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<document id="1EFA76DA7131825FE42C51D8AB882432" ID-DOI="10.3897/zookeys.1212.126536" ID-ZooBank="4A550F74-C788-4282-A731-B8405CA8B8F5" ID-publisher-id="126536" URI-arpha="FC3213C5-BED0-512A-9C14-5F0A39ADDBC7" URI-zoobank="http://zoobank.org/4A550F74-C788-4282-A731-B8405CA8B8F5" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" XM.taxonomicNames_approvedBy="admin" article-type="research-article" checkinTime="1726239268422" checkinUser="pensoft" docAuthor="Srisonchai, Ruttapon, Likhitrakarn, Natdanai, Sutcharit, Chirasak &amp; Wesener, Thomas" docDate="2024" docId="F69B1A1DC0F254338EC1CF3319469C6A" docLanguage="en" docName="ZooKeys 1212: 29-64" docOrigin="ZooKeys 1212" docSource="https://zookeys.pensoft.net/article/126536/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Zephronia macula Srisonchai &amp; Wesener 2024, sp. nov." docType="treatment" docUuid="84892BB6-C242-4C03-8639-39F8A0D97683" docUuidSource="ZooBank" docVersion="2" dtd-version="3.0" lastPageNumber="64" masterDocId="FC3213C5BED0512A9C145F0A39ADDBC7" masterDocTitle="Integrative taxonomy reveals two new giant pill-millipedes of the genus Zephronia Gray, 1832 from eastern Thailand (Diplopoda, Sphaerotheriida, Zephroniidae)" masterLastPageNumber="64" masterPageNumber="29" pageNumber="29" updateTime="1726249556959" updateUser="admin">
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<mods:title id="069EAE2E1F00352F57C6756239330141">Integrative taxonomy reveals two new giant pill-millipedes of the genus Zephronia Gray, 1832 from eastern Thailand (Diplopoda, Sphaerotheriida, Zephroniidae)</mods:title>
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<mods:name id="BEEDE6CC4B9CD225B184B8A4DDEB0511" type="personal">
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<mods:namePart id="C94166F3781C0953569B8359A8AC54D9">Srisonchai, Ruttapon</mods:namePart>
<mods:nameIdentifier id="8FBB8A715ABF3EAB773235EFE935BC5A" type="ORCID">0000-0002-7142-0999</mods:nameIdentifier>
<mods:affiliation id="173AC66A456ED07AC5359BB587AE773C">Department of Biology, Faculty of Science, Khon Kaen University, Khon Kaen 40002, Thailand</mods:affiliation>
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<mods:namePart id="4A0770BF3AA8D5F8AC6796F46ACDF638">Likhitrakarn, Natdanai</mods:namePart>
<mods:nameIdentifier id="04D897864FCC72FBFA659D04D9C3B32A" type="ORCID">0000-0002-1306-317X</mods:nameIdentifier>
<mods:affiliation id="DE667A3066F6563FC6F1BF3F840E8B2D">Program of Agriculture, Faculty of Agricultural Production, Maejo University, Chiang Mai 50290, Thailand</mods:affiliation>
</mods:name>
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<mods:namePart id="9769FF99CC73E506530AF46357F9E2DE">Sutcharit, Chirasak</mods:namePart>
<mods:nameIdentifier id="9A4D2A1E7A89D7E266D4C2706BA4CBBB" type="ORCID">0000-0001-7670-9540</mods:nameIdentifier>
<mods:affiliation id="B0CF48795253E62B548EE179B83E4A2A">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Phayathai Road, Patumwan, Bangkok 10330, Thailand</mods:affiliation>
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<mods:namePart id="22C7F4FAF133B0965B067931CD964ACF">Wesener, Thomas</mods:namePart>
<mods:nameIdentifier id="E2BEADEB7CBFF42F635980EDEFE70436" type="ORCID">0000-0002-2028-3541</mods:nameIdentifier>
<mods:affiliation id="9E467898CE96E6155BB3B9ABD2FC989F">Zoological Research Museum Alexander Koenig, Leibniz Institute for the Study of Biodiversity Change (LIB), Adenauerallee 160, D- 53113 Bonn, Germany</mods:affiliation>
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<mods:date id="499A02FEE02D14B54082A930523BC878">2024</mods:date>
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<mods:number id="6C46C4E4978C9B380AAC018AC47EC7F9">1212</mods:number>
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<treatment id="F69B1A1DC0F254338EC1CF3319469C6A" ID-arpha="F69B1A1D-C0F2-5433-8EC1-CF3319469C6A" ID-zoobank="https://zoobank.org/84892BB6-C242-4C03-8639-39F8A0D97683" LSID="urn:lsid:zoobank.org:act:84892BB6-C242-4C03-8639-39F8A0D97683" LSID-ZBK="urn:lsid:zoobank.org:act:84892BB6-C242-4C03-8639-39F8A0D97683" httpUri="http://treatment.plazi.org/id/F69B1A1DC0F254338EC1CF3319469C6A">
<subSubSection id="40689C1FAF93413F7CAA3893A1989AF8" type="nomenclature">
<paragraph id="7D5B8783E6E47C1204C3B39D1660DBC6">
<taxonomicName id="AC379E51472C0154EFE8E71CACF8F1B6" authority="Srisonchai &amp; Wesener" authorityName="Srisonchai &amp; Wesener" authorityYear="2024" class="Diplopoda" family="Zephroniidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" phylum="Arthropoda" rank="species" species="macula" status="sp. nov.">
<emphasis id="3E6DC4602B597AE333751327CC893607" italics="true">Zephronia macula</emphasis>
Srisonchai &amp; Wesener
</taxonomicName>
<taxonomicNameLabel id="D899AF72513584F7EA24EC7BCB9D8536" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="38EAAE293B8836E2D1D16E58E5205547" type="description">
<paragraph id="F378D338FB54B95E4AC2115021202529">
<figureCitation id="90A23F663137EFD52001E0FA297AE58B" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Photograph of live specimens of Zephronia macula sp. nov. and habitats A G paratypes (CUMZ - MYR 00016) H habitat at Thaeo Khlong Khlung monastery (granitic rocks) I habitat at Phet Pho Thong cave (limestone). Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1212.126536.figure8" httpUri="https://binary.pensoft.net/fig/1133561">Figs 8</figureCitation>
,
<figureCitation id="3E1A74BE706F398F5A253A2ED18B68C7" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Zephronia macula sp. nov., ♂ paratype (CUMZ - MYR 0016) — SEM A head, collum and antenna, dorsal view B ommatidia, inset: organ of Tömösváry C antenna, anterior view D antennal disc, anterior view E apical cones, lateral view F, I gnathochilarium, posterior and ventral views, respectively G sensory cones on central pad, ventral view H sensory cones on inner palpi, black and white arrows point to different types of cones, ventral view J epipharynx, ventral view, K mandible, mesal view. Abbreviations: 3 it = 3 - combed inner tooth, co = condylus, Cp = central pad, ct = central tooth, et = external tooth, Et = external tooth, Ia = inner area, il = incisura lateralis, ip = inner palpi, ll = lamellae linguales, me = mentum, mp = molar plate, pl = pectinate lamellae, rsp = row of spines, sc = sensory cone, st = stipites. Scale bars in millimetres." figureDoi="10.3897/zookeys.1212.126536.figure9" httpUri="https://binary.pensoft.net/fig/1133562">9</figureCitation>
,
<figureCitation id="089FBF80AFBC887F05BA0DCBB908F4BA" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Zephronia macula sp. nov., ♂ paratype (CUMZ - MYR 0016) — SEM A surface of collum B, C thoracic shield, all in dorsal views (anterior margin and surface, respectively) D G midbody tergite, all in dorsal views (overview, anterior margin, surface, and posterior margin, respectively) H J anal shield, all in dorsal views (overview, anterior margin and cuticular impressions, respectively) K M anal shield, underside, all in ventral views (surface, posterior margin, and overview, respectively). Scale bars in millimetres." figureDoi="10.3897/zookeys.1212.126536.figure10" httpUri="https://binary.pensoft.net/fig/1133563">10</figureCitation>
,
<figureCitation id="46674917B6D5DA863633B1D6D7836981" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Zephronia macula sp. nov. — SEM A G, L ♂ paratype (CUMZ - MYR 0016) J, K ♀ paratype (CUMZ - MYR 0015) A collum B seventh pleurite C subanal plate D legs 1 E stigmatic plate 1 F legs 2 with gonopores G leg 10 (arrow points to femoral ridge) H coxal process on leg 10 I small teeth on femur J leg 2 with vulvae K vulvae L tarsus 10. Abbreviation: bu = bursa, cx = coxa, fe = femur, op = operculum, po = postfemur, pre = prefemur, st-pl = stigmatic plate, ta = tarsus, ti = tibia. Scale bars in millimetres." figureDoi="10.3897/zookeys.1212.126536.figure11" httpUri="https://binary.pensoft.net/fig/1133564">11</figureCitation>
,
<figureCitation id="2DD80B50ED8482D2FFEBBE09577CDEC1" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Zephronia macula sp. nov., ♂ paratype (CUMZ - MYR 0016) — SEM A D overview of telopods (anterior view, lateral view with emphasis on anterior telopod, posterior view and ventral view, respectively) E G anterior telopod (telopoditomeres 24, movable finger, and crenulated teeth, respectively) H J posterior telopod (sclerotize spots on immovable finger, telopoditomeres 24 with emphasis on membranous lobe and crenulated teeth, respectively). Abbreviations: cr-t = crenulated teeth, ml = membranous lobe, scl-s = sclerotized spots. Scale bars in millimetres." figureDoi="10.3897/zookeys.1212.126536.figure12" httpUri="https://binary.pensoft.net/fig/1133565">12</figureCitation>
,
<figureCitation id="9DFD73D52EF07924ED202E6F17702071" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Zephronia chantaburiensis sp. nov., ♂ paratype (ZFMK - MYR 13660), drawings — left posterior telopod A anterior view B posterior view. Abbreviations: cr-t = crenulated teeth, imf = immovable finger, ml = membranous lobe. Scale bar: 0.5 millimetres." figureDoi="10.3897/zookeys.1212.126536.figure13" httpUri="https://binary.pensoft.net/fig/1133566">13</figureCitation>
,
<figureCitation id="AFBEA9C8FD659E19D8352497C859F1CA" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Endoterga of thoracic shields and midbody tergites — SEM A D Zephronia chantaburiensis sp. nov. (♂ paratype, CUMZ - MYR 0014) E H Zephronia macula sp. nov. (♂ paratype, CUMZ - MYR 0016) A, B, E, F thoracic shields C, D, G, H midbody tergites. Abbreviations: cp = cuticular impression, ia = inner area, ma = middle area, oa = outer area, pm = posterior margin. Scale bars in millimetres." figureDoi="10.3897/zookeys.1212.126536.figure14" httpUri="https://binary.pensoft.net/fig/1133567">14 E H</figureCitation>
,
<figureCitation id="F7053359F08F286EE78EA6E80B362866" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Distribution and localities of Zephronia species in Thailand. Number inside the circle indicates species: 1 = Z. chantaburiensis sp. nov.; 2 = Z. chrysomallos; 3 = Z. enghoffi; 4 = Z. erawani; 5 = Z. golovatchi; 6 = Z. lannaensis; 7 = Z. macula sp. nov.; 8 = Z. panhai; 9 = Z. phrain; 10 = Z. siamensis; 11 = Z. viridisoma." figureDoi="10.3897/zookeys.1212.126536.figure15" httpUri="https://binary.pensoft.net/fig/1133568">15</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EYLDI" type="Type specimen">
<paragraph id="A914C3610EC9FE6072FDDB5518E47946">
<heading id="D8AE2C348BEDFB75B3DA0538B8030D32" reason="title">Type specimen.</heading>
</paragraph>
<paragraph id="0CAC9AE3E0F1DB6591D805FB947CC1A1">
<materialsCitation id="389FD71895DB69F1B6B2C9E0B234BFCB" collectingDate="2023-06-14" collectionCode="CUMZ" collectorName="R. Srisonchai &amp; KKUMZ students" country="Thailand" county="Kaeng Hang Maeo District" elevation="60" latitude="12.897695" location="Thamma Sooksawang Temple" longLatPrecision="1" longitude="101.81659" municipality="Khao Wong Kot Cave" specimenCode="MYR 0015" specimenCount="1" specimenCount-male="1" stateProvince="Chantaburi Province" typeStatus="holotype">
<emphasis id="55DC1ED9EAFB1201207D676CF2AFE8F8" bold="true" italics="true">
<typeStatus id="75C2B7A2D98BFB4B881223FD7D25DEE5" type="holotype">Holotype</typeStatus>
</emphasis>
<specimenCount id="3ACF2D236A9F18E96ABCD00946EBADEB" count="1" type="male"></specimenCount>
(
<named-content id="820752D3E7CDD5534411C4BA2D7CF8F9" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/chulalongkorn-university-museum-natural-history" xlink_title="Chulalongkorn University Museum of Zoology">
<collectionCode id="431358AD55F31E0A3F85F9CCF42DC999" country="0" httpUri="http://grbio.org/cool/da58-p0xp" name="Cameroon University, Museum of Zoology" type="">CUMZ</collectionCode>
</named-content>
-
<specimenCode id="EB4122DC12EBCBAC9556D6EB3EFAF57D" collectionCode="CUMZ" country="0" httpUri="http://grbio.org/cool/da58-p0xp" name="Cameroon University, Museum of Zoology" type="">MYR 0015</specimenCode>
);
<collectingCountry id="F3C4CF63BF3C4A5A4C88A4D5F02F57BC" name="Thailand">Thailand</collectingCountry>
,
<collectingRegion id="2982195F7CDB7C91B5058063B58BB8DE" country="Thailand" name="Chanthaburi">Chantaburi Province</collectingRegion>
,
<collectingCounty id="FEE321B2CF6A50941171AC5DC4B794E2">Kaeng Hang Maeo District</collectingCounty>
, near
<collectingMunicipality id="4EE6F6B54129BBB6E0A3259BC12DD38E">Khao Wong Kot Cave</collectingMunicipality>
,
<location id="10ED497FA76D59E74CCD393E8872B706" LSID="urn:lsid:plazi:treatment:F69B1A1DC0F254338EC1CF3319469C6A:10ED497FA76D59E74CCD393E8872B706" country="Thailand" county="Kaeng Hang Maeo District" latitude="12.897695" longLatPrecision="1" longitude="101.81659" municipality="Khao Wong Kot Cave" name="Thamma Sooksawang Temple" stateProvince="Chantaburi Province">Thamma Sooksawang Temple</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EKMDI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[101.816583,12.897694]}">
<geoCoordinate id="F060914781038F0BA2D7B5692D5567DB" degrees="12" direction="north" minutes="53" orientation="latitude" precision="1" seconds="51.7" value="12.897695">12 ° 53 ' 51.7 &quot; N</geoCoordinate>
,
<geoCoordinate id="2A9B7A50387548F3D732B319070A1D22" degrees="101" direction="east" minutes="48" orientation="longitude" precision="1" seconds="59.7" value="101.81659">101 ° 48 ' 59.7 &quot; E</geoCoordinate>
</named-content>
; ca.
<elevation id="C6D8552036CD629CEF06D720976E66AB" metricMagnitude="1" metricUnit="m" metricValue="6.0" unit="m" value="60.0">
<quantity id="36E57AAF75D840E2E252C5A55AA8B76C" metricMagnitude="1" metricUnit="m" metricValue="6.0" unit="m" value="60.0">60 m</quantity>
a. s. l.
</elevation>
;
<date id="80F0FBC19223B8E8904DC771083B2253" value="2023-06-14">
<collectingDate id="BFC9D579A3CBFD8B7A3928EF89968317" value="2023-06-14">14 June 2023</collectingDate>
</date>
; leg.
<collectorName id="DB8D3AE876CFA60B5DF1D62CE1A27228">R. Srisonchai</collectorName>
and
<collectorName id="A8900BFB396E0B1DEE9D9C93F2C8BAB1">
<named-content id="119D59A74BDDDB9B84312FC316501D2B" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/khon-kaen-university" xlink_title="Khon Kaen University Museum of Zoology">KKUMZ</named-content>
students
</collectorName>
</materialsCitation>
.
