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<mods:title id="1DAA2F4DF74993EED7E6EADBEA76F495">Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae)</mods:title>
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<mods:namePart id="F2C592367B7E6FB625AA462C4B49A92C">Shen, Jia-Wei</mods:namePart>
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<treatment id="C0438794D849FFA2FF017134FD9CFF49" ID-DOI="http://doi.org/10.5281/zenodo.3844548" ID-GBIF-Taxon="164123608" ID-Zenodo-Dep="3844548" LSID="urn:lsid:plazi:treatment:C0438794D849FFA2FF017134FD9CFF49" httpUri="http://treatment.plazi.org/id/C0438794D849FFA2FF017134FD9CFF49" lastPageId="13" lastPageNumber="14" pageId="11" pageNumber="12">
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<paragraph id="48553682D849FFA4FF017134FD19F95C" blockId="11.[151,701,1851,1914]" box="[151,701,1851,1878]" pageId="11" pageNumber="12">
<heading id="131D81EED849FFA4FF017134FD19F95C" bold="true" box="[151,701,1851,1878]" fontSize="11" level="1" pageId="11" pageNumber="12" reason="1">
<taxonomicName id="8FEA4D01D849FFA4FF017134FD19F95C" ID-CoL="3CT5B" authority="(Broun, 1914)" baseAuthorityName="Broun" baseAuthorityYear="1914" box="[151,701,1851,1878]" class="Insecta" family="Staphylinidae" genus="Eupines" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="bisulcifrons" subGenus="Byraxis">
<emphasis id="7A9EEA90D849FFA4FF017134FD19F95C" bold="true" box="[151,701,1851,1878]" pageId="11" pageNumber="12">
<emphasis id="7A9EEA90D849FFA4FF017134FF5EF95C" bold="true" box="[151,250,1852,1878]" italics="true" pageId="11" pageNumber="12">Eupines</emphasis>
(
<emphasis id="7A9EEA90D849FFA4FE9A7134FECDF95C" bold="true" box="[268,361,1852,1878]" italics="true" pageId="11" pageNumber="12">Byraxis</emphasis>
)
<emphasis id="7A9EEA90D849FFA4FEEC7133FDADF95C" bold="true" box="[378,521,1851,1878]" italics="true" pageId="11" pageNumber="12">bisulcifrons</emphasis>
(Broun, 1914)
</emphasis>
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</heading>
</paragraph>
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<subSubSection id="00F06509D849FFA4FF017168FECAF970" box="[151,366,1887,1914]" pageId="11" pageNumber="12" type="description">
<paragraph id="48553682D849FFA4FF017168FECAF970" blockId="11.[151,701,1851,1914]" box="[151,366,1887,1914]" pageId="11" pageNumber="12">
<figureCitation id="D0D12A07D849FFA4FF017168FF7FF970" box="[151,219,1887,1914]" captionStart="FIGURE 5" captionStartId="12.[151,250,1553,1578]" captionTargetBox="[169,1416,501,1521]" captionTargetId="figure@12.[161,1426,489,1528]" captionTargetPageId="12" captionText="FIGURE 5. Diagnostic characters of E. (B.) bisulcifrons (Broun, 1914). A) Habitus. B) Abdomen, in ventral view. C) Antenna, in dorsal view. Scale bars: A = 1 mm, B, C = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3821186" httpUri="https://zenodo.org/record/3821186/files/figure.png" pageId="11" pageNumber="12">Fig. 5</figureCitation>
; map
<figureCitation id="D0D12A07D849FFA4FE897168FECAF970" box="[287,366,1888,1914]" captionStart="FIGURE 49" captionStartId="75.[151,250,1945,1970]" captionTargetBox="[207,1385,201,1918]" captionTargetId="figure@75.[193,1394,181,1920]" captionTargetPageId="75" captionText="FIGURE 49. Geographic distributions of E. (B.) anisarthra (Broun, 1914), E. (B.) bisulcifrons (Broun, 1914), E. (B.) brevis sp. n., E. (B.) caesta sp. n., E. (B.) carinata sp. n., E. (B.) coalita sp. n., E. (B.) complector sp. n. and E. (B.) conspicua (Broun, 1893) in New Zealand." figureDoi="http://doi.org/10.5281/zenodo.3821276" httpUri="https://zenodo.org/record/3821276/files/figure.png" pageId="11" pageNumber="12">Fig. 49</figureCitation>
</paragraph>
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<subSubSection id="00F06509D849FFA4FF0171ADFD82F9F0" pageId="11" pageNumber="12" type="reference_group">
<paragraph id="48553682D849FFA4FF0171ADFCCCF9B6" blockId="11.[151,1437,1955,2042]" box="[151,872,1955,1980]" pageId="11" pageNumber="12">
<treatmentCitationGroup id="68FA11ACD849FFA4FF0171ADFCCCF9B6" box="[151,872,1955,1980]" pageId="11" pageNumber="12">
<treatmentCitation id="C94B1093D849FFA4FF0171ADFD8EF9B6" author="Broun, T." box="[151,554,1955,1980]" page="170" pageId="11" pageNumber="12" year="1914">
<taxonomicName id="8FEA4D01D849FFA4FF0171ADFD8EF9B6" ID-CoL="9NSWY" authority="Broun, 1914 b: 170" authorityName="Broun" authorityPageNumber="170" authorityYear="1914" box="[151,554,1955,1980]" class="Insecta" family="Staphylinidae" genus="Byraxis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="bisulcifrons">
<emphasis id="7A9EEA90D849FFA4FF0171ADFEC6F9B6" box="[151,354,1956,1980]" italics="true" pageId="11" pageNumber="12">Byraxis bisulcifrons</emphasis>
<bibRefCitation id="2C7B4B73D849FFA4FEFE71ADFD8EF9B6" author="Broun, T." box="[360,554,1955,1980]" pageId="11" pageNumber="12" pagination="143 - 266" refId="ref58289" refString="Broun, T. (1914 b) Descriptions of new genera and species of Coleoptera. Part III. Bulletin of the New Zealand Institute Wellington, 1 (3), 143 - 266." type="journal article" year="1914">Broun, 1914b: 170</bibRefCitation>
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.
<treatmentCitation id="C94B1093D849FFA4FDA571ABFD54F9B6" author="Hudson, G. V." box="[563,752,1955,1980]" page="364" pageId="11" pageNumber="12" year="1923">
<bibRefCitation id="2C7B4B73D849FFA4FDA571ABFD54F9B6" author="Hudson, G. V." box="[563,752,1955,1980]" pageId="11" pageNumber="12" pagination="353 - 399" refId="ref58506" refString="Hudson, G. V. (1923) An index to New Zealand beetles. Transactions of the New Zealand Institute, 54, 353 - 399." type="journal article" year="1923">Hudson 1923: 364</bibRefCitation>
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,
<treatmentCitation id="C94B1093D849FFA4FD6D71ACFCC0F9B6" author="Hudson, G. V." box="[763,868,1955,1980]" page="185" pageId="11" pageNumber="12" year="1934">
<bibRefCitation id="2C7B4B73D849FFA4FD6D71ACFCC0F9B6" author="Hudson, G. V." box="[763,868,1955,1980]" pageId="11" pageNumber="12" refId="ref58535" refString="Hudson, G. V. (1934) New Zealand beetles and their larvae. An elementary introduction to the study of our native Coleoptera. Wellington Ferguson &amp; Osborn Ltd., Wellington, 236 pp., 17 pls." type="book" year="1934">1934: 185</bibRefCitation>
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.
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<treatmentCitationGroup id="68FA11ACD849FFA4FF0171CBFD82F9F0" pageId="11" pageNumber="12">
<taxonomicName id="8FEA4D01D849FFA4FF0171CBFE69F9D1" ID-CoL="3CT5B" baseAuthorityName="Broun" baseAuthorityYear="1914" box="[151,461,1987,2011]" class="Insecta" family="Staphylinidae" genus="Eupines" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="bisulcifrons" subGenus="Byraxis">
<emphasis id="7A9EEA90D849FFA4FF0171CBFF4DF9D0" box="[151,233,1987,2010]" italics="true" pageId="11" pageNumber="12">Eupines</emphasis>
(
<emphasis id="7A9EEA90D849FFA4FF6F71CBFEE2F9D0" box="[249,326,1987,2010]" italics="true" pageId="11" pageNumber="12">Byraxis</emphasis>
)
<emphasis id="7A9EEA90D849FFA4FEC071CBFE69F9D1" box="[342,461,1987,2011]" italics="true" pageId="11" pageNumber="12">bisulcifrons</emphasis>
</taxonomicName>
: combination and subgeneric assignment by
<treatmentCitation id="C94B1093D849FFA4FC0871CBFB4FF9D1" author="Nomura, S. &amp; Leschen, R. A. B." box="[926,1259,1986,2011]" page="254" pageId="11" pageNumber="12" year="2006">
<bibRefCitation id="2C7B4B73D849FFA4FC0871CBFB4FF9D1" author="Nomura, S. &amp; Leschen, R. A. B." box="[926,1259,1986,2011]" pageId="11" pageNumber="12" pagination="239 - 272" refId="ref58967" refString="Nomura, S. &amp; Leschen, R. A. B. (2006) Faunistic review on the pselaphine species known from New Zealand (Insecta, Coleoptera, Staphylinidae). National Science Museum Monographs, 34, 239 - 272." type="journal article" year="2006">Nomura and Leschen 2006: 254</bibRefCitation>
</treatmentCitation>
.
<treatmentCitation id="C94B1093D849FFA4FB6071CAFF54F9F3" author="Maddison, P." page="417" pageId="11" pageNumber="12" year="2010">
<bibRefCitation id="2C7B4B73D849FFA4FB6071CAFF54F9F3" author="Maddison, P." pageId="11" pageNumber="12" pagination="233 - 467" refId="ref58776" refString="Maddison, P. (2010) Checklist of New Zealand Hexapoda. Order Coleoptera. Chapter Nine: Phylum Arthropoda: Subphylum Hexapoda: Protura, springtails, Diplura, and insects. In: Gordon, D. P. (Ed.), New Zealand inventory of biodiversity. Vol. 2. Kingdom Animalia: Chaetognatha, Ecdysozoa, Ichnofossils. Canterbury University Press, Christchurch, pp. 233 - 467." type="book chapter" year="2010">Maddison 2010: 417</bibRefCitation>
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;
<treatmentCitation id="C94B1093D849FFA4FF6D71E9FD86F9F0" author="Shen, J. - W. &amp; Leschen, R. A. B." box="[251,546,2017,2042]" httpUri="http://treatment.plazi.org/id/2D4AB362C8058D4BFF61F90540D6E483" page="259" pageId="11" pageNumber="12" year="2019">
<bibRefCitation id="2C7B4B73D849FFA4FF6D71E9FD86F9F0" author="Shen, J. - W. &amp; Leschen, R. A. B." box="[251,546,2017,2042]" pageId="11" pageNumber="12" pagination="255 - 302" refId="ref59638" refString="Shen, J. - W. &amp; Leschen, R. A. B. (2019) Catalogue and type designations for New Zealand Goniaceritae (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa, 4614 (2), 255 - 302. https: // doi. org / 10.11646 / zootaxa. 4614.2.2" type="journal article" year="2019">Shen and Leschen 2019: 259</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
</subSubSection>
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<paragraph id="48553682D84EFFA3FF017691FDB4FEB9" blockId="12.[151,1437,152,431]" box="[151,528,152,179]" pageId="12" pageNumber="13">
<materialsCitation id="F8823CDFD84EFFA3FF017691FDA8FEB9" ID-GBIF-Occurrence="2612164382" box="[151,524,152,179]" collectionCode="MC" location="Mount Hutt" pageId="12" pageNumber="13" specimenCount="1" typeStatus="holotype">
<emphasis id="7A9EEA90D84EFFA3FF017691FE9DFEB9" bold="true" box="[151,313,152,179]" pageId="12" pageNumber="13">
<typeStatus id="97518820D84EFFA3FF017691FF76FEB9" box="[151,210,153,179]" pageId="12" pageNumber="13">Type</typeStatus>
locality:
</emphasis>
[
<collectionCode id="2EFBAE47D84EFFA3FED17690FEDCFEB9" box="[327,376,152,179]" country="0" name="Museo de Cipolleti" pageId="12" pageNumber="13">MC</collectionCode>
]
<location id="4D356059D84EFFA3FE137691FDA8FEB9" LSID="urn:lsid:plazi:treatment:C0438794D849FFA2FF017134FD9CFF49:4D356059D84EFFA3FE137691FDA8FEB9" box="[389,524,153,179]" name="Mount Hutt" pageId="12" pageNumber="13">Mount Hutt</location>
</materialsCitation>
.
</paragraph>
<paragraph id="48553682D84EFFA3FF5176B5FD21FEDD" blockId="12.[151,1437,152,431]" box="[199,645,188,215]" pageId="12" pageNumber="13">
<materialsCitation id="F8823CDFD84EFFA3FF5176B5FD25FEDD" ID-GBIF-Occurrence="2612164379" box="[199,641,188,215]" collectionCode="BMNH" pageId="12" pageNumber="13" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="7A9EEA90D84EFFA3FF5176B5FE34FEDD" bold="true" box="[199,400,188,215]" pageId="12" pageNumber="13">
<typeStatus id="97518820D84EFFA3FF5176B5FEA6FEDD" box="[199,258,189,215]" pageId="12" pageNumber="13">Type</typeStatus>
depository:
</emphasis>
<collectionCode id="2EFBAE47D84EFFA3FE0176B5FE4FFEDD" box="[407,491,189,215]" pageId="12" pageNumber="13">BMNH</collectionCode>
LT
<specimenCount id="5EECFD0BD84EFFA3FD8E76B5FD8BFEDD" box="[536,559,189,215]" pageId="12" pageNumber="13" type="male"></specimenCount>
, 1 PLT
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="00F06509D84EFFA3FF5176E8FEDAFFA5" pageId="12" pageNumber="13" type="diagnosis">
<paragraph id="48553682D84EFFA3FF5176E8FEDAFFA5" blockId="12.[151,1437,152,431]" pageId="12" pageNumber="13">
<emphasis id="7A9EEA90D84EFFA3FF5176E8FEE0FEF1" bold="true" box="[199,324,224,251]" pageId="12" pageNumber="13">Diagnosis:</emphasis>
Head lacking vertexal foveae; antennae 10-segmented with 2-segmented antennal club; A4, A6, A7, A8 and A10 unmodified, A5 asymmetrical and triangular, excavated at base, A9 asymmetrical, about 3 × longer than A8, inner apical margin expanded, A10 conical, slightly impressed at base, about 1.8 × longer than A9. Abdominal ventrite 2 with pair of small discal protuberances, inserted with strongly curved small apical setae; ventrite 6 unknown. Protrochanters with minute ventral spine, about one-ninth of protrochanteral width; protibiae and mesotibiae unmodified.
</paragraph>
</subSubSection>
<caption id="1C95660AD84EFFA3FF017019FD12F847" ID-DOI="http://doi.org/10.5281/zenodo.3821186" ID-Zenodo-Dep="3821186" httpUri="https://zenodo.org/record/3821186/files/figure.png" pageId="12" pageNumber="13" startId="12.[151,250,1553,1578]" targetBox="[169,1416,501,1521]" targetPageId="12">
<paragraph id="48553682D84EFFA3FF017019FD12F847" blockId="12.[151,1437,1553,1614]" pageId="12" pageNumber="13">
<emphasis id="7A9EEA90D84EFFA3FF017019FEB6F820" bold="true" box="[151,274,1553,1578]" pageId="12" pageNumber="13">FIGURE 5.</emphasis>
Diagnostic characters of
<emphasis id="7A9EEA90D84EFFA3FD85701BFD86F820" box="[531,546,1555,1578]" italics="true" pageId="12" pageNumber="13">E</emphasis>
.
<taxonomicName id="8FEA4D01D84EFFA3FDB9701AFCCFF820" authority="(Broun, 1914)" baseAuthorityName="Broun" baseAuthorityYear="1914" box="[559,875,1554,1578]" class="Insecta" family="Staphylinidae" genus="Eupines" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="bisulcifrons" subGenus="Byraxis">
(
<emphasis id="7A9EEA90D84EFFA3FDA1701BFDE2F820" box="[567,582,1555,1578]" italics="true" pageId="12" pageNumber="13">B</emphasis>
.)
<emphasis id="7A9EEA90D84EFFA3FDCA701AFD77F820" box="[604,723,1554,1578]" italics="true" pageId="12" pageNumber="13">bisulcifrons</emphasis>
(Broun, 1914)
</taxonomicName>
.
<emphasis id="7A9EEA90D84EFFA3FCE57019FC2AF820" bold="true" box="[883,910,1553,1578]" pageId="12" pageNumber="13">A)</emphasis>
Habitus.
<emphasis id="7A9EEA90D84EFFA3FC797019FBACF820" bold="true" box="[1007,1032,1553,1578]" pageId="12" pageNumber="13">B)</emphasis>
Abdomen, in ventral view.
<emphasis id="7A9EEA90D84EFFA3FAB67019FA9EF820" bold="true" box="[1312,1338,1553,1578]" pageId="12" pageNumber="13">C)</emphasis>
Antenna, in dorsal view. Scale bars: A = 1 mm, B, C = 0.2 mm.
</paragraph>
</caption>
<subSubSection id="00F06509D84EFFA2FF517073FD35FEDD" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13" type="description">
<paragraph id="48553682D84EFFA2FF517073FD35FEDD" blockId="12.[151,1437,1659,2011]" lastBlockId="13.[151,1436,153,323]" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13">
<emphasis id="7A9EEA90D84EFFA3FF517073FED3F89C" bold="true" box="[199,375,1659,1686]" pageId="12" pageNumber="13">Redescription:</emphasis>
Body length
<quantity id="8F129B67D84EFFA3FD997074FDD2F89D" box="[527,630,1660,1687]" metricMagnitude="-3" metricUnit="m" metricValue="1.43" pageId="12" pageNumber="13" unit="mm" value="1.43">1.43 mm</quantity>
; body colour reddish brown with lighter coloured tibiae and tarsi.
<emphasis id="7A9EEA90D84EFFA3FACA7074FA38F89C" bold="true" box="[1372,1436,1660,1686]" pageId="12" pageNumber="13">Head</emphasis>
wider than long; frons slightly narrowed at end, about half width of head at eye level; eyes slightly prominent in dorsal view, with about 22 facets; lacking vertexal foveae; antennal tubercles barely visible; lacking postantennal notches; postantennal cavities large, slightly longer than antennal tubercles in dorsal view; antennae 10-segmented with 2-segmented antennal club; A4, A6, A7, A8 and A10 unmodified, A5 asymmetrical and triangular, excavated at base, A9 asymmetrical, about 3 × longer than A8, inner apical margin expanded, A10 conical and slightly impressed at base, about 1.8 × longer than A9.
<emphasis id="7A9EEA90D84EFFA3FDA6715CFD0DF964" bold="true" box="[560,681,1876,1902]" pageId="12" pageNumber="13">Pronotum</emphasis>
about as long as wide, wider than head at eye level; lacking lateral antebasal foveae.
<emphasis id="7A9EEA90D84EFFA3FEF77170FE09F998" bold="true" box="[353,429,1912,1938]" pageId="12" pageNumber="13">Elytra</emphasis>
slightly wider than long, widest at apical two-thirds; humeral calli barely visible; sutural stria distinctively extending from base to apex.
<emphasis id="7A9EEA90D84EFFA3FD3E7193FCB9F9BC" bold="true" box="[680,797,1947,1974]" pageId="12" pageNumber="13">Abdomen</emphasis>
with tergite 1 about 2.3 × longer than tergite 2; ventrite 2 with pair of small discal protuberances, inserted with strongly curved small apical setae; ventrite 6 unknown.
<emphasis id="7A9EEA90D84EFFA3FAF271C8FA38F9D0" bold="true" box="[1380,1436,1984,2010]" pageId="12" pageNumber="13">Legs</emphasis>
with protrochanters modified with minute ventral spine, about one-ninth of protrochanteral width; protibiae and mesotibiae unmodified.
<emphasis id="7A9EEA90D84FFFA2FE3176B4FDBEFEDD" bold="true" box="[423,538,188,215]" pageId="13" pageNumber="14">Aedeagus</emphasis>
unknown.
</paragraph>
</subSubSection>
<subSubSection id="00F06509D84FFFA2FF5176E8FB43FF15" pageId="13" pageNumber="14" type="discussion">
<paragraph id="48553682D84FFFA2FF5176E8FB43FF15" blockId="13.[151,1436,153,323]" pageId="13" pageNumber="14">
<emphasis id="7A9EEA90D84FFFA2FF5176E8FEF6FEF1" bold="true" box="[199,338,224,251]" pageId="13" pageNumber="14">Comments:</emphasis>
Only two type specimens are known. We did not attempt to dissect the
<typeStatus id="97518820D84FFFA2FBF176E9FB6DFEF1" box="[1127,1225,225,251]" pageId="13" pageNumber="14" type="syntype">syntypes</typeStatus>
since males of this species have distinctive antennal modifications (
<figureCitation id="D0D12A07D84FFFA2FD20770DFCA8FF15" box="[694,780,260,287]" captionStart="FIGURE 5" captionStartId="12.[151,250,1553,1578]" captionTargetBox="[169,1416,501,1521]" captionTargetId="figure@12.[161,1426,489,1528]" captionTargetPageId="12" captionText="FIGURE 5. Diagnostic characters of E. (B.) bisulcifrons (Broun, 1914). A) Habitus. B) Abdomen, in ventral view. C) Antenna, in dorsal view. Scale bars: A = 1 mm, B, C = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3821186" httpUri="https://zenodo.org/record/3821186/files/figure.png" pageId="13" pageNumber="14">Fig. 5B</figureCitation>
) that separate the species from congeners.
</paragraph>
</subSubSection>
<subSubSection id="00F06509D84FFFA2FF517720FD9CFF49" box="[199,568,296,323]" pageId="13" pageNumber="14" type="distribution">
<paragraph id="48553682D84FFFA2FF517720FD9CFF49" blockId="13.[151,1436,153,323]" box="[199,568,296,323]" pageId="13" pageNumber="14">
<emphasis id="7A9EEA90D84FFFA2FF517720FEC7FF49" bold="true" box="[199,355,296,323]" pageId="13" pageNumber="14">Distribution:</emphasis>
South Island. MC.
</paragraph>
</subSubSection>
</treatment>
</document>