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<document id="E328F530C7C120C3E37083E62A02047E" ID-CLB-Dataset="22233" ID-DOI="10.11646/zootaxa.4777.1.1" ID-GBIF-Dataset="75de85af-39c5-41ca-9ccc-4ff7b743eefa" ID-ISSN="1175-5326" ID-Zenodo-Dep="3821174" ID-ZooBank="BF4149EF-478F-4FE4-BE7B-B497272F99C0" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1589267332095" checkinUser="plazi" docAuthor="Shen, Jia-Wei &amp; Leschen, Richard A. B." docDate="2020" docId="C0438794D807FFE8FF01711CFCEFFF6D" docLanguage="en" docName="zootaxa.4777.1.1.pdf.imf" docOrigin="Zootaxa 4777 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Eupines (Byraxis) waikaremoana Shen &amp; Leschen 2020, sp. nov." docType="treatment" docUuid="23F70786-BF69-44D8-88D5-367EEE65C675" docUuidSource="ZooBank" docVersion="6" lastPageNumber="72" masterDocId="3C7AFFECD842FFAFFF967608FFA4FE0A" masterDocTitle="Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae)" masterLastPageNumber="84" masterPageNumber="1" pageNumber="70" updateTime="1698839235073" updateUser="ExternalLinkService">
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<mods:title id="1A5C466814F4ECF91CE674987CB1821D">Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae)</mods:title>
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<mods:name id="3D2C3C1D16D46C786A8692E16FA43FA3" type="personal">
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<mods:namePart id="DAF5AE562F21195A0463F703EECE60CF">Shen, Jia-Wei</mods:namePart>
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<mods:namePart id="ADEDA1D2F863E76E204B7A6EDAA7EA3A">Leschen, Richard A. B.</mods:namePart>
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<mods:title id="8BBC33B254E7D25366C37FBD4B37F434">Zootaxa</mods:title>
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<mods:date id="6383ABC9AB5AE9216C15DDEBC08302DF">2020</mods:date>
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<mods:number id="939D0F7236E79D04D5F825AB50DB36CF">4777</mods:number>
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<treatment id="C0438794D807FFE8FF01711CFCEFFF6D" ID-DOI="http://doi.org/10.5281/zenodo.3844544" ID-GBIF-Taxon="164123621" ID-Zenodo-Dep="3844544" LSID="urn:lsid:zoobank.org:act:23F70786-BF69-44D8-88D5-367EEE65C675" httpUri="http://treatment.plazi.org/id/C0438794D807FFE8FF01711CFCEFFF6D" lastPageId="71" lastPageNumber="72" pageId="69" pageNumber="70">
<subSubSection id="00F06509D807FFEAFF01711CFD36F924" box="[151,658,1812,1839]" pageId="69" pageNumber="70" type="nomenclature">
<paragraph id="48553682D807FFEAFF01711CFD36F924" blockId="69.[151,658,1812,1874]" box="[151,658,1812,1839]" pageId="69" pageNumber="70">
<heading id="131D81EED807FFEAFF01711CFD36F924" bold="true" box="[151,658,1812,1839]" fontSize="11" level="1" pageId="69" pageNumber="70" reason="1">
<emphasis id="7A9EEA90D807FFEAFF01711CFD36F924" bold="true" box="[151,658,1812,1839]" pageId="69" pageNumber="70">
<taxonomicName id="8FEA4D01D807FFEAFF01711CFD89F925" ID-CoL="8Y3QR" authority="Shen &amp; Leschen, 2020" authorityName="Shen &amp; Leschen" authorityYear="2020" box="[151,557,1812,1839]" class="Insecta" family="Staphylinidae" genus="Eupines" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="waikaremoana" status="sp. nov." subGenus="Byraxis">
<emphasis id="7A9EEA90D807FFEAFF01711CFF5EF924" bold="true" box="[151,250,1812,1838]" italics="true" pageId="69" pageNumber="70">Eupines</emphasis>
(
<emphasis id="7A9EEA90D807FFEAFE9A711CFECDF924" bold="true" box="[268,361,1812,1838]" italics="true" pageId="69" pageNumber="70">Byraxis</emphasis>
)
<emphasis id="7A9EEA90D807FFEAFEEC711CFD89F925" bold="true" box="[378,557,1812,1839]" italics="true" pageId="69" pageNumber="70">waikaremoana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="61AD57EBD807FFEAFDA2711CFD36F924" box="[564,658,1812,1838]" pageId="69" pageNumber="70" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="00F06509D807FFEAFF017130FED8F958" box="[151,380,1847,1874]" pageId="69" pageNumber="70" type="description">
<paragraph id="48553682D807FFEAFF017130FED8F958" blockId="69.[151,658,1812,1874]" box="[151,380,1847,1874]" pageId="69" pageNumber="70">
<figureCitation id="D0D12A07D807FFEAFF017130FF4EF958" box="[151,234,1848,1874]" captionStart="FIGURE 46" captionStartId="70.[151,250,1325,1350]" captionTargetBox="[155,1424,279,1294]" captionTargetId="figure@70.[151,1435,273,1300]" captionTargetPageId="70" captionText="FIGURE 46. Diagnostic characters of E. (B.) waikaremoana sp. n. A) Habitus. B) Antenna, in dorsal view. C) Same, in lateral view. D) Protrochanter. E) Ventrite 2. F) Ventrite 6. G) Aedeagus, in dorsal view. H) Same, in lateral view. I) Same, in ventral view. Scale bars: A = 1 mm, BE = 0.2 mm, FI = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3821270" httpUri="https://zenodo.org/record/3821270/files/figure.png" pageId="69" pageNumber="70">Fig. 46</figureCitation>
; map
<figureCitation id="D0D12A07D807FFEAFEBB7130FED8F958" box="[301,380,1847,1874]" captionStart="FIGURE 54" captionStartId="80.[151,250,1945,1970]" captionTargetBox="[203,1384,197,1918]" captionTargetId="figure@80.[189,1398,181,1919]" captionTargetPageId="80" captionText="FIGURE 54. Geographic distributions of E. (B.) pannicula sp. n., E. (B.) petila sp. n., E. (B.) platyarthra (Broun, 1893), E. (B.) protibialis sp. n., E. (B.) setifera (Broun, 1893), E. (B.) waikaremoana sp. n., E. (B.) whirinaki sp. n. and E. (B.) undecim sp. n. in New Zealand." figureDoi="http://doi.org/10.5281/zenodo.3821286" httpUri="https://zenodo.org/record/3821286/files/figure.png" pageId="69" pageNumber="70">Fig. 54</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="00F06509D807FFE9FF017177FC9FFEF1" lastPageId="70" lastPageNumber="71" pageId="69" pageNumber="70" type="diagnosis">
<paragraph id="48553682D807FFE9FF017177FC9FFEF1" blockId="69.[151,1436,1919,2018]" lastBlockId="70.[151,1436,153,251]" lastPageId="70" lastPageNumber="71" pageId="69" pageNumber="70">
<emphasis id="7A9EEA90D807FFEAFF017177FEB0F990" bold="true" box="[151,276,1919,1946]" pageId="69" pageNumber="70">Diagnosis:</emphasis>
Head with vertexal foveae punctiform; antennae 10-segmented with 2-segmented antennal club; A4, A5, A6, A7, A8 and A10 unmodified, A8 slightly shorter than A7, A9 reversed cone-shaped, slightly asymmetrical, with apical spine at ventral side, A10 conical and slightly longer than A9. Abdominal ventrite 2 with pair of min- ute discal protuberances, inserted with strongly curved minute apical setae; ventrite 6 with pair of long and acute median struts, about half of ventral length. Protrochanters with long and acute ventral spine, about three-fourths of protrochanteral width; protibiae and mesotibiae unmodified.
</paragraph>
</subSubSection>
<caption id="1C95660AD804FFE9FF017325FD4BFB87" ID-DOI="http://doi.org/10.5281/zenodo.3821270" ID-Zenodo-Dep="3821270" httpUri="https://zenodo.org/record/3821270/files/figure.png" pageId="70" pageNumber="71" startId="70.[151,250,1325,1350]" targetBox="[155,1424,279,1294]" targetPageId="70">
<paragraph id="48553682D804FFE9FF017325FD4BFB87" blockId="70.[151,1436,1325,1422]" pageId="70" pageNumber="71">
<emphasis id="7A9EEA90D804FFE9FF017325FE84FB4C" bold="true" box="[151,288,1325,1350]" pageId="70" pageNumber="71">FIGURE 46.</emphasis>
Diagnostic characters of
<taxonomicName id="8FEA4D01D804FFE9FDB47326FCA5FB4C" authority="Shen &amp; Leschen, 2020" authorityName="Shen &amp; Leschen" authorityYear="2020" box="[546,769,1326,1350]" class="Insecta" family="Staphylinidae" genus="Eupines" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="70" pageNumber="10" phylum="Arthropoda" rank="species" species="waikaremoana" status="sp. nov." subGenus="Byraxis">
<emphasis id="7A9EEA90D804FFE9FDB47326FD95FB4F" box="[546,561,1326,1349]" italics="true" pageId="70" pageNumber="71">E</emphasis>
. (
<emphasis id="7A9EEA90D804FFE9FDD07326FDF1FB4F" box="[582,597,1326,1349]" italics="true" pageId="70" pageNumber="71">B</emphasis>
.)
<emphasis id="7A9EEA90D804FFE9FDFD7326FCA5FB4C" box="[619,769,1326,1350]" italics="true" pageId="70" pageNumber="71">waikaremoana</emphasis>
</taxonomicName>
<emphasis id="7A9EEA90D804FFE9FC907326FCC4FB4C" bold="true" box="[774,864,1325,1350]" pageId="70" pageNumber="71">
<taxonomicNameLabel id="61AD57EBD804FFE9FC907326FC9BFB4C" box="[774,831,1326,1350]" pageId="70" pageNumber="71" rank="species">sp. n.</taxonomicNameLabel>
A)
</emphasis>
Habitus.
<emphasis id="7A9EEA90D804FFE9FC547325FC7FFB4C" bold="true" box="[962,987,1325,1350]" pageId="70" pageNumber="71">B)</emphasis>
Antenna, in dorsal view.
<emphasis id="7A9EEA90D804FFE9FB4B7325FB5CFB4C" bold="true" box="[1245,1272,1325,1350]" pageId="70" pageNumber="71">C)</emphasis>
Same, in lateral view.
<emphasis id="7A9EEA90D804FFE9FF427359FF4AFB60" bold="true" box="[212,238,1361,1386]" pageId="70" pageNumber="71">D)</emphasis>
Protrochanter.
<emphasis id="7A9EEA90D804FFE9FE1D7359FE07FB60" bold="true" box="[395,419,1361,1386]" pageId="70" pageNumber="71">E)</emphasis>
Ventrite 2.
<emphasis id="7A9EEA90D804FFE9FD8D7359FD97FB60" bold="true" box="[539,563,1361,1386]" pageId="70" pageNumber="71">F)</emphasis>
Ventrite 6.
<emphasis id="7A9EEA90D804FFE9FD3C7359FD61FB60" bold="true" box="[682,709,1361,1386]" pageId="70" pageNumber="71">G)</emphasis>
Aedeagus, in dorsal view.
<emphasis id="7A9EEA90D804FFE9FC4F7359FC50FB60" bold="true" box="[985,1012,1361,1386]" pageId="70" pageNumber="71">H)</emphasis>
Same, in lateral view.
<emphasis id="7A9EEA90D804FFE9FB4B7359FB54FB60" bold="true" box="[1245,1264,1361,1386]" pageId="70" pageNumber="71">I)</emphasis>
Same, in ventral view. Scale bars: A = 1 mm, BE = 0.2 mm, FI = 0.1 mm.
</paragraph>
</caption>
<subSubSection id="00F06509D804FFE9FF5173B3FC04F9C4" pageId="70" pageNumber="71" type="description">
<paragraph id="48553682D804FFE9FF5173B3FC04F9C4" blockId="70.[151,1436,1467,1999]" pageId="70" pageNumber="71">
<emphasis id="7A9EEA90D804FFE9FF5173B3FEFFFBDC" bold="true" box="[199,347,1467,1494]" pageId="70" pageNumber="71">Description:</emphasis>
Body length
<quantity id="8F129B67D804FFE9FE6E73B4FDC5FBDD" box="[504,609,1468,1495]" metricMagnitude="-3" metricUnit="m" metricValue="1.91" pageId="70" pageNumber="71" unit="mm" value="1.91">1.91 mm</quantity>
; body colour reddish brown with lighter coloured tibiae and tarsi.
<emphasis id="7A9EEA90D804FFE9FACA73B4FA38FBDC" bold="true" box="[1372,1436,1468,1494]" pageId="70" pageNumber="71">Head</emphasis>
longer than wide; frons not narrowed at end, about three-fourths width of head at eye level; eyes moderately developed, slightly prominent in dorsal view, with about 27 facets; vertexal foveae punctiform; antennal tubercles weakly developed; lacking postantennal notches; postantennal cavities large, slightly shorter than antennal tubercles in dorsal view; antennae 10-segmented with 2-segmented antennal club; A4, A5, A6, A7, A8 and A10 unmodified, A8 slightly shorter than A7, A9 reversed cone-shaped, slightly asymmetrical, with apical spine at ventral side, A10 conical and slightly longer than A9.
<emphasis id="7A9EEA90D804FFE9FDBA709CFD01F8A4" bold="true" box="[556,677,1684,1710]" pageId="70" pageNumber="71">Pronotum</emphasis>
longer than wide, about as wide as head at eye level; lacking lateral antebasal foveae.
<emphasis id="7A9EEA90D804FFE9FEF470B0FE0AF8D8" bold="true" box="[354,430,1720,1746]" pageId="70" pageNumber="71">Elytra</emphasis>
slightly longer than wide, widest near middle; humeral calli weak, slightly prominent in dorsal view; sutural stria barely visible.
<emphasis id="7A9EEA90D804FFE9FDC870D3FD77F8FC" bold="true" box="[606,723,1755,1782]" pageId="70" pageNumber="71">Abdomen</emphasis>
with tergite 1 about 1.3 × longer than tergite 2; ventrite 2 with pair of minute discal protuberances, inserted with strongly curved minute apical setae; ventrite 6 with pair of long and acute median struts, about half of ventral length.
<emphasis id="7A9EEA90D804FFE9FD73712CFCB9F934" bold="true" box="[741,797,1828,1854]" pageId="70" pageNumber="71">Legs</emphasis>
with protrochanters modified with long and acute ventral spine, about three-fourths of protrochanteral width; protibiae and mesotibiae unmodified.
<emphasis id="7A9EEA90D804FFE9FB33714FFABCF968" bold="true" box="[1189,1304,1863,1890]" pageId="70" pageNumber="71">Aedeagus</emphasis>
about
<quantity id="8F129B67D804FFE9FAFD7140FF67F98D" metricMagnitude="-4" metricUnit="m" metricValue="3.5999999999999996" pageId="70" pageNumber="71" unit="mm" value="0.36">0.36 mm</quantity>
long, with slender parameres curved towards each other at apex, apex bent downwards in lateral view, with five pairs of long apical setae; median lobe with two large sclerites, curved towards each other at apex, widest near middle, right one longer and much wider than left one in dorsal view.
</paragraph>
</subSubSection>
<subSubSection id="00F06509D805FFE8FF517690FCEFFF6D" pageId="71" pageNumber="72" type="discussion">
<paragraph id="48553682D805FFE8FF517690FCEFFF6D" blockId="71.[151,1437,152,359]" pageId="71" pageNumber="72">
<emphasis id="7A9EEA90D805FFE8FF517690FEF6FEB9" bold="true" box="[199,338,152,179]" pageId="71" pageNumber="72">Comments:</emphasis>
<materialsCitation id="F8823CDFD805FFE8FECD7690FCEFFF6D" ID-GBIF-Occurrence="2612164349" collectingDate="1975-11-19" collectionCode="NZAC" collectorName="L. Waikaremoana &amp; G. Kuschel" county="This" location="This" municipality="Etymology" pageId="71" pageNumber="72" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<location id="4D356059D805FFE8FECD7690FE29FEB8" LSID="urn:lsid:plazi:treatment:C0438794D807FFE8FF01711CFCEFFF6D:4D356059D805FFE8FECD7690FE29FEB8" box="[347,397,152,178]" county="This" municipality="Etymology" name="This" pageId="71" pageNumber="72">This</location>
species is most similar to
<taxonomicName id="8FEA4D01D805FFE8FD507691FCDFFEB8" authorityName="Shen &amp; Leschen" authorityYear="2020" box="[710,891,153,179]" class="Insecta" family="Staphylinidae" genus="Eupines" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="carinata" subGenus="Byraxis">
<emphasis id="7A9EEA90D805FFE8FD507691FD7AFEB8" box="[710,734,153,178]" italics="true" pageId="71" pageNumber="72">E.</emphasis>
(
<emphasis id="7A9EEA90D805FFE8FD677691FCAEFEB8" box="[753,778,153,178]" italics="true" pageId="71" pageNumber="72">B.</emphasis>
)
<emphasis id="7A9EEA90D805FFE8FC8A7691FCDFFEB8" box="[796,891,153,178]" italics="true" pageId="71" pageNumber="72">carinata</emphasis>
</taxonomicName>
by sharing the large body size (average length around 2.00 mm) but can be separated by having A10 unmodified (
<figureCitation id="D0D12A07D805FFE8FC1176B4FC4AFEDD" box="[903,1006,188,215]" captionStart="FIGURE 46" captionStartId="70.[151,250,1325,1350]" captionTargetBox="[155,1424,279,1294]" captionTargetId="figure@70.[151,1435,273,1300]" captionTargetPageId="70" captionText="FIGURE 46. Diagnostic characters of E. (B.) waikaremoana sp. n. A) Habitus. B) Antenna, in dorsal view. C) Same, in lateral view. D) Protrochanter. E) Ventrite 2. F) Ventrite 6. G) Aedeagus, in dorsal view. H) Same, in lateral view. I) Same, in ventral view. Scale bars: A = 1 mm, BE = 0.2 mm, FI = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3821270" httpUri="https://zenodo.org/record/3821270/files/figure.png" pageId="71" pageNumber="72">Fig. 46B</figureCitation>
).
<emphasis id="7A9EEA90D805FFE8FF5176E8FEC7FEF1" bold="true" box="[199,355,224,251]" pageId="71" pageNumber="72">
<location id="4D356059D805FFE8FF5176E8FEFAFEF1" LSID="urn:lsid:plazi:treatment:C0438794D807FFE8FF01711CFCEFFF6D:4D356059D805FFE8FF5176E8FEFAFEF1" box="[199,350,224,251]" county="This" municipality="Etymology" name="Distribution" pageId="71" pageNumber="72">Distribution</location>
:
</emphasis>
<location id="4D356059D805FFE8FEFC76E8FE5AFEF0" LSID="urn:lsid:plazi:treatment:C0438794D807FFE8FF01711CFCEFFF6D:4D356059D805FFE8FEFC76E8FE5AFEF0" box="[362,510,224,250]" county="This" municipality="Etymology" name="North Island." pageId="71" pageNumber="72">North Island.</location>
GB.
<emphasis id="7A9EEA90D805FFE8FF51770DFEF6FF15" bold="true" box="[199,338,261,287]" pageId="71" pageNumber="72">
<collectingMunicipality id="A831ACF8D805FFE8FF51770DFEE8FF15" box="[199,332,261,287]" pageId="71" pageNumber="72">Etymology</collectingMunicipality>
:
</emphasis>
<collectingCounty id="A1344E0ED805FFE8FECF770CFE2FFF14" box="[345,395,260,286]" pageId="71" pageNumber="72">This</collectingCounty>
species is named after the
<typeStatus id="97518820D805FFE8FD2E770CFCBEFF14" box="[696,794,260,286]" pageId="71" pageNumber="72" type="holotype">holotype</typeStatus>
locality.
<emphasis id="7A9EEA90D805FFE8FF517721FDC7FF49" bold="true" box="[199,611,296,323]" pageId="71" pageNumber="72">
Type material (
<collectionCode id="2EFBAE47D805FFE8FE147720FE7CFF49" box="[386,472,296,323]" country="New Zealand" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34070" lsid="urn:lsid:biocol.org:col:34070" name="New Zealand Arthropod Collection" pageId="71" pageNumber="72" type="Other institutional type">NZAC</collectionCode>
):
<typeStatus id="97518820D805FFE8FE797721FDFAFF49" box="[495,606,297,323]" pageId="71" pageNumber="72" type="holotype">Holotype</typeStatus>
.
</emphasis>
<specimenCount id="5EECFD0BD805FFE8FDFA7721FD27FF49" box="[620,643,297,323]" pageId="71" pageNumber="72" type="male"></specimenCount>
, specimen entire, glued on point, aedeagus + proleg on acetate card: “GB,
<collectorName id="E51F5354D805FFE8FF4F7745FE05FF6C" box="[217,417,332,359]" pageId="71" pageNumber="72">L. Waikaremoana</collectorName>
,
<date id="3C541042D805FFE8FE3A7745FDE6FF6D" box="[428,578,332,359]" pageId="71" pageNumber="72" value="1975-11-19">
<collectingDate id="2C10E9AAD805FFE8FE3A7745FDE6FF6D" box="[428,578,332,359]" pageId="71" pageNumber="72" value="1975-11-19">19 Nov 1975</collectingDate>
</date>
,
<collectorName id="E51F5354D805FFE8FDDB7745FD6FFF6C" box="[589,715,332,359]" pageId="71" pageNumber="72">G. Kuschel</collectorName>
[in hand]”.
</materialsCitation>
</paragraph>
</subSubSection>
</treatment>
</document>