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<document ID-DOI="10.11646/zootaxa.5263.2.4" ID-GBIF-Dataset="f6701c34-91f2-48c3-9ed1-4dd17fb3fb0d" ID-ISSN="1175-5326" ID-Zenodo-Dep="7801106" ID-ZooBank="4E687740-045F-44C5-810D-68C6EE9072C4" IM.illustrations_approvedBy="diego" IM.taxonomicNames_approvedBy="diego" IM.treatments_approvedBy="diego" approvalRequired="4" approvalRequired_for_textStreams="4" checkinTime="1680678286118" checkinUser="plazi" docAuthor="Qi, Zhi-Hao, He, Hao, Su, Rong-Xiang, Ai, Hong-Mu &amp; Song, Hai-Tian" docDate="2023" docId="03B51633FF8CFFDE5FFB5CE805D8FB8F" docLanguage="en" docName="zootaxa.5263.2.4.pdf" docOrigin="Zootaxa 5263 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5263.2.4" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Lucanus boileaui Planet 1897" docType="key" docVersion="4" lastPageNumber="258" masterDocId="FF8C6E4BFF9EFFCC5F6C5E420045FFC7" masterDocTitle="Revision of the Lucanus boileaui group and notes on. takeoi Adachi, 2020 from the. maculifemoratus group (Coleoptera: Lucanidae: Lucaninae)" masterLastPageNumber="260" masterPageNumber="240" pageNumber="258" updateTime="1681158889771" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title>Revision of the Lucanus boileaui group and notes on. takeoi Adachi, 2020 from the. maculifemoratus group (Coleoptera: Lucanidae: Lucaninae)</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Qi, Zhi-Hao</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0001-8596-0913</mods:nameIdentifier>
<mods:affiliation>College of Plant Protection, Fujian Agriculture and Forestry University, Fuzhou, Fujian, 350002, China. &amp; Fujian Academy of Forestry, Fuzhou, Fujian, 350012, China. &amp; qizhihao 2021 @ 126. com; https: // orcid. org / 0000 - 0001 - 8596 - 0913</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>He, Hao</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0002-8871-9671</mods:nameIdentifier>
<mods:affiliation>College of Life Sciences, Yunnan Normal University, Kunming, Yunnan, 650500, China. &amp; 18835130675 @ 163. com; https: // orcid. org / 0000 - 0002 - 8871 - 9671</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Su, Rong-Xiang</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0002-6575-5776</mods:nameIdentifier>
<mods:affiliation>Fujian Academy of Forestry, Fuzhou, Fujian, 350012, China. &amp; surongxiang 2022 @ 126. com; https: // orcid. org / 0000 - 0002 - 6575 - 5776</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Ai, Hong-Mu</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0002-7754-8256</mods:nameIdentifier>
<mods:affiliation>College of Plant Protection, Fujian Agriculture and Forestry University, Fuzhou, Fujian, 350002, China. &amp; aihongmu @ 163. com; https: // orcid. org / 0000 - 0002 - 7754 - 8256</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Song, Hai-Tian</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0003-1042-7959</mods:nameIdentifier>
<mods:affiliation>Fujian Academy of Forestry, Fuzhou, Fujian, 350012, China. &amp; haitiansong @ 126. com; https: // orcid. org / 0000 - 0003 - 1042 - 7959</mods:affiliation>
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<mods:title>Zootaxa</mods:title>
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<mods:date>2023</mods:date>
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<mods:number>2023-04-05</mods:number>
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<mods:detail type="volume">
<mods:number>5263</mods:number>
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<mods:number>2</mods:number>
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<mods:identifier type="DOI">10.11646/zootaxa.5263.2.4</mods:identifier>
<mods:identifier type="GBIF-Dataset">f6701c34-91f2-48c3-9ed1-4dd17fb3fb0d</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.7801108" ID-Zenodo-Dep="7801108" LSID="urn:lsid:plazi:treatment:03B51633FF8CFFDE5FFB5CE805D8FB8F" httpUri="http://treatment.plazi.org/id/03B51633FF8CFFDE5FFB5CE805D8FB8F" lastPageNumber="258" pageId="18" pageNumber="258">
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<emphasis bold="true" box="[151,690,682,709]" pageId="18" pageNumber="258">
Key to males of the
<taxonomicName ID-CoL="72P3S" authorityName="Planet" authorityYear="1897" box="[398,609,682,709]" class="Insecta" family="Lucanidae" genus="Lucanus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="258" phylum="Arthropoda" rank="species" species="boileaui">
<emphasis bold="true" box="[398,609,682,709]" italics="true" pageId="18" pageNumber="258">Lucanus boileaui</emphasis>
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group
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<keyStep pageId="18" pageNumber="258">
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1. Large sized species that can reach more than
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in length; clypeolabrum tongue shaped and long (
<figureCitation box="[1210,1285,751,773]" captionStart="FIGURE 2" captionStartId="3.[151,250,1896,1920]" captionTargetBox="[151,1431,181,1873]" captionTargetId="figure-17@3.[151,1435,181,1873]" captionTargetPageId="3" captionText="FIGURE 2. Male head (A, G, M, S), clypeolabrum (B, H, N, T), canthus (C, I, O, U) and tibia (D–F, J–L, P–R, V–X) of Lucanus species in dorsal view. A–F, L. jiaozishanus Qi, He, Su &amp; Song, new species (holotype, Kunming, Yunnan); G–L, L. ludivinae Boucher, 1998 (Gongshan, Yunnan); M–R, L. takeoi Adachi, 2020 (Huidong, Sichuan); S–X, L. boileaui Planet, 1897 (Mabian, Sichuan). Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7814599" httpUri="https://zenodo.org/record/7814599/files/figure.png" pageId="18" pageNumber="258">Fig. 2T</figureCitation>
); dorsal surface of the body with long yellow brown pubescence (
<figureCitation box="[693,768,780,802]" captionStart="FIGURE 1" captionStartId="2.[151,250,1840,1864]" captionTargetBox="[158,1432,666,1804]" captionTargetId="figure-216@2.[151,1436,666,1814]" captionTargetPageId="2" captionText="FIGURE 1. Male habitus of Lucanus species. A–B, L. jiaozishanus Qi, He, Su &amp; Song, new species (33.3 mm, holotype, large sized, Kunming, Yunnan); C–D, L. ludivinae Boucher, 1998 (31.3 mm, medium sized, Gongshan, Yunnan); E–F, L. takeoi Adachi, 2020 (29.2 mm, small sized, Huidong, Sichuan); G–H, L. boileaui Planet, 1897 (41.9 mm, small sized, Mabian, Sichuan). A, C, E, G, dorsal view; B, D, F, H, ventral view. Scale bar = 10.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7801110" httpUri="https://zenodo.org/record/7801110/files/figure.png" pageId="18" pageNumber="258">Fig. 1G</figureCitation>
); all tibiae with bright yellowish brown stripes (
<figureCitation box="[1222,1329,780,802]" captionStart="FIGURE 2" captionStartId="3.[151,250,1896,1920]" captionTargetBox="[151,1431,181,1873]" captionTargetId="figure-17@3.[151,1435,181,1873]" captionTargetPageId="3" captionText="FIGURE 2. Male head (A, G, M, S), clypeolabrum (B, H, N, T), canthus (C, I, O, U) and tibia (D–F, J–L, P–R, V–X) of Lucanus species in dorsal view. A–F, L. jiaozishanus Qi, He, Su &amp; Song, new species (holotype, Kunming, Yunnan); G–L, L. ludivinae Boucher, 1998 (Gongshan, Yunnan); M–R, L. takeoi Adachi, 2020 (Huidong, Sichuan); S–X, L. boileaui Planet, 1897 (Mabian, Sichuan). Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7814599" httpUri="https://zenodo.org/record/7814599/files/figure.png" pageId="18" pageNumber="258">Fig. 2V–X</figureCitation>
); flagellum very long (
<figureCitation box="[330,407,810,832]" captionStart="FIGURE 4" captionStartId="5.[151,250,1880,1904]" captionTargetBox="[173,1423,186,1856]" captionTargetId="figure-17@5.[164,1423,181,1856]" captionTargetPageId="5" captionText="FIGURE 4. Aedeagus of Lucanus species (H, L, P, T, apex of flagellum). A, E–H, L. jiaozishanus Qi, He, Su &amp; Song, new species (holotype, Kunming, Yunnan); B, I–L, L. ludivinae Boucher, 1998 (Gongshan, Yunnan); C, M–P, L. takeoi Adachi, 2020 (Huidong, Sichuan); D, Q–T, L. boileaui Planet, 1897 (Mabian, Sichuan). F, J, N, R, dorsal view; A–D, E, I, M, Q, ventral view; G, K, O, S, lateral view. Scale bar for H, L, P, T = 0.2 mm, for the others = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7801114" httpUri="https://zenodo.org/record/7801114/files/figure.png" pageId="18" pageNumber="258">Fig. 4D</figureCitation>
), usually 8–10 times longer than aedeagus...........................
<taxonomicName authority="Planet, 1897" authorityName="Planet" authorityYear="1897" box="[1144,1436,810,832]" class="Insecta" family="Lucanidae" genus="Lucanus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="258" phylum="Arthropoda" rank="species" species="boileaui">
<emphasis box="[1144,1311,810,831]" italics="true" pageId="18" pageNumber="258">Lucanus boileaui</emphasis>
<bibRefCitation author="Planet, L." box="[1316,1436,810,832]" pageId="18" pageNumber="258" pagination="205 - 206" refId="ref8126" refString="Planet, L. (1897) Description de coleopteres nouveaux. Le Naturaliste, 19, 205 - 206." type="journal article" year="1897">Planet, 1897</bibRefCitation>
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</paragraph>
<paragraph blockId="18.[151,1437,751,1096]" pageId="18" pageNumber="258">
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– Smaller sized species than cannot reach more than
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; clypeolabrum not tongue shaped and short (
<figureCitation box="[1248,1357,839,861]" captionStart="FIGURE 2" captionStartId="3.[151,250,1896,1920]" captionTargetBox="[151,1431,181,1873]" captionTargetId="figure-17@3.[151,1435,181,1873]" captionTargetPageId="3" captionText="FIGURE 2. Male head (A, G, M, S), clypeolabrum (B, H, N, T), canthus (C, I, O, U) and tibia (D–F, J–L, P–R, V–X) of Lucanus species in dorsal view. A–F, L. jiaozishanus Qi, He, Su &amp; Song, new species (holotype, Kunming, Yunnan); G–L, L. ludivinae Boucher, 1998 (Gongshan, Yunnan); M–R, L. takeoi Adachi, 2020 (Huidong, Sichuan); S–X, L. boileaui Planet, 1897 (Mabian, Sichuan). Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7814599" httpUri="https://zenodo.org/record/7814599/files/figure.png" pageId="18" pageNumber="258">Fig. 2B, H</figureCitation>
); dorsal surface of the body with fine and visible short pubescence or nearly smooth (
<figureCitation box="[986,1095,868,890]" captionStart="FIGURE 1" captionStartId="2.[151,250,1840,1864]" captionTargetBox="[158,1432,666,1804]" captionTargetId="figure-216@2.[151,1436,666,1814]" captionTargetPageId="2" captionText="FIGURE 1. Male habitus of Lucanus species. A–B, L. jiaozishanus Qi, He, Su &amp; Song, new species (33.3 mm, holotype, large sized, Kunming, Yunnan); C–D, L. ludivinae Boucher, 1998 (31.3 mm, medium sized, Gongshan, Yunnan); E–F, L. takeoi Adachi, 2020 (29.2 mm, small sized, Huidong, Sichuan); G–H, L. boileaui Planet, 1897 (41.9 mm, small sized, Mabian, Sichuan). A, C, E, G, dorsal view; B, D, F, H, ventral view. Scale bar = 10.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7801110" httpUri="https://zenodo.org/record/7801110/files/figure.png" pageId="18" pageNumber="258">Fig. 1A, C</figureCitation>
); all tibiae black (
<figureCitation box="[1273,1426,868,890]" captionStart="FIGURE 2" captionStartId="3.[151,250,1896,1920]" captionTargetBox="[151,1431,181,1873]" captionTargetId="figure-17@3.[151,1435,181,1873]" captionTargetPageId="3" captionText="FIGURE 2. Male head (A, G, M, S), clypeolabrum (B, H, N, T), canthus (C, I, O, U) and tibia (D–F, J–L, P–R, V–X) of Lucanus species in dorsal view. A–F, L. jiaozishanus Qi, He, Su &amp; Song, new species (holotype, Kunming, Yunnan); G–L, L. ludivinae Boucher, 1998 (Gongshan, Yunnan); M–R, L. takeoi Adachi, 2020 (Huidong, Sichuan); S–X, L. boileaui Planet, 1897 (Mabian, Sichuan). Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7814599" httpUri="https://zenodo.org/record/7814599/files/figure.png" pageId="18" pageNumber="258">Fig. 2D–F, J–L</figureCitation>
), without bright yellowish brown stripes; flagellum shorter (
<figureCitation box="[788,894,898,920]" captionStart="FIGURE 4" captionStartId="5.[151,250,1880,1904]" captionTargetBox="[173,1423,186,1856]" captionTargetId="figure-17@5.[164,1423,181,1856]" captionTargetPageId="5" captionText="FIGURE 4. Aedeagus of Lucanus species (H, L, P, T, apex of flagellum). A, E–H, L. jiaozishanus Qi, He, Su &amp; Song, new species (holotype, Kunming, Yunnan); B, I–L, L. ludivinae Boucher, 1998 (Gongshan, Yunnan); C, M–P, L. takeoi Adachi, 2020 (Huidong, Sichuan); D, Q–T, L. boileaui Planet, 1897 (Mabian, Sichuan). F, J, N, R, dorsal view; A–D, E, I, M, Q, ventral view; G, K, O, S, lateral view. Scale bar for H, L, P, T = 0.2 mm, for the others = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7801114" httpUri="https://zenodo.org/record/7801114/files/figure.png" pageId="18" pageNumber="258">Fig. 4A–B</figureCitation>
), about 2–3 times longer than aedeagus............. 2
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<paragraph blockId="18.[151,1437,751,1096]" pageId="18" pageNumber="258">
<keyLead pageId="18" pageNumber="258">
2. Each mandible usually with 3–7 inner teeth (excluding apical fork), major inner tooth behind the middle, distinctly near the base, apical fork developed (
<figureCitation box="[514,595,956,978]" captionStart="FIGURE 2" captionStartId="3.[151,250,1896,1920]" captionTargetBox="[151,1431,181,1873]" captionTargetId="figure-17@3.[151,1435,181,1873]" captionTargetPageId="3" captionText="FIGURE 2. Male head (A, G, M, S), clypeolabrum (B, H, N, T), canthus (C, I, O, U) and tibia (D–F, J–L, P–R, V–X) of Lucanus species in dorsal view. A–F, L. jiaozishanus Qi, He, Su &amp; Song, new species (holotype, Kunming, Yunnan); G–L, L. ludivinae Boucher, 1998 (Gongshan, Yunnan); M–R, L. takeoi Adachi, 2020 (Huidong, Sichuan); S–X, L. boileaui Planet, 1897 (Mabian, Sichuan). Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7814599" httpUri="https://zenodo.org/record/7814599/files/figure.png" pageId="18" pageNumber="258">Fig. 2G</figureCitation>
); dorsal surface of the body with fine and visible short pubescence (
<figureCitation box="[1286,1363,956,978]" captionStart="FIGURE 1" captionStartId="2.[151,250,1840,1864]" captionTargetBox="[158,1432,666,1804]" captionTargetId="figure-216@2.[151,1436,666,1814]" captionTargetPageId="2" captionText="FIGURE 1. Male habitus of Lucanus species. A–B, L. jiaozishanus Qi, He, Su &amp; Song, new species (33.3 mm, holotype, large sized, Kunming, Yunnan); C–D, L. ludivinae Boucher, 1998 (31.3 mm, medium sized, Gongshan, Yunnan); E–F, L. takeoi Adachi, 2020 (29.2 mm, small sized, Huidong, Sichuan); G–H, L. boileaui Planet, 1897 (41.9 mm, small sized, Mabian, Sichuan). A, C, E, G, dorsal view; B, D, F, H, ventral view. Scale bar = 10.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7801110" httpUri="https://zenodo.org/record/7801110/files/figure.png" pageId="18" pageNumber="258">Fig. 1C</figureCitation>
); penis narrower (
<figureCitation box="[326,393,986,1008]" captionStart="FIGURE 4" captionStartId="5.[151,250,1880,1904]" captionTargetBox="[173,1423,186,1856]" captionTargetId="figure-17@5.[164,1423,181,1856]" captionTargetPageId="5" captionText="FIGURE 4. Aedeagus of Lucanus species (H, L, P, T, apex of flagellum). A, E–H, L. jiaozishanus Qi, He, Su &amp; Song, new species (holotype, Kunming, Yunnan); B, I–L, L. ludivinae Boucher, 1998 (Gongshan, Yunnan); C, M–P, L. takeoi Adachi, 2020 (Huidong, Sichuan); D, Q–T, L. boileaui Planet, 1897 (Mabian, Sichuan). F, J, N, R, dorsal view; A–D, E, I, M, Q, ventral view; G, K, O, S, lateral view. Scale bar for H, L, P, T = 0.2 mm, for the others = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7801114" httpUri="https://zenodo.org/record/7801114/files/figure.png" pageId="18" pageNumber="258">Fig. 4I</figureCitation>
); flagellum (
<figureCitation box="[516,590,986,1008]" captionStart="FIGURE 4" captionStartId="5.[151,250,1880,1904]" captionTargetBox="[173,1423,186,1856]" captionTargetId="figure-17@5.[164,1423,181,1856]" captionTargetPageId="5" captionText="FIGURE 4. Aedeagus of Lucanus species (H, L, P, T, apex of flagellum). A, E–H, L. jiaozishanus Qi, He, Su &amp; Song, new species (holotype, Kunming, Yunnan); B, I–L, L. ludivinae Boucher, 1998 (Gongshan, Yunnan); C, M–P, L. takeoi Adachi, 2020 (Huidong, Sichuan); D, Q–T, L. boileaui Planet, 1897 (Mabian, Sichuan). F, J, N, R, dorsal view; A–D, E, I, M, Q, ventral view; G, K, O, S, lateral view. Scale bar for H, L, P, T = 0.2 mm, for the others = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7801114" httpUri="https://zenodo.org/record/7801114/files/figure.png" pageId="18" pageNumber="258">Fig. 4B</figureCitation>
) about 2.5 times longer than aedeagus........... ..
<taxonomicName authority="Boucher, 1998" authorityName="Boucher" authorityYear="1998" box="[1113,1436,986,1008]" class="Insecta" family="Lucanidae" genus="Lucanus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="258" phylum="Arthropoda" rank="species" species="ludivinae">
<emphasis box="[1113,1290,986,1007]" italics="true" pageId="18" pageNumber="258">Lucanus ludivinae</emphasis>
<bibRefCitation author="Boucher, S." box="[1296,1436,986,1008]" pageId="18" pageNumber="258" pagination="177 - 185" refId="ref7749" refString="Boucher, S. (1998) Deux Lucanus inedits des provinces chinoises du Yunnan et du Guangxi (Coleoptera, Lucanidae). Coleopteres, 4, 177 - 185." type="journal article" year="1998">Boucher, 1998</bibRefCitation>
</taxonomicName>
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</paragraph>
<paragraph blockId="18.[151,1437,751,1096]" pageId="18" pageNumber="258">
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– Each mandible usually with 1–4 inner teeth (excluding apical fork), major inner tooth at the middle, apical fork less developed (
<figureCitation box="[233,312,1044,1066]" captionStart="FIGURE 2" captionStartId="3.[151,250,1896,1920]" captionTargetBox="[151,1431,181,1873]" captionTargetId="figure-17@3.[151,1435,181,1873]" captionTargetPageId="3" captionText="FIGURE 2. Male head (A, G, M, S), clypeolabrum (B, H, N, T), canthus (C, I, O, U) and tibia (D–F, J–L, P–R, V–X) of Lucanus species in dorsal view. A–F, L. jiaozishanus Qi, He, Su &amp; Song, new species (holotype, Kunming, Yunnan); G–L, L. ludivinae Boucher, 1998 (Gongshan, Yunnan); M–R, L. takeoi Adachi, 2020 (Huidong, Sichuan); S–X, L. boileaui Planet, 1897 (Mabian, Sichuan). Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7814599" httpUri="https://zenodo.org/record/7814599/files/figure.png" pageId="18" pageNumber="258">Fig. 2A</figureCitation>
); dorsal surface of the body nearly smooth (
<figureCitation box="[747,824,1044,1066]" captionStart="FIGURE 1" captionStartId="2.[151,250,1840,1864]" captionTargetBox="[158,1432,666,1804]" captionTargetId="figure-216@2.[151,1436,666,1814]" captionTargetPageId="2" captionText="FIGURE 1. Male habitus of Lucanus species. A–B, L. jiaozishanus Qi, He, Su &amp; Song, new species (33.3 mm, holotype, large sized, Kunming, Yunnan); C–D, L. ludivinae Boucher, 1998 (31.3 mm, medium sized, Gongshan, Yunnan); E–F, L. takeoi Adachi, 2020 (29.2 mm, small sized, Huidong, Sichuan); G–H, L. boileaui Planet, 1897 (41.9 mm, small sized, Mabian, Sichuan). A, C, E, G, dorsal view; B, D, F, H, ventral view. Scale bar = 10.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7801110" httpUri="https://zenodo.org/record/7801110/files/figure.png" pageId="18" pageNumber="258">Fig. 1A</figureCitation>
), without visible pubescence; penis wider (
<figureCitation box="[1249,1326,1044,1066]" captionStart="FIGURE 4" captionStartId="5.[151,250,1880,1904]" captionTargetBox="[173,1423,186,1856]" captionTargetId="figure-17@5.[164,1423,181,1856]" captionTargetPageId="5" captionText="FIGURE 4. Aedeagus of Lucanus species (H, L, P, T, apex of flagellum). A, E–H, L. jiaozishanus Qi, He, Su &amp; Song, new species (holotype, Kunming, Yunnan); B, I–L, L. ludivinae Boucher, 1998 (Gongshan, Yunnan); C, M–P, L. takeoi Adachi, 2020 (Huidong, Sichuan); D, Q–T, L. boileaui Planet, 1897 (Mabian, Sichuan). F, J, N, R, dorsal view; A–D, E, I, M, Q, ventral view; G, K, O, S, lateral view. Scale bar for H, L, P, T = 0.2 mm, for the others = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7801114" httpUri="https://zenodo.org/record/7801114/files/figure.png" pageId="18" pageNumber="258">Fig. 4E</figureCitation>
); flagellum (
<figureCitation box="[233,308,1074,1096]" captionStart="FIGURE 4" captionStartId="5.[151,250,1880,1904]" captionTargetBox="[173,1423,186,1856]" captionTargetId="figure-17@5.[164,1423,181,1856]" captionTargetPageId="5" captionText="FIGURE 4. Aedeagus of Lucanus species (H, L, P, T, apex of flagellum). A, E–H, L. jiaozishanus Qi, He, Su &amp; Song, new species (holotype, Kunming, Yunnan); B, I–L, L. ludivinae Boucher, 1998 (Gongshan, Yunnan); C, M–P, L. takeoi Adachi, 2020 (Huidong, Sichuan); D, Q–T, L. boileaui Planet, 1897 (Mabian, Sichuan). F, J, N, R, dorsal view; A–D, E, I, M, Q, ventral view; G, K, O, S, lateral view. Scale bar for H, L, P, T = 0.2 mm, for the others = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7801114" httpUri="https://zenodo.org/record/7801114/files/figure.png" pageId="18" pageNumber="258">Fig. 4A</figureCitation>
) about 2.0 times longer than aedeagus....................
<taxonomicName authority="Qi, He, Su" authorityName="Qi, He, Su" box="[912,1226,1074,1096]" class="Insecta" family="Lucanidae" genus="Lucanus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="258" phylum="Arthropoda" rank="species" species="jiaozishanus">
<emphasis box="[912,1119,1074,1095]" italics="true" pageId="18" pageNumber="258">Lucanus jiaozishanus</emphasis>
Qi, He, Su
</taxonomicName>
&amp; Song,
<emphasis bold="true" box="[1318,1437,1074,1096]" pageId="18" pageNumber="258">new species</emphasis>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>
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