624 lines
99 KiB
XML
624 lines
99 KiB
XML
<document id="BAA17337AFCD779D70DFD450DB3A6CA7" ID-DOI="10.11646/zootaxa.4763.2.6" ID-GBIF-Dataset="be821e5b-bf9e-43f9-9d06-7aa95a809e0b" ID-ISSN="1175-5334" ID-Zenodo-Dep="3758264" ID-ZooBank="urn:lsid:zoobank.org:pub:6649FF3C-25C1-4A26-9362-D5945684708C" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1587398495903" checkinUser="carolina" docAuthor="Chainthong, Damrong, Sartori, Michel & Boonsoong, Boonsatien" docDate="2020" docId="03B5265BFFC73F6AFF5FDBC36D49F82D" docLanguage="en" docName="zootaxa.4763.2.6.pdf.imf" docOrigin="Zootaxa 4763 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Stylogomphus thongphaphumensis Chainthong & Sartori & Boonsoong 2020, sp. nov." docType="treatment" docVersion="6" lastPageNumber="237" masterDocId="FF8C5E23FFC63F6CFFC8DA566860FFCD" masterDocTitle="Stylogomphus thongphaphumensis (Odonata: Anisoptera: Gomphidae), a new gomphid dragonfly and the first record of S. malayanus Sasamoto, 2001 from Thailand" masterLastPageNumber="245" masterPageNumber="231" pageNumber="232" updateTime="1698739069520" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="57D7B2B4749AC8256F3DDACEFA8D06D3">Stylogomphus thongphaphumensis (Odonata: Anisoptera: Gomphidae), a new gomphid dragonfly and the first record of S. malayanus Sasamoto, 2001 from Thailand</mods:title>
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<mods:name id="2F599D7981DC9476469B8C6592F1E736" type="personal">
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<mods:roleTerm id="B0D9BB944203813F72C1E05EF867519C">Author</mods:roleTerm>
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<mods:namePart id="3972F9855656CB38D56C9B6C1CCBBE7F">Chainthong, Damrong</mods:namePart>
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<mods:affiliation id="4E3B025F4F4490B8C778C0C0C8B90982">Animal Systematics and Ecology Speciality Research Unit (ASESRU), Department of Zoology, faculty of Science, Kasetsart University</mods:affiliation>
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<mods:namePart id="00C9E76DBF03A51E809C11352675DED1">Sartori, Michel</mods:namePart>
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<mods:affiliation id="281D0A361EDD6DD52D89F228D5509740">Museum of Zoology, Palais de Rumine</mods:affiliation>
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<mods:namePart id="3A80583D5B75CE22BD1BC2307267DF1B">Boonsoong, Boonsatien</mods:namePart>
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<mods:affiliation id="286B80EFF3AF73303C56EC52AE74CD6F">Animal Systematics and Ecology Speciality Research Unit (ASESRU), Department of Zoology, faculty of Science, Kasetsart University</mods:affiliation>
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<mods:nameIdentifier id="B21C3DEABF986F066E38CED1BFABD211" type="email">fscibtb@ku.ac.th</mods:nameIdentifier>
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<mods:title id="DB8CA8B5EA7FA8BBD4553E54B9634AA5">Zootaxa</mods:title>
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<mods:date id="47AE103BAF6141120D449CDE62B59233">2020</mods:date>
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<mods:number id="6783A80846F4C7F97F344181C0E8C114">4763</mods:number>
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<treatment id="03B5265BFFC73F6AFF5FDBC36D49F82D" ID-DOI="http://doi.org/10.5281/zenodo.3806442" ID-GBIF-Taxon="163652867" ID-Zenodo-Dep="3806442" LSID="urn:lsid:plazi:treatment:03B5265BFFC73F6AFF5FDBC36D49F82D" httpUri="http://treatment.plazi.org/id/03B5265BFFC73F6AFF5FDBC36D49F82D" lastPageId="6" lastPageNumber="237" pageId="1" pageNumber="232">
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<paragraph id="8BA3974DFFC73F6DFF5FDBC36AC5FE62" blockId="1.[151,677,405,467]" box="[151,677,405,432]" pageId="1" pageNumber="232">
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<emphasis id="B9684B5FFFC73F6DFF5FDBC36AC5FE62" bold="true" box="[151,677,405,432]" pageId="1" pageNumber="232">
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<taxonomicName id="4C1CECCEFFC73F6DFF5FDBC36A20FE7D" ID-CoL="9X4D2" authority="Chainthong & Sartori & Boonsoong, 2020" authorityName="Chainthong & Sartori & Boonsoong" authorityYear="2020" box="[151,576,405,432]" class="Insecta" family="Gomphidae" genus="Stylogomphus" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="232" phylum="Arthropoda" rank="species" species="thongphaphumensis" status="sp. nov.">
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<emphasis id="B9684B5FFFC73F6DFF5FDBC36A20FE7D" bold="true" box="[151,576,405,432]" italics="true" pageId="1" pageNumber="232">Stylogomphus thongphaphumensis</emphasis>
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<taxonomicNameLabel id="A25BF624FFC73F6DFD8FDBC36AC5FE62" box="[583,677,405,431]" pageId="1" pageNumber="232" rank="species">sp. nov.</taxonomicNameLabel>
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<subSubSection id="C306C4C6FFC73F6DFF5FDBEF698BFE1E" box="[151,491,441,467]" pageId="1" pageNumber="232" type="description">
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<paragraph id="8BA3974DFFC73F6DFF5FDBEF698BFE1E" blockId="1.[151,677,405,467]" box="[151,491,441,467]" pageId="1" pageNumber="232">
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(
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<figureCitation id="13278BC8FFC73F6DFF57DBEF6971FE1E" box="[159,273,441,467]" captionStart-0="FIGURES 1–2" captionStart-1="FIGURES 3–7" captionStart-2="FIGURES 8–9" captionStart-3="FIGURES 10–12" captionStart-4="FIGURES 13–14" captionStartId-0="2.[151,264,1960,1985]" captionStartId-1="3.[151,264,942,967]" captionStartId-2="4.[151,264,1239,1264]" captionStartId-3="5.[151,264,1909,1934]" captionStartId-4="6.[151,264,1127,1152]" captionTargetBox-0="[183,1404,306,1932]" captionTargetBox-1="[243,1345,181,917]" captionTargetBox-2="[219,1366,189,1205]" captionTargetBox-3="[467,1121,273,1885]" captionTargetBox-4="[512,1076,181,1100]" captionTargetId-0="figure@2.[183,1404,306,1932]" captionTargetId-1="figure@3.[243,1345,181,917]" captionTargetId-2="figure@4.[214,1374,181,1213]" captionTargetId-3="figure@5.[466,1122,273,1885]" captionTargetId-4="figure@6.[512,1076,181,1102]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="5" captionTargetPageId-4="6" captionText-0="FIGURES 1–2. Stylogomphus thongphaphumensis sp. nov., holotype ♂ (1) and paratype ♀ (2)." captionText-1="FIGURES 3–7. Stylogomphus thongphaphumensis sp. nov., holotype male, (3) head in frontal view; (4) pterothoracic pattern, in dorsal view; (5) pterothoracic pattern, in lateral view; (6) accessory genitalia, in lateral view; (7) vesica spermalis, in lateral view. Scale = 1 mm (3, 4, 5), 0.5 mm (6, 7)." captionText-2="FIGURES 8–9. Stylogomphus thongphaphumensis sp. nov., holotype male, (8) right-forewing; (9) right-hindwing. Scale = 5 mm." captionText-3="FIGURES 10–12. Stylogomphus thongphaphumensis sp. nov., anal appendages of holotype male; (10) dorsal view; (11) lateral view; (12) ventral view. Scale =1 mm." captionText-4="FIGURES 13–14. Stylogomphus thongphaphumensis sp. nov., (13) paratype female; anal appendages in dorsal view; (14) female vulvar scales in ventral view. Scale = 0.5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.3758266" figureDoi-1="http://doi.org/10.5281/zenodo.3758268" figureDoi-2="http://doi.org/10.5281/zenodo.3758270" figureDoi-3="http://doi.org/10.5281/zenodo.3758272" figureDoi-4="http://doi.org/10.5281/zenodo.3758274" httpUri-0="https://zenodo.org/record/3758266/files/figure.png" httpUri-1="https://zenodo.org/record/3758268/files/figure.png" httpUri-2="https://zenodo.org/record/3758270/files/figure.png" httpUri-3="https://zenodo.org/record/3758272/files/figure.png" httpUri-4="https://zenodo.org/record/3758274/files/figure.png" pageId="1" pageNumber="232">Figs 1–14</figureCitation>
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,
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<figureCitation id="13278BC8FFC73F6DFED6DBEF6907FE1E" box="[286,359,441,467]" captionStart-0="FIGURES 15–17" captionStart-1="FIGURES 18" captionStart-2="FIGURES 19–24" captionStart-3="FIGURES 25–39" captionStartId-0="7.[151,264,1910,1935]" captionStartId-1="8.[151,264,1945,1970]" captionStartId-2="9.[151,264,1309,1334]" captionStartId-3="10.[151,264,1945,1970]" captionTargetBox-0="[338,1250,181,1853]" captionTargetBox-1="[228,1347,189,1909]" captionTargetBox-2="[199,1390,189,1274]" captionTargetBox-3="[349,1238,190,1917]" captionTargetId-0="figure@7.[338,1250,181,1885]" captionTargetId-1="figure@8.[228,1360,181,1920]" captionTargetId-2="figure@9.[189,1398,181,1283]" captionTargetId-3="figure@10.[340,1248,181,1919]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionTargetPageId-2="9" captionTargetPageId-3="10" captionText-0="FIGURES 15–17. Habitat of Stylogomphus thongphaphumensis sp. nov. larva, (15) sampling site where larvae were collected; (16) microhabitat (sandy substrate) where the larvae were found; (17) habitus of larva." captionText-1="FIGURES 18. Larva of Stylogomphus thongphaphumensis sp. nov.; dorsal view. Scale = 1 mm." captionText-2="FIGURES 19–24. Larva of Stylogomphus thongphaphumensis sp. nov., (19) antenna in dorsal view; (20) right mandible in ventral views; (21) left mandible in ventral views; (22) prementum in ventral view; (23) right maxilla in ventral view; (24) anal appendages in dorsal view. Scale = 1 mm (19), 0.2 mm (20, 21, 23), 0.5 mm (22, 24)." captionText-3="FIGURES 25–39. Comparison of male anal appendages of Stylogomphus spp.: (25–27) S. thongphaphumensis sp. nov.; (28– 30) S. annamensis (redrawn from Kompier 2017); (31–33) S. delicatus (redrawn from Kompier 2017); (34–36) S. lawrenceae (redrawn from Sasamoto 2004); (37–39) S. malayanus (redrawn from Sasamoto 2004)." figureDoi-0="http://doi.org/10.5281/zenodo.3758276" figureDoi-1="http://doi.org/10.5281/zenodo.3758278" figureDoi-2="http://doi.org/10.5281/zenodo.3758280" figureDoi-3="http://doi.org/10.5281/zenodo.3758282" httpUri-0="https://zenodo.org/record/3758276/files/figure.png" httpUri-1="https://zenodo.org/record/3758278/files/figure.png" httpUri-2="https://zenodo.org/record/3758280/files/figure.png" httpUri-3="https://zenodo.org/record/3758282/files/figure.png" pageId="1" pageNumber="232">17–27</figureCitation>
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,
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<figureCitation id="13278BC8FFC73F6DFEBADBEF6983FE1E" box="[370,483,441,467]" captionStart="FIGURES 43–48" captionStartId="12.[151,264,1478,1503]" captionTargetBox="[214,1373,181,1452]" captionTargetId="figure@12.[214,1373,181,1452]" captionTargetPageId="12" captionText="FIGURES 43–48. Comparison of larvae of two Stylogomphus spp.: S. malayanus (43, 45, 47), S. thongphaphumensis sp. nov. (44, 46, 48): (43, 44) antenna in dorsal view; (45, 46) abdomen in dorsal views; (47, 48) prementum in ventral view. Scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3758286" httpUri="https://zenodo.org/record/3758286/files/figure.png" pageId="1" pageNumber="232">44, 46, 48</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C306C4C6FFC73F6DFF5FD8566D21FD66" pageId="1" pageNumber="232" type="materials_examined">
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<paragraph id="8BA3974DFFC73F6DFF5FD8566968FDAE" blockId="1.[151,1437,512,2015]" pageId="1" pageNumber="232">
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<emphasis id="B9684B5FFFC73F6DFF5FD85669E1FDD6" bold="true" box="[151,385,512,539]" pageId="1" pageNumber="232">Material examined.</emphasis>
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<materialsCitation id="3B749D10FFC73F6DFE40D8566968FDAE" ID-GBIF-Occurrence="2599923285" collectingDate="2014-12-14" collectingDateMax="2015-04-30" collectingDateMin="2014-12-14" country="Thailand" elevation="206" latitude="14.605556" location="Thong Pha Phum district" longLatPrecision="21" longitude="98.577225" municipality="Huai Khayeng" pageId="1" pageNumber="232" specimenCount="1" specimenCount-male="1" stateProvince="Kanchanaburi" typeStatus="holotype">
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<emphasis id="B9684B5FFFC73F6DFE40D8566AC6FDD6" bold="true" box="[392,678,512,539]" pageId="1" pageNumber="232">
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<collectingCountry id="F30BD7DDFFC73F6DFE40D8566A43FDD6" box="[392,547,512,539]" name="Thailand" pageId="1" pageNumber="232">THAILAND</collectingCountry>
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:
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<typeStatus id="54A729EFFFC73F6DFDF8D8576AC1FDD6" box="[560,673,513,539]" pageId="1" pageNumber="232" type="holotype">Holotype</typeStatus>
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:
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</emphasis>
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<specimenCount id="9D1A5CC4FFC73F6DFD66D8576AB3FDD6" box="[686,723,513,539]" pageId="1" pageNumber="232" type="male">1♂</specimenCount>
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,
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<collectingMunicipality id="6BC70D37FFC73F6DFD28D8576BE6FDD6" box="[736,902,513,539]" pageId="1" pageNumber="232">Huai Khayeng</collectingMunicipality>
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,
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<location id="8EC3C196FFC73F6DFC59D8576CCEFDD6" LSID="urn:lsid:plazi:treatment:03B5265BFFC73F6AFF5FDBC36D49F82D:8EC3C196FFC73F6DFC59D8576CCEFDD6" box="[913,1198,513,539]" country="Thailand" latitude="14.605556" longLatPrecision="21" longitude="98.577225" municipality="Huai Khayeng" name="Thong Pha Phum district" pageId="1" pageNumber="232" stateProvince="Kanchanaburi">Thong Pha Phum district</location>
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,
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<collectingRegion id="49D859AFFFC73F6DFB71D8576D36FDD6" box="[1209,1366,513,539]" country="Thailand" name="Kanchanaburi" pageId="1" pageNumber="232">Kanchanaburi</collectingRegion>
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Prov- ince,
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<geoCoordinate id="EE28F18AFFC73F6DFF1BD8726939FDF2" box="[211,345,548,575]" degrees="14" direction="north" minutes="36" orientation="latitude" pageId="1" pageNumber="232" precision="15" seconds="20" value="14.605556">14°36′20″N</geoCoordinate>
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<geoCoordinate id="EE28F18AFFC73F6DFE97D8726983FDF2" box="[351,483,548,575]" degrees="98" direction="east" minutes="34" orientation="longitude" pageId="1" pageNumber="232" precision="15" seconds="38" value="98.577225">98°34′38″E</geoCoordinate>
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,
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<elevation id="0031707EFFC73F6DFE26D8736A10FDF2" box="[494,624,549,575]" metricMagnitude="2" metricUnit="m" metricValue="2.06" pageId="1" pageNumber="232" unit="m" value="206.0">
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<quantity id="4CE43AA8FFC73F6DFE26D8736A54FDF2" box="[494,564,549,575]" metricMagnitude="2" metricUnit="m" metricValue="2.06" pageId="1" pageNumber="232" unit="m" value="206.0">206 m</quantity>
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a.s.l.
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</elevation>
|
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, larva collected on
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<date id="FFA2B18DFFC73F6DFC80D8736BB3FDF2" box="[840,979,549,575]" pageId="1" pageNumber="232" value="2014-12-14">
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<collectingDate id="EFE64865FFC73F6DFC80D8736BB3FDF2" box="[840,979,549,575]" pageId="1" pageNumber="232" value="2014-12-14">14.XII.2014</collectingDate>
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</date>
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, adult emerged on
|
||
<date id="FFA2B18DFFC73F6DFB6DD8726D43FDF2" box="[1189,1315,548,575]" pageId="1" pageNumber="232" value="2015-04-30">
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||
<collectingDate id="EFE64865FFC73F6DFB6DD8726D43FDF2" box="[1189,1315,548,575]" pageId="1" pageNumber="232" value="2015-04-30">30.IV.2015</collectingDate>
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</date>
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||
, D. Chain- thong leg.
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</materialsCitation>
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</paragraph>
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<paragraph id="8BA3974DFFC73F6DFF0FD83B6D21FD66" blockId="1.[151,1437,512,2015]" pageId="1" pageNumber="232">
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<emphasis id="B9684B5FFFC73F6DFF0FD83B6928FD4A" bold="true" box="[199,328,621,647]" pageId="1" pageNumber="232">
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<typeStatus id="54A729EFFFC73F6DFF0FD83B6923FD4A" box="[199,323,621,647]" pageId="1" pageNumber="232" type="paratype">Paratypes</typeStatus>
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||
:
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</emphasis>
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Nine final stadia larvae, same locality and collection date as
|
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<typeStatus id="54A729EFFFC73F6DFBC1D83B6C0FFD4A" box="[1033,1135,621,647]" pageId="1" pageNumber="232" type="holotype">holotype</typeStatus>
|
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; adults emerged on:
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||
<specimenCount id="9D1A5CC4FFC73F6DFA94D83B6DFCFD4A" box="[1372,1436,621,647]" pageId="1" pageNumber="232" type="male">4 ♂♂</specimenCount>
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<date id="FFA2B18DFFC73F6DFF5FD8C76943FD66" box="[151,291,657,683]" pageId="1" pageNumber="232" value="2014-12-24">24.XII.2014</date>
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,
|
||
<date id="FFA2B18DFFC73F6DFEE6D8C669C1FD66" box="[302,417,656,683]" pageId="1" pageNumber="232" value="2015-03-08">8.III.2015</date>
|
||
,
|
||
<date id="FFA2B18DFFC73F6DFE64D8C66A4DFD66" box="[428,557,656,683]" pageId="1" pageNumber="232" value="2015-03-18">18.III.2015</date>
|
||
,
|
||
<date id="FFA2B18DFFC73F6DFDF0D8C66AD9FD66" box="[568,697,656,683]" pageId="1" pageNumber="232" value="2015-03-29">29.III.2015</date>
|
||
;
|
||
<specimenCount id="9D1A5CC4FFC73F6DFD0DD8C66B64FD66" box="[709,772,656,683]" pageId="1" pageNumber="232" type="female">5 ♀♀</specimenCount>
|
||
<date id="FFA2B18DFFC73F6DFCC3D8C76BF6FD66" box="[779,918,657,683]" pageId="1" pageNumber="232" value="2014-12-20">20.XII.2014</date>
|
||
,
|
||
<date id="FFA2B18DFFC73F6DFC6AD8C76C4DFD66" box="[930,1069,657,683]" pageId="1" pageNumber="232" value="2014-12-24">24.XII.2014</date>
|
||
,
|
||
<date id="FFA2B18DFFC73F6DFBF0D8C66CCFFD66" box="[1080,1199,656,683]" pageId="1" pageNumber="232" value="2015-02-18">18.II.2015</date>
|
||
,
|
||
<date id="FFA2B18DFFC73F6DFB73D8C66D5CFD66" box="[1211,1340,656,683]" pageId="1" pageNumber="232" value="2015-03-26">26.III.2015</date>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C306C4C6FFC73F6DFF0FD8E36AEDFD3E" pageId="1" pageNumber="232" type="etymology">
|
||
<paragraph id="8BA3974DFFC73F6DFF0FD8E36AEDFD3E" blockId="1.[151,1437,512,2015]" pageId="1" pageNumber="232">
|
||
<emphasis id="B9684B5FFFC73F6DFF0FD8E3692EFD02" bold="true" box="[199,334,693,719]" pageId="1" pageNumber="232">Etymology.</emphasis>
|
||
The specific epithet “thongphaphumensis” refers to the district of
|
||
<collectingRegion id="49D859AFFFC73F6DFB94D8E36D08FD02" box="[1116,1384,693,719]" country="Thailand" name="Kanchanaburi" pageId="1" pageNumber="232">Kanchanaburi Province</collectingRegion>
|
||
that contains the
|
||
<typeStatus id="54A729EFFFC73F6DFEEDD88F6935FD3E" box="[293,341,729,755]" pageId="1" pageNumber="232">type</typeStatus>
|
||
locality of the new species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C306C4C6FFC73F6EFF0FD8AA6B0CFED2" lastPageId="2" lastPageNumber="233" pageId="1" pageNumber="232" type="description">
|
||
<paragraph id="8BA3974DFFC73F6DFF0FD8AA6D63FCDA" blockId="1.[151,1437,512,2015]" box="[199,1283,764,791]" pageId="1" pageNumber="232">
|
||
<emphasis id="B9684B5FFFC73F6DFF0FD8AA6A71FCDA" bold="true" box="[199,529,764,791]" pageId="1" pageNumber="232">
|
||
Description of the
|
||
<typeStatus id="54A729EFFFC73F6DFE6BD8AB6A6CFCDA" box="[419,524,765,791]" pageId="1" pageNumber="232" type="holotype">holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
General appearance and color pattern as shown in
|
||
<figureCitation id="13278BC8FFC73F6DFB84D8AB6CD5FCDA" box="[1100,1205,765,791]" captionStart="FIGURES 1–2" captionStartId="2.[151,264,1960,1985]" captionTargetBox="[183,1404,306,1932]" captionTargetId="figure@2.[183,1404,306,1932]" captionTargetPageId="2" captionText="FIGURES 1–2. Stylogomphus thongphaphumensis sp. nov., holotype ♂ (1) and paratype ♀ (2)." figureDoi="http://doi.org/10.5281/zenodo.3758266" httpUri="https://zenodo.org/record/3758266/files/figure.png" pageId="1" pageNumber="232">Figures 1</figureCitation>
|
||
,
|
||
<figureCitation id="13278BC8FFC73F6DFB0BD8AB6C9EFCDA" box="[1219,1278,765,791]" captionStart-0="FIGURES 3–7" captionStart-1="FIGURES 8–9" captionStart-2="FIGURES 10–12" captionStartId-0="3.[151,264,942,967]" captionStartId-1="4.[151,264,1239,1264]" captionStartId-2="5.[151,264,1909,1934]" captionTargetBox-0="[243,1345,181,917]" captionTargetBox-1="[219,1366,189,1205]" captionTargetBox-2="[467,1121,273,1885]" captionTargetId-0="figure@3.[243,1345,181,917]" captionTargetId-1="figure@4.[214,1374,181,1213]" captionTargetId-2="figure@5.[466,1122,273,1885]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionText-0="FIGURES 3–7. Stylogomphus thongphaphumensis sp. nov., holotype male, (3) head in frontal view; (4) pterothoracic pattern, in dorsal view; (5) pterothoracic pattern, in lateral view; (6) accessory genitalia, in lateral view; (7) vesica spermalis, in lateral view. Scale = 1 mm (3, 4, 5), 0.5 mm (6, 7)." captionText-1="FIGURES 8–9. Stylogomphus thongphaphumensis sp. nov., holotype male, (8) right-forewing; (9) right-hindwing. Scale = 5 mm." captionText-2="FIGURES 10–12. Stylogomphus thongphaphumensis sp. nov., anal appendages of holotype male; (10) dorsal view; (11) lateral view; (12) ventral view. Scale =1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.3758268" figureDoi-1="http://doi.org/10.5281/zenodo.3758270" figureDoi-2="http://doi.org/10.5281/zenodo.3758272" httpUri-0="https://zenodo.org/record/3758268/files/figure.png" httpUri-1="https://zenodo.org/record/3758270/files/figure.png" httpUri-2="https://zenodo.org/record/3758272/files/figure.png" pageId="1" pageNumber="232">3–12</figureCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BA3974DFFC73F6DFF0FD9776938FC6A" blockId="1.[151,1437,512,2015]" pageId="1" pageNumber="232">
|
||
<emphasis id="B9684B5FFFC73F6DFF0FD9776964FCF6" box="[199,260,801,827]" italics="true" pageId="1" pageNumber="232">Head</emphasis>
|
||
(
|
||
<figureCitation id="13278BC8FFC73F6DFEDCD977693AFCF6" box="[276,346,801,827]" captionStart="FIGURES 3–7" captionStartId="3.[151,264,942,967]" captionTargetBox="[243,1345,181,917]" captionTargetId="figure@3.[243,1345,181,917]" captionTargetPageId="3" captionText="FIGURES 3–7. Stylogomphus thongphaphumensis sp. nov., holotype male, (3) head in frontal view; (4) pterothoracic pattern, in dorsal view; (5) pterothoracic pattern, in lateral view; (6) accessory genitalia, in lateral view; (7) vesica spermalis, in lateral view. Scale = 1 mm (3, 4, 5), 0.5 mm (6, 7)." figureDoi="http://doi.org/10.5281/zenodo.3758268" httpUri="https://zenodo.org/record/3758268/files/figure.png" pageId="1" pageNumber="232">Fig. 3</figureCitation>
|
||
). Black with yellow markings as follows: median lobe of labium black and lateral lobes yellow; mandibles yellow, labrum with a pair of transverse ellipsoid yellow spots; genae black; anteclypeus yellow; postclypeus black, with single large yellow spots laterally; postfrons with a broad yellow band, antefrons black; occiput and vertex black.
|
||
</paragraph>
|
||
<paragraph id="8BA3974DFFC73F6DFF0FD9E76BFEFB96" blockId="1.[151,1437,512,2015]" pageId="1" pageNumber="232">
|
||
<emphasis id="B9684B5FFFC73F6DFF0FD9E76978FC06" box="[199,280,945,971]" italics="true" pageId="1" pageNumber="232">Thorax</emphasis>
|
||
. Colour pattern as shown in
|
||
<figureCitation id="13278BC8FFC73F6DFD94D9E76A84FC06" box="[604,740,944,971]" captionStart="FIGURES 3–7" captionStartId="3.[151,264,942,967]" captionTargetBox="[243,1345,181,917]" captionTargetId="figure@3.[243,1345,181,917]" captionTargetPageId="3" captionText="FIGURES 3–7. Stylogomphus thongphaphumensis sp. nov., holotype male, (3) head in frontal view; (4) pterothoracic pattern, in dorsal view; (5) pterothoracic pattern, in lateral view; (6) accessory genitalia, in lateral view; (7) vesica spermalis, in lateral view. Scale = 1 mm (3, 4, 5), 0.5 mm (6, 7)." figureDoi="http://doi.org/10.5281/zenodo.3758268" httpUri="https://zenodo.org/record/3758268/files/figure.png" pageId="1" pageNumber="232">Figures 4–5</figureCitation>
|
||
. Prothorax black with yellow laterally. Pterothorax black with yellow markings, as shown in
|
||
<figureCitation id="13278BC8FFC73F6DFE38D9836A31FC22" box="[496,593,980,1007]" captionStart="FIGURES 3–7" captionStartId="3.[151,264,942,967]" captionTargetBox="[243,1345,181,917]" captionTargetId="figure@3.[243,1345,181,917]" captionTargetPageId="3" captionText="FIGURES 3–7. Stylogomphus thongphaphumensis sp. nov., holotype male, (3) head in frontal view; (4) pterothoracic pattern, in dorsal view; (5) pterothoracic pattern, in lateral view; (6) accessory genitalia, in lateral view; (7) vesica spermalis, in lateral view. Scale = 1 mm (3, 4, 5), 0.5 mm (6, 7)." figureDoi="http://doi.org/10.5281/zenodo.3758268" httpUri="https://zenodo.org/record/3758268/files/figure.png" pageId="1" pageNumber="232">Figure 5</figureCitation>
|
||
; mesothoracic collar yellow, except on middle; antehumeral stripe narrow, not connected to mesothoracic collar, almost reaching margin of antealar triangle; mesepimeron yellow with black margins all round; metepisternum black with two yellow spots; metepimeron mostly yellow. Legs black, coxae mostly yellow and inner surface black; femora, tibiae and tarsi black.
|
||
</paragraph>
|
||
<paragraph id="8BA3974DFFC73F6DFF0FDE306A12FB0A" blockId="1.[151,1437,512,2015]" pageId="1" pageNumber="232">
|
||
<emphasis id="B9684B5FFFC73F6DFF0FDE306973FBB2" box="[199,275,1126,1151]" italics="true" pageId="1" pageNumber="232">Wings.</emphasis>
|
||
Hyaline, venation dark brown (
|
||
<figureCitation id="13278BC8FFC73F6DFDBDDE336ABBFBB2" box="[629,731,1125,1151]" captionStart="FIGURES 8–9" captionStartId="4.[151,264,1239,1264]" captionTargetBox="[219,1366,189,1205]" captionTargetId="figure@4.[214,1374,181,1213]" captionTargetPageId="4" captionText="FIGURES 8–9. Stylogomphus thongphaphumensis sp. nov., holotype male, (8) right-forewing; (9) right-hindwing. Scale = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.3758270" httpUri="https://zenodo.org/record/3758270/files/figure.png" pageId="1" pageNumber="232">Figs 8–9</figureCitation>
|
||
); 11 (left) and 10 (right) Ax on Fw, 7 (left) and 8 (right) on Hw, 9 (left) and 8 (right) Px on Fw, 9 (left) and 9 (right) on Hw. Pterostigmata very dark brown, surmounting 3.5 cells in both Fw and Hw; anal triangle 3-celled.
|
||
</paragraph>
|
||
<paragraph id="8BA3974DFFC73F6DFF0FDE876CB0FAFE" blockId="1.[151,1437,512,2015]" pageId="1" pageNumber="232">
|
||
<emphasis id="B9684B5FFFC73F6DFF0FDE876951FB26" box="[199,305,1233,1259]" italics="true" pageId="1" pageNumber="232">Abdomen</emphasis>
|
||
. Abdomen predominantly black, with bright yellow markings, as follows (
|
||
<figureCitation id="13278BC8FFC73F6DFBB3DE876CDEFB26" box="[1147,1214,1233,1259]" captionStart="FIGURES 1–2" captionStartId="2.[151,264,1960,1985]" captionTargetBox="[183,1404,306,1932]" captionTargetId="figure@2.[183,1404,306,1932]" captionTargetPageId="2" captionText="FIGURES 1–2. Stylogomphus thongphaphumensis sp. nov., holotype ♂ (1) and paratype ♀ (2)." figureDoi="http://doi.org/10.5281/zenodo.3758266" httpUri="https://zenodo.org/record/3758266/files/figure.png" pageId="1" pageNumber="232">Fig. 1</figureCitation>
|
||
): S1 mostly yellow laterally, S2 yellow around auricle, another narrow yellow mid-dorsal stripe on S1and S2; S3 with basal mid-dorsal tiny triangular yellow spot, S4–7 with small dorso-lateral yellow markings at base; S8–10 black.
|
||
</paragraph>
|
||
<paragraph id="8BA3974DFFC73F6DFF0FDF6B69DEFA2A" blockId="1.[151,1437,512,2015]" pageId="1" pageNumber="232">
|
||
<emphasis id="B9684B5FFFC73F6DFF0FDF6B69E8FA9A" box="[199,392,1341,1367]" italics="true" pageId="1" pageNumber="232">Anal appendages</emphasis>
|
||
(
|
||
<figureCitation id="13278BC8FFC73F6DFE5EDF6B6A79FA9A" box="[406,537,1341,1367]" captionStart="FIGURES 10–12" captionStartId="5.[151,264,1909,1934]" captionTargetBox="[467,1121,273,1885]" captionTargetId="figure@5.[466,1122,273,1885]" captionTargetPageId="5" captionText="FIGURES 10–12. Stylogomphus thongphaphumensis sp. nov., anal appendages of holotype male; (10) dorsal view; (11) lateral view; (12) ventral view. Scale =1 mm." figureDoi="http://doi.org/10.5281/zenodo.3758272" httpUri="https://zenodo.org/record/3758272/files/figure.png" pageId="1" pageNumber="232">Figs 10–12</figureCitation>
|
||
). Cerci creamy white, approximately 1.5x length of S10, with ventrolateral black tooth about one-fifth from base, short and turned outward; in dorsal view (
|
||
<figureCitation id="13278BC8FFC73F6DFC1EDF376C47FAB6" box="[982,1063,1377,1403]" captionStart="FIGURES 10–12" captionStartId="5.[151,264,1909,1934]" captionTargetBox="[467,1121,273,1885]" captionTargetId="figure@5.[466,1122,273,1885]" captionTargetPageId="5" captionText="FIGURES 10–12. Stylogomphus thongphaphumensis sp. nov., anal appendages of holotype male; (10) dorsal view; (11) lateral view; (12) ventral view. Scale =1 mm." figureDoi="http://doi.org/10.5281/zenodo.3758272" httpUri="https://zenodo.org/record/3758272/files/figure.png" pageId="1" pageNumber="232">Fig. 10</figureCitation>
|
||
) constricted in the middle, apical half slender. Epiproct black, wider than long, approximately 0.7x length of S10 and 0.4x of cerci, lateral lobes diverging with blunt tips; the median cleft U-shaped and shallow in ventral view, covered with minute setae, widening toward apex (
|
||
<figureCitation id="13278BC8FFC73F6DFEF8DF9B69D1FA2A" box="[304,433,1485,1511]" captionStart="FIGURES 10–12" captionStartId="5.[151,264,1909,1934]" captionTargetBox="[467,1121,273,1885]" captionTargetId="figure@5.[466,1122,273,1885]" captionTargetPageId="5" captionText="FIGURES 10–12. Stylogomphus thongphaphumensis sp. nov., anal appendages of holotype male; (10) dorsal view; (11) lateral view; (12) ventral view. Scale =1 mm." figureDoi="http://doi.org/10.5281/zenodo.3758272" httpUri="https://zenodo.org/record/3758272/files/figure.png" pageId="1" pageNumber="232">Figs 11–12</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA3974DFFC73F6DFF0FDFA76941F9BA" blockId="1.[151,1437,512,2015]" pageId="1" pageNumber="232">
|
||
<emphasis id="B9684B5FFFC73F6DFF0FDFA769CCF9C6" box="[199,428,1521,1547]" italics="true" pageId="1" pageNumber="232">Accessory genitalia.</emphasis>
|
||
Anterior and posterior hamulus dark brown, anterior hamulus in lateral view slender and smoothly curved. Posterior hamulus broad, anterior margin with small denticles, inner-curved spine on upper side. Vesica spermalis (
|
||
<figureCitation id="13278BC8FFC73F6DFEA8DC6F69A5F999" box="[352,453,1593,1620]" captionStart="FIGURES 3–7" captionStartId="3.[151,264,942,967]" captionTargetBox="[243,1345,181,917]" captionTargetId="figure@3.[243,1345,181,917]" captionTargetPageId="3" captionText="FIGURES 3–7. Stylogomphus thongphaphumensis sp. nov., holotype male, (3) head in frontal view; (4) pterothoracic pattern, in dorsal view; (5) pterothoracic pattern, in lateral view; (6) accessory genitalia, in lateral view; (7) vesica spermalis, in lateral view. Scale = 1 mm (3, 4, 5), 0.5 mm (6, 7)." figureDoi="http://doi.org/10.5281/zenodo.3758268" httpUri="https://zenodo.org/record/3758268/files/figure.png" pageId="1" pageNumber="232">Figs 6–7</figureCitation>
|
||
), first segment bifid, prepuce of third segment bulging in lateral view, apex of lobe hammer-shaped.
|
||
</paragraph>
|
||
<paragraph id="8BA3974DFFC73F6DFF0FDCD76CE0F956" blockId="1.[151,1437,512,2015]" box="[199,1152,1664,1691]" pageId="1" pageNumber="232">
|
||
<emphasis id="B9684B5FFFC73F6DFF0FDCD769A8F956" bold="true" box="[199,456,1664,1691]" pageId="1" pageNumber="232">Measurements (mm):</emphasis>
|
||
Fw length 23, Hw length 21, abdomen with appendages 28.5.
|
||
</paragraph>
|
||
<paragraph id="8BA3974DFFC73F6DFF0FDCF26DFDF8E6" blockId="1.[151,1437,512,2015]" pageId="1" pageNumber="232">
|
||
<emphasis id="B9684B5FFFC73F6DFF0FDCF26A53F972" bold="true" box="[199,563,1700,1727]" pageId="1" pageNumber="232">
|
||
Variation in
|
||
<typeStatus id="54A729EFFFC73F6DFEA5DCF369B8F972" box="[365,472,1701,1727]" pageId="1" pageNumber="232" type="paratype">paratype</typeStatus>
|
||
males.
|
||
</emphasis>
|
||
Minor variations apparent among the four
|
||
<typeStatus id="54A729EFFFC73F6DFB98DCF36CD0F972" box="[1104,1200,1701,1727]" pageId="1" pageNumber="232" type="paratype">paratype</typeStatus>
|
||
males, when compared with the
|
||
<typeStatus id="54A729EFFFC73F6DFE90DC9F69DEF92E" box="[344,446,1737,1763]" pageId="1" pageNumber="232" type="holotype">holotype</typeStatus>
|
||
. The
|
||
<typeStatus id="54A729EFFFC73F6DFDC3DC9F6A0BF92E" box="[523,619,1737,1763]" pageId="1" pageNumber="232" type="paratype">paratype</typeStatus>
|
||
specimens differ in the number of Ax and Px: 8–11 (left) or 11 (right) Ax on Fw, 8 (left) or 8 (right) on Hw, 8–10 (left) or 9–10 (right) Px on Fw, 6 (left) or 6–8 (right) on Hw. Measurements (mm) are:
|
||
<specimenCount id="9D1A5CC4FFC73F6DFDC2DD476A7FF8E6" box="[522,543,1809,1835]" pageId="1" pageNumber="232" type="male">♂</specimenCount>
|
||
forewing 23–24, hindwing 21–23; abdomen including appendages 27–29.
|
||
</paragraph>
|
||
<paragraph id="8BA3974DFFC73F6EFF0FDD636B0CFED2" blockId="1.[151,1437,512,2015]" lastBlockId="2.[151,1437,153,287]" lastPageId="2" lastPageNumber="233" pageId="1" pageNumber="232">
|
||
<emphasis id="B9684B5FFFC73F6DFF0FDD6369FBF882" bold="true" box="[199,411,1844,1871]" pageId="1" pageNumber="232">
|
||
<typeStatus id="54A729EFFFC73F6DFF0FDD636954F882" box="[199,308,1845,1871]" pageId="1" pageNumber="232" type="paratype">Paratype</typeStatus>
|
||
females.
|
||
</emphasis>
|
||
Body colouration similar to male (
|
||
<figureCitation id="13278BC8FFC73F6DFCD4DD636B00F882" box="[796,864,1845,1871]" captionStart="FIGURES 1–2" captionStartId="2.[151,264,1960,1985]" captionTargetBox="[183,1404,306,1932]" captionTargetId="figure@2.[183,1404,306,1932]" captionTargetPageId="2" captionText="FIGURES 1–2. Stylogomphus thongphaphumensis sp. nov., holotype ♂ (1) and paratype ♀ (2)." figureDoi="http://doi.org/10.5281/zenodo.3758266" httpUri="https://zenodo.org/record/3758266/files/figure.png" pageId="1" pageNumber="232">Fig. 2</figureCitation>
|
||
). Yellow areas little broader than those of male. Occiput black without a protuberance; anteclypeus yellow; postclypeus black with pair of large triangular yellow spots; postfrons with a band narrowly interrupted in middle. Ax 10–11 (left), 9–11 (right) on Fw, 8–9 (right), 8–10 (left) on Hw; Px 8–9 (right), 7–9 (left) on Fw, 9–10 (right), 7– 8 (left) on Hw; abdomen almost cylindrical. Yellow markings similar to those of male: S1 and S2 mostly yellow laterally, S3 with dumbbell-shaped yellow markings in lateral view. S4–7 with a half ring yellow markings; S8–10 black; cerci pale, about same length as S10, with gradually tapering, pointed tip (
|
||
<figureCitation id="13278BC8FFC43F6EFE4FDAEB69B8FF1A" box="[391,472,189,215]" captionStart="FIGURES 13–14" captionStartId="6.[151,264,1127,1152]" captionTargetBox="[512,1076,181,1100]" captionTargetId="figure@6.[512,1076,181,1102]" captionTargetPageId="6" captionText="FIGURES 13–14. Stylogomphus thongphaphumensis sp. nov., (13) paratype female; anal appendages in dorsal view; (14) female vulvar scales in ventral view. Scale = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3758274" httpUri="https://zenodo.org/record/3758274/files/figure.png" pageId="2" pageNumber="233">Fig. 13</figureCitation>
|
||
); valvula vulvae length
|
||
<quantity id="4CE43AA8FFC43F6EFD2BDAEB6B5EFF1A" box="[739,830,189,215]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="2" pageNumber="233" unit="mm" value="0.4">0.4 mm</quantity>
|
||
., dark brown (
|
||
<figureCitation id="13278BC8FFC43F6EFC12DAEB6C4CFF1A" box="[986,1068,189,215]" captionStart="FIGURES 13–14" captionStartId="6.[151,264,1127,1152]" captionTargetBox="[512,1076,181,1100]" captionTargetId="figure@6.[512,1076,181,1102]" captionTargetPageId="6" captionText="FIGURES 13–14. Stylogomphus thongphaphumensis sp. nov., (13) paratype female; anal appendages in dorsal view; (14) female vulvar scales in ventral view. Scale = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3758274" httpUri="https://zenodo.org/record/3758274/files/figure.png" pageId="2" pageNumber="233">Fig. 14</figureCitation>
|
||
), one-third length of S9 sternum, bilobed, with a V-shaped shallow cleft between lobes, the apices blunt and not turned outward. Measurements (mm):
|
||
<specimenCount id="9D1A5CC4FFC43F6EFF5FDB5368CCFED2" box="[151,172,261,287]" pageId="2" pageNumber="233" type="female">♀</specimenCount>
|
||
Fw 23–25, Hw 22–23; abdomen including appendages 28–30.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF63C7C5FFC43F6EFF5FDDFE6C03F80C" ID-DOI="http://doi.org/10.5281/zenodo.3758266" ID-Zenodo-Dep="3758266" box="[151,1123,1960,1985]" httpUri="https://zenodo.org/record/3758266/files/figure.png" pageId="2" pageNumber="233" startId="2.[151,264,1960,1985]" targetBox="[183,1404,306,1932]" targetPageId="2">
|
||
<paragraph id="8BA3974DFFC43F6EFF5FDDFE6C03F80C" blockId="2.[151,1123,1960,1985]" box="[151,1123,1960,1985]" pageId="2" pageNumber="233">
|
||
<emphasis id="B9684B5FFFC43F6EFF5FDDFE695AF80C" bold="true" box="[151,314,1960,1985]" pageId="2" pageNumber="233">FIGURES 1–2.</emphasis>
|
||
<taxonomicName id="4C1CECCEFFC43F6EFE89DDFF6AC0F80C" authority="Chainthong & Sartori & Boonsoong, 2020" authorityName="Chainthong & Sartori & Boonsoong" authorityYear="2020" box="[321,672,1961,1985]" class="Insecta" family="Gomphidae" genus="Stylogomphus" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="233" phylum="Arthropoda" rank="species" species="thongphaphumensis" status="sp. nov.">
|
||
<emphasis id="B9684B5FFFC43F6EFE89DDFF6AC0F80C" box="[321,672,1961,1985]" italics="true" pageId="2" pageNumber="233">Stylogomphus thongphaphumensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25BF624FFC43F6EFD6EDDFF6A97F80D" box="[678,759,1961,1984]" pageId="2" pageNumber="233" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, holotype ♂ (1) and paratype ♀ (2).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF63C7C5FFC53F6FFF5FD9F86A34FBC2" ID-DOI="http://doi.org/10.5281/zenodo.3758268" ID-Zenodo-Dep="3758268" httpUri="https://zenodo.org/record/3758268/files/figure.png" pageId="3" pageNumber="234" startId="3.[151,264,942,967]" targetBox="[243,1345,181,917]" targetPageId="3">
|
||
<paragraph id="8BA3974DFFC53F6FFF5FD9F86A34FBC2" blockId="3.[151,1437,942,1039]" pageId="3" pageNumber="234">
|
||
<emphasis id="B9684B5FFFC53F6FFF5FD9F8695BFC0A" bold="true" box="[151,315,942,967]" pageId="3" pageNumber="234">FIGURES 3–7.</emphasis>
|
||
<taxonomicName id="4C1CECCEFFC53F6FFE8AD9F96AC1FC0A" authority="Chainthong & Sartori & Boonsoong, 2020" authorityName="Chainthong & Sartori & Boonsoong" authorityYear="2020" box="[322,673,943,967]" class="Insecta" family="Gomphidae" genus="Stylogomphus" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="234" phylum="Arthropoda" rank="species" species="thongphaphumensis" status="sp. nov.">
|
||
<emphasis id="B9684B5FFFC53F6FFE8AD9F96AC1FC0A" box="[322,673,943,967]" italics="true" pageId="3" pageNumber="234">Stylogomphus thongphaphumensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25BF624FFC53F6FFD60D9E66A99FC0A" box="[680,761,944,967]" pageId="3" pageNumber="234" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, holotype male, (3) head in frontal view; (4) pterothoracic pattern, in dorsal view; (5) pterothoracic pattern, in lateral view; (6) accessory genitalia, in lateral view; (7) vesica spermalis, in lateral view. Scale = 1 mm (3, 4, 5), 0.5 mm (6, 7).
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C306C4C6FFC53F6FFF0FDE6B6D74F816" pageId="3" pageNumber="234" type="diagnosis">
|
||
<paragraph id="8BA3974DFFC53F6FFF0FDE6B6ABAF92D" blockId="3.[151,1437,1085,2012]" pageId="3" pageNumber="234">
|
||
<emphasis id="B9684B5FFFC53F6FFF0FDE6B69B0FB95" bold="true" box="[199,464,1085,1112]" pageId="3" pageNumber="234">Differential diagnosis:</emphasis>
|
||
When compared with known
|
||
<taxonomicName id="4C1CECCEFFC53F6FFCE8DE686BDDFB95" authorityName="Fraser" authorityYear="1922" box="[800,957,1086,1112]" class="Insecta" family="Gomphidae" genus="Stylogomphus" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="234" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9684B5FFFC53F6FFCE8DE686BDDFB95" box="[800,957,1086,1112]" italics="true" pageId="3" pageNumber="234">Stylogomphus</emphasis>
|
||
</taxonomicName>
|
||
species in Asia, the members of this genus can be divided into two groups by the shape of male anal appendages. The first group has long epiproct as in
|
||
<taxonomicName id="4C1CECCEFFC53F6FFA4FDE3469F1FB52" authority="Asahina, 1968" authorityName="Asahina" authorityYear="1968" class="Insecta" family="Gomphidae" genus="Stylogomphus" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="234" phylum="Arthropoda" rank="species" species="changi">
|
||
<emphasis id="B9684B5FFFC53F6FFA4FDE346883FB6D" italics="true" pageId="3" pageNumber="234">S. changi</emphasis>
|
||
Asahina, 1968
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="4C1CECCEFFC53F6FFE55DED06AF6FB6D" authority="Fraser, 1922" authorityName="Fraser" authorityYear="1922" box="[413,662,1158,1184]" class="Insecta" family="Gomphidae" genus="Stylogomphus" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="234" phylum="Arthropoda" rank="species" species="inglisi">
|
||
<emphasis id="B9684B5FFFC53F6FFE55DED06A60FB6D" box="[413,512,1158,1184]" italics="true" pageId="3" pageNumber="234">S. inglisi</emphasis>
|
||
Fraser, 1922
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="4C1CECCEFFC53F6FFD6BDED06B8EFB52" authority="Asahina, 1966" authorityName="Asahina" authorityYear="1966" box="[675,1006,1157,1184]" class="Insecta" family="Gomphidae" genus="Stylogomphus" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="234" phylum="Arthropoda" rank="species" species="ryukyuanus">
|
||
<emphasis id="B9684B5FFFC53F6FFD6BDED06B21FB6D" box="[675,833,1158,1184]" italics="true" pageId="3" pageNumber="234">S. ryukyuanus</emphasis>
|
||
Asahina, 1966
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="4C1CECCEFFC53F6FFC33DED06CF0FB6D" authorityName="Sasamoto" authorityYear="2001" box="[1019,1168,1158,1184]" class="Insecta" family="Gomphidae" genus="Stylogomphus" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="234" phylum="Arthropoda" rank="species" species="malayanus">
|
||
<emphasis id="B9684B5FFFC53F6FFC33DED06CF0FB6D" box="[1019,1168,1158,1184]" italics="true" pageId="3" pageNumber="234">S. malayanus</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="4C1CECCEFFC53F6FFB68DED0694DFB0E" authority="Yang & Davies, 1996" authorityName="Yang & Davies" authorityYear="1996" class="Insecta" family="Gomphidae" genus="Stylogomphus" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="234" phylum="Arthropoda" rank="species" species="lawrenceae">
|
||
<emphasis id="B9684B5FFFC53F6FFB68DED06D5DFB6D" box="[1184,1341,1158,1184]" italics="true" pageId="3" pageNumber="234">S. lawrenceae</emphasis>
|
||
Yang &
|
||
<bibRefCitation id="EF8DEABCFFC53F6FFF5FDEFF694DFB0E" author="Davies, D. A. L." box="[151,301,1193,1219]" pageId="3" pageNumber="234" pagination="283 - 296" refId="ref5370" refString="Yang, B & Davies, D. A. L. (1996) Two new species and one new subspecies of Gomphidae from southwestern China, with descriptions of larvae and distribution records (Anisoptera). ODΟNatologica, 25 (3), 283 - 296." type="journal article" year="1996">Davies, 1996</bibRefCitation>
|
||
</taxonomicName>
|
||
; and
|
||
<taxonomicName id="4C1CECCEFFC53F6FFEA0DEFC6AFAFB09" authority="Kompier, 2017" authorityName="Kompier" authorityYear="2017" box="[360,666,1193,1220]" class="Insecta" family="Gomphidae" genus="Stylogomphus" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="234" phylum="Arthropoda" rank="species" species="delicatus">
|
||
<emphasis id="B9684B5FFFC53F6FFEA0DEFC6988FB09" box="[360,488,1194,1220]" italics="true" pageId="3" pageNumber="234">S. delicatus</emphasis>
|
||
<bibRefCitation id="EF8DEABCFFC53F6FFE27DEFF6AFAFB09" author="Kompier, T." box="[495,666,1193,1220]" pageId="3" pageNumber="234" pagination="38 - 39" refId="ref5086" refString="https: // doi. org / 10.1080 / 13887890.2004.9748190 Kompier, T. (2017) Two new species of Stylogomphus Fraser, 1922 from Vietnam (Odonata: Gomphidae). Tombo, 59, 46 - 52. Matsuki, K. (1988) Notes on the larva of Stylogomphus from Thailand. Gekkan-Mushi, 212, 38 - 39." type="journal article" year="2017">Kompier, 2017</bibRefCitation>
|
||
</taxonomicName>
|
||
. The second group has short epiproct as in
|
||
<taxonomicName id="4C1CECCEFFC53F6FFBB0DEFC6DF7FB0E" authority="Asahina, 1966" authorityName="Asahina" authorityYear="1966" box="[1144,1431,1193,1220]" class="Insecta" family="Gomphidae" genus="Stylogomphus" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="234" phylum="Arthropoda" rank="species" species="shirozui">
|
||
<emphasis id="B9684B5FFFC53F6FFBB0DEFC6C8DFB09" box="[1144,1261,1194,1220]" italics="true" pageId="3" pageNumber="234">S. shirozui</emphasis>
|
||
Asahina, 1966
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="4C1CECCEFFC53F6FFF5FDE9869C7FB2A" authority="Oguma, 1926" authorityName="Oguma" authorityYear="1926" box="[151,423,1229,1256]" class="Insecta" family="Gomphidae" genus="Stylogomphus" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="234" phylum="Arthropoda" rank="species" species="suzukii">
|
||
<emphasis id="B9684B5FFFC53F6FFF5FDE986962FB25" box="[151,258,1230,1256]" italics="true" pageId="3" pageNumber="234">S. suzukii</emphasis>
|
||
Oguma, 1926
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="4C1CECCEFFC53F6FFE7CDE986AADFB25" authority="Chao, 1954" authorityName="Chao" authorityYear="1954" box="[436,717,1229,1256]" class="Insecta" family="Gomphidae" genus="Stylogomphus" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="234" phylum="Arthropoda" rank="species" species="chunliuae">
|
||
<emphasis id="B9684B5FFFC53F6FFE7CDE986A5FFB25" box="[436,575,1230,1256]" italics="true" pageId="3" pageNumber="234">S. chunliuae</emphasis>
|
||
Chao, 1954
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="4C1CECCEFFC53F6FFD12DE986BBFFB25" authority="Chao, 1954" authorityName="Chao" authorityYear="1954" box="[730,991,1229,1256]" class="Insecta" family="Gomphidae" genus="Stylogomphus" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="234" phylum="Arthropoda" rank="species" species="tantulus">
|
||
<emphasis id="B9684B5FFFC53F6FFD12DE986B31FB25" box="[730,849,1230,1256]" italics="true" pageId="3" pageNumber="234">S. tantulus</emphasis>
|
||
Chao, 1954
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="4C1CECCEFFC53F6FFC24DE986D2AFB25" authority=", Kompier, 2017" authorityName=", Kompier" authorityYear="2017" box="[1004,1354,1229,1256]" class="Insecta" family="Gomphidae" genus="Stylogomphus" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="234" phylum="Arthropoda" rank="species" species="annamensis">
|
||
<emphasis id="B9684B5FFFC53F6FFC24DE986CF5FB2A" box="[1004,1173,1230,1256]" italics="true" pageId="3" pageNumber="234">S. annamensis,</emphasis>
|
||
<bibRefCitation id="EF8DEABCFFC53F6FFB55DE9B6D2AFB25" author="Kompier, T." box="[1181,1354,1229,1256]" pageId="3" pageNumber="234" pagination="38 - 39" refId="ref5086" refString="https: // doi. org / 10.1080 / 13887890.2004.9748190 Kompier, T. (2017) Two new species of Stylogomphus Fraser, 1922 from Vietnam (Odonata: Gomphidae). Tombo, 59, 46 - 52. Matsuki, K. (1988) Notes on the larva of Stylogomphus from Thailand. Gekkan-Mushi, 212, 38 - 39." type="journal article" year="2017">Kompier, 2017</bibRefCitation>
|
||
</taxonomicName>
|
||
; and
|
||
<taxonomicName id="4C1CECCEFFC53F6FFA4FDE986917FAC1" authorityName="Chainthong & Sartori & Boonsoong" authorityYear="2020" class="Insecta" family="Gomphidae" genus="Stylogomphus" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="234" phylum="Arthropoda" rank="species" species="thongphaphumensis">
|
||
<emphasis id="B9684B5FFFC53F6FFA4FDE986917FAC1" italics="true" pageId="3" pageNumber="234">S. thongphaphumensis</emphasis>
|
||
</taxonomicName>
|
||
. The comparison between the new species and other known
|
||
<taxonomicName id="4C1CECCEFFC53F6FFBF0DEA46CB5FAC1" authorityName="Fraser" authorityYear="1922" box="[1080,1237,1266,1292]" class="Insecta" family="Gomphidae" genus="Stylogomphus" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="234" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9684B5FFFC53F6FFBF0DEA46CB5FAC1" box="[1080,1237,1266,1292]" italics="true" pageId="3" pageNumber="234">Stylogomphus</emphasis>
|
||
</taxonomicName>
|
||
species indicates close morphological similarity between
|
||
<taxonomicName id="4C1CECCEFFC53F6FFD95DF406AB3FAFD" authorityName="Asahina" authorityYear="1966" box="[605,723,1302,1328]" class="Insecta" family="Gomphidae" genus="Stylogomphus" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="234" phylum="Arthropoda" rank="species" species="shirozui">
|
||
<emphasis id="B9684B5FFFC53F6FFD95DF406A0BFAFD" box="[605,619,1302,1328]" italics="true" pageId="3" pageNumber="234">S</emphasis>
|
||
.
|
||
<emphasis id="B9684B5FFFC53F6FFDB2DF406AB3FAFD" box="[634,723,1302,1328]" italics="true" pageId="3" pageNumber="234">shirozui</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C1CECCEFFC53F6FFCC4DF406BE3FAFD" authorityName="Chao" authorityYear="1954" box="[780,899,1302,1328]" class="Insecta" family="Gomphidae" genus="Stylogomphus" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="234" phylum="Arthropoda" rank="species" species="tantulus">
|
||
<emphasis id="B9684B5FFFC53F6FFCC4DF406BE3FAFD" box="[780,899,1302,1328]" italics="true" pageId="3" pageNumber="234">S. tantulus</emphasis>
|
||
</taxonomicName>
|
||
in the shape of cerci and epiproct, but epiproct of
|
||
<taxonomicName id="4C1CECCEFFC53F6FFF7FDF6C694EFA99" authorityName="Asahina" authorityYear="1966" box="[183,302,1338,1364]" class="Insecta" family="Gomphidae" genus="Stylogomphus" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="234" phylum="Arthropoda" rank="species" species="shirozui">
|
||
<emphasis id="B9684B5FFFC53F6FFF7FDF6C68A5FA99" box="[183,197,1338,1364]" italics="true" pageId="3" pageNumber="234">S</emphasis>
|
||
.
|
||
<emphasis id="B9684B5FFFC53F6FFF1DDF6C694EFA99" box="[213,302,1338,1364]" italics="true" pageId="3" pageNumber="234">shirozui</emphasis>
|
||
</taxonomicName>
|
||
has pointed tips, and the pterothorax bears Y-shaped markings. The epiproct of
|
||
<taxonomicName id="4C1CECCEFFC53F6FFB0FDF6C6D5FFA99" authorityName="Chao" authorityYear="1954" box="[1223,1343,1338,1364]" class="Insecta" family="Gomphidae" genus="Stylogomphus" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="234" phylum="Arthropoda" rank="species" species="tantulus">
|
||
<emphasis id="B9684B5FFFC53F6FFB0FDF6C6D5FFA99" box="[1223,1343,1338,1364]" italics="true" pageId="3" pageNumber="234">S. tantulus</emphasis>
|
||
</taxonomicName>
|
||
is long, approximately half the length of the cerci, lateral lobes diverging more than
|
||
<taxonomicName id="4C1CECCEFFC53F6FFBCFDF086D66FAB5" authorityName="Chainthong & Sartori & Boonsoong" authorityYear="2020" box="[1031,1286,1374,1400]" class="Insecta" family="Gomphidae" genus="Stylogomphus" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="234" phylum="Arthropoda" rank="species" species="thongphaphumensis">
|
||
<emphasis id="B9684B5FFFC53F6FFBCFDF086D66FAB5" box="[1031,1286,1374,1400]" italics="true" pageId="3" pageNumber="234">S. thongphaphumensis</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EF8DEABCFFC53F6FFADEDF0B6DF7FAB5" author="Chao, H. - F." box="[1302,1431,1373,1400]" pageId="3" pageNumber="234" refId="ref4999" refString="Chao, H. - F. (1990) The gomphid dragonflies of China (Odonata: Gomphidae). The Science and Technology Publishing House," type="book" year="1990">Chao 1990</bibRefCitation>
|
||
; Yang &
|
||
<bibRefCitation id="EF8DEABCFFC53F6FFF31DFD769EAFA56" author="Davies, D. A. L." box="[249,394,1409,1435]" pageId="3" pageNumber="234" pagination="283 - 296" refId="ref5370" refString="Yang, B & Davies, D. A. L. (1996) Two new species and one new subspecies of Gomphidae from southwestern China, with descriptions of larvae and distribution records (Anisoptera). ODΟNatologica, 25 (3), 283 - 296." type="journal article" year="1996">Davies 1996</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF8DEABCFFC53F6FFE50DFD46A29FA51" author="Sasamoto, A." box="[408,585,1410,1436]" pageId="3" pageNumber="234" pagination="14 - 18" refId="ref5221" refString="Sasamoto, A. (2001) Description of a new subspecies of Stylogomphus lawrenceae Yang et Davies, 1996 from the Malay Peninsula (Anisoptera: Gomphidae). Tombo, 43, 14 - 18." type="journal article" year="2001">Sasamoto 2001</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EF8DEABCFFC53F6FFD9EDFD46AF2FA51" author="Sasamoto, A." box="[598,658,1410,1436]" pageId="3" pageNumber="234" pagination="27 - 30" refId="ref5259" refString="Sasamoto, A. (2004) On the true taxonomic status of Stylogomphus lawrenceae malayanus (Anisoptera: Gomphidae). Tombo, 47, 27 - 30." type="journal article" year="2004">2004</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF8DEABCFFC53F6FFD56DFD76B26FA51" author="Kompier, T." box="[670,838,1409,1436]" pageId="3" pageNumber="234" pagination="38 - 39" refId="ref5086" refString="https: // doi. org / 10.1080 / 13887890.2004.9748190 Kompier, T. (2017) Two new species of Stylogomphus Fraser, 1922 from Vietnam (Odonata: Gomphidae). Tombo, 59, 46 - 52. Matsuki, K. (1988) Notes on the larva of Stylogomphus from Thailand. Gekkan-Mushi, 212, 38 - 39." type="journal article" year="2017">Kompier 2017</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF8DEABCFFC53F6FFC9CDFD76BBCFA51" author="Zhang, H." box="[852,988,1409,1436]" pageId="3" pageNumber="234" refId="ref5420" refString="Zhang, H. (2019) Dragonflies and Damselflies of China. Chongqing University Press, Chongqing, 1460 pp." type="book" year="2019">Zhang 2019</bibRefCitation>
|
||
). Within Southeast Asian species (
|
||
<figureCitation id="13278BC8FFC53F6FFAA2DFD76881FA0D" captionStart="FIGURES 25–39" captionStartId="10.[151,264,1945,1970]" captionTargetBox="[349,1238,190,1917]" captionTargetId="figure@10.[340,1248,181,1919]" captionTargetPageId="10" captionText="FIGURES 25–39. Comparison of male anal appendages of Stylogomphus spp.: (25–27) S. thongphaphumensis sp. nov.; (28– 30) S. annamensis (redrawn from Kompier 2017); (31–33) S. delicatus (redrawn from Kompier 2017); (34–36) S. lawrenceae (redrawn from Sasamoto 2004); (37–39) S. malayanus (redrawn from Sasamoto 2004)." figureDoi="http://doi.org/10.5281/zenodo.3758282" httpUri="https://zenodo.org/record/3758282/files/figure.png" pageId="3" pageNumber="234">Figs 25–39</figureCitation>
|
||
),
|
||
<taxonomicName id="4C1CECCEFFC53F6FFF3DDFF06991FA0D" authorityName="Chainthong & Sartori & Boonsoong" authorityYear="2020" box="[245,497,1446,1472]" class="Insecta" family="Gomphidae" genus="Stylogomphus" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="234" phylum="Arthropoda" rank="species" species="thongphaphumensis">
|
||
<emphasis id="B9684B5FFFC53F6FFF3DDFF06991FA0D" box="[245,497,1446,1472]" italics="true" pageId="3" pageNumber="234">S. thongphaphumensis</emphasis>
|
||
</taxonomicName>
|
||
is somewhat more similar to
|
||
<taxonomicName id="4C1CECCEFFC53F6FFCF4DFF06BBCFA72" authorityName=", Kompier" authorityYear="2017" box="[828,988,1446,1472]" class="Insecta" family="Gomphidae" genus="Stylogomphus" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="234" phylum="Arthropoda" rank="species" species="annamensis">
|
||
<emphasis id="B9684B5FFFC53F6FFCF4DFF06BBCFA72" box="[828,988,1446,1472]" italics="true" pageId="3" pageNumber="234">S. annamensis</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry id="F30BD7DDFFC53F6FFBE9DFF36CE0FA72" box="[1057,1152,1445,1471]" name="Vietnam" pageId="3" pageNumber="234">Vietnam</collectingCountry>
|
||
than it is to other species (
|
||
<figureCitation id="13278BC8FFC53F6FFF57DF9F6943FA29" box="[159,291,1481,1508]" captionStart="FIGURES 25–39" captionStartId="10.[151,264,1945,1970]" captionTargetBox="[349,1238,190,1917]" captionTargetId="figure@10.[340,1248,181,1919]" captionTargetPageId="10" captionText="FIGURES 25–39. Comparison of male anal appendages of Stylogomphus spp.: (25–27) S. thongphaphumensis sp. nov.; (28– 30) S. annamensis (redrawn from Kompier 2017); (31–33) S. delicatus (redrawn from Kompier 2017); (34–36) S. lawrenceae (redrawn from Sasamoto 2004); (37–39) S. malayanus (redrawn from Sasamoto 2004)." figureDoi="http://doi.org/10.5281/zenodo.3758282" httpUri="https://zenodo.org/record/3758282/files/figure.png" pageId="3" pageNumber="234">Figs 28–30</figureCitation>
|
||
), closely similar in the details of the shape of cerci, strong lateral basal tooth and strong constriction just past half their length, but
|
||
<taxonomicName id="4C1CECCEFFC53F6FFE25DFB86A8BF9C5" authorityName="Chainthong & Sartori & Boonsoong" authorityYear="2020" box="[493,747,1518,1544]" class="Insecta" family="Gomphidae" genus="Stylogomphus" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="234" phylum="Arthropoda" rank="species" species="thongphaphumensis">
|
||
<emphasis id="B9684B5FFFC53F6FFE25DFB86A8BF9C5" box="[493,747,1518,1544]" italics="true" pageId="3" pageNumber="234">S. thongphaphumensis</emphasis>
|
||
</taxonomicName>
|
||
is readily distinguishable from other species by the shape of the epiproct. The epiproct of
|
||
<taxonomicName id="4C1CECCEFFC53F6FFE16DC446ABAF9E1" authorityName="Chainthong & Sartori & Boonsoong" authorityYear="2020" box="[478,730,1554,1580]" class="Insecta" family="Gomphidae" genus="Stylogomphus" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="234" phylum="Arthropoda" rank="species" species="thongphaphumensis">
|
||
<emphasis id="B9684B5FFFC53F6FFE16DC446ABAF9E1" box="[478,730,1554,1580]" italics="true" pageId="3" pageNumber="234">S. thongphaphumensis</emphasis>
|
||
</taxonomicName>
|
||
is wider than long, with lateral lobes diverging with blunt tips, the median cleft U-shaped and shallow (
|
||
<figureCitation id="13278BC8FFC53F6FFD95DC636A80F99D" box="[605,736,1589,1616]" captionStart="FIGURES 25–39" captionStartId="10.[151,264,1945,1970]" captionTargetBox="[349,1238,190,1917]" captionTargetId="figure@10.[340,1248,181,1919]" captionTargetPageId="10" captionText="FIGURES 25–39. Comparison of male anal appendages of Stylogomphus spp.: (25–27) S. thongphaphumensis sp. nov.; (28– 30) S. annamensis (redrawn from Kompier 2017); (31–33) S. delicatus (redrawn from Kompier 2017); (34–36) S. lawrenceae (redrawn from Sasamoto 2004); (37–39) S. malayanus (redrawn from Sasamoto 2004)." figureDoi="http://doi.org/10.5281/zenodo.3758282" httpUri="https://zenodo.org/record/3758282/files/figure.png" pageId="3" pageNumber="234">Figs 25–27</figureCitation>
|
||
); a short epiproct, shallowly incised into two rounded lobes in
|
||
<taxonomicName id="4C1CECCEFFC53F6FFF5FDC0C6956F9BE" authorityName=", Kompier" authorityYear="2017" box="[151,310,1626,1652]" class="Insecta" family="Gomphidae" genus="Stylogomphus" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="234" phylum="Arthropoda" rank="species" species="annamensis">
|
||
<emphasis id="B9684B5FFFC53F6FFF5FDC0C6956F9BE" box="[151,310,1626,1652]" italics="true" pageId="3" pageNumber="234">S. annamensis</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13278BC8FFC53F6FFE8CDC0F69A9F9B9" box="[324,457,1625,1652]" captionStart="FIGURES 25–39" captionStartId="10.[151,264,1945,1970]" captionTargetBox="[349,1238,190,1917]" captionTargetId="figure@10.[340,1248,181,1919]" captionTargetPageId="10" captionText="FIGURES 25–39. Comparison of male anal appendages of Stylogomphus spp.: (25–27) S. thongphaphumensis sp. nov.; (28– 30) S. annamensis (redrawn from Kompier 2017); (31–33) S. delicatus (redrawn from Kompier 2017); (34–36) S. lawrenceae (redrawn from Sasamoto 2004); (37–39) S. malayanus (redrawn from Sasamoto 2004)." figureDoi="http://doi.org/10.5281/zenodo.3758282" httpUri="https://zenodo.org/record/3758282/files/figure.png" pageId="3" pageNumber="234">Figs 28–30</figureCitation>
|
||
).; the epiproct of
|
||
<taxonomicName id="4C1CECCEFFC53F6FFD4EDC0C6B66F9B9" authorityName="Kompier" authorityYear="2017" box="[646,774,1626,1652]" class="Insecta" family="Gomphidae" genus="Stylogomphus" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="234" phylum="Arthropoda" rank="species" species="delicatus">
|
||
<emphasis id="B9684B5FFFC53F6FFD4EDC0C6B66F9B9" box="[646,774,1626,1652]" italics="true" pageId="3" pageNumber="234">S. delicatus</emphasis>
|
||
</taxonomicName>
|
||
is only a little shorter than the cerci, deeply incised to about 3/5 length, resulting in two pointed halves, the incision parallel-sided and the outer edges convergent (
|
||
<figureCitation id="13278BC8FFC53F6FFAC5DC2B6DEFF955" box="[1293,1423,1661,1688]" captionStart="FIGURES 25–39" captionStartId="10.[151,264,1945,1970]" captionTargetBox="[349,1238,190,1917]" captionTargetId="figure@10.[340,1248,181,1919]" captionTargetPageId="10" captionText="FIGURES 25–39. Comparison of male anal appendages of Stylogomphus spp.: (25–27) S. thongphaphumensis sp. nov.; (28– 30) S. annamensis (redrawn from Kompier 2017); (31–33) S. delicatus (redrawn from Kompier 2017); (34–36) S. lawrenceae (redrawn from Sasamoto 2004); (37–39) S. malayanus (redrawn from Sasamoto 2004)." figureDoi="http://doi.org/10.5281/zenodo.3758282" httpUri="https://zenodo.org/record/3758282/files/figure.png" pageId="3" pageNumber="234">Figs 31–33</figureCitation>
|
||
); deeply incised to about 2/3 of length and diverged apically in
|
||
<taxonomicName id="4C1CECCEFFC53F6FFCF7DCF46BBAF971" authorityName="Yang & Davies" authorityYear="1996" box="[831,986,1698,1724]" class="Insecta" family="Gomphidae" genus="Stylogomphus" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="234" phylum="Arthropoda" rank="species" species="lawrenceae">
|
||
<emphasis id="B9684B5FFFC53F6FFCF7DCF46BBAF971" box="[831,986,1698,1724]" italics="true" pageId="3" pageNumber="234">S. lawrenceae</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13278BC8FFC53F6FFC20DCF76C09F971" box="[1000,1129,1697,1724]" captionStart="FIGURES 25–39" captionStartId="10.[151,264,1945,1970]" captionTargetBox="[349,1238,190,1917]" captionTargetId="figure@10.[340,1248,181,1919]" captionTargetPageId="10" captionText="FIGURES 25–39. Comparison of male anal appendages of Stylogomphus spp.: (25–27) S. thongphaphumensis sp. nov.; (28– 30) S. annamensis (redrawn from Kompier 2017); (31–33) S. delicatus (redrawn from Kompier 2017); (34–36) S. lawrenceae (redrawn from Sasamoto 2004); (37–39) S. malayanus (redrawn from Sasamoto 2004)." figureDoi="http://doi.org/10.5281/zenodo.3758282" httpUri="https://zenodo.org/record/3758282/files/figure.png" pageId="3" pageNumber="234">Figs 34–36</figureCitation>
|
||
); and relatively straight and not diverged apically in
|
||
<taxonomicName id="4C1CECCEFFC53F6FFE6FDC906A5CF92D" authorityName="Sasamoto" authorityYear="2001" box="[423,572,1734,1760]" class="Insecta" family="Gomphidae" genus="Stylogomphus" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="234" phylum="Arthropoda" rank="species" species="malayanus">
|
||
<emphasis id="B9684B5FFFC53F6FFE6FDC906A5CF92D" box="[423,572,1734,1760]" italics="true" pageId="3" pageNumber="234">S. malayanus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13278BC8FFC53F6FFD83DC936AAEF92D" box="[587,718,1733,1760]" captionStart="FIGURES 25–39" captionStartId="10.[151,264,1945,1970]" captionTargetBox="[349,1238,190,1917]" captionTargetId="figure@10.[340,1248,181,1919]" captionTargetPageId="10" captionText="FIGURES 25–39. Comparison of male anal appendages of Stylogomphus spp.: (25–27) S. thongphaphumensis sp. nov.; (28– 30) S. annamensis (redrawn from Kompier 2017); (31–33) S. delicatus (redrawn from Kompier 2017); (34–36) S. lawrenceae (redrawn from Sasamoto 2004); (37–39) S. malayanus (redrawn from Sasamoto 2004)." figureDoi="http://doi.org/10.5281/zenodo.3758282" httpUri="https://zenodo.org/record/3758282/files/figure.png" pageId="3" pageNumber="234">Figs 37–39</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA3974DFFC53F6FFF0FDCBC6D74F816" blockId="3.[151,1437,1085,2012]" pageId="3" pageNumber="234">
|
||
In addition, females of
|
||
<taxonomicName id="4C1CECCEFFC53F6FFE0FDCBC6A01F8C9" authorityName="Yang & Davies" authorityYear="1996" box="[455,609,1770,1796]" class="Insecta" family="Gomphidae" genus="Stylogomphus" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="234" phylum="Arthropoda" rank="species" species="lawrenceae">
|
||
<emphasis id="B9684B5FFFC53F6FFE0FDCBC6A01F8C9" box="[455,609,1770,1796]" italics="true" pageId="3" pageNumber="234">S. lawrenceae</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C1CECCEFFC53F6FFDA6DCBC6B6DF8CE" authorityName=", Kompier" authorityYear="2017" box="[622,781,1770,1796]" class="Insecta" family="Gomphidae" genus="Stylogomphus" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="234" phylum="Arthropoda" rank="species" species="annamensis">
|
||
<emphasis id="B9684B5FFFC53F6FFDA6DCBC6B6DF8CE" box="[622,781,1770,1796]" italics="true" pageId="3" pageNumber="234">S. annamensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C1CECCEFFC53F6FFC89DCBC6BB5F8C9" authorityName="Sasamoto" authorityYear="2001" box="[833,981,1770,1796]" class="Insecta" family="Gomphidae" genus="Stylogomphus" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="234" phylum="Arthropoda" rank="species" species="malayanus">
|
||
<emphasis id="B9684B5FFFC53F6FFC89DCBC6BB5F8C9" box="[833,981,1770,1796]" italics="true" pageId="3" pageNumber="234">S. malayanus</emphasis>
|
||
</taxonomicName>
|
||
are known to date, whereas females of
|
||
<taxonomicName id="4C1CECCEFFC53F6FFA4FDCBC689CF8E5" authorityName="Kompier" authorityYear="2017" class="Insecta" family="Gomphidae" genus="Stylogomphus" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="234" phylum="Arthropoda" rank="species" species="delicatus">
|
||
<emphasis id="B9684B5FFFC53F6FFA4FDCBC689CF8E5" italics="true" pageId="3" pageNumber="234">S. delicatus</emphasis>
|
||
</taxonomicName>
|
||
are still unknown (Yang &
|
||
<bibRefCitation id="EF8DEABCFFC53F6FFDF8DD5B6AD8F8EA" author="Davies, D. A. L." box="[560,696,1805,1831]" pageId="3" pageNumber="234" pagination="283 - 296" refId="ref5370" refString="Yang, B & Davies, D. A. L. (1996) Two new species and one new subspecies of Gomphidae from southwestern China, with descriptions of larvae and distribution records (Anisoptera). ODΟNatologica, 25 (3), 283 - 296." type="journal article" year="1996">Davies1996</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF8DEABCFFC53F6FFD0BDD586B12F8E5" author="Sasamoto, A." box="[707,882,1806,1832]" pageId="3" pageNumber="234" pagination="14 - 18" refId="ref5221" refString="Sasamoto, A. (2001) Description of a new subspecies of Stylogomphus lawrenceae Yang et Davies, 1996 from the Malay Peninsula (Anisoptera: Gomphidae). Tombo, 43, 14 - 18." type="journal article" year="2001">Sasamoto 2001</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EF8DEABCFFC53F6FFCB4DD586BD8F8E5" author="Sasamoto, A." box="[892,952,1806,1832]" pageId="3" pageNumber="234" pagination="27 - 30" refId="ref5259" refString="Sasamoto, A. (2004) On the true taxonomic status of Stylogomphus lawrenceae malayanus (Anisoptera: Gomphidae). Tombo, 47, 27 - 30." type="journal article" year="2004">2004</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF8DEABCFFC53F6FFC0ADD5B6C2BF8E5" author="Zhang, H." box="[962,1099,1805,1832]" pageId="3" pageNumber="234" refId="ref5420" refString="Zhang, H. (2019) Dragonflies and Damselflies of China. Chongqing University Press, Chongqing, 1460 pp." type="book" year="2019">Zhang 2019</bibRefCitation>
|
||
, p. 465). Although the male of
|
||
<taxonomicName id="4C1CECCEFFC53F6FFF5FDD6469F3F881" authorityName="Chainthong & Sartori & Boonsoong" authorityYear="2020" box="[151,403,1842,1868]" class="Insecta" family="Gomphidae" genus="Stylogomphus" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="234" phylum="Arthropoda" rank="species" species="thongphaphumensis">
|
||
<emphasis id="B9684B5FFFC53F6FFF5FDD6469F3F881" box="[151,403,1842,1868]" italics="true" pageId="3" pageNumber="234">S. thongphaphumensis</emphasis>
|
||
</taxonomicName>
|
||
is closer to
|
||
<taxonomicName id="4C1CECCEFFC53F6FFDD3DD646ADBF886" authorityName=", Kompier" authorityYear="2017" box="[539,699,1842,1868]" class="Insecta" family="Gomphidae" genus="Stylogomphus" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="234" phylum="Arthropoda" rank="species" species="annamensis">
|
||
<emphasis id="B9684B5FFFC53F6FFDD3DD646ADBF886" box="[539,699,1842,1868]" italics="true" pageId="3" pageNumber="234">S. annamensis</emphasis>
|
||
</taxonomicName>
|
||
than to any other Southeast Asian species, the females are closest to
|
||
<taxonomicName id="4C1CECCEFFC53F6FFF7CDD00692AF8BD" authorityName="Sasamoto" authorityYear="2001" box="[180,330,1878,1904]" class="Insecta" family="Gomphidae" genus="Stylogomphus" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="234" phylum="Arthropoda" rank="species" species="malayanus">
|
||
<emphasis id="B9684B5FFFC53F6FFF7CDD00692AF8BD" box="[180,330,1878,1904]" italics="true" pageId="3" pageNumber="234">S. malayanus</emphasis>
|
||
</taxonomicName>
|
||
, which has the occiput without a Y-shaped horn-like protuberance. In contrast, the occiput of both
|
||
<taxonomicName id="4C1CECCEFFC53F6FFF5FDD2C6953F859" authorityName="Yang & Davies" authorityYear="1996" box="[151,307,1914,1940]" class="Insecta" family="Gomphidae" genus="Stylogomphus" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="234" phylum="Arthropoda" rank="species" species="lawrenceae">
|
||
<emphasis id="B9684B5FFFC53F6FFF5FDD2C6953F859" box="[151,307,1914,1940]" italics="true" pageId="3" pageNumber="234">S. lawrenceae</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C1CECCEFFC53F6FFEA0DD2C6A67F85E" authorityName=", Kompier" authorityYear="2017" box="[360,519,1914,1940]" class="Insecta" family="Gomphidae" genus="Stylogomphus" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="234" phylum="Arthropoda" rank="species" species="annamensis">
|
||
<emphasis id="B9684B5FFFC53F6FFEA0DD2C6A67F85E" box="[360,519,1914,1940]" italics="true" pageId="3" pageNumber="234">S. annamensis</emphasis>
|
||
</taxonomicName>
|
||
has a Y-shaped horn-like protuberance. The females of
|
||
<taxonomicName id="4C1CECCEFFC53F6FFBBCDD2C6D0FF859" authorityName="Chainthong & Sartori & Boonsoong" authorityYear="2020" box="[1140,1391,1914,1940]" class="Insecta" family="Gomphidae" genus="Stylogomphus" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="234" phylum="Arthropoda" rank="species" species="thongphaphumensis">
|
||
<emphasis id="B9684B5FFFC53F6FFBBCDD2C6D0FF859" box="[1140,1391,1914,1940]" italics="true" pageId="3" pageNumber="234">S. thongphaphumensis</emphasis>
|
||
</taxonomicName>
|
||
can be separated from other Southeast Asian species by the shape of the valvula vulvae, one-third the length of the S9 sternum, with a V-shaped shallow cleft between the lobes and the apices blunt and not turned outward.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF63C7C5FFC23F68FF5FDE8168A6FADE" ID-DOI="http://doi.org/10.5281/zenodo.3758270" ID-Zenodo-Dep="3758270" httpUri="https://zenodo.org/record/3758270/files/figure.png" pageId="4" pageNumber="235" startId="4.[151,264,1239,1264]" targetBox="[219,1366,189,1205]" targetPageId="4">
|
||
<paragraph id="8BA3974DFFC23F68FF5FDE8168A6FADE" blockId="4.[151,1437,1239,1299]" pageId="4" pageNumber="235">
|
||
<emphasis id="B9684B5FFFC23F68FF5FDE81695CFB3D" bold="true" box="[151,316,1239,1264]" pageId="4" pageNumber="235">FIGURES 8–9.</emphasis>
|
||
<taxonomicName id="4C1CECCEFFC23F68FE8BDE816AC4FB22" authority="Chainthong & Sartori & Boonsoong, 2020" authorityName="Chainthong & Sartori & Boonsoong" authorityYear="2020" box="[323,676,1239,1263]" class="Insecta" family="Gomphidae" genus="Stylogomphus" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="235" phylum="Arthropoda" rank="species" species="thongphaphumensis" status="sp. nov.">
|
||
<emphasis id="B9684B5FFFC23F68FE8BDE816AC4FB22" box="[323,676,1239,1263]" italics="true" pageId="4" pageNumber="235">Stylogomphus thongphaphumensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25BF624FFC23F68FD64DE8E6A9EFB22" box="[684,766,1240,1263]" pageId="4" pageNumber="235" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, holotype male, (8) right-forewing; (9) right-hindwing. Scale = 5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C306C4C6FFC23F6AFF0FDF176DFCF929" lastPageId="6" lastPageNumber="237" pageId="4" pageNumber="235" type="description">
|
||
<paragraph id="8BA3974DFFC23F68FF0FDF17696EFA4D" blockId="4.[151,1437,1345,2020]" pageId="4" pageNumber="235">
|
||
<emphasis id="B9684B5FFFC23F68FF0FDF176AE5FA91" bold="true" box="[199,645,1345,1372]" pageId="4" pageNumber="235">Description of the last stadium larva.</emphasis>
|
||
General appearance and details of structures as shown in
|
||
<figureCitation id="13278BC8FFC23F68FADFDF146DF8FA91" box="[1303,1432,1346,1372]" captionStart-0="FIGURES 15–17" captionStart-1="FIGURES 18" captionStart-2="FIGURES 19–24" captionStartId-0="7.[151,264,1910,1935]" captionStartId-1="8.[151,264,1945,1970]" captionStartId-2="9.[151,264,1309,1334]" captionTargetBox-0="[338,1250,181,1853]" captionTargetBox-1="[228,1347,189,1909]" captionTargetBox-2="[199,1390,189,1274]" captionTargetId-0="figure@7.[338,1250,181,1885]" captionTargetId-1="figure@8.[228,1360,181,1920]" captionTargetId-2="figure@9.[189,1398,181,1283]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionTargetPageId-2="9" captionText-0="FIGURES 15–17. Habitat of Stylogomphus thongphaphumensis sp. nov. larva, (15) sampling site where larvae were collected; (16) microhabitat (sandy substrate) where the larvae were found; (17) habitus of larva." captionText-1="FIGURES 18. Larva of Stylogomphus thongphaphumensis sp. nov.; dorsal view. Scale = 1 mm." captionText-2="FIGURES 19–24. Larva of Stylogomphus thongphaphumensis sp. nov., (19) antenna in dorsal view; (20) right mandible in ventral views; (21) left mandible in ventral views; (22) prementum in ventral view; (23) right maxilla in ventral view; (24) anal appendages in dorsal view. Scale = 1 mm (19), 0.2 mm (20, 21, 23), 0.5 mm (22, 24)." figureDoi-0="http://doi.org/10.5281/zenodo.3758276" figureDoi-1="http://doi.org/10.5281/zenodo.3758278" figureDoi-2="http://doi.org/10.5281/zenodo.3758280" httpUri-0="https://zenodo.org/record/3758276/files/figure.png" httpUri-1="https://zenodo.org/record/3758278/files/figure.png" httpUri-2="https://zenodo.org/record/3758280/files/figure.png" pageId="4" pageNumber="235">Figs 17–24</figureCitation>
|
||
,
|
||
<figureCitation id="13278BC8FFC23F68FF5FDF30696AFA4D" box="[151,266,1382,1408]" captionStart="FIGURES 43–48" captionStartId="12.[151,264,1478,1503]" captionTargetBox="[214,1373,181,1452]" captionTargetId="figure@12.[214,1373,181,1452]" captionTargetPageId="12" captionText="FIGURES 43–48. Comparison of larvae of two Stylogomphus spp.: S. malayanus (43, 45, 47), S. thongphaphumensis sp. nov. (44, 46, 48): (43, 44) antenna in dorsal view; (45, 46) abdomen in dorsal views; (47, 48) prementum in ventral view. Scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3758286" httpUri="https://zenodo.org/record/3758286/files/figure.png" pageId="4" pageNumber="235">44, 46, 48</figureCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BA3974DFFC23F68FF0FDFDC6889FA05" blockId="4.[151,1437,1345,2020]" pageId="4" pageNumber="235">
|
||
<emphasis id="B9684B5FFFC23F68FF0FDFDC6930FA69" box="[199,336,1418,1444]" italics="true" pageId="4" pageNumber="235">Colouration</emphasis>
|
||
. Larvae uniformly dark brown and black; only the end of femora, tibiae and tarsi with alternating yellow.
|
||
</paragraph>
|
||
<paragraph id="8BA3974DFFC23F68FF0FDF846BF5F909" blockId="4.[151,1437,1345,2020]" pageId="4" pageNumber="235">
|
||
<emphasis id="B9684B5FFFC23F68FF0FDF846964FA21" box="[199,260,1490,1516]" italics="true" pageId="4" pageNumber="235">Head</emphasis>
|
||
. Head wider than long, frontal part stouter, posterior lobe of the head shorter than the eye length, eyes large and broadest across, with three large ocelli. Antennae four-segmented, first two segments small and rather circular, third segment right-angled and triangular, inner side almost linear with many short flat setae, outer side obtusely curved with a long seta, fourth segment tiny (
|
||
<figureCitation id="13278BC8FFC23F68FCC2DC686B02F995" box="[778,866,1598,1624]" captionStart="FIGURES 19–24" captionStartId="9.[151,264,1309,1334]" captionTargetBox="[199,1390,189,1274]" captionTargetId="figure@9.[189,1398,181,1283]" captionTargetPageId="9" captionText="FIGURES 19–24. Larva of Stylogomphus thongphaphumensis sp. nov., (19) antenna in dorsal view; (20) right mandible in ventral views; (21) left mandible in ventral views; (22) prementum in ventral view; (23) right maxilla in ventral view; (24) anal appendages in dorsal view. Scale = 1 mm (19), 0.2 mm (20, 21, 23), 0.5 mm (22, 24)." figureDoi="http://doi.org/10.5281/zenodo.3758280" httpUri="https://zenodo.org/record/3758280/files/figure.png" pageId="4" pageNumber="235">Figs 19</figureCitation>
|
||
,
|
||
<figureCitation id="13278BC8FFC23F68FCB8DC686BF0F995" box="[880,912,1598,1624]" captionStart="FIGURES 43–48" captionStartId="12.[151,264,1478,1503]" captionTargetBox="[214,1373,181,1452]" captionTargetId="figure@12.[214,1373,181,1452]" captionTargetPageId="12" captionText="FIGURES 43–48. Comparison of larvae of two Stylogomphus spp.: S. malayanus (43, 45, 47), S. thongphaphumensis sp. nov. (44, 46, 48): (43, 44) antenna in dorsal view; (45, 46) abdomen in dorsal views; (47, 48) prementum in ventral view. Scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3758286" httpUri="https://zenodo.org/record/3758286/files/figure.png" pageId="4" pageNumber="235">44</figureCitation>
|
||
). Mandibles as in
|
||
<figureCitation id="13278BC8FFC23F68FBABDC686C82F995" box="[1123,1250,1598,1624]" captionStart="FIGURES 19–24" captionStartId="9.[151,264,1309,1334]" captionTargetBox="[199,1390,189,1274]" captionTargetId="figure@9.[189,1398,181,1283]" captionTargetPageId="9" captionText="FIGURES 19–24. Larva of Stylogomphus thongphaphumensis sp. nov., (19) antenna in dorsal view; (20) right mandible in ventral views; (21) left mandible in ventral views; (22) prementum in ventral view; (23) right maxilla in ventral view; (24) anal appendages in dorsal view. Scale = 1 mm (19), 0.2 mm (20, 21, 23), 0.5 mm (22, 24)." figureDoi="http://doi.org/10.5281/zenodo.3758280" httpUri="https://zenodo.org/record/3758280/files/figure.png" pageId="4" pageNumber="235">Figs 20–21</figureCitation>
|
||
and mandibular formula: L 1234 0 a(m1–5 or m1–6)b / R 1234 y a(m1–3 or m1–4)b with a>b in both mandibles (
|
||
<figureCitation id="13278BC8FFC23F68FB03DC346D2EF9B1" box="[1227,1358,1634,1660]" captionStart="FIGURES 19–24" captionStartId="9.[151,264,1309,1334]" captionTargetBox="[199,1390,189,1274]" captionTargetId="figure@9.[189,1398,181,1283]" captionTargetPageId="9" captionText="FIGURES 19–24. Larva of Stylogomphus thongphaphumensis sp. nov., (19) antenna in dorsal view; (20) right mandible in ventral views; (21) left mandible in ventral views; (22) prementum in ventral view; (23) right maxilla in ventral view; (24) anal appendages in dorsal view. Scale = 1 mm (19), 0.2 mm (20, 21, 23), 0.5 mm (22, 24)." figureDoi="http://doi.org/10.5281/zenodo.3758280" httpUri="https://zenodo.org/record/3758280/files/figure.png" pageId="4" pageNumber="235">Figs 20–21</figureCitation>
|
||
). Maxillae: galeolacinia with seven moderately incurved teeth, three dorsal teeth nearly equal in length, four ventral teeth of different sizes, apical one largest; stipes and palp setose (
|
||
<figureCitation id="13278BC8FFC23F68FCFCDCFC6BE9F909" box="[820,905,1706,1732]" captionStart="FIGURES 19–24" captionStartId="9.[151,264,1309,1334]" captionTargetBox="[199,1390,189,1274]" captionTargetId="figure@9.[189,1398,181,1283]" captionTargetPageId="9" captionText="FIGURES 19–24. Larva of Stylogomphus thongphaphumensis sp. nov., (19) antenna in dorsal view; (20) right mandible in ventral views; (21) left mandible in ventral views; (22) prementum in ventral view; (23) right maxilla in ventral view; (24) anal appendages in dorsal view. Scale = 1 mm (19), 0.2 mm (20, 21, 23), 0.5 mm (22, 24)." figureDoi="http://doi.org/10.5281/zenodo.3758280" httpUri="https://zenodo.org/record/3758280/files/figure.png" pageId="4" pageNumber="235">Fig. 23</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA3974DFFC23F68FF0FDC986A87F851" blockId="4.[151,1437,1345,2020]" pageId="4" pageNumber="235">
|
||
<emphasis id="B9684B5FFFC23F68FF0FDC986944F925" box="[199,292,1742,1768]" italics="true" pageId="4" pageNumber="235">Labium.</emphasis>
|
||
Flat and not protruding when at rest (
|
||
<figureCitation id="13278BC8FFC23F68FD08DC986B77F925" box="[704,791,1742,1768]" captionStart="FIGURES 19–24" captionStartId="9.[151,264,1309,1334]" captionTargetBox="[199,1390,189,1274]" captionTargetId="figure@9.[189,1398,181,1283]" captionTargetPageId="9" captionText="FIGURES 19–24. Larva of Stylogomphus thongphaphumensis sp. nov., (19) antenna in dorsal view; (20) right mandible in ventral views; (21) left mandible in ventral views; (22) prementum in ventral view; (23) right maxilla in ventral view; (24) anal appendages in dorsal view. Scale = 1 mm (19), 0.2 mm (20, 21, 23), 0.5 mm (22, 24)." figureDoi="http://doi.org/10.5281/zenodo.3758280" httpUri="https://zenodo.org/record/3758280/files/figure.png" pageId="4" pageNumber="235">Figs 22</figureCitation>
|
||
,
|
||
<figureCitation id="13278BC8FFC23F68FCE8DC986B20F925" box="[800,832,1742,1768]" captionStart="FIGURES 43–48" captionStartId="12.[151,264,1478,1503]" captionTargetBox="[214,1373,181,1452]" captionTargetId="figure@12.[214,1373,181,1452]" captionTargetPageId="12" captionText="FIGURES 43–48. Comparison of larvae of two Stylogomphus spp.: S. malayanus (43, 45, 47), S. thongphaphumensis sp. nov. (44, 46, 48): (43, 44) antenna in dorsal view; (45, 46) abdomen in dorsal views; (47, 48) prementum in ventral view. Scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3758286" httpUri="https://zenodo.org/record/3758286/files/figure.png" pageId="4" pageNumber="235">48</figureCitation>
|
||
). Prementum-postmentum articulation reaching posterior margin of procoxae. Prementum subrectangular, longer than wide, sides convex, convergent basally, with small teeth and minute setae at lateral margins; apical margin convex, with ventral row of 4–5 short, subquadrate, reddish brown teeth and dorsal rows of whitish piliform setae on apical border; labial palp with uniformly inflexed inner edge, yellowish brown, apical lobe reddish, rounded, its internal margin with row of 5–8 equidistant teeth. Movable hook reddish brown, sharp and moderately incurved.
|
||
</paragraph>
|
||
<paragraph id="8BA3974DFFC23F69FF0FDDF0690DFF36" blockId="4.[151,1437,1345,2020]" lastBlockId="5.[151,1436,153,251]" lastPageId="5" lastPageNumber="236" pageId="4" pageNumber="235">
|
||
<emphasis id="B9684B5FFFC23F68FF0FDDF06940F80D" box="[199,288,1958,1984]" italics="true" pageId="4" pageNumber="235">Thorax.</emphasis>
|
||
Prothorax rectangular, dorsal portion raised at sides forming two crescent-shaped ridges. Wing sheaths parallel, reaching almost the distal margin of S4. Femur and tibia of forelegs almost of equal length. Middle tibia slightly longer than femur. Hind femur longer than tibia. When pressed against the abdomen, the hind leg femur reaches the base of S3. Tarsal formula 2–2–3, tarsi yellowish. Minute short rows of setae scattered along femur and tibia of all six legs.
|
||
</paragraph>
|
||
<caption id="DF63C7C5FFC33F69FF5FDD236A7CF87C" ID-DOI="http://doi.org/10.5281/zenodo.3758272" ID-Zenodo-Dep="3758272" httpUri="https://zenodo.org/record/3758272/files/figure.png" pageId="5" pageNumber="236" startId="5.[151,264,1909,1934]" targetBox="[467,1121,273,1885]" targetPageId="5">
|
||
<paragraph id="8BA3974DFFC33F69FF5FDD236A7CF87C" blockId="5.[151,1436,1909,1970]" pageId="5" pageNumber="236">
|
||
<emphasis id="B9684B5FFFC33F69FF5FDD236934F843" bold="true" box="[151,340,1909,1934]" pageId="5" pageNumber="236">FIGURES 10–12.</emphasis>
|
||
<taxonomicName id="4C1CECCEFFC33F69FE91DD206AD8F843" authority="Chainthong & Sartori & Boonsoong, 2020" authorityName="Chainthong & Sartori & Boonsoong" authorityYear="2020" box="[345,696,1910,1934]" class="Insecta" family="Gomphidae" genus="Stylogomphus" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="236" phylum="Arthropoda" rank="species" species="thongphaphumensis" status="sp. nov.">
|
||
<emphasis id="B9684B5FFFC33F69FE91DD206AD8F843" box="[345,696,1910,1934]" italics="true" pageId="5" pageNumber="236">Stylogomphus thongphaphumensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25BF624FFC33F69FD76DD206B6EF840" box="[702,782,1910,1933]" pageId="5" pageNumber="236" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, anal appendages of holotype male; (10) dorsal view; (11) lateral view; (12) ventral view. Scale =1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF63C7C5FFC03F6AFF5FDE316AC2FB6E" ID-DOI="http://doi.org/10.5281/zenodo.3758274" ID-Zenodo-Dep="3758274" httpUri="https://zenodo.org/record/3758274/files/figure.png" pageId="6" pageNumber="237" startId="6.[151,264,1127,1152]" targetBox="[512,1076,181,1100]" targetPageId="6">
|
||
<paragraph id="8BA3974DFFC03F6AFF5FDE316AC2FB6E" blockId="6.[151,1436,1127,1188]" pageId="6" pageNumber="237">
|
||
<emphasis id="B9684B5FFFC03F6AFF5FDE316935FB4D" bold="true" box="[151,341,1127,1152]" pageId="6" pageNumber="237">FIGURES 13–14.</emphasis>
|
||
<taxonomicName id="4C1CECCEFFC03F6AFE94DE3E6ADCFB4D" authority="Chainthong & Sartori & Boonsoong, 2020" authorityName="Chainthong & Sartori & Boonsoong" authorityYear="2020" box="[348,700,1128,1152]" class="Insecta" family="Gomphidae" genus="Stylogomphus" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="237" phylum="Arthropoda" rank="species" species="thongphaphumensis" status="sp. nov.">
|
||
<emphasis id="B9684B5FFFC03F6AFE94DE3E6ADCFB4D" box="[348,700,1128,1152]" italics="true" pageId="6" pageNumber="237">Stylogomphus thongphaphumensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25BF624FFC03F6AFD0BDE3E6B74FBB2" box="[707,788,1128,1151]" pageId="6" pageNumber="237" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, (13) paratype female; anal appendages in dorsal view; (14) female vulvar scales in ventral view. Scale = 0.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BA3974DFFC03F6AFF0FDE846DFCFAF9" blockId="6.[151,1437,1234,2016]" pageId="6" pageNumber="237">
|
||
<emphasis id="B9684B5FFFC03F6AFF0FDE846951FB21" box="[199,305,1234,1260]" italics="true" pageId="6" pageNumber="237">Abdomen</emphasis>
|
||
. Ovoid, dorsum granular, dark brown, with irregular dark colour markings, with a pair of black middorsal spots on S2–10 (
|
||
<figureCitation id="13278BC8FFC03F6AFE6ADEA0699AFADD" box="[418,506,1270,1296]" captionStart="FIGURES 18" captionStartId="8.[151,264,1945,1970]" captionTargetBox="[228,1347,189,1909]" captionTargetId="figure@8.[228,1360,181,1920]" captionTargetPageId="8" captionText="FIGURES 18. Larva of Stylogomphus thongphaphumensis sp. nov.; dorsal view. Scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3758278" httpUri="https://zenodo.org/record/3758278/files/figure.png" pageId="6" pageNumber="237">Figs 18</figureCitation>
|
||
,
|
||
<figureCitation id="13278BC8FFC03F6AFDC0DEA06A48FADC" box="[520,552,1270,1297]" captionStart="FIGURES 43–48" captionStartId="12.[151,264,1478,1503]" captionTargetBox="[214,1373,181,1452]" captionTargetId="figure@12.[214,1373,181,1452]" captionTargetPageId="12" captionText="FIGURES 43–48. Comparison of larvae of two Stylogomphus spp.: S. malayanus (43, 45, 47), S. thongphaphumensis sp. nov. (44, 46, 48): (43, 44) antenna in dorsal view; (45, 46) abdomen in dorsal views; (47, 48) prementum in ventral view. Scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3758286" httpUri="https://zenodo.org/record/3758286/files/figure.png" pageId="6" pageNumber="237">46</figureCitation>
|
||
). Very tiny lateral spines at S8 and S9, lacking dorsal spines. Anal appendage small (
|
||
<figureCitation id="13278BC8FFC03F6AFF2BDF4C6958FAF9" box="[227,312,1306,1332]" captionStart="FIGURES 19–24" captionStartId="9.[151,264,1309,1334]" captionTargetBox="[199,1390,189,1274]" captionTargetId="figure@9.[189,1398,181,1283]" captionTargetPageId="9" captionText="FIGURES 19–24. Larva of Stylogomphus thongphaphumensis sp. nov., (19) antenna in dorsal view; (20) right mandible in ventral views; (21) left mandible in ventral views; (22) prementum in ventral view; (23) right maxilla in ventral view; (24) anal appendages in dorsal view. Scale = 1 mm (19), 0.2 mm (20, 21, 23), 0.5 mm (22, 24)." figureDoi="http://doi.org/10.5281/zenodo.3758280" httpUri="https://zenodo.org/record/3758280/files/figure.png" pageId="6" pageNumber="237">Fig. 24</figureCitation>
|
||
); cercus about 2/3 of epiproct, paraprocts slightly longer. Epiproct triangular with tubercles, conical.
|
||
</paragraph>
|
||
<paragraph id="8BA3974DFFC03F6AFF0FDF696DE8FA6D" blockId="6.[151,1437,1234,2016]" pageId="6" pageNumber="237">Measurements (mm, n=4): Total body length 15.76–16.37; length of abdomen 9.06–9.67; abdominal maximum width 4.07–4.76; head maximum width 3.38–3.76; length of hind femur 1.94–2.17; antennae third segment length 0.90–1.02; antennae fourth segment length 0.06–0.08; epiproct length 0.49–0.66; cerci length 0.38–0.49; paraprocts length 0.56–0.78.</paragraph>
|
||
<paragraph id="8BA3974DFFC03F6AFF0FDFFC6DFCF929" blockId="6.[151,1437,1234,2016]" pageId="6" pageNumber="237">
|
||
<emphasis id="B9684B5FFFC03F6AFF0FDFFC69DCFA08" bold="true" box="[199,444,1450,1477]" pageId="6" pageNumber="237">Diagnosis of larvae:</emphasis>
|
||
Among the Southeast Asian species, the larvae of
|
||
<taxonomicName id="4C1CECCEFFC03F6AFC20DFFD6CE3FA09" authorityName="Yang & Davies" authorityYear="1996" box="[1000,1155,1450,1477]" class="Insecta" family="Gomphidae" genus="Stylogomphus" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="237" phylum="Arthropoda" rank="species" species="lawrenceae">
|
||
<emphasis id="B9684B5FFFC03F6AFC20DFFD6CE3FA09" box="[1000,1155,1450,1477]" italics="true" pageId="6" pageNumber="237">S. lawrenceae</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C1CECCEFFC03F6AFB7FDFFD6D2BFA09" authorityName="Sasamoto" authorityYear="2001" box="[1207,1355,1450,1477]" class="Insecta" family="Gomphidae" genus="Stylogomphus" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="237" phylum="Arthropoda" rank="species" species="malayanus">
|
||
<emphasis id="B9684B5FFFC03F6AFB7FDFFD6D2BFA09" box="[1207,1355,1450,1477]" italics="true" pageId="6" pageNumber="237">S. malayanus</emphasis>
|
||
</taxonomicName>
|
||
are described (Yang &
|
||
<bibRefCitation id="EF8DEABCFFC03F6AFE9DDF986985FA25" author="Davies, D. A. L." box="[341,485,1486,1512]" pageId="6" pageNumber="237" pagination="283 - 296" refId="ref5370" refString="Yang, B & Davies, D. A. L. (1996) Two new species and one new subspecies of Gomphidae from southwestern China, with descriptions of larvae and distribution records (Anisoptera). ODΟNatologica, 25 (3), 283 - 296." type="journal article" year="1996">Davies 1996</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EF8DEABCFFC03F6AFE38DF986AFCFA25" author="Sasamoto, A." box="[496,668,1486,1512]" pageId="6" pageNumber="237" pagination="14 - 18" refId="ref5221" refString="Sasamoto, A. (2001) Description of a new subspecies of Stylogomphus lawrenceae Yang et Davies, 1996 from the Malay Peninsula (Anisoptera: Gomphidae). Tombo, 43, 14 - 18." type="journal article" year="2001">Sasamoto 2001</bibRefCitation>
|
||
), whereas those of
|
||
<taxonomicName id="4C1CECCEFFC03F6AFCBDDF996C75FA25" authorityName=", Kompier" authorityYear="2017" box="[885,1045,1487,1513]" class="Insecta" family="Gomphidae" genus="Stylogomphus" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="237" phylum="Arthropoda" rank="species" species="annamensis">
|
||
<emphasis id="B9684B5FFFC03F6AFCBDDF996C75FA25" box="[885,1045,1487,1513]" italics="true" pageId="6" pageNumber="237">S. annamensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C1CECCEFFC03F6AFB83DF996CACFA25" authorityName="Kompier" authorityYear="2017" box="[1099,1228,1486,1513]" class="Insecta" family="Gomphidae" genus="Stylogomphus" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="237" phylum="Arthropoda" rank="species" species="delicatus">
|
||
<emphasis id="B9684B5FFFC03F6AFB83DF996CACFA25" box="[1099,1228,1486,1513]" italics="true" pageId="6" pageNumber="237">S. delicatus</emphasis>
|
||
</taxonomicName>
|
||
are still unknown. The larva of
|
||
<taxonomicName id="4C1CECCEFFC03F6AFEEEDFA56A42F9C1" authorityName="Chainthong & Sartori & Boonsoong" authorityYear="2020" box="[294,546,1522,1549]" class="Insecta" family="Gomphidae" genus="Stylogomphus" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="237" phylum="Arthropoda" rank="species" species="thongphaphumensis">
|
||
<emphasis id="B9684B5FFFC03F6AFEEEDFA56A42F9C1" box="[294,546,1522,1549]" italics="true" pageId="6" pageNumber="237">S. thongphaphumensis</emphasis>
|
||
</taxonomicName>
|
||
differs from known species by the presence of palpal teeth and seta on the third antennae. The third antennal segment in
|
||
<taxonomicName id="4C1CECCEFFC03F6AFDA8DC416B3DF9FD" authorityName="Chainthong & Sartori & Boonsoong" authorityYear="2020" box="[608,861,1558,1585]" class="Insecta" family="Gomphidae" genus="Stylogomphus" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="237" phylum="Arthropoda" rank="species" species="thongphaphumensis">
|
||
<emphasis id="B9684B5FFFC03F6AFDA8DC416B3DF9FD" box="[608,861,1558,1585]" italics="true" pageId="6" pageNumber="237">S. thongphaphumensis</emphasis>
|
||
</taxonomicName>
|
||
is right-angled, triangular; the inner side has many short, flat setae, the outer side has a long seta; there are 5–8 palpal teeth, and the abdomen has a pair of black middorsal spots on S2–10. (
|
||
<figureCitation id="13278BC8FFC03F6AFE60DC086A38F9B5" box="[424,600,1630,1656]" captionStart="FIGURES 43–48" captionStartId="12.[151,264,1478,1503]" captionTargetBox="[214,1373,181,1452]" captionTargetId="figure@12.[214,1373,181,1452]" captionTargetPageId="12" captionText="FIGURES 43–48. Comparison of larvae of two Stylogomphus spp.: S. malayanus (43, 45, 47), S. thongphaphumensis sp. nov. (44, 46, 48): (43, 44) antenna in dorsal view; (45, 46) abdomen in dorsal views; (47, 48) prementum in ventral view. Scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3758286" httpUri="https://zenodo.org/record/3758286/files/figure.png" pageId="6" pageNumber="237">Figs 44, 46, 48</figureCitation>
|
||
). The larva of
|
||
<taxonomicName id="4C1CECCEFFC03F6AFD37DC096BF5F9B5" authorityName="Sasamoto" authorityYear="2001" box="[767,917,1630,1657]" class="Insecta" family="Gomphidae" genus="Stylogomphus" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="237" phylum="Arthropoda" rank="species" species="malayanus">
|
||
<emphasis id="B9684B5FFFC03F6AFD37DC096BF5F9B5" box="[767,917,1630,1657]" italics="true" pageId="6" pageNumber="237">S. malayanus</emphasis>
|
||
</taxonomicName>
|
||
has a small number of short, flat setae on the inner side and a long setae on the outer side of the third antennal segment, there are 7–10 palpal teeth, and the abdomen lacks the black mid-dorsal spot (
|
||
<figureCitation id="13278BC8FFC03F6AFDF1DCF06A85F90D" box="[569,741,1701,1728]" captionStart="FIGURES 43–48" captionStartId="12.[151,264,1478,1503]" captionTargetBox="[214,1373,181,1452]" captionTargetId="figure@12.[214,1373,181,1452]" captionTargetPageId="12" captionText="FIGURES 43–48. Comparison of larvae of two Stylogomphus spp.: S. malayanus (43, 45, 47), S. thongphaphumensis sp. nov. (44, 46, 48): (43, 44) antenna in dorsal view; (45, 46) abdomen in dorsal views; (47, 48) prementum in ventral view. Scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3758286" httpUri="https://zenodo.org/record/3758286/files/figure.png" pageId="6" pageNumber="237">Figs 43, 45, 47</figureCitation>
|
||
). Conversely, the larva of
|
||
<taxonomicName id="4C1CECCEFFC03F6AFBC2DCF16CC6F90D" authorityName="Yang & Davies" authorityYear="1996" box="[1034,1190,1702,1729]" class="Insecta" family="Gomphidae" genus="Stylogomphus" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="237" phylum="Arthropoda" rank="species" species="lawrenceae">
|
||
<emphasis id="B9684B5FFFC03F6AFBC2DCF16CC6F90D" box="[1034,1190,1702,1729]" italics="true" pageId="6" pageNumber="237">S. lawrenceae</emphasis>
|
||
</taxonomicName>
|
||
has the third antennal segment with dense flat seta on the inner side and without long setae on the outer side, and there are 7 palpal teeth.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C306C4C6FFC03F6AFF0FDCBB6D49F82D" pageId="6" pageNumber="237" type="biology_ecology">
|
||
<paragraph id="8BA3974DFFC03F6AFF0FDCBB6D49F82D" blockId="6.[151,1437,1234,2016]" pageId="6" pageNumber="237">
|
||
<emphasis id="B9684B5FFFC03F6AFF0FDCBB69A4F8C5" bold="true" box="[199,452,1773,1800]" pageId="6" pageNumber="237">Habitat and Ecology.</emphasis>
|
||
The larvae of
|
||
<taxonomicName id="4C1CECCEFFC03F6AFDA0DCB96B05F8C5" authorityName="Chainthong & Sartori & Boonsoong" authorityYear="2020" box="[616,869,1774,1801]" class="Insecta" family="Gomphidae" genus="Stylogomphus" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="237" phylum="Arthropoda" rank="species" species="thongphaphumensis">
|
||
<emphasis id="B9684B5FFFC03F6AFDA0DCB96B05F8C5" box="[616,869,1774,1801]" italics="true" pageId="6" pageNumber="237">S. thongphaphumensis</emphasis>
|
||
</taxonomicName>
|
||
occur in headwater streams, where they are found in the sandy gravel substrate (
|
||
<figureCitation id="13278BC8FFC03F6AFE2BDD446A06F8E1" box="[483,614,1809,1836]" captionStart="FIGURES 15–17" captionStartId="7.[151,264,1910,1935]" captionTargetBox="[338,1250,181,1853]" captionTargetId="figure@7.[338,1250,181,1885]" captionTargetPageId="7" captionText="FIGURES 15–17. Habitat of Stylogomphus thongphaphumensis sp. nov. larva, (15) sampling site where larvae were collected; (16) microhabitat (sandy substrate) where the larvae were found; (17) habitus of larva." figureDoi="http://doi.org/10.5281/zenodo.3758276" httpUri="https://zenodo.org/record/3758276/files/figure.png" pageId="6" pageNumber="237">Figs 15–16</figureCitation>
|
||
). The stream bed was mostly composed of boulders (5%), cobbles (30%), gravel (30%), sand (30%) and leaf litter (5%). During the odonate larval sampling, environmental and physicochemical parameters were measured. The streams ranged from
|
||
<quantity id="4CE43AA8FFC03F6AFCA3DD0F6B9AF8B8" box="[875,1018,1881,1909]" metricMagnitude="0" metricUnit="m" metricValue="6.21" metricValueMax="6.9" metricValueMin="5.52" pageId="6" pageNumber="237" unit="m" value="6.21" valueMax="6.9" valueMin="5.52">5.52–6.90 m</quantity>
|
||
in width and
|
||
<quantity id="4CE43AA8FFC03F6AFB53DD0F6D4FF8B8" box="[1179,1327,1881,1909]" metricMagnitude="-1" metricUnit="m" metricValue="1.5835" metricValueMax="1.667" metricValueMin="1.5" pageId="6" pageNumber="237" unit="cm" value="15.835" valueMax="16.67" valueMin="15.0">15–16.67 cm</quantity>
|
||
in depth, the water velocity was about
|
||
<quantity id="4CE43AA8FFC03F6AFE2ADD286A11F855" box="[482,625,1918,1944]" metricMagnitude="-1" metricUnit="m" metricValue="4.8" metricValueMax="6.3" metricValueMin="3.3" pageId="6" pageNumber="237" unit="m" value="0.48" valueMax="0.63" valueMin="0.33">0.33–0.63 m</quantity>
|
||
/sec, air temperature ranged from 22.50–30.00°C, the water temperature ranged from 21.74–27.29°C, pH ranged from 6.22–7.95, dissolved oxygen (DO) ranged from
|
||
<quantity id="4CE43AA8FFC03F6AFB7EDDF46D25F871" box="[1206,1349,1954,1980]" metricMagnitude="-6" metricUnit="kg" metricValue="6.66" metricValueMax="7.12" metricValueMin="6.2" pageId="6" pageNumber="237" unit="mg" value="6.66" valueMax="7.12" valueMin="6.2">6.2–7.12 mg</quantity>
|
||
/L, total dissolved solids (TDS) ranged from
|
||
<quantity id="4CE43AA8FFC03F6AFDE7DD936B63F82D" box="[559,771,1989,2016]" metricMagnitude="-4" metricUnit="kg" metricValue="1.9616999999999998" metricValueMax="2.3266999999999998" metricValueMin="1.5966999999999998" pageId="6" pageNumber="237" unit="mg" value="196.17" valueMax="232.67" valueMin="159.67">159.67–232.67 mg</quantity>
|
||
/L, and conductivity from 252.67–372.67 µS/cm.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |