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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1179.107587" ID-Pensoft-Pub="1313-2970-1179-35" ID-Pensoft-UUID="07CEAFC9849852A1AC00767783926142" ID-ZooBank="D9BD50CBB127487981E99421E4F059BA" ModsDocID="1313-2970-1179-35" checkinTime="1694247957851" checkinUser="pensoft" docAuthor="Gao, Biao, Han, Hui-Lin, Kononenko, Vladimir S. &amp; Pan, Zhao-Hui" docDate="2023" docId="7324CE36255A563DA0EA69CB01CEF871" docLanguage="en" docName="ZooKeys 1179: 35-61" docOrigin="ZooKeys 1179" docPubDate="2023-09-08" docSource="http://dx.doi.org/10.3897/zookeys.1179.107587" docTitle="Hermonassa kalamantra Kovacs, G. Ronkay &amp; L. Ronkay 2018" docType="treatment" docVersion="1" id="07CEAFC9849852A1AC00767783926142" lastPageNumber="35" masterDocId="07CEAFC9849852A1AC00767783926142" masterDocTitle="Five new species of the genus Hermonassa Walker, 1865 from Xizang Autonomous Region, China (Lepidoptera, Noctuidae, Noctuinae)" masterLastPageNumber="61" masterPageNumber="35" pageNumber="35" updateTime="1694247957851" updateUser="pensoft">
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<mods:title id="4F439AB89DF53CEB50E02DD266E45514">Five new species of the genus Hermonassa Walker, 1865 from Xizang Autonomous Region, China (Lepidoptera, Noctuidae, Noctuinae)</mods:title>
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<mods:namePart id="06C6E352CD30D376CDDE5F5F53716C33">Gao, Biao</mods:namePart>
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<mods:namePart id="943846C5C4F8A667793309E39516175A">Han, Hui-Lin</mods:namePart>
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<mods:affiliation id="3CF7CCF3CD876B630143F0ACA7240D56">Northeast Forestry University, School of Forestry, Harbin 150040, China</mods:affiliation>
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<mods:namePart id="DCBD9BA06E36A0017A94D24DD1CF951B">Kononenko, Vladimir S.</mods:namePart>
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<mods:affiliation id="AEE46DE6F1885D25A8789988C1869B4D">Northeast Asia Biodiversity Research Center, Northeast Forestry University, Harbin 15004, China</mods:affiliation>
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<mods:namePart id="319C416C1A10AEEE5C0DE3D1EB041BFB">Pan, Zhao-Hui</mods:namePart>
<mods:affiliation id="79AB8A5B1448B07E701570157CD3D18C">Northeast Forestry University, Ministry of Education, Key Laboratory of Sustainable Forest Ecosystem Management, Harbin 150040, China</mods:affiliation>
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<subSubSection id="A57738FF789C9AFA8AA94ABAB8596277" pageId="0" pageNumber="35" type="nomenclature">
<paragraph id="37285F344871DF298AE744DC0A97DFEF" pageId="0" pageNumber="35">
<taxonomicName id="1CC45B1FAC0DD8B80DEAD7259B1B9CF4" LSID="7324CE36-255A-563D-A0EA-69CB01CEF871" authority="Kovacs, G. Ronkay &amp; L. Ronkay, 2018" authorityName="Kovacs, G. Ronkay &amp; L. Ronkay" authorityYear="2018" class="Insecta" family="Noctuidae" genus="Hermonassa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hermonassa kalamantra" order="Lepidoptera" pageId="0" pageNumber="35" phylum="Arthropoda" rank="species" species="kalamantra">
Hermonassa kalamantra
<normalizedToken id="A5A4D9BBDE4884B5D03D42F8ED970AA3" originalValue="Kovács">Kovacs</normalizedToken>
, G. Ronkay &amp; L. Ronkay, 2018
</taxonomicName>
</paragraph>
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<subSubSection id="82AEBEA52FD4E52859BE41F9AE7FAE3E" pageId="0" pageNumber="35" type="description">
<paragraph id="44F2719C59B77327E09894DDABE9D06F" pageId="0" pageNumber="35">
<figureCitation id="656B0C6A0A0F9AA789AD55E9870DBA21" captionStart="Figures 1–10" captionStartId="F1" captionText="Figures 1 - 10. Hermonassa spp., adults 1 H. nigricans sp. nov., ♂, holotype, China 2 Ditto, ♀, paratype 3 H. anthracina, ♂, holotype, China (ZFMK) 4 H. anthracina, ♀, China 5 H. kalamantra, ♂, holotype, Nepal (after Kovacs et al. 2018) 6 H. kalamantra, ♂, China 7 H. conusa sp. nov., ♂, holotype, China 8 Ditto, ♀, paratype, China 9 H. renifera, ♂, holotype, China (NACR) 10 H. shizukoae, ♂, holotype, India (NIAES)." figureDoi="10.3897/zookeys.1179.107587.figures1-10" httpUri="https://binary.pensoft.net/fig/903263" pageId="0" pageNumber="35">Figs 5</figureCitation>
<figureCitation id="338D95C0EE48847F05B5C89308F4038F" captionStart="Figures 1–10" captionStartId="F1" captionText="Figures 1 - 10. Hermonassa spp., adults 1 H. nigricans sp. nov., ♂, holotype, China 2 Ditto, ♀, paratype 3 H. anthracina, ♂, holotype, China (ZFMK) 4 H. anthracina, ♀, China 5 H. kalamantra, ♂, holotype, Nepal (after Kovacs et al. 2018) 6 H. kalamantra, ♂, China 7 H. conusa sp. nov., ♂, holotype, China 8 Ditto, ♀, paratype, China 9 H. renifera, ♂, holotype, China (NACR) 10 H. shizukoae, ♂, holotype, India (NIAES)." figureDoi="10.3897/zookeys.1179.107587.figures1-10" httpUri="https://binary.pensoft.net/fig/903263" pageId="0" pageNumber="35">, 6</figureCitation>
<figureCitation id="8778EDC1D5391F6B048B42E6008C0ADF" captionStart="Figures 11–18" captionStartId="F2" captionText="Figures 11 - 18. Hermonassa spp., male genitalia 11 H. nigricans sp. nov., holotype, China, genit. prep. GB- 101 - 1 12 H. nigricans sp. nov., Nepal (referred as H. anthracina, after Sugi, 1995) 13 H. anthracina, paratype, China (ZFMK); 14 H. anthracina, China (after Kovacs et al. 2018) 15 H. anthracina, China, genit. prep. hhl- 3490 - 1 16 H. kalamantra, paratype, Nepal (after Kovacs et al. 2018) 17 H. kalamantra, China, genit. prep. hhl- 3489 - 1 18 H. kalamantra, China, genit. prep. hhl- 3488 - 1." figureDoi="10.3897/zookeys.1179.107587.figures11-18" httpUri="https://binary.pensoft.net/fig/903264" pageId="0" pageNumber="35">, 16-18</figureCitation>
<figureCitation id="31E0B933BEECACB859DDA259DD6C5B12" captionStart="Figures 19–27" captionStartId="F3" captionText="Figures 19 - 27. Hermonassa spp., male (19 - 22) and female (23 - 27) genitalia 19 H. conusa sp. nov., holotype, genit. prep. hhl- 3492 - 1 20 H. renifera, paratype (after Chen 1991) 21 H. shizukoae, holotype, India (after Sugi 1995) 22 H. shizukoae, Nepal (after Kovacs et al. 2018) 23 H. nigricans, paratype, genit. prep. hhl- 3493 - 2 24 H. anthracina, China, (after Kovacs et al. 2018) 25 H. kalamantra, Nepal (after Kovacs et al. 2018) 26 H. conusa sp. nov., paratype, China, genit. prep. hhl- 3491 - 2 27 H. shizukoae, paratype, India (after Sugi 1995)." figureDoi="10.3897/zookeys.1179.107587.figures19-27" httpUri="https://binary.pensoft.net/fig/903265" pageId="0" pageNumber="35">, 25</figureCitation>
</paragraph>
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<subSubSection id="F132B2745F4F78C302D805C1EE2911BB" pageId="0" pageNumber="35" type="reference_group">
<paragraph id="EEA6B4507A57A374679ABF8B20E3E8A5" pageId="0" pageNumber="35">
<taxonomicName id="6ECE6DAF8227FDC2830BDA3157C2247C" authorityName="Kovacs, G. Ronkay &amp; L. Ronkay" authorityYear="2018" class="Insecta" family="Noctuidae" genus="Hermonassa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hermonassa kalamantra" order="Lepidoptera" pageId="0" pageNumber="35" phylum="Arthropoda" rank="species" species="kalamantra">Hermonassa kalamantra</taxonomicName>
<normalizedToken id="5350888EF93F31718F6C44D96A99566C" originalValue="Kovács">Kovacs</normalizedToken>
, G. Ronkay &amp; L. Ronkay, 2018, Revue suisse de Zoologie 126 (2): 298, pl. 1, figs 5-8, genitalia figs 3-5.
</paragraph>
</subSubSection>
<subSubSection id="C10F80A859D815411E3728EC16B8E5EF" pageId="0" pageNumber="35" type="type material">
<paragraph id="EF71198ECC48D19EAD1AD40F89B65EA4" pageId="0" pageNumber="35">Type material.</paragraph>
<paragraph id="E3AF6270FF1544552F18A68EF0477104" pageId="0" pageNumber="35">
<emphasis id="63A4FFBF748D602AD0AB97ED47D9606F" bold="true" italics="true" pageId="0" pageNumber="35">Holotype</emphasis>
: ♂, Nepal, Annapurna Himal, Mesokantu pass, 4200 m. Deposited in the collection of HNHM, Budapest, Hungary (not examined).
</paragraph>
</subSubSection>
<subSubSection id="16656815914A79F40159BDB6A44C9C5C" pageId="0" pageNumber="35" type="materials_examined">
<paragraph id="6598C374C87EDEDF09057C15CFA3E187" pageId="0" pageNumber="35">Other material examined.</paragraph>
<paragraph id="3F9F4873A0D62A38DBC6347F1B4BA572" pageId="0" pageNumber="35">
<materialsCitation id="70050C3275183AAEBE8DC941B1CDDE22" collectingDate="2015-07-28" collectorName="H. L. Han" country="China" elevation="3300" location="Mt. Sejila" municipality="Linzhi City" specimenCount="2" specimenCount-male="2" stateProvince="Xizang">
<collectingCountry id="C48028EA47ECA563A37DF4D823CE0D00" name="China">China</collectingCountry>
,
<specimenCount id="4A048D1570D59C6BCD2709B401C1A82F" type="male">2 ♂</specimenCount>
, Aut. Reg.
<collectingRegion id="354B220BA2123BCB625D5FCF9C322A29" country="China" name="Xizang">Xizang</collectingRegion>
,
<collectingMunicipality id="B3B17C63895BDAB76977A2F59CE5F11F">Linzhi City</collectingMunicipality>
,
<location id="A948B5ADB7ACC1ABB6C0114BBE5F93E1" LSID="urn:lsid:plazi:treatment:7324CE36255A563DA0EA69CB01CEF871:A948B5ADB7ACC1ABB6C0114BBE5F93E1" country="China" municipality="Linzhi City" name="Mt. Sejila" stateProvince="Xizang">Mt. Sejila</location>
,
<quantity id="90A2572A84A865A3B0FA0547E8D086C6" metricMagnitude="3" metricUnit="m" metricValue="3.3" unit="m" value="3300.0">
<elevation id="EA0AEE368D8A9FD59FD5530A51FEE439" metricMagnitude="3" metricUnit="m" metricValue="3.3" unit="m" value="3300.0">3300 m</elevation>
</quantity>
,
<collectingDate id="40B41225EE5D147905CC1E42980309DE" value="2015-07-28">28 July 2015</collectingDate>
,
<collectorName id="C22B64DC1248B6FDBBE83293DB6A7316">H.L. Han</collectorName>
leg., genit. prep. hhl-3488-1, hhl-3489-1 (coll. NEFU)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="83E431618CE413D315F9AA51FDD9F852" pageId="0" pageNumber="35" type="distribution">
<paragraph id="1A50B1888CB75FECF88952C4FB261869" pageId="0" pageNumber="35">Distribution and biology.</paragraph>
<paragraph id="6EC73273203B43BA47308EF5EEEC2DE4" pageId="0" pageNumber="35">
<taxonomicName id="57A6C8C6F1B818F57BF393B4A9A7D7DD" authorityName="Kovacs, G. Ronkay &amp; L. Ronkay" authorityYear="2018" class="Insecta" family="Noctuidae" genus="Hermonassa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hermonassa kalamantra" order="Lepidoptera" pageId="0" pageNumber="35" phylum="Arthropoda" rank="species" species="kalamantra">
<emphasis id="1D568DB60A948F353116CBAFDF1B01A4" italics="true" pageId="0" pageNumber="35">Hermonassa kalamantra</emphasis>
</taxonomicName>
occurs in Nepal and the southwest China (Aut. Reg. Xizang). Judging from the collecting data in Nepal it is a common species at altitudes of 2000-4000 m; adults flying from May to September (
<bibRefCitation id="6E63A9715264E19E4C55595825F5CE31" author="Kovacs, ST" journalOrPublisher="Revue Suisse de Zoologie" pageId="0" pageNumber="35" pagination="297 - 319" refId="B16" refString="Kovacs, ST, Ronkay, G, Ronkay, L, 2018. Revision of the genus Hermonassa Walker, 1865 (Lepidoptera, Noctuidae, Noctuinae). I. The anthracina and dispila species groups, with descriptions of four new species. Revue Suisse de Zoologie 126 (2): 297 - 319" title="Revision of the genus Hermonassa Walker, 1865 (Lepidoptera, Noctuidae, Noctuinae). I. The anthracina and dispila species groups, with descriptions of four new species." volume="126" year="2018">
<normalizedToken id="30108C2738263FF7DE8F8FA8EF1A7BF3" originalValue="Kovács">Kovacs</normalizedToken>
et al. 2018
</bibRefCitation>
). In Aut. Reg. Xizang it is known from only two specimens and appears rare compared with
<taxonomicName id="F56525713B339FB424B107916D74396B" family="Noctuidae" kingdom="Animalia" lsidName="H. anthracina" order="Lepidoptera" pageId="0" pageNumber="35" rank="species" species="anthracina">
<emphasis id="501B46A3D46A2EF0FE436B22D96F4202" italics="true" pageId="0" pageNumber="35">H. anthracina</emphasis>
</taxonomicName>
and
<taxonomicName id="2DEAEED6998D0B46664C9531215CAC45" family="Noctuidae" kingdom="Animalia" lsidName="H. nigricans" order="Lepidoptera" pageId="0" pageNumber="35" rank="species" species="nigricans">
<emphasis id="8213B7D55E6D0AB4021EF0B6093D70FC" italics="true" pageId="0" pageNumber="35">H. nigricans</emphasis>
</taxonomicName>
sp. nov. adults have been collected in late July.
</paragraph>
<caption id="4350B827A73CEDC5C0913F5CB6002740" doi="10.3897/zookeys.1179.107587.figures11-18" httpUri="https://binary.pensoft.net/fig/903264" pageId="0" pageNumber="35" start="Figures 11–18" startId="F2">
<paragraph id="032CF0787616D1BE9719F7D1D4AFBB15" pageId="0" pageNumber="35">
<emphasis id="467F1F0A7B29F31946AB90343A03C024" bold="true" pageId="0" pageNumber="35">Figures 11-18.</emphasis>
<taxonomicName id="26A350AE3802083567A3670C5D0600EB" authorityName="Walker" authorityYear="1865" class="Insecta" family="Noctuidae" genus="Hermonassa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hermonassa" order="Lepidoptera" pageId="0" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="53F2AA98A4AEA50B127D0011831ED537" italics="true" pageId="0" pageNumber="35">Hermonassa</emphasis>
</taxonomicName>
spp., male genitalia
<emphasis id="83F0127C963E8D700C6671B40D40F8CB" bold="true" pageId="0" pageNumber="35">11</emphasis>
<taxonomicName id="2F395A796D8038106E7E19277C2C64B0" family="Noctuidae" kingdom="Animalia" lsidName="H. nigricans" order="Lepidoptera" pageId="0" pageNumber="35" rank="species" species="nigricans">
<emphasis id="A87480FED0170FE9AA5F33836C2DD3FF" italics="true" pageId="0" pageNumber="35">H. nigricans</emphasis>
</taxonomicName>
sp. nov., holotype, China, genit. prep. GB-101-1
<emphasis id="692BD366A3311DC69B846CCCB03E24A4" bold="true" pageId="0" pageNumber="35">12</emphasis>
<taxonomicName id="D6426E6CDE827DAF5CEBD887CD82DC2E" family="Noctuidae" kingdom="Animalia" lsidName="H. nigricans" order="Lepidoptera" pageId="0" pageNumber="35" rank="species" species="nigricans">
<emphasis id="F604988A7FFCD136F13A1F0044BECA6F" italics="true" pageId="0" pageNumber="35">H. nigricans</emphasis>
</taxonomicName>
sp. nov., Nepal (referred as
<taxonomicName id="A03F525F6B9136C404B2C6D466B276EC" family="Noctuidae" kingdom="Animalia" lsidName="H. anthracina" order="Lepidoptera" pageId="0" pageNumber="35" rank="species" species="anthracina">
<emphasis id="064E5612696AC2A03FB1E2814F1AA149" italics="true" pageId="0" pageNumber="35">H. anthracina</emphasis>
</taxonomicName>
, after Sugi, 1995)
<emphasis id="AE6EA79421AC4C7DB87BAA92536275BC" bold="true" pageId="0" pageNumber="35">13</emphasis>
<taxonomicName id="F2B9EE0B5773807C93535991DA1129E2" family="Noctuidae" kingdom="Animalia" lsidName="H. anthracina" order="Lepidoptera" pageId="0" pageNumber="35" rank="species" species="anthracina">
<emphasis id="791A741FBDA2C011C465F2610CDD4BBB" italics="true" pageId="0" pageNumber="35">H. anthracina</emphasis>
</taxonomicName>
, paratype, China (ZFMK);
<emphasis id="0C5B4353483D0ECCA1EA0E5B6CBCBBDC" bold="true" pageId="0" pageNumber="35">14</emphasis>
<taxonomicName id="9B0EE137D2232CE7215FD369F8B1A2D0" family="Noctuidae" kingdom="Animalia" lsidName="H. anthracina" order="Lepidoptera" pageId="0" pageNumber="35" rank="species" species="anthracina">
<emphasis id="A2B7EDBE92C108CD39F14A890B6A72C6" italics="true" pageId="0" pageNumber="35">H. anthracina</emphasis>
</taxonomicName>
, China (after
<bibRefCitation id="F48F37C9D50EDE211D7252E08143EDF1" author="Kovacs, ST" journalOrPublisher="Revue Suisse de Zoologie" pageId="0" pageNumber="35" pagination="297 - 319" refId="B16" refString="Kovacs, ST, Ronkay, G, Ronkay, L, 2018. Revision of the genus Hermonassa Walker, 1865 (Lepidoptera, Noctuidae, Noctuinae). I. The anthracina and dispila species groups, with descriptions of four new species. Revue Suisse de Zoologie 126 (2): 297 - 319" title="Revision of the genus Hermonassa Walker, 1865 (Lepidoptera, Noctuidae, Noctuinae). I. The anthracina and dispila species groups, with descriptions of four new species." volume="126" year="2018">
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et al. 2018
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)
<emphasis id="39AEE5161F3A18699D6E2806C91C521E" bold="true" pageId="0" pageNumber="35">15</emphasis>
<taxonomicName id="8BFA4A7675B5A7D6674CB51637499455" family="Noctuidae" kingdom="Animalia" lsidName="H. anthracina" order="Lepidoptera" pageId="0" pageNumber="35" rank="species" species="anthracina">
<emphasis id="E0CB4240591C39789A31451358B31547" italics="true" pageId="0" pageNumber="35">H. anthracina</emphasis>
</taxonomicName>
, China, genit. prep. hhl-3490-1
<emphasis id="1A6A580C1FE031B64A216A89D9109F20" bold="true" pageId="0" pageNumber="35">16</emphasis>
<taxonomicName id="A427AA8233DB50881EAA0C6ABCC579C9" family="Noctuidae" kingdom="Animalia" lsidName="H. kalamantra" order="Lepidoptera" pageId="0" pageNumber="35" rank="species" species="kalamantra">
<emphasis id="D38A20A19413CA27F36A719B9C1303ED" italics="true" pageId="0" pageNumber="35">H. kalamantra</emphasis>
</taxonomicName>
, paratype, Nepal (after
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et al. 2018
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)
<emphasis id="4E54F1C7D8CEB80C60116A40DBB372CA" bold="true" pageId="0" pageNumber="35">17</emphasis>
<taxonomicName id="F35924BFEB3DC2ABE944F3B1718E2305" family="Noctuidae" kingdom="Animalia" lsidName="H. kalamantra" order="Lepidoptera" pageId="0" pageNumber="35" rank="species" species="kalamantra">
<emphasis id="8A207DE5CA2C24D0CF462BFE16580A06" italics="true" pageId="0" pageNumber="35">H. kalamantra</emphasis>
</taxonomicName>
, China, genit. prep. hhl-3489-1
<emphasis id="D0A4E14A4D3808FE871CBBFB7BC15898" bold="true" pageId="0" pageNumber="35">18</emphasis>
<taxonomicName id="86712C365ED735116D16ABD26E4461CA" family="Noctuidae" kingdom="Animalia" lsidName="H. kalamantra" order="Lepidoptera" pageId="0" pageNumber="35" rank="species" species="kalamantra">
<emphasis id="4CB53A1912215FEEECDE6B7D2F089E4B" italics="true" pageId="0" pageNumber="35">H. kalamantra</emphasis>
</taxonomicName>
, China, genit. prep. hhl-3488-1.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="7DB87A292460DCF96B6EF10BF9D1B1C5" pageId="0" pageNumber="35" type="remarks">
<paragraph id="D0ACDA6B111072790B593C91E46F4905" pageId="0" pageNumber="35">Remark.</paragraph>
<paragraph id="399583689FBC6AC8D8ED123C0BF36EA0" pageId="0" pageNumber="35">The species is reported for China for the first time.</paragraph>
</subSubSection>
<subSubSection id="F0E03916B176A121F87BF088230D2111" pageId="0" pageNumber="35" type="the dispila species group">
<paragraph id="19FD0BD2A052E03052AF3711B2756DD0" pageId="0" pageNumber="35">
The
<emphasis id="8AE9E306477C2BF0CDA01090D0CFE692" italics="true" pageId="0" pageNumber="35">dispila</emphasis>
species group
</paragraph>
<paragraph id="4096CB900D1B9B3C3BE05BCC62F45933" pageId="0" pageNumber="35">
The members of the
<emphasis id="CA4F92BD340E6D1C3133BB66F7CDD204" italics="true" pageId="0" pageNumber="35">dispila</emphasis>
species group are externally similar to taxa of the
<taxonomicName id="C60CCB2A46EF5128E8FADDC83C9D4D0D" authorityName="Boursin" authorityYear="1967" class="Insecta" family="Noctuidae" genus="Hermonassa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hermonassa anthracina" order="Lepidoptera" pageId="0" pageNumber="35" phylum="Arthropoda" rank="species" species="anthracina">
<emphasis id="AEAD256C8AAC18BC0F25EB253D6F7390" italics="true" pageId="0" pageNumber="35">Hermonassa anthracina</emphasis>
</taxonomicName>
group, while some species have narrower forewing and different wing colouration. As mentioned by
<bibRefCitation id="D2B2AFB4E41DBE4921D1DAFB8999E874" author="Kovacs, ST" journalOrPublisher="Revue Suisse de Zoologie" pageId="0" pageNumber="35" pagination="297 - 319" refId="B16" refString="Kovacs, ST, Ronkay, G, Ronkay, L, 2018. Revision of the genus Hermonassa Walker, 1865 (Lepidoptera, Noctuidae, Noctuinae). I. The anthracina and dispila species groups, with descriptions of four new species. Revue Suisse de Zoologie 126 (2): 297 - 319" title="Revision of the genus Hermonassa Walker, 1865 (Lepidoptera, Noctuidae, Noctuinae). I. The anthracina and dispila species groups, with descriptions of four new species." volume="126" year="2018">
<normalizedToken id="15993EE9491FC6B3A19B72A6863F9EC4" originalValue="Kovács">Kovacs</normalizedToken>
et al. (2018
</bibRefCitation>
: 301) they are morphologically more heterogeneous and diverse: &quot;The genitalia of these species show different trends of change in certain features which led to the often remarkable differences between the male genital capsule of the different lineages of the clade. Due to the basically uniform external appearance and the genital features of the entire species group, as well as partly overlapping differential characters, the taxa of the species group can be arranged into different lineages.&quot; Those authors listed 12 species in the
<emphasis id="9CA78CEC655FE1AC6519EA7B96971EFC" italics="true" pageId="0" pageNumber="35">dispila</emphasis>
species group; however, the exact number of species and selection of the main lineages of this group are likely will be clarified in the course of further revisions.
</paragraph>
</subSubSection>
</treatment>
</document>
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