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<mods:title id="2F0232EBD84E9C1648E35649A0A759CB">New records of eutherian mammals from the Goler Formation (Tiffanian, Paleocene) of California and their biostratigraphic and paleobiogeographic implications</mods:title>
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<taxonomicName id="17DA4D5C142A517FFEFB1B2EFC15F9C1" authority="McKenna and Lofgren, 2003" authorityName="McKenna and Lofgren" authorityYear="2003" box="[369,975,1617,1644]" class="Mammalia" family="Arctocyonidae" genus="Mimotricentes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Procreodi" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="tedfordi">
<emphasis id="E2AEEACD142A517FFEFB1B2EFDACF9C6" box="[369,630,1617,1643]" italics="true" pageId="18" pageNumber="19">Mimotricentes tedfordi</emphasis>
<bibRefCitation id="B44B4B2E142A517FFDF41B2EFC15F9C1" author="McKenna, M. C. &amp; D. L. Lofgren" box="[638,975,1617,1644]" pageId="18" pageNumber="19" pagination="632 - 643" refId="ref30562" refString="McKenna, M. C., and D. L. Lofgren. 2003. Mimotricentes tedfordi, a new arctocyonid from the Late Paleocene of California. In L. J. Flynn (editor), Vertebrate fossils and their context: contributions in honor of Richard H. Tedford. Bulletin of the American Museum of Natural History 279: 632 - 643." type="journal article" year="2003">McKenna and Lofgren, 2003</bibRefCitation>
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<paragraph id="D06536DF142A517FFF221BF6FB12F90E" blockId="18.[120,1224,1673,1737]" box="[168,1224,1673,1700]" pageId="18" pageNumber="19">
<materialsCitation id="60B23C82142A517FFF221BF6FB12F90E" accessionNumber="V94133" box="[168,1224,1673,1700]" collectionCode="RAM" pageId="18" pageNumber="19" specimenCode="RAM 6908" specimenCount="1" typeStatus="holotype">
<typeStatus id="0F61887D142A517FFF221BF6FEF2F909" box="[168,296,1674,1700]" pageId="18" pageNumber="19" type="holotype">HOLOTYPE</typeStatus>
:
<specimenCode id="807C9EA4142A517FFEB21BF6FE64F90E" box="[312,446,1673,1700]" collectionCode="RAM" pageId="18" pageNumber="19">RAM 6908</specimenCode>
, right maxilla with P4M3 from
<collectionCode id="B6CBAE1A142A517FFCDA1BF6FC4BF909" box="[848,913,1674,1700]" pageId="18" pageNumber="19">RAM</collectionCode>
locality
<accessionNumber id="CF89AB3C142A517FFC701BF6FB82F909" box="[1018,1112,1674,1700]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/V94133" pageId="18" pageNumber="19" type="EnaNcbi">V94133</accessionNumber>
, member
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</paragraph>
<paragraph id="D06536DF142A517FFFF21BD3FE2CF964" blockId="18.[120,1224,1673,1737]" box="[120,502,1711,1737]" pageId="18" pageNumber="19">
4a, Goler Formation,
<collectingRegion id="121EF83D142A517FFEFD1BD3FE2BF964" box="[375,497,1711,1737]" country="United States of America" name="California" pageId="18" pageNumber="19">California</collectingRegion>
.
</paragraph>
<caption id="84A56657142B517EFFF21907FCE3FAD1" ID-DOI="http://doi.org/10.5281/zenodo.5365761" ID-Zenodo-Dep="5365761" httpUri="https://zenodo.org/record/5365761/files/figure.png" pageId="19" pageNumber="20" startId="19.[120,206,1147,1168]" targetBox="[130,1220,230,1117]" targetPageId="19">
<paragraph id="D06536DF142B517EFFF21907FCE3FAD1" blockId="19.[120,1228,1145,1404]" pageId="19" pageNumber="20">
FIGURE 6. Dentition of oxyclaenids and arctocyonids.
<emphasis id="E2AEEACD142B517EFD271905FD1FFB3C" bold="true" box="[685,709,1145,1169]" pageId="19" pageNumber="20">A.</emphasis>
Occlusal view of RAM 9041, right m1 of
<taxonomicName id="17DA4D5C142B517EFBF91905FF0CFB03" class="Mammalia" family="Arctocyonidae" genus="Thryptacodon" kingdom="Animalia" order="Procreodi" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="E2AEEACD142B517EFBF91905FF69FB02" italics="true" pageId="19" pageNumber="20">Thryptacodon</emphasis>
sp.
</taxonomicName>
from RAM locality V200510.
<emphasis id="E2AEEACD142B517EFD8619EBFDF8FB02" bold="true" box="[524,546,1175,1199]" pageId="19" pageNumber="20">B.</emphasis>
Occlusal view of RAM 15622, right dentary fragment with talonid of p4, and m12 of
<taxonomicName id="17DA4D5C142B517EFEB719C9FDC4FB61" authorityName="McKenna and Lofgren" authorityYear="2003" box="[317,542,1204,1228]" class="Mammalia" family="Arctocyonidae" genus="Mimotricentes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Procreodi" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="tedfordi">
<emphasis id="E2AEEACD142B517EFEB719C9FDC4FB61" box="[317,542,1204,1228]" italics="true" pageId="19" pageNumber="20">Mimotricentes tedfordi</emphasis>
</taxonomicName>
from RAM locality V201014;
<emphasis id="E2AEEACD142B517EFCDA19C8FCB2FB61" bold="true" box="[848,872,1204,1228]" pageId="19" pageNumber="20">C.</emphasis>
oblique labial view of RAM 15622.
<emphasis id="E2AEEACD142B517EFFF219AEFF4BFB47" bold="true" box="[120,145,1234,1258]" pageId="19" pageNumber="20">D.</emphasis>
Occlusal view of RAM 9670, left dentary fragment of
<taxonomicName id="17DA4D5C142B517EFD4F19AEFC72FB44" authorityName="McKenna and Lofgren" authorityYear="2003" box="[709,936,1233,1257]" class="Mammalia" family="Arctocyonidae" genus="Mimotricentes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Procreodi" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="tedfordi">
<emphasis id="E2AEEACD142B517EFD4F19AEFC72FB44" box="[709,936,1233,1257]" italics="true" pageId="19" pageNumber="20">Mimotricentes tedfordi</emphasis>
</taxonomicName>
with damaged m1 and base of p4 from RAM locality V200510.
<emphasis id="E2AEEACD142B517EFE6C1993FE21FAAA" bold="true" box="[486,507,1263,1287]" pageId="19" pageNumber="20">E.</emphasis>
Occlusal view of RAM 6928, right dentary fragment of
<taxonomicName id="17DA4D5C142B517EFBB7198CFF1FFA89" authorityName="McKenna and Lofgren" authorityYear="2003" class="Mammalia" family="Arctocyonidae" genus="Mimotricentes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Procreodi" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="tedfordi">
<emphasis id="E2AEEACD142B517EFBB7198CFF1FFA89" italics="true" pageId="19" pageNumber="20">Mimotricentes tedfordi</emphasis>
</taxonomicName>
with m3 talonid and base of trigonid from RAM locality V200001.
<emphasis id="E2AEEACD142B517EFC031870FC40FA89" bold="true" box="[905,922,1292,1316]" pageId="19" pageNumber="20">F.</emphasis>
Occlusal view of RAM 9040, right M1 of
<taxonomicName id="17DA4D5C142B517EFF7C1855FE5FFAEC" authorityName="Gingerich" authorityYear="1979" box="[246,389,1321,1345]" class="Mammalia" family="Arctocyonidae" genus="Lambertocyon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Procreodi" pageId="19" pageNumber="20" phylum="Chordata" rank="genus">
<emphasis id="E2AEEACD142B517EFF7C1855FE5FFAEC" box="[246,389,1321,1345]" italics="true" pageId="19" pageNumber="20">Lambertocyon</emphasis>
</taxonomicName>
cf.
<taxonomicName id="17DA4D5C142B517EFE211856FDFFFAEC" authorityName="Gunnell" authorityYear="1994" box="[427,549,1321,1345]" class="Mammalia" family="Plesiadapidae" genus="Plesiadapis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Primates" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="gingerichi">
<emphasis id="E2AEEACD142B517EFE211856FDFFFAEC" box="[427,549,1321,1345]" italics="true" pageId="19" pageNumber="20">L. gingerichi</emphasis>
</taxonomicName>
from RAM locality V200001.
<emphasis id="E2AEEACD142B517EFCD41855FCADFAEC" bold="true" box="[862,887,1321,1345]" pageId="19" pageNumber="20">G.</emphasis>
Occlusal view of RAM 9043, left p3 fragment of
<taxonomicName id="17DA4D5C142B517EFE92183BFE7DFAF2" authorityName="Gingerich" authorityYear="1979" box="[280,423,1351,1375]" class="Mammalia" family="Arctocyonidae" genus="Lambertocyon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Procreodi" pageId="19" pageNumber="20" phylum="Chordata" rank="genus">
<emphasis id="E2AEEACD142B517EFE92183BFE7DFAF2" box="[280,423,1351,1375]" italics="true" pageId="19" pageNumber="20">Lambertocyon</emphasis>
</taxonomicName>
cf.
<taxonomicName id="17DA4D5C142B517EFE441834FD93FAF2" authorityName="Gunnell" authorityYear="1994" box="[462,585,1351,1375]" class="Mammalia" family="Plesiadapidae" genus="Plesiadapis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Primates" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="gingerichi">
<emphasis id="E2AEEACD142B517EFE441834FD93FAF2" box="[462,585,1351,1375]" italics="true" pageId="19" pageNumber="20">L. gingerichi</emphasis>
</taxonomicName>
from RAM locality V200001.
<emphasis id="E2AEEACD142B517EFC09183BFC47FAF2" bold="true" box="[899,925,1351,1375]" pageId="19" pageNumber="20">H.</emphasis>
Occlusal view of RAM 9660, premolar of undetermined arctocyonid from RAM locality V200613.
