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<document id="FB74628D5C9CE2E5D3456B57F792B60F" ID-CLB-Dataset="264199" ID-DOI="10.11646/zootaxa.5332.1.1" ID-GBIF-Dataset="66e0ec51-d494-43d4-965e-a2cd1462ef54" ID-ISSN="1175-5326" ID-Zenodo-Dep="8261502" ID-ZooBank="424F7439-4095-46A5-93E3-C4130E3B6D9A" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="juliana" checkinTime="1692350092877" checkinUser="plazi" docAuthor="POORANI, J." docDate="2023" docId="03C74162148746D5BDDF3001FE4FFE37" docLanguage="en" docName="zootaxa.5332.1.1.pdf" docOrigin="Zootaxa 5332 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5332.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Oenopia smetanai Canepari" docType="treatment" docVersion="8" lastPageNumber="227" masterDocId="FFFE391A14584637BD48367CFFCDFF9B" masterDocTitle="An illustrated guide to lady beetles (Coleoptera: Coccinellidae) of the Indian Subcontinent. Part 1. Tribe Coccinellini" masterLastPageNumber="307" masterPageNumber="1" pageNumber="224" updateTime="1699593964221" updateUser="ExternalLinkService">
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<mods:title id="B07AE4F2B96C59F8D8A5F09C89584FEF">An illustrated guide to lady beetles (Coleoptera: Coccinellidae) of the Indian Subcontinent. Part 1. Tribe Coccinellini</mods:title>
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<treatment id="03C74162148746D5BDDF3001FE4FFE37" ID-DOI="http://doi.org/10.5281/zenodo.8273858" ID-GBIF-Taxon="212449986" ID-Zenodo-Dep="8273858" LSID="urn:lsid:plazi:treatment:03C74162148746D5BDDF3001FE4FFE37" httpUri="http://treatment.plazi.org/id/03C74162148746D5BDDF3001FE4FFE37" lastPageId="226" lastPageNumber="227" pageId="223" pageNumber="224">
<subSubSection id="C374A3FF148746E8BDDF3001FE22F90C" box="[151,495,1661,1687]" pageId="223" pageNumber="224" type="nomenclature">
<paragraph id="8BD1F074148746E8BDDF3001FE22F90C" blockId="223.[151,495,1661,1723]" box="[151,495,1661,1687]" pageId="223" pageNumber="224">
<heading id="D0994718148746E8BDDF3001FE22F90C" bold="true" box="[151,495,1661,1687]" fontSize="11" level="2" pageId="223" pageNumber="224" reason="4">
<taxonomicName id="4C6E8BF7148746E8BDDF3001FE22F90C" authority="Canepari" authorityName="Canepari" box="[151,495,1661,1687]" class="Insecta" family="Coccinellidae" genus="Oenopia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="223" pageNumber="224" phylum="Arthropoda" rank="species" species="smetanai">
<emphasis id="B91A2C66148746E8BDDF3001FE22F90C" bold="true" box="[151,495,1661,1687]" pageId="223" pageNumber="224">
<emphasis id="B91A2C66148746E8BDDF3001FEBFF90C" bold="true" box="[151,370,1661,1687]" italics="true" pageId="223" pageNumber="224">Oenopia smetanai</emphasis>
Canepari
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C374A3FF148746E8BDDF30DDFEC5F920" box="[151,264,1697,1723]" pageId="223" pageNumber="224" type="description">
<paragraph id="8BD1F074148746E8BDDF30DDFEC5F920" blockId="223.[151,495,1661,1723]" box="[151,264,1697,1723]" pageId="223" pageNumber="224">
(
