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<document id="5EBB40A2A83A78AD092EDB152611A98F" ID-CLB-Dataset="31279" ID-DOI="10.11646/zootaxa.4358.2.6" ID-GBIF-Dataset="ab4e1d4d-3b0a-430f-8e9a-4bfc8e991a4e" ID-ISSN="1175-5326" ID-Zenodo-Dep="1068376" ID-ZooBank="C21198D8-7962-4C81-A1C5-CB9BD826CAB8" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1511986867004" checkinUser="plazi" docAuthor="Jałoszyński, Paweł" docDate="2017" docId="583087E8C97AFFEBFF3EFD9F593EAC44" docLanguage="en" docName="zootaxa.4358.2.6.pdf.imf" docOrigin="Zootaxa 4358 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Euconnus" docType="treatment" docVersion="7" masterDocId="A409FF90C97DFFE2FFA9FFF15A68AD04" masterDocTitle="Taxonomy of ' Euconnus complex'. Part XVI. Alloconophron Franz transferred to Anhoraeomorphus Franz as subgenus, with notes on systematic position of Noctophus Casey (Coleoptera, Staphylinidae, Scydmaeninae)" updateTime="1698505123509" updateUser="ExternalLinkService">
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<mods:title id="F355937DE66D2BE4D349D32B8ACE2120">Taxonomy of ' Euconnus complex'. Part XVI. Alloconophron Franz transferred to Anhoraeomorphus Franz as subgenus, with notes on systematic position of Noctophus Casey (Coleoptera, Staphylinidae, Scydmaeninae)</mods:title>
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<treatment id="583087E8C97AFFEBFF3EFD9F593EAC44" ID-DOI="http://doi.org/10.5281/zenodo.6044490" ID-GBIF-Taxon="137581163" ID-Zenodo-Dep="6044490" LSID="urn:lsid:plazi:treatment:583087E8C97AFFEBFF3EFD9F593EAC44" httpUri="http://treatment.plazi.org/id/583087E8C97AFFEBFF3EFD9F593EAC44" lastPageId="9" pageId="7">
<subSubSection id="98836575C97AFFE5FF3EFD9F5E2AAF8C" box="[151,1090,622,648]" pageId="7" type="nomenclature">
<paragraph id="D02636FEC97AFFE5FF3EFD9F5E2AAF8C" blockId="7.[151,1090,622,648]" box="[151,1090,622,648]" pageId="7">
<heading id="8B6E8192C97AFFE5FF3EFD9F5E2AAF8C" bold="true" box="[151,1090,622,648]" fontSize="11" level="1" pageId="7" reason="1">
<emphasis id="E2EDEAECC97AFFE5FF3EFD9F5E2AAF8C" bold="true" box="[151,1090,622,648]" pageId="7">
<keyStep id="6B6D2E5BC97AFFE5FF3EFD9F5E2AAF8C" box="[151,1090,622,648]" pageId="7">
<keyLead id="6B6895CBC97AFFE5FF3EFD9F5E2AAF8C" box="[151,1090,622,648]" pageId="7">
2. Morphological structures and taxonomic status of
<taxonomicName id="17994D7DC97AFFE5FC80FD9F5E2AAF8C" authority="(Noctophus)" baseAuthorityName="Noctophus" baseAuthorityYear="2017" box="[809,1090,622,648]" class="Insecta" family="Scydmaenidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="7" phylum="Arthropoda" rank="genus">
<emphasis id="E2EDEAECC97AFFE5FC80FD9F59CDAF8C" bold="true" box="[809,933,622,648]" italics="true" pageId="7">Euconnus</emphasis>
(
<emphasis id="E2EDEAECC97AFFE5FC1FFD9F5E50AF8C" bold="true" box="[950,1080,622,648]" italics="true" pageId="7">Noctophus</emphasis>
)
</taxonomicName>
</keyLead>
</keyStep>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="98836575C97AFFE5FF3EFD445941AE86" pageId="7" type="diagnosis">
<paragraph id="D02636FEC97AFFE5FF3EFD445941AE86" blockId="7.[151,1436,693,898]" pageId="7">
<taxonomicName id="17994D7DC97AFFE5FF3EFD445B67AFC9" authorityName="Casey" authorityYear="1897" box="[151,271,693,717]" class="Insecta" family="Staphylinidae" genus="Noctophus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" phylum="Arthropoda" rank="genus">
<emphasis id="E2EDEAECC97AFFE5FF3EFD445B67AFC9" box="[151,271,693,717]" italics="true" pageId="7">Noctophus</emphasis>
</taxonomicName>
(
<figureCitation id="48A22A7BC97AFFE5FEB4FD445B1CAFCA" box="[285,372,693,718]" captionStart="FIGURES 1416" captionStartId="7.[151,264,1896,1919]" captionTargetBox="[158,1436,974,1849]" captionTargetId="figure@7.[151,1436,974,1849]" captionTargetPageId="7" captionText="FIGURES 1416. Morphological structures of Euconnus (Noctophus) schmitti (Casey). Dorsal habitus of holotype female (14); head, prothorax and elytral base in dorsal view (15); head, prothorax and anterior portion of pterothorax in ventral view (16). Abbreviations: abp, antebasal pit; bef, basal elytral fovea; bst, basisternal part of prosternum; hr, hypostomal ridge; hyr, hypomeral ridge; ptp, posterior tentorial pit; nss, notosternal suture; smn, submentum." httpUri="https://zenodo.org/record/1068388/files/figure.png" pageId="7">Figs 14</figureCitation>
¯18) differs from
<taxonomicName id="17994D7DC97AFFE5FD93FD445962AFC9" authorityName="Jałoszyński" authorityYear="2017" box="[570,778,693,717]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" phylum="Arthropoda" rank="genus">
<emphasis id="E2EDEAECC97AFFE5FD93FD445962AFC9" box="[570,778,693,717]" italics="true" pageId="7">Anhoraeomorphus</emphasis>
</taxonomicName>
merely in a short, round (and not elongate) anterior part of head; vertex not bulging posterodorsally; emarginate labrum; hypostomal ridges running posteromesally and not connected at middle; antennae gradually thickened (and not with a delimited club); an uneven number of pronotal antebasal pits, with a distinct median pit (lacking in
<taxonomicName id="17994D7DC97AFFE5FCB6FCD05998AE3D" authorityName="Jałoszyński" authorityYear="2017" box="[799,1008,801,825]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" phylum="Arthropoda" rank="genus">
<emphasis id="E2EDEAECC97AFFE5FCB6FCD05998AE3D" box="[799,1008,801,825]" italics="true" pageId="7">Anhoraeomorphus</emphasis>
</taxonomicName>
); and protibiae modified. All these characters are known only in a female, males of
<taxonomicName id="17994D7DC97AFFE5FD1BFCB45942AE59" authorityName="Casey" authorityYear="1897" box="[690,810,837,861]" class="Insecta" family="Staphylinidae" genus="Noctophus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" phylum="Arthropoda" rank="genus">
<emphasis id="E2EDEAECC97AFFE5FD1BFCB45942AE59" box="[690,810,837,861]" italics="true" pageId="7">Noctophus</emphasis>
</taxonomicName>
remain unknown. Moreover, the
<typeStatus id="0F22885CC97AFFE5FB37FCB45F68AE5A" box="[1182,1280,837,862]" pageId="7" type="holotype">holotype</typeStatus>
of
<taxonomicName id="17994D7DC97AFFE5FA8DFCB75FF4AE59" baseAuthorityName="Casey" baseAuthorityYear="1897" box="[1316,1436,837,861]" class="Insecta" family="Scydmaenidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="7" phylum="Arthropoda" rank="species" species="schmitti">
<emphasis id="E2EDEAECC97AFFE5FA8DFCB75FF4AE59" box="[1316,1436,837,861]" italics="true" pageId="7">E. schmitti</emphasis>
</taxonomicName>
has lost both maxillary palps during previous preparations.
</paragraph>
</subSubSection>
<caption id="84E66676C97AFFE5FF3EF8995996AADE" httpUri="https://zenodo.org/record/1068388/files/figure.png" pageId="7" targetBox="[158,1436,974,1849]" targetPageId="7">
<paragraph id="D02636FEC97AFFE5FF3EF8995996AADE" blockId="7.[151,1436,1896,2010]" pageId="7">
<emphasis id="E2EDEAECC97AFFE5FF3EF8995B3FAA7A" bold="true" box="[151,343,1896,1919]" pageId="7">FIGURES 1416.</emphasis>
Morphological structures of
<taxonomicName id="17994D7DC97AFFE5FD23F89B5E42AA7B" authority="(Casey)" authorityName="Casey" baseAuthorityName="Casey" baseAuthorityYear="1897" box="[650,1066,1897,1919]" class="Insecta" family="Scydmaenidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="7" phylum="Arthropoda" rank="species" species="schmitti" subGenus="Noctophus">
<emphasis id="E2EDEAECC97AFFE5FD23F89B5886AA7B" box="[650,750,1898,1919]" italics="true" pageId="7">Euconnus</emphasis>
(
<emphasis id="E2EDEAECC97AFFE5FCA9F8985904AA7B" box="[768,876,1897,1919]" italics="true" pageId="7">Noctophus</emphasis>
)
<emphasis id="E2EDEAECC97AFFE5FCD7F89859A6AA7B" box="[894,974,1897,1919]" italics="true" pageId="7">schmitti</emphasis>
(Casey)
</taxonomicName>
. Dorsal habitus of holotype female (14); head, prothorax and elytral base in dorsal view (15); head, prothorax and anterior portion of pterothorax in ventral view (16). Abbreviations: abp, antebasal pit; bef, basal elytral fovea; bst, basisternal part of prosternum; hr, hypostomal ridge; hyr, hypomeral ridge; ptp, posterior tentorial pit; nss, notosternal suture; smn, submentum.
</paragraph>
</caption>
<caption id="84E66676C975FFEAFF3EFAF45E99A85D" httpUri="https://zenodo.org/record/1068390/files/figure.png" pageId="8" targetBox="[151,1436,193,1264]" targetPageId="8">
<paragraph id="D02636FEC975FFEAFF3EFAF45E99A85D" blockId="8.[151,1436,1285,1369]" pageId="8">
<emphasis id="E2EDEAECC975FFEAFF3EFAF45B3DA81F" bold="true" box="[151,341,1285,1308]" pageId="8">FIGURES 1719.</emphasis>
Morphological structures of
<taxonomicName id="17994D7DC975FFEAFD2DFAF65E74A818" authority="(Casey)" authorityName="Casey" baseAuthorityName="Casey" baseAuthorityYear="1897" box="[644,1052,1286,1308]" class="Insecta" family="Scydmaenidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="8" phylum="Arthropoda" rank="species" species="schmitti" subGenus="Noctophus">
<emphasis id="E2EDEAECC975FFEAFD2DFAF65880A818" box="[644,744,1287,1308]" italics="true" pageId="8">Euconnus</emphasis>
(
<emphasis id="E2EDEAECC975FFEAFD51FAF7590CA818" box="[760,868,1286,1308]" italics="true" pageId="8">Noctophus</emphasis>
)
<emphasis id="E2EDEAECC975FFEAFCDCFAF759ADA818" box="[885,965,1286,1308]" italics="true" pageId="8">schmitti</emphasis>
(Casey)
</taxonomicName>
(17¯18) and original set of labels of holotype female (19). Pterothorax and abdomen in ventral view (17); left protibia and tarsus in ventral view. Abbreviations: ar, anterior ridge; msvp, mesoventral intercoxal process; mtvp, metaventral intercoxal process; pcr, procoxal rest.
</paragraph>
</caption>
<subSubSection id="98836575C975FFEAFF6EFA715835AB49" pageId="8" type="discussion">
<paragraph id="D02636FEC975FFEAFF6EFA715835AB49" blockId="8.[151,1436,1408,2009]" pageId="8">
<emphasis id="E2EDEAECC975FFEAFF6EFA715B37A89D" bold="true" box="[199,351,1408,1433]" pageId="8">Conclusions.</emphasis>
<taxonomicName id="17994D7DC975FFEAFECFFA705BB6A89D" authorityName="Casey" authorityYear="1897" box="[358,478,1409,1433]" class="Insecta" family="Staphylinidae" genus="Noctophus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" phylum="Arthropoda" rank="genus">
<emphasis id="E2EDEAECC975FFEAFECFFA705BB6A89D" box="[358,478,1409,1433]" italics="true" pageId="8">Noctophus</emphasis>
</taxonomicName>
must be excluded from
<taxonomicName id="17994D7DC975FFEAFD5BFA705908A89C" baseAuthorityName="Jaloszynski" baseAuthorityYear="2017" box="[754,864,1409,1432]" class="Insecta" family="Scydmaenidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="8" phylum="Arthropoda" rank="genus">
<emphasis id="E2EDEAECC975FFEAFD5BFA705908A89C" box="[754,864,1409,1432]" italics="true" pageId="8">Euconnus</emphasis>
</taxonomicName>
on the basis of a narrow, subtriangular and deeply notched at middle metaventral process.
