361 lines
36 KiB
XML
361 lines
36 KiB
XML
<document ID-DOI="http://doi.org/10.5281/zenodo.3790668" ID-GBIF-Dataset="6b003da1-30a4-4961-839d-53f2f79ae563" ID-GBIF-Taxon="163748825" ID-ISSN="1313–2970" ID-Zenodo-Dep="3790668" ID-ZooBank="A447D3B6-5387-44B6-AC1B-A05988000E43" approvalRequired="18" approvalRequired_for_taxonomicNames="7" approvalRequired_for_textStreams="11" checkinTime="1587964501893" checkinUser="plazi" docAuthor="Huys, Rony & Lee, Wonchoel" docDate="2009" docId="8F78FF1D1618CB4EC380181BFBB1D6C8" docLanguage="en" docName="ZK_article_2079.pdf" docOrigin="ZooKeys 23 (23)" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@763ee217" docStyleName="zookeys.2008.journal_article" docTitle="Pseudonychocamptus Lang 1944, comb. n." docType="key" docVersion="4" lastPageId="17" lastPageNumber="18" masterDocId="734187651608CB5FC30C1B58FFE2D275" masterDocTitle="Proposal of Marbefia, gen. n. and Inermiphonte, gen. n., including updated keys to the species of Pseudonychocamptus Lang, 1944 and Paralaophonte Lang, 1948 (Copepoda, Harpacticoida, Laophontidae)" masterLastPageNumber="38" masterPageNumber="1" pageId="16" pageNumber="17" updateTime="1643568717307" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Proposal of Marbefia, gen. n. and Inermiphonte, gen. n., including updated keys to the species of Pseudonychocamptus Lang, 1944 and Paralaophonte Lang, 1948 (Copepoda, Harpacticoida, Laophontidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Huys, Rony</mods:namePart>
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<mods:affiliation>Natural History Museum, London, United Kingdom</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Lee, Wonchoel</mods:namePart>
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<mods:affiliation>Hanyang University, Laboratory of Biodiversity, Seoul, Korea, South</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2009</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2009-10-06</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>23</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>23</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>38</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.3897/zookeys.23.168</mods:identifier>
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<mods:identifier type="GBIF-Dataset">6b003da1-30a4-4961-839d-53f2f79ae563</mods:identifier>
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<mods:identifier type="ISSN">1313–2970</mods:identifier>
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<mods:identifier type="Zenodo-Dep">576545</mods:identifier>
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<mods:identifier type="ZooBank">A447D3B6-5387-44B6-AC1B-A05988000E43</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.3790668" ID-GBIF-Taxon="163748825" ID-Zenodo-Dep="3790668" LSID="urn:lsid:plazi:treatment:8F78FF1D1618CB4EC380181BFBB1D6C8" httpUri="http://treatment.plazi.org/id/8F78FF1D1618CB4EC380181BFBB1D6C8" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17">
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<subSubSection lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17" type="nomenclature">
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<paragraph blockId="16.[140,784,835,862]" box="[140,784,835,862]" pageId="16" pageNumber="17">
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<heading bold="true" box="[140,784,835,862]" fontSize="11" level="1" pageId="16" pageNumber="17" reason="1">
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<emphasis bold="true" box="[140,784,835,862]" pageId="16" pageNumber="17">
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Key to the species of
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<taxonomicName authority="Lang, 1944" authorityName="Lang" authorityYear="1944" box="[391,784,835,862]" class="Maxillopoda" family="Laophontidae" genus="Pseudonychocamptus" kingdom="Animalia" order="Harpacticoida" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">
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<emphasis bold="true" box="[391,641,835,862]" italics="true" pageId="16" pageNumber="17">Pseudonychocamptus</emphasis>
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<bibRefCitation author="Lang K" box="[648,784,835,861]" pageId="16" pageNumber="17" refId="ref19213" refString="Lang K (1944) Monographie der Harpacticiden (Vorlaufige Mitteilung). Almqvist & Wiksells Boktryckeri Ab, Uppsala, 39 pp." type="book" year="1944">Lang, 1944</bibRefCitation>
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph blockId="16.[140,1108,906,1672]" pageId="16" pageNumber="17">
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<bibRefCitation author="Lang K" box="[140,275,906,932]" pageId="16" pageNumber="17" refId="ref19213" refString="Lang K (1944) Monographie der Harpacticiden (Vorlaufige Mitteilung). Almqvist & Wiksells Boktryckeri Ab, Uppsala, 39 pp." type="book" year="1944">Lang (1944</bibRefCitation>
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,
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<bibRefCitation author="Lang K" box="[287,347,906,932]" pageId="16" pageNumber="17" refId="ref19237" refString="Lang K (1948) Monographie der Harpacticiden. Hakan Ohlsson, Lund, 1682 pp. (2 volumes)." type="book" year="1948">1948</bibRefCitation>
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) proposed the genus
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<taxonomicName box="[591,769,906,932]" class="Maxillopoda" family="Laophontidae" genus="Pseudolaophonte" kingdom="Animalia" order="Harpacticoida" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">