<figureCitation id="1355ECF1148746E8BDD730DDFF32F920" box="[159,255,1697,1723]" captionStart="FIGURE 161" captionStartId="225.[151,250,1943,1967]" captionTargetBox="[301,1285,181,1918]" captionTargetId="figure-15@225.[301,1285,181,1919]" captionTargetPageId="225" captionText="FIGURE 161. Oenopia smetanai Canepari: ad. adult, variants (BMNH); e. nominate form, dorsal view; fh. male genitalia: f. tegmen, lateral view; g. tegmen, ventral view; h. penis." figureDoi="http://doi.org/10.5281/zenodo.8262073" httpUri="https://zenodo.org/record/8262073/files/figure.png" pageId="223" pageNumber="224">Fig. 161</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C374A3FF148746E8BDDF309AFF39F886" pageId="223" pageNumber="224" type="reference_group">
<paragraph id="8BD1F074148746E8BDDF309AFF39F886" blockId="223.[151,1437,1766,1821]" pageId="223" pageNumber="224">
<treatmentCitationGroup id="AB7ED75A148746E8BDDF309AFF39F886" pageId="223" pageNumber="224">
<taxonomicName id="4C6E8BF7148746E8BDDF309AFDD0F965" authority="Canepari, 1997: 53" authorityName="Canepari" authorityPageNumber="53" authorityYear="1997" box="[151,541,1766,1790]" class="Insecta" family="Coccinellidae" genus="Oenopia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="223" pageNumber="224" phylum="Arthropoda" rank="species" species="smetanai">
<emphasis id="B91A2C66148746E8BDDF309AFE9CF966" box="[151,337,1766,1789]" italics="true" pageId="223" pageNumber="224">Oenopia smetanai</emphasis>
Canepari, 1997: 53
</taxonomicName>
(
<typeStatus id="54D54ED6148746E8BF64309AFD41F965" box="[556,652,1766,1790]" pageId="223" pageNumber="224" type="holotype">Holotype</typeStatus>
male, MHNG; Type locality:
<collectingCountry id="F379B0E4148746E8BE8B309AFBCCF965" box="[963,1025,1766,1790]" name="Nepal" pageId="223" pageNumber="224">Nepal</collectingCountry>
).—
<treatmentCitation id="0ACFD665148746E8B96D309AFB39F965" author="Poorani, J." box="[1061,1268,1766,1790]" page="337" pageId="223" pageNumber="224" year="2002">
<bibRefCitation id="EFFF8D85148746E8B96D309AFB39F965" author="Poorani, J." box="[1061,1268,1766,1790]" pageId="223" pageNumber="224" pagination="307 - 383" refId="ref118931" refString="Poorani, J. (2002 a) An annotated checklist of the Coccinellidae (excluding Epilachninae) of the Indian subregion. Oriental Insects, 36, 307 - 383. https: // doi. org / 10.1080 / 00305316.2002.10417335" type="journal article" year="2002">Poorani 2002a: 337</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0ACFD665148746E8B9B7309AFF3DF886" author="Poorani, J." page="104" pageId="223" pageNumber="224" year="2002">
<bibRefCitation id="EFFF8D85148746E8B9B7309AFF3DF886" author="Poorani, J." pageId="223" pageNumber="224" pagination="97 - 116" refId="ref118973" refString="Poorani, J. (2002 b) A review of the genus Oenopia Mulsant (Coleoptera: Coccinellidae) of the Indian subcontinent, with description of a new species. Oriental Insects, 36, 97 - 116. https: // doi. org / 10.1080 / 00305316.2002.10417328" type="journal article" year="2002">Poorani 2002b: 104</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C374A3FF148746E8BDDF3136FDFCF86E" pageId="223" pageNumber="224" type="diagnosis">
<paragraph id="8BD1F074148746E8BDDF3136FDFCF86E" blockId="223.[151,1437,1866,2037]" pageId="223" pageNumber="224">
<emphasis id="B91A2C66148746E8BDDF3136FEDFF8FF" bold="true" box="[151,274,1866,1892]" pageId="223" pageNumber="224">Diagnosis.</emphasis>
Length: 2.503.00 mm. Form small, broad oval, dorsum moderately convex, glabrous. Ground colour pale creamy yellow; pronotum with a pair of oblique oval median black spots above posterior margin; scutellar shield black; elytra with a vase-shaped elongate sutural macula and three black spots on each elytron in the nominate form (
<figureCitation id="1355ECF1148746E8BD9731CBFEA7F84A" box="[223,362,1975,2001]" captionStart="FIGURE 161" captionStartId="225.[151,250,1943,1967]" captionTargetBox="[301,1285,181,1918]" captionTargetId="figure-15@225.[301,1285,181,1919]" captionTargetPageId="225" captionText="FIGURE 161. Oenopia smetanai Canepari: ad. adult, variants (BMNH); e. nominate form, dorsal view; fh. male genitalia: f. tegmen, lateral view; g. tegmen, ventral view; h. penis." figureDoi="http://doi.org/10.5281/zenodo.8262073" httpUri="https://zenodo.org/record/8262073/files/figure.png" pageId="223" pageNumber="224">Fig. 161c, e</figureCitation>
), elytral pattern variable with the spots fused or reduced as illustrated (
<figureCitation id="1355ECF1148746E8B9E831CBFA81F84A" box="[1184,1356,1975,2001]" captionStart="FIGURE 161" captionStartId="225.[151,250,1943,1967]" captionTargetBox="[301,1285,181,1918]" captionTargetId="figure-15@225.[301,1285,181,1919]" captionTargetPageId="225" captionText="FIGURE 161. Oenopia smetanai Canepari: ad. adult, variants (BMNH); e. nominate form, dorsal view; fh. male genitalia: f. tegmen, lateral view; g. tegmen, ventral view; h. penis." figureDoi="http://doi.org/10.5281/zenodo.8262073" httpUri="https://zenodo.org/record/8262073/files/figure.png" pageId="223" pageNumber="224">Fig. 161a, b, d</figureCitation>
). Male genitalia (
<figureCitation id="1355ECF1148746E8BC4F31A7FE41F86E" box="[263,396,2011,2037]" captionStart="FIGURE 161" captionStartId="225.[151,250,1943,1967]" captionTargetBox="[301,1285,181,1918]" captionTargetId="figure-15@225.[301,1285,181,1919]" captionTargetPageId="225" captionText="FIGURE 161. Oenopia smetanai Canepari: ad. adult, variants (BMNH); e. nominate form, dorsal view; fh. male genitalia: f. tegmen, lateral view; g. tegmen, ventral view; h. penis." figureDoi="http://doi.org/10.5281/zenodo.8262073" httpUri="https://zenodo.org/record/8262073/files/figure.png" pageId="223" pageNumber="224">Fig. 161fh</figureCitation>
) as illustrated.
</paragraph>
</subSubSection>
<caption id="DF11A0FC14B846D7BDDF31EBFED0F863" ID-DOI="http://doi.org/10.5281/zenodo.8262071" ID-Zenodo-Dep="8262071" httpUri="https://zenodo.org/record/8262071/files/figure.png" pageId="224" pageNumber="225" startId="224.[151,250,1943,1967]" targetBox="[212,1364,192,1911]" targetPageId="224" targetType="figure">
<paragraph id="8BD1F07414B846D7BDDF31EBFED0F863" blockId="224.[151,1436,1943,2040]" pageId="224" pageNumber="225">
<emphasis id="B91A2C6614B846D7BDDF31EBFEE0F834" bold="true" box="[151,301,1943,1967]" pageId="224" pageNumber="225">FIGURE 160.</emphasis>
<taxonomicName id="4C6E8BF714B846D7BC7C31E4FDA9F82B" authority="(Mulsant)" baseAuthorityName="Mulsant" box="[308,612,1944,1968]" class="Insecta" family="Coccinellidae" genus="Oenopia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="224" pageNumber="225" phylum="Arthropoda" rank="species" species="signatella">
<emphasis id="B91A2C6614B846D7BC7C31E4FE3BF82B" box="[308,502,1944,1968]" italics="true" pageId="224" pageNumber="225">Oenopia signatella</emphasis>
(Mulsant)
</taxonomicName>
: a. adult, dorsal view; b. abdominal postcoxal line, male; c. abdominal postcoxal line, female; dg. male genitalia: d. tegmen, lateral view; e. tegmen, ventral view; f. penis; g. penis apex; h. female genitalia; i. spermatheca.
</paragraph>
</caption>
<caption id="DF11A0FC14B946D6BDDF31EBFD08F848" ID-DOI="http://doi.org/10.5281/zenodo.8262073" ID-Zenodo-Dep="8262073" httpUri="https://zenodo.org/record/8262073/files/figure.png" pageId="225" pageNumber="226" startId="225.[151,250,1943,1967]" targetBox="[301,1285,181,1918]" targetPageId="225" targetType="figure">
<paragraph id="8BD1F07414B946D6BDDF31EBFD08F848" blockId="225.[151,1437,1943,2004]" pageId="225" pageNumber="226">
<emphasis id="B91A2C6614B946D6BDDF31EBFEE1F834" bold="true" box="[151,300,1943,1967]" pageId="225" pageNumber="226">FIGURE 161.</emphasis>
<taxonomicName id="4C6E8BF714B946D6BC7A31E4FD9CF834" authority="Canepari" authorityName="Canepari" box="[306,593,1943,1967]" class="Insecta" family="Coccinellidae" genus="Oenopia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="225" pageNumber="226" phylum="Arthropoda" rank="species" species="smetanai">
<emphasis id="B91A2C6614B946D6BC7A31E4FE27F834" box="[306,490,1944,1967]" italics="true" pageId="225" pageNumber="226">Oenopia smetanai</emphasis>
Canepari
</taxonomicName>
: ad. adult, variants (BMNH); e. nominate form, dorsal view; fh. male genitalia: f. tegmen, lateral view; g. tegmen, ventral view; h. penis.
</paragraph>
</caption>
<subSubSection id="C374A3FF14BA46D5BD8F36EAFCD3FF4E" pageId="226" pageNumber="227" type="distribution">
<paragraph id="8BD1F07414BA46D5BD8F36EAFC2DFF2A" blockId="226.[151,1436,150,429]" box="[199,992,150,177]" pageId="226" pageNumber="227">
<emphasis id="B91A2C6614BA46D5BD8F36EAFEACFF2B" bold="true" box="[199,353,150,176]" pageId="226" pageNumber="227">Distribution.</emphasis>
<collectingCountry id="F379B0E414BA46D5BC2F36EBFE6CFF2A" box="[359,417,151,177]" name="India" pageId="226" pageNumber="227">India</collectingCountry>
(
<collectingRegion id="49AA3E9614BA46D5BCF836EBFD8FFF2A" box="[432,578,151,177]" country="India" name="Uttarakhand" pageId="226" pageNumber="227">Uttarakhand</collectingRegion>
); Eastern Himalayas;
<collectingCountry id="F379B0E414BA46D5BE7D36EBFCB6FF2A" box="[821,891,151,177]" name="Nepal" pageId="226" pageNumber="227">Nepal</collectingCountry>
;
<collectingCountry id="F379B0E414BA46D5BECF36EBFC11FF2A" box="[903,988,151,177]" name="Bhutan" pageId="226" pageNumber="227">Bhutan</collectingCountry>
.
</paragraph>
<paragraph id="8BD1F07414BA46D5BD8F36C6FCD3FF4E" blockId="226.[151,1436,150,429]" box="[199,798,186,213]" pageId="226" pageNumber="227">
<emphasis id="B91A2C6614BA46D5BD8F36C6FE0CFF4E" bold="true" box="[199,449,186,213]" pageId="226" pageNumber="227">Seasonal occurrence.</emphasis>
Collected in April (label data).
</paragraph>
</subSubSection>
<subSubSection id="C374A3FF14BA46D5BD8F36A3FE4FFE37" pageId="226" pageNumber="227" type="discussion">
<paragraph id="8BD1F07414BA46D5BD8F36A3FE4FFE37" blockId="226.[151,1436,150,429]" pageId="226" pageNumber="227">
<emphasis id="B91A2C6614BA46D5BD8F36A3FEDCFF62" bold="true" box="[199,273,223,249]" pageId="226" pageNumber="227">Notes.</emphasis>
Canepari (2012) synonymized
<taxonomicName id="4C6E8BF714BA46D5BF2736A3FD3BFF63" box="[623,758,223,249]" class="Insecta" family="Coccinellidae" genus="Oenopia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="226" pageNumber="227" phylum="Arthropoda" rank="species" species="smetanai">
<emphasis id="B91A2C6614BA46D5BF2736A3FD3BFF63" box="[623,758,223,249]" italics="true" pageId="226" pageNumber="227">O. smetanai</emphasis>
</taxonomicName>
with
<taxonomicName id="4C6E8BF714BA46D5BE7C36A3FC2AFF62" authorityName="Jing" authorityYear="1986" box="[820,999,223,249]" class="Insecta" family="Coccinellidae" genus="Oenopia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="226" pageNumber="227" phylum="Arthropoda" rank="species" species="flavidbrunna">
<emphasis id="B91A2C6614BA46D5BE7C36A3FC2AFF62" box="[820,999,223,249]" italics="true" pageId="226" pageNumber="227">O. flavidbrunna</emphasis>
</taxonomicName>
, apparently unaware that
<bibRefCitation id="EFFF8D8514BA46D5B84D36A3FA51FF62" author="Kovar, I." box="[1285,1436,223,249]" pageId="226" pageNumber="227" pagination="1807" refId="ref116365" refString="Kovar, I. (2007) Family Coccinellidae Latreille, 1807. In: L ˆ bl, I. &amp; Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 4. Apollo Books, Stenstrup, pp. 568 - 631." type="book chapter" year="2007">Kovář (2007)</bibRefCitation>
treated
<taxonomicName id="4C6E8BF714BA46D5BDA1377FFE56FE86" authorityName="Jing" authorityYear="1986" box="[233,411,259,285]" class="Insecta" family="Coccinellidae" genus="Oenopia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="226" pageNumber="227" phylum="Arthropoda" rank="species" species="flavidbrunna">
<emphasis id="B91A2C6614BA46D5BDA1377FFE56FE86" box="[233,411,259,285]" italics="true" pageId="226" pageNumber="227">O. flavidbrunna</emphasis>
</taxonomicName>
as a junior synonym of
<taxonomicName id="4C6E8BF714BA46D5BFE9377FFCC7FE86" box="[673,778,259,285]" class="Insecta" family="Coccinellidae" genus="Oenopia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="226" pageNumber="227" phylum="Arthropoda" rank="species" species="billieti">
<emphasis id="B91A2C6614BA46D5BFE9377FFCC7FE86" box="[673,778,259,285]" italics="true" pageId="226" pageNumber="227">O. billieti</emphasis>
</taxonomicName>
which is also incorrect. Here
<taxonomicName id="4C6E8BF714BA46D5B919377FFAD1FE87" box="[1105,1308,259,285]" class="Insecta" family="Coccinellidae" genus="Oenopia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="226" pageNumber="227" phylum="Arthropoda" rank="species" species="smetanai">
<emphasis id="B91A2C6614BA46D5B919377FFAD1FE87" box="[1105,1308,259,285]" italics="true" pageId="226" pageNumber="227">Oenopia smetanai</emphasis>
</taxonomicName>
is removed from synonymy with
<taxonomicName id="4C6E8BF714BA46D5BCC4375BFCCEFEDA" authority="Jing, 1986" authorityName="Jing" authorityYear="1986" box="[396,771,295,321]" class="Insecta" family="Coccinellidae" genus="Oenopia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="226" pageNumber="227" phylum="Arthropoda" rank="species" species="flavidbrunna">
<emphasis id="B91A2C6614BA46D5BCC4375BFD48FEDA" box="[396,645,295,321]" italics="true" pageId="226" pageNumber="227">Oenopia flavidbrunna</emphasis>
<bibRefCitation id="EFFF8D8514BA46D5BFC5375BFCCEFEDA" author="Jing, X. L." box="[653,771,295,321]" pageId="226" pageNumber="227" pagination="203 - 207" refId="ref114545" refString="Jing, X. L. (1986) Three new species of Oenopia Mulsant (Coccinellidae: Coleoptera). Acta Entomologica Sinica, 29 (2), 203 - 207." type="journal article" year="1986">Jing, 1986</bibRefCitation>
</taxonomicName>
and treated as a valid species (
<emphasis id="B91A2C6614BA46D5B92D375BFAC5FEDB" bold="true" box="[1125,1288,294,321]" pageId="226" pageNumber="227">status revised</emphasis>
) because the male genitalia of
<taxonomicName id="4C6E8BF714BA46D5BC143737FDDDFEFE" authorityName="Jing" authorityYear="1986" box="[348,528,331,357]" class="Insecta" family="Coccinellidae" genus="Oenopia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="226" pageNumber="227" phylum="Arthropoda" rank="species" species="flavidbrunna">
<emphasis id="B91A2C6614BA46D5BC143737FDDDFEFE" box="[348,528,331,357]" italics="true" pageId="226" pageNumber="227">O. flavidbrunna</emphasis>
</taxonomicName>
illustrated by
<bibRefCitation id="EFFF8D8514BA46D5BFFA3737FCFEFEFE" author="Jing, X. L." box="[690,819,331,357]" pageId="226" pageNumber="227" pagination="203 - 207" refId="ref114545" refString="Jing, X. L. (1986) Three new species of Oenopia Mulsant (Coccinellidae: Coleoptera). Acta Entomologica Sinica, 29 (2), 203 - 207." type="journal article" year="1986">Jing (1986)</bibRefCitation>
are totally different from those of
<taxonomicName id="4C6E8BF714BA46D5B9F33737FA8FFEFF" box="[1211,1346,331,357]" class="Insecta" family="Coccinellidae" genus="Oenopia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="226" pageNumber="227" phylum="Arthropoda" rank="species" species="smetanai">
<emphasis id="B91A2C6614BA46D5B9F33737FA8FFEFF" box="[1211,1346,331,357]" italics="true" pageId="226" pageNumber="227">O. smetanai</emphasis>
</taxonomicName>
and the body outline of
<taxonomicName id="4C6E8BF714BA46D5BC0F3713FE34FE12" authorityName="Jing" authorityYear="1986" box="[327,505,367,393]" class="Insecta" family="Coccinellidae" genus="Oenopia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="226" pageNumber="227" phylum="Arthropoda" rank="species" species="flavidbrunna">
<emphasis id="B91A2C6614BA46D5BC0F3713FE34FE12" box="[327,505,367,393]" italics="true" pageId="226" pageNumber="227">O. flavidbrunna</emphasis>
</taxonomicName>
illustrated by
<bibRefCitation id="EFFF8D8514BA46D5BFDF3713FCCAFE12" author="Yu, G." box="[663,775,367,393]" pageId="226" pageNumber="227" refId="ref124811" refString="Yu, G. (2010) Chinese Lady Beetles (The Subfamily Coccinellinae). China Scientific Press, Beijing, 180 pp. [in Chinese]" type="book" year="2010">Yu (2010)</bibRefCitation>
is also more elongate oval and different from the short oval form of
<emphasis id="B91A2C6614BA46D5BDBB37EFFE4FFE37" box="[243,386,403,429]" italics="true" pageId="226" pageNumber="227">
<taxonomicName id="4C6E8BF714BA46D5BDBB37EFFEB3FE37" box="[243,382,403,429]" class="Insecta" family="Coccinellidae" genus="Oenopia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="226" pageNumber="227" phylum="Arthropoda" rank="species" species="smetanai">O. smetanai</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>