436 lines
37 KiB
XML
436 lines
37 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.151.50538" ID-GBIF-Dataset="a62873a2-c4e1-4c6d-9baa-5c5fb94e7348" ID-PMC="PMC7329918" ID-Pensoft-Pub="1314-2003-151-107" ID-Pensoft-UUID="42D1EB2DF0545744B92BFCCCB395C45B" ID-PubMed="32655282" ModsDocID="1314-2003-151-107" checkinTime="1593036331120" checkinUser="pensoft" docAuthor="Sylvester, Steven P., Cuta-Alarcon, Lia E., Bravo-Pedraza, William J. & Soreng, Robert J." docDate="2020" docId="638D5321DB6B5F9EBE332D3FCBF41B79" docLanguage="en" docName="PhytoKeys 151: 107-160" docOrigin="PhytoKeys 151" docSource="http://dx.doi.org/10.3897/phytokeys.151.50538" docTitle="Agrostis perennans (Walter) Tuck. s. l., Amer. J. Sci. Arts 45: 44. 1843" docType="treatment" docVersion="4" id="42D1EB2DF0545744B92BFCCCB395C45B" lastPageNumber="107" masterDocId="42D1EB2DF0545744B92BFCCCB395C45B" masterDocTitle="Agrostis and Podagrostis (Agrostidinae, Poaceae) from paramos of Boyaca, Colombia: synoptic taxonomy including a key to Colombian species" masterLastPageNumber="160" masterPageNumber="107" pageNumber="107" updateTime="1668139837814" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Agrostis and Podagrostis (Agrostidinae, Poaceae) from paramos of Boyaca, Colombia: synoptic taxonomy including a key to Colombian species</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Sylvester, Steven P.</mods:namePart>
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<mods:affiliation>College of Biology and the Environment, Nanjing Forestry University, Long Pan Road No. 159, Nanjing, 210037, China & Royal Botanic Gardens, Kew, Richmond, Surrey, TW 9 3 AE, UK</mods:affiliation>
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<mods:nameIdentifier type="orcid">https://orcid.org/0000-0001-5577-8782</mods:nameIdentifier>
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<mods:nameIdentifier type="email">steven_sylvester@hotmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Cuta-Alarcon, Lia E.</mods:namePart>
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<mods:affiliation>Grupo Sistematica Biologica, Herbario UPTC, Escuela de Biologia, Facultad de Ciencias, Universidad Pedagogica y Tecnologica de Colombia, Avenida Central del Norte 39 - 115, Tunja-Boyaca, Colombia</mods:affiliation>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Bravo-Pedraza, William J.</mods:namePart>
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<mods:affiliation>Grupo Sistematica Biologica, Herbario UPTC, Escuela de Biologia, Facultad de Ciencias, Universidad Pedagogica y Tecnologica de Colombia, Avenida Central del Norte 39 - 115, Tunja-Boyaca, Colombia</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Soreng, Robert J.</mods:namePart>
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<mods:affiliation>Department of Botany, National Museum of Natural History, Smithsonian Institution, Washington DC 20560, USA</mods:affiliation>
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<mods:nameIdentifier type="orcid">https://orcid.org/0000-0002-8358-4915</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>PhytoKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2020</mods:date>
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<mods:detail type="volume">
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<mods:number>151</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>107</mods:start>
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<mods:end>160</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/phytokeys.151.50538</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/phytokeys.151.50538</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1314-2003-151-107</mods:identifier>
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<mods:identifier type="Pensoft-UUID">42D1EB2DF0545744B92BFCCCB395C45B</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="164730853" LSID="urn:lsid:plazi:treatment:638D5321DB6B5F9EBE332D3FCBF41B79" httpUri="http://treatment.plazi.org/id/638D5321DB6B5F9EBE332D3FCBF41B79" lastPageNumber="107" pageId="0" pageNumber="107">
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<subSubSection pageId="0" pageNumber="107" type="nomenclature">
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<paragraph pageId="0" pageNumber="107">
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<taxonomicName LSID="638D5321-DB6B-5F9E-BE33-2D3FCBF41B79" authority="(Walter) Tuck. s. l., Amer. J. Sci. Arts 45: 44. 1843" authorityName="(Walter) Tuck. s. l., Amer. J. Sci. Arts 45: 44." authorityYear="1843" class="Liliopsida" family="Poaceae" genus="Agrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Agrostis perennans" order="Poales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="perennans">Agrostis perennans (Walter) Tuck. s.l., Amer. J. Sci. Arts 45: 44. 1843</taxonomicName>
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<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Agrostis perennans s. l. A spikelet, glumes in lateral view, with floret in dorsal view, detached and raised slightly above the glumes B whole plant. Images of Vorontsova 2247 (FMB)." figureDoi="10.3897/phytokeys.151.50538.figure8" httpUri="https://binary.pensoft.net/fig/424846" pageId="0" pageNumber="107">Fig. 8</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="107" type="reference_group">
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<paragraph pageId="0" pageNumber="107">
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<taxonomicName authorityName="Walter" authorityYear="1788" class="Liliopsida" family="Poaceae" genus="Cornucopiae" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Cornucopiae perennans" order="Poales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="perennans">Cornucopiae perennans</taxonomicName>
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Walter, Fl. Carol. [Walter] 74. 1788.
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</paragraph>
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<paragraph pageId="0" pageNumber="107">
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<taxonomicName class="Liliopsida" family="Poaceae" genus="Agrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Agrostis cornucopiae" order="Poales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="cornucopiae">Agrostis cornucopiae</taxonomicName>
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Sm.,
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<normalizedToken originalValue="Gentleman’s">Gentleman's</normalizedToken>
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Mag. 59 (2): 873. 1789, nom. illeg. superfl., also Smith in Frasier, Short Hist. 2,pl. Nov. 25. 1789.
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</paragraph>
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<paragraph pageId="0" pageNumber="107">
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<taxonomicName class="Liliopsida" family="Poaceae" genus="Agrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Agrostis cornucopiae" order="Poales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="cornucopiae">Agrostis cornucopiae</taxonomicName>
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Lam., Tabl. Encycl. 161. 1791, nom. illeg. superfl.
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</paragraph>
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<paragraph pageId="0" pageNumber="107">
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<taxonomicName class="Liliopsida" family="Poaceae" genus="Agrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Agrostis elegans" order="Poales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="elegans">Agrostis elegans</taxonomicName>
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(Walter) Salisb., Prodr. Stirp. Chap. Allerton 25. 1796, nom. illeg. superfl.
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</paragraph>
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<paragraph pageId="0" pageNumber="107">
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<taxonomicName class="Liliopsida" family="Poaceae" genus="Agrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Agrostis anomala" order="Poales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="anomala">Agrostis anomala</taxonomicName>
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Willd., Sp. Pl. 1: 370. 1797, nom. illeg. superfl.
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</paragraph>
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<paragraph pageId="0" pageNumber="107">
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<taxonomicName class="Liliopsida" family="Poaceae" genus="Trichodium" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Trichodium decumbens" order="Poales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="decumbens">Trichodium decumbens</taxonomicName>
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(Walter) Michx., Fl. Bor.-Amer. (Michaux) 1: 42. 1803, nom. illeg. superfl.
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</paragraph>
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<paragraph pageId="0" pageNumber="107">
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<taxonomicName authorityName="Elliott" authorityYear="1816" baseAuthorityName="Walter" class="Liliopsida" family="Poaceae" genus="Trichodium" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Trichodium perennans" order="Poales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="perennans">Trichodium perennans</taxonomicName>
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(Walter) Elliott, Sketch Bot. S. Carolina [Elliott] 1: 99. 1816.
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</paragraph>
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<paragraph pageId="0" pageNumber="107">
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<taxonomicName authorityName="Alph.Wood" authorityYear="1861" baseAuthorityName="Walter" class="Liliopsida" family="Poaceae" genus="Agrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Agrostis scabra subsp. var. var. perennans" order="Poales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="variety" species="scabra" subSpecies="var." variety="perennans">Agrostis scabra var. perennans</taxonomicName>
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(Walter) Alph. Wood, Class-book Bot. (ed. 1861) 774. 1861.
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</paragraph>
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<paragraph pageId="0" pageNumber="107">
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=
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<taxonomicName class="Liliopsida" family="Poaceae" genus="Agrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Agrostis fasciculata" order="Poales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="fasciculata">Agrostis fasciculata</taxonomicName>
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(Kunth) Roem. & Schult., Syst. Veg., ed. 15 bis [Roemer & Schultes] 2: 362. 1817.
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<taxonomicName authorityName="Kunth" authorityYear="1816" class="Liliopsida" family="Poaceae" genus="Vilfa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Vilfa fasciculata" order="Poales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="fasciculata">Vilfa fasciculata</taxonomicName>
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Kunth, Nov. Gen. Sp. [H.B.K.] 1: 139. 1816. Type: Ecuador. Pichincha, May, F.W.H.A. von Humboldt & A.J.A. Bonpland s.n. (holotype: P (P00669402 [image!]; isotypes: HAL (HAL0106915 [image!]), LE-TRIN (LE-TRIN1610.01 fragm. ex P), P (P00740549 [image!]; P00740548 [image!]), US (US556249 fragm. ex P)).
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</paragraph>
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<paragraph pageId="0" pageNumber="107">
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=
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<taxonomicName class="Liliopsida" family="Poaceae" genus="Agrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Agrostis humboldtiana" order="Poales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="humboldtiana">Agrostis humboldtiana</taxonomicName>
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Steud., Nomencl. Bot. [Steudel], ed. 2. i. 40. 1840.
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<taxonomicName class="Liliopsida" family="Poaceae" genus="Agrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Agrostis pulchella" order="Poales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="pulchella">Agrostis pulchella</taxonomicName>
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Kunth, Enum. Pl. 1: 223. 1833, hom. illeg., non Roth, 1817, nom. superfl.
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<taxonomicName class="Liliopsida" family="Poaceae" genus="Agrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Agrostis elegans" order="Poales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="elegans">Agrostis elegans</taxonomicName>
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(Kunth) Roem. & Schult., Syst. Veg., ed. 15 bis [Roemer & Schultes] 2: 362. 1817, hom. illeg., non. Salisb. 1796.
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<taxonomicName authorityName="Kunth" authorityYear="1816" class="Liliopsida" family="Poaceae" genus="Vilfa" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Vilfa elegans" order="Poales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="elegans">Vilfa elegans</taxonomicName>
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Kunth, Nov. Gen. Sp. [H.B.K.] 1: 139. 1816. Type: Ecuador. [Crescit in planitie Cochapamba, in regione temperata regni Quitensis, alt. 1340 hexap. Floret Majo] Herbier de Amerique equatoriale, A.J.A. Bonpland & F.W.H.A. von Humboldt 3010 (holotype: P (P00669401 [image!]); isotypes: BAA (BAA-Col. typus 4256), BM (BM000938529 [image!]; BM000938530 [image!]), LE-TRIN (LE-TRIN1644.01 ex hrbr. Humb.), P (P00740586 [image!]; P00740588 [image!]; P00740587 [image!])).
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</paragraph>
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<paragraph pageId="0" pageNumber="107">
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=
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<taxonomicName class="Liliopsida" family="Poaceae" genus="Agrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Agrostis weberbaueri" order="Poales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="weberbaueri">Agrostis weberbaueri</taxonomicName>
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Mez, Repert. Spec. Nov. Regni Veg. 18 (1-3): 1. 1922. Type: PERU. Huacapistana and Monson, Weberbauer s.n. (holotype: [not found]; isotypes: [not found]). Many other heterotypic synonyms.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="107" type="materials_examined">
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<paragraph pageId="0" pageNumber="107">Type.</paragraph>
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<paragraph pageId="0" pageNumber="107">
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USA. South Carolina: Richmond County, Fort Jackson Military Reservation, found at the intersection of Cut Off Rd. and Fire Break 49, 11 July 1995, K.B. Kelly, Jr. & J.B. Nelson 254 (neotype, designated by
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<bibRefCitation author="Ward, DB" journalOrPublisher="Journal of the Botanical Research Institute of Texas" pageId="0" pageNumber="107" pagination="1091 - 1100" refId="B40" refString="Ward, DB, 2007. Thomas Walter typification project, IV: Neotypes and epitypes for 43 Walter names, of genera A through C. Journal of the Botanical Research Institute of Texas 1 (2): 1091 - 1100" title="Thomas Walter typification project, IV: Neotypes and epitypes for 43 Walter names, of genera A through C." volume="1" year="2007">Ward (2007</bibRefCitation>
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: 1099): GH (GH00247994 [image!])).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="107" type="description">
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<paragraph pageId="0" pageNumber="107">Description.</paragraph>
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<paragraph pageId="0" pageNumber="107">
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<emphasis bold="true" pageId="0" pageNumber="107">Perennial herbs</emphasis>
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, laxly to densely tufted, sometimes stooling or with short ascending pseudostolons that have the appearance of rhizomes on herbarium sheets.
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<emphasis bold="true" pageId="0" pageNumber="107">Tillers</emphasis>
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extravaginal, with cataphylls present.
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<emphasis bold="true" pageId="0" pageNumber="107">Culms</emphasis>
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(21-)33-64(-100) cm tall, erect or decumbent to subgeniculate at their base, delicate to fairly firm, with 0-2(-3) nodes exerted at flowering, smooth to rarely scaberulous.
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<emphasis bold="true" pageId="0" pageNumber="107">Leaves</emphasis>
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mainly basal early in the flowering season but tending to become mostly cauline with maturity, glabrous, smooth or scaberulous;
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<emphasis bold="true" pageId="0" pageNumber="107">ligules</emphasis>
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0.5-5(-7) mm long, of basal leaves and tillers 0.5-2.5 mm long, of upper culm 3-5(-7) mm long, truncate to obtuse-triangular, slightly to strongly decurrent with the sheath, abaxial surface scabrous to scaberulous, rarely smooth;
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<emphasis bold="true" pageId="0" pageNumber="107">blades</emphasis>
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(3-)6-15 cm long, (1-)1.5-3.5(-5) mm wide, usually flat or conduplicate and lax to slightly firm, sometimes involute and rigid in the basal leaves, smooth throughout or scaberulous on margins and sometimes surfaces, apices acute.
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<emphasis bold="true" pageId="0" pageNumber="107">Panicles</emphasis>
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(3.5-)10-22 cm long, 2-11 cm wide, open, lax, ovoid to pyramidal, slightly to usually greatly exerted from the basal foliage, lateral branches without spikelets near their base and for a large distance, long, ascending, spreading, to somewhat divergent and not held close to the central inflorescence axis, central axis and panicle branches smooth or lightly scaberulous;
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<emphasis bold="true" pageId="0" pageNumber="107">pedicels</emphasis>
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1-4.5 mm long, usually longer than their spikelets, not or slightly dilated at their apex, smooth or lightly scaberulous.
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<emphasis bold="true" pageId="0" pageNumber="107">Spikelets</emphasis>
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(not including awn, if present) (1.5-)1.8-2.5(-3.2) mm long;
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<emphasis bold="true" pageId="0" pageNumber="107">glumes</emphasis>
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subequal, 1-veined, keels scabrous in the distal
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<normalizedToken originalValue="½">1/2</normalizedToken>
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-1/3, apices acute;
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<emphasis bold="true" pageId="0" pageNumber="107">floret</emphasis>
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usually 1/2-2/3 the length of the glumes, rarely longer;
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<emphasis bold="true" pageId="0" pageNumber="107">calluses</emphasis>
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lightly pilose with 2 sparse lateral tufts of short hairs;
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<emphasis bold="true" pageId="0" pageNumber="107">lemmas</emphasis>
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1.5-2 mm long, glabrous, smooth, 5-veined, apex acute to more-or-less truncate, denticulate, muticous, mucronate or exceptionally with a short awn to 0.5 mm long (to 1.4-1.9 mm long in Argentina;
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<bibRefCitation author="Rugolo de Agrasar, ZE" editor="Zuloaga, FO" journalOrPublisher="Poaceae: Pooideae (Vol. 3 Tomo 2). IBODA-IMBIV, CONICET, Buenos Aires, Argentina" pageId="0" pageNumber="107" pagination="102 - 125" refId="B30" refString="Rugolo de Agrasar, ZE, 2012. Agrostis L. In: Zuloaga, FO, Rugolo de Agrasar, ZE, Anton, AM, Eds., Flora Argentina. Poaceae: Pooideae (Vol. 3 Tomo 2). IBODA-IMBIV, CONICET, Buenos Aires, Argentina: 102 - 125" title="Agrostis L." volumeTitle="Flora Argentina." year="2012">
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<normalizedToken originalValue="Rúgolo">Rugolo</normalizedToken>
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de Agrasar 2012
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</bibRefCitation>
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), inserted in the upper (
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<normalizedToken originalValue="½">1/2</normalizedToken>
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) 1/3 of the dorsal keel, usually not or rarely only slightly surpassing the glumes, straight, not twisted, weak and falling easily;
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<emphasis bold="true" pageId="0" pageNumber="107">paleas</emphasis>
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absent or to 0.5 mm long, usually <
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<normalizedToken originalValue="¼">1/4</normalizedToken>
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the length of the lemma;
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<emphasis bold="true" pageId="0" pageNumber="107">rachilla</emphasis>
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absent;
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<emphasis bold="true" pageId="0" pageNumber="107">anthers</emphasis>
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0.7-1 mm long.
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</paragraph>
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<caption doi="10.3897/phytokeys.151.50538.figure8" httpUri="https://binary.pensoft.net/fig/424846" pageId="0" pageNumber="107" start="Figure 8" startId="F8">
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<paragraph pageId="0" pageNumber="107">
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<emphasis bold="true" pageId="0" pageNumber="107">Figure 8.</emphasis>
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<taxonomicName class="Liliopsida" family="Poaceae" genus="Agrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Agrostis perennans" order="Poales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="perennans">
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<emphasis italics="true" pageId="0" pageNumber="107">Agrostis perennans</emphasis>
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</taxonomicName>
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s.l.
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<emphasis bold="true" pageId="0" pageNumber="107">A</emphasis>
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spikelet, glumes in lateral view, with floret in dorsal view, detached and raised slightly above the glumes
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<emphasis bold="true" pageId="0" pageNumber="107">B</emphasis>
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whole plant. Images of Vorontsova 2247 (FMB).
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="107" type="distribution">
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<paragraph pageId="0" pageNumber="107">Distribution and ecology.</paragraph>
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<paragraph pageId="0" pageNumber="107">Stretching from Alaska, Canada and USA, through Central America and the Caribbean, to Argentina and Chile of South America. The species has an exceptionally large ecological amplitude, being found in maritime dunes, wetlands, and grasslands from sea level to> 4000 m alt.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="107" type="materials_examined">
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<paragraph pageId="0" pageNumber="107">Other specimens examined.</paragraph>
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<paragraph pageId="0" pageNumber="107">
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<emphasis bold="true" pageId="0" pageNumber="107">Colombia</emphasis>
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.
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<emphasis bold="true" pageId="0" pageNumber="107">
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<normalizedToken originalValue="Boyacá">Boyaca</normalizedToken>
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</emphasis>
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: Munic.
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<normalizedToken originalValue="Belén">Belen</normalizedToken>
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,
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<normalizedToken originalValue="Páramo">Paramo</normalizedToken>
|
||
de La Rusia, near
|
||
<normalizedToken originalValue="Páramo">Paramo</normalizedToken>
|
||
El Consuelo, unprotected private land, somewhat disturbed
|
||
<normalizedToken originalValue="páramo">paramo</normalizedToken>
|
||
grazed by horses and rodents, with
|
||
<taxonomicName authorityName="Cuatrec" authorityYear="1973" class="Magnoliopsida" family="Asteraceae" genus="Espeletia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Espeletia boyacensis" order="Asterales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="boyacensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="107">Espeletia boyacensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName lsidName="E. discoides" pageId="0" pageNumber="107" rank="species" species="discoides">
|
||
<emphasis italics="true" pageId="0" pageNumber="107">E. discoides</emphasis>
|
||
</taxonomicName>
|
||
, near path,
|
||
<geoCoordinate degrees="6.02415" direction="north" orientation="latitude" precision="1" value="6.02415">6.02415N</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="72.57289" direction="west" orientation="longitude" precision="1" value="-72.57289">72.57289W</geoCoordinate>
|
||
, 3831, 22 Nov. 2017, M. Vorontsova 2247 (FMB, K, SI, UPTC, US); Munic. Duitama,
|
||
<normalizedToken originalValue="Páramo">Paramo</normalizedToken>
|
||
de Agueros. junto a la casa, muy perturbado, pastizales junto a la
|
||
<normalizedToken originalValue="plantación">plantacion</normalizedToken>
|
||
de pinos,
|
||
<geoCoordinate degrees="8.91069" direction="north" orientation="latitude" precision="1" value="8.91069">8.91069N</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="73.07219" direction="west" orientation="longitude" precision="1" value="-73.07219">73.07219W</geoCoordinate>
|
||
, 3377 m alt., 28 Oct. 2017, S.P. Sylvester 3039 (FMB, K, SI, UPTC, US); Munic. Duitama,
|
||
<normalizedToken originalValue="Páramo">Paramo</normalizedToken>
|
||
de La Rusia, Guanenta Alto
|
||
<normalizedToken originalValue="Río">Rio</normalizedToken>
|
||
Fonce National Park, top of the ridge
|
||
<normalizedToken originalValue="Peña">Pena</normalizedToken>
|
||
Negra just below military base, ridge along the top of a steep rocky landscape,
|
||
<geoCoordinate degrees="5.58389" direction="north" orientation="latitude" precision="1" value="5.58389">5.58389N</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="73.053263" direction="west" orientation="longitude" precision="1" value="-73.05326">73.053263W</geoCoordinate>
|
||
, 3970 m alt., open
|
||
<normalizedToken originalValue="Páramo">Paramo</normalizedToken>
|
||
with
|
||
<taxonomicName class="Magnoliopsida" family="Asteraceae" genus="Espeletia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Espeletia cachaluensis" order="Asterales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="cachaluensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="107">Espeletia cachaluensis</emphasis>
|
||
</taxonomicName>
|
||
, no signs of grazing, 21 Nov. 2017, M. Vorontsova 2203 (US); Munic. Duitama,
|
||
<normalizedToken originalValue="Páramo">Paramo</normalizedToken>
|
||
de la Rusia, Vereda El Carmen,
|
||
<geoCoordinate degrees="5.95333" direction="north" orientation="latitude" precision="1" value="5.95333">5.95333N</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="73.11019" direction="west" orientation="longitude" precision="1" value="-73.11019">73.11019W</geoCoordinate>
|
||
, 3445 m alt., 30 Oct. 2017, S.P. Sylvester 3064 (FMB, K, SI, US); Munic. Duitama,
|
||
<normalizedToken originalValue="Páramo">Paramo</normalizedToken>
|
||
de la Rusia,
|
||
<normalizedToken originalValue="vía">via</normalizedToken>
|
||
que conduce a vereda de
|
||
<normalizedToken originalValue="Avendaños">Avendanos</normalizedToken>
|
||
,
|
||
<geoCoordinate degrees="5.95011" direction="north" orientation="latitude" precision="1" value="5.95011">5.95011N</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="73.09097" direction="west" orientation="longitude" precision="1" value="-73.09097">73.09097W</geoCoordinate>
|
||
, 3795 m alt., 1 Oct. 2017, S.P. Sylvester 3002 (COL, FMB, K, SI, US);
|
||
<geoCoordinate degrees="5.93246" direction="north" orientation="latitude" precision="1" value="5.93246">5.93246N</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="73.0798" direction="west" orientation="longitude" precision="5" value="-73.0798">73.0798W</geoCoordinate>
|
||
, 3726 m alt., 4 Oct. 2017, S.P. Sylvester 3036 (COL, K, SI, UPTC, US); Munic. Sotaquira, protected area of
|
||
<normalizedToken originalValue="Páramo">Paramo</normalizedToken>
|
||
El Valle,
|
||
<normalizedToken originalValue="páramo">paramo</normalizedToken>
|
||
just above riverine forest,
|
||
<geoCoordinate degrees="5.4605" direction="north" orientation="latitude" precision="5" value="5.4605">5.4605N</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="73.21507" direction="west" orientation="longitude" precision="1" value="-73.21507">73.21507W</geoCoordinate>
|
||
, 3220 m alt., 17 Nov. 2017, M. Vorontsova 2161 (K, SI, US); Sierra Nevada del Cocuy, Alto Valle Lagunillas,
|
||
<normalizedToken originalValue="páramo">paramo</normalizedToken>
|
||
pantanoso al sur de la Laguna Cuadrada, [
|
||
<geoCoordinate degrees="6.3619" direction="north" orientation="latitude" precision="5" value="6.3619">6.3619N</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="72.3345" direction="west" orientation="longitude" precision="5" value="-72.3345">72.3345W</geoCoordinate>
|
||
], 4060 m alt., 26 Sep. 1972, A.M. Cleef 5579 (US2785750).
|
||
<emphasis bold="true" pageId="0" pageNumber="107">
|
||
<normalizedToken originalValue="Nariño">Narino</normalizedToken>
|
||
</emphasis>
|
||
:
|
||
<normalizedToken originalValue="Páramo">Paramo</normalizedToken>
|
||
del
|
||
<normalizedToken originalValue="Tábano">Tabano</normalizedToken>
|
||
, alto de la Cordillera, entre Pasto y El Escano, vertiente occidental, [
|
||
<geoCoordinate degrees="1.1756" direction="north" orientation="latitude" precision="5" value="1.1756">1.1756N</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="77.1867" direction="west" orientation="longitude" precision="5" value="-77.1867">77.1867W</geoCoordinate>
|
||
], 3200 m alt., 11 Jan. 1941, J. Cuatrecasas 11908-A (US1798782).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="107" type="notes">
|
||
<paragraph pageId="0" pageNumber="107">Notes.</paragraph>
|
||
<paragraph pageId="0" pageNumber="107">
|
||
<taxonomicName class="Liliopsida" family="Poaceae" genus="Agrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Agrostis perennans" order="Poales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="perennans">
|
||
<emphasis italics="true" pageId="0" pageNumber="107">Agrostis perennans</emphasis>
|
||
</taxonomicName>
|
||
appears to be a
|
||
<normalizedToken originalValue="‘grab-bag’">'grab-bag'</normalizedToken>
|
||
of many different taxa that come out in many different places amongst other taxa of
|
||
<taxonomicName class="Liliopsida" family="Poaceae" genus="Agrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Agrostis" order="Poales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="107">Agrostis</emphasis>
|
||
</taxonomicName>
|
||
in molecular phylogenies (Konstantin Romaschenko pers. communication), with the species complex needing urgent revision. Specimens found in
|
||
<normalizedToken originalValue="Boyacán">Boyacan</normalizedToken>
|
||
<normalizedToken originalValue="páramos">paramos</normalizedToken>
|
||
correspond in almost all characteristics with
|
||
<taxonomicName class="Liliopsida" family="Poaceae" genus="Agrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Agrostis perennans" order="Poales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="perennans">
|
||
<emphasis italics="true" pageId="0" pageNumber="107">Agrostis perennans</emphasis>
|
||
</taxonomicName>
|
||
s.s. apart from sometimes being found with shorter spikelets (c. 1.5-2 mm long) and, what appear to be, short ascending rhizomes/pseudostolons, with these characteristics also found on type material of
|
||
<taxonomicName lsidName="A. fasciculata" pageId="0" pageNumber="107" rank="species" species="fasciculata">
|
||
<emphasis italics="true" pageId="0" pageNumber="107">A. fasciculata</emphasis>
|
||
</taxonomicName>
|
||
, which is here considered a synonym of
|
||
<taxonomicName lsidName="A. perennans" pageId="0" pageNumber="107" rank="species" species="perennans">
|
||
<emphasis italics="true" pageId="0" pageNumber="107">A. perennans</emphasis>
|
||
</taxonomicName>
|
||
until comprehensive systematic research is undertaken on the species complex. Cataphyllous extravaginal shoots are found on the neotype of
|
||
<taxonomicName lsidName="A. perennans" pageId="0" pageNumber="107" rank="species" species="perennans">
|
||
<emphasis italics="true" pageId="0" pageNumber="107">A. perennans</emphasis>
|
||
</taxonomicName>
|
||
as well as other type material of species considered synonyms of
|
||
<taxonomicName lsidName="A. perennans" pageId="0" pageNumber="107" rank="species" species="perennans">
|
||
<emphasis italics="true" pageId="0" pageNumber="107">A. perennans</emphasis>
|
||
</taxonomicName>
|
||
, e.g.
|
||
<taxonomicName lsidName="A. decumbens" pageId="0" pageNumber="107" rank="species" species="decumbens">
|
||
<emphasis italics="true" pageId="0" pageNumber="107">A. decumbens</emphasis>
|
||
</taxonomicName>
|
||
E. Fourn., with it plausible that these
|
||
<normalizedToken originalValue="‘rhizomes’">'rhizomes'</normalizedToken>
|
||
are short pseudostolons that arise from growing through moss or between rocks.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="107">
|
||
Type specimens and specimens from North America, Central America, and northwest South America have lemmas without awns or with awns under-developed, to 0.5 mm long, straight and easily falling. Specimens noted by
|
||
<bibRefCitation author="Rugolo de Agrasar, ZE" editor="Zuloaga, FO" journalOrPublisher="Poaceae: Pooideae (Vol. 3 Tomo 2). IBODA-IMBIV, CONICET, Buenos Aires, Argentina" pageId="0" pageNumber="107" pagination="102 - 125" refId="B30" refString="Rugolo de Agrasar, ZE, 2012. Agrostis L. In: Zuloaga, FO, Rugolo de Agrasar, ZE, Anton, AM, Eds., Flora Argentina. Poaceae: Pooideae (Vol. 3 Tomo 2). IBODA-IMBIV, CONICET, Buenos Aires, Argentina: 102 - 125" title="Agrostis L." volumeTitle="Flora Argentina." year="2012">
|
||
<normalizedToken originalValue="Rúgolo">Rugolo</normalizedToken>
|
||
de Agrasar (2012
|
||
</bibRefCitation>
|
||
: 119) to have awns to 1.4 mm long, or exceptionally to 1.9 mm long, may be a distinct taxon, but further research is needed to ascertain this.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="107" type="similar species">
|
||
<paragraph pageId="0" pageNumber="107">Similar species.</paragraph>
|
||
<paragraph pageId="0" pageNumber="107">
|
||
<taxonomicName class="Liliopsida" family="Poaceae" genus="Agrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Agrostis mertensii" order="Poales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="mertensii">
|
||
<emphasis italics="true" pageId="0" pageNumber="107">Agrostis mertensii</emphasis>
|
||
</taxonomicName>
|
||
, which is principally differentiated in having a well-developed, flexuose to geniculate, twisted awn (1.8-)2.5-4.5 mm long, inserted in the middle or lower third of the lemma, and which is greatly exerted from the glumes (vs. unawned or exceptionally with a short straight awn to 0.5 mm long, inserted in the upper (
|
||
<normalizedToken originalValue="½">1/2</normalizedToken>
|
||
) third of the dorsal keel, usually not or rarely only slightly surpassing the glumes in
|
||
<taxonomicName lsidName="A. perennans" pageId="0" pageNumber="107" rank="species" species="perennans">
|
||
<emphasis italics="true" pageId="0" pageNumber="107">A. perennans</emphasis>
|
||
</taxonomicName>
|
||
s.l. specimens from northwest South America, Central America and North America).
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="107">
|
||
<taxonomicName class="Liliopsida" family="Poaceae" genus="Agrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Agrostis subrepens" order="Poales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="subrepens">
|
||
<emphasis italics="true" pageId="0" pageNumber="107">Agrostis subrepens</emphasis>
|
||
</taxonomicName>
|
||
, here considered to be excluded from Departamento
|
||
<normalizedToken originalValue="Boyacá">Boyaca</normalizedToken>
|
||
and possibly Colombia (see notes under 'Excluded
|
||
<normalizedToken originalValue="species’">species'</normalizedToken>
|
||
<taxonomicName lsidName="A. subrepens" pageId="0" pageNumber="107" rank="species" species="subrepens">
|
||
<emphasis italics="true" pageId="0" pageNumber="107">A. subrepens</emphasis>
|
||
</taxonomicName>
|
||
), sometimes bears certain similar, but not equal, characteristics such as a) plants often stooling with notable pseudostolons and appearing long rhizomatous on herbarium sheets; b) culms 50-100 cm tall, slightly creeping or decumbent at their base but erect towards their apex; c) panicles open, lax, 5-10 cm wide; d) lemma unawned or awn to 0.5 mm long and straight, not twisted, inserted in the middle or upper 1/3 of the lemma or subapical; e) palea absent or <
|
||
<normalizedToken originalValue="¼">1/4</normalizedToken>
|
||
the length of the lemma.
|
||
<taxonomicName class="Liliopsida" family="Poaceae" genus="Agrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Agrostis subrepens" order="Poales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="subrepens">
|
||
<emphasis italics="true" pageId="0" pageNumber="107">Agrostis subrepens</emphasis>
|
||
</taxonomicName>
|
||
can be differentiated from
|
||
<taxonomicName lsidName="A. perennans" pageId="0" pageNumber="107" rank="species" species="perennans">
|
||
<emphasis italics="true" pageId="0" pageNumber="107">A. perennans</emphasis>
|
||
</taxonomicName>
|
||
s.l. by a) upper culm ligules 1-2.2 mm long, obtuse (vs. 3-5(-7) mm long, truncate to obtuse-triangular in
|
||
<taxonomicName lsidName="A. perennans" pageId="0" pageNumber="107" rank="species" species="perennans">
|
||
<emphasis italics="true" pageId="0" pageNumber="107">A. perennans</emphasis>
|
||
</taxonomicName>
|
||
s.l.) b) leaf blades 0.5-1 mm wide in diameter as rolled or folded, convolute, involute, or conduplicate, rigid, surfaces scabrous (vs. leaf blades (1-)1.5-3.5(-5) mm wide, usually flat or conduplicate and lax to slightly firm, sometimes involute and rigid in the basal leaves, smooth throughout or scaberulous on margins and sometimes surfaces in
|
||
<taxonomicName lsidName="A. perennans" pageId="0" pageNumber="107" rank="species" species="perennans">
|
||
<emphasis italics="true" pageId="0" pageNumber="107">A. perennans</emphasis>
|
||
</taxonomicName>
|
||
s.l.); anthers 1-1.3 mm long (vs. 0.7-1 mm long in
|
||
<taxonomicName lsidName="A. perennans" pageId="0" pageNumber="107" rank="species" species="perennans">
|
||
<emphasis italics="true" pageId="0" pageNumber="107">A. perennans</emphasis>
|
||
</taxonomicName>
|
||
s.l.)
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="107">
|
||
<taxonomicName class="Liliopsida" family="Poaceae" genus="Agrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Agrostis turrialbae" order="Poales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="turrialbae">
|
||
<emphasis italics="true" pageId="0" pageNumber="107">Agrostis turrialbae</emphasis>
|
||
</taxonomicName>
|
||
, here considered to be excluded from Departamento
|
||
<normalizedToken originalValue="Boyacá">Boyaca</normalizedToken>
|
||
and possibly Colombia, bears similarities and may possibly be part of the
|
||
<taxonomicName lsidName="A. perennans" pageId="0" pageNumber="107" rank="species" species="perennans">
|
||
<emphasis italics="true" pageId="0" pageNumber="107">A. perennans</emphasis>
|
||
</taxonomicName>
|
||
s.l. species complex (see notes under 'Excluded
|
||
<normalizedToken originalValue="species’">species'</normalizedToken>
|
||
<taxonomicName lsidName="A. turrialbae" pageId="0" pageNumber="107" rank="species" species="turrialbae">
|
||
<emphasis italics="true" pageId="0" pageNumber="107">A. turrialbae</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="107">
|
||
<taxonomicName class="Liliopsida" family="Poaceae" genus="Agrostis" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Agrostis vinealis" order="Poales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="vinealis">
|
||
<emphasis italics="true" pageId="0" pageNumber="107">Agrostis vinealis</emphasis>
|
||
</taxonomicName>
|
||
, of Eurasian origin which is not recorded for Colombia, but found introduced and naturalized in Argentina and Chile (
|
||
<bibRefCitation author="Soreng, RJ" editor="Soreng, RJ" journalOrPublisher="Contributions from the United States National Herbarium" pageId="0" pageNumber="107" pagination="42 - 89" refId="B33" refString="Soreng, RJ, Peterson, PM, 2003. Agrostis. In: Soreng, RJ, Peterson, PM, Davidse, G, Judziewicz, EJ, Zuloaga, FO, Filguieras, TS, Morrone, O, Eds., Catalogue of New World Grasses (Poaceae): IV. Subfamily Pooideae. Contributions from the United States National Herbarium 48: 42 - 89" title="Agrostis." volume="48" volumeTitle="Catalogue of New World Grasses (Poaceae): IV. Subfamily Pooideae." year="2003">Soreng and Peterson 2003</bibRefCitation>
|
||
;
|
||
<bibRefCitation author="Rugolo de Agrasar, ZE" editor="Zuloaga, FO" journalOrPublisher="Poaceae: Pooideae (Vol. 3 Tomo 2). IBODA-IMBIV, CONICET, Buenos Aires, Argentina" pageId="0" pageNumber="107" pagination="102 - 125" refId="B30" refString="Rugolo de Agrasar, ZE, 2012. Agrostis L. In: Zuloaga, FO, Rugolo de Agrasar, ZE, Anton, AM, Eds., Flora Argentina. Poaceae: Pooideae (Vol. 3 Tomo 2). IBODA-IMBIV, CONICET, Buenos Aires, Argentina: 102 - 125" title="Agrostis L." volumeTitle="Flora Argentina." year="2012">
|
||
<normalizedToken originalValue="Rúgolo">Rugolo</normalizedToken>
|
||
de Agrasar 2012
|
||
</bibRefCitation>
|
||
), bears certain similarities (see also comments regarding this species under
|
||
<taxonomicName lsidName="A. imberbis" pageId="0" pageNumber="107" rank="species" species="imberbis">A. cf. imberbis</taxonomicName>
|
||
). It can be differentiated by having a) lemmas usually with a persistent, geniculate and twisted awn to 4 mm long, inserted near the base (sometimes absent); b) rhizomes conspicuous, long, lateral tending, and usually covered in bracts; c) panicles contracted and rather dense before and after flowering.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |