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<mods:title id="25DEA343471B40E35A3EF08E5FF950F9">The taxonomy of Indian gorgonians: an assessment of the descriptive records of gorgonians (Anthozoa: Octocorallia: Alcyonacea) recorded as occurring in the territorial waters of India, along with neighbouring regions and the Andaman and Nicobar Islands, and the highlighting of perceived unethical practice</mods:title>
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<paragraph id="8B9E07577B27FF9DFF56FA9BFDCEFAB3" blockId="49.[151,560,1378,1405]" box="[151,560,1378,1405]" pageId="49" pageNumber="50">
<heading id="D0D6B03B7B27FF9DFF56FA9BFDCEFAB3" bold="true" box="[151,560,1378,1405]" fontSize="11" level="2" pageId="49" pageNumber="50" reason="4">
<taxonomicName id="4C217CD47B27FF9DFF56FA9BFDCEFAB3" ID-CoL="L5V4" authority="Whitelegge, 1897" authorityName="Whitelegge" authorityYear="1897" box="[151,560,1378,1405]" class="Anthozoa" family="Plexauridae" genus="Bebryce" kingdom="Animalia" order="Alcyonacea" pageId="49" pageNumber="50" phylum="Cnidaria" rank="species" species="studeri">
<emphasis id="B955DB457B27FF9DFF56FA9BFDCEFAB3" bold="true" box="[151,560,1378,1405]" pageId="49" pageNumber="50">
<emphasis id="B955DB457B27FF9DFF56FA9BFEADFAB2" bold="true" box="[151,339,1378,1405]" italics="true" pageId="49" pageNumber="50">Bebryce studeri</emphasis>
<bibRefCitation id="EFB07AA67B27FF9DFE9AFA9BFDCEFAB3" author="Whitelegge, T." box="[347,560,1378,1404]" pageId="49" pageNumber="50" pagination="307 - 320" refId="ref110659" refString="Whitelegge, T. (1897) The Alcyonaria of Funafuti. Part II. Memoirs of the Australian Museum, 3 (5), 307 - 320. https: // doi. org / 10.3853 / j. 0067 - 1967.3.1897.498" type="journal article" year="1897">Whitelegge, 1897</bibRefCitation>
</emphasis>
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</paragraph>
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<treatmentCitation id="0A8021467B27FF9DFF56FA5EFDCCFA70" author="Whitelegge, T." box="[151,562,1447,1471]" page="314" pageId="49" pageNumber="50" year="1897">
<taxonomicName id="4C217CD47B27FF9DFF56FA5EFDCCFA70" ID-CoL="L5V4" authority="Whitelegge, 1897: 314" authorityName="Whitelegge" authorityPageNumber="314" authorityYear="1897" box="[151,562,1447,1471]" class="Anthozoa" family="Plexauridae" genus="Bebryce" kingdom="Animalia" order="Alcyonacea" pageId="49" pageNumber="50" phylum="Cnidaria" rank="species" species="studeri">
<emphasis id="B955DB457B27FF9DFF56FA5EFEC7FA70" box="[151,313,1447,1471]" italics="true" pageId="49" pageNumber="50">Bebryce studeri</emphasis>
<bibRefCitation id="EFB07AA67B27FF9DFE82FA5EFDCCFA70" author="Whitelegge, T." box="[323,562,1447,1471]" pageId="49" pageNumber="50" pagination="307 - 320" refId="ref110659" refString="Whitelegge, T. (1897) The Alcyonaria of Funafuti. Part II. Memoirs of the Australian Museum, 3 (5), 307 - 320. https: // doi. org / 10.3853 / j. 0067 - 1967.3.1897.498" type="journal article" year="1897">Whitelegge, 1897: 314</bibRefCitation>
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, pl. 17, fig. 2125 (Funafuti):
<treatmentCitation id="0A8021467B27FF9DFCB7FA5EFBBEFA70" author="Grasshoff, M." box="[886,1088,1447,1471]" page="63" pageId="49" pageNumber="50" year="1999">
<bibRefCitation id="EFB07AA67B27FF9DFCB7FA5EFBBEFA70" author="Grasshoff, M." box="[886,1088,1447,1471]" pageId="49" pageNumber="50" pagination="1 - 121" refId="ref101429" refString="Grasshoff, M. (1999) The shallow water gorgonians of New Caledonia and adjacent islands (Coelenterata: Octocorallia). Senckenbergiana biologica, 78 (1 / 2), 1 - 121." type="journal article" year="1999">Grasshoff 1999: 63</bibRefCitation>
</treatmentCitation>
, figs. 106108 (
<collectingCountry id="F33647C77B27FF9DFB2DFA5EFA6FFA70" box="[1260,1425,1447,1471]" name="New Caledonia" pageId="49" pageNumber="50">New Caledonia</collectingCountry>
);
<treatmentCitation id="0A8021467B27FF9DFF06FA3FFDE6FA11" author="Bayer, F. M. &amp; Ofwegen, L. V." box="[199,536,1478,1502]" page="346" pageId="49" pageNumber="50" year="2016">
<bibRefCitation id="EFB07AA67B27FF9DFF06FA3FFDE6FA11" author="Bayer, F. M. &amp; Ofwegen, L. V." box="[199,536,1478,1502]" pageId="49" pageNumber="50" pagination="301 - 358" refId="ref99006" refString="Bayer, F. M. &amp; Ofwegen, L. V. (2016) The type specimens of Bebryce (Cnidaria, Octocorallia, Plexauridae) re-examined, with emphasis on the sclerites. Zootaxa, 4083 (3), 301 - 358. https: // doi. org / 10.11646 / zootaxa. 4083.3.1" type="journal article" year="2016">Bayer &amp; van Ofwegen 2016: 346</bibRefCitation>
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(
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description).
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</paragraph>
<paragraph id="8B9E07577B27FF9DFF56F9F5FC0EF9E9" blockId="49.[151,1008,1548,1575]" box="[151,1008,1548,1575]" pageId="49" pageNumber="50">Opinion: There is not enough evidence that this species occurs in the region.</paragraph>
</subSubSection>
<subSubSection id="C33B54DC7B27FF9EFF56F9ADFE15FEAA" lastPageId="50" lastPageNumber="51" pageId="49" pageNumber="50" type="description">
<paragraph id="8B9E07577B27FF9DFF56F9ADFED7F9A1" blockId="49.[151,1436,1620,1755]" box="[151,297,1620,1646]" pageId="49" pageNumber="50">Justification:</paragraph>
<paragraph id="8B9E07577B27FF9DFF06F981FD93F915" blockId="49.[151,1436,1620,1755]" pageId="49" pageNumber="50">
<emphasis id="B955DB457B27FF9DFF06F981FC64F95D" box="[199,922,1656,1682]" italics="true" pageId="49" pageNumber="50">These Indian records seem to be either invalid or unconfirmable</emphasis>
:
<bibRefCitation id="EFB07AA67B27FF9DFC68F981FB4EF95D" author="Fernando, S. A." box="[937,1200,1656,1682]" pageId="49" pageNumber="50" refId="ref100707" refString="Fernando, S. A. (2011) Monograph on Gorgonids (Sea fans) of India. Centre of Advanced Study in Marine Biology, Faculty of Marine Sciences, Parangipettai, 145 pp." type="book" year="2011">Fernando, 2011: 4243</bibRefCitation>
, pl. 19, fig. 11f (SE coast of
<collectingCountry id="F33647C77B27FF9DFF37F965FECAF979" box="[246,308,1692,1718]" name="India" pageId="49" pageNumber="50">India</collectingCountry>
);
<bibRefCitation id="EFB07AA67B27FF9DFE89F964FDB0F979" author="Kumar, J. S. Y. &amp; Raghunathan, C. &amp; Raghuraman, R. &amp; Sreeraj, C. R. &amp; Venkataraman, K." box="[328,590,1692,1719]" pageId="49" pageNumber="50" refId="ref103651" refString="Kumar, J. S. Y., Raghunathan, C., Raghuraman, R., Sreeraj, C. R. &amp; Venkataraman, K. (2014 a) Handbook on Gorgonians (Octocorallia) of Andaman and Nicobar Islands. Director, Zoological Survey of India, Kolkata, 119 pp." type="book" year="2014">
Kumar
<emphasis id="B955DB457B27FF9DFE5CF964FE30F979" box="[413,462,1692,1718]" italics="true" pageId="49" pageNumber="50">et al</emphasis>
. 2014a: 72
</bibRefCitation>
, pl. 33, fig. AC (Havelock);
<bibRefCitation id="EFB07AA67B27FF9DFC5DF965FB4FF979" author="Fernando, S. &amp; Venkataraman, K. &amp; Raghunathan, C." box="[924,1201,1692,1719]" pageId="49" pageNumber="50" refId="ref100744" refString="Fernando, S. A, Venkataraman, K. &amp; Raghunathan, C. (2017) Gorgonians of Indian sea. Director, Zoological Survey of India, Kolkata, 330 pp. Available from: https: // www. nhbs. com / (accessed 21 March 2022)" type="book" year="2017">
Fernando
<emphasis id="B955DB457B27FF9DFBCCF964FBC0F979" box="[1037,1086,1692,1718]" italics="true" pageId="49" pageNumber="50">et al</emphasis>
. 2017: 87
</bibRefCitation>
, pl. 38, fig. AC (SE coast of
<collectingCountry id="F33647C77B27FF9DFF37F939FECDF915" box="[246,307,1728,1754]" name="India" pageId="49" pageNumber="50">India</collectingCountry>
; Andaman and Nicobar Is.).
</paragraph>
<paragraph id="8B9E07577B27FF9EFF06F8FAFE15FEAA" blockId="49.[151,1436,1795,2037]" lastBlockId="50.[151,1437,151,357]" lastPageId="50" lastPageNumber="51" pageId="49" pageNumber="50">
<emphasis id="B955DB457B27FF9DFF06F8FAFE65F8D2" box="[199,411,1795,1821]" italics="true" pageId="49" pageNumber="50">Literature analysis</emphasis>
: In the account of the species given by
<bibRefCitation id="EFB07AA67B27FF9DFC96F8FAFBEFF8D2" author="Fernando, S. A." box="[855,1041,1795,1821]" pageId="49" pageNumber="50" refId="ref100707" refString="Fernando, S. A. (2011) Monograph on Gorgonids (Sea fans) of India. Centre of Advanced Study in Marine Biology, Faculty of Marine Sciences, Parangipettai, 145 pp." type="book" year="2011">Fernando (2011)</bibRefCitation>
and
<bibRefCitation id="EFB07AA67B27FF9DFB89F8FAFABCF8D2" author="Fernando, S. &amp; Venkataraman, K. &amp; Raghunathan, C." box="[1096,1346,1795,1821]" pageId="49" pageNumber="50" refId="ref100744" refString="Fernando, S. A, Venkataraman, K. &amp; Raghunathan, C. (2017) Gorgonians of Indian sea. Director, Zoological Survey of India, Kolkata, 330 pp. Available from: https: // www. nhbs. com / (accessed 21 March 2022)" type="book" year="2017">
Fernando
<emphasis id="B955DB457B27FF9DFB79F8FAFB14F8D2" box="[1208,1258,1795,1821]" italics="true" pageId="49" pageNumber="50">et al</emphasis>
. (2017)
</bibRefCitation>
the text is identical, but the illustrations are different and all but one of the specimens examined (ZSI/ANC: 6064) are the same. However, in the account of the species by
<bibRefCitation id="EFB07AA67B27FF9DFD70F8B2FC67F8AA" author="Kumar, J. S. Y. &amp; Raghunathan, C. &amp; Raghuraman, R. &amp; Sreeraj, C. R. &amp; Venkataraman, K." box="[689,921,1867,1893]" pageId="49" pageNumber="50" refId="ref103651" refString="Kumar, J. S. Y., Raghunathan, C., Raghuraman, R., Sreeraj, C. R. &amp; Venkataraman, K. (2014 a) Handbook on Gorgonians (Octocorallia) of Andaman and Nicobar Islands. Director, Zoological Survey of India, Kolkata, 119 pp." type="book" year="2014">
Kumar
<emphasis id="B955DB457B27FF9DFCC5F8B2FCC2F8AA" box="[772,828,1867,1893]" italics="true" pageId="49" pageNumber="50">et al.</emphasis>
(2014a)
</bibRefCitation>
and
<bibRefCitation id="EFB07AA67B27FF9DFC0CF8B2FB3BF8AA" author="Fernando, S. &amp; Venkataraman, K. &amp; Raghunathan, C." box="[973,1221,1867,1893]" pageId="49" pageNumber="50" refId="ref100744" refString="Fernando, S. A, Venkataraman, K. &amp; Raghunathan, C. (2017) Gorgonians of Indian sea. Director, Zoological Survey of India, Kolkata, 330 pp. Available from: https: // www. nhbs. com / (accessed 21 March 2022)" type="book" year="2017">
Fernando
<emphasis id="B955DB457B27FF9DFBFCF8B2FB90F8AA" box="[1085,1134,1867,1893]" italics="true" pageId="49" pageNumber="50">et al</emphasis>
. (2017)
</bibRefCitation>
the text is different but the illustrations are identical and must be of ZSI/ANC: 6064, which is the only specimen featured by
<bibRefCitation id="EFB07AA67B27FF9DFAF2F896FEE9F862" author="Kumar, J. S. Y. &amp; Raghunathan, C. &amp; Raghuraman, R. &amp; Sreeraj, C. R. &amp; Venkataraman, K." pageId="49" pageNumber="50" refId="ref103651" refString="Kumar, J. S. Y., Raghunathan, C., Raghuraman, R., Sreeraj, C. R. &amp; Venkataraman, K. (2014 a) Handbook on Gorgonians (Octocorallia) of Andaman and Nicobar Islands. Director, Zoological Survey of India, Kolkata, 119 pp." type="book" year="2014">
Kumar
<emphasis id="B955DB457B27FF9DFA49F896FF53F862" italics="true" pageId="49" pageNumber="50">et al</emphasis>
. (2014a)
</bibRefCitation>
. The few sclerites illustrated by
<bibRefCitation id="EFB07AA67B27FF9DFD48F86AFC87F862" author="Kumar, J. S. Y. &amp; Raghunathan, C. &amp; Raghuraman, R. &amp; Sreeraj, C. R. &amp; Venkataraman, K." box="[649,889,1939,1965]" pageId="49" pageNumber="50" refId="ref103651" refString="Kumar, J. S. Y., Raghunathan, C., Raghuraman, R., Sreeraj, C. R. &amp; Venkataraman, K. (2014 a) Handbook on Gorgonians (Octocorallia) of Andaman and Nicobar Islands. Director, Zoological Survey of India, Kolkata, 119 pp." type="book" year="2014">
Kumar
<emphasis id="B955DB457B27FF9DFD21F86AFCE7F862" box="[736,793,1939,1965]" italics="true" pageId="49" pageNumber="50">et al.</emphasis>
(2014a)
</bibRefCitation>
are very different from those of the
<typeStatus id="549AB9F57B27FF9DFADAF86AFA83F862" box="[1307,1405,1939,1965]" pageId="49" pageNumber="50" type="holotype">holotype</typeStatus>
as figured by
<bibRefCitation id="EFB07AA67B27FF9DFED1F84EFDD1F81E" author="Bayer, F. M. &amp; Ofwegen, L. V." box="[272,559,1975,2001]" pageId="49" pageNumber="50" pagination="301 - 358" refId="ref99006" refString="Bayer, F. M. &amp; Ofwegen, L. V. (2016) The type specimens of Bebryce (Cnidaria, Octocorallia, Plexauridae) re-examined, with emphasis on the sclerites. Zootaxa, 4083 (3), 301 - 358. https: // doi. org / 10.11646 / zootaxa. 4083.3.1" type="journal article" year="2016">Bayer &amp; Ofwegen (2016)</bibRefCitation>
, but the true nature of the Indian specimen is unknown as the description of the sclerites by Kumar
<emphasis id="B955DB457B27FF9DFEB2F822FE55F83A" box="[371,427,2011,2037]" italics="true" pageId="49" pageNumber="50">et al.</emphasis>
(apart from one line) is taken word-for-word from
<bibRefCitation id="EFB07AA67B27FF9DFC2DF822FB50F83A" author="Grasshoff, M." box="[1004,1198,2011,2037]" pageId="49" pageNumber="50" pagination="1 - 121" refId="ref101429" refString="Grasshoff, M. (1999) The shallow water gorgonians of New Caledonia and adjacent islands (Coelenterata: Octocorallia). Senckenbergiana biologica, 78 (1 / 2), 1 - 121." type="journal article" year="1999">Grasshoffs 1999</bibRefCitation>
text. The description given by
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Fernando
<emphasis id="B955DB457B24FF9EFEB1FF6EFE5CFF7E" box="[368,418,151,177]" italics="true" pageId="50" pageNumber="51">et al</emphasis>
. (2017)
</bibRefCitation>
clearly refers to the illustrations in
<bibRefCitation id="EFB07AA67B24FF9EFC48FF6EFBB8FF7E" author="Fernando, S. A." box="[905,1094,151,177]" pageId="50" pageNumber="51" refId="ref100707" refString="Fernando, S. A. (2011) Monograph on Gorgonids (Sea fans) of India. Centre of Advanced Study in Marine Biology, Faculty of Marine Sciences, Parangipettai, 145 pp." type="book" year="2011">Fernando (2011)</bibRefCitation>
, but the sclerite images are far too small and far too crowded to be able to see sufficient detail in order to recognise the species. However, in plate 19, fig. 1e of
<bibRefCitation id="EFB07AA67B24FF9EFEEAFF26FE19FF36" author="Fernando, S. A." box="[299,487,223,249]" pageId="50" pageNumber="51" refId="ref100707" refString="Fernando, S. A. (2011) Monograph on Gorgonids (Sea fans) of India. Centre of Advanced Study in Marine Biology, Faculty of Marine Sciences, Parangipettai, 145 pp." type="book" year="2011">Fernando (2011)</bibRefCitation>
, the large collaret sclerite is quite unlike those of the
<typeStatus id="549AB9F57B24FF9EFBF4FF26FB69FF36" box="[1077,1175,223,249]" pageId="50" pageNumber="51" type="holotype">holotype</typeStatus>
(see
<bibRefCitation id="EFB07AA67B24FF9EFB11FF26FF2DFED2" author="Bayer, F. M. &amp; Ofwegen, L. V." pageId="50" pageNumber="51" pagination="301 - 358" refId="ref99006" refString="Bayer, F. M. &amp; Ofwegen, L. V. (2016) The type specimens of Bebryce (Cnidaria, Octocorallia, Plexauridae) re-examined, with emphasis on the sclerites. Zootaxa, 4083 (3), 301 - 358. https: // doi. org / 10.11646 / zootaxa. 4083.3.1" type="journal article" year="2016">Bayer &amp; Ofwegen 2016</bibRefCitation>
), and there are warty ovals with a waist, a little like those in
<taxonomicName id="4C217CD47B24FF9EFCBFFEFAFB72FED2" authority="Bayer, 1981" authorityName="Bayer" authorityYear="1981" box="[894,1164,259,285]" class="Anthozoa" family="Plexauridae" genus="Bebryce" kingdom="Animalia" order="Alcyonacea" pageId="50" pageNumber="51" phylum="Cnidaria" rank="species" species="crucifera">
<emphasis id="B955DB457B24FF9EFCBFFEFAFC03FED2" box="[894,1021,259,285]" italics="true" pageId="50" pageNumber="51">B crucifera</emphasis>
<bibRefCitation id="EFB07AA67B24FF9EFBC5FEFAFB72FED2" author="Bayer, F. M." box="[1028,1164,259,285]" pageId="50" pageNumber="51" pagination="3 - 11" refId="ref98812" refString="Bayer, F. M. (1981) Status of knowledge of octocorals of world seas. Seminarios de Biologia Marinha, Academia Brasileira de Ciencias, 1981, 3 - 11." type="journal article" year="1981">Bayer, 1981</bibRefCitation>
</taxonomicName>
(see
<bibRefCitation id="EFB07AA67B24FF9EFB06FEFAFF2DFE8E" author="Bayer, F. M. &amp; Ofwegen, L. V." pageId="50" pageNumber="51" pagination="301 - 358" refId="ref99006" refString="Bayer, F. M. &amp; Ofwegen, L. V. (2016) The type specimens of Bebryce (Cnidaria, Octocorallia, Plexauridae) re-examined, with emphasis on the sclerites. Zootaxa, 4083 (3), 301 - 358. https: // doi. org / 10.11646 / zootaxa. 4083.3.1" type="journal article" year="2016">Bayer &amp; Ofwegen, 2016</bibRefCitation>
: fig. 11d), that do not occur at all in
<taxonomicName id="4C217CD47B24FF9EFDA3FEDEFD30FE8E" authorityName="Whitelegge" authorityYear="1897" box="[610,718,295,321]" class="Anthozoa" family="Plexauridae" genus="Bebryce" kingdom="Animalia" order="Alcyonacea" pageId="50" pageNumber="51" phylum="Cnidaria" rank="species" species="studeri">
<emphasis id="B955DB457B24FF9EFDA3FEDEFD30FE8E" box="[610,718,295,321]" italics="true" pageId="50" pageNumber="51">B. studeri</emphasis>
</taxonomicName>
.
<bibRefCitation id="EFB07AA67B24FF9EFD1AFEDEFC49FE8E" author="Kumar, J. S. Y. &amp; Raghunathan, C. &amp; Venkataraman, K." box="[731,951,294,321]" pageId="50" pageNumber="51" pagination="15 - 33" refId="ref103709" refString="Kumar, J. S. Y., Raghunathan, C. &amp; Venkataraman, K. (2015) Abundance of shallow water octocorals in the Andaman and Nicobar Archipelago, India. In: Venkataraman, K. &amp; Sivaperuman, C. (Eds.), Marine faunal diversity in India: taxonomy, ecology and conservation. London Academic Press, London, pp. 15 - 33. https: // doi. org / 10.1016 / B 978 - 0 - 12 - 801948 - 1.00002 - 1" type="book chapter" year="2015">
Kumar
<emphasis id="B955DB457B24FF9EFCEEFEDEFC9EFE8E" box="[815,864,295,321]" italics="true" pageId="50" pageNumber="51">et al</emphasis>
. (2015)
</bibRefCitation>
just lists the species and presents the image of the ZSI/ANC: 6064 colony.
</paragraph>
</subSubSection>
</treatment>
</document>