</paragraph>
</caption>
<paragraph id="D06536DF142B517EFF2218EEFDBFF9EC" blockId="19.[120,1224,1425,1788]" pageId="19" pageNumber="20">
REFERRED SPECIMENS:
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15622, right dentary fragment with talonid of p4, and m12 from
<collectionCode id="B6CBAE1A142B517EFF3118CBFF26FA7C" box="[187,252,1463,1489]" pageId="19" pageNumber="20">RAM</collectionCode>
locality V201014, member 4a (
<figureCitation id="48E12A5A142B517EFDF918CBFD3DFA7F" box="[627,743,1463,1490]" captionStart="FIGURE 6" captionStartId="19.[120,206,1147,1168]" captionTargetBox="[130,1220,230,1117]" captionTargetId="figure-173@19.[205,692,583,855]" captionTargetPageId="19" captionText="FIGURE 6. Dentition of oxyclaenids and arctocyonids. A. Occlusal view of RAM 9041, right m1 of Thryptacodon sp. from RAM locality V200510. B. Occlusal view of RAM 15622, right dentary fragment with talonid of p4, and m12 of Mimotricentes tedfordi from RAM locality V201014; C. oblique labial view of RAM 15622. D. Occlusal view of RAM 9670, left dentary fragment of Mimotricentes tedfordi with damaged m1 and base of p4 from RAM locality V200510. E. Occlusal view of RAM 6928, right dentary fragment of Mimotricentes tedfordi with m3 talonid and base of trigonid from RAM locality V200001. F. Occlusal view of RAM 9040, right M1 of Lambertocyon cf. L. gingerichi from RAM locality V200001. G. Occlusal view of RAM 9043, left p3 fragment of Lambertocyon cf. L. gingerichi from RAM locality V200001. H. Occlusal view of RAM 9660, premolar of undetermined arctocyonid from RAM locality V200613." figureDoi="http://doi.org/10.5281/zenodo.5365761" httpUri="https://zenodo.org/record/5365761/files/figure.png" pageId="19" pageNumber="20">fig. 6BC</figureCitation>
);
<collectionCode id="B6CBAE1A142B517EFD7718CBFCE4FA7C" box="[765,830,1463,1489]" pageId="19" pageNumber="20">RAM</collectionCode>
9670, left dentary fragment with the base of p4 and damaged m1 from
<collectionCode id="B6CBAE1A142B517EFDA518A0FDAAFA5B" box="[559,624,1500,1526]" pageId="19" pageNumber="20">RAM</collectionCode>
locality V200510, member 4a (
<figureCitation id="48E12A5A142B517EFC5018A0FBF7FA5A" box="[986,1069,1500,1527]" captionStart="FIGURE 6" captionStartId="19.[120,206,1147,1168]" captionTargetBox="[130,1220,230,1117]" captionTargetId="figure-173@19.[205,692,583,855]" captionTargetPageId="19" captionText="FIGURE 6. Dentition of oxyclaenids and arctocyonids. A. Occlusal view of RAM 9041, right m1 of Thryptacodon sp. from RAM locality V200510. B. Occlusal view of RAM 15622, right dentary fragment with talonid of p4, and m12 of Mimotricentes tedfordi from RAM locality V201014; C. oblique labial view of RAM 15622. D. Occlusal view of RAM 9670, left dentary fragment of Mimotricentes tedfordi with damaged m1 and base of p4 from RAM locality V200510. E. Occlusal view of RAM 6928, right dentary fragment of Mimotricentes tedfordi with m3 talonid and base of trigonid from RAM locality V200001. F. Occlusal view of RAM 9040, right M1 of Lambertocyon cf. L. gingerichi from RAM locality V200001. G. Occlusal view of RAM 9043, left p3 fragment of Lambertocyon cf. L. gingerichi from RAM locality V200001. H. Occlusal view of RAM 9660, premolar of undetermined arctocyonid from RAM locality V200613." figureDoi="http://doi.org/10.5281/zenodo.5365761" httpUri="https://zenodo.org/record/5365761/files/figure.png" pageId="19" pageNumber="20">fig. 6D</figureCitation>
);
<collectionCode id="B6CBAE1A142B517EFBC818A0FB59FA5B" box="[1090,1155,1500,1526]" pageId="19" pageNumber="20">RAM</collectionCode>
6928, right dentary fragment with m3 talonid and base of trigonid (
<figureCitation id="48E12A5A142B517EFCCE1B7DFC4AF9B1" box="[836,912,1537,1564]" captionStart="FIGURE 6" captionStartId="19.[120,206,1147,1168]" captionTargetBox="[130,1220,230,1117]" captionTargetId="figure-173@19.[205,692,583,855]" captionTargetPageId="19" captionText="FIGURE 6. Dentition of oxyclaenids and arctocyonids. A. Occlusal view of RAM 9041, right m1 of Thryptacodon sp. from RAM locality V200510. B. Occlusal view of RAM 15622, right dentary fragment with talonid of p4, and m12 of Mimotricentes tedfordi from RAM locality V201014; C. oblique labial view of RAM 15622. D. Occlusal view of RAM 9670, left dentary fragment of Mimotricentes tedfordi with damaged m1 and base of p4 from RAM locality V200510. E. Occlusal view of RAM 6928, right dentary fragment of Mimotricentes tedfordi with m3 talonid and base of trigonid from RAM locality V200001. F. Occlusal view of RAM 9040, right M1 of Lambertocyon cf. L. gingerichi from RAM locality V200001. G. Occlusal view of RAM 9043, left p3 fragment of Lambertocyon cf. L. gingerichi from RAM locality V200001. H. Occlusal view of RAM 9660, premolar of undetermined arctocyonid from RAM locality V200613." figureDoi="http://doi.org/10.5281/zenodo.5365761" httpUri="https://zenodo.org/record/5365761/files/figure.png" pageId="19" pageNumber="20">fig. 6E</figureCitation>
), and
<collectionCode id="B6CBAE1A142B517EFC531B7EFBC0F9B1" box="[985,1050,1538,1564]" pageId="19" pageNumber="20">RAM</collectionCode>
15333, left M2 from
<collectionCode id="B6CBAE1A142B517EFF301B5BFF21F9EC" box="[186,251,1575,1601]" pageId="19" pageNumber="20">RAM</collectionCode>
locality V200001, member 4a.
</paragraph>
</subSubSection>
<subSubSection id="98C06554142B5179FF221B30FC54FE16" lastPageId="20" lastPageNumber="21" pageId="19" pageNumber="20" type="discussion">
<paragraph id="D06536DF142B517EFF221B30FCC9F956" blockId="19.[120,1224,1425,1788]" pageId="19" pageNumber="20">
DISCUSSION: The
<typeStatus id="0F61887D142B517EFEF21B30FE05F9CB" box="[376,479,1612,1638]" pageId="19" pageNumber="20" type="holotype">holotype</typeStatus>
of
<taxonomicName id="17DA4D5C142B517EFD8D1B31FD52F9CB" authorityName="McKenna and Lofgren" authorityYear="2003" box="[519,648,1612,1638]" class="Mammalia" family="Arctocyonidae" genus="Mimotricentes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Procreodi" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="tedfordi">
<emphasis id="E2AEEACD142B517EFD8D1B31FD52F9CB" box="[519,648,1612,1638]" italics="true" pageId="19" pageNumber="20">M. tedfordi</emphasis>
</taxonomicName>
, a maxillary fragment with P4-M3 (RAM 6908), from
<accessionNumber id="CF89AB3C142B517EFF331B0EFECDF921" box="[185,279,1650,1676]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/V94133" pageId="19" pageNumber="20" type="EnaNcbi">V94133</accessionNumber>
, high in member 4a, was described by
<bibRefCitation id="B44B4B2E142B517EFD6A1B0EFBE4F921" author="McKenna, M. C. &amp; D. L. Lofgren" box="[736,1086,1649,1676]" pageId="19" pageNumber="20" pagination="632 - 643" refId="ref30562" refString="McKenna, M. C., and D. L. Lofgren. 2003. Mimotricentes tedfordi, a new arctocyonid from the Late Paleocene of California. In L. J. Flynn (editor), Vertebrate fossils and their context: contributions in honor of Richard H. Tedford. Bulletin of the American Museum of Natural History 279: 632 - 643." type="journal article" year="2003">McKenna and Lofgren (2003)</bibRefCitation>
, who noted that
<taxonomicName id="17DA4D5C142B517EFF251BE4FEEBF91C" authorityName="McKenna and Lofgren" authorityYear="2003" box="[175,305,1687,1713]" class="Mammalia" family="Arctocyonidae" genus="Mimotricentes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Procreodi" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="tedfordi">
<emphasis id="E2AEEACD142B517EFF251BE4FEEBF91C" box="[175,305,1687,1713]" italics="true" pageId="19" pageNumber="20">M. tedfordi</emphasis>
</taxonomicName>
was significantly smaller than
<taxonomicName id="17DA4D5C142B517EFD291BE4FC8BF91C" baseAuthorityName="Cope" baseAuthorityYear="1881" box="[675,849,1687,1713]" class="Mammalia" family="Arctocyonidae" genus="Mimotricentes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Procreodi" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="subtrigonus">
<emphasis id="E2AEEACD142B517EFD291BE4FC8BF91C" box="[675,849,1687,1713]" italics="true" pageId="19" pageNumber="20">M. subtrigonus</emphasis>
</taxonomicName>
and all other described species of
<taxonomicName id="17DA4D5C142B517EFF121BC1FEE6F97B" authorityName="Simpson" authorityYear="1937" box="[152,316,1725,1750]" class="Mammalia" family="Arctocyonidae" genus="Mimotricentes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Procreodi" pageId="19" pageNumber="20" phylum="Chordata" rank="genus">
<emphasis id="E2AEEACD142B517EFF121BC1FEE6F97B" box="[152,316,1725,1750]" italics="true" pageId="19" pageNumber="20">Mimotricentes</emphasis>
</taxonomicName>
and
<taxonomicName id="17DA4D5C142B517EFEF21BC0FDC7F97B" authorityName="Gingerich" authorityYear="1979" box="[376,541,1724,1750]" class="Mammalia" family="Arctocyonidae" genus="Lambertocyon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Procreodi" pageId="19" pageNumber="20" phylum="Chordata" rank="genus">
<emphasis id="E2AEEACD142B517EFEF21BC0FDC7F97B" box="[376,541,1724,1750]" italics="true" pageId="19" pageNumber="20">Lambertocyon</emphasis>
</taxonomicName>
. The
<typeStatus id="0F61887D142B517EFDEA1BC0FD1DF97B" box="[608,711,1724,1750]" pageId="19" pageNumber="20" type="holotype">holotype</typeStatus>
was the only known specimen of
<taxonomicName id="17DA4D5C142B517EFBD41BC1FEC7F956" authorityName="McKenna and Lofgren" authorityYear="2003" class="Mammalia" family="Arctocyonidae" genus="Mimotricentes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Procreodi" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="tedfordi">
<emphasis id="E2AEEACD142B517EFBD41BC1FEC7F956" italics="true" pageId="19" pageNumber="20">Mimotricentes tedfordi</emphasis>
</taxonomicName>
, so the lower dentition was not described.
</paragraph>
<paragraph id="D06536DF142C5179FF221D9BFC54FE16" blockId="20.[120,1225,230,1788]" pageId="20" pageNumber="21">
In 2010, a dentary fragment (RAM 15622) was recovered from V201014, a site low in member 4a, about
<quantity id="17229B3A142C5179FED01C70FE43FE8B" box="[346,409,268,294]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="20" pageNumber="21" unit="km" value="3.0">3 km</quantity>
south of the type locality of
<taxonomicName id="17DA4D5C142C5179FD7C1C71FCA3FE8B" authorityName="McKenna and Lofgren" authorityYear="2003" box="[758,889,268,294]" class="Mammalia" family="Arctocyonidae" genus="Mimotricentes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Procreodi" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="tedfordi">
<emphasis id="E2AEEACD142C5179FD7C1C71FCA3FE8B" box="[758,889,268,294]" italics="true" pageId="20" pageNumber="21">M. tedfordi</emphasis>
</taxonomicName>
. RAM 15622 represents the lower dentition of a small arctocyonid that almost certainly is
<taxonomicName id="17DA4D5C142C5179FCC41C4EFC14FEE6" authorityName="McKenna and Lofgren" authorityYear="2003" box="[846,974,305,331]" class="Mammalia" family="Arctocyonidae" genus="Mimotricentes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Procreodi" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="tedfordi">
<emphasis id="E2AEEACD142C5179FCC41C4EFC14FEE6" box="[846,974,305,331]" italics="true" pageId="20" pageNumber="21">M. tedfordi</emphasis>
</taxonomicName>
. Two other fragmentary arctocyonid dentaries (RAM 6928 and RAM 9670), collected prior to 2008, probably also represent
<taxonomicName id="17DA4D5C142C5179FF661C01FEB4FE3B" authorityName="McKenna and Lofgren" authorityYear="2003" box="[236,366,380,406]" class="Mammalia" family="Arctocyonidae" genus="Mimotricentes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Procreodi" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="tedfordi">
<emphasis id="E2AEEACD142C5179FF661C01FEB4FE3B" box="[236,366,380,406]" italics="true" pageId="20" pageNumber="21">M. tedfordi</emphasis>
</taxonomicName>
. Also, an isolated M2 (RAM 15333), found in 2012, is nearly identical to the M2 of the
<typeStatus id="0F61887D142C5179FEA81CDDFE56FE16" box="[290,396,417,443]" pageId="20" pageNumber="21" type="holotype">holotype</typeStatus>
. These
<specimenCount id="C6DCFD56142C5179FE681CDDFD4CFE16" box="[482,662,417,443]" pageId="20" pageNumber="21" type="generic">four specimens</specimenCount>
are described below.
</paragraph>
</subSubSection>
<subSubSection id="98C06554142C5178FF221CBBFCB2FC02" lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21" type="description">
<paragraph id="D06536DF142C5179FF221CBBFCADFDFC" blockId="20.[120,1225,230,1788]" pageId="20" pageNumber="21">
<materialsCitation id="60B23C82142C5179FF221CBBFCA9FDFC" accessionNumber="RAM 15333" collectionCode="RAM" collectorName="The M" pageId="20" pageNumber="21" specimenCode="RAM 6908" specimenCount="2" typeStatus="holotype">
DESCRIPTION:
<collectorName id="7D2F5309142C5179FED31CBBFE73FE4C" box="[345,425,455,481]" pageId="20" pageNumber="21">The M</collectorName>
2 of the
<typeStatus id="0F61887D142C5179FD981CBBFDA3FE4C" box="[530,633,455,481]" pageId="20" pageNumber="21" type="holotype">holotype</typeStatus>
of
<taxonomicName id="17DA4D5C142C5179FD2E1CB4FCF2FE4C" authorityName="McKenna and Lofgren" authorityYear="2003" box="[676,808,455,481]" class="Mammalia" family="Arctocyonidae" genus="Mimotricentes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Procreodi" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="tedfordi">
<emphasis id="E2AEEACD142C5179FD2E1CB4FCF2FE4C" box="[676,808,455,481]" italics="true" pageId="20" pageNumber="21">M. tedfordi</emphasis>
</taxonomicName>
(
<specimenCode id="807C9EA4142C5179FCB61CBBFC1BFE4F" box="[828,961,455,482]" collectionCode="RAM" pageId="20" pageNumber="21">RAM 6908</specimenCode>
) and
<collectionCode id="B6CBAE1A142C5179FB831CBBFB90FE4C" box="[1033,1098,455,481]" pageId="20" pageNumber="21">RAM</collectionCode>
15333 are similar in all features, their unusually large crestlike parastyles being especially noteworthy. Also, the length of
<collectionCode id="B6CBAE1A142C5179FEDC1F6EFE4DFD81" box="[342,407,530,556]" pageId="20" pageNumber="21">RAM</collectionCode>
15333 is
<quantity id="17229B3A142C5179FD8F1F6DFDB7FD81" box="[517,621,529,556]" metricMagnitude="-3" metricUnit="m" metricValue="4.54" pageId="20" pageNumber="21" unit="mm" value="4.54">4.54 mm</quantity>
and the width
<quantity id="17229B3A142C5179FC941F6DFC52FD81" box="[798,904,529,556]" metricMagnitude="-3" metricUnit="m" metricValue="5.19" pageId="20" pageNumber="21" unit="mm" value="5.19">5.19 mm</quantity>
, dimensions similar to that of the M2 of the
<typeStatus id="0F61887D142C5179FECB1F4BFE72FDFC" box="[321,424,567,593]" pageId="20" pageNumber="21" type="holotype">holotype</typeStatus>
(length
<quantity id="17229B3A142C5179FD861F4BFDAFFDFC" box="[524,629,567,593]" metricMagnitude="-3" metricUnit="m" metricValue="4.62" pageId="20" pageNumber="21" unit="mm" value="4.62">4.62 mm</quantity>
and width
<quantity id="17229B3A142C5179FD771F4BFCB0FDFC" box="[765,874,567,594]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="20" pageNumber="21" unit="mm" value="5.5">5.50 mm</quantity>
)
</materialsCitation>
.
</paragraph>
<paragraph id="D06536DF142C5179FF221F20FB52FBBC" blockId="20.[120,1225,230,1788]" pageId="20" pageNumber="21">
RAM 15622 is a dentary fragment with a broken p4, and m12, with only the talonid and posterior wall of the trigonid of the p4 preserved (
<figureCitation id="48E12A5A142C5179FD471FFDFC9BFD31" box="[717,833,641,668]" captionStart="FIGURE 6" captionStartId="19.[120,206,1147,1168]" captionTargetBox="[130,1220,230,1117]" captionTargetId="figure-173@19.[205,692,583,855]" captionTargetPageId="19" captionText="FIGURE 6. Dentition of oxyclaenids and arctocyonids. A. Occlusal view of RAM 9041, right m1 of Thryptacodon sp. from RAM locality V200510. B. Occlusal view of RAM 15622, right dentary fragment with talonid of p4, and m12 of Mimotricentes tedfordi from RAM locality V201014; C. oblique labial view of RAM 15622. D. Occlusal view of RAM 9670, left dentary fragment of Mimotricentes tedfordi with damaged m1 and base of p4 from RAM locality V200510. E. Occlusal view of RAM 6928, right dentary fragment of Mimotricentes tedfordi with m3 talonid and base of trigonid from RAM locality V200001. F. Occlusal view of RAM 9040, right M1 of Lambertocyon cf. L. gingerichi from RAM locality V200001. G. Occlusal view of RAM 9043, left p3 fragment of Lambertocyon cf. L. gingerichi from RAM locality V200001. H. Occlusal view of RAM 9660, premolar of undetermined arctocyonid from RAM locality V200613." figureDoi="http://doi.org/10.5281/zenodo.5365761" httpUri="https://zenodo.org/record/5365761/files/figure.png" pageId="20" pageNumber="21">fig. 6BC</figureCitation>
). The p4 has a low,
<quantity id="17229B3A142C5179FBA11FFEFB5FFD31" box="[1067,1157,642,668]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="20" pageNumber="21" unit="mm" value="1.2">1.2 mm</quantity>
long, posterior shelf, which has a weakly developed posteromedial cusp connected to the trigonid wall by a small crest, features often seen in arctocyonids. The metaconid and protoconid of m1 are subequal in size and height, both larger than the low, labially shifted paraconid, and a protocristid and paracrista are present. The precingulid and a partial labial cingulid are present (labial cingulid terminates posteriorly at ectoflexid). The cristid obliqua is well developed and terminates at a large hypoconid, and a small but deep talonid basin, with steep posterior and labial slopes is present. A small posterolingually positioned entoconid and hypoconulid are closely appressed, with the hypoconulid slightly posterior to the transversely aligned hypoconid and entoconid. The m2 is similar to m
<quantity id="17229B3A142C5179FDCF1EAEFDB5FC46" box="[581,623,977,1004]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="20" pageNumber="21" unit="in" value="1.0">1 in</quantity>
most respects, but m2 is more quadrate in outline and has a more labially shifted paraconid. Depth of the mandible below m1 is
<quantity id="17229B3A142C5179FB921E8BFB5EFBBC" box="[1048,1156,1015,1042]" metricMagnitude="-3" metricUnit="m" metricValue="6.15" pageId="20" pageNumber="21" unit="mm" value="6.15">6.15 mm</quantity>
.
</paragraph>
<paragraph id="D06536DF142C5179FF221960FB7DF9B1" blockId="20.[120,1225,230,1788]" pageId="20" pageNumber="21">
The p4 of RAM 9670 is sheared off just above the gum line and the anterior part of the tooth is missing (
<figureCitation id="48E12A5A142C5179FEC0193DFE7AFBF1" box="[330,416,1089,1116]" captionStart="FIGURE 6" captionStartId="19.[120,206,1147,1168]" captionTargetBox="[130,1220,230,1117]" captionTargetId="figure-173@19.[205,692,583,855]" captionTargetPageId="19" captionText="FIGURE 6. Dentition of oxyclaenids and arctocyonids. A. Occlusal view of RAM 9041, right m1 of Thryptacodon sp. from RAM locality V200510. B. Occlusal view of RAM 15622, right dentary fragment with talonid of p4, and m12 of Mimotricentes tedfordi from RAM locality V201014; C. oblique labial view of RAM 15622. D. Occlusal view of RAM 9670, left dentary fragment of Mimotricentes tedfordi with damaged m1 and base of p4 from RAM locality V200510. E. Occlusal view of RAM 6928, right dentary fragment of Mimotricentes tedfordi with m3 talonid and base of trigonid from RAM locality V200001. F. Occlusal view of RAM 9040, right M1 of Lambertocyon cf. L. gingerichi from RAM locality V200001. G. Occlusal view of RAM 9043, left p3 fragment of Lambertocyon cf. L. gingerichi from RAM locality V200001. H. Occlusal view of RAM 9660, premolar of undetermined arctocyonid from RAM locality V200613." figureDoi="http://doi.org/10.5281/zenodo.5365761" httpUri="https://zenodo.org/record/5365761/files/figure.png" pageId="20" pageNumber="21">fig. 6D</figureCitation>
), but its length was at least
<quantity id="17229B3A142C5179FD79193DFC88FBF1" box="[755,850,1089,1116]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="20" pageNumber="21" unit="mm" value="3.5">3.5 mm</quantity>
, and width about
<quantity id="17229B3A142C5179FBBA193DFB55FBF1" box="[1072,1167,1089,1116]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="20" pageNumber="21" unit="mm" value="2.5">2.5 mm</quantity>
. The talonid of m1 is well preserved but the trigonid crown is missing due to breakage. The dimensions of the m1 of RAM 9670 are similar to those of the m1 of RAM 15622, but the talonid of RAM 9670 has crenulated enamel, a broader and shallower basin, and greater spacing of the entoconid and hypoconulid, differences suggesting they could represent different taxa. However, significant variation in lower molar morphology is well documented in
<taxonomicName id="17DA4D5C142C5179FBA91981FF27FA91" baseAuthorityName="Cope" baseAuthorityYear="1881" class="Mammalia" family="Arctocyonidae" genus="Mimotricentes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Procreodi" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="subtrigonus">
<emphasis id="E2AEEACD142C5179FBA91981FF27FA91" italics="true" pageId="20" pageNumber="21">Mimotricentes subtrigonus</emphasis>
</taxonomicName>
, a larger species than
<taxonomicName id="17DA4D5C142C5179FE74185EFDA5FA91" authorityName="McKenna and Lofgren" authorityYear="2003" box="[510,639,1314,1340]" class="Mammalia" family="Arctocyonidae" genus="Mimotricentes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Procreodi" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="tedfordi">
<emphasis id="E2AEEACD142C5179FE74185EFDA5FA91" box="[510,639,1314,1340]" italics="true" pageId="20" pageNumber="21">M. tedfordi</emphasis>
</taxonomicName>
(
<tableCitation id="9D580364142C5179FD04185EFD05FA91" box="[654,735,1314,1340]" captionStart="TABLE 7" captionStartId="21.[121,195,224,248]" captionText="TABLE 7. Measurements of the lower dentition of Mimotricentes tedfordi from the Goler Formation and Mimotricentes subtrigonus from Swain Quarry, Fort Union Formation, Wyoming. Measurements of M. subtrigonus from Rigby (1980: table 38). Measurements of M. tedfordi from RAM 9670 (p4m1), RAM 6928 (m3), and RAM 15622 (p4m2)." pageId="20" pageNumber="21">table 7</tableCitation>
), from Swain Quarry in
<collectingRegion id="121EF83D142C5179FC75185EFBAFFA91" box="[1023,1141,1314,1340]" country="United States of America" name="Wyoming" pageId="20" pageNumber="21">Wyoming</collectingRegion>
, where sample sizes of isolated teeth can exceed
<specimenCount id="C6DCFD56142C5179FDE4183BFCCAFACC" box="[622,784,1351,1377]" pageId="20" pageNumber="21" type="generic">60 specimens</specimenCount>
(
<bibRefCitation id="B44B4B2E142C5179FCAE183BFC69FACC" author="Rigby, J. K., Jr." box="[804,947,1351,1377]" pageId="20" pageNumber="21" pagination="1 - 179" refId="ref31055" refString="Rigby, J. K., Jr. 1980. Swain Quarry of the Fort Union Formation, middle Paleocene (Torrejonian), Carbon County, Wyoming: Geologic setting and mammalian fauna. Evolutionary Monographs 3: 1 - 179." type="journal article" year="1980">Rigby, 1980</bibRefCitation>
: table 38). Teeth of
<taxonomicName id="17DA4D5C142C5179FB2D1834FF24FA2B" baseAuthorityName="Cope" baseAuthorityYear="1881" class="Mammalia" family="Arctocyonidae" genus="Mimotricentes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Procreodi" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="subtrigonus">
<emphasis id="E2AEEACD142C5179FB2D1834FF24FA2B" italics="true" pageId="20" pageNumber="21">M. subtrigonus</emphasis>
</taxonomicName>
exhibit considerable variation in talonid development, as the proximity of the hypoconuild and entoconid can vary from nearly fused to well separated with the hypoconulid medially located (
<bibRefCitation id="B44B4B2E142C5179FEC118CBFE02FA7C" author="Rigby, J. K., Jr." box="[331,472,1463,1489]" pageId="20" pageNumber="21" pagination="1 - 179" refId="ref31055" refString="Rigby, J. K., Jr. 1980. Swain Quarry of the Fort Union Formation, middle Paleocene (Torrejonian), Carbon County, Wyoming: Geologic setting and mammalian fauna. Evolutionary Monographs 3: 1 - 179." type="journal article" year="1980">Rigby, 1980</bibRefCitation>
). Also, some m1s from Swain Quarry in the AMNH collections lack crenulated talonid enamel, while in others it is well developed. If this amount of variation is also present in
<taxonomicName id="17DA4D5C142C5179FECD1B7EFDC2F9B1" authority=", RAM" authorityName="RAM" box="[327,536,1538,1564]" class="Mammalia" family="Arctocyonidae" genus="Mimotricentes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Procreodi" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="tedfordi">
<emphasis id="E2AEEACD142C5179FECD1B7EFE12F9B1" box="[327,456,1538,1564]" italics="true" pageId="20" pageNumber="21">M. tedfordi</emphasis>
, RAM
</taxonomicName>
9670 likely represents the same species as RAM 15622.
</paragraph>
<paragraph id="D06536DF142C5178FF221B5BFCB2FC02" blockId="20.[120,1225,230,1788]" lastBlockId="21.[120,1224,841,943]" lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21">
RAM 6928 is a dentary fragment with a damaged m3 that may also represent
<taxonomicName id="17DA4D5C142C5179FBCA1B54FB18F9EC" box="[1088,1218,1575,1601]" class="Mammalia" family="Arctocyonidae" genus="Mimotricentes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Procreodi" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="tedfordi">
<emphasis id="E2AEEACD142C5179FBCA1B54FB18F9EC" box="[1088,1218,1575,1601]" italics="true" pageId="20" pageNumber="21">M. tedfordi</emphasis>
</taxonomicName>
. RAM 6928 has an intact m3 talonid, but its trigonid is broken at the gum line (
<figureCitation id="48E12A5A142C5179FBBE1B30FB58F9CA" box="[1076,1154,1612,1639]" captionStart="FIGURE 6" captionStartId="19.[120,206,1147,1168]" captionTargetBox="[130,1220,230,1117]" captionTargetId="figure-173@19.[205,692,583,855]" captionTargetPageId="19" captionText="FIGURE 6. Dentition of oxyclaenids and arctocyonids. A. Occlusal view of RAM 9041, right m1 of Thryptacodon sp. from RAM locality V200510. B. Occlusal view of RAM 15622, right dentary fragment with talonid of p4, and m12 of Mimotricentes tedfordi from RAM locality V201014; C. oblique labial view of RAM 15622. D. Occlusal view of RAM 9670, left dentary fragment of Mimotricentes tedfordi with damaged m1 and base of p4 from RAM locality V200510. E. Occlusal view of RAM 6928, right dentary fragment of Mimotricentes tedfordi with m3 talonid and base of trigonid from RAM locality V200001. F. Occlusal view of RAM 9040, right M1 of Lambertocyon cf. L. gingerichi from RAM locality V200001. G. Occlusal view of RAM 9043, left p3 fragment of Lambertocyon cf. L. gingerichi from RAM locality V200001. H. Occlusal view of RAM 9660, premolar of undetermined arctocyonid from RAM locality V200613." figureDoi="http://doi.org/10.5281/zenodo.5365761" httpUri="https://zenodo.org/record/5365761/files/figure.png" pageId="20" pageNumber="21">fig. 6E</figureCitation>
). The talonid has a low hypoconulid, an entoconid crest with three cuspules, and a large hypoconid. The talonid basin of m3 is broad and shallow and opens lingually through a small talonid notch. The morphology of the m3 talonid in
<taxonomicName id="17DA4D5C142C5179FD161BC1FC15F97B" baseAuthorityName="Cope" baseAuthorityYear="1881" box="[668,975,1724,1750]" class="Mammalia" family="Arctocyonidae" genus="Mimotricentes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Procreodi" pageId="20" pageNumber="21" phylum="Chordata" rank="species" species="subtrigonus">
<emphasis id="E2AEEACD142C5179FD161BC1FC15F97B" box="[668,975,1724,1750]" italics="true" pageId="20" pageNumber="21">Mimotricentes subtrigonus</emphasis>
</taxonomicName>
from Swain Quarry, varies even more than the m1 talonid, so much so that the m3 is “posteriorly too variable for taxonomic use” (
<bibRefCitation id="B44B4B2E142D5178FEB51E35FDDFFCCE" author="Rigby, J. K., Jr." box="[319,517,841,867]" pageId="21" pageNumber="22" pagination="1 - 179" refId="ref31055" refString="Rigby, J. K., Jr. 1980. Swain Quarry of the Fort Union Formation, middle Paleocene (Torrejonian), Carbon County, Wyoming: Geologic setting and mammalian fauna. Evolutionary Monographs 3: 1 - 179." type="journal article" year="1980">Rigby, 1980: 104</bibRefCitation>
). Specimens of
<taxonomicName id="17DA4D5C142D5178FD361E36FCB3FCCE" baseAuthorityName="Cope" baseAuthorityYear="1881" box="[700,873,841,867]" class="Mammalia" family="Arctocyonidae" genus="Mimotricentes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Procreodi" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="subtrigonus">
<emphasis id="E2AEEACD142D5178FD361E36FCB3FCCE" box="[700,873,841,867]" italics="true" pageId="21" pageNumber="22">M. subtrigonus</emphasis>
</taxonomicName>
observed at the AMNH from Swain Quarry have m3 talonids of similar construction as RAM 6928, but are much larger (
<tableCitation id="9D580364142D5178FF0A1EE8FF09FC02" box="[128,211,916,943]" captionStart="TABLE 7" captionStartId="21.[121,195,224,248]" captionText="TABLE 7. Measurements of the lower dentition of Mimotricentes tedfordi from the Goler Formation and Mimotricentes subtrigonus from Swain Quarry, Fort Union Formation, Wyoming. Measurements of M. subtrigonus from Rigby (1980: table 38). Measurements of M. tedfordi from RAM 9670 (p4m1), RAM 6928 (m3), and RAM 15622 (p4m2)." pageId="21" pageNumber="22">table 7</tableCitation>
). Thus, RAM 6928 probably also represents
<taxonomicName id="17DA4D5C142D5178FD551EE9FCBBFC03" box="[735,865,916,942]" class="Mammalia" family="Arctocyonidae" genus="Mimotricentes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Procreodi" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="tedfordi">
<emphasis id="E2AEEACD142D5178FD551EE9FCBBFC03" box="[735,865,916,942]" italics="true" pageId="21" pageNumber="22">M. tedfordi</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>