<materialsCitation id="68C8E2790DF8952EBDC53D39E1A55AC7" collectingDate="2023-06-14" collectionCode="CUMZ" collectorName="R. Srisonchai" country="Thailand" county="Kaeng Hang Maeo District" elevation="60" latitude="12.897695" location="Thamma Sooksawang Temple" longLatPrecision="1" longitude="101.81659" municipality="Khao Wong Kot Cave" specimenCode="MYR 0016" specimenCount="127" specimenCount-female="74" specimenCount-male="53" stateProvince="Chanthaburi" typeStatus="paratype">
<emphasis id="D5EA838308F853B9496B4219FF025F7F" bold="true">
<emphasis id="6B20712C44D41584310EF6BCF0AE8D79" bold="true" italics="true">
<typeStatus id="6617E084A32934A6ACF7D3468E5CF844" type="paratype">Paratypes</typeStatus>
</emphasis>
.
</emphasis>
<specimenCount id="5EE184DB61D0765212CC1C6B1608976F" count="53" type="male">53 ♂</specimenCount>
,
<specimenCount id="263C5E07CE03BEFAB0708436AC80E680" count="74" type="female">74 ♀</specimenCount>
(
<named-content id="78BB779D8AEC40EF2607398DA985C81E" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/chulalongkorn-university-museum-natural-history" xlink_title="Chulalongkorn University Museum of Zoology">
<collectionCode id="6A9EAFFBFCE0A0D7234C4845DCFFF88D" country="0" httpUri="http://grbio.org/cool/da58-p0xp" name="Cameroon University, Museum of Zoology" type="">CUMZ</collectionCode>
</named-content>
-
<specimenCode id="6C77810D1E02BD422CA3BE865C4FB8F4" collectionCode="CUMZ" country="0" httpUri="http://grbio.org/cool/da58-p0xp" name="Cameroon University, Museum of Zoology" type="">MYR 0016</specimenCode>
), same data as holotype
</materialsCitation>
; •
<materialsCitation id="9DA040FDDBE28B92CD3D1F27B0CE7F62" collectingDate="2023-06-14" collectionCode="NHMD" collectorName="R. Srisonchai" country="Thailand" county="Kaeng Hang Maeo District" elevation="60" latitude="12.897695" location="Thamma Sooksawang Temple" longLatPrecision="1" longitude="101.81659" municipality="Khao Wong Kot Cave" specimenCode="NHMD 1184696" specimenCount="4" specimenCount-female="2" specimenCount-male="2" stateProvince="Chanthaburi" typeStatus="paratype">
<specimenCount id="051F3DE5BD4B31E72B38CE99C4E45823" count="2" type="male">2 ♂</specimenCount>
,
<specimenCount id="45A04148C91932B3F40FE8B37EC48E1C" count="2" type="female">2 ♀</specimenCount>
(
<specimenCode id="6B45379A10523D3A31A60B9127EBACF1" collectionCode="NHMD">NHMD 1184696</specimenCode>
), same data as holotype
</materialsCitation>
; •
<materialsCitation id="E8F257F2C712FAA016F3A5AF18CA2CF2" collectingDate="2023-06-14" collectionCode="NHMW" collectorName="R. Srisonchai" country="Thailand" county="Kaeng Hang Maeo District" elevation="60" latitude="12.897695" location="Thamma Sooksawang Temple" longLatPrecision="1" longitude="101.81659" municipality="Khao Wong Kot Cave" specimenCode="NHMW 10437" specimenCount="4" specimenCount-female="2" specimenCount-male="2" stateProvince="Chanthaburi" typeStatus="paratype">
<specimenCount id="484857F7F8D82E9BBA50A4CFA0D839C1" count="2" type="male">2 ♂</specimenCount>
,
<specimenCount id="2BBC0986FFA28014EF854A5FEF5156EF" count="2" type="female">2 ♀</specimenCount>
(
<specimenCode id="AC8E4308FBAEBA2B4F91F58F8667B2FF" collectionCode="NHMW" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" type="Museum">NHMW 10437</specimenCode>
), same data as holotype
</materialsCitation>
; •
<materialsCitation id="E005C9B96DC1B3E7510E19E90E012646" collectingDate="2023-06-14" collectionCode="ZFMK" collectorName="R. Srisonchai" country="Thailand" county="Kaeng Hang Maeo District" elevation="60" latitude="12.897695" location="Thamma Sooksawang Temple" longLatPrecision="1" longitude="101.81659" municipality="Khao Wong Kot Cave" specimenCode="MYR 13660" specimenCount="4" specimenCount-female="2" specimenCount-male="2" stateProvince="Chanthaburi" typeStatus="paratype">
<specimenCount id="71B8B96382CBD5FDEC1BD9C423CC769F" count="2" type="male">2 ♂</specimenCount>
,
<specimenCount id="9117ACC49D94DD22905411F1F9718E45" count="2" type="female">2 ♀</specimenCount>
(
<named-content id="4854EFD59C12F3FD62ED368222B28085" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/zoologisches-forschungsmuseum-alexander-koenig" xlink_title="Zoological Research Museum Koenig">
<collectionCode id="8CDCEA62FAA66F1F4C69C3EB1651A4A2" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" type="Museum">ZFMK</collectionCode>
</named-content>
-
<specimenCode id="82F2E9BB9F3F40599C711D6B257DF13C" collectionCode="ZFMK" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" type="Museum">MYR 13660</specimenCode>
), same data as holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0E6MDI" type="material">
<paragraph id="429CF1AD1A7F2A2F87B7FB3FBBC2FE93">
<heading id="916A28C5E908DCC8518E11BA49A7648D" reason="title">Additional material.</heading>
</paragraph>
<paragraph id="04D38A114FE69FF13F79EB6967B2D3C8">
<materialsCitation id="1AD3E0CC1B677DAAC08774376E51D328" collectingDate="2023-06-14" collectionCode="CUMZ" collectorName="R. Srisonchai &amp; KKUMZ students" country="Thailand" county="Kaeng Hang Maeo District" elevation="60" latitude="12.897695" location="Thamma Sooksawang Temple" longLatPrecision="1" longitude="101.81659" municipality="Khao Wong Kot Cave" specimenCount="29" specimenCount-juvenile="29" stateProvince="Chantaburi Province">
<specimenCount id="517345C6C395E390E5EBA55C9510106C" count="29" type="juvenile">29 juveniles</specimenCount>
(
<named-content id="752A5A147B80ADF45C2DB604381021B0" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/chulalongkorn-university-museum-natural-history" xlink_title="Chulalongkorn University Museum of Zoology">
<collectionCode id="FB52141F5254D966E7C621C61AF8E7BA" country="0" httpUri="http://grbio.org/cool/da58-p0xp" name="Cameroon University, Museum of Zoology" type="">CUMZ</collectionCode>
</named-content>
);
<collectingCountry id="4FFB5CE27287B8EE263B93CE6569E9D0" name="Thailand">Thailand</collectingCountry>
,
<collectingRegion id="E8BBF1435A359904731829F6EF7984F1" country="Thailand" name="Chanthaburi">Chantaburi Province</collectingRegion>
,
<collectingCounty id="7922FF92324E5632174B843A74DA13CA">Kaeng Hang Maeo District</collectingCounty>
, near
<collectingMunicipality id="51DCE2B8E25B460DE2BA08D7C8CD3A51">Khao Wong Kot Cave</collectingMunicipality>
,
<location id="B8058066F8F2E328ACEB8542723F6137" LSID="urn:lsid:plazi:treatment:F69B1A1DC0F254338EC1CF3319469C6A:B8058066F8F2E328ACEB8542723F6137" country="Thailand" county="Kaeng Hang Maeo District" latitude="12.897695" longLatPrecision="1" longitude="101.81659" municipality="Khao Wong Kot Cave" name="Thamma Sooksawang Temple" stateProvince="Chantaburi Province">Thamma Sooksawang Temple</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EMNDI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[101.816583,12.897694]}">
<geoCoordinate id="781A189FB8E4B4EA4ECD65758290F3B0" degrees="12" direction="north" minutes="53" orientation="latitude" precision="1" seconds="51.7" value="12.897695">12 ° 53 ' 51.7 &quot; N</geoCoordinate>
,
<geoCoordinate id="87FD7E6556DA927A8371C7D2BEC11DF4" degrees="101" direction="east" minutes="48" orientation="longitude" precision="1" seconds="59.7" value="101.81659">101 ° 48 ' 59.7 &quot; E</geoCoordinate>
</named-content>
; ca.
<elevation id="53191AAF570255AA622E2EE4712E0970" metricMagnitude="1" metricUnit="m" metricValue="6.0" unit="m" value="60.0">
<quantity id="7F562AEF447B78A3B934B2DD0D22811D" metricMagnitude="1" metricUnit="m" metricValue="6.0" unit="m" value="60.0">60 m</quantity>
a. s. l.
</elevation>
;
<date id="4763F284AC5807BC212AEA3BB6FFD167" value="2023-06-14">
<collectingDate id="A2B28792897DD9AF9F05D98779D4368B" value="2023-06-14">14 June 2023</collectingDate>
</date>
; leg.
<collectorName id="73FD3DA1F0AA305C971C0FE284EF2139">R. Srisonchai</collectorName>
and
<collectorName id="5D2D42D30AE90458763FD78065322C80">
<named-content id="20B487DF5CE3B8D4E4684E4C091DA2CD" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/khon-kaen-university" xlink_title="Khon Kaen University Museum of Zoology">KKUMZ</named-content>
students
</collectorName>
</materialsCitation>
; •
<materialsCitation id="09C16180DA9F2F237BDDC1F8B2D8648F" collectingDate="2023-06-14" collectionCode="CUMZ" collectorName="R. Srisonchai &amp; KKUMZ students" country="Thailand" county="Tha Mai District" elevation="53" latitude="12.763056" location="Wat Khao Sukim" longLatPrecision="21" longitude="102.03223" municipality="Wat Khao Sukim" specimenCount="57" specimenCount-female="31" specimenCount-male="26" stateProvince="Chantaburi Province">
<specimenCount id="FA08A20936859F6A2C01D99E24402736" count="26" type="male">26 ♂</specimenCount>
,
<specimenCount id="8E472149E135BF87BFD55B3F657E503F" count="31" type="female">31 ♀</specimenCount>
(
<named-content id="8570569A33E77F415C8B4A6C057A7B7E" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/chulalongkorn-university-museum-natural-history" xlink_title="Chulalongkorn University Museum of Zoology">
<collectionCode id="8F95370E36C4AC1E16240A4DA7FD31B5" country="0" httpUri="http://grbio.org/cool/da58-p0xp" name="Cameroon University, Museum of Zoology" type="">CUMZ</collectionCode>
</named-content>
);
<collectingCountry id="F9D11536D84F5FB2B38AE63F8FF98A18" name="Thailand">Thailand</collectingCountry>
,
<collectingRegion id="1F04FED4A849A5D947D349189B9A06C5" country="Thailand" name="Chanthaburi">Chantaburi Province</collectingRegion>
,
<collectingCounty id="DDDBF310E69FBCE5446B141282C354A5">Tha Mai District</collectingCounty>
,
<location id="DCE53DD0AF7E8613E65338AB6D134A0D" LSID="urn:lsid:plazi:treatment:F69B1A1DC0F254338EC1CF3319469C6A:DCE53DD0AF7E8613E65338AB6D134A0D" country="Thailand" county="Tha Mai District" latitude="12.763056" longLatPrecision="21" longitude="102.03223" municipality="Wat Khao Sukim" name="Wat Khao Sukim" stateProvince="Chantaburi Province">Wat Khao Sukim</location>
(Khao Sukim Temple);
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E3NDI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[102.032222,12.763056]}">
<geoCoordinate id="08D99E195808DAC217A627CA24F5C1CE" degrees="12" direction="north" minutes="45" orientation="latitude" precision="15" seconds="47" value="12.763056">12 ° 45 ' 47 &quot; N</geoCoordinate>
,
<geoCoordinate id="C99519881E4BB312245D8E9C55D63CE0" degrees="102" direction="east" minutes="01" orientation="longitude" precision="15" seconds="56" value="102.03223">102 ° 01 ' 56 &quot; E</geoCoordinate>
</named-content>
; ca.
<elevation id="FA135D0D4F742638DCC388A8CE4A2AAA" metricMagnitude="1" metricUnit="m" metricValue="5.3" unit="m" value="53.0">
<quantity id="C943C4D473DB457CFADB91D17FDACB00" metricMagnitude="1" metricUnit="m" metricValue="5.3" unit="m" value="53.0">53 m</quantity>
a. s. l.
</elevation>
;
<date id="229A66F2A055A6CDF2C58DD5B3422E53" value="2023-06-14">
<collectingDate id="9C49C62435D4F6FBB63C1C430C2F1050" value="2023-06-14">14 June 2023</collectingDate>
</date>
; leg.
<collectorName id="D60705062D653B7A17B05D7441F967B4">R. Srisonchai</collectorName>
and
<collectorName id="FE145D3975375FAE64CB67749DA6DB43">
<named-content id="D652828E834A1260B58E5B8C6AF6CD0E" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/khon-kaen-university" xlink_title="Khon Kaen University Museum of Zoology">KKUMZ</named-content>
students
</collectorName>
</materialsCitation>
; •
<materialsCitation id="6A05ECE3B49F414E90867999513A69B8" collectingDate="2023-06-14" collectionCode="CUMZ" collectorName="R. Srisonchai &amp; KKUMZ students" country="Thailand" elevation="74" latitude="12.481472" location="Thaeo Klong Khlung Monastery" longLatPrecision="1" longitude="102.218414" municipality="Khlung District" specimenCount="44" specimenCount-female="26" specimenCount-male="18" stateProvince="Chantaburi Province">
<specimenCount id="BAEC05D66BBE7FDA5B9D3BF88DC9C4C2" count="18" type="male">18 ♂</specimenCount>
,
<specimenCount id="61DA29C4632463BA5979B4CE2212EFBB" count="26" type="female">26 ♀</specimenCount>
(
<named-content id="E1DAD37C5CD826351BA2ACD4B388B095" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/chulalongkorn-university-museum-natural-history" xlink_title="Chulalongkorn University Museum of Zoology">
<collectionCode id="7F116B9A99F6038248969CFAF6018957" country="0" httpUri="http://grbio.org/cool/da58-p0xp" name="Cameroon University, Museum of Zoology" type="">CUMZ</collectionCode>
</named-content>
);
<collectingCountry id="1318284924511086600B875913C0604D" name="Thailand">Thailand</collectingCountry>
,
<collectingRegion id="B0DF65850025FFB0322ED9741CABF15F" country="Thailand" name="Chanthaburi">Chantaburi Province</collectingRegion>
,
<collectingMunicipality id="BD28B814F9C732F95DD0DA800FB1FCAA">Khlung District</collectingMunicipality>
,
<location id="4622EC1BBAC1C15B67B639F36233C8D6" LSID="urn:lsid:plazi:treatment:F69B1A1DC0F254338EC1CF3319469C6A:4622EC1BBAC1C15B67B639F36233C8D6" country="Thailand" latitude="12.481472" longLatPrecision="1" longitude="102.218414" municipality="Khlung District" name="Thaeo Klong Khlung Monastery" stateProvince="Chantaburi Province">Thaeo Klong Khlung Monastery</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EMODI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[102.218417,12.481472]}">
<geoCoordinate id="9364E7941BABE3691DF9FB3853E88B69" degrees="12" direction="north" minutes="28" orientation="latitude" precision="1" seconds="53.3" value="12.481472">12 ° 28 ' 53.3 &quot; N</geoCoordinate>
,
<geoCoordinate id="642C0DB1A67D477B26DCA20AD824F0A4" degrees="102" direction="east" minutes="13" orientation="longitude" precision="1" seconds="06.3" value="102.218414">102 ° 13 ' 06.3 &quot; E</geoCoordinate>
</named-content>
; ca.
<elevation id="560BFC104916C899FFF858B8AA5257DE" metricMagnitude="1" metricUnit="m" metricValue="7.4" unit="m" value="74.0">
<quantity id="A0035CFFAB788FB68B082D84BCCB693F" metricMagnitude="1" metricUnit="m" metricValue="7.4" unit="m" value="74.0">74 m</quantity>
a. s. l.
</elevation>
;
<date id="2E4392283005DCD9398EE43F093EDFCC" value="2023-06-14">
<collectingDate id="238FF400D9CA0D11C9B081CECE98F329" value="2023-06-14">14 June 2023</collectingDate>
</date>
; leg.
<collectorName id="6ED0CBED35C18D3E0047C9B9D9FED301">R. Srisonchai</collectorName>
and
<collectorName id="963773C3AD0C04F6A0F1222C07CD94E7">
<named-content id="A98E752E82074793AB15FC175A3F9981" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/khon-kaen-university" xlink_title="Khon Kaen University Museum of Zoology">KKUMZ</named-content>
students
</collectorName>
</materialsCitation>
; •
<materialsCitation id="57B0BC5DD06AC671E152B4503F106342" collectingDate="2023-06-13" collectionCode="CUMZ" collectorName="R. Srisonchai &amp; KKUMZ students" country="Thailand" elevation="110" latitude="12.8525" location="Wat Khao Banchob" longLatPrecision="1" longitude="102.20416" municipality="Makham District" specimenCount="138" specimenCount-female="65" specimenCount-juvenile="47" specimenCount-male="26" stateProvince="Chantaburi Province">
<specimenCount id="BAF59C309DF0FD02CB675820F6477F84" count="26" type="male">26 ♂</specimenCount>
,
<specimenCount id="F66415459FE65855CABD07F1A23B7F8D" count="65" type="female">65 ♀</specimenCount>
;
<specimenCount id="F59557D1BD7140EEBCD0744740457567" count="47" type="juvenile">47 juveniles</specimenCount>
(
<named-content id="EF1CAAB093D0A49831AD13A07E956F26" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/chulalongkorn-university-museum-natural-history" xlink_title="Chulalongkorn University Museum of Zoology">
<collectionCode id="D4EB1511F9A9D51FE12AF75D19EB9619" country="0" httpUri="http://grbio.org/cool/da58-p0xp" name="Cameroon University, Museum of Zoology" type="">CUMZ</collectionCode>
</named-content>
);
<collectingCountry id="A53F83F0721676C8CA2B2255AB9A09F0" name="Thailand">Thailand</collectingCountry>
,
<collectingRegion id="F5A01DD2397F3159B412135269358F75" country="Thailand" name="Chanthaburi">Chantaburi Province</collectingRegion>
,
<collectingMunicipality id="097A42E20E0C7640360022D91857290B">Makham District</collectingMunicipality>
,
<location id="16B181596916A5E82111423AFE5BB69A" LSID="urn:lsid:plazi:treatment:F69B1A1DC0F254338EC1CF3319469C6A:16B181596916A5E82111423AFE5BB69A" country="Thailand" latitude="12.8525" longLatPrecision="1" longitude="102.20416" municipality="Makham District" name="Wat Khao Banchob" stateProvince="Chantaburi Province">Wat Khao Banchob</location>
,
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E3ODI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[102.204167,12.852500]}">
<geoCoordinate id="C0B0C0DF73DD93F640FDB3C68D27E904" degrees="12" direction="north" minutes="51" orientation="latitude" precision="1" seconds="09.0" value="12.8525">12 ° 51 ' 09.0 &quot; N</geoCoordinate>
,
<geoCoordinate id="21E274DB5A5BB4AB8AE6760B644BF7F4" degrees="102" direction="east" minutes="12" orientation="longitude" precision="1" seconds="15.0" value="102.20416">102 ° 12 ' 15.0 &quot; E</geoCoordinate>
</named-content>
; ca.
<elevation id="B1B0991936BED201535305964C5B7653" metricMagnitude="2" metricUnit="m" metricValue="1.1" unit="m" value="110.0">
<quantity id="6CB4F5487D8C403A91BE6B2F0419EE21" metricMagnitude="2" metricUnit="m" metricValue="1.1" unit="m" value="110.0">110 m</quantity>
a. s. l.
</elevation>
;
<date id="04E6BB30DF183041CDCE32D72C87025C" value="2023-06-13">
<collectingDate id="F97F660D550EC590E76CF089872B994A" value="2023-06-13">13 June 2023</collectingDate>
</date>
; leg.
<collectorName id="70BBAEFCDEADDC082C3E4A72E34EA9E9">R. Srisonchai</collectorName>
and
<collectorName id="55B95397372513A70E6183EE20D9856C">
<named-content id="AC197EEB903614FC4EFB2A75F116081F" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/khon-kaen-university" xlink_title="Khon Kaen University Museum of Zoology">KKUMZ</named-content>
students
</collectorName>
</materialsCitation>
<materialsCitation id="796B0608CF702687E1FF113F6AB7FEE8" collectingDate="2023-06-15" collectionCode="CUMZ" collectorName="R. Srisonchai &amp; KKUMZ students" country="Thailand" elevation="160" latitude="13.2485285" location="Wat Khao Yai Aran Khiri" longLatPrecision="1" longitude="101.62489" municipality="Bo Thong District" specimenCount="76" specimenCount-female="43" specimenCount-male="33" stateProvince="Chonburi Province">
<specimenCount id="17C9FA986A92729D241D55F3BC9BB69C" count="33" type="male">33 ♂</specimenCount>
,
<specimenCount id="A2558BB23F8772AD4A01B9334F4E4D3D" count="43" type="female">43 ♀</specimenCount>
(
<named-content id="4D55F74419B992162901BFC1A57819EA" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/chulalongkorn-university-museum-natural-history" xlink_title="Chulalongkorn University Museum of Zoology">
<collectionCode id="6E60720BEF6DF8DF81C3C8758B2A83A2" country="0" httpUri="http://grbio.org/cool/da58-p0xp" name="Cameroon University, Museum of Zoology" type="">CUMZ</collectionCode>
</named-content>
);
<collectingCountry id="906233BA64002127782F31F1BF856FAA" name="Thailand">Thailand</collectingCountry>
,
<collectingRegion id="4CD55875F76C4B3D0FC846E60632389B" country="Thailand" name="Chon Buri">Chonburi Province</collectingRegion>
,
<collectingMunicipality id="4521A3F6B01B6FDBFEBD0F838B3BF44E">Bo Thong District</collectingMunicipality>
,
<location id="E3A53341FD6004E578A10F0531BE85B5" LSID="urn:lsid:plazi:treatment:F69B1A1DC0F254338EC1CF3319469C6A:E3A53341FD6004E578A10F0531BE85B5" country="Thailand" latitude="13.2485285" longLatPrecision="1" longitude="101.62489" municipality="Bo Thong District" name="Wat Khao Yai Aran Khiri" stateProvince="Chonburi Province">Wat Khao Yai Aran Khiri</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EMPDI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[101.624889,13.248528]}">
<geoCoordinate id="28F21D9C88EB163F58FEBD10E836CFBD" degrees="13" direction="north" minutes="14" orientation="latitude" precision="1" seconds="54.7" value="13.2485285">13 ° 14 ' 54.7 &quot; N</geoCoordinate>
,
<geoCoordinate id="A6A87DFB57B4DF96DC14FB52101C211C" degrees="101" direction="east" minutes="37" orientation="longitude" precision="1" seconds="29.6" value="101.62489">101 ° 37 ' 29.6 &quot; E</geoCoordinate>
</named-content>
; ca.
<elevation id="1C5ACE5BEE4F807839B0A7C0D516ECAD" metricMagnitude="2" metricUnit="m" metricValue="1.6" unit="m" value="160.0">
<quantity id="C6157FBB8E222294F2F9BE0BAE833773" metricMagnitude="2" metricUnit="m" metricValue="1.6" unit="m" value="160.0">160 m</quantity>
a. s. l.
</elevation>
;
<date id="DA7C1F74E4FB93B943A58894CC054DF6" value="2023-06-15">
<collectingDate id="60C1CE813B0B73D374362E2D66D1562F" value="2023-06-15">15 June 2023</collectingDate>
</date>
; leg.
<collectorName id="D9321BF4DDA1E712646923B8B77EFA38">R. Srisonchai</collectorName>
and
<collectorName id="69FF85C225C467A8901A028B06048FF9">
<named-content id="851DFD30BCD69B5A4266A7D412A405D5" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/khon-kaen-university" xlink_title="Khon Kaen University Museum of Zoology">KKUMZ</named-content>
students
</collectorName>
</materialsCitation>
; •
<materialsCitation id="5D01D3C8F0074D27BCCBA79971F485CF" collectingDate="2023-06-14" collectionCode="CUMZ" collectorName="R. Srisonchai &amp; KKUMZ students" country="Thailand" county="Makham District" elevation="80" latitude="12.700916" location="Wat Pa Theprangsi" longLatPrecision="1" longitude="101.53814" municipality="Wat Pa Theprangsi" specimenCount="91" specimenCount-female="25" specimenCount-male="66" stateProvince="Rayong Province">
<specimenCount id="F5618B1CE4E47BFF4B67CADAD94B07DF" count="66" type="male">66 ♂</specimenCount>
,
<specimenCount id="1AFA6DED3796FB412DBAFAE374AE4765" count="25" type="female">25 ♀</specimenCount>
(
<named-content id="A81BBF5437C6CB9BC3FB92521096146E" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/chulalongkorn-university-museum-natural-history" xlink_title="Chulalongkorn University Museum of Zoology">
<collectionCode id="9FA40B2CF9B3BDF46ED8A9948DE4B95C" country="0" httpUri="http://grbio.org/cool/da58-p0xp" name="Cameroon University, Museum of Zoology" type="">CUMZ</collectionCode>
</named-content>
);
<collectingCountry id="996ABE96E228105D368C446CFD4DA8C1" name="Thailand">Thailand</collectingCountry>
,
<collectingRegion id="7BD2468AF06A6A6AE10E9791386F5D75" country="Thailand" name="Rayong">Rayong Province</collectingRegion>
,
<collectingCounty id="9923058CBA0C9E85D3952BA7A321A5D3">Makham District</collectingCounty>
,
<location id="3B6794F7581034C7CEE9E326D371378C" LSID="urn:lsid:plazi:treatment:F69B1A1DC0F254338EC1CF3319469C6A:3B6794F7581034C7CEE9E326D371378C" country="Thailand" county="Makham District" latitude="12.700916" longLatPrecision="1" longitude="101.53814" municipality="Wat Pa Theprangsi" name="Wat Pa Theprangsi" stateProvince="Rayong Province">Wat Pa Theprangsi</location>
(= Wat Khao Hin Tang);
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E3PDI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[101.538139,12.700917]}">
<geoCoordinate id="D834A464605521A0DC398083916979CF" degrees="12" direction="north" minutes="42" orientation="latitude" precision="1" seconds="03.3" value="12.700916">12 ° 42 ' 03.3 &quot; N</geoCoordinate>
,
<geoCoordinate id="E531B92F303E821778774F6BF379A119" degrees="101" direction="east" minutes="32" orientation="longitude" precision="1" seconds="17.3" value="101.53814">101 ° 32 ' 17.3 &quot; E</geoCoordinate>
</named-content>
; ca.
<elevation id="749A287C302B600CF8E3EB14A0507BD9" metricMagnitude="1" metricUnit="m" metricValue="8.0" unit="m" value="80.0">
<quantity id="9B3C942B178A0FE3339DE2DDAD82D0ED" metricMagnitude="1" metricUnit="m" metricValue="8.0" unit="m" value="80.0">80 m</quantity>
a. s. l.
</elevation>
;
<date id="11BB392F16A1149D12181DFC98092BD9" value="2023-06-14">
<collectingDate id="BD54147C0406CABFD8DA6278F09C27AB" value="2023-06-14">14 June 2023</collectingDate>
</date>
; leg.
<collectorName id="488ABADDC198E270FFA77ED4FF2E5896">R. Srisonchai</collectorName>
and
<collectorName id="949A6EE03EBA4E23F2B6BD1971B5F994">
<named-content id="3AA185205B900C6E51523A314FF77816" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/khon-kaen-university" xlink_title="Khon Kaen University Museum of Zoology">KKUMZ</named-content>
students
</collectorName>
</materialsCitation>
; •
<materialsCitation id="86EBA7CA9DD4EA3A06FA2440F5D8E9E1" collectingDate="2023-06-12" collectionCode="CUMZ" collectorName="R. Srisonchai &amp; KKUMZ students" country="Thailand" elevation="245" latitude="13.414861" location="Phet Pho Thong Cave" longLatPrecision="1" longitude="102.326416" municipality="Khlong Hat District" specimenCount="12" specimenCount-juvenile="7" specimenCount-male="5" stateProvince="Sa Kaeo Province">
<specimenCount id="6CF3F925B665B87291878DCF648F603A" count="5" type="male">5 ♂</specimenCount>
,
<specimenCount id="9D23AA7FF23110DD32978D22058A5201" count="7" type="juvenile">7 juveniles</specimenCount>
(
<named-content id="B0662DD11377744C2E1085C3257B924F" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/chulalongkorn-university-museum-natural-history" xlink_title="Chulalongkorn University Museum of Zoology">
<collectionCode id="F992CA7F83323A622146AEFB272CF736" country="0" httpUri="http://grbio.org/cool/da58-p0xp" name="Cameroon University, Museum of Zoology" type="">CUMZ</collectionCode>
</named-content>
);
<collectingCountry id="3325EC81210FF422BEC62A100E0D51A5" name="Thailand">Thailand</collectingCountry>
,
<collectingRegion id="6303998B78CE92BCC4EA0434A8E52EC8" country="Thailand" name="Sa Kaeo">Sa Kaeo Province</collectingRegion>
,
<collectingMunicipality id="CE49A4D01E8D10E72108F7C7D08F854F">Khlong Hat District</collectingMunicipality>
,
<location id="735BECE2D6330198B5183BA19EACF5CE" LSID="urn:lsid:plazi:treatment:F69B1A1DC0F254338EC1CF3319469C6A:735BECE2D6330198B5183BA19EACF5CE" country="Thailand" latitude="13.414861" longLatPrecision="1" longitude="102.326416" municipality="Khlong Hat District" name="Phet Pho Thong Cave" stateProvince="Sa Kaeo Province">Phet Pho Thong Cave</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EMQDI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[102.326417,13.414861]}">
<geoCoordinate id="23B08E52E38E9D842547041C2C2D00B5" degrees="13" direction="north" minutes="24" orientation="latitude" precision="1" seconds="53.5" value="13.414861">13 ° 24 ' 53.5 &quot; N</geoCoordinate>
,
<geoCoordinate id="80EFFCA19BBEBD232002EEF56DC2EF9E" degrees="102" direction="east" minutes="19" orientation="longitude" precision="1" seconds="35.1" value="102.326416">102 ° 19 ' 35.1 &quot; E</geoCoordinate>
</named-content>
; ca.
<elevation id="9F117303AAC763D0BE33AF099AAE8111" metricMagnitude="2" metricUnit="m" metricValue="2.45" unit="m" value="245.0">
<quantity id="ACD99070B0FC308F17534FC3DCB9D333" metricMagnitude="2" metricUnit="m" metricValue="2.45" unit="m" value="245.0">245 m</quantity>
a. s. l.
</elevation>
;
<date id="E104375ABD2D68BD1FFA1308A3CD7E4E" value="2023-06-12">
<collectingDate id="817C0C8A479DFD60AA2DF2AF0005CA76" value="2023-06-12">12 June 2023</collectingDate>
</date>
; leg.