<taxonomicName id="17994D7DC975FFEAFDF4FA5458BDA8B9" authorityName="Casey" authorityYear="1897" box="[605,725,1445,1469]" class="Insecta" family="Staphylinidae" genus="Noctophus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" phylum="Arthropoda" rank="genus">
<emphasis id="E2EDEAECC975FFEAFDF4FA5458BDA8B9" box="[605,725,1445,1469]" italics="true" pageId="8">Noctophus</emphasis>
</taxonomicName>
, although very similar to
<taxonomicName id="17994D7DC975FFEAFC56FA545EA7A8B9" authorityName="Jałoszyński" authorityYear="2017" box="[1023,1231,1445,1469]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" phylum="Arthropoda" rank="genus">
<emphasis id="E2EDEAECC975FFEAFC56FA545EA7A8B9" box="[1023,1231,1445,1469]" italics="true" pageId="8">Anhoraeomorphus</emphasis>
</taxonomicName>
(especially to the subgenus
<taxonomicName id="17994D7DC975FFEAFEAEFA385BC6A8E5" authorityName="Franz" authorityYear="1986" box="[263,430,1481,1505]" class="Insecta" family="Staphylinidae" genus="Alloconophron" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" phylum="Arthropoda" rank="genus">
<emphasis id="E2EDEAECC975FFEAFEAEFA385BC6A8E5" box="[263,430,1481,1505]" italics="true" pageId="8">Alloconophron</emphasis>
</taxonomicName>
due to complete hypomeral ridges), differs in the hypostomal ridges, which do not form the unique single transverse ridge seen in the latter genus. Besides, I am reluctant to place in one genus species that are known to occur in
<collectingCountry id="A88E766EC975FFEAFE37F9E1584DAB2D" box="[414,549,1552,1577]" name="Madagascar" pageId="8">Madagascar</collectingCountry>
and in the eastern
<collectingCountry id="A88E766EC975FFEAFCAAF9E159F7AB2D" box="[771,927,1552,1577]" name="United States of America" pageId="8">United States</collectingCountry>
; at least not until phylogenetic evidence has been found for such a close relationship.
</paragraph>
</subSubSection>
<subSubSection id="98836575C975FFEAFF6EF9A85820AA69" pageId="8" type="diagnosis">
<paragraph id="D02636FEC975FFEAFF6EF9A85820AA69" blockId="8.[151,1436,1408,2009]" pageId="8">
<taxonomicName id="17994D7DC975FFEAFF6EF9A85B57AB75" authorityName="Casey" authorityYear="1897" box="[199,319,1625,1649]" class="Insecta" family="Staphylinidae" genus="Noctophus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" phylum="Arthropoda" rank="genus">
<emphasis id="E2EDEAECC975FFEAFF6EF9A85B57AB75" box="[199,319,1625,1649]" italics="true" pageId="8">Noctophus</emphasis>
</taxonomicName>
is also very similar to the exclusively Southern Hemisphere
<taxonomicName id="17994D7DC975FFEAFBA2F9A85F76AB75" authority="Broun, 1893" authorityName="Broun" authorityYear="1893" box="[1035,1310,1625,1649]" class="Insecta" family="Staphylinidae" genus="Sciacharis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" phylum="Arthropoda" rank="genus">
<emphasis id="E2EDEAECC975FFEAFBA2F9A85EE9AB75" box="[1035,1153,1625,1649]" italics="true" pageId="8">Sciacharis</emphasis>
<bibRefCitation id="B4084B0FC975FFEAFB22F9A85F76AB75" author="Broun" box="[1163,1310,1625,1649]" pageId="8" refString="Broun, T. (1893) Manual of the New Zealand Coleoptera. Parts V, VI, VII. New Zealand Institute, Wellington, xvii, 975 - 1504." type="book chapter" year="1893">Broun, 1893</bibRefCitation>
</taxonomicName>
. The latter genus also has unconnected hypomeral ridges, two small asetose basal foveae on each elytron, and nearly identical ventral structures of the pterothorax (illustrated in
<bibRefCitation id="B4084B0FC975FFEAFD54F951598BABBD" author="Jaloszynski" box="[765,995,1696,1721]" pageId="8" refString="Jaloszynski, P. (2014) Comparative morphology and taxonomy of Sciacharis s. str., Chathamaenus Franz and Maorinus Franz (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa, 3794 (2), 222 - 236. https: // doi. org / 10.11646 / zootaxa. 3794.2.2" type="journal article" year="2014" yearSuffix="a">Jałoszyński (2014a)</bibRefCitation>
and
<bibRefCitation id="B4084B0FC975FFEAFB8EF9515FF1ABBD" author="Jaloszynski" box="[1063,1433,1696,1721]" pageId="8" refString="Jaloszynski, P. &amp; Newton, A. F., Jr. (2017) Taxonomy of ' Euconnus complex'. Part XV. Disentangling the Euconnus - Sciacharis knot (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa, 4344 (3), 401 ¯ 443. http: // dx. doi. org / 10.11646 / zootaxa. 4344.3.1" type="journal article" year="2017">Jałoszyński &amp; Newton (2017))</bibRefCitation>
. However, there are a few minor differences, as the head in
<taxonomicName id="17994D7DC975FFEAFCF8F93459A1ABD9" authorityName="Casey" authorityYear="1897" box="[849,969,1733,1757]" class="Insecta" family="Staphylinidae" genus="Noctophus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" phylum="Arthropoda" rank="genus">
<emphasis id="E2EDEAECC975FFEAFCF8F93459A1ABD9" box="[849,969,1733,1757]" italics="true" pageId="8">Noctophus</emphasis>
</taxonomicName>
is very short (length = width) and very convex; it is neither 'anthiciform' (i.e., subpentagonal and often flattened) as in
<taxonomicName id="17994D7DC975FFEAFBA2F9185EE9AA05" authority="Jałoszyński, 2017" authorityName="Jałoszyński" authorityYear="2017" box="[1035,1153,1769,1793]" class="Insecta" family="Staphylinidae" genus="Sciacharis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="E2EDEAECC975FFEAFBA2F9185EE9AA05" box="[1035,1153,1769,1793]" italics="true" pageId="8">Sciacharis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F9DE5797C975FFEAFB2EF9185EACAA04" box="[1159,1220,1769,1792]" pageId="8" sensu="stricto">s. str.</taxonomicNameLabel>
, nor elongate, as in many
<taxonomicName id="17994D7DC975FFEAFF74F8FC5BB5AA21" authority="(Maorinus)" authorityName="Jałoszyński" authorityYear="2017" baseAuthorityName="Maorinus" box="[221,477,1805,1829]" class="Insecta" family="Staphylinidae" genus="Sciacharis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" phylum="Arthropoda" rank="genus">
<emphasis id="E2EDEAECC975FFEAFF74F8FC5B3BAA21" box="[221,339,1805,1829]" italics="true" pageId="8">Sciacharis</emphasis>
(
<emphasis id="E2EDEAECC975FFEAFECCF8FC5BBAAA20" box="[357,466,1805,1828]" italics="true" pageId="8">Maorinus</emphasis>
)
</taxonomicName>
; also the antebasal pronotal pits in
<taxonomicName id="17994D7DC975FFEAFCDBF8FC5982AA21" authorityName="Casey" authorityYear="1897" box="[882,1002,1805,1829]" class="Insecta" family="Staphylinidae" genus="Noctophus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" phylum="Arthropoda" rank="genus">
<emphasis id="E2EDEAECC975FFEAFCDBF8FC5982AA21" box="[882,1002,1805,1829]" italics="true" pageId="8">Noctophus</emphasis>
</taxonomicName>
are very small, broadly separated and their number is uneven, whereas those in
<taxonomicName id="17994D7DC975FFEAFD2CF8C05893AA4D" authorityName="Jałoszyński" authorityYear="2017" box="[645,763,1841,1865]" class="Insecta" family="Staphylinidae" genus="Sciacharis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" phylum="Arthropoda" rank="genus">
<emphasis id="E2EDEAECC975FFEAFD2CF8C05893AA4D" box="[645,763,1841,1865]" italics="true" pageId="8">Sciacharis</emphasis>
</taxonomicName>
are paired, typically large and the inner pair is narrowly separated at middle or nearly adjacent.
</paragraph>
</subSubSection>
<subSubSection id="98836575C975FFEBFF6EF889593EAC44" lastPageId="9" pageId="8" type="discussion">
<paragraph id="D02636FEC975FFEBFF6EF889593EAC44" blockId="8.[151,1436,1408,2009]" lastBlockId="9.[151,1436,151,789]" lastPageId="9" pageId="8">
Based on numerous similarities between
<taxonomicName id="17994D7DC975FFEAFD3FF8885966AA95" authorityName="Casey" authorityYear="1897" box="[662,782,1913,1937]" class="Insecta" family="Staphylinidae" genus="Noctophus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" phylum="Arthropoda" rank="genus">
<emphasis id="E2EDEAECC975FFEAFD3FF8885966AA95" box="[662,782,1913,1937]" italics="true" pageId="8">Noctophus</emphasis>
</taxonomicName>
and
<taxonomicName id="17994D7DC975FFEAFCECF88859D3AA95" authorityName="Jałoszyński" authorityYear="2017" box="[837,955,1913,1937]" class="Insecta" family="Staphylinidae" genus="Sciacharis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" phylum="Arthropoda" rank="genus">
<emphasis id="E2EDEAECC975FFEAFCECF88859D3AA95" box="[837,955,1913,1937]" italics="true" pageId="8">Sciacharis</emphasis>
</taxonomicName>
, the former could be placed as a subgenus of the latter. The distribution (
<taxonomicName id="17994D7DC975FFEAFE40F86C5809AAB1" authorityName="Casey" authorityYear="1897" box="[489,609,1949,1973]" class="Insecta" family="Staphylinidae" genus="Noctophus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" phylum="Arthropoda" rank="genus">
<emphasis id="E2EDEAECC975FFEAFE40F86C5809AAB1" box="[489,609,1949,1973]" italics="true" pageId="8">Noctophus</emphasis>
</taxonomicName>
in North America,
<taxonomicName id="17994D7DC975FFEAFC95F86C59DAAAB1" authorityName="Jałoszyński" authorityYear="2017" box="[828,946,1949,1973]" class="Insecta" family="Staphylinidae" genus="Sciacharis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" phylum="Arthropoda" rank="genus">
<emphasis id="E2EDEAECC975FFEAFC95F86C59DAAAB1" box="[828,946,1949,1973]" italics="true" pageId="8">Sciacharis</emphasis>
</taxonomicName>
in
<collectingCountry id="A88E766EC975FFEAFC7FF86C5E04AAB1" box="[982,1132,1948,1973]" name="New Zealand" pageId="8">New Zealand</collectingCountry>
and
<collectingCountry id="A88E766EC975FFEAFB0AF86D5F62AAB1" box="[1187,1290,1948,1973]" name="Australia" pageId="8">Australia</collectingCountry>
(
<bibRefCitation id="B4084B0FC975FFEAFAB0F86D5B04AADD" author="Jaloszynski" pageId="8" refString="Jaloszynski, P. &amp; Newton, A. F., Jr. (2017) Taxonomy of ' Euconnus complex'. Part XV. Disentangling the Euconnus - Sciacharis knot (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa, 4344 (3), 401 ¯ 443. http: // dx. doi. org / 10.11646 / zootaxa. 4344.3.1" type="journal article" year="2017">Jałoszyński &amp; Newton (2017))</bibRefCitation>
does not support such a placement, although some unrevised species of
<taxonomicName id="17994D7DC975FFEAFB1AF8305F41AADD" authorityName="Jałoszyński" authorityYear="2017" box="[1203,1321,1985,2009]" class="Insecta" family="Staphylinidae" genus="Sciacharis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" phylum="Arthropoda" rank="genus">
<emphasis id="E2EDEAECC975FFEAFB1AF8305F41AADD" box="[1203,1321,1985,2009]" italics="true" pageId="8">Sciacharis</emphasis>
</taxonomicName>
may hide under the name
<taxonomicName id="17994D7DC974FFEBFEFBFF685BA8ADB4" baseAuthorityName="Jaloszynski" baseAuthorityYear="2017" box="[338,448,153,176]" class="Insecta" family="Scydmaenidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="9" phylum="Arthropoda" rank="genus">
<emphasis id="E2EDEAECC974FFEBFEFBFF685BA8ADB4" box="[338,448,153,176]" italics="true" pageId="9">Euconnus</emphasis>
</taxonomicName>
and may have a broader distribution. The maxillary palps and male genital structures may be important to clarify the status of
<taxonomicName id="17994D7DC974FFEBFDD8FF4D5881ADD0" authorityName="Casey" authorityYear="1897" box="[625,745,188,212]" class="Insecta" family="Staphylinidae" genus="Noctophus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" phylum="Arthropoda" rank="genus">
<emphasis id="E2EDEAECC974FFEBFDD8FF4D5881ADD0" box="[625,745,188,212]" italics="true" pageId="9">Noctophus</emphasis>
</taxonomicName>
, especially the aedeagus, which in
<taxonomicName id="17994D7DC974FFEBFB28FF4D5E9FADD0" authorityName="Jałoszyński" authorityYear="2017" box="[1153,1271,188,212]" class="Insecta" family="Staphylinidae" genus="Sciacharis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" phylum="Arthropoda" rank="genus">
<emphasis id="E2EDEAECC974FFEBFB28FF4D5E9FADD0" box="[1153,1271,188,212]" italics="true" pageId="9">Sciacharis</emphasis>
</taxonomicName>
is remarkably thin-walled and with a weakly sclerotized endophallus. As long as males remain unknown, the problem of relationships between
<taxonomicName id="17994D7DC974FFEBFE3BFEF55862AC18" authorityName="Casey" authorityYear="1897" box="[402,522,260,284]" class="Insecta" family="Staphylinidae" genus="Noctophus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" phylum="Arthropoda" rank="genus">
<emphasis id="E2EDEAECC974FFEBFE3BFEF55862AC18" box="[402,522,260,284]" italics="true" pageId="9">Noctophus</emphasis>
</taxonomicName>
and
<taxonomicName id="17994D7DC974FFEBFDE9FEF558DEAC18" authorityName="Jałoszyński" authorityYear="2017" box="[576,694,260,284]" class="Insecta" family="Staphylinidae" genus="Sciacharis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" phylum="Arthropoda" rank="genus">
<emphasis id="E2EDEAECC974FFEBFDE9FEF558DEAC18" box="[576,694,260,284]" italics="true" pageId="9">Sciacharis</emphasis>
</taxonomicName>
cannot be solved.
<emphasis id="E2EDEAECC974FFEBFC21FEF5593EAC44" bold="true" pageId="9">
Tentatively,
<taxonomicName id="17994D7DC974FFEBFBB0FEF25EFDAC18" authorityName="Casey" authorityYear="1897" box="[1049,1173,259,284]" class="Insecta" family="Staphylinidae" genus="Noctophus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" phylum="Arthropoda" rank="genus">
<emphasis id="E2EDEAECC974FFEBFBB0FEF25EFDAC18" bold="true" box="[1049,1173,259,284]" italics="true" pageId="9">Noctophus</emphasis>
</taxonomicName>
is here removed from
<taxonomicName id="17994D7DC974FFEBFF3EFED95B65AC3B" baseAuthorityName="Jaloszynski" baseAuthorityYear="2017" box="[151,269,296,319]" class="Insecta" family="Scydmaenidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="9" phylum="Arthropoda" rank="genus">
<emphasis id="E2EDEAECC974FFEBFF3EFED95B65AC3B" bold="true" box="[151,269,296,319]" italics="true" pageId="9">Euconnus</emphasis>
</taxonomicName>
and restituted as a genus, pending further study.
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>