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Genus
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<emphasis id="B91A2C661420464FBDA63370FEA1FABD" bold="true" box="[238,364,1292,1318]" italics="true" pageId="120" pageNumber="121">Harmonia</emphasis>
Mulsant
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<emphasis id="B91A2C661420464FBDDF332DFF32FAF3" box="[151,255,1361,1384]" italics="true" pageId="120" pageNumber="121">Harmonia</emphasis>
<bibRefCitation id="EFFF8D851420464FBC40332CFE18FAF2" author="Mulsant, E." box="[264,469,1360,1385]" pageId="120" pageNumber="121" refId="ref117813" refString="Mulsant, E. (1846) Securipalpes. Vol. 4. Histoire Naturelle des Coleopteres de France, Paris, 380 pp., 1 pl." type="book" year="1846">Mulsant, 1846: 108</bibRefCitation>
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;
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<bibRefCitation id="EFFF8D851420464FBCA9332CFDF2FAF3" author="Mulsant, E." box="[481,575,1360,1384]" pageId="120" pageNumber="121" pagination="1 - 1104" refId="ref117842" refString="Mulsant, E. (1850) Species des Coleopteres Trimeres Securipalpes. Annales des Sciences Physiques et Naturelles, d'Agriculture et d'Industrie, publiees par la Societe nationale d'Agriculture, etc., de Lyon, Deuxieme Serie, 2, 1 - 1104. https: // doi. org / 10.5962 / bhl. title. 8953" type="journal article" year="1850">1850: 75</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0ACFD6651420464FBF04332CFD6BFAF3" author="Mulsant, E." box="[588,678,1360,1384]" page="55" pageId="120" pageNumber="121" year="1866">
<bibRefCitation id="EFFF8D851420464FBF04332CFD6BFAF3" author="Mulsant, E." box="[588,678,1360,1384]" pageId="120" pageNumber="121" refId="ref117962" refString="Mulsant, E. (1866) Monographie des Coccinellides. 1 re Partie Coccinelliens. Savy et Deyrolle, Paris, 294 pp." type="book" year="1866">1866: 55</bibRefCitation>
</treatmentCitation>
.—Crotch 1871: 3;
<treatmentCitation id="0ACFD6651420464FBE3B332CFBD1FAF2" author="Mader, L." box="[883,1052,1360,1385]" page="19" pageId="120" pageNumber="121" year="1926">
<bibRefCitation id="EFFF8D851420464FBE3B332CFBD1FAF2" author="Mader, L." box="[883,1052,1360,1385]" pageId="120" pageNumber="121" pagination="289 - 336" refId="ref116941" refString="Mader, L. (1926 - 1934) Evidenz der palaarktischen Coccinelliden nnd ihrer aberrationen in Wort und Bild, l-Teil, Epilachnini, Coccinellini, Halyziini und Synonychini. Zeitschrift des Vereins der Naturbeobachter und Sammler, Wien, 1926 (1), 1927 (2), 1928 (3), 1929 (4), 1930 (5), 1931 (6), 1932 (7), 1933 (8), 1934 (9), 1 - 24, 25 - 48, 49 - 76, 77 - 124, 124 - 168, 169 - 204, 205 - 244, 245 - 288, 289 - 336." type="journal article" year="1926">Mader 1926: 19</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0ACFD6651420464FB960332CFACFFAF2" author="Timberlake, P. H." box="[1064,1282,1360,1385]" page="17" pageId="120" pageNumber="121" year="1943">
<bibRefCitation id="EFFF8D851420464FB960332CFACFFAF2" author="Timberlake, P. H." box="[1064,1282,1360,1385]" pageId="120" pageNumber="121" pagination="1 - 67" refId="ref123429" refString="Timberlake, P. H. (1943) The Coccinellidae or lady beetles of the Koebele collection - Part I. Hawaiian Planters' Record, 47, 1 - 67." type="journal article" year="1943">Timberlake 1943: 17</bibRefCitation>
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;
<treatmentCitation id="0ACFD6651420464FB847332DFF2EFA1C" author="Watson, W. Y." page="48" pageId="120" pageNumber="121" year="1956">
<bibRefCitation id="EFFF8D851420464FB847332DFF2EFA1C" author="Watson, W. Y." pageId="120" pageNumber="121" pagination="1 - 52" refId="ref123932" refString="Watson, W. Y. (1956) A study of the phylogeny of the genera of the tribe Coccinellini (Coleoptera). Contributions of the Royal Ontario Museum, Division of Zoology and Palaentology, 42, 1 - 52. https: // doi. org / 10.5962 / bhl. title. 52186" type="journal article" year="1956">Watson 1956: 48</bibRefCitation>
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;
<treatmentCitation id="0ACFD6651420464FBDB93313FDE6FA1C" author="Iablokoff-Khnzorian, S. M." box="[241,555,1391,1415]" page="71" pageId="120" pageNumber="121" year="1979">
<bibRefCitation id="EFFF8D851420464FBDB93313FDE6FA1C" author="Iablokoff-Khnzorian, S. M." box="[241,555,1391,1415]" pageId="120" pageNumber="121" pagination="37 - 75" refId="ref114003" refString="Iablokoff-Khnzorian, S. M. (1979) Genera der Palaarktischen Coccinellini (Coleoptera: Coccinellidae). Entomologische Blatter fur Biologie und Systematik der Kafer, 75 (1 - 2), 37 - 75." type="journal article" year="1979">Iablokoff-Khnzorian 1979: 71</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0ACFD6651420464FBF713313FD6BFA1C" author="Iablokoff-Khnzorian, S. M." box="[569,678,1391,1415]" page="456" pageId="120" pageNumber="121" year="1982">
