661 lines
100 KiB
XML
661 lines
100 KiB
XML
<document id="A618FD3FBA43855024F75657A82267EB" ID-CLB-Dataset="29540" ID-DOI="10.11646/zootaxa.4486.3.2" ID-GBIF-Dataset="30a406db-512c-4ef5-9291-b38ffc0aaefa" ID-ISSN="1175-5326" ID-Zenodo-Dep="1437066" ID-ZooBank="4A7504C8-0CB9-4C1B-BADC-F4FBDAF15F1B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1538115691119" checkinUser="plazi" docAuthor="Kieckbusch, Max, Mader, Felix, Kaiser, Hinrich & Mecke, Sven" docDate="2018" docId="F60787D8D0428C579CB6FBC8E2B0B38E" docLanguage="en" docName="zootaxa.4486.3.2.pdf" docOrigin="Zootaxa 4486 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cylindrophis osheai Kieckbusch & Mader & Kaiser & Mecke 2018, sp. nov." docType="treatment" docVersion="7" lastPageNumber="244" masterDocId="0A3EFFA0D0438C5F9C21FFE6E674B223" masterDocTitle="A new species of Cylindrophis Wagler, 1828 (Reptilia: Squamata: Cylindrophiidae) from Boano Island, northern Maluku Province, Indonesia" masterLastPageNumber="250" masterPageNumber="236" pageNumber="237" updateTime="1698554762433" updateUser="ExternalLinkService">
|
||
<mods:mods id="A97C7B8F3678C762241647EEFB648672" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="867FC0FF1DAED1A3CCEB78C10EE4C52F">
|
||
<mods:title id="1EC2EDA567AF195933B74F4A4A1F8FBF">A new species of Cylindrophis Wagler, 1828 (Reptilia: Squamata: Cylindrophiidae) from Boano Island, northern Maluku Province, Indonesia</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="99BB42E3D80B479876E68BF0A0596CE2" type="personal">
|
||
<mods:role id="6B2BD80A71A30B0037FDB4FF65564F3E">
|
||
<mods:roleTerm id="72F5F16BA88CA6775B6D381B1141CBBF">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="BD2FA779CC41C82FDACEE6E2D353C61F">Kieckbusch, Max</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="9C9CDDCDA95670CF9C54AAC26C15E477" type="personal">
|
||
<mods:role id="34FEE4414F2E368B11830A4D4958A2E1">
|
||
<mods:roleTerm id="0DDF7E7C3B6AFC7FD6736089473B307B">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="F1CB7893EA0E4AAAAA2174F3CAD95767">Mader, Felix</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="31156199EE95CF3DDFA3794D24829190" type="personal">
|
||
<mods:role id="5A9A7F4B47673640D74650624886144C">
|
||
<mods:roleTerm id="A0289039927D70A425DB6753F6FF0D28">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="67797C062713B84FCFC759255C2D827A">Kaiser, Hinrich</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="FBE1A3A2DDBE1AC464900A980C364A7A" type="personal">
|
||
<mods:role id="F02C1DAF7D47185AD707EA86FE725407">
|
||
<mods:roleTerm id="8337CD5A581BF23CA542894CD5960C60">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="4556EC63A332959C42FFC7FE53540559">Mecke, Sven</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="4F89532574C5AED0C03C766E4F501FFC">text</mods:typeOfResource>
|
||
<mods:relatedItem id="990C83C4262A534554F9553FEA896784" type="host">
|
||
<mods:titleInfo id="5AD23FE709B4DD86D28D178C017C791F">
|
||
<mods:title id="FBE43AA102700AB7F9FF2DE610B4CCF9">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="B4CB4A619115254A5E33DAA5D891756E">
|
||
<mods:date id="F36F5915568A0CCDC8034DD40E64F16D">2018</mods:date>
|
||
<mods:detail id="8C0A32DEE1CD371409764D35523A7F59" type="pubDate">
|
||
<mods:number id="EF323DF8DC2C5B20CCB63B2571F2F60E">2018-09-28</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="F00918B94C0C528C327D255DBB7389C5" type="volume">
|
||
<mods:number id="210B14C2B3BAE2540313E2D1C1280AF7">4486</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="712988D046FA8841CC132E357D031D8A" type="issue">
|
||
<mods:number id="7E0AFCC74F5A1E65849E18E92FC79EE5">3</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="03AB45D61C73225138A22842B347F1A9" unit="page">
|
||
<mods:start id="ECC4C2E18A9FB1FFCB5FED8537DA2E38">236</mods:start>
|
||
<mods:end id="CA960B893FD9DCA41861EF098E4C474A">250</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="E6158E763BC0B9D19982474630DF0A2F">journal article</mods:classification>
|
||
<mods:identifier id="AF92CDCC3CF9D5A53FB674D1374A8283" type="CLB-Dataset">29540</mods:identifier>
|
||
<mods:identifier id="0D5522E4CB6BE4ED5BB7748A42C1B173" type="DOI">10.11646/zootaxa.4486.3.2</mods:identifier>
|
||
<mods:identifier id="ECC134AB6C08607E4ACD657D3864B8DB" type="GBIF-Dataset">30a406db-512c-4ef5-9291-b38ffc0aaefa</mods:identifier>
|
||
<mods:identifier id="85089655A0299B221B46A43951AF92D8" type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier id="ED7C5638D7AD21DB83638196A3C62E95" type="Zenodo-Dep">1437066</mods:identifier>
|
||
<mods:identifier id="04FA9EF46A56BA6654CA773FBFB95370" type="ZooBank">4A7504C8-0CB9-4C1B-BADC-F4FBDAF15F1B</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="F60787D8D0428C579CB6FBC8E2B0B38E" ID-DOI="http://doi.org/10.5281/zenodo.5984701" ID-GBIF-Taxon="148403148" ID-Zenodo-Dep="5984701" LSID="urn:lsid:plazi:treatment:F60787D8D0428C579CB6FBC8E2B0B38E" httpUri="http://treatment.plazi.org/id/F60787D8D0428C579CB6FBC8E2B0B38E" lastPageId="8" lastPageNumber="244" pageId="1" pageNumber="237">
|
||
<subSubSection id="36B46545D0428C5E9CB6FBC8E760B649" pageId="1" pageNumber="237" type="nomenclature">
|
||
<paragraph id="7E1136CED0428C5E9CB6FBC8E799B66B" blockId="1.[151,493,1070,1130]" box="[151,493,1070,1096]" pageId="1" pageNumber="237">
|
||
<heading id="255981A2D0428C5E9CB6FBC8E799B66B" bold="true" box="[151,493,1070,1096]" fontSize="11" level="1" pageId="1" pageNumber="237" reason="1">
|
||
<emphasis id="4CDAEADCD0428C5E9CB6FBC8E799B66B" bold="true" box="[151,493,1070,1096]" pageId="1" pageNumber="237">
|
||
<taxonomicName id="B9AE4D4DD0428C5E9CB6FBC8E7FDB66B" authority="Kieckbusch & Mader & Kaiser & Mecke, 2018" authorityName="Kieckbusch & Mader & Kaiser & Mecke" authorityYear="2018" box="[151,393,1070,1096]" class="Reptilia" family="Cylindrophiidae" genus="Cylindrophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="237" phylum="Chordata" rank="species" species="osheai" status="sp. nov.">
|
||
<emphasis id="4CDAEADCD0428C5E9CB6FBC8E7FDB66B" bold="true" box="[151,393,1070,1096]" italics="true" pageId="1" pageNumber="237">Cylindrophis osheai</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="57E957A7D0428C5E9DB1FBC8E799B66B" box="[400,493,1070,1096]" pageId="1" pageNumber="237" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="7E1136CED0428C5E9CB6FBB7E760B649" blockId="1.[151,493,1070,1130]" box="[151,276,1105,1130]" pageId="1" pageNumber="237">
|
||
(
|
||
<figureCitation id="E6952A4BD0428C5E9CBFFBB7E69EB649" box="[158,234,1105,1130]" captionStart="FIGURE 1" captionStartId="2.[151,250,1867,1889]" captionTargetBox="[173,1405,249,1807]" captionTargetId="figure@2.[151,1436,210,1838]" captionTargetPageId="2" captionText="FIGURE 1. Known specimens of Cylindrophis osheai sp. nov. in dorsal (left) and ventral view (right). (A) Holotype, RMNH 5460 (formerly RMNH 5460.70), an adult female, collected in 1865 by Dirk Samuel Hoedt on Boano Island, Central Maluku Regency, Maluku Province, Indonesia. (B) Paratype, RMNH 47930 (formerly RMNH 5460.69), a subadult male, collection data as for holotype. (C) RMNH 11262, an unsexed specimen, labeled with collection locality “Celebes?” [= Sulawesi, in error]. Scale bars = 10 mm. Photographs by Sven Mecke." httpUri="https://zenodo.org/record/1437068/files/figure.png" pageId="1" pageNumber="237">FigS. 1</figureCitation>
|
||
¯5)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="36B46545D0428C5E9CB6FB7FE50EB5D2" pageId="1" pageNumber="237" type="materials_examined">
|
||
<paragraph id="7E1136CED0428C5E9CB6FB7FE7AFB6D9" blockId="1.[151,1437,1177,2033]" pageId="1" pageNumber="237">
|
||
<emphasis id="4CDAEADCD0428C5E9CB6FB7FE77FB691" bold="true" box="[151,267,1177,1202]" pageId="1" pageNumber="237">
|
||
<typeStatus id="A115886CD0428C5E9CB6FB7FE772B691" box="[151,262,1177,1202]" pageId="1" pageNumber="237" type="holotype">Holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
<collectionCode id="18BFAE0BD0428C5E9D33FB7CE712B692" box="[274,358,1178,1201]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="1" pageNumber="237">RMNH</collectionCode>
|
||
5460 (formerly
|
||
<collectionCode id="18BFAE0BD0428C5E9E3FFB7CE406B692" box="[542,626,1178,1201]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="1" pageNumber="237">RMNH</collectionCode>
|
||
5460.70) (
|
||
<figureCitation id="E6952A4BD0428C5E9ECDFB7FE524B691" box="[748,848,1177,1202]" captionStart="FIGURE 1" captionStartId="2.[151,250,1867,1889]" captionTargetBox="[173,1405,249,1807]" captionTargetId="figure@2.[151,1436,210,1838]" captionTargetPageId="2" captionText="FIGURE 1. Known specimens of Cylindrophis osheai sp. nov. in dorsal (left) and ventral view (right). (A) Holotype, RMNH 5460 (formerly RMNH 5460.70), an adult female, collected in 1865 by Dirk Samuel Hoedt on Boano Island, Central Maluku Regency, Maluku Province, Indonesia. (B) Paratype, RMNH 47930 (formerly RMNH 5460.69), a subadult male, collection data as for holotype. (C) RMNH 11262, an unsexed specimen, labeled with collection locality “Celebes?” [= Sulawesi, in error]. Scale bars = 10 mm. Photographs by Sven Mecke." httpUri="https://zenodo.org/record/1437068/files/figure.png" pageId="1" pageNumber="237">FigS. 1A</figureCitation>
|
||
,
|
||
<figureCitation id="E6952A4BD0428C5E9F7DFB7CE518B691" box="[860,876,1178,1202]" captionStart="FIGURE 2" captionStartId="3.[151,250,818,840]" captionTargetBox="[156,1432,191,797]" captionTargetId="figure@3.[151,1436,180,807]" captionTargetPageId="3" captionText="FIGURE 2. Holotype of Cylindrophis osheai sp. nov. (RMNH 5460). Head in (A) dorsal, (B) lateral, and (C) ventral view; (D) tail in ventral view showing subcaudals. Scale bar = 40 mm. Drawings by Felix Mader based on photographs by Sven Mecke." httpUri="https://zenodo.org/record/1437070/files/figure.png" pageId="1" pageNumber="237">2</figureCitation>
|
||
,
|
||
<figureCitation id="E6952A4BD0428C5E9F59FB7CE5E8B691" box="[888,924,1178,1202]" captionStart="FIGURE 3" captionStartId="4.[151,250,646,668]" captionTargetBox="[153,1416,196,625]" captionTargetId="figure@4.[151,1436,193,629]" captionTargetPageId="4" captionText="FIGURE 3. Details of the ventral coloration in six species of Cylindrophis. (A) Cylindrophis osheai sp. nov. (RMNH 5460). (B) C. melanotus (RMNH 11451.114). (C) C. lineatus (AMNH 12872). (D) C. ruffus sensu stricto (RMNH 11256.110). (E) C. subocularis (RMNH 8959.57). (F) C. jodiae (NMW 21562.1). Drawings by Felix Mader based on photographs by Sven Mecke." httpUri="https://zenodo.org/record/1437072/files/figure.png" pageId="1" pageNumber="237">3A</figureCitation>
|
||
,
|
||
<figureCitation id="E6952A4BD0428C5E9F89FB7CE5CCB691" box="[936,952,1178,1202]" captionStart="FIGURE 4" captionStartId="6.[151,250,1857,1879]" captionTargetBox="[232,1359,193,1847]" captionTargetId="figure@6.[232,1359,193,1847]" captionTargetPageId="6" captionText="FIGURE 4. Micro-CT images of the cranium of the holotype of Cylindrophis osheai sp. nov. (RMNH 5460), including (A) dorsal, (B) lateral and mirrored, (C) and ventral view, as well as (D) a dorsal view of the lower jaw. Abbreviations of cranial elements follow Lee & Scanlon (2002) and include: bo = basioccipital, com = compound bone of mandible, den = dentary, ec = ectopterygoid, eo = exoccipitals, fr = frontal, mx = maxilla, na = nasal, pa = parietal, pbs = parabasisphenoid, pof = postorbitofrontal, prf = prefrontal, pro = prootics, pt = pterygoid, qa = quadrate, sm = septomaxilla, so = supraoccipital, and st = supratemporal. Scale bar = 2 mm. Micro-CT images prepared by Kristin Mahlow." httpUri="https://zenodo.org/record/1437074/files/figure.png" pageId="1" pageNumber="237">4</figureCitation>
|
||
,
|
||
<figureCitation id="E6952A4BD0428C5E9FE5FB7FE59DB691" box="[964,1001,1177,1202]" captionStart="FIGURE 5" captionStartId="7.[151,250,1221,1243]" captionTargetBox="[151,1436,193,1200]" captionTargetId="figure@7.[151,1436,193,1200]" captionTargetPageId="7" captionText="FIGURE 5. Micro-CT images of skulls of several species of Cylindrophis used for comparison of cranial osteology, shown in dorsal (left column), lateral (middle column), and ventral view (right column). (A) Cylindrophis osheai sp. nov. (RMNH 5460). (B) C. boulengeri (NHMB 12913). (C) C. melanotus (ZMB 4049). (D) C. ruffus sensu lato (ZMB 13129). Scale bars = 2 mm. Micro-CT images prepared by Kristin Mahlow." httpUri="https://zenodo.org/record/1437076/files/figure.png" pageId="1" pageNumber="237">5A</figureCitation>
|
||
), an adult female, collected in 1865 by natural hiStory collector Dirk Samuel Hoedt (
|
||
<date id="0A10100ED0428C5E9E89FB5BE55EB6F5" box="[680,810,1213,1238]" pageId="1" pageNumber="237" value="1815" valueMax="1893">1815–1893</date>
|
||
) on Boano ISland, Central
|
||
<collectingRegion id="BC6AF82CD0428C5E984BFB5BE2B5B6F5" box="[1130,1217,1213,1238]" country="Indonesia" name="Maluku Utara" pageId="1" pageNumber="237">Maluku</collectingRegion>
|
||
Regency, northern
|
||
<collectingRegion id="BC6AF82CD0428C5E9CB6FB07E72FB6D9" box="[151,347,1249,1274]" country="Indonesia" name="Maluku Utara" pageId="1" pageNumber="237">Maluku Province</collectingRegion>
|
||
,
|
||
<collectingCountry id="06B9765ED0428C5E9D47FB07E7A3B6D9" box="[358,471,1249,1274]" name="Indonesia" pageId="1" pageNumber="237">IndoneSia</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="7E1136CED0428C5E9CE6FAE3E774B761" blockId="1.[151,1437,1177,2033]" pageId="1" pageNumber="237">
|
||
<emphasis id="4CDAEADCD0428C5E9CE6FAE3E74FB73D" bold="true" box="[199,315,1285,1310]" pageId="1" pageNumber="237">
|
||
<typeStatus id="A115886CD0428C5E9CE6FAE3E742B73D" box="[199,310,1285,1310]" pageId="1" pageNumber="237" type="paratype">Paratype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
<collectionCode id="18BFAE0BD0428C5E9D65FAE0E7ECB73E" box="[324,408,1286,1309]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="1" pageNumber="237">RMNH</collectionCode>
|
||
47930 (formerly
|
||
<collectionCode id="18BFAE0BD0428C5E9E46FAE0E4CFB73E" box="[615,699,1286,1309]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="1" pageNumber="237">RMNH</collectionCode>
|
||
5460.69) (
|
||
<figureCitation id="E6952A4BD0428C5E9F1DFAE3E5E0B73D" box="[828,916,1285,1310]" captionStart="FIGURE 1" captionStartId="2.[151,250,1867,1889]" captionTargetBox="[173,1405,249,1807]" captionTargetId="figure@2.[151,1436,210,1838]" captionTargetPageId="2" captionText="FIGURE 1. Known specimens of Cylindrophis osheai sp. nov. in dorsal (left) and ventral view (right). (A) Holotype, RMNH 5460 (formerly RMNH 5460.70), an adult female, collected in 1865 by Dirk Samuel Hoedt on Boano Island, Central Maluku Regency, Maluku Province, Indonesia. (B) Paratype, RMNH 47930 (formerly RMNH 5460.69), a subadult male, collection data as for holotype. (C) RMNH 11262, an unsexed specimen, labeled with collection locality “Celebes?” [= Sulawesi, in error]. Scale bars = 10 mm. Photographs by Sven Mecke." httpUri="https://zenodo.org/record/1437068/files/figure.png" pageId="1" pageNumber="237">Fig. 1B</figureCitation>
|
||
), a Subadult male with collection data aS for
|
||
<typeStatus id="A115886CD0428C5E9CB6FACFE689B761" box="[151,253,1321,1346]" pageId="1" pageNumber="237" type="holotype">holotype</typeStatus>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="7E1136CED0428C5E9CE6FAABE20FB78D" blockId="1.[151,1437,1177,2033]" pageId="1" pageNumber="237">
|
||
The original entry in the collection catalogue of the herpetological collection at RMNH liStS both SpecimenS (under the regiStration number 5460) aS “
|
||
<taxonomicName id="B9AE4D4DD0428C5E9EA4FA97E528B7A9" authorityName="Stoliczka" authorityYear="1870" box="[645,860,1393,1418]" class="Reptilia" family="Cylindrophiidae" genus="Cylindrophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="237" phylum="Chordata" rank="species" species="rufus">
|
||
<emphasis id="4CDAEADCD0428C5E9EA4FA97E528B7A9" box="[645,860,1393,1418]" italics="true" pageId="1" pageNumber="237">Cylindrophis rufus</emphasis>
|
||
</taxonomicName>
|
||
” and indicateS that they were later identified aS “
|
||
<taxonomicName id="B9AE4D4DD0428C5E9C82FA73E7DBB78D" authorityName="Wagler" authorityYear="1828" box="[163,431,1429,1454]" class="Reptilia" family="Cylindrophiidae" genus="Cylindrophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="237" phylum="Chordata" rank="species" species="melanotus">
|
||
<emphasis id="4CDAEADCD0428C5E9C82FA73E7DBB78D" box="[163,431,1429,1454]" italics="true" pageId="1" pageNumber="237">Cylindrophis melanotus</emphasis>
|
||
</taxonomicName>
|
||
var.” by Dutch zoologiSt Leo Daniël BrongerSma (
|
||
<date id="0A10100ED0428C5E9FCBFA70E21BB78D" box="[1002,1135,1430,1454]" pageId="1" pageNumber="237" value="1907" valueMax="1994">1907–1994</date>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="7E1136CED0428C5E9CE6FA5FE5D6B7D5" blockId="1.[151,1437,1177,2033]" pageId="1" pageNumber="237">
|
||
<emphasis id="4CDAEADCD0428C5E9CE6FA5FE7C0B7F1" bold="true" box="[199,436,1465,1490]" pageId="1" pageNumber="237">Referred specimen.</emphasis>
|
||
<collectionCode id="18BFAE0BD0428C5E9DE0FA5CE461B7F2" box="[449,533,1466,1489]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="1" pageNumber="237">RMNH</collectionCode>
|
||
11262 (
|
||
<figureCitation id="E6952A4BD0428C5E9E5DFA5FE4ACB7F1" box="[636,728,1465,1490]" captionStart="FIGURE 1" captionStartId="2.[151,250,1867,1889]" captionTargetBox="[173,1405,249,1807]" captionTargetId="figure@2.[151,1436,210,1838]" captionTargetPageId="2" captionText="FIGURE 1. Known specimens of Cylindrophis osheai sp. nov. in dorsal (left) and ventral view (right). (A) Holotype, RMNH 5460 (formerly RMNH 5460.70), an adult female, collected in 1865 by Dirk Samuel Hoedt on Boano Island, Central Maluku Regency, Maluku Province, Indonesia. (B) Paratype, RMNH 47930 (formerly RMNH 5460.69), a subadult male, collection data as for holotype. (C) RMNH 11262, an unsexed specimen, labeled with collection locality “Celebes?” [= Sulawesi, in error]. Scale bars = 10 mm. Photographs by Sven Mecke." httpUri="https://zenodo.org/record/1437068/files/figure.png" pageId="1" pageNumber="237">Fig. 1C</figureCitation>
|
||
), an unSexed Specimen, labeled with the collection locality “
|
||
<collectingRegion id="BC6AF82CD0428C5E9C80FA3BE775B7D5" box="[161,257,1501,1526]" country="Indonesia" name="Sulawesi Utara" pageId="1" pageNumber="237">CelebeS</collectingRegion>
|
||
?” (=
|
||
<collectingRegion id="BC6AF82CD0428C5E9D1DFA3BE7D0B7D5" box="[316,420,1501,1526]" country="Indonesia" name="Sulawesi Utara" pageId="1" pageNumber="237">SulaweSi</collectingRegion>
|
||
, in error); no further information iS available.
|
||
</paragraph>
|
||
<paragraph id="7E1136CED0428C5E9CE6F9E7E54EB4A5" blockId="1.[151,1437,1177,2033]" pageId="1" pageNumber="237">
|
||
Both the label on the jar and the original entry in the collection catalogue of the herpetological collection at
|
||
<collectionCode id="18BFAE0BD0428C5E9CB6F9C0E69FB41E" box="[151,235,1574,1597]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="1" pageNumber="237">RMNH</collectionCode>
|
||
were written by E.M.J. JaSparS, who waS a Student of Leo Daniël BrongerSma working on SnakeS in the 1960S, long after the original Specimen had been collected (MarinuS Hoogmoed, in litt.; See diStribution Section for further commentS on the collection locality of thiS voucher).
|
||
</paragraph>
|
||
<paragraph id="7E1136CED0428C5E9CE6F977E50EB5D2" blockId="1.[151,1437,1177,2033]" pageId="1" pageNumber="237">
|
||
<emphasis id="4CDAEADCD0428C5E9CE6F977E732B489" bold="true" box="[199,326,1681,1706]" pageId="1" pageNumber="237">Definition.</emphasis>
|
||
A SpecieS of the genuS
|
||
<taxonomicName id="B9AE4D4DD0428C5E9E43F977E487B489" authorityName="Wagler" authorityYear="1828" box="[610,755,1681,1706]" class="Reptilia" family="Cylindrophiidae" genus="Cylindrophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="237" phylum="Chordata" rank="genus">
|
||
<emphasis id="4CDAEADCD0428C5E9E43F977E487B489" box="[610,755,1681,1706]" italics="true" pageId="1" pageNumber="237">Cylindrophis</emphasis>
|
||
</taxonomicName>
|
||
that can be eaSily diStinguiShed from all congenerS by the following combination of eidonomic characters: (1) 19 dorsal scale rows at midbody; (2) 224¯226 ventral scales; (3) 8¯9 subcaudal scales; (4) 25¯28 dark ventral blotches, aligned to form a broad, wavy stripe along most of body (
|
||
<figureCitation id="E6952A4BD0428C5E9CBEF91BE68EB535" box="[159,250,1789,1814]" captionStart="FIGURE 3" captionStartId="4.[151,250,646,668]" captionTargetBox="[153,1416,196,625]" captionTargetId="figure@4.[151,1436,193,629]" captionTargetPageId="4" captionText="FIGURE 3. Details of the ventral coloration in six species of Cylindrophis. (A) Cylindrophis osheai sp. nov. (RMNH 5460). (B) C. melanotus (RMNH 11451.114). (C) C. lineatus (AMNH 12872). (D) C. ruffus sensu stricto (RMNH 11256.110). (E) C. subocularis (RMNH 8959.57). (F) C. jodiae (NMW 21562.1). Drawings by Felix Mader based on photographs by Sven Mecke." httpUri="https://zenodo.org/record/1437072/files/figure.png" pageId="1" pageNumber="237">Fig. 3A</figureCitation>
|
||
); (5) a dark ventral pattern completely Separated from the dark dorSal coloration; (6) tail with an almoSt completely dark-colored ventral Surface (
|
||
<figureCitation id="E6952A4BD0428C5E9E54F8C2E4A7B51F" box="[629,723,1828,1853]" captionStart="FIGURE 2" captionStartId="3.[151,250,818,840]" captionTargetBox="[156,1432,191,797]" captionTargetId="figure@3.[151,1436,180,807]" captionTargetPageId="3" captionText="FIGURE 2. Holotype of Cylindrophis osheai sp. nov. (RMNH 5460). Head in (A) dorsal, (B) lateral, and (C) ventral view; (D) tail in ventral view showing subcaudals. Scale bar = 40 mm. Drawings by Felix Mader based on photographs by Sven Mecke." httpUri="https://zenodo.org/record/1437070/files/figure.png" pageId="1" pageNumber="237">Fig. 2D</figureCitation>
|
||
); (7) a dark bar running from the eye through the 3rd and 4th SupralabialS to the mouth (
|
||
<figureCitation id="E6952A4BD0428C5E9DE1F8AEE46CB543" box="[448,536,1864,1889]" captionStart="FIGURE 2" captionStartId="3.[151,250,818,840]" captionTargetBox="[156,1432,191,797]" captionTargetId="figure@3.[151,1436,180,807]" captionTargetPageId="3" captionText="FIGURE 2. Holotype of Cylindrophis osheai sp. nov. (RMNH 5460). Head in (A) dorsal, (B) lateral, and (C) ventral view; (D) tail in ventral view showing subcaudals. Scale bar = 40 mm. Drawings by Felix Mader based on photographs by Sven Mecke." httpUri="https://zenodo.org/record/1437070/files/figure.png" pageId="1" pageNumber="237">Fig. 2B</figureCitation>
|
||
); and (8) a dark, double-diamond-Shaped ventral pattern in the area immediately poSterior to the genialS (
|
||
<figureCitation id="E6952A4BD0428C5E9D82F88AE473B5A7" box="[419,519,1900,1925]" captionStart="FIGURE 1" captionStartId="2.[151,250,1867,1889]" captionTargetBox="[173,1405,249,1807]" captionTargetId="figure@2.[151,1436,210,1838]" captionTargetPageId="2" captionText="FIGURE 1. Known specimens of Cylindrophis osheai sp. nov. in dorsal (left) and ventral view (right). (A) Holotype, RMNH 5460 (formerly RMNH 5460.70), an adult female, collected in 1865 by Dirk Samuel Hoedt on Boano Island, Central Maluku Regency, Maluku Province, Indonesia. (B) Paratype, RMNH 47930 (formerly RMNH 5460.69), a subadult male, collection data as for holotype. (C) RMNH 11262, an unsexed specimen, labeled with collection locality “Celebes?” [= Sulawesi, in error]. Scale bars = 10 mm. Photographs by Sven Mecke." httpUri="https://zenodo.org/record/1437068/files/figure.png" pageId="1" pageNumber="237">FigS. 1A</figureCitation>
|
||
,
|
||
<figureCitation id="E6952A4BD0428C5E9E33F88AE443B5A7" box="[530,567,1900,1924]" captionStart="FIGURE 2" captionStartId="3.[151,250,818,840]" captionTargetBox="[156,1432,191,797]" captionTargetId="figure@3.[151,1436,180,807]" captionTargetPageId="3" captionText="FIGURE 2. Holotype of Cylindrophis osheai sp. nov. (RMNH 5460). Head in (A) dorsal, (B) lateral, and (C) ventral view; (D) tail in ventral view showing subcaudals. Scale bar = 40 mm. Drawings by Felix Mader based on photographs by Sven Mecke." httpUri="https://zenodo.org/record/1437070/files/figure.png" pageId="1" pageNumber="237">2C</figureCitation>
|
||
). Cranial characteriSticS (
|
||
<figureCitation id="E6952A4BD0428C5E9F70F88AE5D4B5A7" box="[849,928,1900,1925]" captionStart="FIGURE 4" captionStartId="6.[151,250,1857,1879]" captionTargetBox="[232,1359,193,1847]" captionTargetId="figure@6.[232,1359,193,1847]" captionTargetPageId="6" captionText="FIGURE 4. Micro-CT images of the cranium of the holotype of Cylindrophis osheai sp. nov. (RMNH 5460), including (A) dorsal, (B) lateral and mirrored, (C) and ventral view, as well as (D) a dorsal view of the lower jaw. Abbreviations of cranial elements follow Lee & Scanlon (2002) and include: bo = basioccipital, com = compound bone of mandible, den = dentary, ec = ectopterygoid, eo = exoccipitals, fr = frontal, mx = maxilla, na = nasal, pa = parietal, pbs = parabasisphenoid, pof = postorbitofrontal, prf = prefrontal, pro = prootics, pt = pterygoid, qa = quadrate, sm = septomaxilla, so = supraoccipital, and st = supratemporal. Scale bar = 2 mm. Micro-CT images prepared by Kristin Mahlow." httpUri="https://zenodo.org/record/1437074/files/figure.png" pageId="1" pageNumber="237">FigS. 4</figureCitation>
|
||
,
|
||
<figureCitation id="E6952A4BD0428C5E9F8DF88AE5BBB5A6" box="[940,975,1900,1925]" captionStart="FIGURE 5" captionStartId="7.[151,250,1221,1243]" captionTargetBox="[151,1436,193,1200]" captionTargetId="figure@7.[151,1436,193,1200]" captionTargetPageId="7" captionText="FIGURE 5. Micro-CT images of skulls of several species of Cylindrophis used for comparison of cranial osteology, shown in dorsal (left column), lateral (middle column), and ventral view (right column). (A) Cylindrophis osheai sp. nov. (RMNH 5460). (B) C. boulengeri (NHMB 12913). (C) C. melanotus (ZMB 4049). (D) C. ruffus sensu lato (ZMB 13129). Scale bars = 2 mm. Micro-CT images prepared by Kristin Mahlow." httpUri="https://zenodo.org/record/1437076/files/figure.png" pageId="1" pageNumber="237">5A</figureCitation>
|
||
) of thiS taxon include (1) well-developed poStorbitofrontalS that project laterally well beyond the maxilla in dorSal view, creating a characteriStic, horn-like appearance; (2) a broad parietal with a bulbouS middle Section; (3) 10–12 maxillary teeth; (4) nine palatine teeth; (5) 7–9 teeth on the pterygoid; and (6) 11–13 teeth on the dentary.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="2AD16646D0418C5D9CB6F8ADE4A9B5F8" httpUri="https://zenodo.org/record/1437068/files/figure.png" pageId="2" pageNumber="238" startId="2.[151,250,1867,1889]" targetBox="[173,1405,249,1807]" targetPageId="2">
|
||
<paragraph id="7E1136CED0418C5D9CB6F8ADE4A9B5F8" blockId="2.[151,1436,1867,2011]" pageId="2" pageNumber="238">
|
||
<emphasis id="4CDAEADCD0418C5D9CB6F8ADE767B542" bold="true" box="[151,275,1867,1889]" pageId="2" pageNumber="238">FIGURE 1.</emphasis>
|
||
Known specimens of
|
||
<taxonomicName id="B9AE4D4DD0418C5D9DD9F8ADE4B0B542" authority="Kieckbusch & Mader & Kaiser & Mecke, 2018" authorityName="Kieckbusch & Mader & Kaiser & Mecke" authorityYear="2018" box="[504,708,1867,1889]" class="Reptilia" family="Cylindrophiidae" genus="Cylindrophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="238" phylum="Chordata" rank="species" species="osheai" status="sp. nov.">
|
||
<emphasis id="4CDAEADCD0418C5D9DD9F8ADE4B0B542" box="[504,708,1867,1889]" italics="true" pageId="2" pageNumber="238">Cylindrophis osheai</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="57E957A7D0418C5D9EEAF8ADE56FB542" box="[715,795,1867,1889]" pageId="2" pageNumber="238" rank="species">
|
||
<emphasis id="4CDAEADCD0418C5D9EEAF8ADE56FB542" bold="true" box="[715,795,1867,1889]" pageId="2" pageNumber="238">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
in dorsal (left) and ventral view (right). (A) Holotype, RMNH 5460 (formerly RMNH 5460.70), an adult female, collected in 1865 by Dirk Samuel Hoedt on Boano Island, Central Maluku Regency, Maluku Province, Indonesia. (B) Paratype, RMNH 47930 (formerly RMNH 5460.69), a subadult male, collection data as for holotype. (C) RMNH 11262, an unsexed specimen, labeled with collection locality “Celebes?” [= Sulawesi, in error]. Scale bars = 10 mm. Photographs by Sven Mecke.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="2AD16646D0408C5C9CB6FCD4E3E6B144" httpUri="https://zenodo.org/record/1437070/files/figure.png" pageId="3" pageNumber="239" startId="3.[151,250,818,840]" targetBox="[156,1432,191,797]" targetPageId="3">
|
||
<paragraph id="7E1136CED0408C5C9CB6FCD4E3E6B144" blockId="3.[151,1436,818,871]" pageId="3" pageNumber="239">
|
||
<emphasis id="4CDAEADCD0408C5C9CB6FCD4E766B16B" bold="true" box="[151,274,818,840]" pageId="3" pageNumber="239">FIGURE 2.</emphasis>
|
||
Holotype of
|
||
<taxonomicName id="B9AE4D4DD0408C5C9DB6FCD4E417B16B" authority="Kieckbusch & Mader & Kaiser & Mecke, 2018" authorityName="Kieckbusch & Mader & Kaiser & Mecke" authorityYear="2018" box="[407,611,818,840]" class="Reptilia" family="Cylindrophiidae" genus="Cylindrophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="239" phylum="Chordata" rank="species" species="osheai" status="sp. nov.">
|
||
<emphasis id="4CDAEADCD0408C5C9DB6FCD4E417B16B" box="[407,611,818,840]" italics="true" pageId="3" pageNumber="239">Cylindrophis osheai</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="57E957A7D0408C5C9E48FCD4E4CDB16B" box="[617,697,818,840]" pageId="3" pageNumber="239" rank="species">
|
||
<emphasis id="4CDAEADCD0408C5C9E48FCD4E4CDB16B" bold="true" box="[617,697,818,840]" pageId="3" pageNumber="239">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(RMNH 5460). Head in (A) dorsal, (B) lateral, and (C) ventral view; (D) tail in ventral view showing subcaudals. Scale bar = 40 mm. Drawings by Felix Mader based on photographs by Sven Mecke.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="36B46545D0408C589CE6FC68E373B412" lastPageId="7" lastPageNumber="243" pageId="3" pageNumber="239" type="description">
|
||
<paragraph id="7E1136CED0408C5C9CE6FC68E240B6E9" blockId="3.[151,1437,910,2018]" pageId="3" pageNumber="239">
|
||
<emphasis id="4CDAEADCD0408C5C9CE6FC68E43EB184" bold="true" box="[199,586,910,935]" pageId="3" pageNumber="239">Comparisons with other species.</emphasis>
|
||
<taxonomicName id="B9AE4D4DD0408C5C9E73FC68E540B184" authorityName="Kieckbusch & Mader & Kaiser & Mecke" authorityYear="2018" box="[594,820,910,935]" class="Reptilia" family="Cylindrophiidae" genus="Cylindrophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="239" phylum="Chordata" rank="species" species="osheai">
|
||
<emphasis id="4CDAEADCD0408C5C9E73FC68E540B184" box="[594,820,910,935]" italics="true" pageId="3" pageNumber="239">Cylindrophis osheai</emphasis>
|
||
</taxonomicName>
|
||
differS from all congenerS by a combination of unique color pattern characterS, including a light Snout bordered by a dark head cap with a bar running from the eye through the 3rd and 4th SupralabialS to the mouth (only preSent elSewhere in the genuS in a majority of
|
||
<taxonomicName id="B9AE4D4DD0408C5C9929FC3CE3E8B1D1" authorityName="Wagler" authorityYear="1828" box="[1288,1436,985,1010]" class="Reptilia" family="Cylindrophiidae" genus="Cylindrophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="239" phylum="Chordata" rank="species" species="melanotus">
|
||
<emphasis id="4CDAEADCD0408C5C9929FC3CE36FB1D2" box="[1288,1307,986,1009]" italics="true" pageId="3" pageNumber="239">C</emphasis>
|
||
.
|
||
<emphasis id="4CDAEADCD0408C5C9908FC3FE3E8B1D1" box="[1321,1436,985,1010]" italics="true" pageId="3" pageNumber="239">melanotus</emphasis>
|
||
</taxonomicName>
|
||
SpecimenS); a dark, double-diamond-Shaped ventral blotch in the area immediately poSterior to the genialS (
|
||
<figureCitation id="E6952A4BD0408C5C994EFC1BE6CFB61A" captionStart="FIGURE 2" captionStartId="3.[151,250,818,840]" captionTargetBox="[156,1432,191,797]" captionTargetId="figure@3.[151,1436,180,807]" captionTargetPageId="3" captionText="FIGURE 2. Holotype of Cylindrophis osheai sp. nov. (RMNH 5460). Head in (A) dorsal, (B) lateral, and (C) ventral view; (D) tail in ventral view showing subcaudals. Scale bar = 40 mm. Drawings by Felix Mader based on photographs by Sven Mecke." httpUri="https://zenodo.org/record/1437070/files/figure.png" pageId="3" pageNumber="239">Fig. 2C</figureCitation>
|
||
); 25¯28 dark ventral blotches aligned to form a broad wavy stripe for most part of the body (
|
||
<figureCitation id="E6952A4BD0408C5C98E3FBC7E36FB61A" box="[1218,1307,1057,1081]" captionStart="FIGURE 3" captionStartId="4.[151,250,646,668]" captionTargetBox="[153,1416,196,625]" captionTargetId="figure@4.[151,1436,193,629]" captionTargetPageId="4" captionText="FIGURE 3. Details of the ventral coloration in six species of Cylindrophis. (A) Cylindrophis osheai sp. nov. (RMNH 5460). (B) C. melanotus (RMNH 11451.114). (C) C. lineatus (AMNH 12872). (D) C. ruffus sensu stricto (RMNH 11256.110). (E) C. subocularis (RMNH 8959.57). (F) C. jodiae (NMW 21562.1). Drawings by Felix Mader based on photographs by Sven Mecke." httpUri="https://zenodo.org/record/1437072/files/figure.png" pageId="3" pageNumber="239">Fig. 3A</figureCitation>
|
||
; the ventral pattern of other taxa iS Shown in
|
||
<figureCitation id="E6952A4BD0408C5C9E3EFBA3E4ECB67D" box="[543,664,1093,1118]" captionStart="FIGURE 3" captionStartId="4.[151,250,646,668]" captionTargetBox="[153,1416,196,625]" captionTargetId="figure@4.[151,1436,193,629]" captionTargetPageId="4" captionText="FIGURE 3. Details of the ventral coloration in six species of Cylindrophis. (A) Cylindrophis osheai sp. nov. (RMNH 5460). (B) C. melanotus (RMNH 11451.114). (C) C. lineatus (AMNH 12872). (D) C. ruffus sensu stricto (RMNH 11256.110). (E) C. subocularis (RMNH 8959.57). (F) C. jodiae (NMW 21562.1). Drawings by Felix Mader based on photographs by Sven Mecke." httpUri="https://zenodo.org/record/1437072/files/figure.png" pageId="3" pageNumber="239">Fig. 3B–F</figureCitation>
|
||
); a dark ventral pattern completely Separated from the dark dorSal coloration; and an almoSt completely dark ventral Surface of the tail, with only the laSt Subcaudal and terminal Spine in a light color (
|
||
<figureCitation id="E6952A4BD0408C5C9DB1FB6BE79CB685" box="[400,488,1165,1190]" captionStart="FIGURE 2" captionStartId="3.[151,250,818,840]" captionTargetBox="[156,1432,191,797]" captionTargetId="figure@3.[151,1436,180,807]" captionTargetPageId="3" captionText="FIGURE 2. Holotype of Cylindrophis osheai sp. nov. (RMNH 5460). Head in (A) dorsal, (B) lateral, and (C) ventral view; (D) tail in ventral view showing subcaudals. Scale bar = 40 mm. Drawings by Felix Mader based on photographs by Sven Mecke." httpUri="https://zenodo.org/record/1437070/files/figure.png" pageId="3" pageNumber="239">Fig. 2D</figureCitation>
|
||
) (dark coloration coverS 3/4–5/6 of tail length, whereaS in
|
||
<taxonomicName id="B9AE4D4DD0408C5C98A2FB68E363B685" authorityName="Wagler" authorityYear="1828" box="[1155,1303,1165,1190]" class="Reptilia" family="Cylindrophiidae" genus="Cylindrophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="239" phylum="Chordata" rank="species" species="melanotus">
|
||
<emphasis id="4CDAEADCD0408C5C98A2FB68E2E2B686" box="[1155,1174,1166,1189]" italics="true" pageId="3" pageNumber="239">C</emphasis>
|
||
.
|
||
<emphasis id="4CDAEADCD0408C5C9885FB6BE363B685" box="[1188,1303,1165,1190]" italics="true" pageId="3" pageNumber="239">melanotus</emphasis>
|
||
</taxonomicName>
|
||
78% of the SpecimenS we examined had leSS than half of the tail length Showing a dark color).
|
||
</paragraph>
|
||
<paragraph id="7E1136CED0408C5C9CE6FB30E232B5C1" blockId="3.[151,1437,910,2018]" pageId="3" pageNumber="239">
|
||
In the following individual compariSonS of external morphology, the characterS for
|
||
<taxonomicName id="B9AE4D4DD0408C5C9895FB33E3E8B6CD" authorityName="Kieckbusch & Mader & Kaiser & Mecke" authorityYear="2018" box="[1204,1436,1237,1262]" class="Reptilia" family="Cylindrophiidae" genus="Cylindrophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="239" phylum="Chordata" rank="species" species="osheai">
|
||
<emphasis id="4CDAEADCD0408C5C9895FB33E3E8B6CD" box="[1204,1436,1237,1262]" italics="true" pageId="3" pageNumber="239">Cylindrophis osheai</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<emphasis id="4CDAEADCD0408C5C9C81FB1CE6DAB732" box="[160,174,1274,1297]" italics="true" pageId="3" pageNumber="239">n</emphasis>
|
||
= 3) are provided in parentheSeS.
|
||
<taxonomicName id="B9AE4D4DD0408C5C9E75FB1FE524B732" authorityName="Boulenger" authorityYear="1920" box="[596,848,1273,1298]" class="Reptilia" family="Cylindrophiidae" genus="Cylindrophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="239" phylum="Chordata" rank="species" species="aruensis">
|
||
<emphasis id="4CDAEADCD0408C5C9E75FB1FE524B732" box="[596,848,1273,1298]" italics="true" pageId="3" pageNumber="239">Cylindrophis aruensis</emphasis>
|
||
</taxonomicName>
|
||
poSSeSSeS 23 (19) dorSal Scale rowS at midbody, 176¯179 (224¯226) ventrals, and six (8¯9) subcaudals (
|
||
<bibRefCitation id="1A3F4B3FD0408C5C9F70FAFBE260B715" box="[849,1044,1309,1334]" pageId="3" pageNumber="239" refId="ref7388" refString="Boulenger, G. A. (1920) Descriptions of four new snakes in the collection of the British Museum. Annals and Magazine of Natural History, Series 9, 6 (31), 108 ‾ 111. https: // doi. org / 10.1080 / 00222932008632417" type="journal article">Boulenger 1920</bibRefCitation>
|
||
; McDowell 1975).
|
||
<taxonomicName id="B9AE4D4DD0408C5C992AFAFBE764B779" authorityName="Roux" authorityYear="1911" class="Reptilia" family="Cylindrophiidae" genus="Cylindrophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="239" phylum="Chordata" rank="species" species="boulengeri">
|
||
<emphasis id="4CDAEADCD0408C5C992AFAFBE764B779" italics="true" pageId="3" pageNumber="239">Cylindrophis boulengeri</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<emphasis id="4CDAEADCD0408C5C9D01FAA4E73CB77A" box="[288,328,1346,1369]" italics="true" pageId="3" pageNumber="239">n =</emphasis>
|
||
4) poSSeSSeS 195¯204 (224¯226) ventralS, 5¯6 (8¯9) SubcaudalS, and a light collar (collar abSent).
|
||
<taxonomicName id="B9AE4D4DD0408C5C9CB6FA83E441B75D" authority="Smith, 1943" authorityName="Smith" authorityYear="1943" box="[151,565,1381,1406]" class="Reptilia" family="Cylindrophiidae" genus="Cylindrophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="239" phylum="Chordata" rank="species" species="burmanus">
|
||
<emphasis id="4CDAEADCD0408C5C9CB6FA83E7D5B75D" box="[151,417,1381,1406]" italics="true" pageId="3" pageNumber="239">Cylindrophis burmanus</emphasis>
|
||
<bibRefCitation id="1A3F4B3FD0408C5C9D8BFA83E441B75D" box="[426,565,1381,1406]" pageId="3" pageNumber="239" refId="ref8316" refString="Smith, M. A. (1943) The Fauna of British India, Ceylon and Burma, including the Whole of the Indo-Chinese Subregion. Reptilia and Amphibia. Vol. III. Serpentes. Taylor and Francis, London, 583 pp." type="book">Smith, 1943</bibRefCitation>
|
||
</taxonomicName>
|
||
(
|
||
<emphasis id="4CDAEADCD0408C5C9E69FA81E406B75D" box="[584,626,1383,1406]" italics="true" pageId="3" pageNumber="239">n =</emphasis>
|
||
21) poSSeSSeS 190¯221 (224¯226) ventralS and 4¯7 (8¯9) SubcaudalS.
|
||
<taxonomicName id="B9AE4D4DD0408C5C9CB6FA6FE41BB781" authority="Stuebing, 1994" authorityName="Stuebing" authorityYear="1994" box="[151,623,1417,1442]" class="Reptilia" family="Cylindrophiidae" genus="Cylindrophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="239" phylum="Chordata" rank="species" species="engkariensis">
|
||
<emphasis id="4CDAEADCD0408C5C9CB6FA6FE7C9B781" box="[151,445,1417,1442]" italics="true" pageId="3" pageNumber="239">Cylindrophis engkariensis</emphasis>
|
||
<bibRefCitation id="1A3F4B3FD0408C5C9DE4FA6FE41BB781" box="[453,623,1417,1442]" pageId="3" pageNumber="239" refId="ref8556" refString="Stuebing, R. (1994) A new species of Cylindrophis (Serpentes: Cylindrophiidae) from Sarawak, Western Borneo. Raffles Bulletin of Zoology, 42 (4), 967 ‾ 973." type="journal article">Stuebing, 1994</bibRefCitation>
|
||
</taxonomicName>
|
||
(
|
||
<emphasis id="4CDAEADCD0408C5C9EA1FA6CE4DDB782" box="[640,681,1418,1441]" italics="true" pageId="3" pageNumber="239">n =</emphasis>
|
||
1) poSSeSSeS 17 (19) dorSal Scale rowS at midbody, 234 (224¯226) ventralS, five (8¯9) SubcaudalS, and rugoSe (Smooth) dorSalS on the tail.
|
||
<taxonomicName id="B9AE4D4DD0408C5C9F98FA4BE311B7E5" authority="Boulenger, 1896" authorityName="Boulenger" authorityYear="1896" box="[953,1381,1453,1478]" class="Reptilia" family="Cylindrophiidae" genus="Cylindrophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="239" phylum="Chordata" rank="species" species="isolepis">
|
||
<emphasis id="4CDAEADCD0408C5C9F98FA4BE2D1B7E5" box="[953,1189,1453,1478]" italics="true" pageId="3" pageNumber="239">Cylindrophis isolepis</emphasis>
|
||
<bibRefCitation id="1A3F4B3FD0408C5C988DFA4BE311B7E5" box="[1196,1381,1453,1478]" pageId="3" pageNumber="239" refId="ref7268" refString="Boulenger, G. A. (1896) Descriptions of new reptiles and batrachians obtained by Mr. Alfred Everett in Celebes and Jampea. Annals and Magazine of Natural History, Series 6, 18 (103), 62 ‾ 64. https: // doi. org / 10.1080 / 00222939608680409" type="journal article">Boulenger, 1896</bibRefCitation>
|
||
</taxonomicName>
|
||
(
|
||
<emphasis id="4CDAEADCD0408C5C9955FA49E3E8B7E5" box="[1396,1436,1455,1478]" italics="true" pageId="3" pageNumber="239">n =</emphasis>
|
||
2) poSSeSSeS 21 (19) dorSal Scale rowS at midbody, 5¯6 (8¯9) SubcaudalS, and haS the naSalS Separated by the roStral (naSalS in contact).
|
||
<taxonomicName id="B9AE4D4DD0408C5C9D4EFA13E2D0B42D" authority="AmaraSinghe, Ineich, Campbell & Hallermann, 2015" authorityName="AmaraSinghe, Ineich, Campbell & Hallermann" authorityYear="2015" box="[367,1188,1525,1550]" class="Reptilia" family="Cylindrophiidae" genus="Cylindrophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="239" phylum="Chordata" rank="species" species="jodiae">
|
||
<emphasis id="4CDAEADCD0408C5C9D4EFA13E439B42D" box="[367,589,1525,1550]" italics="true" pageId="3" pageNumber="239">Cylindrophis jodiae</emphasis>
|
||
AmaraSinghe,
|
||
<bibRefCitation id="1A3F4B3FD0408C5C9ED6FA13E2D0B42D" box="[759,1188,1525,1550]" pageId="3" pageNumber="239" refId="ref7119" refString="Amarasinghe, A. A. T., Campbell, P. D., Hallermann, J., Sidik, I., Supriatna, J. & Ineich, I. (2015) Two new species of the genus Cylindrophis Wagler, 1828 (Squamata: Cylindrophiidae) from Southeast Asia. Amphibian & Reptile Conservation, 9 (1), 34 ‾ 51." type="journal article">Ineich, Campbell & Hallermann, 2015</bibRefCitation>
|
||
</taxonomicName>
|
||
(
|
||
<emphasis id="4CDAEADCD0408C5C9895FA11E2A8B42D" box="[1204,1244,1527,1550]" italics="true" pageId="3" pageNumber="239">n =</emphasis>
|
||
90) poSSeSSeS 21 (19) dorSal Scale rowS at midbody, 177¯201 (224¯226) ventralS, and 5¯7 (8¯9) SubcaudalS.
|
||
<taxonomicName id="B9AE4D4DD0408C5C988BF9FFE735B475" authority="Blanford, 1881" authorityName="Blanford" authorityYear="1881" class="Reptilia" family="Cylindrophiidae" genus="Cylindrophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="239" phylum="Chordata" rank="species" species="lineatus">
|
||
<emphasis id="4CDAEADCD0408C5C988BF9FFE3E8B411" box="[1194,1436,1561,1586]" italics="true" pageId="3" pageNumber="239">Cylindrophis lineatus</emphasis>
|
||
<bibRefCitation id="1A3F4B3FD0408C5C9CB6F9DBE735B475" box="[151,321,1597,1622]" pageId="3" pageNumber="239" refId="ref7190" refString="Blanford, W. T. (1881) On a collection of reptiles and frogs chiefly from Singapore. Proceedings of the Zoological Society of London, 1881, 215 ‾ 226. https: // doi. org / 10.5962 / bhl. title. 101596" type="book">Blanford, 1881</bibRefCitation>
|
||
</taxonomicName>
|
||
(
|
||
<emphasis id="4CDAEADCD0408C5C9D73F9D9E70EB475" box="[338,378,1599,1622]" italics="true" pageId="3" pageNumber="239">n =</emphasis>
|
||
1) poSSeSSeS 21 (19) dorSal Scale rowS at midbody, 210¯218 (224¯226) ventralS, and a dorSal pattern of StripeS (StripeS abSent) (See alSo AmaraSinghe
|
||
<emphasis id="4CDAEADCD0408C5C9F04F984E52DB459" box="[805,857,1633,1658]" italics="true" pageId="3" pageNumber="239">et al</emphasis>
|
||
., 2015).
|
||
<taxonomicName id="B9AE4D4DD0408C5C9FE3F987E2A4B459" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[962,1232,1633,1658]" class="Reptilia" family="Cylindrophiidae" genus="Cylindrophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="239" phylum="Chordata" rank="species" species="maculatus">
|
||
<emphasis id="4CDAEADCD0408C5C9FE3F987E2A4B459" box="[962,1232,1633,1658]" italics="true" pageId="3" pageNumber="239">Cylindrophis maculatus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="1A3F4B3FD0408C5C98C0F987E3E0B459" box="[1249,1428,1633,1658]" pageId="3" pageNumber="239" refId="ref8013" refString="Linnaeus, C. (1758) Systema Naturae per Regna Tria Naturae Secundum Classes, Ordines, Genera, Species, cum Characteribus, Differentiis, Synonymis, Locis. Tomus I. Editio Decima, Reformata. Salvius, Holmiae [Stockholm], 4 + 824 pp." type="book">LinnaeuS, 1758</bibRefCitation>
|
||
) (
|
||
<emphasis id="4CDAEADCD0408C5C9C81F961E6BDB4BD" box="[160,201,1671,1694]" italics="true" pageId="3" pageNumber="239">n =</emphasis>
|
||
37) haS 4¯7 (8¯9) SubcaudalS and a dorSal pattern of paired reddiSh brown blotcheS encloSed by dark ground color (no encloSed blotcheS).
|
||
<taxonomicName id="B9AE4D4DD0408C5C9DC6F94FE487B4E1" authorityName="Wagler" authorityYear="1828" box="[487,755,1705,1730]" class="Reptilia" family="Cylindrophiidae" genus="Cylindrophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="239" phylum="Chordata" rank="species" species="melanotus">
|
||
<emphasis id="4CDAEADCD0408C5C9DC6F94FE487B4E1" box="[487,755,1705,1730]" italics="true" pageId="3" pageNumber="239">Cylindrophis melanotus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<emphasis id="4CDAEADCD0408C5C9F24F94CE55BB4E2" box="[773,815,1706,1729]" italics="true" pageId="3" pageNumber="239">n =</emphasis>
|
||
39) possesses 228¯266 (224¯226) ventrals; Moluccan populationS (
|
||
<emphasis id="4CDAEADCD0408C5C9D0CF929E74FB4C5" box="[301,315,1743,1766]" italics="true" pageId="3" pageNumber="239">n</emphasis>
|
||
= 3) poSSeSS 232¯251 (224–226) ventralS.
|
||
<taxonomicName id="B9AE4D4DD0408C5C9F19F92BE34FB4C5" authority="Boulenger, 1897" authorityName="Boulenger" authorityYear="1897" box="[824,1339,1741,1766]" class="Reptilia" family="Cylindrophiidae" genus="Cylindrophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="239" phylum="Chordata" rank="species" species="opisthorhodus">
|
||
<emphasis id="4CDAEADCD0408C5C9F19F92BE200B4C5" box="[824,1140,1741,1766]" italics="true" pageId="3" pageNumber="239">Cylindrophis opisthorhodus</emphasis>
|
||
<bibRefCitation id="1A3F4B3FD0408C5C985FF92BE34FB4C5" box="[1150,1339,1741,1766]" pageId="3" pageNumber="239" refId="ref7323" refString="Boulenger, G. A. (1897) List of the reptiles and batrachians collected by Mr. Alfred Everett in Lombok, Flores, Sumba and Savu, with descriptions of new species. Annals and Magazine of Natural History, Series 6, 19 (113), 503 ‾ 509. https: // doi. org / 10.1080 / 00222939708680570" type="journal article">Boulenger, 1897</bibRefCitation>
|
||
</taxonomicName>
|
||
(
|
||
<emphasis id="4CDAEADCD0408C5C996EF929E30EB4C5" box="[1359,1402,1743,1766]" italics="true" pageId="3" pageNumber="239">n =</emphasis>
|
||
6) poSSeSSeS 23 (19) dorSal Scale rowS at midbody, 185¯210 (224¯226) ventralS, 4¯7 (8¯9) SubcaudalS, and a light dorSum with dark SpeckleS forming two paravertebral rowS and occaSionally a diScontinuouS vertebral line (no SpeckleS or vertebral line on dorSum) (See alSo AmaraSinghe
|
||
<emphasis id="4CDAEADCD0408C5C9F7CF8DCE5ECB571" box="[861,920,1849,1874]" italics="true" pageId="3" pageNumber="239">et al.</emphasis>
|
||
, 2015).
|
||
<taxonomicName id="B9AE4D4DD0408C5C9FDAF8DFE6A1B555" authority="(Laurenti, 1768) SenSu" authorityName="SenSu" baseAuthorityName="Laurenti" baseAuthorityYear="1768" class="Reptilia" family="Cylindrophiidae" genus="Cylindrophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="239" phylum="Chordata" rank="species" species="ruffus">
|
||
<emphasis id="4CDAEADCD0408C5C9FDAF8DFE2A3B571" box="[1019,1239,1849,1874]" italics="true" pageId="3" pageNumber="239">Cylindrophis ruffus</emphasis>
|
||
(
|
||
<bibRefCitation id="1A3F4B3FD0408C5C98C8F8DFE3E0B571" box="[1257,1428,1849,1874]" pageId="3" pageNumber="239" refId="ref7926" refString="Laurenti, J. N. (1768) Specimen Medicum, Exhibens Synopsin Reptilium Emendatam cum Experimentis Circa Venena et Antidota Reptilium Austracorum, Quod Authoritate et Consensu. Johann Thomas von Trattner, Vienna, 217 pp." type="book">Laurenti, 1768</bibRefCitation>
|
||
)
|
||
<taxonomicNameLabel id="57E957A7D0408C5C9CB6F8B8E6A1B555" box="[151,213,1886,1910]" pageId="3" pageNumber="239" sensu="lato">SenSu</taxonomicNameLabel>
|
||
</taxonomicName>
|
||
lato (aS defined by KieckbuSch
|
||
<emphasis id="4CDAEADCD0408C5C9E47F8B9E4D1B555" box="[614,677,1885,1910]" italics="true" pageId="3" pageNumber="239">et al.</emphasis>
|
||
2016;
|
||
<emphasis id="4CDAEADCD0408C5C9EDEF8B9E55AB555" box="[767,814,1887,1910]" italics="true" pageId="3" pageNumber="239">n =</emphasis>
|
||
243) poSSeSSeS 5¯7 (8¯9) SubcaudalS.
|
||
<taxonomicName id="B9AE4D4DD0408C5C992AF8BBE249B5B9" authority="KieckbuSch, Mecke, Hartmann, Ehrmantraut, O'Shea & KaiSer, 2016" authorityName="KieckbuSch, Mecke, Hartmann, Ehrmantraut, O'Shea & KaiSer" authorityYear="2016" class="Reptilia" family="Cylindrophiidae" genus="Cylindrophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="239" phylum="Chordata" rank="species" species="subocularis">
|
||
<emphasis id="4CDAEADCD0408C5C992AF8BBE76EB5B9" italics="true" pageId="3" pageNumber="239">Cylindrophis subocularis</emphasis>
|
||
KieckbuSch, Mecke, Hartmann, Ehrmantraut, O’Shea &
|
||
<bibRefCitation id="1A3F4B3FD0408C5C9F8CF867E249B5B9" box="[941,1085,1921,1946]" pageId="3" pageNumber="239" refId="ref7801" refString="Kieckbusch, M., Mecke, S., Hartmann, L., Ehrmantraut, L., O'Shea, M. & Kaiser, H. (2016) An inconspicuous, conspicuous new species of Asian pipesnake, genus Cylindrophis (Reptilia: Squamata: Cylindrophiidae), from the south coast of Jawa Tengah, Java, Indonesia, and an overview of the tangled taxonomic history of C. ruffus (Laurenti, 1768). Zootaxa, 4093 (1), 1 ‾ 25. https: // doi. org / 10.11646 / zootaxa. 4093.1.1" type="journal article">KaiSer, 2016</bibRefCitation>
|
||
</taxonomicName>
|
||
(
|
||
<emphasis id="4CDAEADCD0408C5C9871F864E20FB5BA" box="[1104,1147,1922,1945]" italics="true" pageId="3" pageNumber="239">n =</emphasis>
|
||
9) poSSeSSeS a Subocular Scale (Subocular Scale abSent), 190¯196 (224¯226) ventralS, and 6¯7 (8¯9) SubcaudalS.
|
||
<taxonomicName id="B9AE4D4DD0408C5C9849F843E311B59D" authorityName="Smith & Sidik" authorityYear="1998" box="[1128,1381,1957,1982]" class="Reptilia" family="Cylindrophiidae" genus="Cylindrophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="239" phylum="Chordata" rank="species" species="yamdena">
|
||
<emphasis id="4CDAEADCD0408C5C9849F843E311B59D" box="[1128,1381,1957,1982]" italics="true" pageId="3" pageNumber="239">Cylindrophis yamdena</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<emphasis id="4CDAEADCD0408C5C9955F841E3E8B59D" box="[1396,1436,1959,1982]" italics="true" pageId="3" pageNumber="239">n =</emphasis>
|
||
5) poSSeSSeS 21 (19) dorSal Scale rowS at midbody and 183¯195 (224¯226) ventralS.
|
||
</paragraph>
|
||
<caption id="2AD16646D0478C5B9CB6FD60E697B0DB" httpUri="https://zenodo.org/record/1437072/files/figure.png" pageId="4" pageNumber="240" startId="4.[151,250,646,668]" targetBox="[153,1416,196,625]" targetPageId="4">
|
||
<paragraph id="7E1136CED0478C5B9CB6FD60E697B0DB" blockId="4.[151,1437,646,760]" pageId="4" pageNumber="240">
|
||
<emphasis id="4CDAEADCD0478C5B9CB6FD60E767B0BF" bold="true" box="[151,275,646,668]" pageId="4" pageNumber="240">FIGURE 3.</emphasis>
|
||
Details of the ventral coloration in six species of
|
||
<taxonomicName id="B9AE4D4DD0478C5B9F37FD60E5EDB0BF" authorityName="Wagler" authorityYear="1828" box="[790,921,646,668]" class="Reptilia" family="Cylindrophiidae" genus="Cylindrophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="240" phylum="Chordata" rank="genus">
|
||
<emphasis id="4CDAEADCD0478C5B9F37FD60E5EDB0BF" box="[790,921,646,668]" italics="true" pageId="4" pageNumber="240">Cylindrophis</emphasis>
|
||
</taxonomicName>
|
||
. (A)
|
||
<taxonomicName id="B9AE4D4DD0478C5B9FF0FD60E2EAB0BF" authority="Kieckbusch & Mader & Kaiser & Mecke, 2018" authorityName="Kieckbusch & Mader & Kaiser & Mecke" authorityYear="2018" box="[977,1182,646,668]" class="Reptilia" family="Cylindrophiidae" genus="Cylindrophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="240" phylum="Chordata" rank="species" species="osheai" status="sp. nov.">
|
||
<emphasis id="4CDAEADCD0478C5B9FF0FD60E2EAB0BF" box="[977,1182,646,668]" italics="true" pageId="4" pageNumber="240">Cylindrophis osheai</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="57E957A7D0478C5B9884FD60E283B0BF" box="[1189,1271,646,668]" pageId="4" pageNumber="240" rank="species">
|
||
<emphasis id="4CDAEADCD0478C5B9884FD60E283B0BF" bold="true" box="[1189,1271,646,668]" pageId="4" pageNumber="240">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(RMNH 5460). (B)
|
||
<taxonomicName id="B9AE4D4DD0478C5B9CE0FD43E73DB099" authorityName="Wagler" authorityYear="1828" box="[193,329,676,698]" class="Reptilia" family="Cylindrophiidae" genus="Cylindrophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="240" phylum="Chordata" rank="species" species="melanotus">
|
||
<emphasis id="4CDAEADCD0478C5B9CE0FD43E6A6B099" box="[193,210,677,698]" italics="true" pageId="4" pageNumber="240">C</emphasis>
|
||
.
|
||
<emphasis id="4CDAEADCD0478C5B9CC0FD42E73DB099" box="[225,329,676,698]" italics="true" pageId="4" pageNumber="240">melanotus</emphasis>
|
||
</taxonomicName>
|
||
(RMNH 11451.114). (C)
|
||
<taxonomicName id="B9AE4D4DD0478C5B9E78FD43E4BDB099" authorityName="Blanford" authorityYear="1881" box="[601,713,676,698]" class="Reptilia" family="Cylindrophiidae" genus="Cylindrophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="240" phylum="Chordata" rank="species" species="lineatus">
|
||
<emphasis id="4CDAEADCD0478C5B9E78FD43E41EB099" box="[601,618,677,698]" italics="true" pageId="4" pageNumber="240">C</emphasis>
|
||
.
|
||
<emphasis id="4CDAEADCD0478C5B9E58FD42E4BDB099" box="[633,713,676,698]" italics="true" pageId="4" pageNumber="240">lineatus</emphasis>
|
||
</taxonomicName>
|
||
(AMNH 12872). (D)
|
||
<taxonomicName id="B9AE4D4DD0478C5B9F93FD43E239B099" authority="sensu" authorityName="SenSu" baseAuthorityName="Laurenti" baseAuthorityYear="1768" box="[946,1101,676,698]" class="Reptilia" family="Cylindrophiidae" genus="Cylindrophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="240" phylum="Chordata" rank="species" species="ruffus">
|
||
<emphasis id="4CDAEADCD0478C5B9F93FD43E5B7B099" box="[946,963,677,698]" italics="true" pageId="4" pageNumber="240">C</emphasis>
|
||
.
|
||
<emphasis id="4CDAEADCD0478C5B9FF3FD42E279B099" box="[978,1037,676,698]" italics="true" pageId="4" pageNumber="240">ruffus</emphasis>
|
||
<taxonomicNameLabel id="57E957A7D0478C5B9834FD43E239B099" box="[1045,1101,677,698]" pageId="4" pageNumber="240" sensu="stricto">sensu</taxonomicNameLabel>
|
||
</taxonomicName>
|
||
stricto (RMNH 11256.110). (E)
|
||
<taxonomicName id="B9AE4D4DD0478C5B9CB6FD22E759B0FA" authorityName="KieckbuSch, Mecke, Hartmann, Ehrmantraut, O'Shea & KaiSer" authorityYear="2016" box="[151,301,707,729]" class="Reptilia" family="Cylindrophiidae" genus="Cylindrophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="240" phylum="Chordata" rank="species" species="subocularis">
|
||
<emphasis id="4CDAEADCD0478C5B9CB6FD22E6DCB0FA" box="[151,168,708,729]" italics="true" pageId="4" pageNumber="240">C</emphasis>
|
||
.
|
||
<emphasis id="4CDAEADCD0478C5B9C96FD25E759B0FA" box="[183,301,707,729]" italics="true" pageId="4" pageNumber="240">subocularis</emphasis>
|
||
</taxonomicName>
|
||
(RMNH 8959.57). (F)
|
||
<taxonomicName id="B9AE4D4DD0478C5B9E05FD22E4F7B0FA" authorityName="AmaraSinghe, Ineich, Campbell & Hallermann" authorityYear="2015" box="[548,643,707,729]" class="Reptilia" family="Cylindrophiidae" genus="Cylindrophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="240" phylum="Chordata" rank="species" species="jodiae">
|
||
<emphasis id="4CDAEADCD0478C5B9E05FD22E441B0FA" box="[548,565,708,729]" italics="true" pageId="4" pageNumber="240">C</emphasis>
|
||
.
|
||
<emphasis id="4CDAEADCD0478C5B9E65FD25E4F7B0FA" box="[580,643,707,729]" italics="true" pageId="4" pageNumber="240">jodiae</emphasis>
|
||
</taxonomicName>
|
||
(NMW 21562.1). Drawings by Felix Mader based on photographs by Sven Mecke.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="7E1136CED0478C5B9CE6FCC2E4ADB782" blockId="4.[151,1437,804,2023]" pageId="4" pageNumber="240">
|
||
The following compariSon of cranial characterS were made between the
|
||
<typeStatus id="A115886CD0478C5B9FD4FCC2E223B11E" box="[1013,1111,804,829]" pageId="4" pageNumber="240" type="holotype">holotype</typeStatus>
|
||
(RMNH 5460) and
|
||
<typeStatus id="A115886CD0478C5B991DFCC3E3E8B11E" box="[1340,1436,805,829]" pageId="4" pageNumber="240" type="paratype">paratype</typeStatus>
|
||
(RMNH 47930) of
|
||
<taxonomicName id="B9AE4D4DD0478C5B9D5DFCAEE415B142" authorityName="Kieckbusch & Mader & Kaiser & Mecke" authorityYear="2018" box="[380,609,840,865]" class="Reptilia" family="Cylindrophiidae" genus="Cylindrophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="240" phylum="Chordata" rank="species" species="osheai">
|
||
<emphasis id="4CDAEADCD0478C5B9D5DFCAEE415B142" box="[380,609,840,865]" italics="true" pageId="4" pageNumber="240">Cylindrophis osheai</emphasis>
|
||
</taxonomicName>
|
||
and SpecimenS of
|
||
<taxonomicName id="B9AE4D4DD0478C5B9F60FCAFE5AAB142" authorityName="Roux" authorityYear="1911" box="[833,990,840,865]" class="Reptilia" family="Cylindrophiidae" genus="Cylindrophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="240" phylum="Chordata" rank="species" species="boulengeri">
|
||
<emphasis id="4CDAEADCD0478C5B9F60FCAFE520B143" box="[833,852,841,864]" italics="true" pageId="4" pageNumber="240">C</emphasis>
|
||
.
|
||
<emphasis id="4CDAEADCD0478C5B9F44FCAEE5AAB142" box="[869,990,840,865]" italics="true" pageId="4" pageNumber="240">boulengeri</emphasis>
|
||
</taxonomicName>
|
||
(NMBA 12913),
|
||
<taxonomicName id="B9AE4D4DD0478C5B9892FCAFE33FB142" authorityName="Wagler" authorityYear="1828" box="[1203,1355,840,865]" class="Reptilia" family="Cylindrophiidae" genus="Cylindrophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="240" phylum="Chordata" rank="species" species="melanotus">
|
||
<emphasis id="4CDAEADCD0478C5B9892FCAFE2B2B143" box="[1203,1222,841,864]" italics="true" pageId="4" pageNumber="240">C</emphasis>
|
||
.
|
||
<emphasis id="4CDAEADCD0478C5B98F9FCAEE33FB142" box="[1240,1355,840,865]" italics="true" pageId="4" pageNumber="240">melanotus</emphasis>
|
||
</taxonomicName>
|
||
(ZMB 4049), aS well aS a voucher of
|
||
<taxonomicName id="B9AE4D4DD0478C5B9E21FC8BE4D8B1A6" authority="SenSu" authorityName="SenSu" baseAuthorityName="Laurenti" baseAuthorityYear="1768" box="[512,684,877,901]" class="Reptilia" family="Cylindrophiidae" genus="Cylindrophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="240" phylum="Chordata" rank="species" species="ruffus">
|
||
<emphasis id="4CDAEADCD0478C5B9E21FC8BE411B1A6" box="[512,613,877,901]" italics="true" pageId="4" pageNumber="240">C. ruffus</emphasis>
|
||
<taxonomicNameLabel id="57E957A7D0478C5B9E4FFC8BE4D8B1A6" box="[622,684,877,901]" pageId="4" pageNumber="240" sensu="lato">SenSu</taxonomicNameLabel>
|
||
</taxonomicName>
|
||
lato (ZMB 13129) (
|
||
<figureCitation id="E6952A4BD0478C5B9FBAFC8AE596B1A6" box="[923,994,876,901]" captionStart="FIGURE 5" captionStartId="7.[151,250,1221,1243]" captionTargetBox="[151,1436,193,1200]" captionTargetId="figure@7.[151,1436,193,1200]" captionTargetPageId="7" captionText="FIGURE 5. Micro-CT images of skulls of several species of Cylindrophis used for comparison of cranial osteology, shown in dorsal (left column), lateral (middle column), and ventral view (right column). (A) Cylindrophis osheai sp. nov. (RMNH 5460). (B) C. boulengeri (NHMB 12913). (C) C. melanotus (ZMB 4049). (D) C. ruffus sensu lato (ZMB 13129). Scale bars = 2 mm. Micro-CT images prepared by Kristin Mahlow." httpUri="https://zenodo.org/record/1437076/files/figure.png" pageId="4" pageNumber="240">Fig. 5</figureCitation>
|
||
).
|
||
<collectingCountry id="06B9765ED0478C5B9FDBFC8BE26DB1A6" box="[1018,1049,877,901]" name="American Samoa" pageId="4" pageNumber="240">AS</collectingCountry>
|
||
for eidonomic characterS, cranial featureS of
|
||
<taxonomicName id="B9AE4D4DD0478C5B9D3CFC77E7F8B18A" authorityName="Kieckbusch & Mader & Kaiser & Mecke" authorityYear="2018" box="[285,396,912,937]" class="Reptilia" family="Cylindrophiidae" genus="Cylindrophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="240" phylum="Chordata" rank="species" species="osheai">
|
||
<emphasis id="4CDAEADCD0478C5B9D3CFC77E744B18B" box="[285,304,913,936]" italics="true" pageId="4" pageNumber="240">C</emphasis>
|
||
.
|
||
<emphasis id="4CDAEADCD0478C5B9D62FC76E7F8B18A" box="[323,396,912,937]" italics="true" pageId="4" pageNumber="240">osheai</emphasis>
|
||
</taxonomicName>
|
||
are liSted in parentheSeS.
|
||
<taxonomicName id="B9AE4D4DD0478C5B9EE2FC76E5ADB18A" authorityName="Roux" authorityYear="1911" box="[707,985,912,937]" class="Reptilia" family="Cylindrophiidae" genus="Cylindrophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="240" phylum="Chordata" rank="species" species="boulengeri">
|
||
<emphasis id="4CDAEADCD0478C5B9EE2FC76E5ADB18A" box="[707,985,912,937]" italics="true" pageId="4" pageNumber="240">Cylindrophis boulengeri</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="E6952A4BD0478C5B9FCCFC76E23CB18A" box="[1005,1096,912,937]" captionStart="FIGURE 5" captionStartId="7.[151,250,1221,1243]" captionTargetBox="[151,1436,193,1200]" captionTargetId="figure@7.[151,1436,193,1200]" captionTargetPageId="7" captionText="FIGURE 5. Micro-CT images of skulls of several species of Cylindrophis used for comparison of cranial osteology, shown in dorsal (left column), lateral (middle column), and ventral view (right column). (A) Cylindrophis osheai sp. nov. (RMNH 5460). (B) C. boulengeri (NHMB 12913). (C) C. melanotus (ZMB 4049). (D) C. ruffus sensu lato (ZMB 13129). Scale bars = 2 mm. Micro-CT images prepared by Kristin Mahlow." httpUri="https://zenodo.org/record/1437076/files/figure.png" pageId="4" pageNumber="240">Fig. 5B</figureCitation>
|
||
) differS by having a broader cranium in dorSal view (Slender and long in dorSal view); Smaller prefrontalS in compariSon to frontalS (frontalS Slightly Smaller than prefrontalS); very Small poStorbitofrontalS that do not project beyond the maxilla in dorSal view (well-developed poStorbitofrontalS projecting laterally well beyond the maxilla, creating a characteriStic hornlike appearance on the SideS of the cranium); a very Slender parietal with a leSS pronounced bulbouS middle Section (broad parietal with obviouS bulbouS middle Section); Seven (nine) palatine teeth; and Six (7–9) teeth on the pterygoid.
|
||
<taxonomicName id="B9AE4D4DD0478C5B9D35FB8EE456B6A2" authorityName="Wagler" authorityYear="1828" box="[276,546,1128,1153]" class="Reptilia" family="Cylindrophiidae" genus="Cylindrophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="240" phylum="Chordata" rank="species" species="melanotus">
|
||
<emphasis id="4CDAEADCD0478C5B9D35FB8EE456B6A2" box="[276,546,1128,1153]" italics="true" pageId="4" pageNumber="240">Cylindrophis melanotus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="E6952A4BD0478C5B9E14FB8EE4FAB6A2" box="[565,654,1128,1153]" captionStart="FIGURE 5" captionStartId="7.[151,250,1221,1243]" captionTargetBox="[151,1436,193,1200]" captionTargetId="figure@7.[151,1436,193,1200]" captionTargetPageId="7" captionText="FIGURE 5. Micro-CT images of skulls of several species of Cylindrophis used for comparison of cranial osteology, shown in dorsal (left column), lateral (middle column), and ventral view (right column). (A) Cylindrophis osheai sp. nov. (RMNH 5460). (B) C. boulengeri (NHMB 12913). (C) C. melanotus (ZMB 4049). (D) C. ruffus sensu lato (ZMB 13129). Scale bars = 2 mm. Micro-CT images prepared by Kristin Mahlow." httpUri="https://zenodo.org/record/1437076/files/figure.png" pageId="4" pageNumber="240">Fig. 5C</figureCitation>
|
||
) differS by having very Small poStorbitofrontalS that do not project beyond the maxilla in dorSal view (well-developed poStorbitofrontalS projecting laterally well beyond the maxilla, creating a characteriStic horn-like appearance on the SideS of the cranium); a very broad parietal that haS an obviouS bulbouS middle Section (leSS broad parietal with leSS bulbouS middle Section); Seven (nine) palatine teeth; and five (7–9) teeth on the pterygoid.
|
||
<taxonomicName id="B9AE4D4DD0478C5B9DC4FB1EE4B4B732" authorityName="SenSu" baseAuthorityName="Laurenti" baseAuthorityYear="1768" box="[485,704,1272,1297]" class="Reptilia" family="Cylindrophiidae" genus="Cylindrophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="240" phylum="Chordata" rank="species" species="ruffus">
|
||
<emphasis id="4CDAEADCD0478C5B9DC4FB1EE4B4B732" box="[485,704,1272,1297]" italics="true" pageId="4" pageNumber="240">Cylindrophis ruffus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="E6952A4BD0478C5B9EF0FB1EE55FB732" box="[721,811,1272,1297]" captionStart="FIGURE 5" captionStartId="7.[151,250,1221,1243]" captionTargetBox="[151,1436,193,1200]" captionTargetId="figure@7.[151,1436,193,1200]" captionTargetPageId="7" captionText="FIGURE 5. Micro-CT images of skulls of several species of Cylindrophis used for comparison of cranial osteology, shown in dorsal (left column), lateral (middle column), and ventral view (right column). (A) Cylindrophis osheai sp. nov. (RMNH 5460). (B) C. boulengeri (NHMB 12913). (C) C. melanotus (ZMB 4049). (D) C. ruffus sensu lato (ZMB 13129). Scale bars = 2 mm. Micro-CT images prepared by Kristin Mahlow." httpUri="https://zenodo.org/record/1437076/files/figure.png" pageId="4" pageNumber="240">Fig. 5D</figureCitation>
|
||
) differS by having a Short and broad cranium in dorSal view (Slender and long cranium); Slightly Smaller prefrontalS in compariSon to the frontalS (frontalS Slightly Smaller than prefrontalS); Small poStorbitofrontalS that do not project beyond the maxilla in dorSal view (well-developed poStorbitofrontalS projecting laterally well beyond the maxilla, creating a characteriStic horn-like appearance on the SideS of the cranium); and 6–7 (nine) palatine teeth.
|
||
</paragraph>
|
||
<paragraph id="7E1136CED0478C5A9CE6FA4AE79FB3AE" blockId="4.[151,1437,804,2023]" lastBlockId="5.[151,1437,151,1990]" lastPageId="5" lastPageNumber="241" pageId="4" pageNumber="240">
|
||
<emphasis id="4CDAEADCD0478C5B9CE6FA4AE464B7E6" bold="true" box="[199,528,1452,1477]" pageId="4" pageNumber="240">
|
||
Description of the
|
||
<typeStatus id="A115886CD0478C5B9D83FA4AE47FB7E6" box="[418,523,1452,1477]" pageId="4" pageNumber="240" type="holotype">holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
<emphasis id="4CDAEADCD0478C5B9E37FA48E562B7E6" box="[534,790,1452,1477]" italics="true" pageId="4" pageNumber="240">Metrics and pholidosis</emphasis>
|
||
. An adult female (RMNH 5460) with
|
||
<quantity id="B9569B2BD0478C5B98E2FA4BE354B7E7" box="[1219,1312,1453,1477]" metricMagnitude="-1" metricUnit="m" metricValue="4.72" pageId="4" pageNumber="240" unit="mm" value="472.0">472 mm</quantity>
|
||
Snout-vent length (SVL) +
|
||
<quantity id="B9569B2BD0478C5B9D68FA37E7ECB7CB" box="[329,408,1489,1513]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" pageId="4" pageNumber="240" unit="mm" value="14.0">14 mm</quantity>
|
||
tail length =
|
||
<quantity id="B9569B2BD0478C5B9E0DFA36E4FDB7CB" box="[556,649,1488,1513]" metricMagnitude="-1" metricUnit="m" metricValue="4.86" pageId="4" pageNumber="240" unit="mm" value="486.0">486 mm</quantity>
|
||
total length; tail very Short (3% of SVL); head not diStinct from body; body cylindrical, body diameter
|
||
<quantity id="B9569B2BD0478C5B9E30FA12E40DB42F" box="[529,633,1524,1549]" metricMagnitude="-2" metricUnit="m" metricValue="1.46" pageId="4" pageNumber="240" unit="mm" value="14.6">14.6 mm</quantity>
|
||
(3% of SVL); head length (HL)
|
||
<quantity id="B9569B2BD0478C5B9821FA13E21CB42F" box="[1024,1128,1525,1549]" metricMagnitude="-2" metricUnit="m" metricValue="1.28" pageId="4" pageNumber="240" unit="mm" value="12.8">12.8 mm</quantity>
|
||
(3% of SVL); head width
|
||
<quantity id="B9569B2BD0478C5B9CB6F9FFE69AB413" box="[151,238,1561,1585]" metricMagnitude="-3" metricUnit="m" metricValue="9.4" pageId="4" pageNumber="240" unit="mm" value="9.4">9.4 mm</quantity>
|
||
(73% of HL); Snout rounded in dorSal and lateral view (
|
||
<figureCitation id="E6952A4BD0478C5B9F4CF9FEE58AB412" box="[877,1022,1560,1585]" captionStart="FIGURE 2" captionStartId="3.[151,250,818,840]" captionTargetBox="[156,1432,191,797]" captionTargetId="figure@3.[151,1436,180,807]" captionTargetPageId="3" captionText="FIGURE 2. Holotype of Cylindrophis osheai sp. nov. (RMNH 5460). Head in (A) dorsal, (B) lateral, and (C) ventral view; (D) tail in ventral view showing subcaudals. Scale bar = 40 mm. Drawings by Felix Mader based on photographs by Sven Mecke." httpUri="https://zenodo.org/record/1437070/files/figure.png" pageId="4" pageNumber="240">Fig. 2A & B</figureCitation>
|
||
); Snout length (SL)
|
||
<quantity id="B9569B2BD0478C5B98C3F9FEE34DB413" box="[1250,1337,1560,1585]" metricMagnitude="-3" metricUnit="m" metricValue="5.6" pageId="4" pageNumber="240" unit="mm" value="5.6">5.6 mm</quantity>
|
||
(44% of HL), Snout width
|
||
<quantity id="B9569B2BD0478C5B9D7DF9DAE7C6B477" box="[348,434,1596,1621]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" pageId="4" pageNumber="240" unit="mm" value="4.6">4.6 mm</quantity>
|
||
(82% of SL); eye diameter
|
||
<quantity id="B9569B2BD0478C5B9EC6F9DBE548B477" box="[743,828,1597,1621]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="4" pageNumber="240" unit="mm" value="1.2">1.2 mm</quantity>
|
||
(9% of HL); pupil round; interorbital diStance
|
||
<quantity id="B9569B2BD0478C5B9966F9DAE3E9B477" box="[1351,1437,1596,1621]" metricMagnitude="-3" metricUnit="m" metricValue="5.4" pageId="4" pageNumber="240" unit="mm" value="5.4">5.4 mm</quantity>
|
||
(42% of HL); naSo-orbital diStance
|
||
<quantity id="B9569B2BD0478C5B9E06F986E40AB45B" box="[551,638,1632,1657]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="4" pageNumber="240" unit="mm" value="3.5">3.5 mm</quantity>
|
||
(27% of HL); internarial diStance
|
||
<quantity id="B9569B2BD0478C5B9822F987E228B45B" box="[1027,1116,1633,1657]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="4" pageNumber="240" unit="mm" value="3.1">3.1 mm</quantity>
|
||
; pelvic SpurS Slightly viSible externally, but mainly retracted in poucheS lateral to cloacal plate, covered by ScaleS; 19 dorSal Scale rowS one head length behind head, 19 dorSal Scale rowS at midbody, 19 dorSal Scale rowS one head length before cloaca, ScaleS Smooth, apical pitS abSent; 224 ventral ScaleS; nine undivided Subcaudal ScaleS (except the 1
|
||
<taxonomicName id="B9AE4D4DD0478C5B9895F92AE284B4CB" box="[1204,1264,1740,1768]" pageId="4" pageNumber="240" rank="subSpecies" subSpecies="and">st and</taxonomicName>
|
||
9th Subcaudal), tail capped by a terminal Spine; cloacal plate paired; roStral clearly viSible from above, triangular, wider than high, height
|
||
<quantity id="B9569B2BD0478C5B9CC9F8F1E730B50C" box="[232,324,1815,1840]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="4" pageNumber="240" unit="mm" value="2.5">2.5 mm</quantity>
|
||
, width
|
||
<quantity id="B9569B2BD0478C5B9DBEF8FEE788B50C" box="[415,508,1816,1840]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="4" pageNumber="240" unit="mm" value="2.7">2.7 mm</quantity>
|
||
; two irregularly pentagonal naSalS, length
|
||
<quantity id="B9569B2BD0478C5B9FD1F8FEE238B50C" box="[1008,1100,1816,1840]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="4" pageNumber="240" unit="mm" value="2.2">2.2 mm</quantity>
|
||
, width
|
||
<quantity id="B9569B2BD0478C5B9887F8FEE370B50C" box="[1190,1284,1816,1840]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="4" pageNumber="240" unit="mm" value="2.9">2.9 mm</quantity>
|
||
; naSal Suture SiniStral with reSpect to prefrontal Suture; noStril poSitioned cloSe to the Suture of the naSal with the firSt Supralabial; poStnaSal abSent; loreal abSent, irregularly hexagonal prefrontalS in contact with 2nd and 3rd SupralabialS; preocular abSent; Subocular abSent; one rectangular poStocular, length
|
||
<quantity id="B9569B2BD0478C5B9F76F861E5C7B5BD" box="[855,947,1927,1951]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="4" pageNumber="240" unit="mm" value="0.7">0.7 mm</quantity>
|
||
, height
|
||
<quantity id="B9569B2BD0478C5B9835F861E205B5BD" box="[1044,1137,1927,1951]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="4" pageNumber="240" unit="mm" value="1.1">1.1 mm</quantity>
|
||
; temporal formula 1 + 2, anterior temporal larger than lower poSterior temporal, almoSt equal in Size to upper poSterior temporal, anterior temporal length
|
||
<quantity id="B9569B2BD0478C5B9D6EF829E7DCB5C5" box="[335,424,1999,2023]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="4" pageNumber="240" unit="mm" value="2.7">2.7 mm</quantity>
|
||
, height
|
||
<quantity id="B9569B2BD0478C5B9E20F829E42EB5C5" box="[513,602,1999,2023]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="4" pageNumber="240" unit="mm" value="2.8">2.8 mm</quantity>
|
||
, upper poSterior temporal length 3.0 mm, height
|
||
<quantity id="B9569B2BD0478C5B98A3F828E2AFB5C5" box="[1154,1243,1998,2023]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="4" pageNumber="240" unit="mm" value="2.6">2.6 mm</quantity>
|
||
; Six SupralabialS, 1st SmalleSt, 3rd–5th largeSt and equal in Size, 2nd and 6th equal in Size and Slightly Smaller than 3rd–5th, the 3rd and 4th in contact with orbit; Six infralabialS, 1st–3rd in contact with firSt pair of genialS, 1
|
||
<taxonomicName id="B9AE4D4DD0468C5A9811FF5BE204B2F9" box="[1072,1136,189,218]" pageId="5" pageNumber="241" rank="subSpecies" subSpecies="pair">st pair</taxonomicName>
|
||
of infralabialS in contact, preventing contact of mental with firSt pair of genialS; mental triangular, wider than high, height
|
||
<quantity id="B9569B2BD0468C5A98EEFF03E353B2DE" box="[1231,1319,229,254]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="5" pageNumber="241" unit="mm" value="1.5">1.5 mm</quantity>
|
||
, width
|
||
<quantity id="B9569B2BD0468C5A9958FF00E6B2B302" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="5" pageNumber="241" unit="mm" value="2.3">2.3 mm</quantity>
|
||
; two pairS of genialS, anterior genialS length 3.0 mm, width 2.0 mm, poSterior genialS length
|
||
<quantity id="B9569B2BD0468C5A98F1FEECE35CB302" box="[1232,1320,266,290]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="5" pageNumber="241" unit="mm" value="2.3">2.3 mm</quantity>
|
||
, width
|
||
<quantity id="B9569B2BD0468C5A9958FEECE6B2B366" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="5" pageNumber="241" unit="mm" value="1.8">1.8 mm</quantity>
|
||
; mental groove preSent, length
|
||
<quantity id="B9569B2BD0468C5A9E1BFECBE4E3B366" box="[570,663,301,326]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" pageId="5" pageNumber="241" unit="mm" value="4.6">4.6 mm</quantity>
|
||
; prefrontal hexagonal, length
|
||
<quantity id="B9569B2BD0468C5A9FD7FEC8E226B366" box="[1014,1106,302,326]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="5" pageNumber="241" unit="mm" value="3.1">3.1 mm</quantity>
|
||
, width
|
||
<quantity id="B9569B2BD0468C5A988CFECBE37EB366" box="[1197,1290,301,326]" metricMagnitude="-3" metricUnit="m" metricValue="3.6" pageId="5" pageNumber="241" unit="mm" value="3.6">3.6 mm</quantity>
|
||
; Supraocular pentagonal, length
|
||
<quantity id="B9569B2BD0468C5A9D4FFEB7E7B2B34A" box="[366,454,337,362]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="5" pageNumber="241" unit="mm" value="3.5">3.5 mm</quantity>
|
||
, width
|
||
<quantity id="B9569B2BD0468C5A9E3BFEB4E400B34A" box="[538,628,338,362]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="5" pageNumber="241" unit="mm" value="2.8">2.8 mm</quantity>
|
||
; frontal rectangular, length
|
||
<quantity id="B9569B2BD0468C5A9F8AFEB4E270B34A" box="[939,1028,338,362]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="5" pageNumber="241" unit="mm" value="3.3">3.3 mm</quantity>
|
||
, width
|
||
<quantity id="B9569B2BD0468C5A9876FEB4E2C5B34A" box="[1111,1201,338,362]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="5" pageNumber="241" unit="mm" value="2.9">2.9 mm</quantity>
|
||
; parietal rectangular, length
|
||
<quantity id="B9569B2BD0468C5A9CC5FE90E748B3AE" box="[228,316,374,398]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="5" pageNumber="241" unit="mm" value="2.9">2.9 mm</quantity>
|
||
, width
|
||
<quantity id="B9569B2BD0468C5A9DAFFE93E793B3AE" box="[398,487,373,398]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="5" pageNumber="241" unit="mm" value="2.5">2.5 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="7E1136CED0468C5A9CE6FE7FE5FDB411" blockId="5.[151,1437,151,1990]" pageId="5" pageNumber="241">
|
||
<emphasis id="4CDAEADCD0468C5A9CE6FE7FE7E4B391" box="[199,400,409,434]" italics="true" pageId="5" pageNumber="241">Cranial osteology</emphasis>
|
||
. A Single, Small, and toothleSS median premaxilla preSent at the anteriormoSt point of the Snout, triangle-Shaped in dorSal view, in looSe contact with the naSalS, maxilla, and Septomaxilla; paired naSalS elongated in dorSal view, almoSt three timeS aS long aS broad, medially in looSe contact, outer edgeS of naSalS lacerated, poSteriorly contacting anterior edgeS of frontalS; paired, Subpentangular frontalS flattened dorSally, about two timeS aS long aS broad, Slightly broader but Significantly Shorter than naSalS, and in looSe medial contact, contacting prefrontalS ventrally with anterolateral edge; prefrontal contacting naSal dorSally in lateral view and the dorSal Surface of the maxilla ventrally; poStorbitofrontalS well developed, articulate looSely with the lateral edgeS of the frontalS, and, in dorSal view, projecting laterally well beyond the maxilla, creating a characteriStic horn-like appearance on the SideS of the cranium; fuSed parietalS in cloSe contact with frontalS anteriorly, ventrally in contact with the parabaSiSphenoid, poSterolateral borderS in contact with prooticS and SupratemporalS, poSterior border in cloSe contact with Supraoccipital, flat in dorSal view, except for a median creSt, concave-Shaped in the anterior and convex-Shaped in the poSterior portion with a bulbouS middle Section, poSSeSSing long proceSSeS at the anterior edge framing the frontalS, creSt of fuSed parietalS more diStinct at poSterior portion; Single Supraoccipital Somewhat wing-Shaped, much broader than long, bearing a diStinct keel that iS more Strongly developed than parietal keel, laterally in contact with prooticS and poSteriorly with exoccipitalS; paired prooticS largely elongated in dorSal view, Subrectangular in lateral view, encloSing the poSterior portion of the fuSed parietalS together with the two SupratemporalS, ventrally in contact with parabaSiSphenoid and baSioccipital, each prootic with two large foramina, prooticS connecting the poSterolateral dorSal part of the Skull with the quadrate; SupratemporalS curved, encloSing parietal, poSterior edgeS of prooticS, Supraoccipital, and exoccipitalS in dorSal view; exoccipitalS very Short in dorSal view, forming poSterolateral wallS of braincaSe, joined together by a Short Suture, fuSed with the baSioccipital ventrally at their poSteriormoSt portion; baSioccipital Subpentagonal, in cloSe contact with the parabaSiSphenoid anteroventrally, forming a tranSverSe Suture; parabaSiSphenoid extending to roStral area anteriorly, in looSe contact with palatineS and frontalS; maxilla Slightly curved, with a dorSal proceSS articulating with ventral Surface of prefrontal, articulating with Septomaxilla anterodorSally, poSterior portion of maxilla articulating with ectopterygoid, connecting maxilla with pterygoid, maxilla broadeSt anteriorly in lateral view, two large foramina preSent, each maxilla bearing 11 Slightly recurved teeth, no maxillary diaStema preSent; ectopterygoid without teeth; palatine narrow, bearing nine teeth, poSterior end of each palatine articulating with anterior end of pterygoid; pterygoid bearing Seven teeth; quadrate articulating with poSterolateral border of SupratemporalS, itS ventral Surface articulating with the mandibular condyle; mandible compoSed of three boneS; anterior portion of jaw formed by a dentary originally bearing 11|13 Slightly recurved teeth (SometimeS SocketS are preSent only), dentary about half the length of the mandible; poSterior part of the jaw and the compound bone of the mandible without teeth; coronoid located at the level of articulation of dentary with mandible, juSt below ectopterygoid and the anterior proceSS of the pterygoid (See
|
||
<figureCitation id="E6952A4BD0468C5A9F16F9FFE508B411" box="[823,892,1561,1586]" captionStart="FIGURE 4" captionStartId="6.[151,250,1857,1879]" captionTargetBox="[232,1359,193,1847]" captionTargetId="figure@6.[232,1359,193,1847]" captionTargetPageId="6" captionText="FIGURE 4. Micro-CT images of the cranium of the holotype of Cylindrophis osheai sp. nov. (RMNH 5460), including (A) dorsal, (B) lateral and mirrored, (C) and ventral view, as well as (D) a dorsal view of the lower jaw. Abbreviations of cranial elements follow Lee & Scanlon (2002) and include: bo = basioccipital, com = compound bone of mandible, den = dentary, ec = ectopterygoid, eo = exoccipitals, fr = frontal, mx = maxilla, na = nasal, pa = parietal, pbs = parabasisphenoid, pof = postorbitofrontal, prf = prefrontal, pro = prootics, pt = pterygoid, qa = quadrate, sm = septomaxilla, so = supraoccipital, and st = supratemporal. Scale bar = 2 mm. Micro-CT images prepared by Kristin Mahlow." httpUri="https://zenodo.org/record/1437074/files/figure.png" pageId="5" pageNumber="241">Fig. 4</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="7E1136CED0468C5A9CE6F9DBE408B5E5" blockId="5.[151,1437,151,1990]" pageId="5" pageNumber="241">
|
||
<emphasis id="4CDAEADCD0468C5A9CE6F9DBE7A6B475" box="[199,466,1597,1622]" italics="true" pageId="5" pageNumber="241">Coloration and pattern</emphasis>
|
||
(
|
||
<emphasis id="4CDAEADCD0468C5A9DC7F9D8E5E3B476" box="[486,919,1597,1622]" italics="true" pageId="5" pageNumber="241">after 152 years in liquid preservative</emphasis>
|
||
). DorSal Surface of trunk and tail uniformly Sepia (279), but faded on Some partS of the dorSum; dorSal ground color reaching mid-lateral region, lower lateral and ventrolateral regionS Pale Buff (1), a majority of pale ScaleS with Sepia (279) colored anterior edgeS. Snout Pale Buff (1) with diffuSe Sepia (279) markingS on naSalS, roStral, and 1
|
||
<taxonomicName id="B9AE4D4DD0468C5A9FF5F94EE264B4E6" box="[980,1040,1704,1733]" pageId="5" pageNumber="241" rank="subSpecies" subSpecies="and">st and</taxonomicName>
|
||
2nd SupralabialS; a Sepia (279) bar running from the eye through the 3rd and 4th SupralabialS to the mouth, bordering the pale Snout; ventral Surface of the head Pale Buff (1) with a diffuSe Sepia (279) colored blotch on the anterior and poSterior genialS, which iS followed by a characteriStic double-diamond-Shaped ventral pattern. Ventral Surface of trunk Pale Buff (1), with 28 Olive Brown (278) blotcheS forming a broad wavy Stripe for almoSt the entire length of the body; the dark ventral pattern completely Separated from the dark dorSal coloration; cloacal region and ventral Surface of tail covered by a large Sepia (279) colored blotch, beginning four ventralS anterior to the cloacal plate, ending on 8th Subcaudal; 9th Subcaudal and terminal Spine Pale Buff (1).
|
||
</paragraph>
|
||
<caption id="2AD16646D0458C599CB6F8A7E27FB5D2" httpUri="https://zenodo.org/record/1437074/files/figure.png" pageId="6" pageNumber="242" startId="6.[151,250,1857,1879]" targetBox="[232,1359,193,1847]" targetPageId="6">
|
||
<paragraph id="7E1136CED0458C599CB6F8A7E27FB5D2" blockId="6.[151,1436,1857,2033]" pageId="6" pageNumber="242">
|
||
<emphasis id="4CDAEADCD0458C599CB6F8A7E760B574" bold="true" box="[151,276,1857,1879]" pageId="6" pageNumber="242">FIGURE 4.</emphasis>
|
||
Micro-CT images of the cranium of the holotype of
|
||
<taxonomicName id="B9AE4D4DD0458C599F1AF8A7E27CB574" authority="Kieckbusch & Mader & Kaiser & Mecke, 2018" authorityName="Kieckbusch & Mader & Kaiser & Mecke" authorityYear="2018" box="[827,1032,1857,1879]" class="Reptilia" family="Cylindrophiidae" genus="Cylindrophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="242" phylum="Chordata" rank="species" species="osheai" status="sp. nov.">
|
||
<emphasis id="4CDAEADCD0458C599F1AF8A7E27CB574" box="[827,1032,1857,1879]" italics="true" pageId="6" pageNumber="242">Cylindrophis osheai</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="57E957A7D0458C599830F8A7E217B574" box="[1041,1123,1857,1879]" pageId="6" pageNumber="242" rank="species">
|
||
<emphasis id="4CDAEADCD0458C599830F8A7E217B574" bold="true" box="[1041,1123,1857,1879]" pageId="6" pageNumber="242">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(RMNH 5460), including (A) dorsal, (B) lateral and mirrored, (C) and ventral view, as well as (D) a dorsal view of the lower jaw. Abbreviations of cranial elements follow Lee & Scanlon (2002) and include: bo = basioccipital, com = compound bone of mandible, den = dentary, ec = ectopterygoid, eo = exoccipitals, fr = frontal, mx = maxilla, na = nasal, pa = parietal, pbs = parabasisphenoid, pof = postorbitofrontal, prf = prefrontal, pro = prootics, pt = pterygoid, qa = quadrate, sm = septomaxilla, so = supraoccipital, and st = supratemporal. Scale bar = 2 mm. Micro-CT images prepared by Kristin Mahlow.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="2AD16646D0448C589CB6FB23E403B714" httpUri="https://zenodo.org/record/1437076/files/figure.png" pageId="7" pageNumber="243" startId="7.[151,250,1221,1243]" targetBox="[151,1436,193,1200]" targetPageId="7">
|
||
<paragraph id="7E1136CED0448C589CB6FB23E403B714" blockId="7.[151,1437,1221,1335]" pageId="7" pageNumber="243">
|
||
<emphasis id="4CDAEADCD0448C589CB6FB23E767B6F8" bold="true" box="[151,275,1221,1243]" pageId="7" pageNumber="243">FIGURE 5.</emphasis>
|
||
Micro-CT images of skulls of several species of
|
||
<taxonomicName id="B9AE4D4DD0448C589F28FB23E5F8B6F8" authorityName="Wagler" authorityYear="1828" box="[777,908,1221,1243]" class="Reptilia" family="Cylindrophiidae" genus="Cylindrophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="243" phylum="Chordata" rank="genus">
|
||
<emphasis id="4CDAEADCD0448C589F28FB23E5F8B6F8" box="[777,908,1221,1243]" italics="true" pageId="7" pageNumber="243">Cylindrophis</emphasis>
|
||
</taxonomicName>
|
||
used for comparison of cranial osteology, shown in dorsal (left column), lateral (middle column), and ventral view (right column). (A)
|
||
<taxonomicName id="B9AE4D4DD0448C589FF8FB02E2D1B6D9" authority="Kieckbusch & Mader & Kaiser & Mecke, 2018" authorityName="Kieckbusch & Mader & Kaiser & Mecke" authorityYear="2018" box="[985,1189,1252,1274]" class="Reptilia" family="Cylindrophiidae" genus="Cylindrophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="243" phylum="Chordata" rank="species" species="osheai" status="sp. nov.">
|
||
<emphasis id="4CDAEADCD0448C589FF8FB02E2D1B6D9" box="[985,1189,1252,1274]" italics="true" pageId="7" pageNumber="243">Cylindrophis osheai</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="57E957A7D0448C58988BFB02E28EB6D9" box="[1194,1274,1252,1274]" pageId="7" pageNumber="243" rank="species">
|
||
<emphasis id="4CDAEADCD0448C58988BFB02E28EB6D9" bold="true" box="[1194,1274,1252,1274]" pageId="7" pageNumber="243">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(RMNH 5460). (B)
|
||
<taxonomicName id="B9AE4D4DD0448C589C9EFAE5E73FB73B" authorityName="Roux" authorityYear="1911" box="[191,331,1282,1304]" class="Reptilia" family="Cylindrophiidae" genus="Cylindrophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="243" phylum="Chordata" rank="species" species="boulengeri">
|
||
<emphasis id="4CDAEADCD0448C589C9EFAE5E73FB73B" box="[191,331,1282,1304]" italics="true" pageId="7" pageNumber="243">C. boulengeri</emphasis>
|
||
</taxonomicName>
|
||
(NHMB 12913). (C)
|
||
<taxonomicName id="B9AE4D4DD0448C589E0AFAE5E4C5B73B" authorityName="Wagler" authorityYear="1828" box="[555,689,1282,1304]" class="Reptilia" family="Cylindrophiidae" genus="Cylindrophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="243" phylum="Chordata" rank="species" species="melanotus">
|
||
<emphasis id="4CDAEADCD0448C589E0AFAE5E4C5B73B" box="[555,689,1282,1304]" italics="true" pageId="7" pageNumber="243">C. melanotus</emphasis>
|
||
</taxonomicName>
|
||
(ZMB 4049). (D)
|
||
<taxonomicName id="B9AE4D4DD0448C589F51FAE5E27CB73B" authority="sensu" authorityName="SenSu" baseAuthorityName="Laurenti" baseAuthorityYear="1768" box="[880,1032,1282,1304]" class="Reptilia" family="Cylindrophiidae" genus="Cylindrophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="240" phylum="Chordata" rank="species" species="ruffus">
|
||
<emphasis id="4CDAEADCD0448C589F51FAE5E5BDB73B" box="[880,969,1282,1304]" italics="true" pageId="7" pageNumber="243">C. ruffus</emphasis>
|
||
<taxonomicNameLabel id="57E957A7D0448C589FF1FAE5E27CB73B" box="[976,1032,1283,1304]" pageId="7" pageNumber="243" sensu="lato">sensu</taxonomicNameLabel>
|
||
</taxonomicName>
|
||
lato (ZMB 13129). Scale bars = 2 mm. Micro-CT images prepared by Kristin Mahlow.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="7E1136CED0448C589CE6FA82E373B412" blockId="7.[151,1437,1380,2017]" pageId="7" pageNumber="243">
|
||
<emphasis id="4CDAEADCD0448C589CE6FA82E7ADB75E" bold="true" box="[199,473,1380,1405]" pageId="7" pageNumber="243">Intraspecific variation.</emphasis>
|
||
The
|
||
<typeStatus id="A115886CD0448C589E33FA83E406B75E" box="[530,626,1381,1405]" pageId="7" pageNumber="243" type="paratype">paratype</typeStatus>
|
||
RMNH 47930 (Subadult male) differS from the
|
||
<typeStatus id="A115886CD0448C5898ACFA82E29BB75E" box="[1165,1263,1380,1405]" pageId="7" pageNumber="243" type="holotype">holotype</typeStatus>
|
||
(RMNH 5460) aS followS:
|
||
<quantity id="B9569B2BD0448C589D38FA6EE702B783" box="[281,374,1416,1441]" metricMagnitude="-1" metricUnit="m" metricValue="3.5799999999999996" pageId="7" pageNumber="243" unit="mm" value="358.0">358 mm</quantity>
|
||
SVL +
|
||
<quantity id="B9569B2BD0448C589DF0FA6EE46BB783" box="[465,543,1416,1440]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" pageId="7" pageNumber="243" unit="mm" value="11.0">11 mm</quantity>
|
||
tail length =
|
||
<quantity id="B9569B2BD0448C589E97FA6EE567B783" box="[694,787,1416,1441]" metricMagnitude="-1" metricUnit="m" metricValue="3.69" pageId="7" pageNumber="243" unit="mm" value="369.0">369 mm</quantity>
|
||
total length; 226 ventralS; 6|5 infralabialS; preSence of 31 Olive Brown (278), tranSverSe and alternating blotcheS on the Sepia (279) colored dorSum. RMNH 11262 (unSexed Specimen) differS from the
|
||
<typeStatus id="A115886CD0448C589DEBFA36E458B7CA" box="[458,556,1488,1513]" pageId="7" pageNumber="243" type="holotype">holotype</typeStatus>
|
||
(RMNH 5460) aS followS:
|
||
<quantity id="B9569B2BD0448C589F45FA36E5B6B7CB" box="[868,962,1488,1513]" metricMagnitude="-1" metricUnit="m" metricValue="4.6899999999999995" pageId="7" pageNumber="243" unit="mm" value="469.0">469 mm</quantity>
|
||
SVL +
|
||
<quantity id="B9569B2BD0448C58983FFA36E219B7CB" box="[1054,1133,1488,1513]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="7" pageNumber="243" unit="mm" value="15.0">15 mm</quantity>
|
||
tail length =
|
||
<quantity id="B9569B2BD0448C589924FA36E317B7CB" box="[1285,1379,1488,1512]" metricMagnitude="-1" metricUnit="m" metricValue="4.84" pageId="7" pageNumber="243" unit="mm" value="484.0">484 mm</quantity>
|
||
total length; 225 ventralS; eight SubcaudalS. With reSpect to cranial oSteology the
|
||
<typeStatus id="A115886CD0448C589821FA13E214B42E" box="[1024,1120,1525,1549]" pageId="7" pageNumber="243" type="paratype">paratype</typeStatus>
|
||
RMNH 47930 differS from the
|
||
<typeStatus id="A115886CD0448C589CE1F9FEE756B412" box="[192,290,1560,1585]" pageId="7" pageNumber="243" type="holotype">holotype</typeStatus>
|
||
aS followS: 12|10 maxillary teeth, 9|8 teeth on the pterygoid, and 12 teeth on the dentary.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="36B46545D0448C589CE6F9DAE594B5BB" pageId="7" pageNumber="243" type="etymology">
|
||
<paragraph id="7E1136CED0448C589CE6F9DAE335B572" blockId="7.[151,1437,1380,2017]" pageId="7" pageNumber="243">
|
||
<emphasis id="4CDAEADCD0448C589CE6F9DAE73AB476" bold="true" box="[199,334,1596,1621]" pageId="7" pageNumber="243">Etymology.</emphasis>
|
||
We deScribe the new SpecieS to honor our colleague and friend Mark O’Shea for hiS many contributionS to Snake biology and taxonomy. Mark haS authored Several herpetology bookS and a plethora of Scientific articleS, dealing eSpecially with the herpetofauna of SoutheaSt ASia’S iSlandS. He haS alSo contributed much to improving an underStanding of reptileS and particularly enhancing the reputation of SnakeS among a broader audience via hiS media work and hiS popular lectureS. Mark haS inSpired many young herpetologiStS to purSue StudieS of amphibianS and reptileS. He continueS to do So, having recently been appointed to the poSition of ProfeSSor of Herpetology at the UniverSity of
|
||
<collectingRegion id="BC6AF82CD0448C589EBBF8F2E53AB50E" box="[666,846,1812,1837]" country="United Kingdom" name="Wolverhampton" pageId="7" pageNumber="243">Wolverhampton</collectingRegion>
|
||
in the
|
||
<collectingCountry id="06B9765ED0448C589FBAF8F3E5A6B50F" box="[923,978,1813,1836]" name="United Kingdom" pageId="7" pageNumber="243">U.K.</collectingCountry>
|
||
The Specific epithet iS a patronym in the genitive caSe of a Latinized maSculine noun and iS correctly pronounced oh-Shay-eye [phonetic: əʊ-ʃɪə-aɪ].
|
||
</paragraph>
|
||
<paragraph id="7E1136CED0448C589CE6F8BBE594B5BB" blockId="7.[151,1437,1380,2017]" pageId="7" pageNumber="243">We propoSe the following colloquial nameS: O’Shea’S pipe Snake, Boano pipeSnake (E), Ular Pipa O’Shea, Ular Pipa Boano (I), O’Shea-WalzenSchlange, Boano-WalzenSchlange (G).</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="36B46545D0448C579CE6F842E2B0B38E" lastPageId="8" lastPageNumber="244" pageId="7" pageNumber="243" type="distribution">
|
||
<paragraph id="7E1136CED0448C579CE6F842E2BEB346" blockId="7.[151,1437,1380,2017]" lastBlockId="8.[151,1436,151,429]" lastPageId="8" lastPageNumber="244" pageId="7" pageNumber="243">
|
||
<emphasis id="4CDAEADCD0448C589CE6F842E714B59E" bold="true" box="[199,352,1956,1981]" pageId="7" pageNumber="243">Distribution.</emphasis>
|
||
<taxonomicName id="B9AE4D4DD0448C589D47F842E433B59E" authorityName="Kieckbusch & Mader & Kaiser & Mecke" authorityYear="2018" box="[358,583,1956,1981]" class="Reptilia" family="Cylindrophiidae" genus="Cylindrophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="243" phylum="Chordata" rank="species" species="osheai">
|
||
<emphasis id="4CDAEADCD0448C589D47F842E433B59E" box="[358,583,1956,1981]" italics="true" pageId="7" pageNumber="243">Cylindrophis osheai</emphasis>
|
||
</taxonomicName>
|
||
iS only known from Boano, an
|
||
<collectingCountry id="06B9765ED0448C589F87F842E59DB59E" box="[934,1001,1956,1981]" name="Iceland" pageId="7" pageNumber="243">iSland</collectingCountry>
|
||
with a land area of ca.
|
||
<quantity id="B9569B2BD0448C5898CBF842E34AB59E" box="[1258,1342,1956,1981]" metricMagnitude="5" metricUnit="m" metricValue="1.25" pageId="7" pageNumber="243" unit="km" value="125.0">125 km</quantity>
|
||
² (
|
||
<bibRefCitation id="1A3F4B3FD0448C589975F842E761B5C3" pageId="7" pageNumber="243" refId="ref7439" refString="Collar, N. J., Crosby, M. J. & Stattersfield, A. J. (1994) Birds to Watch 2: The World List of Threatened Birds. BirdLife International, Cambridge, 407 pp." type="book">
|
||
Collar
|
||
<emphasis id="4CDAEADCD0448C589CB6F82FE6BFB5C2" box="[151,203,1992,2017]" italics="true" pageId="7" pageNumber="243">et al</emphasis>
|
||
. 1994
|
||
</bibRefCitation>
|
||
) lying off the northweStern coaSt of Seram (
|
||
<figureCitation id="E6952A4BD0448C589F3CF82EE511B5C2" box="[797,869,1992,2017]" captionStart="FIGURE 6" captionStartId="8.[151,250,1725,1747]" captionTargetBox="[151,1436,483,1704]" captionTargetId="figure@8.[151,1436,483,1704]" captionTargetPageId="8" captionText="FIGURE 6. Distribution of Cylindrophis in the Moluccas, showing C. aruensis (triangle), C. boulengeri (diamond), C. melanotus (inverted triangles), C. osheai sp. nov. (open circle), and C. yamdena (closed circle). Base map modified from Wikipedia (© NordNordWest / Wikimedia Commons) by Max Kieckbusch." httpUri="https://zenodo.org/record/1437078/files/figure.png" pageId="7" pageNumber="243">Fig. 6</figureCitation>
|
||
). Although one of the three known SpecimenS iS labeled aS having the locality “
|
||
<collectingRegion id="BC6AF82CD04B8C579DDDFF71E42FB293" box="[508,603,151,176]" country="Indonesia" name="Sulawesi Utara" pageId="8" pageNumber="244">CelebeS</collectingRegion>
|
||
?” [=
|
||
<collectingRegion id="BC6AF82CD04B8C579EBDFF71E573B293" box="[668,775,151,176]" country="Indonesia" name="Sulawesi Utara" pageId="8" pageNumber="244">SulaweSi</collectingRegion>
|
||
], thiS notation waS made by E.M.J. JaSparS long after the original Specimen had been collected. JaSparS’ uSe of a queStion mark behind the locality in combination with the abSence of Similar SpecimenS from mainland
|
||
<collectingRegion id="BC6AF82CD04B8C579EBAFF39E574B2DB" box="[667,768,223,248]" country="Indonesia" name="Sulawesi Utara" pageId="8" pageNumber="244">SulaweSi</collectingRegion>
|
||
(33 SpecimenS from
|
||
<collectingRegion id="BC6AF82CD04B8C579FD2FF39E22CB2DB" box="[1011,1112,223,248]" country="Indonesia" name="Sulawesi Utara" pageId="8" pageNumber="244">SulaweSi</collectingRegion>
|
||
examined) indicate that thiS locality iS in error. IntereStingly, there are no recordS for thiS SpecieS from nearby Seram, an
|
||
<collectingCountry id="06B9765ED04B8C5798E8FEE2E37BB33E" box="[1225,1295,260,285]" name="Iceland" pageId="8" pageNumber="244">iSland</collectingCountry>
|
||
, on which a herpetofauna Survey waS relatively recently completed along itS northern coaSt (
|
||
<bibRefCitation id="1A3F4B3FD04B8C579834FEC1E375B363" box="[1045,1281,295,320]" pageId="8" pageNumber="244" refId="ref7527" refString="Edgar, P. W. & Lilley, R. P. H. (1993) Herpetofauna survey of Manusela National Park. In: Edwards, I. D., Macdonald, A. & Proctor, J. (Eds.) Natural History of Seram. Intercept Ltd., Hampshire, pp. 131 ‾ 141." type="journal article">Edgar & Lilley 1993</bibRefCitation>
|
||
), and where a Significant amount of collecting waS done in the paSt (e.g.,
|
||
<bibRefCitation id="1A3F4B3FD04B8C579F0CFEAAE5B2B346" box="[813,966,332,357]" pageId="8" pageNumber="244" refId="ref7238" refString="Bleeker, P. (1860 b) Over de Reptilien-Fauna van van [sic] Ceram. Natuurkundig Tijdschrift voor Nederlandsch Indie, 22, 35 - 38." type="journal article">Bleeker 1860</bibRefCitation>
|
||
; von
|
||
<bibRefCitation id="1A3F4B3FD04B8C579823FEAAE2C9B346" box="[1026,1213,332,357]" pageId="8" pageNumber="244" refId="ref8710" refString="von Rosenberg, C. B. H. (1867) Beschrijving van eenige gedeelten van Ceram. Tijdschrift voor Indische Taal-, Land- en Volkenkunde, 16, 97 - 182." type="journal article">RoSenberg 1867</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="7E1136CED04B8C579CE6FE89E2B0B38E" blockId="8.[151,1436,151,429]" pageId="8" pageNumber="244">
|
||
<emphasis id="4CDAEADCD04B8C579CE6FE89E7DFB3AB" bold="true" box="[199,427,367,392]" pageId="8" pageNumber="244">Red List Category.</emphasis>
|
||
Under IUCN criteria thiS SpecieS Should be liSted aS Data Deficient becauSe not enough information iS available about itS population Size, diStribution, or the impact of poSSible threatS.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |