153/files/figure.png" pageId="11" pageNumber="12">Fig. 17</figureCitation>
); anterior margin with a row of small thin setae. Mandibles (
<figureCitation id="130845F84403FFFBCDDB70D0FB86F931" box="[962,1061,1762,1788]" captionStart="FIGURES 2632" captionStartId="16.[151,269,1832,1856]" captionTargetBox="[206,1387,216,1762]" captionTargetId="figure-68@16.[159,1444,194,1782]" captionTargetPageId="16" captionText="FIGURES 2632. Right mandible of Oriental Teloganodidae. Fig. 26: Teloganodes tuberculatus; Fig. 27: T. jacobusi; Fig. 28: Dudgeodes hutanis; Fig. 29: D. pescadori (shape of the new mandible in dotted line); Fig. 30: D. celebensis; Fig. 31: D. ulmeri; Fig. 32: Derlethina eloisae. All same scale." figureDoi="http://doi.org/10.5281/zenodo.5241155" httpUri="https://zenodo.org/record/5241155/files/figure.png" pageId="11" pageNumber="12">Figs. 26</figureCitation>
,
<figureCitation id="130845F84403FFFBCA2870D0FBECF931" box="[1073,1103,1762,1788]" captionStart="FIGURES 3339" captionStartId="17.[151,269,1831,1855]" captionTargetBox="[200,1380,212,1766]" captionTargetId="figure-69@17.[171,1456,197,1785]" captionTargetPageId="17" captionText="FIGURES 3339. Left mandible of Oriental Teloganodidae. Fig. 33: Teloganodes tuberculatus; Fig. 34: T. jacobusi; Fig. 35: Dudgeodes hutanis; Fig. 36: D. pescadori (shape of the new mandible in dotted line); Fig. 37: D. celebensis; Fig. 38: D. ulmeri; Fig. 39: Derlethina eloisae. All same scale." figureDoi="http://doi.org/10.5281/zenodo.5241159" httpUri="https://zenodo.org/record/5241159/files/figure.png" pageId="11" pageNumber="12">33</figureCitation>
) slender with 2 thin setae in middle of outer margin; right mandible (
<figureCitation id="130845F84403FFFBCC777138FD60F8E9" box="[622,707,1802,1828]" captionStart="FIGURES 2632" captionStartId="16.[151,269,1832,1856]" captionTargetBox="[206,1387,216,1762]" captionTargetId="figure-68@16.[159,1444,194,1782]" captionTargetPageId="16" captionText="FIGURES 2632. Right mandible of Oriental Teloganodidae. Fig. 26: Teloganodes tuberculatus; Fig. 27: T. jacobusi; Fig. 28: Dudgeodes hutanis; Fig. 29: D. pescadori (shape of the new mandible in dotted line); Fig. 30: D. celebensis; Fig. 31: D. ulmeri; Fig. 32: Derlethina eloisae. All same scale." figureDoi="http://doi.org/10.5281/zenodo.5241155" httpUri="https://zenodo.org/record/5241155/files/figure.png" pageId="11" pageNumber="12">Fig. 26</figureCitation>
) with outer incisor with inner tooth; inner incisor with 2 teeth; prostheca reduced to a cluster of thin setae; a row of long, thin setae below mola; left mandible (
<figureCitation id="130845F84403FFFBCB1B7100FAF4F881" box="[1282,1367,1842,1868]" captionStart="FIGURES 3339" captionStartId="17.[151,269,1831,1855]" captionTargetBox="[200,1380,212,1766]" captionTargetId="figure-69@17.[171,1456,197,1785]" captionTargetPageId="17" captionText="FIGURES 3339. Left mandible of Oriental Teloganodidae. Fig. 33: Teloganodes tuberculatus; Fig. 34: T. jacobusi; Fig. 35: Dudgeodes hutanis; Fig. 36: D. pescadori (shape of the new mandible in dotted line); Fig. 37: D. celebensis; Fig. 38: D. ulmeri; Fig. 39: Derlethina eloisae. All same scale." figureDoi="http://doi.org/10.5281/zenodo.5241159" httpUri="https://zenodo.org/record/5241159/files/figure.png" pageId="11" pageNumber="12">Fig. 33</figureCitation>
) with long outer incisor rounded; inner tooth close to inner incisor; inner incisor with 3 teeth, prostheca well developed with cluster of small setae. Maxillae (
<figureCitation id="130845F84403FFFBCC8C71B0FD48F851" box="[661,747,1922,1948]" captionStart="FIGURES 4046" captionStartId="18.[151,269,1781,1805]" captionTargetBox="[260,1412,191,1697]" captionTargetId="figure-70@18.[151,1436,156,1744]" captionTargetPageId="18" captionText="FIGURES 4046. Maxillae (40, 41) and detail of the setae on the apico-ventral part (4246) of Oriental Teloganodidae. Fig. 40: Teloganodes tuberculatus; Fig. 41: Dudgeodes hutanis; Fig. 42: T. jacobusi; Fig. 43: T. kodai; Fig. 44: D. pescadori; Fig. 45: D. ulmeri; Fig. 46: Derlethina eloisae." figureDoi="http://doi.org/10.5281/zenodo.5241161" httpUri="https://zenodo.org/record/5241161/files/figure.png" pageId="11" pageNumber="12">Fig. 40</figureCitation>
) slender, with a well developed canine and 2 dentisetae; 3 long inner setae apically and a cluster of long, simple setae at crown; inner margin at base of lacinia with one long feathered seta dorsally and row of 34 short feathered setae ventrally; maxillary palp reduced to a simple seta. Submentum moderately developed laterally; glossae and paraglossae partially fused; glossae rhomboid and paraglossae slightly falciform (similar to
<figureCitation id="130845F84404FFFCCCA676F2FCBAFF17" box="[703,793,192,218]" captionStart="FIGURES 4758" captionStartId="21.[151,269,1874,1898]" captionTargetBox="[186,1404,554,1862]" captionTargetId="figure-209@21.[151,1436,542,1868]" captionTargetPageId="21" captionText="FIGURES 4758. Hypopharynx (4749), labium (50) and labial palp (5158) of Oriental Teloganodidae. Fig. 47: Teloganodes jacobusi (setae on the left superlingua not drawn); Fig. 48: T. hubbardi (half right); Fig. 49: Dudgeodes hutanis (half left); Fig. 50: T. jacobusi; Fig. 51: T. tuberculatus; Fig. 52: T. hubbardi; Fig. 53: D. hutanis; Fig. 54: D. stephani; Fig. 55: D. celebensis; Fig. 56: D. ulmeri; Fig. 57: D. pescadori; Fig. 58; Derlethina eloisae. Hypopharynx at same scale; labium and labial palps at same scale." figureDoi="http://doi.org/10.5281/zenodo.5241167" httpUri="https://zenodo.org/record/5241167/files/figure.png" pageId="12" pageNumber="13">Fig. 50</figureCitation>
); labial palp three-segmented, segment 1 subequal in length to segments 2 and 3 combined (
<figureCitation id="130845F84404FFFCCC4276DAFD11FECF" box="[603,690,232,258]" captionStart="FIGURES 4758" captionStartId="21.[151,269,1874,1898]" captionTargetBox="[186,1404,554,1862]" captionTargetId="figure-209@21.[151,1436,542,1868]" captionTargetPageId="21" captionText="FIGURES 4758. Hypopharynx (4749), labium (50) and labial palp (5158) of Oriental Teloganodidae. Fig. 47: Teloganodes jacobusi (setae on the left superlingua not drawn); Fig. 48: T. hubbardi (half right); Fig. 49: Dudgeodes hutanis (half left); Fig. 50: T. jacobusi; Fig. 51: T. tuberculatus; Fig. 52: T. hubbardi; Fig. 53: D. hutanis; Fig. 54: D. stephani; Fig. 55: D. celebensis; Fig. 56: D. ulmeri; Fig. 57: D. pescadori; Fig. 58; Derlethina eloisae. Hypopharynx at same scale; labium and labial palps at same scale." figureDoi="http://doi.org/10.5281/zenodo.5241167" httpUri="https://zenodo.org/record/5241167/files/figure.png" pageId="12" pageNumber="13">Fig. 51</figureCitation>
).
</paragraph>
<caption id="DF4C09F54404FFFCCE8E71B1FBA7F82D" ID-DOI="http://doi.org/10.5281/zenodo.5241143" ID-Zenodo-Dep="5241143" httpUri="https://zenodo.org/record/5241143/files/figure.png" pageId="12" pageNumber="13" startId="12.[151,255,1923,1947]" targetBox="[402,1190,310,1897]" targetPageId="12">
<paragraph id="8B8C597D4404FFFCCE8E71B1FBA7F82D" blockId="12.[151,1436,1923,2016]" pageId="12" pageNumber="13">
<emphasis id="B947856F4404FFFCCE8E71B1FE91F856" bold="true" box="[151,306,1923,1947]" pageId="12" pageNumber="13">FIGURE 7, 8.</emphasis>
Fig. 7: Phylogram of the single most parsimonious tree reconstructed: synapomorphies are represented by black boxes, number above the box: character as in table 1, number below the box: character state. Fig. 8: Bootstrap 50% majority-rule consensus tree: bootstrap support values indicated before each node.
</paragraph>
</caption>
<caption id="DF4C09F54405FFFDCE8E70E5FE6CF8F9" ID-DOI="http://doi.org/10.5281/zenodo.5241147" ID-Zenodo-Dep="5241147" httpUri="https://zenodo.org/record/5241147/files/figure.png" pageId="13" pageNumber="14" startId="13.[151,255,1751,1775]" targetBox="[186,1395,186,1715]" targetPageId="13">
<paragraph id="8B8C597D4405FFFDCE8E70E5FE6CF8F9" blockId="13.[151,1436,1751,1844]" pageId="13" pageNumber="14">
<emphasis id="B947856F4405FFFDCE8E70E5FEE7F922" bold="true" box="[151,324,1751,1775]" pageId="13" pageNumber="14">FIGURE 913.</emphasis>
<taxonomicName id="4C3322FE4405FFFDCF5770EAFE5FF922" box="[334,508,1752,1775]" class="Insecta" family="Teloganodidae" genus="Dudgeodes" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B947856F4405FFFDCF5770EAFE65F922" box="[334,454,1752,1775]" italics="true" pageId="13" pageNumber="14">Dudgeodes</emphasis>
spp.
</taxonomicName>
Fig. 9: Female subimago forewing of
<taxonomicName id="4C3322FE4405FFFDCDB570EAFBDFF922" box="[940,1148,1752,1775]" class="Insecta" family="Teloganodidae" genus="Dudgeodes" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="hutanis">
<emphasis id="B947856F4405FFFDCDB570EAFBDFF922" box="[940,1148,1752,1775]" italics="true" pageId="13" pageNumber="14">Dudgeodes hutanis</emphasis>
</taxonomicName>
; Fig. 10: Hindwing of
<taxonomicName id="4C3322FE4405FFFDCB9B70EAFF46F8DF" class="Insecta" family="Teloganodidae" genus="Dudgeodes" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="hutanis">
<emphasis id="B947856F4405FFFDCB9B70EAFF46F8DF" italics="true" pageId="13" pageNumber="14">D. hutanis</emphasis>
</taxonomicName>
; Fig. 11: Male imago forewing of
<taxonomicName id="4C3322FE4405FFFDCC4D70C9FC98F8DF" authorityName="Sartori &amp; Peters &amp; Hubbard" authorityYear="2008" box="[596,827,1787,1810]" class="Insecta" family="Teloganodidae" genus="Dudgeodes" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="pescadori">
<emphasis id="B947856F4405FFFDCC4D70C9FC98F8DF" box="[596,827,1787,1810]" italics="true" pageId="13" pageNumber="14">Dudgeodes pescadori</emphasis>
</taxonomicName>
; Fig. 12: Hindwing of
<taxonomicName id="4C3322FE4405FFFDCA3670C9FB1BF8DF" authorityName="Sartori &amp; Peters &amp; Hubbard" authorityYear="2008" box="[1071,1208,1787,1810]" class="Insecta" family="Teloganodidae" genus="Dudgeodes" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="pescadori">
<emphasis id="B947856F4405FFFDCA3670C9FB1BF8DF" box="[1071,1208,1787,1810]" italics="true" pageId="13" pageNumber="14">D. pescadori</emphasis>
</taxonomicName>
; Fig. 13: Genitalia in ventral view of
<taxonomicName id="4C3322FE4405FFFDCF26712FFE6BF8F9" authorityName="Sartori &amp; Peters &amp; Hubbard" authorityYear="2008" box="[319,456,1821,1844]" class="Insecta" family="Teloganodidae" genus="Dudgeodes" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="pescadori">
<emphasis id="B947856F4405FFFDCF26712FFE6BF8F9" box="[319,456,1821,1844]" italics="true" pageId="13" pageNumber="14">D. pescadori</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="8B8C597D4405FFFECEDF7150FE26FEEF" blockId="13.[151,1436,1890,2028]" lastBlockId="14.[151,1436,152,290]" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14">
Prothorax with 6 rounded tubercles on dorsal face; mesothorax with 5. All legs similar in shape and ornamentation. Femur (
<figureCitation id="130845F84405FFFDCF6071BAFE6CF86F" box="[377,463,1928,1954]" captionStart="FIGURES 5964" captionStartId="23.[151,269,1824,1848]" captionTargetBox="[179,1393,223,1780]" captionTargetId="figure-78@23.[151,1436,197,1785]" captionTargetPageId="23" captionText="FIGURES 5964. Forefemur (59, 60, 61, 63) and setae on the transverse row (62, 64) of Teloganodes spp. Insertion of the setae on the margins indicated by dots; only some setae drawn. Fig. 59: T. tuberculatus; Fig. 60: T. kodai; Figs. 61 62: T. jacobusi; Figs. 6364: T. hubbardi. Femora at same scale; setae at same scale." figureDoi="http://doi.org/10.5281/zenodo.5241169" httpUri="https://zenodo.org/record/5241169/files/figure.png" pageId="13" pageNumber="14">Fig. 59</figureCitation>
) moderately slender, ca. 2.2 times longer than wide; outer margin covered by thin and long setae together with 67 sharp, thick and pointed setae; submarginal row of short and blunt setae almost absent in foreleg, but distinct on middle and hind femora; inner margin with a row of long and thin setae, the row continuing on dorsal face, close to articulation with trochanter. Tibia with a row of long and thin setae, starting on proximal part close to outer margin and ending in distal part close to inner margin. Tarsal claw hooked, bearing 3 blunt teeth medially, and 2 pointed teeth subapically; apex of claw with two rows of small setae laterally.
</paragraph>
<caption id="DF4C09F54406FFFECE8E7209FC73FB9E" ID-DOI="http://doi.org/10.5281/zenodo.5241149" ID-Zenodo-Dep="5241149" box="[151,976,1083,1107]" httpUri="https://zenodo.org/record/5241149/files/figure.png" pageId="14" pageNumber="15" startId="14.[151,269,1083,1107]" targetBox="[183,1409,343,1047]" targetPageId="14">
<paragraph id="8B8C597D4406FFFECE8E7209FC73FB9E" blockId="14.[151,976,1083,1107]" box="[151,976,1083,1107]" pageId="14" pageNumber="15">
<emphasis id="B947856F4406FFFECE8E7209FEFEFB9E" bold="true" box="[151,349,1083,1107]" pageId="14" pageNumber="15">FIGURES 1415.</emphasis>
<taxonomicName id="4C3322FE4406FFFECF7D720EFD8AFB9E" box="[356,553,1084,1107]" class="Insecta" family="Teloganodidae" genus="Derlethina" kingdom="Animalia" order="Ephemeroptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="eloisae">
<emphasis id="B947856F4406FFFECF7D720EFD8AFB9E" box="[356,553,1084,1107]" italics="true" pageId="14" pageNumber="15">Derlethina eloisae</emphasis>
</taxonomicName>
. Fig. 14: Forewing; Fig. 15: Hindwing.
</paragraph>
</caption>
<paragraph id="8B8C597D4406FFFECEDC72B5FED0FA5C" blockId="14.[151,1437,1159,1665]" pageId="14" pageNumber="15">
All abdominal terga with well developed median tubercle, slightly increasing in size posteriorly (
<figureCitation id="130845F84406FFFECB7572B5FF64FB04" captionStart="FIGURES 120125" captionStartId="33.[151,269,1811,1835]" captionTargetBox="[151,1436,197,1784]" captionTargetId="figure-49@33.[151,1436,197,1784]" captionTargetPageId="33" captionText="FIGURES 120125. Holotypes of Teloganodes spp. in dorsal view (120, 123), abdomen in ventral view (121, 124) and in lateral view (122, 125). Figs. 120122: T. tuberculatus; Figs. 123125: T. kodai." figureDoi="http://doi.org/10.5281/zenodo.5241188" httpUri="https://zenodo.org/record/5241188/files/figure.png" pageId="14" pageNumber="15">Fig. 122</figureCitation>
). No posterolateral projections on segments IIVII, slightly marked on segments VIII and IX (
<figureCitation id="130845F84406FFFECB3E729DFA2DFB04" box="[1319,1422,1199,1225]" captionStart="FIGURES 120125" captionStartId="33.[151,269,1811,1835]" captionTargetBox="[151,1436,197,1784]" captionTargetId="figure-49@33.[151,1436,197,1784]" captionTargetPageId="33" captionText="FIGURES 120125. Holotypes of Teloganodes spp. in dorsal view (120, 123), abdomen in ventral view (121, 124) and in lateral view (122, 125). Figs. 120122: T. tuberculatus; Figs. 123125: T. kodai." figureDoi="http://doi.org/10.5281/zenodo.5241188" httpUri="https://zenodo.org/record/5241188/files/figure.png" pageId="14" pageNumber="15">Fig. 121</figureCitation>
). Lateral margins of terga with short and thin setae. Gills (Figs. 8286) on abdominal segments IIVI. Gill II with dorsal lamella operculate and covering others, oval with margin entire; gills IIIV with dorsal lamella incised medially; gill VI with dorsal lamella reduced and entire; ventral lobe flabellate, well developed and purple on gills IIV. Cerci with a whorl of spines every 23 segments; spines shorter than length of corresponding segment.
</paragraph>
<paragraph id="8B8C597D4406FFFECEDF73ADFE46FA74" blockId="14.[151,1437,1159,1665]" box="[198,485,1439,1465]" pageId="14" pageNumber="15">Winged stages unknown</paragraph>
</subSubSection>
<subSubSection id="C3290AF64406FFFECEDF73F4FEBEF9FC" pageId="14" pageNumber="15" type="diagnosis">
<paragraph id="8B8C597D4406FFFECEDF73F4FEBEF9FC" blockId="14.[151,1437,1159,1665]" pageId="14" pageNumber="15">
<emphasis id="B947856F4406FFFECEDF73F4FEE5FA2D" bold="true" box="[198,326,1478,1504]" pageId="14" pageNumber="15">Diagnosis.</emphasis>
The nymph of
<taxonomicName id="4C3322FE4406FFFECFE573F5FD0AFA2D" authorityName="Sartori &amp; Peters &amp; Hubbard" authorityYear="2008" box="[508,681,1479,1504]" class="Insecta" family="Teloganodidae" genus="Teloganodes" kingdom="Animalia" order="Ephemeroptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="tuberculatus">
<emphasis id="B947856F4406FFFECFE573F5FD0AFA2D" box="[508,681,1479,1504]" italics="true" pageId="14" pageNumber="15">T. tuberculatus</emphasis>
</taxonomicName>
can be easily distinguished from that of
<taxonomicName id="4C3322FE4406FFFECA9073F5FAA2FA2D" authorityName="Wang &amp; McCafferty" authorityYear="1996" box="[1161,1281,1479,1504]" class="Insecta" family="Teloganodidae" genus="Macafertiella" kingdom="Animalia" order="Ephemeroptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="insignis">
<emphasis id="B947856F4406FFFECA9073F5FAA2FA2D" box="[1161,1281,1479,1504]" italics="true" pageId="14" pageNumber="15">T. insignis</emphasis>
</taxonomicName>
by the shape of the labrum, the number of setae on the dorsal margin of the mandibles and the shape and ornamentation of the femora.
</paragraph>
</subSubSection>
<subSubSection id="C3290AF64406FFFECEDF700CFE14F94C" pageId="14" pageNumber="15" type="etymology">
<paragraph id="8B8C597D4406FFFECEDF700CFE14F94C" blockId="14.[151,1437,1159,1665]" pageId="14" pageNumber="15">
<emphasis id="B947856F4406FFFECEDF700CFEF0F995" bold="true" box="[198,339,1598,1624]" pageId="14" pageNumber="15">Etymology.</emphasis>
The epithet “tuberculatus” refers to the well developed tubercles found on the thorax and abdomen of this species.
</paragraph>
</subSubSection>
</treatment>
</document>