<bibRefCitation id="EFFF8D851420464FBF713313FD6BFA1C" author="Iablokoff-Khnzorian, S. M." box="[569,678,1391,1415]" pageId="120" pageNumber="121" refId="ref114042" refString="Iablokoff-Khnzorian, S. M. (1982) Les Coccinelles Coleopteres-Coccinellidae. Tribu Coccinellini des regions Palearctique et Orientale. Boubee, Paris, 568 pp. [in French]" type="book" year="1982">1982: 456</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
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<subSubSection id="C374A3FF1420464FBFFA330CFB88FA3D" pageId="120" pageNumber="121" type="type_taxon">
<paragraph id="8BD1F0741420464FBFFA330CFB88FA3D" blockId="120.[151,1437,1360,1845]" pageId="120" pageNumber="121">
<typeStatus id="54D54ED61420464FBFFA330CFD29FA13" box="[690,740,1392,1416]" pageId="120" pageNumber="121">Type</typeStatus>
species:
<taxonomicName id="4C6E8BF71420464FBE003313FAC7FA1C" authority="Schaller, 1783" authorityName="Schaller" authorityYear="1783" box="[840,1290,1391,1415]" class="Insecta" family="Coccinellidae" genus="Coccinella" kingdom="Animalia" order="Coleoptera" pageId="120" pageNumber="121" phylum="Arthropoda" rank="species" species="marginepunctata">
<emphasis id="B91A2C661420464FBE003313FBA1FA1C" box="[840,1132,1391,1415]" italics="true" pageId="120" pageNumber="121">Coccinella marginepunctata</emphasis>
Schaller, 1783
</taxonomicName>
(=
<taxonomicName id="4C6E8BF71420464FB87C330CFDE1FA3D" authority="Pontoppidan, 1763" authorityName="Pontoppidan" authorityYear="1763" class="Insecta" family="Coccinellidae" genus="Harmonia" kingdom="Animalia" order="Coleoptera" pageId="120" pageNumber="121" phylum="Arthropoda" rank="species" species="quadripunctata">
<emphasis id="B91A2C661420464FB87C330CFEAEFA3D" italics="true" pageId="120" pageNumber="121">Harmonia quadripunctata</emphasis>
<bibRefCitation id="EFFF8D851420464FBC2233F2FDE1FA3D" author="Pontoppidan, E." box="[362,556,1422,1446]" pageId="120" pageNumber="121" refId="ref118814" refString="Pontoppidan, E. (1763) Den Danske Atlas. Tomus 1, 4 °. Godiche, Ki ˆ benhavn, 669 pp." type="book" year="1763">Pontoppidan, 1763</bibRefCitation>
</taxonomicName>
), by subsequent designation of
<treatmentCitation id="0ACFD6651420464FBE2333F2FB8CFA3D" author="Timberlake, P. H." box="[875,1089,1422,1446]" page="17" pageId="120" pageNumber="121" year="1943">
<bibRefCitation id="EFFF8D851420464FBE2333F2FB8CFA3D" author="Timberlake, P. H." box="[875,1089,1422,1446]" pageId="120" pageNumber="121" pagination="1 - 67" refId="ref123429" refString="Timberlake, P. H. (1943) The Coccinellidae or lady beetles of the Koebele collection - Part I. Hawaiian Planters' Record, 47, 1 - 67." type="journal article" year="1943">Timberlake 1943: 17</bibRefCitation>
</treatmentCitation>
.
</paragraph>
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<subSubSection id="C374A3FF1420464FBDDF33D1FBA2F8AF" pageId="120" pageNumber="121" type="reference_group">
<paragraph id="8BD1F0741420464FBDDF33D1FDF2FA78" blockId="120.[151,1437,1360,1845]" pageId="120" pageNumber="121">
<treatmentCitationGroup id="AB7ED75A1420464FBDDF33D1FDF2FA78" pageId="120" pageNumber="121">
<treatmentCitation id="0ACFD6651420464FBDDF33D1FE6FFA5E" author="Mulsant, E." box="[151,418,1452,1477]" page="241" pageId="120" pageNumber="121" year="1850">
<taxonomicName id="4C6E8BF71420464FBDDF33D1FE6FFA5E" authority="Mulsant, 1850: 241" authorityName="Mulsant" authorityPageNumber="241" authorityYear="1850" box="[151,418,1452,1477]" class="Insecta" genus="Leis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="120" pageNumber="121" phylum="Arthropoda" rank="genus">
<emphasis id="B91A2C661420464FBDDF33D1FF0CFA5F" box="[151,193,1453,1476]" italics="true" pageId="120" pageNumber="121">Leis</emphasis>
<bibRefCitation id="EFFF8D851420464FBD8633D0FE6FFA5E" author="Mulsant, E." box="[206,418,1452,1477]" pageId="120" pageNumber="121" pagination="1 - 1104" refId="ref117842" refString="Mulsant, E. (1850) Species des Coleopteres Trimeres Securipalpes. Annales des Sciences Physiques et Naturelles, d'Agriculture et d'Industrie, publiees par la Societe nationale d'Agriculture, etc., de Lyon, Deuxieme Serie, 2, 1 - 1104. https: // doi. org / 10.5962 / bhl. title. 8953" type="journal article" year="1850">Mulsant, 1850: 241</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
<typeStatus id="54D54ED61420464FBCFA33D1FE29FA5E" box="[434,484,1453,1477]" pageId="120" pageNumber="121">Type</typeStatus>
species:
<taxonomicName id="4C6E8BF71420464FBF0533D0FC58FA5F" authority="Fabricius" authorityName="Fabricius" box="[589,917,1452,1476]" class="Insecta" family="Coccinellidae" genus="Coccinella" kingdom="Animalia" order="Coleoptera" pageId="120" pageNumber="121" phylum="Arthropoda" rank="species" species="dimidiata">
<emphasis id="B91A2C661420464FBF0533D0FCE5FA5F" box="[589,808,1452,1476]" italics="true" pageId="120" pageNumber="121">Coccinella dimidiata</emphasis>
Fabricius
</taxonomicName>
, by subsequent designation (Crotch 1874: 119). Synonymized by
<treatmentCitation id="0ACFD6651420464FBC3133B7FDF6FA78" author="Miyatake, M." box="[377,571,1483,1507]" page="60" pageId="120" pageNumber="121" year="1965">
<bibRefCitation id="EFFF8D851420464FBC3133B7FDF6FA78" author="Miyatake, M." box="[377,571,1483,1507]" pageId="120" pageNumber="121" pagination="50 - 74" refId="ref117611" refString="Miyatake, M. (1965) Some Coccinellidae (excluding Scymnini) of Formosa (Coleoptera). Special Bulletin of the Lepidopterists' Society of Japan, 1, 50 - 74." type="journal article" year="1965">Miyatake 1965: 60</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
<paragraph id="8BD1F0741420464FBDDF3396FD7DF9BA" blockId="120.[151,1437,1360,1845]" pageId="120" pageNumber="121">
<treatmentCitationGroup id="AB7ED75A1420464FBDDF3396FD7DF9BA" pageId="120" pageNumber="121">
<treatmentCitation id="0ACFD6651420464FBDDF3396FE73F999" author="Mulsant, E." box="[151,446,1514,1538]" page="159" pageId="120" pageNumber="121" year="1853">
<taxonomicName id="4C6E8BF71420464FBDDF3396FE73F999" ID-CoL="37NS" authority="Mulsant, 1853 a: 159" authorityName="Mulsant" authorityPageNumber="159" authorityYear="1853" box="[151,446,1514,1538]" class="Florideophyceae" family="Balliaceae" genus="Ballia" kingdom="Plantae" order="Balliales" pageId="120" pageNumber="121" phylum="Rhodophyta" rank="genus">
<emphasis id="B91A2C661420464FBDDF3396FF18F999" box="[151,213,1514,1538]" italics="true" pageId="120" pageNumber="121">Ballia</emphasis>
<bibRefCitation id="EFFF8D851420464FBDA83396FE73F999" author="Mulsant, E." box="[224,446,1514,1538]" pageId="120" pageNumber="121" pagination="129 - 333" refId="ref117902" refString="Mulsant, E. (1853 a) Supplement a la monographie des coleopteres trimeres securipalpes. Annales de la Societe Linneenne de Lyon, New Series, 1, 129 - 333." type="journal article" year="1853">Mulsant, 1853a: 159</bibRefCitation>
</taxonomicName>
</treatmentCitation>
;
<typeStatus id="54D54ED61420464FBC843396FE33F999" box="[460,510,1514,1538]" pageId="120" pageNumber="121">Type</typeStatus>
species:
<taxonomicName id="4C6E8BF71420464FBF2D3396FC41F999" authority="Mulsant" authorityName="Mulsant" box="[613,908,1514,1538]" class="Florideophyceae" family="Balliaceae" genus="Ballia" kingdom="Plantae" order="Balliales" pageId="120" pageNumber="121" phylum="Rhodophyta" rank="species" species="christophori">
<emphasis id="B91A2C661420464FBF2D3396FCE6F999" box="[613,811,1514,1538]" italics="true" pageId="120" pageNumber="121">Ballia christophori</emphasis>
Mulsant
</taxonomicName>
, by subsequent designation of Crotch 1874: 126. Synonymized by
<treatmentCitation id="0ACFD6651420464FBC313074FD61F9BA" author="Iablokoff-Khnzorian, S. M." box="[377,684,1544,1569]" page="71" pageId="120" pageNumber="121" year="1979">
<bibRefCitation id="EFFF8D851420464FBC313074FD61F9BA" author="Iablokoff-Khnzorian, S. M." box="[377,684,1544,1569]" pageId="120" pageNumber="121" pagination="37 - 75" refId="ref114003" refString="Iablokoff-Khnzorian, S. M. (1979) Genera der Palaarktischen Coccinellini (Coleoptera: Coccinellidae). Entomologische Blatter fur Biologie und Systematik der Kafer, 75 (1 - 2), 37 - 75." type="journal article" year="1979">Iablokoff-Khnzorian 1979: 71</bibRefCitation>
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.
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<paragraph id="8BD1F0741420464FBDDF305BFBEDF9C5" blockId="120.[151,1437,1360,1845]" pageId="120" pageNumber="121">
<treatmentCitationGroup id="AB7ED75A1420464FBDDF305BFBEDF9C5" pageId="120" pageNumber="121">
<taxonomicName id="4C6E8BF71420464FBDDF305BFE52F9A4" authority="Crotch, 1871: 6" authorityName="Crotch" authorityPageNumber="6" authorityYear="1871" box="[151,415,1575,1599]" class="Insecta" family="Coccinellidae" genus="Callineda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="120" pageNumber="121" phylum="Arthropoda" rank="genus">
<emphasis id="B91A2C661420464FBDDF305BFF36F9A4" box="[151,251,1575,1599]" italics="true" pageId="120" pageNumber="121">Callineda</emphasis>
Crotch, 1871: 6
</taxonomicName>
.—
<treatmentCitation id="0ACFD6651420464FBCF4305BFD54F9A4" author="Timberlake, P. H." box="[444,665,1575,1599]" page="17" pageId="120" pageNumber="121" year="1943">
<bibRefCitation id="EFFF8D851420464FBCF4305BFD54F9A4" author="Timberlake, P. H." box="[444,665,1575,1599]" pageId="120" pageNumber="121" pagination="1 - 67" refId="ref123429" refString="Timberlake, P. H. (1943) The Coccinellidae or lady beetles of the Koebele collection - Part I. Hawaiian Planters' Record, 47, 1 - 67." type="journal article" year="1943">Timberlake, 1943: 17</bibRefCitation>
</treatmentCitation>
(not Crotch, 1874: 122).
<typeStatus id="54D54ED61420464FBED43054FC03F9DB" box="[924,974,1576,1600]" pageId="120" pageNumber="121">Type</typeStatus>
species:
<taxonomicName id="4C6E8BF71420464FB963305BFA54F9A4" authority="Fabricius" authorityName="Fabricius" box="[1067,1433,1575,1599]" class="Insecta" family="Coccinellidae" genus="Coccinella" kingdom="Animalia" order="Coleoptera" pageId="120" pageNumber="121" phylum="Arthropoda" rank="species" species="sedecimnotata">
<emphasis id="B91A2C661420464FB963305BFAFCF9A4" box="[1067,1329,1575,1599]" italics="true" pageId="120" pageNumber="121">Coccinella sedecimnotata</emphasis>
Fabricius
</taxonomicName>
, by subsequent designation of
<treatmentCitation id="0ACFD6651420464FBCBC303AFD46F9C5" author="Rye, E. C." box="[500,651,1606,1630]" page="329" pageId="120" pageNumber="121" year="1873">
<bibRefCitation id="EFFF8D851420464FBCBC303AFD46F9C5" author="Rye, E. C." box="[500,651,1606,1630]" pageId="120" pageNumber="121" pagination="1871" refId="ref120977" refString="Rye, E. C. (1873) Insecta. In: Zoological Record 1871. Vol. 8. John van Voorst, London. [unknown pagination]" type="book chapter" year="1873">Rye 1873: 329</bibRefCitation>
</treatmentCitation>
. Synonymized by
<treatmentCitation id="0ACFD6651420464FBE0E303AFBD1F9C5" author="Timberlake, P. H." box="[838,1052,1606,1630]" page="17" pageId="120" pageNumber="121" year="1943">
<bibRefCitation id="EFFF8D851420464FBE0E303AFBD1F9C5" author="Timberlake, P. H." box="[838,1052,1606,1630]" pageId="120" pageNumber="121" pagination="1 - 67" refId="ref123429" refString="Timberlake, P. H. (1943) The Coccinellidae or lady beetles of the Koebele collection - Part I. Hawaiian Planters' Record, 47, 1 - 67." type="journal article" year="1943">Timberlake 1943: 17</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BD1F0741420464FBDDF3018FCFFF900" blockId="120.[151,1437,1360,1845]" pageId="120" pageNumber="121">
<treatmentCitationGroup id="AB7ED75A1420464FBDDF3018FCFFF900" pageId="120" pageNumber="121">
<taxonomicName id="4C6E8BF71420464FBDDF3018FE7CF9E7" authority="Crotch, 1874: 118" authorityName="Crotch" authorityPageNumber="118" authorityYear="1874" box="[151,433,1636,1661]" class="Insecta" family="Coccinellidae" genus="Stictoleis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="120" pageNumber="121" phylum="Arthropoda" rank="genus">
<emphasis id="B91A2C661420464FBDDF3018FF39F9E7" box="[151,244,1636,1660]" italics="true" pageId="120" pageNumber="121">Stictoleis</emphasis>
Crotch, 1874: 118
</taxonomicName>
.—
<treatmentCitation id="0ACFD6651420464FBC873018FD05F9E7" author="Korschefsky, R." box="[463,712,1636,1661]" page="275" pageId="120" pageNumber="121" year="1932">
<bibRefCitation id="EFFF8D851420464FBC873018FD05F9E7" author="Korschefsky, R." box="[463,712,1636,1661]" pageId="120" pageNumber="121" refId="ref116133" refString="Korschefsky, R. (1932) Coleopterorum Catalogus. Pars 120. Coccinellidae II. W. Junk, Berlin, 435 pp." type="book" year="1932">Korschefsky, 1932: 275</bibRefCitation>
</treatmentCitation>
.
<typeStatus id="54D54ED61420464FBF9A3019FCC9F9E6" box="[722,772,1637,1661]" pageId="120" pageNumber="121">Type</typeStatus>
species:
<taxonomicName id="4C6E8BF71420464FBE2A3018FAEFF9E7" authority="Guerin-Meneville 1835" authorityName="Guerin-Meneville" authorityYear="1835" box="[866,1314,1636,1660]" class="Insecta" family="Coccinellidae" genus="Coccinella" kingdom="Animalia" order="Coleoptera" pageId="120" pageNumber="121" phylum="Arthropoda" rank="species" species="corypha">
<emphasis id="B91A2C661420464FBE2A3018FBE7F9E7" box="[866,1066,1636,1660]" italics="true" pageId="120" pageNumber="121">Coccinella corypha</emphasis>
<bibRefCitation id="EFFF8D851420464FB9793018FAEFF9E7" author="Guerin-Meneville, F. E." box="[1073,1314,1636,1660]" pageId="120" pageNumber="121" pagination="1829 - 1837" refId="ref113312" refString="Guerin-Meneville, F. E. (1835) Iconographie du Regne Animale de Cuvier. Vol. 2. Insectes. J. B. Balliere, Libraire de l'Academie Royale de Medicine, Paris, 104 pls. [pls. 1829 - 1837, text 1844]" type="journal article" year="1835">Guérin-Méneville 1835</bibRefCitation>
</taxonomicName>
, by original designation. Synonymized by
<treatmentCitation id="0ACFD6651420464FBCB230FFFCE0F900" author="Iablokoff-Khnzorian, S. M." box="[506,813,1667,1691]" page="71" pageId="120" pageNumber="121" year="1979">
<bibRefCitation id="EFFF8D851420464FBCB230FFFCE0F900" author="Iablokoff-Khnzorian, S. M." box="[506,813,1667,1691]" pageId="120" pageNumber="121" pagination="37 - 75" refId="ref114003" refString="Iablokoff-Khnzorian, S. M. (1979) Genera der Palaarktischen Coccinellini (Coleoptera: Coccinellidae). Entomologische Blatter fur Biologie und Systematik der Kafer, 75 (1 - 2), 37 - 75." type="journal article" year="1979">Iablokoff-Khnzorian 1979: 71</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BD1F0741420464FBDDF30DEFE76F942" blockId="120.[151,1437,1360,1845]" pageId="120" pageNumber="121">
<treatmentCitationGroup id="AB7ED75A1420464FBDDF30DEFE76F942" pageId="120" pageNumber="121">
<taxonomicName id="4C6E8BF71420464FBDDF30DEFE10F921" authority="Crotch, 1874: 122" authorityName="Crotch" authorityPageNumber="122" authorityYear="1874" box="[151,477,1698,1722]" class="Insecta" family="Coccinellidae" genus="Ptychanatis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="120" pageNumber="121" phylum="Arthropoda" rank="genus">
<emphasis id="B91A2C661420464FBDDF30DEFEC2F921" box="[151,271,1698,1722]" italics="true" pageId="120" pageNumber="121">Ptychanatis</emphasis>
Crotch, 1874: 122
</taxonomicName>
.
<typeStatus id="54D54ED61420464FBCA330DEFDD0F921" box="[491,541,1698,1722]" pageId="120" pageNumber="121">Type</typeStatus>
species:
<taxonomicName id="4C6E8BF71420464FBFCB30DEFC17F921" authority="Pallas, 1773" authorityName="Pallas" authorityYear="1773" box="[643,986,1698,1722]" class="Insecta" family="Coccinellidae" genus="Coccinella" kingdom="Animalia" order="Coleoptera" pageId="120" pageNumber="121" phylum="Arthropoda" rank="species" species="axyridis">
<emphasis id="B91A2C661420464FBFCB30DEFC80F921" box="[643,845,1698,1722]" italics="true" pageId="120" pageNumber="121">Coccinella axyridis</emphasis>
<bibRefCitation id="EFFF8D851420464FBE1030DEFC17F921" author="Pallas, P. S." box="[856,986,1698,1722]" pageId="120" pageNumber="121" refId="ref118401" refString="Pallas, P. S. (1773) Reise durch verschiedene Provinzen des Russischen Reichs. Zweyter Theil. zweytes Buch vom Jahr 1771. Kayserliche Academie der Wissenschaften, St. Petersburg. [unknown pagination]" type="book" year="1773">Pallas, 1773</bibRefCitation>
</taxonomicName>
, by original designation. Synonymized by
<treatmentCitation id="0ACFD6651420464FBD8F30BCFE7AF942" author="Korschefsky, R." box="[199,439,1728,1753]" page="440" pageId="120" pageNumber="121" year="1932">
<bibRefCitation id="EFFF8D851420464FBD8F30BCFE7AF942" author="Korschefsky, R." box="[199,439,1728,1753]" pageId="120" pageNumber="121" refId="ref116133" refString="Korschefsky, R. (1932) Coleopterorum Catalogus. Pars 120. Coccinellidae II. W. Junk, Berlin, 435 pp." type="book" year="1932">Korschefsky 1932: 440</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BD1F0741420464FBDDF30A3FC9FF96C" blockId="120.[151,1437,1360,1845]" box="[151,850,1759,1783]" pageId="120" pageNumber="121">
<treatmentCitationGroup id="AB7ED75A1420464FBDDF30A3FC9FF96C" box="[151,850,1759,1783]" pageId="120" pageNumber="121">
<taxonomicName id="4C6E8BF71420464FBDDF30A3FE4AF96C" box="[151,391,1759,1783]" class="Insecta" family="Coccinellidae" genus="Coccinella" kingdom="Animalia" order="Coleoptera" pageId="120" pageNumber="121" phylum="Arthropoda" rank="subGenus" subGenus="Harmonia">
<emphasis id="B91A2C661420464FBDDF30A3FEC8F96C" box="[151,261,1759,1783]" italics="true" pageId="120" pageNumber="121">Coccinella</emphasis>
(
<emphasis id="B91A2C661420464FBC5C309CFEB1F96C" box="[276,380,1760,1783]" italics="true" pageId="120" pageNumber="121">Harmonia</emphasis>
)
</taxonomicName>
: Chapuis 1876: 179;
<treatmentCitation id="0ACFD6651420464FBF1630A3FC83F96C" author="Korschefsky, R." box="[606,846,1759,1783]" page="439" pageId="120" pageNumber="121" year="1932">
<bibRefCitation id="EFFF8D851420464FBF1630A3FC83F96C" author="Korschefsky, R." box="[606,846,1759,1783]" pageId="120" pageNumber="121" refId="ref116133" refString="Korschefsky, R. (1932) Coleopterorum Catalogus. Pars 120. Coccinellidae II. W. Junk, Berlin, 435 pp." type="book" year="1932">Korschefsky 1932: 439</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BD1F0741420464FBDDF3082FBA2F8AF" blockId="120.[151,1437,1360,1845]" pageId="120" pageNumber="121">
<treatmentCitationGroup id="AB7ED75A1420464FBDDF3082FBA2F8AF" pageId="120" pageNumber="121">
<treatmentCitation id="0ACFD6651420464FBDDF3082FDCCF88D" author="Timberlake, P. H." box="[151,513,1790,1814]" page="17" pageId="120" pageNumber="121" year="1943">
<taxonomicName id="4C6E8BF71420464FBDDF3082FDCCF88D" authority="Timberlake, 1943: 17" authorityName="Timberlake" authorityPageNumber="17" authorityYear="1943" box="[151,513,1790,1814]" class="Insecta" family="Coccinellidae" genus="Rhopaloneda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="120" pageNumber="121" phylum="Arthropoda" rank="genus">
<emphasis id="B91A2C661420464FBDDF3082FED3F88D" box="[151,286,1790,1814]" italics="true" pageId="120" pageNumber="121">Rhopaloneda</emphasis>
<bibRefCitation id="EFFF8D851420464FBC6D3082FDCCF88D" author="Timberlake, P. H." box="[293,513,1790,1814]" pageId="120" pageNumber="121" pagination="1 - 67" refId="ref123429" refString="Timberlake, P. H. (1943) The Coccinellidae or lady beetles of the Koebele collection - Part I. Hawaiian Planters' Record, 47, 1 - 67." type="journal article" year="1943">Timberlake, 1943: 17</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(replacement name for
<taxonomicName id="4C6E8BF71420464FBFB03082FBE6F88D" authority="sensu Crotch, 1874" authorityName="Crotch" authorityYear="1874" box="[760,1067,1790,1814]" class="Insecta" family="Coccinellidae" genus="Callineda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="120" pageNumber="121" phylum="Arthropoda" rank="genus">
<emphasis id="B91A2C661420464FBFB03082FC91F88D" box="[760,860,1790,1814]" italics="true" pageId="120" pageNumber="121">Callineda</emphasis>
sensu Crotch, 1874
</taxonomicName>
, not 1871).
<typeStatus id="54D54ED61420464FB9EF3082FB14F88D" box="[1191,1241,1790,1814]" pageId="120" pageNumber="121">Type</typeStatus>
species:
<taxonomicName id="4C6E8BF71420464FB8703082FEB4F8AF" authority="Crotch" authorityName="Crotch" class="Insecta" family="Coccinellidae" genus="Callineda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="120" pageNumber="121" phylum="Arthropoda" rank="species" species="decussata">
<emphasis id="B91A2C661420464FB8703082FEE6F8AF" italics="true" pageId="120" pageNumber="121">Callineda decussata</emphasis>
Crotch
</taxonomicName>
, by original designation. Synonymized by
<treatmentCitation id="0ACFD6651420464FBE633160FBA6F8AF" author="Iablokoff-Khnzorian, S. M." box="[811,1131,1820,1845]" page="458" pageId="120" pageNumber="121" year="1982">
<bibRefCitation id="EFFF8D851420464FBE633160FBA6F8AF" author="Iablokoff-Khnzorian, S. M." box="[811,1131,1820,1845]" pageId="120" pageNumber="121" refId="ref114042" refString="Iablokoff-Khnzorian, S. M. (1982) Les Coccinelles Coleopteres-Coccinellidae. Tribu Coccinellini des regions Palearctique et Orientale. Boubee, Paris, 568 pp. [in French]" type="book" year="1982">Iablokoff-Khnzorian 1982: 458</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<caption id="DF11A0FC1421464EBDDF30FDFEB9F927" ID-DOI="http://doi.org/10.5281/zenodo.8261775" ID-Zenodo-Dep="8261775" httpUri="https://zenodo.org/record/8261775/files/figure.png" pageId="121" pageNumber="122" startId="121.[151,250,1665,1689]" targetBox="[152,1436,181,1630]" targetPageId="121" targetType="figure">
<paragraph id="8BD1F0741421464EBDDF30FDFEB9F927" blockId="121.[151,1436,1665,1725]" pageId="121" pageNumber="122">
<emphasis id="B91A2C661421464EBDDF30FDFED2F902" bold="true" box="[151,287,1665,1689]" pageId="121" pageNumber="122">FIGURE 83.</emphasis>
<taxonomicName id="4C6E8BF71421464EBC6C30FDFDA3F902" authority="Semenow" authorityName="Semenow" box="[292,622,1665,1689]" class="Insecta" family="Coccinellidae" genus="Halyzia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="121" pageNumber="122" phylum="Arthropoda" rank="species" species="tschitscherini">
<emphasis id="B91A2C661421464EBC6C30FDFDCCF902" box="[292,513,1665,1689]" italics="true" pageId="121" pageNumber="122">Halyzia tschitscherini</emphasis>
Semenow
</taxonomicName>
: a–f. live adult; g. adult, dorsal view; h–j. male genitalia: h. tegmen, ventral view; i. penis; j. penis apex.
</paragraph>
</caption>
<caption id="DF11A0FC1422464DBDDF31EBFBF4F84F" ID-DOI="http://doi.org/10.5281/zenodo.8261779" ID-Zenodo-Dep="8261779" httpUri="https://zenodo.org/record/8261779/files/figure.png" pageId="122" pageNumber="123" startId="122.[151,250,1943,1967]" targetBox="[170,1407,192,1920]" targetPageId="122" targetType="figure">
<paragraph id="8BD1F0741422464DBDDF31EBFBF4F84F" blockId="122.[151,1437,1943,2004]" pageId="122" pageNumber="123">
<emphasis id="B91A2C661422464DBDDF31EBFEEEF834" bold="true" box="[151,291,1943,1967]" pageId="122" pageNumber="123">FIGURE 84.</emphasis>
<taxonomicName id="4C6E8BF71422464DBC6531E4FD4DF82B" authority="Semenow" authorityName="Semenow" box="[301,640,1944,1968]" class="Insecta" family="Coccinellidae" genus="Halyzia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="122" pageNumber="123" phylum="Arthropoda" rank="species" species="tschitscherini">
<emphasis id="B91A2C661422464DBC6531E4FDC2F82B" box="[301,527,1944,1968]" italics="true" pageId="122" pageNumber="123">Halyzia tschitscherini</emphasis>
Semenow
</taxonomicName>
: a. head; b. abdomen, male; c–f. male genitalia: c. tegmen, lateral view; d. tegmen, ventral view; e. penis; f. penis apex; g. female genitalia; h. spermatheca (enlarged).
</paragraph>
</caption>
<subSubSection id="C374A3FF1423464CBDDF36EAFEB4FE6E" pageId="123" pageNumber="124" type="diagnosis">
<paragraph id="8BD1F0741423464CBDDF36EAFEB4FE6E" blockId="123.[151,1437,150,753]" pageId="123" pageNumber="124">
<emphasis id="B91A2C661423464CBDDF36EAFEDFFF2B" bold="true" box="[151,274,150,176]" pageId="123" pageNumber="124">Diagnosis.</emphasis>
Size medium to large, coloration yellow, orange or reddish with various patterns. Form elongate oval (erstwhile subgenus
<taxonomicName id="4C6E8BF71423464CBCC836C7FD54FF4E" authority="Crotch 1874" authorityName="Crotch" authorityYear="1874" box="[384,665,187,213]" class="Insecta" family="Coccinellidae" genus="Ptychanatis" kingdom="Animalia" order="Coleoptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="subGenus" subGenus="Ptychanatis">
<emphasis id="B91A2C661423464CBCC836C7FDC9FF4E" box="[384,516,187,213]" italics="true" pageId="123" pageNumber="124">Ptychanatis</emphasis>
Crotch 1874
</taxonomicName>
) or strongly rounded and convex (erstwhile subgenus
<taxonomicName id="4C6E8BF71423464CB84B36C7FF1EFF62" authority="Mulsant, 1850" authorityName="Mulsant" authorityYear="1850" class="Insecta" genus="Leis" kingdom="Animalia" order="Coleoptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="subGenus" subGenus="Leis">
<emphasis id="B91A2C661423464CB84B36C7FAFFFF4F" box="[1283,1330,187,212]" italics="true" pageId="123" pageNumber="124">Leis</emphasis>
<bibRefCitation id="EFFF8D851423464CB87136C7FF1EFF62" author="Mulsant, E." pageId="123" pageNumber="124" pagination="1 - 1104" refId="ref117842" refString="Mulsant, E. (1850) Species des Coleopteres Trimeres Securipalpes. Annales des Sciences Physiques et Naturelles, d'Agriculture et d'Industrie, publiees par la Societe nationale d'Agriculture, etc., de Lyon, Deuxieme Serie, 2, 1 - 1104. https: // doi. org / 10.5962 / bhl. title. 8953" type="journal article" year="1850">Mulsant, 1850</bibRefCitation>
</taxonomicName>
). Head with anteroclypeal margin straight between lateral projections. Inner margins of eyes strongly divergent towards apex, separated by at least twice the eye width. Antenna 11-segmented, terminal antennomere apically obliquely truncate and asymmetrically transverse. Prothoracic hypomeron without foveae. Prosternal intercoxal process with carinae. Mesoventrite with anterior margin very shallowly emarginate. Abdomen with six visible ventrites, abdominal postcoxal line incomplete with an oblique, associate dividing line that is usually curved and apically merged with the main line. Mid and hind tibia without spurs. Ovipositor with coxites shaped like club handles; spermatheca with a narrow / reduced ramus and a well-developed / produced nodulus, infundibulum present and distinct.
</paragraph>
</subSubSection>
<subSubSection id="C374A3FF1423464CBD8F3782FAFCFDA6" pageId="123" pageNumber="124" type="distribution">
<paragraph id="8BD1F0741423464CBD8F3782FAFCFDA6" blockId="123.[151,1437,150,753]" pageId="123" pageNumber="124">
<emphasis id="B91A2C661423464CBD8F3782FEACFD83" bold="true" box="[199,353,510,536]" pageId="123" pageNumber="124">Distribution.</emphasis>
This is a moderately large genus with about 30 species (Coutanceau 2008), distributed “throughout the Old World, particularly abundant in the
<collectingRegion id="49AA3E961423464CBFC8345FFD11FDA6" box="[640,732,547,573]" country="Saudi Arabia" name="Ash Sharqiyah" pageId="123" pageNumber="124">Oriental</collectingRegion>
and Australo-Pacific regions” (
<bibRefCitation id="EFFF8D851423464CB9743458FAE8FDA6" author="Slipinski, A. &amp; Li, J. &amp; Pang, H." box="[1084,1317,547,575]" pageId="123" pageNumber="124" refId="ref122780" refString="Slipinski, A., Li, J. &amp; Pang, H. (2020) Ladybird Beetles of the Australo-Pacific Region Coleoptera: Coccinellidae: Coccinellini. CSIRO Publishing, Clayton, Victoria, 240 pp. https: // doi. org / 10.1071 / 9781486303885" type="book" year="2020">
Ślipiński
<emphasis id="B91A2C661423464CB9E1345FFB17FDA6" box="[1193,1242,547,573]" italics="true" pageId="123" pageNumber="124">et al</emphasis>
. 2020
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C374A3FF1423464CBD8F343AFE42FD32" pageId="123" pageNumber="124" type="discussion">
<paragraph id="8BD1F0741423464CBD8F343AFE42FD32" blockId="123.[151,1437,150,753]" pageId="123" pageNumber="124">
<emphasis id="B91A2C661423464CBD8F343AFEF0FDFB" bold="true" box="[199,317,582,608]" pageId="123" pageNumber="124">Affinities.</emphasis>
<bibRefCitation id="EFFF8D851423464CBC09343BFDA8FDFA" author="Tomaszewska, W. &amp; Escalona, H. E. &amp; Hartley, D. &amp; Li, J. &amp; Wang, X. &amp; Li, H. S. &amp; Pang, H. &amp; Slipinski, A. &amp; Zwick, A." box="[321,613,583,609]" pageId="123" pageNumber="124" pagination="611 - 631" refId="ref123461" refString="Tomaszewska, W., Escalona, H. E., Hartley, D., Li, J., Wang, X., Li, H. S., Pang, H., Slipinski, A. &amp; Zwick, A. (2021) Phylogeny of true ladybird beetles (Coccinellidae: Coccinellini) reveals pervasive convergent evolution and a rapid Cenozoic radiation. Systematic Entomology, 46, 611 - 631. https: // doi. org / 10.1111 / syen. 12479" type="journal article" year="2021">
Tomaszewska
<emphasis id="B91A2C661423464CBCAA343BFDDDFDFA" box="[482,528,583,609]" italics="true" pageId="123" pageNumber="124">et al</emphasis>
. (2021)
</bibRefCitation>
included
<taxonomicName id="4C6E8BF71423464CBF86343BFC97FDFA" box="[718,858,583,609]" class="Insecta" family="Coccinellidae" genus="Hippodamia" kingdom="Animalia" order="Coleoptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="genus">
<emphasis id="B91A2C661423464CBF86343BFC97FDFA" box="[718,858,583,609]" italics="true" pageId="123" pageNumber="124">Hippodamia</emphasis>
</taxonomicName>
,
<taxonomicName id="4C6E8BF71423464CBE2D343BFC15FDFB" authorityName="Mulsant" authorityYear="1846" box="[869,984,583,608]" class="Insecta" family="Coccinellidae" genus="Harmonia" kingdom="Animalia" order="Coleoptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="genus">
<emphasis id="B91A2C661423464CBE2D343BFC15FDFB" box="[869,984,583,608]" italics="true" pageId="123" pageNumber="124">Harmonia</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6E8BF71423464CB941343BFB0EFDFA" authority="Weise" authorityName="Weise" authorityYear="1899" box="[1033,1219,583,609]" class="Insecta" family="Coccinellidae" genus="Aphidecta" kingdom="Animalia" order="Coleoptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="genus">
<emphasis id="B91A2C661423464CB941343BFBB6FDFA" box="[1033,1147,583,609]" italics="true" pageId="123" pageNumber="124">Aphidecta</emphasis>
Weise
</taxonomicName>
in the
<emphasis id="B91A2C661423464CB84F343BFF15FD1E" italics="true" pageId="123" pageNumber="124">Hippodamiagroup</emphasis>
of genera. They recovered
<taxonomicName id="4C6E8BF71423464CBF543417FD65FD1E" box="[540,680,619,645]" class="Insecta" family="Coccinellidae" genus="Hippodamia" kingdom="Animalia" order="Coleoptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="genus">
<emphasis id="B91A2C661423464CBF543417FD65FD1E" box="[540,680,619,645]" italics="true" pageId="123" pageNumber="124">Hippodamia</emphasis>
</taxonomicName>
+
<taxonomicName id="4C6E8BF71423464CBF843417FCF2FD1F" authorityName="Mulsant" authorityYear="1846" box="[716,831,619,644]" class="Insecta" family="Coccinellidae" genus="Harmonia" kingdom="Animalia" order="Coleoptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="genus">
<emphasis id="B91A2C661423464CBF843417FCF2FD1F" box="[716,831,619,644]" italics="true" pageId="123" pageNumber="124">Harmonia</emphasis>
</taxonomicName>
in a subclade with moderate support and
<taxonomicName id="4C6E8BF71423464CB8633417FA50FD1E" authorityName="Weise" authorityYear="1899" box="[1323,1437,619,645]" class="Insecta" family="Coccinellidae" genus="Aphidecta" kingdom="Animalia" order="Coleoptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="genus">
<emphasis id="B91A2C661423464CB8633417FA50FD1E" box="[1323,1437,619,645]" italics="true" pageId="123" pageNumber="124">Aphidecta</emphasis>
</taxonomicName>
formed a sister-group.
</paragraph>
</subSubSection>
<subSubSection id="C374A3FF1423464CBD8F34CFFD70FD6A" pageId="123" pageNumber="124" type="taxon_list">
<paragraph id="8BD1F0741423464CBD8F34CFFD70FD6A" blockId="123.[151,1437,150,753]" pageId="123" pageNumber="124">
<emphasis id="B91A2C661423464CBD8F34CFFE5FFD57" bold="true" box="[199,402,690,717]" pageId="123" pageNumber="124">Included species.</emphasis>
Eight species are present in the Indian region, one of them,
<taxonomicName id="4C6E8BF71423464CB97534CFFA95FD56" authority="POORANI, 2023" authorityName="Poorani" authorityYear="2023" box="[1085,1368,691,717]" class="Insecta" family="Coccinellidae" genus="Harmonia" kingdom="Animalia" order="Coleoptera" pageId="123" pageNumber="124" phylum="Arthropoda" rank="species" species="andamanensis" status="sp. nov.">
<emphasis id="B91A2C661423464CB97534CFFA95FD56" box="[1085,1368,691,717]" italics="true" pageId="123" pageNumber="124">Harmonia andamanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A229911D1423464CB82834CFFA51FD56" box="[1376,1436,691,717]" pageId="123" pageNumber="124" rank="species">sp. n.</taxonomicNameLabel>
(described here), is endemic to South Andamans.
</paragraph>
</subSubSection>
</treatment>
</document>
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