159 lines
18 KiB
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159 lines
18 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.155.1779" ID-GBIF-Dataset="f8c61767-8004-48b6-bbdc-c9a86455d930" ID-PMC="PMC3238047" ID-Pensoft-Pub="1313-2970-155-1" ID-PubMed="22291510" ModsDocAuthor="" ModsDocDate="2011" ModsDocID="1313-2970-155-1" ModsDocOrigin="ZooKeys 155" ModsDocTitle="Review of subtribe Singilina Jeannel, 1949, of the Middle East and Central Asia (Coleoptera, Carabidae, Lebiini)" checkinTime="1451249489653" checkinUser="pensoft" docAuthor="Anichtchenko, Alexander" docDate="2011" docId="6379CBBB4CEDD07683EC1D27339329D3" docLanguage="en" docName="ZooKeys 155: 1-50" docOrigin="ZooKeys 155" docSource="http://dx.doi.org/10.3897/zookeys.155.1779" docTitle="Singilis mesopotamicus Pic 1901" docType="treatment" docVersion="3" lastPageNumber="17" masterDocId="FFC0FFCFFFBDFF9F7458F904FF88FFFB" masterDocTitle="Review of subtribe Singilina Jeannel, 1949, of the Middle East and Central Asia (Coleoptera, Carabidae, Lebiini)" masterLastPageNumber="50" masterPageNumber="1" pageNumber="15" updateTime="1668152993135" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Review of subtribe Singilina Jeannel, 1949, of the Middle East and Central Asia (Coleoptera, Carabidae, Lebiini)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Anichtchenko, Alexander</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2011</mods:date>
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<mods:detail type="volume">
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<mods:number>155</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>50</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.155.1779</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.155.1779</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-155-1</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="152032655" LSID="urn:lsid:plazi:treatment:6379CBBB4CEDD07683EC1D27339329D3" httpUri="http://treatment.plazi.org/id/6379CBBB4CEDD07683EC1D27339329D3" lastPageId="16" lastPageNumber="17" pageId="14" pageNumber="15">
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<subSubSection pageId="14" pageNumber="15" type="nomenclature">
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<paragraph pageId="14" pageNumber="15">
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<taxonomicName LSID="http://species-id.net/wiki/Singilis_mesopotamicus" authority="Pic, 1901" authorityName="Pic" authorityYear="1901" class="Insecta" family="Carabidae" genus="Singilis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Singilis mesopotamicus" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="mesopotamicus">Singilis mesopotamicus Pic, 1901</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="14" pageNumber="15" type="reference_group">
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<paragraph pageId="14" pageNumber="15">
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<taxonomicName class="Insecta" family="Carabidae" genus="Singilis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Singilis (Phloeozeteus) plagiata" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="plagiata" subGenus="Phloeozeteus">Singilis (Phloeozeteus) plagiata</taxonomicName>
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var. mesopotamica
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<bibRefCitation author="Pic, M" journalOrPublisher="L'Echange, Revue Linneenne" pageId="35" pageNumber="36" pagination="89 - 91" title="Coleopteres nouveaux d'Orient, Armenie et Tunisie." volume="204" year="1901">Pic 1901</bibRefCitation>
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: 89
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</paragraph>
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<paragraph pageId="14" pageNumber="15">
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<taxonomicName class="Insecta" family="Carabidae" genus="Singilis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Singilis (Phloezetaeus) apicalis" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="apicalis" subGenus="Phloezetaeus">Singilis (Phloezetaeus) apicalis</taxonomicName>
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<bibRefCitation pageId="14" pageNumber="15">
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<normalizedToken originalValue="Jedlička">Jedlicka</normalizedToken>
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1956
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</bibRefCitation>
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: 204, syn. n. (synonymy presumed; type of
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<taxonomicName class="Insecta" family="Carabidae" genus="Singilis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Singilis mesopotamicus" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="mesopotamicus">Singilis mesopotamicus</taxonomicName>
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Pic, 1901, unavailable).
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</paragraph>
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<paragraph pageId="14" pageNumber="15">
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<taxonomicName class="Insecta" family="Carabidae" genus="Phloeozetus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Phloeozetus plagiatius subsp. mesopotamicus" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="subSpecies" species="plagiatius" subSpecies="mesopotamicus">Phloeozetus plagiatius mesopotamicus</taxonomicName>
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:
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<bibRefCitation pageId="14" pageNumber="15">Kabak 2003</bibRefCitation>
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: 438
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15" type="material examined">
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<paragraph pageId="14" pageNumber="15">Material examined.</paragraph>
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<paragraph lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15">
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AFGHANISTAN: S Jalalabad, S Agam 2000 m, 23.XII.1970, Kabakov leg. (1♀, ZIN); 20 km NO Kabul, 2000m, 10.IV.1970 Kabakov (1♂ 3♀♀, ZIN)
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<pageBreakToken pageId="15" pageNumber="16" start="start">;</pageBreakToken>
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same locality, 15.IV.1971 (1♂, ZIN); Ghazni W Moqur 2300m, 5.X.1972, Kabakov (1♂ 2♀♀, ZIN); same locality, 11.IX.1972 (1♀, ZIN); Kondahar, Ghbargay 2000 m, 20.III.1973, Kabakov (2♀♀, ZIN); Ghor Saghar 2500 m, 14.VIII.1970 (1♂, ZIN); Heart, NW Adraskan 26.X.1971 (1♂, ZIN); same locality, 20.XI.1071 (1♀, ZIN). IRAN: Chaharmahal-va-Bakhuyari prov., 10 km E Chaman Goli, 7-8.VI.2008, 3500m, Anichtchenko A. leg. (2♂♂, AAC); Kerman, Qohrud mts., 10 km E Korin, 3500m, 13.V.2007, Anichtchenko A. leg. (2♀♀, AAC); S Iran, Kerman prov., Balt area, Korin v., 17-19.V.2008, A. Klimenko leg. (1♂, AAC); Kerman prov., 15 km E Korin, 2800-3300m, Kuh-e-Lalehzar mt., 17-19.V.2008, Anichtchenko A. leg. (1♂, AAC); Kerman, Sargad, 24-26.VI.1898, N. Zarudnyi leg. (1♀, ZIN); same locality, 25-27.VIII.1898 (1♂, ZIN); same locality, 30.IV.1901 (1♀, ZIN); Perse, Bender-Bouchir, Dr. Bussieres, 1905 (1♀, MNHN); S Iran, 65 km
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<normalizedToken originalValue="nördl">noerdl</normalizedToken>
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. Bandar-Abbas, 30.III.1972, Exped. Mus. Vind. (1 ex, NHMW). IRAQ: Bagdad, coll.
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<normalizedToken originalValue="Kálalová">Kalalova</normalizedToken>
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(6♂♂ 4♀♀, NMPC); Bagdad, Irak, coll.
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<normalizedToken originalValue="Kálalová">Kalalova</normalizedToken>
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(4 ex., NMPC); Bagdad, Ex Musaeo H.W. Bates 1892, Museum Paris, 1952, Coll. R.
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<normalizedToken originalValue="Oberthür">Oberthuer</normalizedToken>
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(2♂♂ 2♀♀, MNHN). PAKISTAN: NW Pakistan, prov. Swat, 71°90' L/ 35°70' B, Madyan, 1400 m, am Licht, 19.
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<normalizedToken originalValue="VI–">VI-</normalizedToken>
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4.VII.1971, leg. C. Holzschuh (2♂♂, NMPC); Northwest frontier Prov. Barseen, 35°21'42N, 73°12'42E, 900m, at light, No 26, 21.VII.1998, G. Csorba & L. Ronkay (1♂ 1♀, NHMW). TURKEY: Abanc-Sae, Preczmann, Aspock, Radda, 26.V.67 (1♀, NHMW).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="15" pageNumber="16" type="diagnosis">
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<paragraph pageId="15" pageNumber="16">Diagnosis.</paragraph>
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<paragraph pageId="15" pageNumber="16">
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Very similar in both external and aedeagal anatomy to
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<taxonomicName class="Insecta" family="Carabidae" genus="Singilis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Singilis turcicus" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="turcicus">Singilis turcicus</taxonomicName>
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. The widespread
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<taxonomicName class="Insecta" family="Carabidae" genus="Singilis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Singilis mesopotamicus" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="mesopotamicus">Singilis mesopotamicus</taxonomicName>
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somewhat varies in size, pronotal punctation, and elytral pattern. Endophallic anatomy in each species is rather uniform throughout its range, and the differences are minor and require further study.
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<taxonomicName class="Insecta" family="Carabidae" genus="Singilis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Singilis turcicus" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="turcicus">Singilis turcicus</taxonomicName>
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tends to be smaller, with elytra shorter and wider than in
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<taxonomicName class="Insecta" family="Carabidae" genus="Singilis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Singilis mesopotamicus" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="mesopotamicus">Singilis mesopotamicus</taxonomicName>
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, while the latter has a more transverse pronotum. In
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<taxonomicName class="Insecta" family="Carabidae" genus="Singilis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Singilis turcicus" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="turcicus">Singilis turcicus</taxonomicName>
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, the elytral spot is located immediately behind the middle, often restricted to innermost intervals, and never reaches elytral apices that are widely red-brown; while in
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<taxonomicName class="Insecta" family="Carabidae" genus="Singilis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Singilis mesopotamicus" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="mesopotamicus">Singilis mesopotamicus</taxonomicName>
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the spot is a bit farther posteriorly and usually reaches the apices (sometimes narrowly red-brown). Endophallic spine size is clearly different; small spines more numerous than in
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<taxonomicName class="Insecta" family="Carabidae" genus="Singilis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Singilis turcicus" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="turcicus">Singilis turcicus</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16" type="description">
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<paragraph pageId="15" pageNumber="16">Redescription.</paragraph>
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<paragraph pageId="15" pageNumber="16">Length 5.0-6.3 mm. Yellowish brown, elytra behind the middle with piceous spot that may reach apical and lateral margins but sometimes not extended beyond interval 6 and often leaving elytral apices brownish red (Figs 14-15).</paragraph>
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<paragraph pageId="15" pageNumber="16">Head microsculptured and deeply irregularly punctate, punctures sometimes almost confluent in frontal depressions, on the frons separated by more than 3 diameters. Clypeus with a few punctures near base. Eyes large and bulging, with no setae at posterior margin. Second supraocular seta located just anterad the posterior margin of eye. Temples short and smooth. Scape with a very long seta and a few short thin setae; pedicel with a band of apical setae; antennomere III with two bands of setae (at mid-length and at apex). Antennae pubescent from the basal third of antennomere IV.</paragraph>
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<paragraph lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16">
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Pronotum pale yellow, sometimes shinier than head, 1.16-1.18 times as wide as head, 1.4-1.47 times as wide as long, widest just behind the marginal setae. Anterior margin straight, anterior angles effaced to slightly marked, sides very broadly and evenly
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<pageBreakToken pageId="16" pageNumber="17" start="start">rounded</pageBreakToken>
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, slightly sinuate towards posterior angles, which are rectangular (normally) to subacute. Punctation, especially on disc, normally sparser and more delicate than on head; confluent and rugose towards lateral and posterior margins. Lateral explanate margin rapidly widens basad lateral setae, broad and flat at base. Posterior pore right in front of angle. Basal grooves shallow, punctate. Pronotal base extended in a rounded median lobe. Furrow variable. Microsculpture isodiametric.