<collectorName id="94DF66BD116012FDA3F405B753441471">R. Srisonchai</collectorName>
and
<collectorName id="C7F8A0247A9B634E4EE610C6FFE16CEE">
<named-content id="E5AFB92DCFA644FCC328B23DD3D9AB10" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/khon-kaen-university" xlink_title="Khon Kaen University Museum of Zoology">KKUMZ</named-content>
students
</collectorName>
</materialsCitation>
; •
<materialsCitation id="998E2D2FDAA18874B36366DF96BB7AE2" collectingDate="2014-08" collectionCode="CUMZ" collectorName="ASRU members" country="Thailand" elevation="70" latitude="13.786639" location="Wat Tham Khao Maka" longLatPrecision="1" longitude="101.94772" municipality="Mueang Sra Kaeo District" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Sra Kaeo Province">
<specimenCount id="B4889A28D4C2A2311D57530072E3CE40" count="1" type="male">1 ♂</specimenCount>
,
<specimenCount id="F678699ACA68200D1D5718836B5AD9BE" count="2" type="female">2 ♀</specimenCount>
(
<named-content id="11D466A177F1E5FD36FD9A379AB41824" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/chulalongkorn-university-museum-natural-history" xlink_title="Chulalongkorn University Museum of Zoology">
<collectionCode id="E6630FC12D35F6A7FF319C24CBF33EF9" country="0" httpUri="http://grbio.org/cool/da58-p0xp" name="Cameroon University, Museum of Zoology" type="">CUMZ</collectionCode>
</named-content>
);
<collectingCountry id="67D99EBBED5F240D314680AC0495AC07" name="Thailand">Thailand</collectingCountry>
,
<collectingRegion id="62D2DB04F7D1AB94E13D115517DBB40E" inferredBySuffix="true">Sra Kaeo Province</collectingRegion>
,
<collectingMunicipality id="499B1F41304B97AAEB9C4447EBF83EF1">Mueang Sra Kaeo District</collectingMunicipality>
,
<location id="A02ED503B3823A30E84993A078B49E4C" LSID="urn:lsid:plazi:treatment:F69B1A1DC0F254338EC1CF3319469C6A:A02ED503B3823A30E84993A078B49E4C" country="Thailand" latitude="13.786639" longLatPrecision="1" longitude="101.94772" municipality="Mueang Sra Kaeo District" name="Wat Tham Khao Maka" stateProvince="Sra Kaeo Province">Wat Tham Khao Maka</location>
;
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E3QDI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[101.947722,13.786639]}">
<geoCoordinate id="20E8FDD11BC10977ACA46B68A10AD2F5" degrees="13" direction="north" minutes="47" orientation="latitude" precision="1" seconds="11.9" value="13.786639">13 ° 47 ' 11.9 &quot; N</geoCoordinate>
,
<geoCoordinate id="B0B2712509A5E2E0D56E87D966AC983B" degrees="101" direction="east" minutes="56" orientation="longitude" precision="1" seconds="51.8" value="101.94772">101 ° 56 ' 51.8 &quot; E</geoCoordinate>
</named-content>
; ca.
<elevation id="EE874EE889A72E292AE5876E02E74672" metricMagnitude="1" metricUnit="m" metricValue="7.0" unit="m" value="70.0">
<quantity id="AFEDEC0A99625D748E51391E68ADCAC1" metricMagnitude="1" metricUnit="m" metricValue="7.0" unit="m" value="70.0">70 m</quantity>
a. s. l.
</elevation>
;
<date id="F902BB2A1167993C49053D1B64B22B25" value="2014-08">
<collectingDate id="F9A522294BAA3524252188FF65E31B50" value="2014-08">August 2014</collectingDate>
</date>
; leg.
<collectorName id="AAEFFE6AC20574A281D594B75DD4CADA">ASRU members</collectorName>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EBRDI" type="diagnosis">
<paragraph id="D153714A16DC1A25B1E70ABB31364E8E">
<heading id="B9C1E84FD05606CC6314849F6B166692" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="F35A69CE8614ECBE2DE7939D2B129A8F">
The position of the organ of Tömösváry in this small
<taxonomicName id="83C721158B1BC89F75CBE363CD5B02FA" authorityName="Gray" authorityYear="1832" class="Diplopoda" family="Zephroniidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" phylum="Arthropoda" rank="genus">
<emphasis id="86B5943AE694AAD6CFD62FFE09AC2232" italics="true">Zephronia</emphasis>
</taxonomicName>
with an axe-shaped antennomere 6 identifies
<taxonomicName id="D71F0D12DDB9EB9348F20E7EE04917AA" authorityName="Srisonchai &amp; Wesener" authorityYear="2024" class="Diplopoda" family="Zephroniidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" phylum="Arthropoda" rank="species" species="macula" status="sp. nov.">
<emphasis id="50AC7A137F959527A6DCEF59BB64B90B" italics="true">Z. macula</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9FD1CBC38CE2050801F4597B4BBE6AB7" rank="species">sp. nov.</taxonomicNameLabel>
as a member of the
<taxonomicName id="DA77E5D32D191D657C319B6CE83C731B" authorityName="Gray" authorityYear="1832" class="Diplopoda" family="Zephroniidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="45F242D7D9A68207B0A360CA9943BEA6" italics="true">Zephronia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E7B1A6C8DDB388789CDCF5E8B8365E15" sensu="stricto">s. s.</taxonomicNameLabel>
species-group (
<bibRefCitation id="9056E041687670682AF4C72093A111EB" DOI="10.11646/zootaxa.4459.3.7" author="Semenyuk" etAl="et al." firstAuthor="Semenyuk" issue="3" journalOrPublisher="Zootaxa" pagination="535-550" refId="B27" refString="Semenyuk I, Golovatch SI, Wesener T (2018) Four new species of giant pill-millipedes from Vietnam (Sphaerotheriida, Zephroniidae). Zootaxa 4459 (3): 535550. https://doi.org/10.11646/zootaxa.4459.3.7" title="Four new species of giant pill-millipedes from Vietnam (Sphaerotheriida, Zephroniidae)." volume="4459" year="2018">Semenyuk et al. 2018</bibRefCitation>
).
<taxonomicName id="D1781D55AA2A7877897C51F0F3DA1798" authorityName="Srisonchai &amp; Wesener" authorityYear="2024" class="Diplopoda" family="Zephroniidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" phylum="Arthropoda" rank="species" species="macula" status="sp. nov.">
<emphasis id="BA9D2604689B45EC85F877F515AF98D0" italics="true">Z. macula</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FE1FEA2BD7607F0C782A5D5945D20B08" rank="species">sp. nov.</taxonomicNameLabel>
differs from all other known species of the group except for the syntopic
<taxonomicName id="2EC94296304D85E73A2EE5A1DD7DE262" authorityName="Srisonchai &amp; Wesener" authorityYear="2024" class="Diplopoda" family="Zephroniidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" phylum="Arthropoda" rank="species" species="chantaburiensis" status="sp. nov.">
<emphasis id="6F253E883D350761062B912EE1615C55" italics="true">Z. chantaburiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="98A296CFC3F38AA11E8E1F986CDB3661" rank="species">sp. nov.</taxonomicNameLabel>
, in the presence of only a single apical spine on the tarsi of legs 421 (at least 2 or
<quantity id="8964265A2B5EC4AC1A1E5A5CA1438F07" metricMagnitude="-2" metricUnit="m" metricValue="7.62" unit="in" value="3.0">3 in</quantity>
the other
<taxonomicName id="0B9606B7429AE78B7212CFAE710D1BE0" authorityName="Gray" authorityYear="1832" class="Diplopoda" family="Zephroniidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" phylum="Arthropoda" rank="genus">
<emphasis id="5446C3AD92144EE55330B76B9E5FB4A4" italics="true">Zephronia</emphasis>
</taxonomicName>
).
<taxonomicName id="55C4BF1F16E7ED3F7E212F88271C99D7" authorityName="Srisonchai &amp; Wesener" authorityYear="2024" class="Diplopoda" family="Zephroniidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" phylum="Arthropoda" rank="species" species="macula" status="sp. nov.">
<emphasis id="94FACA3F4462699BC72691DCF9BF0A8E" italics="true">Z. macula</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E575C9816F364E969793BC8EFAC5882A" rank="species">sp. nov.</taxonomicNameLabel>
differs from
<taxonomicName id="B23028EE8EC8C7AB97962F9C90FAAE95" authorityName="Srisonchai &amp; Wesener" authorityYear="2024" class="Diplopoda" family="Zephroniidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" phylum="Arthropoda" rank="species" species="chantaburiensis" status="sp. nov.">
<emphasis id="45E202882A5D7FB81B8CEA15B668E71C" italics="true">Z. chantaburiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="67C4EB48BE96365F991ACC79A346F8F9" rank="species">sp. nov.</taxonomicNameLabel>
by having a combination of distinct characters, viz. tergite with dark or greenish dark spots, femur of walking legs less strongly widened, slightly longer than wide (wider than long in
<taxonomicName id="E2E9A944829C341E4873967184BD1CF3" authorityName="Srisonchai &amp; Wesener" authorityYear="2024" class="Diplopoda" family="Zephroniidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" phylum="Arthropoda" rank="species" species="chantaburiensis" status="sp. nov.">
<emphasis id="A9E9154C0EE92094326B43A1C0EDB7C2" italics="true">Z. chantaburiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3B378EC006886BB268F96EB6D5844382" rank="species">sp. nov.</taxonomicNameLabel>
), bristles of midbody endotergum reaching to posterior margin, operculum of female vulva not protruding and round, and immovable finger (process) of telopoditomere 2 of the anterior telopod more curved and much longer. Genetically distant from others by a p-distance of the
<abbrev id="ABBRID0ELUDI" xlink_title="mitochondrial cytochrome c oxidase subunit I">COI</abbrev>
barcoding fragment of 17.9325.13 %.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EPUDI" type="description">
<paragraph id="186D43ACF3EFA5CB98045875B8220D9F">
<heading id="7B63723F32D527B51D277B4964F8E652" reason="title">Description.</heading>
</paragraph>
<paragraph id="498CDFEBE253A7D3843F5B54850344C2">
<emphasis id="4C713FB69D981CFB4393DE88A0057922" bold="true" italics="true">Measurements</emphasis>
: Male
<typeStatus id="FB2C30D13E2D77E5B2C4AD65680EBFEC" type="holotype">holotype</typeStatus>
. Body length
<quantity id="3BDBD2B9AEE6DBC2D67A2EB498E4C8A3" metricMagnitude="-2" metricUnit="m" metricValue="1.8" unit="mm" value="18.0">18 mm</quantity>
. Width, of thoracic shield
<quantity id="96AEBB63A0AB595935411D853F5536C3" metricMagnitude="-3" metricUnit="m" metricValue="8.0" unit="mm" value="8.0">8 mm</quantity>
, of tergite 7 =
<quantity id="264330FA90A43879B988396DA77DAEC9" metricMagnitude="-3" metricUnit="m" metricValue="9.0" unit="mm" value="9.0">9 mm</quantity>
(= broadest). Height of tergite 7 =
<quantity id="A34AD1B1B3DF0715BEFF92E0853AC45B" metricMagnitude="-3" metricUnit="m" metricValue="6.0" unit="mm" value="6.0">6 mm</quantity>
(= highest). Males: body length =
<quantity id="1BB9BB874F034BFA9C20050D37D420A6" metricMagnitude="-2" metricUnit="m" metricValue="1.9" metricValueMax="2.0" metricValueMin="1.8" unit="mm" value="19.0" valueMax="20.0" valueMin="18.0">1820 mm</quantity>
. Width, of thoracic shield =
<quantity id="EA108CD35DFF6112B86F745426CF1358" metricMagnitude="-3" metricUnit="m" metricValue="8.5" metricValueMax="9.0" metricValueMin="8.0" unit="mm" value="8.5" valueMax="9.0" valueMin="8.0">89 mm</quantity>
, of tergite 7 =
<quantity id="E5FEA81896C00DC196F29094512BD49F" metricMagnitude="-3" metricUnit="m" metricValue="9.0" metricValueMax="10.0" metricValueMin="8.0" unit="mm" value="9.0" valueMax="10.0" valueMin="8.0">810 mm</quantity>
. Height of tergite 7 =
<quantity id="DC6834E4BD7E7F2E04477142CB8F666C" metricMagnitude="-3" metricUnit="m" metricValue="6.5" metricValueMax="7.0" metricValueMin="6.0" unit="mm" value="6.5" valueMax="7.0" valueMin="6.0">67 mm</quantity>
. Females: body length =
<quantity id="188DBBEFA4EAD2A5844E44EB1345BCEB" metricMagnitude="-2" metricUnit="m" metricValue="1.9" metricValueMax="2.0" metricValueMin="1.8" unit="mm" value="19.0" valueMax="20.0" valueMin="18.0">1820 mm</quantity>
. Width, of thoracic shield =
<quantity id="8B3BCD0EECFD063859191A6E439D6225" metricMagnitude="-3" metricUnit="m" metricValue="8.5" metricValueMax="9.0" metricValueMin="8.0" unit="mm" value="8.5" valueMax="9.0" valueMin="8.0">89 mm</quantity>
, of tergite 7 =
<quantity id="D36F9D1654D321A99F452B03B6198C7E" metricMagnitude="-3" metricUnit="m" metricValue="9.5" metricValueMax="10.0" metricValueMin="9.0" unit="mm" value="9.5" valueMax="10.0" valueMin="9.0">910 mm</quantity>
. Height of tergite 7 =
<quantity id="9568CD82C6BB23F8DA84F5906EE2AF56" metricMagnitude="-3" metricUnit="m" metricValue="7.0" metricValueMax="8.0" metricValueMin="6.0" unit="mm" value="7.0" valueMax="8.0" valueMin="6.0">68 mm</quantity>
(= highest).
</paragraph>
<paragraph id="9245108BD62F4AC32366DEA7234335FE">
<emphasis id="35DA5B279A10BC44E69EE83C894298FA" bold="true" italics="true">Colouration</emphasis>
(Fig.
<figureCitation id="CD58831A574B842479F1C133559BB91A" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Photograph of live specimens of Zephronia macula sp. nov. and habitats A G paratypes (CUMZ - MYR 00016) H habitat at Thaeo Khlong Khlung monastery (granitic rocks) I habitat at Phet Pho Thong cave (limestone). Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1212.126536.figure8" httpUri="https://binary.pensoft.net/fig/1133561">8 A G</figureCitation>
): Specimen in life with body of brown / greenish brown / reddish brown colour, tergite with dark or greenish dark spots. Head, antenna, and collum dark brown. Thoracic shield, tergites and anal shield brown. Colour in alcohol after one year faded to brown.
</paragraph>
<caption id="5A5D4FF783379C4E073741C94CC69A12" ID-DOI="10.3897/zookeys.1212.126536.figure8" ID-arpha="7546A88E-938D-5E50-BAE7-B8C7F65D57EF" httpUri="https://binary.pensoft.net/fig/1133561" startId="F8">
<paragraph id="912629C256C727D85F5D5FFFFABC0A7D">
<label id="A95C50F19F12DAF9E0701C60322EDE4D">Figure 8.</label>
</paragraph>
<paragraph id="8BBBBE26DC019F0C14880CCC582A006C">
Photograph of live specimens of
<taxonomicName id="281FF67D0E6AF4E4204CE1D8D1411F20" authorityName="Srisonchai &amp; Wesener" authorityYear="2024" class="Diplopoda" family="Zephroniidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" phylum="Arthropoda" rank="species" species="macula" status="sp. nov.">
<emphasis id="DFC964ABADE8E101BEA3A187C379CD1F" italics="true">Zephronia macula</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B5E753EA9728A18B36465F4F968FE0C7" rank="species">sp. nov.</taxonomicNameLabel>
and habitats
<emphasis id="48DD64B9BB962316F700CCDFA4FD1720" bold="true">A G</emphasis>
paratypes (
<named-content id="D9D9E27E5DE44340EB78C63D6EE7B52B" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/chulalongkorn-university-museum-natural-history" xlink_title="Chulalongkorn University Museum of Zoology">CUMZ</named-content>
- MYR 00016)
<emphasis id="B6B63F42967B5A39501D2A4937C56EB6" bold="true">H</emphasis>
habitat at Thaeo Khlong Khlung monastery (granitic rocks)
<emphasis id="8CDB3C2EF6C8292613B6978964A30FFC" bold="true">I</emphasis>
habitat at Phet Pho Thong cave (limestone). Scale bars: 0.5 mm.
</paragraph>
</caption>
<paragraph id="E5C9CE5C1BB9A27FB94C5C5802D6DFD2">
<emphasis id="3E0E2AC72DB3A2546DE8360C940AA3A5" bold="true" italics="true">Head</emphasis>
(Fig.
<figureCitation id="24CA6471DEF2F20689FB1CFA3191D77E" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Zephronia macula sp. nov., ♂ paratype (CUMZ - MYR 0016) — SEM A head, collum and antenna, dorsal view B ommatidia, inset: organ of Tömösváry C antenna, anterior view D antennal disc, anterior view E apical cones, lateral view F, I gnathochilarium, posterior and ventral views, respectively G sensory cones on central pad, ventral view H sensory cones on inner palpi, black and white arrows point to different types of cones, ventral view J epipharynx, ventral view, K mandible, mesal view. Abbreviations: 3 it = 3 - combed inner tooth, co = condylus, Cp = central pad, ct = central tooth, et = external tooth, Et = external tooth, Ia = inner area, il = incisura lateralis, ip = inner palpi, ll = lamellae linguales, me = mentum, mp = molar plate, pl = pectinate lamellae, rsp = row of spines, sc = sensory cone, st = stipites. Scale bars in millimetres." figureDoi="10.3897/zookeys.1212.126536.figure9" httpUri="https://binary.pensoft.net/fig/1133562">9 A, B</figureCitation>
): Trapeziform, densely setose; each seta located inside small pit. With 5565 ommatidia (ocelli) in males and
<quantity id="3C0B6F6B9A6F9C31E37EDCC38C3FDE09" metricMagnitude="0" metricUnit="m" metricValue="1.6637" metricValueMax="1.778" metricValueMin="1.5493999999999999" unit="in" value="65.5" valueMax="70.0" valueMin="61.0">6170 in</quantity>
females. At rim of antennal groove with aberrant ommatidium. Organ of Tömösváry located near base of antenna, clearly separated from eye field. No sclerotized crest / ridge between antennal socket and eye field.
</paragraph>
<caption id="10707867513CAAC8F985C5D77EDCD960" ID-DOI="10.3897/zookeys.1212.126536.figure9" ID-arpha="6B7ED5F9-FFD2-5928-A779-96A9FACD00C0" httpUri="https://binary.pensoft.net/fig/1133562" startId="F9">
<paragraph id="5195198D8693C4F4C8B1DEF56D57AECA">
<label id="A2562A110146A7491B331A5821668B98">Figure 9.</label>
</paragraph>
<paragraph id="BF5942CED1E657E9EC7446D552AEAB01">
<taxonomicName id="AF78708B6DAEBC693A02DFCC8C2E5A3B" authorityName="Srisonchai &amp; Wesener" authorityYear="2024" class="Diplopoda" family="Zephroniidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" phylum="Arthropoda" rank="species" species="macula" status="sp. nov.">
<emphasis id="49A48CE95EF7109042D6556080C4A2C9" italics="true">Zephronia macula</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CCA90470911435EE7A2DA2D294E23DD6" rank="species">sp. nov.</taxonomicNameLabel>
, ♂ paratype (
<named-content id="D2489C490CF96B4E5CA24727544869E6" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/chulalongkorn-university-museum-natural-history" xlink_title="Chulalongkorn University Museum of Zoology">CUMZ</named-content>
- MYR 0016) —
<abbrev id="ABBRID0EIZDI" xlink_title="scanning electron microscopy">SEM</abbrev>
<emphasis id="07DC6BAE41AB2C6D1158BD76ECC7C8D8" bold="true">A</emphasis>
head, collum and antenna, dorsal view
<emphasis id="A3628DBD2C1A7A1142E3230E2E90213B" bold="true">B</emphasis>
ommatidia, inset: organ of Tömösváry
<emphasis id="E69E715BE9E29ED15D85202A02CDB2B4" bold="true">C</emphasis>
antenna, anterior view
<emphasis id="336A2E685530F4A41A54ED28921D5A4A" bold="true">D</emphasis>
antennal disc, anterior view
<emphasis id="3F6D8832F0ADE186878D9DF8A8981EC7" bold="true">E</emphasis>
apical cones, lateral view
<emphasis id="205B9845917DC3FBD6B52B760E5CE241" bold="true">F, I</emphasis>
gnathochilarium, posterior and ventral views, respectively
<emphasis id="13DBA65BFF821276E5FB2FEA6DBDE612" bold="true">G</emphasis>
sensory cones on central pad, ventral view
<emphasis id="42262450A124DA692E7A694FA8487CAF" bold="true">H</emphasis>
sensory cones on inner palpi, black and white arrows point to different types of cones, ventral view
<emphasis id="AEF9619BB5BB0C419F8E3DB274724517" bold="true">J</emphasis>
epipharynx, ventral view,
<emphasis id="CC31B3B6ED58BDB33590CB906BC9239D" bold="true">K</emphasis>
mandible, mesal view. Abbreviations:
<abbrev id="ABBRID0EA1DI" xlink_title="3-combed inner tooth of mandible">3 it</abbrev>
= 3 - combed inner tooth,
<abbrev id="ABBRID0EE1DI" xlink_title="condylus of mandible">co</abbrev>
= condylus,
<abbrev id="ABBRID0EI1DI" xlink_title="central pad of gnathochilarium">Cp</abbrev>
= central pad,
<abbrev id="ABBRID0EM1DI" xlink_title="central tooth">ct</abbrev>
= central tooth,
<abbrev id="ABBRID0EQ1DI" xlink_title="external tooth of epipharynx">et</abbrev>
= external tooth,
<abbrev id="ABBRID0EU1DI" xlink_title="external tooth of mandibular gnathal lobe">Et</abbrev>
= external tooth,
<abbrev id="ABBRID0EY1DI" xlink_title="inner area of mandible">Ia</abbrev>
= inner area,
<abbrev id="ABBRID0E31DI" xlink_title="incisura lateralis of the head">il</abbrev>
= incisura lateralis,
<abbrev id="ABBRID0EA2DI" xlink_title="inner palpi of gnathochilarium">ip</abbrev>
= inner palpi,
<abbrev id="ABBRID0EE2DI" xlink_title="lamellae lingulales of gnathochilarium">ll</abbrev>
= lamellae linguales,
<abbrev id="ABBRID0EI2DI" xlink_title="mentum of gnathochilarium">me</abbrev>
= mentum,
<abbrev id="ABBRID0EM2DI" xlink_title="molar plate of mandible">mp</abbrev>
= molar plate,
<abbrev id="ABBRID0EQ2DI" xlink_title="pectinate lamellae of mandible">pl</abbrev>
= pectinate lamellae,
<abbrev id="ABBRID0EU2DI" xlink_title="row of spines">rsp</abbrev>
= row of spines,
<abbrev id="ABBRID0EY2DI" xlink_title="sensory cone">sc</abbrev>
= sensory cone,
<abbrev id="ABBRID0E32DI" xlink_title="stipites of gnathochilarium">st</abbrev>
= stipites. Scale bars in millimetres.
</paragraph>
</caption>
<paragraph id="1984F7D74C9E098F26C8AE956C381258">
<emphasis id="8E3C3EF4E39ACE9C27D3FC26B64EFC26" bold="true" italics="true">Antennae</emphasis>
(Figs
<figureCitation id="E9917342454390B3F2819FC0AC0B8516" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Photograph of live specimens of Zephronia macula sp. nov. and habitats A G paratypes (CUMZ - MYR 00016) H habitat at Thaeo Khlong Khlung monastery (granitic rocks) I habitat at Phet Pho Thong cave (limestone). Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1212.126536.figure8" httpUri="https://binary.pensoft.net/fig/1133561">8 C, D</figureCitation>
,
<figureCitation id="4EB1FAAFD127BD25D50C9ECD76791E25" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Zephronia macula sp. nov., ♂ paratype (CUMZ - MYR 0016) — SEM A head, collum and antenna, dorsal view B ommatidia, inset: organ of Tömösváry C antenna, anterior view D antennal disc, anterior view E apical cones, lateral view F, I gnathochilarium, posterior and ventral views, respectively G sensory cones on central pad, ventral view H sensory cones on inner palpi, black and white arrows point to different types of cones, ventral view J epipharynx, ventral view, K mandible, mesal view. Abbreviations: 3 it = 3 - combed inner tooth, co = condylus, Cp = central pad, ct = central tooth, et = external tooth, Et = external tooth, Ia = inner area, il = incisura lateralis, ip = inner palpi, ll = lamellae linguales, me = mentum, mp = molar plate, pl = pectinate lamellae, rsp = row of spines, sc = sensory cone, st = stipites. Scale bars in millimetres." figureDoi="10.3897/zookeys.1212.126536.figure9" httpUri="https://binary.pensoft.net/fig/1133562">9 A, C E</figureCitation>
): Short and stout, covered by long and dense setae; last antennomere reaching back to leg pair 2 or 3 when stretched ventrally. Lengths of antennomeres: 1 = 2 = 3 = 4 &lt;5 &lt;&lt; 6. Antennomere 6 strongly flattened apically, axe-shaped; apically with sensilla basiconica. Apical disc slightly concave, with 4857 apical cones (male) or 3643 (female).
</paragraph>
<paragraph id="2371C2DB6A8F8328CBBFB05F8B3A2DF2">
<emphasis id="0F9DC406E63F287460E6E3BDCA7C3C0D" bold="true" italics="true">Epipharynx</emphasis>
(Fig.
<figureCitation id="4825B480CCB8C734B365A4647E4AB172" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Zephronia macula sp. nov., ♂ paratype (CUMZ - MYR 0016) — SEM A head, collum and antenna, dorsal view B ommatidia, inset: organ of Tömösváry C antenna, anterior view D antennal disc, anterior view E apical cones, lateral view F, I gnathochilarium, posterior and ventral views, respectively G sensory cones on central pad, ventral view H sensory cones on inner palpi, black and white arrows point to different types of cones, ventral view J epipharynx, ventral view, K mandible, mesal view. Abbreviations: 3 it = 3 - combed inner tooth, co = condylus, Cp = central pad, ct = central tooth, et = external tooth, Et = external tooth, Ia = inner area, il = incisura lateralis, ip = inner palpi, ll = lamellae linguales, me = mentum, mp = molar plate, pl = pectinate lamellae, rsp = row of spines, sc = sensory cone, st = stipites. Scale bars in millimetres." figureDoi="10.3897/zookeys.1212.126536.figure9" httpUri="https://binary.pensoft.net/fig/1133562">9 J</figureCitation>
): With a large central tooth (
<abbrev id="ABBRID0E63DI" xlink_title="central tooth">ct</abbrev>
); inner tooth conspicuous and swollen; laterally with group of long external teeth (
<abbrev id="ABBRID0ED4DI" xlink_title="external tooth of epipharynx">et</abbrev>
); inner area with a single row of fringed spines (
<abbrev id="ABBRID0EH4DI" xlink_title="row of spines">rsp</abbrev>
) on each side.
</paragraph>
<paragraph id="17D09F318827CA8711A74D263D694240">
<emphasis id="1AF08ACD124E1DEEA560B591020E124D" bold="true" italics="true">Gnathochilarium</emphasis>
(Fig.
<figureCitation id="01F42882D29C0D469CBE4560F23F1F24" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Zephronia macula sp. nov., ♂ paratype (CUMZ - MYR 0016) — SEM A head, collum and antenna, dorsal view B ommatidia, inset: organ of Tömösváry C antenna, anterior view D antennal disc, anterior view E apical cones, lateral view F, I gnathochilarium, posterior and ventral views, respectively G sensory cones on central pad, ventral view H sensory cones on inner palpi, black and white arrows point to different types of cones, ventral view J epipharynx, ventral view, K mandible, mesal view. Abbreviations: 3 it = 3 - combed inner tooth, co = condylus, Cp = central pad, ct = central tooth, et = external tooth, Et = external tooth, Ia = inner area, il = incisura lateralis, ip = inner palpi, ll = lamellae linguales, me = mentum, mp = molar plate, pl = pectinate lamellae, rsp = row of spines, sc = sensory cone, st = stipites. Scale bars in millimetres." figureDoi="10.3897/zookeys.1212.126536.figure9" httpUri="https://binary.pensoft.net/fig/1133562">9 F I</figureCitation>
): As in
<taxonomicName id="3CCB4522ED892265EFF519A27B09CF1D" authorityName="Srisonchai &amp; Wesener" authorityYear="2024" class="Diplopoda" family="Zephroniidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" phylum="Arthropoda" rank="species" species="chantaburiensis" status="sp. nov.">
<emphasis id="FF62604F688DB30D2463D89AE97136C1" italics="true">Z. chantaburiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7F94CA2572EE5B37390B1137CCF37B5A" rank="species">sp. nov.</taxonomicNameLabel>
Lamellae linguales (
<abbrev id="ABBRID0EB5DI" xlink_title="lamellae lingulales of gnathochilarium">ll</abbrev>
) oval, apically concave, with long setae. Central pads (
<abbrev id="ABBRID0EF5DI" xlink_title="central pad of gnathochilarium">Cp</abbrev>
) modified, with numerous “ pillows ” of sensory cones (
<abbrev id="ABBRID0EJ5DI" xlink_title="sensory cone">sc</abbrev>
) (Fig.
<figureCitation id="C4D9FA703A6EC4CA6A3DCEE9BFE4FE8A" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Zephronia macula sp. nov., ♂ paratype (CUMZ - MYR 0016) — SEM A head, collum and antenna, dorsal view B ommatidia, inset: organ of Tömösváry C antenna, anterior view D antennal disc, anterior view E apical cones, lateral view F, I gnathochilarium, posterior and ventral views, respectively G sensory cones on central pad, ventral view H sensory cones on inner palpi, black and white arrows point to different types of cones, ventral view J epipharynx, ventral view, K mandible, mesal view. Abbreviations: 3 it = 3 - combed inner tooth, co = condylus, Cp = central pad, ct = central tooth, et = external tooth, Et = external tooth, Ia = inner area, il = incisura lateralis, ip = inner palpi, ll = lamellae linguales, me = mentum, mp = molar plate, pl = pectinate lamellae, rsp = row of spines, sc = sensory cone, st = stipites. Scale bars in millimetres." figureDoi="10.3897/zookeys.1212.126536.figure9" httpUri="https://binary.pensoft.net/fig/1133562">9 H</figureCitation>
); two different types of sensory cones (one with a pillow and another without a pillow). Stipites (
<abbrev id="ABBRID0ER5DI" xlink_title="stipites of gnathochilarium">st</abbrev>
) large and stout, densely setose. Mentum (
<abbrev id="ABBRID0EV5DI" xlink_title="mentum of gnathochilarium">me</abbrev>
) large and broad, fused, with long setae. Lateral palpi inconspicuous. Inner palpi (
<abbrev id="ABBRID0EZ5DI" xlink_title="inner palpi of gnathochilarium">ip</abbrev>
) with sensory cones (
<abbrev id="ABBRID0E45DI" xlink_title="sensory cone">sc</abbrev>
) arranged in single field (Fig.
<figureCitation id="BD973E2C740AD3089695885E404789AE" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Zephronia macula sp. nov., ♂ paratype (CUMZ - MYR 0016) — SEM A head, collum and antenna, dorsal view B ommatidia, inset: organ of Tömösváry C antenna, anterior view D antennal disc, anterior view E apical cones, lateral view F, I gnathochilarium, posterior and ventral views, respectively G sensory cones on central pad, ventral view H sensory cones on inner palpi, black and white arrows point to different types of cones, ventral view J epipharynx, ventral view, K mandible, mesal view. Abbreviations: 3 it = 3 - combed inner tooth, co = condylus, Cp = central pad, ct = central tooth, et = external tooth, Et = external tooth, Ia = inner area, il = incisura lateralis, ip = inner palpi, ll = lamellae linguales, me = mentum, mp = molar plate, pl = pectinate lamellae, rsp = row of spines, sc = sensory cone, st = stipites. Scale bars in millimetres." figureDoi="10.3897/zookeys.1212.126536.figure9" httpUri="https://binary.pensoft.net/fig/1133562">9 G</figureCitation>
).
</paragraph>
<paragraph id="DA9083073B10820684BE157080D02A34">
<emphasis id="1C5B47E7AC7ECA204FE9792BF432DDC8" bold="true">
<emphasis id="089E69768F8917EEE1C4DC59FD35A26B" bold="true" italics="true">Mandibles</emphasis>
(
<emphasis id="40494F0B04CE78343D56F546ACB8B070" bold="true" italics="true">gnathal lobe</emphasis>
)
</emphasis>
(Fig.
<figureCitation id="BDBE5E2C1D9B871FBA2FDE1E71C7145E" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Zephronia macula sp. nov., ♂ paratype (CUMZ - MYR 0016) — SEM A head, collum and antenna, dorsal view B ommatidia, inset: organ of Tömösváry C antenna, anterior view D antennal disc, anterior view E apical cones, lateral view F, I gnathochilarium, posterior and ventral views, respectively G sensory cones on central pad, ventral view H sensory cones on inner palpi, black and white arrows point to different types of cones, ventral view J epipharynx, ventral view, K mandible, mesal view. Abbreviations: 3 it = 3 - combed inner tooth, co = condylus, Cp = central pad, ct = central tooth, et = external tooth, Et = external tooth, Ia = inner area, il = incisura lateralis, ip = inner palpi, ll = lamellae linguales, me = mentum, mp = molar plate, pl = pectinate lamellae, rsp = row of spines, sc = sensory cone, st = stipites. Scale bars in millimetres." figureDoi="10.3897/zookeys.1212.126536.figure9" httpUri="https://binary.pensoft.net/fig/1133562">9 K</figureCitation>
): With undivided external tooth (
<abbrev id="ABBRID0ES6DI" xlink_title="external tooth of mandibular gnathal lobe">Et</abbrev>
) and with prominent 3 - combed inner tooth (
<abbrev id="ABBRID0EW6DI" xlink_title="3-combed inner tooth of mandible">3 it</abbrev>
). With 56 pectinate lamellae (
<abbrev id="ABBRID0E16DI" xlink_title="pectinate lamellae of mandible">pl</abbrev>
). Inner area (
<abbrev id="ABBRID0E56DI" xlink_title="inner area of mandible">Ia</abbrev>
) with group of long and tiny teeth, spine-like. Molar plate (
<abbrev id="ABBRID0EDAAK" xlink_title="molar plate of mandible">mp</abbrev>
) flat, velvet-like; lacking a membranous fringe. Condylus (
<abbrev id="ABBRID0EHAAK" xlink_title="condylus of mandible">co</abbrev>
) conspicuous, apically with 2 distinct ridges.
</paragraph>
<paragraph id="AF11FD37F6EE5400764B04892B2F07E8">
<emphasis id="C28642A38BE62E4F765C5A6E9F6120FF" bold="true" italics="true">Tegument</emphasis>
(Figs
<figureCitation id="9FB35655B540EB0E5C6E1DA3ADF28E55" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Photograph of live specimens of Zephronia macula sp. nov. and habitats A G paratypes (CUMZ - MYR 00016) H habitat at Thaeo Khlong Khlung monastery (granitic rocks) I habitat at Phet Pho Thong cave (limestone). Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1212.126536.figure8" httpUri="https://binary.pensoft.net/fig/1133561">8 A G</figureCitation>
,
<figureCitation id="CCB00C65D1FD334303F2C26E4A1746D6" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Zephronia macula sp. nov., ♂ paratype (CUMZ - MYR 0016) — SEM A surface of collum B, C thoracic shield, all in dorsal views (anterior margin and surface, respectively) D G midbody tergite, all in dorsal views (overview, anterior margin, surface, and posterior margin, respectively) H J anal shield, all in dorsal views (overview, anterior margin and cuticular impressions, respectively) K M anal shield, underside, all in ventral views (surface, posterior margin, and overview, respectively). Scale bars in millimetres." figureDoi="10.3897/zookeys.1212.126536.figure10" httpUri="https://binary.pensoft.net/fig/1133563">10</figureCitation>
): Dull; collum, thoracic shield, tergite and anal shield with tiny golden setae; each seta located in small pit. Anterior margins of midbody tergite and of anal shield with lower number of setae than posterior margins. Posterior rim on dorsal and ventral side of anal shield with a few small setae.
</paragraph>
<caption id="ACFFC793AF9F47A0022F284483D167D0" ID-DOI="10.3897/zookeys.1212.126536.figure10" ID-arpha="92A30B93-1E4F-5CF4-95B5-74956FB8AEA8" httpUri="https://binary.pensoft.net/fig/1133563" startId="F10">
<paragraph id="F2E5BBA230641393A3FF93F615B5D82C">
<label id="6EC01FB57493937A8B8E01DE6324B25A">Figure 10.</label>
</paragraph>
<paragraph id="47D97C0C345162D6BFD8D9D268AFA70F">
<taxonomicName id="D730CBF8C22B30D815D37F764996FE91" authorityName="Srisonchai &amp; Wesener" authorityYear="2024" class="Diplopoda" family="Zephroniidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" phylum="Arthropoda" rank="species" species="macula" status="sp. nov.">
<emphasis id="8BBFC662DEA25D67DF397F5AF469D690" italics="true">Zephronia macula</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1537A94C5F6BF84E28C984CC1CDC8F64" rank="species">sp. nov.</taxonomicNameLabel>
, ♂ paratype (
<named-content id="A4B2F57A802FE530A3374B45129F8812" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/chulalongkorn-university-museum-natural-history" xlink_title="Chulalongkorn University Museum of Zoology">CUMZ</named-content>
- MYR 0016) —
<abbrev id="ABBRID0EDCAK" xlink_title="scanning electron microscopy">SEM</abbrev>
<emphasis id="35CE99FF0BD74166D9B3F626B2836919" bold="true">A</emphasis>
surface of collum
<emphasis id="9506C86EAB1D1EF6DE1945208DA1FADD" bold="true">B, C</emphasis>
thoracic shield, all in dorsal views (anterior margin and surface, respectively)
<emphasis id="202957225487D568D7E1382386098BBF" bold="true">D G</emphasis>
midbody tergite, all in dorsal views (overview, anterior margin, surface, and posterior margin, respectively)
<emphasis id="CAA6F6BB3FC767C32007CADADED4A06F" bold="true">H J</emphasis>
anal shield, all in dorsal views (overview, anterior margin and cuticular impressions, respectively)
<emphasis id="2101CCE88425BBA75A30D68F86FCC7EC" bold="true">K M</emphasis>
anal shield, underside, all in ventral views (surface, posterior margin, and overview, respectively). Scale bars in millimetres.
</paragraph>
</caption>
<paragraph id="4EB85AC24A4EBA80D72A6C8D0C358B48">
<emphasis id="1183D98C28EEF87D92A617AFD29231F7" bold="true" italics="true">Collum</emphasis>
(Figs
<figureCitation id="FCF47BAFB8741E29BF08AF6BABC82B52" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Photograph of live specimens of Zephronia macula sp. nov. and habitats A G paratypes (CUMZ - MYR 00016) H habitat at Thaeo Khlong Khlung monastery (granitic rocks) I habitat at Phet Pho Thong cave (limestone). Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1212.126536.figure8" httpUri="https://binary.pensoft.net/fig/1133561">8 C, D</figureCitation>
,
<figureCitation id="830424205D7CA638A0A21EE584A47DA6" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Zephronia macula sp. nov. — SEM A G, L ♂ paratype (CUMZ - MYR 0016) J, K ♀ paratype (CUMZ - MYR 0015) A collum B seventh pleurite C subanal plate D legs 1 E stigmatic plate 1 F legs 2 with gonopores G leg 10 (arrow points to femoral ridge) H coxal process on leg 10 I small teeth on femur J leg 2 with vulvae K vulvae L tarsus 10. Abbreviation: bu = bursa, cx = coxa, fe = femur, op = operculum, po = postfemur, pre = prefemur, st-pl = stigmatic plate, ta = tarsus, ti = tibia. Scale bars in millimetres." figureDoi="10.3897/zookeys.1212.126536.figure11" httpUri="https://binary.pensoft.net/fig/1133564">11 A</figureCitation>
): Sub-semicircular; tip of lateral margin obtuse, covered with fine setae.
</paragraph>
<caption id="78F5B2BC9FA9FEE45F12A170CE860990" ID-DOI="10.3897/zookeys.1212.126536.figure11" ID-arpha="5D052990-499A-571B-B75C-D719545400EE" httpUri="https://binary.pensoft.net/fig/1133564" startId="F11">
<paragraph id="A1A81ED35572D4CABE8795C25D3CF47A">
<label id="D008B1142CE08C39C15E62F76497EC4A">Figure 11.</label>
</paragraph>
<paragraph id="636176FB2F61D8F0279A4D03B448D84E">
<taxonomicName id="DCC9A20603FF3CC23F23AAB7750ADB94" authorityName="Srisonchai &amp; Wesener" authorityYear="2024" class="Diplopoda" family="Zephroniidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" phylum="Arthropoda" rank="species" species="macula" status="sp. nov.">
<emphasis id="E6EDBE328C335B4636BE9B7B5F0CC328" italics="true">Zephronia macula</emphasis>
</taxonomicName>
<taxonomicNameLabel id="713056AFD7FF59EAE2F04B039D5489C7" rank="species">sp. nov.</taxonomicNameLabel>
<abbrev id="ABBRID0EBEAK" xlink_title="scanning electron microscopy">SEM</abbrev>
<emphasis id="046421459B155F6E233D9F4B4FBE8418" bold="true">A G, L</emphasis>
♂ paratype (
<named-content id="CC1F2C817FBA9CA5CE9FDDCE46D333BA" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/chulalongkorn-university-museum-natural-history" xlink_title="Chulalongkorn University Museum of Zoology">CUMZ</named-content>
- MYR 0016)
<emphasis id="C0EDC4052F00B9228B4FF9844B0653B6" bold="true">J, K</emphasis>
♀ paratype (
<named-content id="A3C4F0C5E0C4A5100E9FB3C559A5A413" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/chulalongkorn-university-museum-natural-history" xlink_title="Chulalongkorn University Museum of Zoology">CUMZ</named-content>
- MYR 0015)
<emphasis id="050B138EAFC0D3C30C300EAAC29676A9" bold="true">A</emphasis>
collum
<emphasis id="1E2752DD3929D15145B066E5591A4E13" bold="true">B</emphasis>
seventh pleurite
<emphasis id="B8CE008BD14D1A3FE95FB5961C080D9B" bold="true">C</emphasis>
subanal plate
<emphasis id="AAD064CAE4B1CF151761790587015CF1" bold="true">D</emphasis>
legs 1
<emphasis id="3AF19E769C83260DE6DA9E285A01FB52" bold="true">E</emphasis>
stigmatic plate 1
<emphasis id="258734E27737CA706A0124910EFD497C" bold="true">F</emphasis>
legs 2 with gonopores
<emphasis id="2336FF5E346112A89855FCF9F93DBFEB" bold="true">G</emphasis>
leg 10 (arrow points to femoral ridge)
<emphasis id="050B13965D80370189281FF4C3E4F838" bold="true">H</emphasis>
coxal process on leg 10
<emphasis id="87195597C9C3B6DFF2C78EC2D668232C" bold="true">I</emphasis>
small teeth on femur
<emphasis id="7FC962F8521CB94119CF03270F8931D1" bold="true">J</emphasis>
leg 2 with vulvae
<emphasis id="BC41DC76AFBF14975712657BA4A2BC5F" bold="true">K</emphasis>
vulvae
<emphasis id="07EABFF123161B3F6B78A94554D83041" bold="true">L</emphasis>
tarsus 10. Abbreviation:
<abbrev id="ABBRID0EJFAK" xlink_title="bursa of vulva">bu</abbrev>
= bursa,
<abbrev id="ABBRID0ENFAK" xlink_title="coxa">cx</abbrev>
= coxa,
<abbrev id="ABBRID0ERFAK" xlink_title="femur">fe</abbrev>
= femur,
<abbrev id="ABBRID0EVFAK" xlink_title="operculum of vulva">op</abbrev>
= operculum,
<abbrev id="ABBRID0EZFAK" xlink_title="postfemur">po</abbrev>
= postfemur,
<abbrev id="ABBRID0E4FAK" xlink_title="prefemur">pre</abbrev>
= prefemur,
<abbrev id="ABBRID0EBGAK" xlink_title="stigmatic plate">st-pl</abbrev>
= stigmatic plate,
<abbrev id="ABBRID0EFGAK" xlink_title="tarsus">ta</abbrev>
= tarsus,
<abbrev id="ABBRID0EJGAK" xlink_title="tibia">ti</abbrev>
= tibia. Scale bars in millimetres.
</paragraph>
</caption>
<paragraph id="1B50A13B8D1141204B5B7A16EDDBBF83">
<emphasis id="B868E8098622C8B46FCAB0FEE254DD5C" bold="true" italics="true">Thoracic shield</emphasis>
(Fig.
<figureCitation id="CC2EE88AC07E8AFC96D54F72CCD91B06" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Zephronia macula sp. nov., ♂ paratype (CUMZ - MYR 0016) — SEM A surface of collum B, C thoracic shield, all in dorsal views (anterior margin and surface, respectively) D G midbody tergite, all in dorsal views (overview, anterior margin, surface, and posterior margin, respectively) H J anal shield, all in dorsal views (overview, anterior margin and cuticular impressions, respectively) K M anal shield, underside, all in ventral views (surface, posterior margin, and overview, respectively). Scale bars in millimetres." figureDoi="10.3897/zookeys.1212.126536.figure10" httpUri="https://binary.pensoft.net/fig/1133563">10 B</figureCitation>
): With a shallow, large, wide groove separated by a large and long ridge. Slope towards groove without keels. Groove smooth, without setae; on a ridge with a few setae.
</paragraph>
<paragraph id="E56C06464ABFC188C84A7C8E97FA931B">
<emphasis id="1C4589EAC932249FBF15AAED03B8C759" bold="true" italics="true">Midbody tergite</emphasis>
(Fig.
<figureCitation id="F2E62C7DA6941067696237824F676FE4" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Zephronia macula sp. nov., ♂ paratype (CUMZ - MYR 0016) — SEM A surface of collum B, C thoracic shield, all in dorsal views (anterior margin and surface, respectively) D G midbody tergite, all in dorsal views (overview, anterior margin, surface, and posterior margin, respectively) H J anal shield, all in dorsal views (overview, anterior margin and cuticular impressions, respectively) K M anal shield, underside, all in ventral views (surface, posterior margin, and overview, respectively). Scale bars in millimetres." figureDoi="10.3897/zookeys.1212.126536.figure10" httpUri="https://binary.pensoft.net/fig/1133563">10 D G</figureCitation>
): With a row of oval impressions; anterior margin with tiny tubercles. Inner area with a crenate barrier; a wide and shallow groove. Tips of midbody paratergites projecting ventro-posteriad.
</paragraph>
<paragraph id="9E69D45A2D018188FEB9CC9D72C65626">
<emphasis id="5E123FE6579CE722E03B56A8CAFBE677" bold="true" italics="true">Anal shield</emphasis>
(Fig.
<figureCitation id="8B226E214ED41207E976C7723F956F14" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Zephronia macula sp. nov., ♂ paratype (CUMZ - MYR 0016) — SEM A surface of collum B, C thoracic shield, all in dorsal views (anterior margin and surface, respectively) D G midbody tergite, all in dorsal views (overview, anterior margin, surface, and posterior margin, respectively) H J anal shield, all in dorsal views (overview, anterior margin and cuticular impressions, respectively) K M anal shield, underside, all in ventral views (surface, posterior margin, and overview, respectively). Scale bars in millimetres." figureDoi="10.3897/zookeys.1212.126536.figure10" httpUri="https://binary.pensoft.net/fig/1133563">10 H M</figureCitation>
): In female large and well-rounded, in male slenderer. With a row of oval impressions, edge of anterior margin with numerous tiny tubercles (sometimes arranged into rows). Inner area with a row of oval impressions, forming a crenate barrier; with a wide and shallow groove.
</paragraph>
<paragraph id="6B405B0427D7BFD23FA49B05D31D879A">
<emphasis id="27734FB16D4584271ECA7573B9427C00" bold="true" italics="true">Endotergum of thoracic shield and midbody tergite</emphasis>
(Fig.
<figureCitation id="E0CA87B30B8D80BBC79B685B463AAEC4" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Endoterga of thoracic shields and midbody tergites — SEM A D Zephronia chantaburiensis sp. nov. (♂ paratype, CUMZ - MYR 0014) E H Zephronia macula sp. nov. (♂ paratype, CUMZ - MYR 0016) A, B, E, F thoracic shields C, D, G, H midbody tergites. Abbreviations: cp = cuticular impression, ia = inner area, ma = middle area, oa = outer area, pm = posterior margin. Scale bars in millimetres." figureDoi="10.3897/zookeys.1212.126536.figure14" httpUri="https://binary.pensoft.net/fig/1133567">14 E H</figureCitation>
): Thoracic shield similar to midbody tergites. Overall surface smooth. Posterior margin (
<abbrev id="ABBRID0EQIAK" xlink_title="posterior margin of endotergum">pm</abbrev>
) flat, not modified. Outer area (os) without setae. Bristles arranged in one row; tip of longest bristles reaching to posterior margin (Fig.
<figureCitation id="061029553D2359C8B6822B58C6412BAA" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Endoterga of thoracic shields and midbody tergites — SEM A D Zephronia chantaburiensis sp. nov. (♂ paratype, CUMZ - MYR 0014) E H Zephronia macula sp. nov. (♂ paratype, CUMZ - MYR 0016) A, B, E, F thoracic shields C, D, G, H midbody tergites. Abbreviations: cp = cuticular impression, ia = inner area, ma = middle area, oa = outer area, pm = posterior margin. Scale bars in millimetres." figureDoi="10.3897/zookeys.1212.126536.figure14" httpUri="https://binary.pensoft.net/fig/1133567">14 G, H</figureCitation>
showing short bristles). Middle area (
<abbrev id="ABBRID0EYIAK" xlink_title="middle area of endotergum">ma</abbrev>
) with a single row of conspicuous, oval cuticular impressions; distance between impressions twice as long as individual diameter. Inner area (
<abbrev id="ABBRID0E3IAK" xlink_title="inner area of endotergum">ia</abbrev>
) without tubercles or setae.
</paragraph>
<paragraph id="624660DA3D299A37439CFFCA18634156">
<emphasis id="E64F8E7D213B6A6A75C7C1FD76F04299" bold="true" italics="true">Pleurite (laterotergite)</emphasis>
(Fig.
<figureCitation id="FB7D0083E969FA0026D9FAC31153E269" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Zephronia macula sp. nov. — SEM A G, L ♂ paratype (CUMZ - MYR 0016) J, K ♀ paratype (CUMZ - MYR 0015) A collum B seventh pleurite C subanal plate D legs 1 E stigmatic plate 1 F legs 2 with gonopores G leg 10 (arrow points to femoral ridge) H coxal process on leg 10 I small teeth on femur J leg 2 with vulvae K vulvae L tarsus 10. Abbreviation: bu = bursa, cx = coxa, fe = femur, op = operculum, po = postfemur, pre = prefemur, st-pl = stigmatic plate, ta = tarsus, ti = tibia. Scale bars in millimetres." figureDoi="10.3897/zookeys.1212.126536.figure11" httpUri="https://binary.pensoft.net/fig/1133564">11 B</figureCitation>
): Densely setose. First pleurite boomerang-shaped; apical margin attenuated; strongly projecting into sharp tip. Pleurite 2 projecting into an obtuse tip. Remaining pleurites flat and wide, lamella-like, apical margin extended.
</paragraph>
<paragraph id="C6CC485DA25DFDECC157063C1FDD628B">
<emphasis id="52B80B5B79672CC5E2F637B5AD2B26BC" bold="true" italics="true">Subanal plate of female</emphasis>
(Fig.
<figureCitation id="02E1F40E2C7C09802A3FA2A01A584946" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Zephronia macula sp. nov. — SEM A G, L ♂ paratype (CUMZ - MYR 0016) J, K ♀ paratype (CUMZ - MYR 0015) A collum B seventh pleurite C subanal plate D legs 1 E stigmatic plate 1 F legs 2 with gonopores G leg 10 (arrow points to femoral ridge) H coxal process on leg 10 I small teeth on femur J leg 2 with vulvae K vulvae L tarsus 10. Abbreviation: bu = bursa, cx = coxa, fe = femur, op = operculum, po = postfemur, pre = prefemur, st-pl = stigmatic plate, ta = tarsus, ti = tibia. Scale bars in millimetres." figureDoi="10.3897/zookeys.1212.126536.figure11" httpUri="https://binary.pensoft.net/fig/1133564">11 C</figureCitation>
): Sparsely setose. Large and broad (sometimes quite narrow), semicircular; apical margin slightly concave (rarely truncate / obtuse).
</paragraph>
<paragraph id="2E5DB1132ACB773A64AFC7F29537C26C">
<emphasis id="D1DDD7D68B04A53E6226523908CEFC32" bold="true" italics="true">Stigmatic plates</emphasis>
(Fig.
<figureCitation id="376BBA6199FE102189BCB55BC2A1B4AB" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Zephronia macula sp. nov. — SEM A G, L ♂ paratype (CUMZ - MYR 0016) J, K ♀ paratype (CUMZ - MYR 0015) A collum B seventh pleurite C subanal plate D legs 1 E stigmatic plate 1 F legs 2 with gonopores G leg 10 (arrow points to femoral ridge) H coxal process on leg 10 I small teeth on femur J leg 2 with vulvae K vulvae L tarsus 10. Abbreviation: bu = bursa, cx = coxa, fe = femur, op = operculum, po = postfemur, pre = prefemur, st-pl = stigmatic plate, ta = tarsus, ti = tibia. Scale bars in millimetres." figureDoi="10.3897/zookeys.1212.126536.figure11" httpUri="https://binary.pensoft.net/fig/1133564">11 D, E</figureCitation>
): First stigmatic plate large, slender; apex well-rounded, slightly curved towards basal part of prefemur. Second stigmatic plates in both male (Fig.
<figureCitation id="179A77D8441DA62C44C614FEDFF36753" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Zephronia macula sp. nov. — SEM A G, L ♂ paratype (CUMZ - MYR 0016) J, K ♀ paratype (CUMZ - MYR 0015) A collum B seventh pleurite C subanal plate D legs 1 E stigmatic plate 1 F legs 2 with gonopores G leg 10 (arrow points to femoral ridge) H coxal process on leg 10 I small teeth on femur J leg 2 with vulvae K vulvae L tarsus 10. Abbreviation: bu = bursa, cx = coxa, fe = femur, op = operculum, po = postfemur, pre = prefemur, st-pl = stigmatic plate, ta = tarsus, ti = tibia. Scale bars in millimetres." figureDoi="10.3897/zookeys.1212.126536.figure11" httpUri="https://binary.pensoft.net/fig/1133564">11 F</figureCitation>
) and female (Fig.
<figureCitation id="D7DB43F71D069DEEA24E462289D4CDE7" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Zephronia macula sp. nov. — SEM A G, L ♂ paratype (CUMZ - MYR 0016) J, K ♀ paratype (CUMZ - MYR 0015) A collum B seventh pleurite C subanal plate D legs 1 E stigmatic plate 1 F legs 2 with gonopores G leg 10 (arrow points to femoral ridge) H coxal process on leg 10 I small teeth on femur J leg 2 with vulvae K vulvae L tarsus 10. Abbreviation: bu = bursa, cx = coxa, fe = femur, op = operculum, po = postfemur, pre = prefemur, st-pl = stigmatic plate, ta = tarsus, ti = tibia. Scale bars in millimetres." figureDoi="10.3897/zookeys.1212.126536.figure11" httpUri="https://binary.pensoft.net/fig/1133564">11 J</figureCitation>
) only very weakly curved towards coxa.
</paragraph>
<paragraph id="D6BE20D44C480CBC1D8AEC50AAB663A8">
<emphasis id="16FD2DDF6B923788C2BA9F16AB96ED24" bold="true" italics="true">Legs</emphasis>
(Fig.
<figureCitation id="B5E56BAA4BF48EC9BA15FBF1EC8C69E0" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Zephronia macula sp. nov. — SEM A G, L ♂ paratype (CUMZ - MYR 0016) J, K ♀ paratype (CUMZ - MYR 0015) A collum B seventh pleurite C subanal plate D legs 1 E stigmatic plate 1 F legs 2 with gonopores G leg 10 (arrow points to femoral ridge) H coxal process on leg 10 I small teeth on femur J leg 2 with vulvae K vulvae L tarsus 10. Abbreviation: bu = bursa, cx = coxa, fe = femur, op = operculum, po = postfemur, pre = prefemur, st-pl = stigmatic plate, ta = tarsus, ti = tibia. Scale bars in millimetres." figureDoi="10.3897/zookeys.1212.126536.figure11" httpUri="https://binary.pensoft.net/fig/1133564">11 D L</figureCitation>
): All podomeres with long setae. Coxa (
<abbrev id="ABBRID0EUKAK" xlink_title="coxa">cx</abbrev>
) large, as long as length of prefemur; coxal process absent in first and second legs; legs 321 marginally with dentate process (tooth), conspicuous, broad. Prefemur (
<abbrev id="ABBRID0EYKAK" xlink_title="prefemur">pre</abbrev>
) stout; apico-mesally with weak projection; mesal margin with conspicuous and small spines. Femur (
<abbrev id="ABBRID0E3KAK" xlink_title="femur">fe</abbrev>
) short and stout, as long as length of prefemur; with a long ridge in all leg-pairs; apico-mesally with strong spines, forming a dentate margin. Postfemur (
<abbrev id="ABBRID0EALAK" xlink_title="postfemur">po</abbrev>
) and tibia (
<abbrev id="ABBRID0EELAK" xlink_title="tibia">ti</abbrev>
) quite short. Tarsus (
<abbrev id="ABBRID0EILAK" xlink_title="tarsus">ta</abbrev>
) of midbody legs quite short; as long as length of femur + postfemur; first two leg-pairs without an apical spine; leg-pair 1 with 1 ventral spine; leg-pair 2 with 3 ventral spines; leg-pair 3 with 46 ventral spines and 1 apical spine; leg pairs 4 with 68 ventral spines and 1 apical spine; leg pairs 521 with 79 ventral spines and 1 apical spine. In leg 9, length of femur equal to width, tarsus 4 × longer than wide. Claw normal, at base with a notch, conspicuous.
</paragraph>
<paragraph id="E0BCE32E738E95F320973FEAB92162A2">
<emphasis id="EFCB77E53FB82FF1CE50C41275BAC068" bold="true" italics="true">Male sexual characters</emphasis>
(Fig.
<figureCitation id="1ACC8AE218D0AFC8693CCD6910E78390" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Zephronia macula sp. nov. — SEM A G, L ♂ paratype (CUMZ - MYR 0016) J, K ♀ paratype (CUMZ - MYR 0015) A collum B seventh pleurite C subanal plate D legs 1 E stigmatic plate 1 F legs 2 with gonopores G leg 10 (arrow points to femoral ridge) H coxal process on leg 10 I small teeth on femur J leg 2 with vulvae K vulvae L tarsus 10. Abbreviation: bu = bursa, cx = coxa, fe = femur, op = operculum, po = postfemur, pre = prefemur, st-pl = stigmatic plate, ta = tarsus, ti = tibia. Scale bars in millimetres." figureDoi="10.3897/zookeys.1212.126536.figure11" httpUri="https://binary.pensoft.net/fig/1133564">11 F</figureCitation>
): Gonopore large, with long setae; with divided sclerotized plates, triangular.
</paragraph>
<paragraph id="9924B1CE7132F15A1A9A518C744BE782">
<emphasis id="BBB7E6C6311F8D8264CA27EC8852B6CE" bold="true" italics="true">Anterior telopods</emphasis>
(Fig.
<figureCitation id="310ADB82265DEF5C3EE1CF7A67DF390D" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Zephronia macula sp. nov., ♂ paratype (CUMZ - MYR 0016) — SEM A D overview of telopods (anterior view, lateral view with emphasis on anterior telopod, posterior view and ventral view, respectively) E G anterior telopod (telopoditomeres 24, movable finger, and crenulated teeth, respectively) H J posterior telopod (sclerotize spots on immovable finger, telopoditomeres 24 with emphasis on membranous lobe and crenulated teeth, respectively). Abbreviations: cr-t = crenulated teeth, ml = membranous lobe, scl-s = sclerotized spots. Scale bars in millimetres." figureDoi="10.3897/zookeys.1212.126536.figure12" httpUri="https://binary.pensoft.net/fig/1133565">12 A, B, E G</figureCitation>
): First telopoditomere stout, rectangular. Telopoditomere 2 large. Immovable finger (process of telopoditomere 2) quite long; as long as telopoditomere 3; clearly seen in posterior and anterior views; strongly curved; tip obtuse, directed anteriad and close to basal part of telopoditomere 4; with a large membranous lobe; with sclerotized spots located at inner margin. Telopoditomere 3 longer than telopoditomere 4; clearly demarcated from telopoditomere 4 by conspicuous suture; apically with crenulated teeth (
<abbrev id="ABBRID0ECMAK" xlink_title="crenulated teeth on telopods">cr-t</abbrev>
). Tepoloditomere 4 apically with a few setae located in the side pits.
</paragraph>
<caption id="3FAE11DE1B16130364EC36FCB4CD66CD" ID-DOI="10.3897/zookeys.1212.126536.figure12" ID-arpha="2FABAD41-FF7F-5CD5-ACF9-F0A65346C749" httpUri="https://binary.pensoft.net/fig/1133565" startId="F12">
<paragraph id="59C13137DE53DC4C0BF474B132E76038">
<label id="B2DD6F950E7AAFD26144354ABACE366E">Figure 12.</label>
</paragraph>
<paragraph id="12381644F272EF641231D75DA2413A88">
<taxonomicName id="12644F66BDB4A01F5429A7AB20BD02F7" authorityName="Srisonchai &amp; Wesener" authorityYear="2024" class="Diplopoda" family="Zephroniidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" phylum="Arthropoda" rank="species" species="macula" status="sp. nov.">
<emphasis id="5591CA7C56516FAB91CB89FBE7295926" italics="true">Zephronia macula</emphasis>
</taxonomicName>
<taxonomicNameLabel id="54CE9E4376F2D77EAFB1C94A0974BA7E" rank="species">sp. nov.</taxonomicNameLabel>
, ♂ paratype (
<named-content id="97D8E3D206816E6D377195ECF5C0FC13" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/chulalongkorn-university-museum-natural-history" xlink_title="Chulalongkorn University Museum of Zoology">CUMZ</named-content>
- MYR 0016) —
<abbrev id="ABBRID0E4MAK" xlink_title="scanning electron microscopy">SEM</abbrev>
<emphasis id="B7FFDB3AFFEE576D34484C27707057BA" bold="true">A D</emphasis>
overview of telopods (anterior view, lateral view with emphasis on anterior telopod, posterior view and ventral view, respectively)
<emphasis id="F3A5C57B3700046542BB404F54E9CCAA" bold="true">E G</emphasis>
anterior telopod (telopoditomeres 24, movable finger, and crenulated teeth, respectively)
<emphasis id="94595834EDEDC9917F91FD828E69EB00" bold="true">H J</emphasis>
posterior telopod (sclerotize spots on immovable finger, telopoditomeres 24 with emphasis on membranous lobe and crenulated teeth, respectively). Abbreviations:
<abbrev id="ABBRID0EHNAK" xlink_title="crenulated teeth on telopods">cr-t</abbrev>
= crenulated teeth,
<abbrev id="ABBRID0ELNAK" xlink_title="membranous lobe">ml</abbrev>
= membranous lobe,
<abbrev id="ABBRID0EPNAK" xlink_title="sclerotized spots">scl-s</abbrev>
= sclerotized spots. Scale bars in millimetres.
</paragraph>
</caption>
<paragraph id="CACE7662E23BD55EBA1A89DD8A3671DF">
<emphasis id="05F965A58E48BBF9BDAB997DA8663E25" bold="true" italics="true">Posterior telopods</emphasis>
(Figs
<figureCitation id="A45742E929BBAAD78D059BA1157A634A" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Zephronia macula sp. nov., ♂ paratype (CUMZ - MYR 0016) — SEM A D overview of telopods (anterior view, lateral view with emphasis on anterior telopod, posterior view and ventral view, respectively) E G anterior telopod (telopoditomeres 24, movable finger, and crenulated teeth, respectively) H J posterior telopod (sclerotize spots on immovable finger, telopoditomeres 24 with emphasis on membranous lobe and crenulated teeth, respectively). Abbreviations: cr-t = crenulated teeth, ml = membranous lobe, scl-s = sclerotized spots. Scale bars in millimetres." figureDoi="10.3897/zookeys.1212.126536.figure12" httpUri="https://binary.pensoft.net/fig/1133565">12 A D, H J</figureCitation>
;
<figureCitation id="CF372B9C11A546B6D8E0B4E30701BA2A" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Zephronia chantaburiensis sp. nov., ♂ paratype (ZFMK - MYR 13660), drawings — left posterior telopod A anterior view B posterior view. Abbreviations: cr-t = crenulated teeth, imf = immovable finger, ml = membranous lobe. Scale bar: 0.5 millimetres." figureDoi="10.3897/zookeys.1212.126536.figure13" httpUri="https://binary.pensoft.net/fig/1133566">13</figureCitation>
): With 4 telopoditomeres. First telopoditomere rectangular, stout; slightly longer than wide. Telopoditomere 2 large and stout. Immovable finger (process of telopoditomere 2) long and slender, 3 × longer than wide; equal in length to movable finger (= combination of telopoditomeres 3 + 4); attenuate near tip; tip obtuse,
<emphasis id="4C8109DD5A2F76EB0C56D7285FD74D69" italics="true">in-situ</emphasis>
curving anteriad; inner margin with conspicuous sclerotized spots (
<abbrev id="ABBRID0EJOAK" xlink_title="sclerotized spots">scl-s</abbrev>
) in a blackish ledge and two large membranous lobes (
<abbrev id="ABBRID0ENOAK" xlink_title="membranous lobe">ml</abbrev>
). Telopoditomere 3 long, 2.5 × longer than wide, inner margin with a long membranous ledge and basally with a single spine. Telopoditomere 4 quite short; 3.5 × shorter than telopoditomere 3; slightly tapering toward apex; with two spines located on membranous ledge at inner margin; with 9 or 10 large crenulated teeth (
<abbrev id="ABBRID0EROAK" xlink_title="crenulated teeth on telopods">cr-t</abbrev>
) posteriorly. Telopoditomeres 1 mostly glabrous, telopoditomere
<quantity id="4E0D3DA7454129776A0FA3DEC976FAF5" metricMagnitude="-2" metricUnit="m" metricValue="5.08" unit="in" value="2.0">2 in</quantity>
posterior view mostly glabrous, in anterior view covered by setae. Telopoditomeres 3 and
<quantity id="BE4D5A72883177686DC0E6F670DDB021" metricMagnitude="-1" metricUnit="m" metricValue="1.016" unit="in" value="4.0">4 in</quantity>
posterior view glabrous; at basal part of telopoditomere
<quantity id="87AD18CC952C1A1F5E85E4CD27EF2B5B" metricMagnitude="-2" metricUnit="m" metricValue="7.62" unit="in" value="3.0">3 in</quantity>
anterior view sparsely setose. Inner horns with sharp-edged tips, slightly curved caudad.
</paragraph>
<caption id="E53814DAEFAD47E2EB817DFB492B2F7B" ID-DOI="10.3897/zookeys.1212.126536.figure13" ID-arpha="4E363BE5-B720-5CFC-A2A2-BAB48BC04B51" httpUri="https://binary.pensoft.net/fig/1133566" startId="F13">
<paragraph id="07468DCE5343E886C83961FA2DAE62B6">
<label id="2A293818AFA5629E9211837A3F60F9EE">Figure 13.</label>
</paragraph>
<paragraph id="6C7274F23B40687E020B21563F0367ED">
<taxonomicName id="66FF7143839F536BCF1886BB20CAB314" authorityName="Srisonchai &amp; Wesener" authorityYear="2024" class="Diplopoda" family="Zephroniidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" phylum="Arthropoda" rank="species" species="chantaburiensis" status="sp. nov.">
<emphasis id="1D4CE9380C11B299721F2A0F401C14B3" italics="true">Zephronia chantaburiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="422330D5109EE7739F764E321E3C341A" rank="species">sp. nov.</taxonomicNameLabel>
, ♂ paratype (
<named-content id="4E9A2484AE22435AA1664BCD9ADBCCDD" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/zoologisches-forschungsmuseum-alexander-koenig" xlink_title="Zoological Research Museum Koenig">ZFMK</named-content>
- MYR 13660), drawings — left posterior telopod
<emphasis id="E4C1A9776CBA80198A5D19FB0C28D163" bold="true">A</emphasis>
anterior view
<emphasis id="A403C763BBEB428A8C481419CDB6E372" bold="true">B</emphasis>
posterior view. Abbreviations:
<abbrev id="ABBRID0EQPAK" xlink_title="crenulated teeth on telopods">cr-t</abbrev>
= crenulated teeth,
<abbrev id="ABBRID0EUPAK" xlink_title="immovable finger of telopod">imf</abbrev>
= immovable finger,
<abbrev id="ABBRID0EYPAK" xlink_title="membranous lobe">ml</abbrev>
= membranous lobe. Scale bar: 0.5 millimetres.
</paragraph>
</caption>
<caption id="4103FF296991B1134FD1F7D795B5A974" ID-DOI="10.3897/zookeys.1212.126536.figure14" ID-arpha="B7F341B7-5367-503A-8DCD-30A8984C1821" httpUri="https://binary.pensoft.net/fig/1133567" startId="F14">
<paragraph id="A9818D4BB205B730616C45AA1B00148A">
<label id="8BDDA1423D0D5D7903E6F03205E78EBC">Figure 14.</label>
</paragraph>
<paragraph id="85D56DF700527574CE39D25654942628">
Endoterga of thoracic shields and midbody tergites —
<abbrev id="ABBRID0EJQAK" xlink_title="scanning electron microscopy">SEM</abbrev>
<emphasis id="6B2D0C5DE16166EA46AD2E982B9E5DA7" bold="true">A D</emphasis>
<taxonomicName id="BE85BD8075D147AB5E2AD0F44775A842" authorityName="Srisonchai &amp; Wesener" authorityYear="2024" class="Diplopoda" family="Zephroniidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" phylum="Arthropoda" rank="species" species="chantaburiensis" status="sp. nov.">
<emphasis id="7759D0F65AC9559B94705CA1576E48FB" italics="true">Zephronia chantaburiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4E775F42AE2921AE417D0A727ACEBDB8" rank="species">sp. nov.</taxonomicNameLabel>
(♂ paratype,
<named-content id="80EB2EB50898A6DBF15DEB55159F64D0" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/chulalongkorn-university-museum-natural-history" xlink_title="Chulalongkorn University Museum of Zoology">CUMZ</named-content>
- MYR 0014)
<emphasis id="C331D34FCC2AEF80660008F3CF1EE436" bold="true">E H</emphasis>
<taxonomicName id="51266DCA85FB88D21E7C5373E95D005A" authorityName="Srisonchai &amp; Wesener" authorityYear="2024" class="Diplopoda" family="Zephroniidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" phylum="Arthropoda" rank="species" species="macula" status="sp. nov.">
<emphasis id="241714050F41E4785DF710B4E1810D46" italics="true">Zephronia macula</emphasis>
</taxonomicName>
<taxonomicNameLabel id="878E6F6DD70D1DF5FBDBC771349C5186" rank="species">sp. nov.</taxonomicNameLabel>
(♂ paratype,
<named-content id="01455BF618D3F7C415719F90ED5CD706" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/chulalongkorn-university-museum-natural-history" xlink_title="Chulalongkorn University Museum of Zoology">CUMZ</named-content>
- MYR 0016)
<emphasis id="02E6882AD0F3088E3F77EA566CFBE272" bold="true">A, B, E, F</emphasis>
thoracic shields
<emphasis id="447F7AC234B064D1CAFC3860172B0BC0" bold="true">C, D, G, H</emphasis>
midbody tergites. Abbreviations:
<abbrev id="ABBRID0ETRAK" xlink_title="cuticular impression of endotergum">cp</abbrev>
= cuticular impression,
<abbrev id="ABBRID0EXRAK" xlink_title="inner area of endotergum">ia</abbrev>
= inner area,
<abbrev id="ABBRID0E2RAK" xlink_title="middle area of endotergum">ma</abbrev>
= middle area, oa = outer area,
<abbrev id="ABBRID0E6RAK" xlink_title="posterior margin of endotergum">pm</abbrev>
= posterior margin. Scale bars in millimetres.
</paragraph>
</caption>
<caption id="AE79B2B5FCD70AF7671874FDFC06C92E" ID-DOI="10.3897/zookeys.1212.126536.figure15" ID-arpha="A3F4B178-D332-5CEA-A01B-8AE34FB759BA" httpUri="https://binary.pensoft.net/fig/1133568" startId="F15">
<paragraph id="55203BE588E7B434C6537DB6A6BED353">
<label id="15ED1BB21D0D8FC577AA784374122D97">Figure 15.</label>
</paragraph>
<paragraph id="45837C7F7DEF82CA47358912745D56D4">
Distribution and localities of
<taxonomicName id="18B1B80088A5F23B8AC9FF13F78627C8">
<emphasis id="B8B78BFFE874BC5ED545C742F002A815" italics="true">Zephronia</emphasis>
species
</taxonomicName>
in Thailand. Number inside the circle indicates species: 1 =
<taxonomicName id="41B5825F9914F3213CCEFBBFF1EB3374" authorityName="Srisonchai &amp; Wesener" authorityYear="2024" class="Diplopoda" family="Zephroniidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" phylum="Arthropoda" rank="species" species="chantaburiensis" status="sp. nov.">
<emphasis id="D4D7D3EDE3CF9561396D5192F78C9219" italics="true">Z. chantaburiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="43825F2417A2D0FCB9B3F8CF444CCCD8" rank="species">sp. nov.</taxonomicNameLabel>
; 2 =
<taxonomicName id="2550246AD689AFD62DAA6CA21C869229" authorityName="Bhansali &amp; Wesener" authorityYear="2022" class="Diplopoda" family="Zephroniidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" phylum="Arthropoda" rank="species" species="chrysomallos">
<emphasis id="BA82D02A4A55D33F1EE0908B1BE21260" italics="true">Z. chrysomallos</emphasis>
</taxonomicName>
; 3 =
<taxonomicName id="D28924C7D96D6F6A413C03634D9D5604" authorityName="Srisonchai &amp; Likhitrakarn" authorityYear="2021" class="Diplopoda" family="Zephroniidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" phylum="Arthropoda" rank="species" species="enghoffi">
<emphasis id="CFEA28D9DCF623B6DD3A65090F5FC8B2" italics="true">Z. enghoffi</emphasis>
</taxonomicName>
; 4 =
<taxonomicName id="6A69419178E3D98C1AE742FFA49383AB" authorityName="Bhansali &amp; Wesener" authorityYear="2022" class="Diplopoda" family="Zephroniidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" phylum="Arthropoda" rank="species" species="erawani">
<emphasis id="A41F5A8A07D9DFEAE135CA06610DD80D" italics="true">Z. erawani</emphasis>
</taxonomicName>
; 5 =
<taxonomicName id="0ACF5330C2B18FD0D593A87A8F7CA258" authorityName="Srisonchai, Sutcharit &amp; Likhitrakarn" authorityYear="2021" class="Diplopoda" family="Zephroniidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" phylum="Arthropoda" rank="species" species="golovatchi">
<emphasis id="1E9993A177018CF8FF74C01CE47D476B" italics="true">Z. golovatchi</emphasis>
</taxonomicName>
; 6 =
<taxonomicName id="A14F2B7F593306E5B532B220FF5D0C96" authorityName="Likhitrakarn &amp; Golovatch" authorityYear="2021" class="Diplopoda" family="Zephroniidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" phylum="Arthropoda" rank="species" species="lannaensis">
<emphasis id="B47F07B41E13A49A3BFAA318E94E50DF" italics="true">Z. lannaensis</emphasis>
</taxonomicName>
; 7 =
<taxonomicName id="6263C1CD3D4568C9C244E41812CB8F6A" authorityName="Srisonchai &amp; Wesener" authorityYear="2024" class="Diplopoda" family="Zephroniidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" phylum="Arthropoda" rank="species" species="macula" status="sp. nov.">
<emphasis id="905C54B1636948AF2ECDAB8B2401AEB1" italics="true">Z. macula</emphasis>
</taxonomicName>
<taxonomicNameLabel id="60F90430C3565EF9F72E918C94DB1E14" rank="species">sp. nov.</taxonomicNameLabel>
; 8 =
<taxonomicName id="13FB4A00CE27AD16493AF1EF8B83E1E3" authorityName="Srisonchai, Sutcharit &amp; Likhitrakarn" authorityYear="2021" class="Diplopoda" family="Zephroniidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" phylum="Arthropoda" rank="species" species="panhai">
<emphasis id="57293436A2D1732C732E356DA6088C22" italics="true">Z. panhai</emphasis>
</taxonomicName>
; 9 =
<taxonomicName id="080578796D358049A599E4C6014D4B82" authorityName="Likhitrakarn &amp; Golovatch" authorityYear="2021" class="Diplopoda" family="Zephroniidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" phylum="Arthropoda" rank="species" species="phrain">
<emphasis id="71D35CBE41AEA63B99E6D788AAE5E481" italics="true">Z. phrain</emphasis>
</taxonomicName>
; 10 =
<taxonomicName id="8AE0B41A299F9DA4C5A30656A02A342E" authorityName="Hirst" authorityYear="1907" class="Diplopoda" family="Zephroniidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" phylum="Arthropoda" rank="species" species="siamensis">
<emphasis id="B866C576ADE0881208E6521DE2B954E0" italics="true">Z. siamensis</emphasis>
</taxonomicName>
; 11 =
<taxonomicName id="6179076E14ACB75D3BD448AD96E46DB6" authorityName="Rosenmejer &amp; Wesener" authorityYear="2021" class="Diplopoda" family="Zephroniidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" phylum="Arthropoda" rank="species" species="viridisoma">
<emphasis id="3E0C0F2252AEFDB16A08C1F4E69D2945" italics="true">Z. viridisoma</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="5D322C79A389565CAF3DC54DB98F50CA">
<emphasis id="64D53B684A270DDE5FF37D66281CC7FA" bold="true" italics="true">Female sexual characters</emphasis>
(Fig.
<figureCitation id="6129DD77C710F4EAFC90380550579827" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Zephronia macula sp. nov. — SEM A G, L ♂ paratype (CUMZ - MYR 0016) J, K ♀ paratype (CUMZ - MYR 0015) A collum B seventh pleurite C subanal plate D legs 1 E stigmatic plate 1 F legs 2 with gonopores G leg 10 (arrow points to femoral ridge) H coxal process on leg 10 I small teeth on femur J leg 2 with vulvae K vulvae L tarsus 10. Abbreviation: bu = bursa, cx = coxa, fe = femur, op = operculum, po = postfemur, pre = prefemur, st-pl = stigmatic plate, ta = tarsus, ti = tibia. Scale bars in millimetres." figureDoi="10.3897/zookeys.1212.126536.figure11" httpUri="https://binary.pensoft.net/fig/1133564">11 J, K</figureCitation>
): Vulva large and stout, sparsely setose; covering almost 1 / 3 of coxa; located at mesal margin; mesally extended to base of prefemur. Operculum (
<abbrev id="ABBRID0EAXAK" xlink_title="operculum of vulva">op</abbrev>
) broad and well-rounded; swollen dorsoventrally, mesal margin not protruding; tip of operculum truncate. Bursa (
<abbrev id="ABBRID0EEXAK" xlink_title="bursa of vulva">bu</abbrev>
) large, completely demarcated from operculum by a triangular groove.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EIXAK" type="remarks">
<paragraph id="1B4DC66E07BDA522E8822E8F4BC76E8F">
<heading id="209B877F27756986911980EE1D9CBB38" reason="title">Remarks.</heading>
</paragraph>
<paragraph id="E24DCF871E10E9B9C17690BFCD818AF4">Intrapopulational variation was found in which most specimens have a brown / reddish brown (majority) colour, while the others are greenish brown (minority). In addition, the posterior margin of subanal scale is in most specimens wide, whereas some specimens possess a quite narrow margin.</paragraph>
</subSubSection>
<subSubSection id="SECID0ENXAK" type="distribution">
<paragraph id="D576ECCDA3F425CAAA49509E3364464D">
<heading id="74B6B971545679FFAAA4847B8A114550" reason="title">Distribution and habitat.</heading>
</paragraph>
<paragraph id="B47D9BC966AB6F66788B696804AA0F24">
This species is widely distributed across eastern
<collectingCountry id="72155D6A345B75E173D3381211480249" name="Thailand">Thailand</collectingCountry>
(&gt;
<quantity id="FB69BAD5F166BCD91FB9BB64CB31311C" metricMagnitude="5" metricUnit="m" metricValue="1.5" unit="km" value="150.0">150 km</quantity>
). Most specimens were collected from locations with granitic rock habitats, while a few localities were in forested limestone habitats (Fig.
<figureCitation id="DBC96D08BCF77736EB1260B969E3AE0B" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Photograph of live specimens of Zephronia macula sp. nov. and habitats A G paratypes (CUMZ - MYR 00016) H habitat at Thaeo Khlong Khlung monastery (granitic rocks) I habitat at Phet Pho Thong cave (limestone). Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1212.126536.figure8" httpUri="https://binary.pensoft.net/fig/1133561">8 H, I</figureCitation>
). All individuals were encountered during the day under rotten logs and in leaf litters.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EXXAK" type="etymology">
<paragraph id="0E4CF566F015361D8E02EF241A28093F">
<heading id="CB27AC9FE2D0187CA27B2B4E0D82170B" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="FFA0BBDFBBA5BD76257CBE9DB56277A2">The name is a noun referring to the pattern of spots on the body.</paragraph>
</subSubSection>
</treatment>
</document>