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<emphasis box="[591,769,906,932]" italics="true" pageId="16" pageNumber="17">Pseudolaophonte</emphasis>
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</taxonomicName>
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for four species previously allocated to the genus
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<taxonomicName authority="Philippi, 1840" authorityName="Philippi" authorityYear="1840" box="[372,660,941,967]" class="Maxillopoda" family="Laophontidae" genus="Laophonte" kingdom="Animalia" order="Harpacticoida" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">
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<emphasis box="[372,485,941,967]" italics="true" pageId="16" pageNumber="17">Laophonte</emphasis>
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<bibRefCitation author="Philippi A" box="[493,660,941,967]" pageId="16" pageNumber="17" pagination="188 - 190" refId="ref20024" refString="Philippi A (1840) Zoologische Bemerkungen (Fortsetzung). IV. Kurze Charakteristik mehrerer neuer Genera aus der Familie der Copepoden. Archiv fur Naturgeschichte 6: 188 - 190." type="journal article" year="1840">Philippi, 1840</bibRefCitation>
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</taxonomicName>
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:
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<taxonomicName authority="Boeck, 1873" authorityName="Boeck" authorityYear="1873" box="[671,1008,941,967]" class="Maxillopoda" family="Laophontidae" genus="Laophonte" kingdom="Animalia" order="Harpacticoida" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="koreni">
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<emphasis box="[671,858,941,967]" italics="true" pageId="16" pageNumber="17">Laophonte koreni</emphasis>
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<bibRefCitation author="Boeck A" box="[866,1008,941,967]" pageId="16" pageNumber="17" pagination="35 - 60" refId="ref17768" refString="Boeck A (1873) Nye Slaegter og Arter af Saltvands-Copepoder. Forhandlinger i Videnskabsselskabet i Kristiania 1872: 35 - 60." type="journal article" year="1873">Boeck, 1873</bibRefCitation>
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</taxonomicName>
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(
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<typeStatus box="[1023,1073,941,967]" pageId="16" pageNumber="17">type</typeStatus>
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by original designation),
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<taxonomicName authority="T. Scott, 1903" authorityName="T. Scott" authorityYear="1903" box="[391,671,976,1003]" class="Maxillopoda" family="Laophontidae" genus="Laophonte" kingdom="Animalia" order="Harpacticoida" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="gracilis">
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<emphasis box="[391,497,976,1002]" italics="true" pageId="16" pageNumber="17">L. gracilis</emphasis>
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T.
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<bibRefCitation author="Scott T" box="[536,671,976,1002]" pageId="16" pageNumber="17" pagination="109 - 135" refId="ref20637" refString="Scott T (1903) On some new and rare Crustacea collected at various times in connection with the investigations of the Fishery Board for Scotland. Reports of the Fishery Board for Scotland, Edinburgh 21 (3): 109 - 135." type="journal article" year="1903">Scott, 1903</bibRefCitation>
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</taxonomicName>
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,
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<taxonomicName authority="Sars, 1908 a" authorityName="Sars" authorityYear="1908" box="[685,947,976,1002]" class="Maxillopoda" family="Laophontidae" genus="Laophonte" kingdom="Animalia" order="Harpacticoida" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="proxima">
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<emphasis box="[685,807,977,1002]" italics="true" pageId="16" pageNumber="17">L. proxima</emphasis>
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<bibRefCitation author="Sars GO" box="[815,947,976,1002]" pageId="16" pageNumber="17" pagination="241 - 256" refId="ref20226" refString="Sars GO (1908 a) Copepoda Harpacticoida. Parts XXI & XXII. Laophontidae (continued). An account of the Crustacea of Norway, with short descriptions and figures of all the species 5: 241 - 256 + pls. 161 - 176." type="journal article" year="1908">Sars, 1908a</bibRefCitation>
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</taxonomicName>
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and
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<taxonomicName authority="Sars, 1920 a" authorityName="Sars" authorityYear="1920" class="Maxillopoda" family="Laophontidae" genus="Laophonte" kingdom="Animalia" order="Harpacticoida" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="abbreviata">
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<emphasis italics="true" pageId="16" pageNumber="17">L. abbreviata</emphasis>
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<bibRefCitation author="Sars GO" box="[203,338,1012,1038]" pageId="16" pageNumber="17" pagination="53 - 72" refId="ref20419" refString="Sars GO (1920 a) Copepoda Supplement. Parts V & VI. Harpacticoida (continued). An account of the Crustacea of Norway, with short descriptions and figures of all the species 7: 53 - 72 + pls. 33 - 48." type="journal article" year="1920">Sars, 1920a</bibRefCitation>
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</taxonomicName>
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.
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<bibRefCitation author="Noodt W" box="[350,510,1011,1038]" pageId="16" pageNumber="17" pagination="105 - 142" refId="ref19739" refString="Noodt W (1952) Marine Harpacticiden (Cop.) aus dem eulitoralen Sandstrand der Insel Sylt. Abhandlungen der mathematisch-naturwissenschaftlichen Klasse. Akademie der Wissenschaften und der Literatur in Mainz 1952 (3): 105 - 142." type="journal article" year="1952">Noodt (1952)</bibRefCitation>
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removed
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<taxonomicName authorityName="T. Scott" authorityYear="1903" box="[623,729,1011,1037]" class="Maxillopoda" family="Laophontidae" genus="Laophonte" kingdom="Animalia" order="Harpacticoida" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="gracilis">
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<emphasis box="[623,729,1011,1037]" italics="true" pageId="16" pageNumber="17">L. gracilis</emphasis>
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</taxonomicName>
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and designated it as the
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<typeStatus box="[1014,1062,1012,1038]" pageId="16" pageNumber="17">type</typeStatus>
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(by original designation) of a new genus
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<taxonomicName authority="Noodt, 1952" authorityName="Noodt" authorityYear="1952" box="[556,791,1046,1073]" class="Maxillopoda" family="Laophontidae" genus="Pilifera" kingdom="Animalia" order="Harpacticoida" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">
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<emphasis box="[556,635,1047,1073]" italics="true" pageId="16" pageNumber="17">Pilifera</emphasis>
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<bibRefCitation author="Noodt W" box="[643,791,1046,1073]" pageId="16" pageNumber="17" pagination="105 - 142" refId="ref19739" refString="Noodt W (1952) Marine Harpacticiden (Cop.) aus dem eulitoralen Sandstrand der Insel Sylt. Abhandlungen der mathematisch-naturwissenschaftlichen Klasse. Akademie der Wissenschaften und der Literatur in Mainz 1952 (3): 105 - 142." type="journal article" year="1952">Noodt, 1952</bibRefCitation>
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</taxonomicName>
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which has remained monotypic since its proposal. New species were added to
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<taxonomicName box="[708,931,1082,1108]" class="Maxillopoda" family="Laophontidae" genus="Pseudonychocamptus" kingdom="Animalia" order="Harpacticoida" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">
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<emphasis box="[708,931,1082,1108]" italics="true" pageId="16" pageNumber="17">Pseudonychocamptus</emphasis>
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</taxonomicName>
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by
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<bibRefCitation author="Lang K" box="[970,1104,1082,1109]" pageId="16" pageNumber="17" pagination="1 - 560" refId="ref19259" refString="Lang K (1965) Copepoda Harpacticoida from the Californian Pacific coast. Kungliga Svenska Vetenskapsakademiens Handlingar (4) 10 (2): 1 - 560." type="journal article" year="1965">Lang (1965</bibRefCitation>
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:
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<taxonomicName authorityName="Lang" authorityYear="1965" box="[140,314,1118,1143]" class="Maxillopoda" family="Laophontidae" genus="Pseudonychocamptus" kingdom="Animalia" order="Harpacticoida" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="paraproximus">
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<emphasis box="[140,314,1118,1143]" italics="true" pageId="16" pageNumber="17">P. paraproximus</emphasis>
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</taxonomicName>
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and
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<taxonomicName authorityName="Lang" authorityYear="1965" box="[373,480,1117,1143]" class="Maxillopoda" family="Laophontidae" genus="Pseudonychocamptus" kingdom="Animalia" order="Harpacticoida" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="spinifer">
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<emphasis box="[373,480,1117,1143]" italics="true" pageId="16" pageNumber="17">P. spinifer</emphasis>
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</taxonomicName>
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),
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<bibRefCitation author="Hamond R" box="[505,689,1117,1143]" pageId="16" pageNumber="17" pagination="172 - 176" refId="ref18534" refString="Hamond R (1968) Pseudonychocamptus carthyi n. sp. (Harpacticoida) from Hunstanton, Norfolk. Crustaceana supplement 1: 172 - 176." type="journal article" year="1968">Hamond (1968</bibRefCitation>
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:
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<taxonomicName authorityName="Hamond" authorityYear="1968" box="[701,802,1117,1143]" class="Maxillopoda" family="Laophontidae" genus="Pseudonychocamptus" kingdom="Animalia" order="Harpacticoida" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="carthyi">
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<emphasis box="[701,802,1117,1143]" italics="true" pageId="16" pageNumber="17">P. carthyi</emphasis>
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</taxonomicName>
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),
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<bibRefCitation author="Apostolov A & Petkovski T" pageId="16" pageNumber="17" pagination="24 - 35" refId="ref17648" refString="Apostolov A, Petkovski T (1980) Copepodes Harpacticoides de la mer Adriatique (region de Rovigne). Acta zoologica bulgarica 15: 24 - 35." type="journal article" year="1980">Apostolov and Petkovski (1980</bibRefCitation>
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:
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<taxonomicName authorityName="Apostolov & Petkovski" authorityYear="1980" box="[222,347,1153,1178]" class="Maxillopoda" family="Laophontidae" genus="Pseudonychocamptus" kingdom="Animalia" order="Harpacticoida" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="marinovi">
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<emphasis box="[222,347,1153,1178]" italics="true" pageId="16" pageNumber="17">P. marinovi</emphasis>
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</taxonomicName>
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),
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<bibRefCitation author="Ceccherelli VU" box="[371,576,1152,1178]" pageId="16" pageNumber="17" pagination="155 - 171" refId="ref17934" refString="Ceccherelli VU (1988) Trois nouvelles especes de Copepodes Harpacticoides des lagunes de l'Adriatique du Nord, Italie. Vie et Milieu 38 (2): 155 - 171." type="journal article" year="1988">Ceccherelli (1988</bibRefCitation>
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:
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<taxonomicName authorityName="Ceccherelli" authorityYear="1988" box="[587,708,1152,1178]" class="Maxillopoda" family="Laophontidae" genus="Pseudonychocamptus" kingdom="Animalia" order="Harpacticoida" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="colomboi">
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<emphasis box="[587,708,1152,1178]" italics="true" pageId="16" pageNumber="17">P. colomboi</emphasis>
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</taxonomicName>
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) and
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<bibRefCitation author="Apostolov A" box="[774,967,1152,1178]" pageId="16" pageNumber="17" pagination="5 - 33" refId="ref17614" refString="Apostolov A (2008) Harpacticoides (Crustacea, Copepoda) de la mer Egee (plages de Kavala, Grece du nord). Historia naturalis bulgarica 19: 5 - 33." type="journal article" year="2008">Apostolov (2008</bibRefCitation>
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:
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<taxonomicName authorityName="Huys & Lee" authorityYear="2009" baseAuthorityName="Apostolov" baseAuthorityYear="2008" box="[978,1090,1152,1178]" class="Maxillopoda" family="Laophontidae" genus="Paralaophonte" kingdom="Animalia" order="Harpacticoida" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="kolarovi">
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<emphasis box="[978,1090,1152,1178]" italics="true" pageId="16" pageNumber="17">P. kolarovi</emphasis>
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</taxonomicName>
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).
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</paragraph>
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<paragraph blockId="16.[140,1108,906,1672]" lastBlockId="17.[140,1108,167,545]" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17">
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Both
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<bibRefCitation author="Sars GO" box="[249,387,1188,1214]" pageId="16" pageNumber="17" pagination="241 - 256" refId="ref20226" refString="Sars GO (1908 a) Copepoda Harpacticoida. Parts XXI & XXII. Laophontidae (continued). An account of the Crustacea of Norway, with short descriptions and figures of all the species 5: 241 - 256 + pls. 161 - 176." type="journal article" year="1908">Sars (1908a)</bibRefCitation>
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and
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<bibRefCitation author="Wilson CB" box="[446,609,1187,1214]" pageId="16" pageNumber="17" pagination="1 - 635" refId="ref21130" refString="Wilson CB (1932) The copepods of the Woods Hole region, Massachusetts. Bulletin of the United States National Museum 158: 1 - 635." type="journal article" year="1932">Wilson (1932)</bibRefCitation>
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described
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<taxonomicName authorityName="Sars" authorityYear="1908" box="[730,935,1187,1213]" class="Maxillopoda" family="Laophontidae" genus="Laophonte" kingdom="Animalia" order="Harpacticoida" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="proxima">
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<emphasis box="[730,935,1187,1213]" italics="true" pageId="16" pageNumber="17">Laophonte proxima</emphasis>
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</taxonomicName>
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on the basis of females only (the latter author from a freshwater locality!) whereas
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<bibRefCitation author="Klie W" box="[875,1002,1223,1249]" pageId="16" pageNumber="17" pagination="329 - 386" refId="ref19072" refString="Klie W (1929) Die Copepoda Harpacticoida der sudlichen und westlichen Ostsee mit besonderer Berucksichtigung der Sandfauna der Kieler Bucht. Zoologische Jahrbucher fur Systematik 57 (3 - 4): 329 - 386." type="journal article" year="1929">Klie (1929)</bibRefCitation>
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provided the first illustrations of the male, including the P5 which he figured with one seta on the endopodal lobe.
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<bibRefCitation author="Lang K" box="[325,463,1293,1320]" pageId="16" pageNumber="17" pagination="1 - 560" refId="ref19259" refString="Lang K (1965) Copepoda Harpacticoida from the Californian Pacific coast. Kungliga Svenska Vetenskapsakademiens Handlingar (4) 10 (2): 1 - 560." type="journal article" year="1965">Lang (1965)</bibRefCitation>
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distinguished the closely related
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<taxonomicName class="Maxillopoda" family="Laophontidae" genus="Pseudonychocamptus" kingdom="Animalia" order="Harpacticoida" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="proximus">
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<emphasis italics="true" pageId="16" pageNumber="17">Pseudonychocamptus proximus</emphasis>
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</taxonomicName>
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and
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<taxonomicName authorityName="Lang" authorityYear="1965" box="[247,414,1329,1354]" class="Maxillopoda" family="Laophontidae" genus="Pseudonychocamptus" kingdom="Animalia" order="Harpacticoida" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="paraproximus">
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<emphasis box="[247,414,1329,1354]" italics="true" pageId="16" pageNumber="17">P. paraproximus</emphasis>
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</taxonomicName>
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on the basis of morphometric differences in the distal segment of the P4 exopod and P5 baseoendopod in the female and the P5 exopod in the male. An additional differentiating character used in his key referred to the number of setae on the male P5 baseoendopod.
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<bibRefCitation author="Hamond R" box="[410,593,1434,1461]" pageId="16" pageNumber="17" pagination="172 - 176" refId="ref18534" refString="Hamond R (1968) Pseudonychocamptus carthyi n. sp. (Harpacticoida) from Hunstanton, Norfolk. Crustaceana supplement 1: 172 - 176." type="journal article" year="1968">Hamond (1968)</bibRefCitation>
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and
|
||
<bibRefCitation author="Ceccherelli VU" box="[650,853,1434,1460]" pageId="16" pageNumber="17" pagination="155 - 171" refId="ref17934" refString="Ceccherelli VU (1988) Trois nouvelles especes de Copepodes Harpacticoides des lagunes de l'Adriatique du Nord, Italie. Vie et Milieu 38 (2): 155 - 171." type="journal article" year="1988">Ceccherelli (1988)</bibRefCitation>
|
||
followed Lang’s judgement, however
|
||
<bibRefCitation author="Mielke W" box="[307,466,1469,1496]" pageId="16" pageNumber="17" pagination="1 - 134" refId="ref19479" refString="Mielke W (1975) Systematik der Copepoda eines Sandstrandes der Nordseeinsel Sylt. Mikrofauna des Meeresbodens 52: 1 - 134." type="journal article" year="1975">Mielke (1975)</bibRefCitation>
|
||
, in his redescription of the male of
|
||
<taxonomicName box="[847,966,1470,1495]" class="Maxillopoda" family="Laophontidae" genus="Pseudonychocamptus" kingdom="Animalia" order="Harpacticoida" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="proximus">
|
||
<emphasis box="[847,966,1470,1495]" italics="true" pageId="16" pageNumber="17">P. proximus</emphasis>
|
||
</taxonomicName>
|
||
, pointed out that the latter has two setae on the P5 baseoendopod (as in all other congeners), rendering the distinctiveness of
|
||
<taxonomicName authorityName="Lang" authorityYear="1965" box="[372,539,1540,1565]" class="Maxillopoda" family="Laophontidae" genus="Pseudonychocamptus" kingdom="Animalia" order="Harpacticoida" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="paraproximus">
|
||
<emphasis box="[372,539,1540,1565]" italics="true" pageId="16" pageNumber="17">P. paraproximus</emphasis>
|
||
</taxonomicName>
|
||
doubtful. Although we have followed
|
||
<bibRefCitation author="Bodin P" box="[958,1107,1539,1566]" pageId="16" pageNumber="17" pagination="1 - 304" refId="ref17734" refString="Bodin P (1997) Catalogue of the new marine harpacticoid copepods (1997 edition). Documents de Travail, Institut royal des Sciences naturelles de Belgique 89: 1 - 304." type="journal article" year="1997">Bodin (1997)</bibRefCitation>
|
||
and
|
||
<bibRefCitation author="Wells JBJ" box="[190,333,1575,1601]" pageId="16" pageNumber="17" pagination="1 - 872" refId="ref20947" refString="Wells JBJ (2007) An annotated checklist and keys to the species of Copepoda Harpacticoida (Crustacea). Zootaxa 1568: 1 - 872." type="journal article" year="2007">Wells (2007)</bibRefCitation>
|
||
in considering the latter a species of uncertain status (here ranked as
|
||
<emphasis box="[140,327,1610,1636]" italics="true" pageId="16" pageNumber="17">species inquirenda</emphasis>
|
||
), we have nevertheless included it in the key below, based on the interspecific differences displayed in the length/width ratio of the male P5 exopod (
|
||
<tableCitation box="[1007,1094,1645,1671]" captionStart="Table 2" captionStartId="18.[140,200,166,188]" captionText="Table 2. Setal number and morphometric differences between species of Pseudonychocamptus (CR = caudal ramus; exp = exopod; enp = endopod; benp = baseoendopod).Length/width ratios based on Chislenko (1967)a, Sars (1908a)b, Sars (1920a)c, Lang (1965)d, Letova (1982)e, Apostolov and Petkovski (1980)f and Ceccherelli (1988)g." pageId="16" pageNumber="17">Table 2</tableCitation>
|
||
). Recently,
|
||
<bibRefCitation author="Apostolov A" box="[245,436,167,193]" pageId="17" pageNumber="18" pagination="5 - 33" refId="ref17614" refString="Apostolov A (2008) Harpacticoides (Crustacea, Copepoda) de la mer Egee (plages de Kavala, Grece du nord). Historia naturalis bulgarica 19: 5 - 33." type="journal article" year="2008">Apostolov (2008)</bibRefCitation>
|
||
added a new species,
|
||
<taxonomicName authorityName="Huys & Lee" authorityYear="2009" baseAuthorityName="Apostolov" baseAuthorityYear="2008" box="[672,780,167,193]" class="Maxillopoda" family="Laophontidae" genus="Paralaophonte" kingdom="Animalia" order="Harpacticoida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="kolarovi">
|
||
<emphasis box="[672,780,167,193]" italics="true" pageId="17" pageNumber="18">P. kolarovi</emphasis>
|
||
</taxonomicName>
|
||
, based on two males collected from the Kavala beach (
|
||
<collectingCountry box="[405,483,202,228]" name="Greece" pageId="17" pageNumber="18">Greece</collectingCountry>
|
||
) in the
|
||
<collectingRegion box="[573,652,202,228]" country="Greece" name="Voreio Aigaio" pageId="17" pageNumber="18">Aegean</collectingRegion>
|
||
Sea. The author claimed that the species occupied an isolated position in the genus on account of the structure of the caudal rami and P1–P5. The 2-segmented P4 endopod (with one inner and two distal setae), the presence of 5 setae on the P5 exopod and the sexually dimorphic distal inner spine on the P2 endopod clearly exclude
|
||
<taxonomicName authorityName="Huys & Lee" authorityYear="2009" baseAuthorityName="Apostolov" baseAuthorityYear="2008" box="[414,524,343,369]" class="Maxillopoda" family="Laophontidae" genus="Paralaophonte" kingdom="Animalia" order="Harpacticoida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="kolarovi">
|
||
<emphasis box="[414,524,343,369]" italics="true" pageId="17" pageNumber="18">P. kolarovi</emphasis>
|
||
</taxonomicName>
|
||
from the genus
|
||
<taxonomicName box="[710,925,343,369]" class="Maxillopoda" family="Laophontidae" genus="Pseudonychocamptus" kingdom="Animalia" order="Harpacticoida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[710,925,343,369]" italics="true" pageId="17" pageNumber="18">Pseudonychocamptus</emphasis>
|
||
</taxonomicName>
|
||
and particularly the latter character unequivocally points to a relationship with the genera
|
||
<taxonomicName authority="Lang, 1944" authorityName="Lang" authorityYear="1944" class="Maxillopoda" family="Laophontidae" genus="Paralaophonte" kingdom="Animalia" order="Harpacticoida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[959,1107,378,404]" italics="true" pageId="17" pageNumber="18">Paralaophonte</emphasis>
|
||
<bibRefCitation author="Lang K" box="[140,267,413,440]" pageId="17" pageNumber="18" refId="ref19213" refString="Lang K (1944) Monographie der Harpacticiden (Vorlaufige Mitteilung). Almqvist & Wiksells Boktryckeri Ab, Uppsala, 39 pp." type="book" year="1944">Lang, 1944</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName authority="Jakobi, 1953" authorityName="Jakobi" authorityYear="1953" box="[326,638,413,440]" class="Maxillopoda" family="Laophontidae" genus="Loureirophonte" kingdom="Animalia" order="Harpacticoida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[326,484,413,439]" italics="true" pageId="17" pageNumber="18">Loureirophonte</emphasis>
|
||
<bibRefCitation author="Jakobi H" box="[493,638,413,440]" pageId="17" pageNumber="18" pagination="47 - 60" refId="ref19016" refString="Jakobi H (1953) Novos Laophontidae (Copepoda-Crustacea) da costa Brasileira. (Neue Laophontiden von der Brasilianischen Kuste). Dusenia 4: 47 - 60." type="journal article" year="1953">Jakobi, 1953</bibRefCitation>
|
||
</taxonomicName>
|
||
. The species is here formally placed in the genus
|
||
<taxonomicName authorityName="Lang" authorityYear="1944" box="[210,358,448,474]" class="Maxillopoda" family="Laophontidae" genus="Paralaophonte" kingdom="Animalia" order="Harpacticoida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[210,358,448,474]" italics="true" pageId="17" pageNumber="18">Paralaophonte</emphasis>
|
||
</taxonomicName>
|
||
as
|
||
<taxonomicName authorityName="Huys & Lee" authorityYear="2009" baseAuthorityName="Apostolov" baseAuthorityYear="2008" box="[394,634,448,474]" class="Maxillopoda" family="Laophontidae" genus="Paralaophonte" kingdom="Animalia" order="Harpacticoida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="kolarovi" status="comb. n.">
|
||
<emphasis box="[394,634,448,474]" italics="true" pageId="17" pageNumber="18">Paralaophonte kolarovi</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel box="[648,745,448,475]" pageId="17" pageNumber="18" rank="species">comb. n.</taxonomicNameLabel>
|
||
(see below). The six valid species currently recognized in the genus, and the problematic species
|
||
<taxonomicName authorityName="Lang" authorityYear="1965" box="[814,980,484,509]" class="Maxillopoda" family="Laophontidae" genus="Pseudonychocamptus" kingdom="Animalia" order="Harpacticoida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="paraproximus">
|
||
<emphasis box="[814,980,484,509]" italics="true" pageId="17" pageNumber="18">P. paraproximus</emphasis>
|
||
</taxonomicName>
|
||
, can be differentiated by the key below and the character states tabulated in
|
||
<tableCitation box="[848,931,519,545]" captionStart="Table 2" captionStartId="18.[140,200,166,188]" captionText="Table 2. Setal number and morphometric differences between species of Pseudonychocamptus (CR = caudal ramus; exp = exopod; enp = endopod; benp = baseoendopod).Length/width ratios based on Chislenko (1967)a, Sars (1908a)b, Sars (1920a)c, Lang (1965)d, Letova (1982)e, Apostolov and Petkovski (1980)f and Ceccherelli (1988)g." pageId="17" pageNumber="18">Table 2</tableCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="17" pageNumber="18" type="key">
|
||
<keyStep pageId="17" pageNumber="18">
|
||
<paragraph blockId="17.[140,1108,589,1215]" box="[140,1107,589,617]" pageId="17" pageNumber="18">
|
||
<keyLead box="[140,1107,589,617]" pageId="17" pageNumber="18">
|
||
1. P3 enp-2 with 4 setae in
|
||
<specimenCount box="[528,543,590,616]" pageId="17" pageNumber="18" type="female">♀</specimenCount>
|
||
and 2 setae in ♁ ...............................................
|
||
<emphasis bold="true" box="[1092,1107,589,615]" pageId="17" pageNumber="18">2</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="17.[140,1108,589,1215]" box="[140,1107,624,652]" pageId="17" pageNumber="18">
|
||
<keyLead box="[140,1107,624,652]" pageId="17" pageNumber="18">
|
||
– P3 enp-2 with 5 setae in
|
||
<specimenCount box="[528,543,625,651]" pageId="17" pageNumber="18" type="female">♀</specimenCount>
|
||
and 3–4 setae in ♁ ...........................................
|
||
<emphasis bold="true" box="[1092,1107,624,650]" pageId="17" pageNumber="18">3</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="17" pageNumber="18">
|
||
<paragraph blockId="17.[140,1108,589,1215]" pageId="17" pageNumber="18">
|
||
<keyLead pageId="17" pageNumber="18">
|
||
2. P5 exopod
|
||
<specimenCount box="[372,387,660,686]" pageId="17" pageNumber="18" type="female">♀</specimenCount>
|
||
at least twice longer than wide, with 5 setae; P5 exopod ♁ with straight outer margin ....................................................................
|
||
<taxonomicName authorityName="Boeck" authorityYear="1873" box="[989,1093,694,721]" class="Maxillopoda" family="Laophontidae" genus="Laophonte" kingdom="Animalia" order="Harpacticoida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="koreni">
|
||
<emphasis bold="true" box="[989,1093,694,721]" italics="true" pageId="17" pageNumber="18">P. koreni</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[1100,1108,694,708]" pageId="17" pageNumber="18">
|
||
<superScript attach="right" box="[1100,1108,694,708]" fontSize="6" pageId="17" pageNumber="18">a</superScript>
|
||
</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="17.[140,1108,589,1215]" pageId="17" pageNumber="18">
|
||
<keyLead pageId="17" pageNumber="18">
|
||
– P5 exopod
|
||
<specimenCount box="[376,391,730,756]" pageId="17" pageNumber="18" type="female">♀</specimenCount>
|
||
about 1.3 times longer than wide, with 6 setae; P5 exopod ♁ with convex outer margin .............................................................
|
||
<taxonomicName authorityName="Lang" authorityYear="1965" box="[989,1107,765,791]" class="Maxillopoda" family="Laophontidae" genus="Pseudonychocamptus" kingdom="Animalia" order="Harpacticoida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="spinifer">
|
||
<emphasis bold="true" box="[989,1107,765,791]" italics="true" pageId="17" pageNumber="18">P. spinifer</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="17" pageNumber="18">
|
||
<paragraph blockId="17.[140,1108,589,1215]" box="[140,1107,800,827]" pageId="17" pageNumber="18">
|
||
<keyLead box="[140,1107,800,827]" pageId="17" pageNumber="18">
|
||
3. Caudal ramus longer than wide ..................................................................
|
||
<emphasis bold="true" box="[1092,1107,800,826]" pageId="17" pageNumber="18">4</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="17.[140,1108,589,1215]" box="[140,1107,835,862]" pageId="17" pageNumber="18">
|
||
<keyLead box="[140,1107,835,862]" pageId="17" pageNumber="18">
|
||
– Caudal ramus wider than long ...................................................
|
||
<taxonomicName authorityName="Apostolov & Petkovski" authorityYear="1980" box="[970,1107,835,862]" class="Maxillopoda" family="Laophontidae" genus="Pseudonychocamptus" kingdom="Animalia" order="Harpacticoida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="marinovi">
|
||
<emphasis bold="true" box="[970,1107,835,862]" italics="true" pageId="17" pageNumber="18">P. marinovi</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="17" pageNumber="18">
|
||
<paragraph blockId="17.[140,1108,589,1215]" pageId="17" pageNumber="18">
|
||
<keyLead pageId="17" pageNumber="18">
|
||
4. P5 exopod
|
||
<specimenCount box="[374,389,871,897]" pageId="17" pageNumber="18" type="female">♀</specimenCount>
|
||
with 1 naked and 5 pinnate setae; P2 enp-2, P3 enp-2 and P5 exopod of ♁ with 4 setae ............................................................................
|
||
<emphasis bold="true" box="[1092,1107,906,932]" pageId="17" pageNumber="18">5</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="17.[140,1108,589,1215]" pageId="17" pageNumber="18">
|
||
<keyLead pageId="17" pageNumber="18">
|
||
– P5 exopod
|
||
<specimenCount box="[376,391,942,968]" pageId="17" pageNumber="18" type="female">♀</specimenCount>
|
||
with 6 naked setae; P2 enp-2, P3 enp-2 and P5 exopod of ♁ with 3 setae.................................................................................
|
||
<taxonomicName authorityName="Ceccherelli" authorityYear="1988" box="[973,1107,976,1003]" class="Maxillopoda" family="Laophontidae" genus="Pseudonychocamptus" kingdom="Animalia" order="Harpacticoida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="colomboi">
|
||
<emphasis bold="true" box="[973,1107,976,1003]" italics="true" pageId="17" pageNumber="18">P. colomboi</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="17" pageNumber="18">
|
||
<paragraph blockId="17.[140,1108,589,1215]" pageId="17" pageNumber="18">
|
||
<keyLead pageId="17" pageNumber="18">
|
||
5. Body elongate, length 3–4 times maximum width of cephalothorax; all setae of P5 exopod
|
||
<specimenCount box="[403,418,1047,1073]" pageId="17" pageNumber="18" type="female">♀</specimenCount>
|
||
marginal............................................................................
|
||
<emphasis bold="true" box="[1092,1107,1046,1072]" pageId="17" pageNumber="18">6</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="17.[140,1108,589,1215]" pageId="17" pageNumber="18">
|
||
<keyLead pageId="17" pageNumber="18">
|
||
– Body short, length about 1.85 times maximum width of cephalothorax; P5 exopod
|
||
<specimenCount box="[335,350,1118,1144]" pageId="17" pageNumber="18" type="female">♀</specimenCount>
|
||
with 4 marginal and 2 surface setae .......................
|
||
<taxonomicName box="[940,1108,1116,1143]" class="Maxillopoda" family="Laophontidae" genus="Paralaophonte" kingdom="Animalia" order="Harpacticoida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="abbreviatus">
|
||
<emphasis bold="true" box="[940,1108,1116,1143]" italics="true" pageId="17" pageNumber="18">P. abbreviatus</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="17" pageNumber="18">
|
||
<paragraph blockId="17.[140,1108,589,1215]" box="[140,1107,1152,1180]" pageId="17" pageNumber="18">
|
||
<keyLead box="[140,1107,1152,1180]" pageId="17" pageNumber="18">
|
||
6. P5 exopod ♁ about 2.7 times longer than wide .........................
|
||
<taxonomicName box="[970,1107,1152,1179]" class="Maxillopoda" family="Laophontidae" genus="Pseudonychocamptus" kingdom="Animalia" order="Harpacticoida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="proximus">
|
||
<emphasis bold="true" box="[970,1107,1152,1179]" italics="true" pageId="17" pageNumber="18">P. proximus</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="17.[140,1108,589,1215]" box="[140,1107,1187,1215]" pageId="17" pageNumber="18">
|
||
<keyLead box="[140,1107,1187,1215]" pageId="17" pageNumber="18">
|
||
– P5 exopod ♁ about 3.25 times longer than wide ................
|
||
<taxonomicName authorityName="Lang" authorityYear="1965" box="[914,1107,1187,1214]" class="Maxillopoda" family="Laophontidae" genus="Pseudonychocamptus" kingdom="Animalia" order="Harpacticoida" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="paraproximus">
|
||
<emphasis bold="true" box="[914,1107,1187,1214]" italics="true" pageId="17" pageNumber="18">P. paraproximus</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |