treatments-xml/data/C0/74/F0/C074F028A4EB58D6BCEB22EFF5A1CD76.xml

<document ID-DOI="http://dx.doi.org/10.3897/asp.80.e84013" ID-Pensoft-Pub="1864-8312-80-575" ID-Pensoft-UUID="81B59BA932BC5DAF90815FD08F5820A6" ID-ZooBank="4A6669D57DDD4AE0A8B455724B9C594A" ModsDocID="1864-8312-80-575" checkinTime="1668813875820" checkinUser="pensoft" docAuthor="Linsky, Marek, Ciamporova-Zatovicova, Zuzana, Lassova, Kristina &amp; Ciampor Jr, Fedor" docDate="2022" docId="C074F028A4EB58D6BCEB22EFF5A1CD76" docLanguage="en" docName="ArthropodSystPhyl 80: 575-602" docOrigin="Arthropod Systematics &amp; amp; Phylogeny 80" docPubDate="2022-11-18" docSource="http://dx.doi.org/10.3897/asp.80.e84013" docTitle="Rumilara leleupi Linský &amp; Čiamporová-Zaťovičová &amp; Laššová &amp; Čiampor Jr 2022, comb. nov." docType="treatment" docVersion="1" id="81B59BA932BC5DAF90815FD08F5820A6" lastPageNumber="575" masterDocId="81B59BA932BC5DAF90815FD08F5820A6" masterDocTitle="Molecular phylogeny of the riffle beetle genus Hexanchorus revealed a presence of a new genus (Coleoptera: Elmidae)" masterLastPageNumber="602" masterPageNumber="575" pageNumber="575" updateTime="1668813875820" updateUser="pensoft">
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<mods:titleInfo>
<mods:title>Molecular phylogeny of the riffle beetle genus Hexanchorus revealed a presence of a new genus (Coleoptera: Elmidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Linsky, Marek</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-7206-2380</mods:nameIdentifier>
<mods:affiliation>Plant Science and Biodiversity Centre, Slovak Academy of Sciences, Du ́ bravska ́ cesta 9, SK- 84523, Bratislava, Slovakia &amp; Department of Zoology, Faculty of Natural Sciences, Comenius University, Mlynska ́ dolina B- 1, SK- 84215 Bratislava, Slovakia</mods:affiliation>
</mods:name>
<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ciamporova-Zatovicova, Zuzana</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-0506-6212</mods:nameIdentifier>
<mods:affiliation>Plant Science and Biodiversity Centre, Slovak Academy of Sciences, Du ́ bravska ́ cesta 9, SK- 84523, Bratislava, Slovakia &amp; Department of Zoology, Faculty of Natural Sciences, Comenius University, Mlynska ́ dolina B- 1, SK- 84215 Bratislava, Slovakia</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Lassova, Kristina</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-7175-2109</mods:nameIdentifier>
<mods:affiliation>Plant Science and Biodiversity Centre, Slovak Academy of Sciences, Du ́ bravska ́ cesta 9, SK- 84523, Bratislava, Slovakia</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ciampor Jr, Fedor</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-6269-3592</mods:nameIdentifier>
<mods:affiliation>Plant Science and Biodiversity Centre, Slovak Academy of Sciences, Du ́ bravska ́ cesta 9, SK- 84523, Bratislava, Slovakia &amp; Department of Zoology, Faculty of Natural Sciences, Comenius University, Mlynska ́ dolina B- 1, SK- 84215 Bratislava, Slovakia</mods:affiliation>
<mods:nameIdentifier type="email">f.ciampor@savba.sk</mods:nameIdentifier>
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<mods:title>Arthropod Systematics &amp; amp; Phylogeny</mods:title>
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<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-11-18</mods:number>
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<mods:detail type="volume">
<mods:number>80</mods:number>
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<mods:start>575</mods:start>
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<mods:url>http://dx.doi.org/10.3897/asp.80.e84013</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/asp.80.e84013</mods:identifier>
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<subSection pageId="0" pageNumber="575" type="3. results">
<treatment LSID="urn:lsid:plazi:treatment:C074F028A4EB58D6BCEB22EFF5A1CD76" httpUri="http://treatment.plazi.org/id/C074F028A4EB58D6BCEB22EFF5A1CD76" lastPageNumber="575" pageId="0" pageNumber="575">
<subSubSection pageId="0" pageNumber="575" type="nomenclature">
<paragraph pageId="0" pageNumber="575">
3.2.7. 
<taxonomicName LSID="C074F028-A4EB-58D6-BCEB-22EFF5A1CD76" authority="(Deleve, 1968)" authorityName="Linský &amp; Čiamporová-Zaťovičová &amp; Laššová &amp; Čiampor Jr" authorityYear="2022" baseAuthorityName="Deleve" baseAuthorityYear="1968" family="Elmidae" genus="Rumilara" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Rumilara leleupi" order="Coleoptera" pageId="0" pageNumber="575" rank="species" species="leleupi" status="comb. nov.">
Rumilara leleupi (
<normalizedToken originalValue="Delève">Deleve</normalizedToken>
, 1968)
</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="575">comb. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="575" type="description">
<paragraph pageId="0" pageNumber="575">
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dorsal habiti of Rumilara gen. nov. species: a R. obscura sp. nov. male holotype; b R. suppressa sp. nov. male holotype; c R. riberai sp. nov. male holotype; d R. paterna sp. nov. male holotype; e R. leleupi comb. nov. male holotype. Scale: 0.5 mm." figureDoi="10.3897/asp.80.e84013.figure2" httpUri="https://binary.pensoft.net/fig/770425" pageId="0" pageNumber="575">Figs 2e</figureCitation>
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Lateral habiti of Rumilara gen. nov. species. a R. obscura sp. nov. male holotype; b R. suppressa sp. nov. male holotype; c R. riberai sp. nov. male holotype; d R. paterna sp. nov. male holotype; e R. leleupi comb. nov. male holotype. Scale: 0.5 mm." figureDoi="10.3897/asp.80.e84013.figure3" httpUri="https://binary.pensoft.net/fig/770426" pageId="0" pageNumber="575">, 3e</figureCitation>
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Antennae of Rumilara gen. nov. and Hexanchorus: a R. obscura sp. nov.; b R. suppressa sp. nov.; c R. riberai sp. nov.; d R. paterna sp. nov.; e R. leleupi comb. nov.; f H. mcdiarmidi; g H. virilis; h H. shannoni; i H. angeli; j H. rostratus; k H. cordillierae; l H. onorei; m H. tarsalis; n H. usitatus; o H. gracilipes; p H. caraibus; q H. crinitus; r H. dimorphus; s H. emarginatus; t H. browni. (a, k based on dissected antennae; b-j, l-r based on photographs, not to scale, setae omitted)." figureDoi="10.3897/asp.80.e84013.figure5" httpUri="https://binary.pensoft.net/fig/770428" pageId="0" pageNumber="575">, 5e</figureCitation>
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Pronota of Rumilara gen. nov. species: a R. obscura sp. nov. male holotype; b R. suppressa sp. nov. male holotype; c R. riberai sp. nov. male holotype; d R. paterna sp. nov. male holotype; e R. leleupi comb. nov. male holotype. Scale: 0.2 mm." figureDoi="10.3897/asp.80.e84013.figure6" httpUri="https://binary.pensoft.net/fig/770429" pageId="0" pageNumber="575">, 6e</figureCitation>
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Ventral habiti of Rumilara gen. nov. species: a R. obscura sp. nov. male paratype; b R. suppressa sp. nov. male paratype; c R. riberai sp. nov. female paratype; d R. paterna sp. nov. paratype; e R. leleupi comb. nov. male holotype. Scale: 0.5 mm." figureDoi="10.3897/asp.80.e84013.figure7" httpUri="https://binary.pensoft.net/fig/770430" pageId="0" pageNumber="575">, 7e</figureCitation>
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Abdomens of Rumilara gen. nov. species: a R. obscura sp. nov. male holotype; b R. suppressa sp. nov. male holotype; c R. riberai sp. nov. male holotype; d R. paterna sp. nov. male holotype; e R. leleupi comb. nov. male holotype. 0.2 mm." figureDoi="10.3897/asp.80.e84013.figure8" httpUri="https://binary.pensoft.net/fig/770431" pageId="0" pageNumber="575">, 8e</figureCitation>
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Rumilara leleupi comb. nov.: a aedeagus, left lateral view; b aedeagus, ventral view; c aedeagus, right lateral view; d male sternite VIII. Scale: 0.1 mm." figureDoi="10.3897/asp.80.e84013.figure15" httpUri="https://binary.pensoft.net/fig/770438" pageId="0" pageNumber="575">, 15</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="575" type="reference_group">
<paragraph pageId="0" pageNumber="575">
<taxonomicName authorityName="Linský &amp; Čiamporová-Zaťovičová &amp; Laššová &amp; Čiampor Jr" authorityYear="2022" baseAuthorityName="Linský &amp; Čiamporová-Zaťovičová &amp; Laššová &amp; Čiampor Jr" baseAuthorityYear="2022" class="Insecta" family="Elmidae" genus="Hexanchorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hexanchorus leleupi" order="Coleoptera" pageId="0" pageNumber="575" phylum="Arthropoda" rank="species" species="leleupi" status="comb. nov.">Hexanchorus leleupi</taxonomicName>
<normalizedToken originalValue="Delève">Deleve</normalizedToken>
, 1968: 214.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="575" type="type locality">
<paragraph pageId="0" pageNumber="575">Type locality.</paragraph>
<paragraph pageId="0" pageNumber="575">Ecuador, [Napo Province], Papallacta, 3.300m a.s.l.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="575" type="materials_examined">
<paragraph pageId="0" pageNumber="575">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="575">
<emphasis bold="true" italics="true" pageId="0" pageNumber="575">Type material</emphasis>
: 
<materialsCitation collectingDate="1965-03" collectorName="N. Leleup, R. I. Sc." country="Ecuador" determinerName="J. Deleve" elevation="3" location="Papallacta" specimenCount="♂" typeStatus="Holotype">
<typeStatus>Holotype</typeStatus>
<specimenCount>♂</specimenCount>
(RBINS) 
<normalizedToken originalValue="“♂”">
&quot; 
<specimenCount>♂</specimenCount>
&quot;
</normalizedToken>
// 
<normalizedToken originalValue="“Holotype”">
&quot; 
<typeStatus>Holotype</typeStatus>
&quot;
</normalizedToken>
[red] // &quot; 
<collectingCountry name="Ecuador">Ecuador</collectingCountry>
<location LSID="urn:lsid:plazi:treatment:C074F028A4EB58D6BCEB22EFF5A1CD76:5A76B9D7E5218230ADC6DEAE9829B562" country="Ecuador" name="Papallacta">Papallacta</location>
<collectingDate value="1965-03">III-1965</collectingDate>
. 
<collectorName>N. Leleup</collectorName>
&quot; // &quot;Ruisseau torrentueux 
<quantity metricMagnitude="0" metricUnit="m" metricValue="3.3" unit="m" value="3.3">
<elevation metricMagnitude="0" metricUnit="m" metricValue="3.3" unit="m" value="3.3">3.300 m</elevation>
</quantity>
&quot; // &quot; 
<collectorName>R. I. Sc.</collectorName>
N. B. I. G. 23.948&quot; // &quot; 
<determinerName>
J. 
<normalizedToken originalValue="Delève">Deleve</normalizedToken>
</determinerName>
det., 1966 
<taxonomicName authorityName="Linský &amp; Čiamporová-Zaťovičová &amp; Laššová &amp; Čiampor Jr" authorityYear="2022" baseAuthorityName="Linský &amp; Čiamporová-Zaťovičová &amp; Laššová &amp; Čiampor Jr" baseAuthorityYear="2022" class="Insecta" family="Elmidae" genus="Hexanchorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hexanchorus leleupi" order="Coleoptera" pageId="0" pageNumber="575" phylum="Arthropoda" rank="species" species="leleupi">
<emphasis italics="true" pageId="0" pageNumber="575">Hexanchorus leleupi</emphasis>
</taxonomicName>
n.sp.&quot; // &quot; 
<taxonomicName authorityName="Linský &amp; Čiamporová-Zaťovičová &amp; Laššová &amp; Čiampor Jr" authorityYear="2022" baseAuthorityName="Deleve" baseAuthorityYear="1968" family="Elmidae" genus="Rumilara" kingdom="Animalia" lsidName="Rumilara leleupi" order="Coleoptera" pageId="0" pageNumber="575" rank="species" species="leleupi">
<emphasis italics="true" pageId="0" pageNumber="575">Rumilara leleupi</emphasis>
</taxonomicName>
(
<normalizedToken originalValue="Delève">Deleve</normalizedToken>
, 1968)&quot; [appended].
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="575" type="condition">
<paragraph pageId="0" pageNumber="575">Condition.</paragraph>
<paragraph pageId="0" pageNumber="575">Generally, in good condition, except for following parts of legs that are missing: last segment of right protarsus, last two segments of left protarsus, left midleg (only coxa and trochanter present), and last two segments of left metatarsus.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="575" type="adult diagnosis">
<paragraph pageId="0" pageNumber="575">Adult diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="575">The species can be distinguished by a combination of following characters: 1) head and clypeus fully clothed by pubescence; 2) femora fully pubescent; 3) elytra with dense pubescence arranged in stripes; 4) aedeagus without rod-like sclerite; 5) parameres strongly asymmetrical.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="575" type="description">
<paragraph pageId="0" pageNumber="575">Adult diagnostic redescription.</paragraph>
<paragraph pageId="0" pageNumber="575">
Male. 
<emphasis bold="true" italics="true" pageId="0" pageNumber="575">Colour</emphasis>
: Body (Figs 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dorsal habiti of Rumilara gen. nov. species: a R. obscura sp. nov. male holotype; b R. suppressa sp. nov. male holotype; c R. riberai sp. nov. male holotype; d R. paterna sp. nov. male holotype; e R. leleupi comb. nov. male holotype. Scale: 0.5 mm." figureDoi="10.3897/asp.80.e84013.figure2" httpUri="https://binary.pensoft.net/fig/770425" pageId="0" pageNumber="575">2e</figureCitation>
, 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Lateral habiti of Rumilara gen. nov. species. a R. obscura sp. nov. male holotype; b R. suppressa sp. nov. male holotype; c R. riberai sp. nov. male holotype; d R. paterna sp. nov. male holotype; e R. leleupi comb. nov. male holotype. Scale: 0.5 mm." figureDoi="10.3897/asp.80.e84013.figure3" httpUri="https://binary.pensoft.net/fig/770426" pageId="0" pageNumber="575">3e</figureCitation>
) black, tarsal claws pale brown with reddish tincture. 
<emphasis bold="true" italics="true" pageId="0" pageNumber="575">Pubescence</emphasis>
: Head and clypeus fully clothed with short, prone pubescence, most densely set behind eyes. Pronotum (Fig. 
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Pronota of Rumilara gen. nov. species: a R. obscura sp. nov. male holotype; b R. suppressa sp. nov. male holotype; c R. riberai sp. nov. male holotype; d R. paterna sp. nov. male holotype; e R. leleupi comb. nov. male holotype. Scale: 0.2 mm." figureDoi="10.3897/asp.80.e84013.figure6" httpUri="https://binary.pensoft.net/fig/770429" pageId="0" pageNumber="575">6e</figureCitation>
) covered by a short, moderately dense, prone pubescence. Elytra with short, recumbent pubescence, most densely arranged in stripes along intervals, and with numerous moderately long, curved, semi-erect, hair-like setae. Femora, pro- and metatibiae densely covered with moderately long, recumbent, hair-like setae; mesotibiae without pubescence. Metaventrite readily clothed with short, prone pubescence. 
<emphasis bold="true" italics="true" pageId="0" pageNumber="575">Surface</emphasis>
: Head, clypeus and pronotum sparsely microreticulate, concealed by pubescence. Elytra with rows of punctures deeply impressed; third interval slightly elevated in anterior 1/5; intervals 2-4 depressed from anterior 1/5 to 2/5 in lateral view. Prosternal process (Fig. 
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Ventral habiti of Rumilara gen. nov. species: a R. obscura sp. nov. male paratype; b R. suppressa sp. nov. male paratype; c R. riberai sp. nov. female paratype; d R. paterna sp. nov. paratype; e R. leleupi comb. nov. male holotype. Scale: 0.5 mm." figureDoi="10.3897/asp.80.e84013.figure7" httpUri="https://binary.pensoft.net/fig/770430" pageId="0" pageNumber="575">7e</figureCitation>
) subparallel between procoxae, then subtriangular with rounded apex. 
<emphasis bold="true" italics="true" pageId="0" pageNumber="575">Aedeagus</emphasis>
(Figs 
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Rumilara leleupi comb. nov.: a aedeagus, left lateral view; b aedeagus, ventral view; c aedeagus, right lateral view; d male sternite VIII. Scale: 0.1 mm." figureDoi="10.3897/asp.80.e84013.figure15" httpUri="https://binary.pensoft.net/fig/770438" pageId="0" pageNumber="575">15a-c</figureCitation>
) without long, dorsal, rod-like sclerite on penis; parameres asymmetrical, left one distinctly longer. - Female unknown.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="575" type="description">
<paragraph pageId="0" pageNumber="575">Measurements.</paragraph>
<paragraph pageId="0" pageNumber="575">♂ - CL: 3.28 mm; PL: 0.92 mm; PW: 0.95 mm; EL: 2.36 mm; EW: 1.37 mm.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="575" type="larva">
<paragraph pageId="0" pageNumber="575">Larva</paragraph>
<paragraph pageId="0" pageNumber="575">unknown.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="575" type="distribution">
<paragraph pageId="0" pageNumber="575">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="575">So far known only from the type locality in Napo Province.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="575" type="remarks">
<paragraph pageId="0" pageNumber="575">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="575">
<bibRefCitation author="Deleve, J" journalOrPublisher="Zootaxa" pageId="0" pageNumber="575" refId="B11" refString="Deleve, J, 1968. IV. - ColeopteraElminthidae. In: Leleup N, Leleup J (Eds) Mission zoologique belge aux iles Galapagos et en Ecuador. Resultats scientifiques 1. Institut Royal des Sciences Naturelles de Belgique, Bruxelles, pp. 211-272." title="IV. - ColeopteraElminthidae. In: Leleup N, Leleup J (Eds) Mission zoologique belge aux iles Galapagos et en Ecuador. Resultats scientifiques 1. Institut Royal des Sciences Naturelles de Belgique, Bruxelles, pp. 211 - 272." year="1968">
<normalizedToken originalValue="Delève">Deleve</normalizedToken>
(1968)
</bibRefCitation>
very likely interpreted distinctly shorter left paramere as broken and illustrated it with the same length as the right one. There are no signs of artificial ending of the left paramere, and even their shapes are dissimilar.
</paragraph>
</subSubSection>
</treatment>
</subSection>
</document>