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</paragraph>
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<paragraph pageId="16" pageNumber="17">Elytra 1.45-1.52 times as long as width in the middle. Intervals 1, 3, 5 and 7 each with minute setae and two irregular rows of pores from base to apex, often poorly visible. Even intervals may also bear a few setae in apical areas. Interval 7 convex and narrower than the adjacent intervals. Microsculpture nearly isodiametric. Apices slightly sinuate. Striae narrow and slightly punctate. Intervals slightly convex at base and flat behind the middle.</paragraph>
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<paragraph pageId="16" pageNumber="17">Legs brownish yellow. Metatarsomere V with 3-4 pairs of ventral setae. Propleura slightly wavy rugose towards sternum and coxae. Claws with 5 teeth. Venter brownish yellow throughout. All abdominal sterna pubescent; pubescence as short as on metatarsomere IV.</paragraph>
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<paragraph pageId="16" pageNumber="17">Aedeagus - Fig. 37. Aedeagal median lobe slender, ventral surface straight at midlength, apical fifth of lobe slightly downturned, tip long.</paragraph>
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</subSubSection>
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<subSubSection pageId="16" pageNumber="17" type="variation">
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<paragraph pageId="16" pageNumber="17">Variation.</paragraph>
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<paragraph pageId="16" pageNumber="17">Varies in size, elytral pattern (dark spot may be reduced), shape of posterior pronotal angles (usually acute and prominent but may be rectangular and less prominent). In almost all specimens from Afghanistan the apical spot reaches elytral apices, and the series of punctures in intervals 5 and 7 are shorter. Pakistani specimens have more convex elytral intervals.</paragraph>
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</subSubSection>
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<subSubSection pageId="16" pageNumber="17" type="comments">
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<paragraph pageId="16" pageNumber="17">Comments.</paragraph>
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<paragraph pageId="16" pageNumber="17">
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<bibRefCitation pageId="16" pageNumber="17">
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<normalizedToken originalValue="Jedlička">Jedlicka</normalizedToken>
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(1956
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</bibRefCitation>
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: 204) described
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<taxonomicName class="Insecta" family="Carabidae" genus="Singilis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Singilis apicalis" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="apicalis">Singilis apicalis</taxonomicName>
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from four syntypes (all NMPC): Afghanistan, Nuristan, Bashgultal (3♀♀); Asmar (1♀). Recent catalogues place this species in
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<taxonomicName class="Insecta" family="Carabidae" genus="Agatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Agatus" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">Agatus</taxonomicName>
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(
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<bibRefCitation pageId="16" pageNumber="17">Kabak 2003</bibRefCitation>
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;
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<bibRefCitation author="Lorenz, W" journalOrPublisher="Privately published, W. Lorenz, Tutzing, Germany" pageId="35" pageNumber="36" title="Nomina Carabidarum, a directory of the scientific names of ground beetles." year="2005">Lorenz 2005</bibRefCitation>
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).
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</paragraph>
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<paragraph pageId="16" pageNumber="17">
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<taxonomicName class="Insecta" family="Carabidae" genus="Singilis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Singilis plagiatus subsp. mesopotamicus" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="subSpecies" species="plagiatus" subSpecies="mesopotamicus">Singilis plagiatus mesopotamicus</taxonomicName>
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Pic, 1901, was described from Baghdad. The original description says that the type is in coll. Pic (MNHN), but my attempts to locate it have been unsuccessful. However, 4 specimens of
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<taxonomicName class="Insecta" family="Carabidae" genus="Singilis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Singilis apicalis" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="apicalis">Singilis apicalis</taxonomicName>
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labeled "Bagdad" I've found in MNHN match the original description of
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<taxonomicName class="Insecta" family="Carabidae" genus="Singilis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Singilis plagiatus subsp. mesopotamicus" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="subSpecies" species="plagiatus" subSpecies="mesopotamicus">Singilis plagiatus mesopotamicus</taxonomicName>
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. All other
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<taxonomicName class="Insecta" family="Carabidae" genus="Singilis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Singilis" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">Singilis</taxonomicName>
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specimens (14 total) from Baghdad area I have examined were
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<taxonomicName class="Insecta" family="Carabidae" genus="Singilis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Singilis apicalis" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="apicalis">Singilis apicalis</taxonomicName>
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. Therefore I presume
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<taxonomicName class="Insecta" family="Carabidae" genus="Singilis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Singilis apicalis" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="apicalis">Singilis apicalis</taxonomicName>
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to be synonymous with
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<taxonomicName class="Insecta" family="Carabidae" genus="Singilis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Singilis mesopotamicus" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="mesopotamicus">Singilis mesopotamicus</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="16" pageNumber="17" type="distribution">
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<paragraph pageId="16" pageNumber="17">Distribution.</paragraph>
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<paragraph pageId="16" pageNumber="17">Afghanistan, Iran, Iraq (country record), Pakistan (country record), Turkey (country record), Turkmenistan.</paragraph>
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</subSubSection>
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</treatment>
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</document> |