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<document id="5BF38D5339F7BC44FB65FDE599335943" ID-DOI="10.11646/zootaxa.5372.1.1" ID-ISSN="1175-5334" ID-Zenodo-Dep="10146342" ID-ZooBank="1300D562-3E0B-4F72-933F-8FA7D28F6853" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1700186674367" checkinUser="plazi" docAuthor="Mendonça, Maria Thayane Da Silva, Silva, Valéria Juliete Da &amp; Fernandes, José Antônio Marin" docDate="2023" docId="414B87B5FFDA9841FF63DB1D77FDFC9C" docLanguage="en" docName="zootaxa.5372.1.1.pdf" docOrigin="Zootaxa 5372 (1)" docSource="https://www.mapress.com/zt/article/download/zootaxa.5372.1.1/52272" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Edessa (Edessa) accola Mendonça &amp; Silva &amp; Fernandes 2023, sp. n." docType="treatment" docVersion="2" lastPageNumber="12" masterDocId="BD72FFCDFFD3984AFFF4DE477513FFC1" masterDocTitle="Description of fifty-one new species and new taxonomic arrangement for the E. sexdens group of the subgenus Edessa (Heteroptera, Pentatomidae, Edessinae, Edessa)" masterLastPageNumber="128" masterPageNumber="1" pageNumber="10" updateTime="1700188109802" updateUser="ExternalLinkService">
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<mods:title id="EFFA01BED0DBE1027BFBE233C518B842">Description of fifty-one new species and new taxonomic arrangement for the E. sexdens group of the subgenus Edessa (Heteroptera, Pentatomidae, Edessinae, Edessa)</mods:title>
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<mods:namePart id="A05A847E7A76561F7396429132B8D1F3">Mendonça, Maria Thayane Da Silva</mods:namePart>
<mods:affiliation id="B61F01F82855158515F57D47D23AE0B9">Universidade Federal do Pará, Instituto de Ciências Biológicas, Av. Augusto Correa # 1 66075 - 110, Belém, Pará, Brazil &amp; Programa de Pós-Graduação em Zoologia UFPA / MPEG</mods:affiliation>
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<mods:namePart id="5DBCEDC04E6FB6ACA5422AB71472E1F3">Silva, Valéria Juliete Da</mods:namePart>
<mods:nameIdentifier id="4F14EBA563B0C5F2F947AFE9BBBEBB72" type="ORCID">0000-0002-1845-5228</mods:nameIdentifier>
<mods:affiliation id="EA6ADAD0BB4437C3A1881D5310005196">Universidade Federal do Pará, Instituto de Ciências Biológicas, Av. Augusto Correa # 1 66075 - 110, Belém, Pará, Brazil &amp; valeriajuliete @ hotmail. com; https: // orcid. org / 0000 - 0002 - 1845 - 5228</mods:affiliation>
<mods:nameIdentifier id="5D21697871FAADDE1AF02E76BCB1D702" type="email">valeriajuliete@hotmail.com</mods:nameIdentifier>
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<mods:namePart id="90235F8A51FA8E1DAC117084E92EF753">Fernandes, José Antônio Marin</mods:namePart>
<mods:nameIdentifier id="DFFB5C9CDF688ACF58FFEE547657036F" type="ORCID">0000-0001-7450-5296</mods:nameIdentifier>
<mods:affiliation id="261124F9F94E3337689586ADB29DD161">Universidade Federal do Pará, Instituto de Ciências Biológicas, Av. Augusto Correa # 1 66075 - 110, Belém, Pará, Brazil &amp; Universidade Federal do Pará, Instituto de Ciências Biológicas, Av. Augusto Correa # 1 66075 - 110, Belém, Pará, Brazil &amp; Corresponding author: joseamf @ ufpa. br; https: // orcid. org / 0000 - 0001 - 7450 - 5296</mods:affiliation>
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<mods:title id="28F1729753988D3A3FD1FB2926ED5E69">Zootaxa</mods:title>
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<mods:date id="3C9FE6CF3729E9471DDE19D8F43ED5BE">2023</mods:date>
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<mods:number id="D2D67D8B4F184E4B84CB05F7D661A3A2">2023-11-14</mods:number>
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<mods:number id="D280CA7F431CE67D1AB76102FE1BFCE5">5372</mods:number>
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<mods:number id="E588A6EA6C9772CB1118098FBA633DC3">1</mods:number>
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<treatment id="414B87B5FFDA9841FF63DB1D77FDFC9C" LSID="urn:lsid:plazi:treatment:414B87B5FFDA9841FF63DB1D77FDFC9C" httpUri="http://treatment.plazi.org/id/414B87B5FFDA9841FF63DB1D77FDFC9C" lastPageId="11" lastPageNumber="12" pageId="9" pageNumber="10">
<subSubSection id="81F86528FFDA9843FF63DB1D74E8FAB4" box="[151,507,1370,1397]" pageId="9" pageNumber="10" type="nomenclature">
<paragraph id="C95D36A3FFDA9843FF63DB1D74E8FAB4" blockId="9.[151,507,1370,1434]" box="[151,507,1370,1397]" pageId="9" pageNumber="10">
<heading id="921581CFFFDA9843FF63DB1D74E8FAB4" bold="true" box="[151,507,1370,1397]" fontSize="11" level="2" pageId="9" pageNumber="10" reason="4">
<emphasis id="FB96EAB1FFDA9843FF63DB1D74E8FAB4" bold="true" box="[151,507,1370,1397]" pageId="9" pageNumber="10">
<taxonomicName id="0EE24D20FFDA9843FF63DB1D74A2FAB4" authority="Mendonça &amp; Silva &amp; Fernandes, 2023" authorityName="Mendonça &amp; Silva &amp; Fernandes" authorityYear="2023" box="[151,433,1370,1397]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="accola" status="sp. nov." subGenus="Edessa">
<emphasis id="FB96EAB1FFDA9843FF63DB1D74A2FAB4" bold="true" box="[151,433,1370,1397]" italics="true" pageId="9" pageNumber="10">Edessa (Edessa) accola</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E0A557CAFFDA9843FE4CDB1C74E8FAB4" box="[440,507,1371,1397]" pageId="9" pageNumber="10" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="81F86528FFDA9843FF63DB3874B2FA58" box="[151,417,1407,1434]" pageId="9" pageNumber="10" type="description">
<paragraph id="C95D36A3FFDA9843FF63DB3874B2FA58" blockId="9.[151,507,1370,1434]" box="[151,417,1407,1434]" pageId="9" pageNumber="10">
(
<figureCitation id="51D92A26FFDA9843FF6BDB387419FA58" box="[159,266,1407,1433]" captionStart="FIGURE 1" captionStartId="10.[152,255,1943,1967]" captionTargetBox="[185,1408,189,1913]" captionTargetId="figure-22@10.[179,1408,181,1919]" captionTargetPageId="10" captionText="FIGURE 1. AF. Edessa (E.) accola sp. n.: AE, pygophore in dorsal, dorsolateral, ventral, posterior and posterolateral views, respectively; F, metasternal process. dr—dorsal rim; vr—ventral rim; proc—proctiger; pa—paramere; gp—superior processes of the genital cup. Scale= 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146345" httpUri="https://zenodo.org/record/10146345/files/figure.png" pageId="9" pageNumber="10">Figures 1</figureCitation>
,
<figureCitation id="51D92A26FFDA9843FEECDB387463FA58" box="[280,368,1407,1434]" captionStart="FIGURE 54" captionStartId="107.[152,255,1943,1967]" captionTargetBox="[250,1338,187,1913]" captionTargetId="figure-20@107.[244,1344,181,1919]" captionTargetPageId="107" captionText="FIGURE 54. AF. Dorsal and ventral surface: AB, Edessa (E.) accola sp. n.; CD, Edessa (E.) amapaensis sp. n.—yellow lines indicate where the length (L) and width (W) measurements of the humeral angles were taken; EF, Edessa (E.) amaurata sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146457" httpUri="https://zenodo.org/record/10146457/files/figure.png" pageId="9" pageNumber="10">54 AB</figureCitation>
,
<figureCitation id="51D92A26FFDA9843FE88DB38748AFA58" box="[380,409,1407,1433]" captionStart="FIGURE 72" captionStartId="125.[152,255,1071,1095]" captionTargetBox="[195,1391,181,1040]" captionTargetId="figure-20@125.[195,1391,181,1050]" captionTargetPageId="125" captionText="FIGURE 72. Distribution map of the species." figureDoi="http://doi.org/10.5281/zenodo.10146495" httpUri="https://zenodo.org/record/10146495/files/figure.png" pageId="9" pageNumber="10">72</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="81F86528FFDA9843FF63DB8071E3FA20" box="[151,1264,1479,1506]" pageId="9" pageNumber="10" type="etymology">
<paragraph id="C95D36A3FFDA9843FF63DB8071E3FA20" blockId="9.[151,1437,1479,2017]" box="[151,1264,1479,1506]" pageId="9" pageNumber="10">
<emphasis id="FB96EAB1FFDA9843FF63DB80740BFA20" bold="true" box="[151,280,1479,1505]" pageId="9" pageNumber="10">Etymology</emphasis>
. This name refers to the similarity with
<taxonomicName id="0EE24D20FFDA9843FD2FDB8F76E0FA20" authority="Stal, 1872" authorityName="Stal" authorityYear="1872" box="[731,1011,1479,1506]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="fallenii" subGenus="Edessa">
<emphasis id="FB96EAB1FFDA9843FD2FDB8F7669FA20" box="[731,890,1479,1506]" italics="true" pageId="9" pageNumber="10">E. (E.) fallenii</emphasis>
<bibRefCitation id="AD734B52FFDA9843FC76DB8076E0FA20" author="Stal, C." box="[898,1011,1479,1506]" pageId="9" pageNumber="10" pagination="1 - 159" refId="ref75136" refString="Stal, C. (1872) Enumeratio Hemipterorum. Bildrag till em f ˆ rteckning ˆ fver alla hittilis kanda Hemiptera, Jemte systematiska meddelanden. 2. Kongliga Svenska Vetenskaps - Akademiens Handlingar, Neue Folge, 10 (4), 1 - 159. https: // doi. org / 10.5962 / bhl. title. 12549" type="journal article" year="1872">Stål, 1872</bibRefCitation>
</taxonomicName>
(L.
<taxonomicName id="0EE24D20FFDA9843FBD7DB80717DFA20" authorityName="Mendonça &amp; Silva &amp; Fernandes" authorityYear="2023" box="[1059,1134,1479,1505]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="accola">
<emphasis id="FB96EAB1FFDA9843FBD7DB80717DFA20" box="[1059,1134,1479,1505]" italics="true" pageId="9" pageNumber="10">accola</emphasis>
</taxonomicName>
, neighbor).
</paragraph>
</subSubSection>
<subSubSection id="81F86528FFDA9843FF33DBAB71AAF9C7" box="[199,1209,1516,1542]" pageId="9" pageNumber="10" type="materials_examined">
<paragraph id="C95D36A3FFDA9843FF33DBAB71AAF9C7" blockId="9.[151,1437,1479,2017]" box="[199,1209,1516,1542]" pageId="9" pageNumber="10">
<emphasis id="FB96EAB1FFDA9843FF33DBAB74A2F9C7" bold="true" box="[199,433,1516,1542]" pageId="9" pageNumber="10">Material examined.</emphasis>
<materialsCitation id="798A3CFEFFDA9843FE43DBAB71AAF9C7" box="[439,1209,1516,1542]" collectionCode="CMNH" country="Brazil" location="Rio de Janeiro" pageId="9" pageNumber="10" specimenCount="1" stateProvince="Rio de Janeiro" typeStatus="holotype">
<typeStatus id="16598801FFDA9843FE43DBAB770CF9C7" box="[439,543,1516,1542]" pageId="9" pageNumber="10" type="holotype">Holotype</typeStatus>
male.
<emphasis id="FB96EAB1FFDA9843FD9FDBAB7688F9C7" bold="true" box="[619,923,1516,1542]" pageId="9" pageNumber="10">
<collectingCountry id="B1F57633FFDA9843FD9FDBAB77C5F9C7" box="[619,726,1516,1542]" name="Brazil" pageId="9" pageNumber="10">BRAZIL</collectingCountry>
,
<collectingRegion id="0B26F841FFDA9843FD10DBAB7685F9C7" box="[740,918,1516,1542]" country="Brazil" name="Rio de Janeiro" pageId="9" pageNumber="10">Rio de Janeiro</collectingRegion>
:
</emphasis>
<location id="CC3D6078FFDA9843FC56DBAB7157F9C7" LSID="urn:lsid:plazi:treatment:414B87B5FFDA9841FF63DB1D77FDFC9C:CC3D6078FFDA9843FC56DBAB7157F9C7" box="[930,1092,1516,1542]" country="Brazil" name="Rio de Janeiro" pageId="9" pageNumber="10" stateProvince="Rio de Janeiro">Rio de Janeiro</location>
(
<collectionCode id="AFF3AE66FFDA9843FBA7DBAB71BEF9C7" box="[1107,1197,1516,1542]" country="USA" httpUri="http://grbio.org/cool/daw7-8h44" name="The Cleveland Museum of Natural History" pageId="9" pageNumber="10" type="Museum">CMNH</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="81F86528FFDA9843FF33D8567489F9B5" pageId="9" pageNumber="10" type="description">
<paragraph id="C95D36A3FFDA9843FF33D8567489F9B5" blockId="9.[151,1437,1479,2017]" pageId="9" pageNumber="10">
<emphasis id="FB96EAB1FFDA9843FF33D8567465F9EA" bold="true" box="[199,374,1553,1579]" pageId="9" pageNumber="10">Measurements</emphasis>
(n= 1). Total length: 16.4; head length: 1.4; head width: 3.2; pronotum length: 3.8; pronotum width: 11.3; scutellum length: 8.2; scutellum width: 6.6; abdominal width: 10.1; length antennomeres: I: 0.9; II: 1.3; III: 1.6; IV: 3.5; V: 4.0.
</paragraph>
</subSubSection>
<subSubSection id="81F86528FFDA9841FF33D839768AFDFC" lastPageId="11" lastPageNumber="12" pageId="9" pageNumber="10" type="diagnosis">
<paragraph id="C95D36A3FFDA9841FF33D839768AFDFC" blockId="9.[151,1437,1479,2017]" lastBlockId="11.[151,1437,151,861]" lastPageId="11" lastPageNumber="12" pageId="9" pageNumber="10">
<emphasis id="FB96EAB1FFDA9843FF33D8397451F959" bold="true" box="[199,322,1662,1688]" pageId="9" pageNumber="10">Diagnosis.</emphasis>
Large (
<quantity id="0E1A9B46FFDA9843FE6ED8397717F959" box="[410,516,1662,1688]" metricMagnitude="-2" metricUnit="m" metricValue="1.64" pageId="9" pageNumber="10" unit="mm" value="16.4">16.4 mm</quantity>
). Dorsal body surface green (
<figureCitation id="51D92A26FFDA9843FCA6D83976D2F959" box="[850,961,1662,1689]" captionStart="FIGURE 54" captionStartId="107.[152,255,1943,1967]" captionTargetBox="[250,1338,187,1913]" captionTargetId="figure-20@107.[244,1344,181,1919]" captionTargetPageId="107" captionText="FIGURE 54. AF. Dorsal and ventral surface: AB, Edessa (E.) accola sp. n.; CD, Edessa (E.) amapaensis sp. n.—yellow lines indicate where the length (L) and width (W) measurements of the humeral angles were taken; EF, Edessa (E.) amaurata sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146457" httpUri="https://zenodo.org/record/10146457/files/figure.png" pageId="9" pageNumber="10">Fig. 54 A</figureCitation>
). Ventral surface dark reddish brown with transversal smudged reddish band on thorax and abdomen (
<figureCitation id="51D92A26FFDA9843FCEBD8E57694F97C" box="[799,903,1698,1725]" captionStart="FIGURE 54" captionStartId="107.[152,255,1943,1967]" captionTargetBox="[250,1338,187,1913]" captionTargetId="figure-20@107.[244,1344,181,1919]" captionTargetPageId="107" captionText="FIGURE 54. AF. Dorsal and ventral surface: AB, Edessa (E.) accola sp. n.; CD, Edessa (E.) amapaensis sp. n.—yellow lines indicate where the length (L) and width (W) measurements of the humeral angles were taken; EF, Edessa (E.) amaurata sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146457" httpUri="https://zenodo.org/record/10146457/files/figure.png" pageId="9" pageNumber="10">Fig. 54 B</figureCitation>
). Antennae brown (
<figureCitation id="51D92A26FFDA9843FBA8D8E571F4F97C" box="[1116,1255,1698,1725]" captionStart="FIGURE 54" captionStartId="107.[152,255,1943,1967]" captionTargetBox="[250,1338,187,1913]" captionTargetId="figure-20@107.[244,1344,181,1919]" captionTargetPageId="107" captionText="FIGURE 54. AF. Dorsal and ventral surface: AB, Edessa (E.) accola sp. n.; CD, Edessa (E.) amapaensis sp. n.—yellow lines indicate where the length (L) and width (W) measurements of the humeral angles were taken; EF, Edessa (E.) amaurata sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146457" httpUri="https://zenodo.org/record/10146457/files/figure.png" pageId="9" pageNumber="10">Fig. 54AB</figureCitation>
). Pronotum with concolorous with the surrounding surface punctures (
<figureCitation id="51D92A26FFDA9843FD18D8807649F920" box="[748,858,1735,1762]" captionStart="FIGURE 54" captionStartId="107.[152,255,1943,1967]" captionTargetBox="[250,1338,187,1913]" captionTargetId="figure-20@107.[244,1344,181,1919]" captionTargetPageId="107" captionText="FIGURE 54. AF. Dorsal and ventral surface: AB, Edessa (E.) accola sp. n.; CD, Edessa (E.) amapaensis sp. n.—yellow lines indicate where the length (L) and width (W) measurements of the humeral angles were taken; EF, Edessa (E.) amaurata sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146457" httpUri="https://zenodo.org/record/10146457/files/figure.png" pageId="9" pageNumber="10">Fig. 54 A</figureCitation>
). Humeral angles short (1.6 times wider than long); apex with light brown spot restricted to the angles in dorsal view, ventrally without spot; slightly bent posteriorly (
<figureCitation id="51D92A26FFDA9843FF6BD957743EF8EB" box="[159,301,1808,1835]" captionStart="FIGURE 54" captionStartId="107.[152,255,1943,1967]" captionTargetBox="[250,1338,187,1913]" captionTargetId="figure-20@107.[244,1344,181,1919]" captionTargetPageId="107" captionText="FIGURE 54. AF. Dorsal and ventral surface: AB, Edessa (E.) accola sp. n.; CD, Edessa (E.) amapaensis sp. n.—yellow lines indicate where the length (L) and width (W) measurements of the humeral angles were taken; EF, Edessa (E.) amaurata sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146457" httpUri="https://zenodo.org/record/10146457/files/figure.png" pageId="9" pageNumber="10">Fig. 54 AB</figureCitation>
). Scutellum with concolorous with the surrounding surface punctures; apex not reaching the end of the coria (
<figureCitation id="51D92A26FFDA9843FF2BD972745DF88E" box="[223,334,1844,1871]" captionStart="FIGURE 54" captionStartId="107.[152,255,1943,1967]" captionTargetBox="[250,1338,187,1913]" captionTargetId="figure-20@107.[244,1344,181,1919]" captionTargetPageId="107" captionText="FIGURE 54. AF. Dorsal and ventral surface: AB, Edessa (E.) accola sp. n.; CD, Edessa (E.) amapaensis sp. n.—yellow lines indicate where the length (L) and width (W) measurements of the humeral angles were taken; EF, Edessa (E.) amaurata sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146457" httpUri="https://zenodo.org/record/10146457/files/figure.png" pageId="9" pageNumber="10">Fig. 54 A</figureCitation>
). Coria with all veins concolorous with the surrounding surface (
<figureCitation id="51D92A26FFDA9843FBDDD9727184F88E" box="[1065,1175,1844,1871]" captionStart="FIGURE 54" captionStartId="107.[152,255,1943,1967]" captionTargetBox="[250,1338,187,1913]" captionTargetId="figure-20@107.[244,1344,181,1919]" captionTargetPageId="107" captionText="FIGURE 54. AF. Dorsal and ventral surface: AB, Edessa (E.) accola sp. n.; CD, Edessa (E.) amapaensis sp. n.—yellow lines indicate where the length (L) and width (W) measurements of the humeral angles were taken; EF, Edessa (E.) amaurata sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146457" httpUri="https://zenodo.org/record/10146457/files/figure.png" pageId="9" pageNumber="10">Fig. 54 A</figureCitation>
). Connexival segments without spot, with a pair of concavities (
<figureCitation id="51D92A26FFDA9843FD97D91E77C0F8B2" box="[611,723,1881,1908]" captionStart="FIGURE 54" captionStartId="107.[152,255,1943,1967]" captionTargetBox="[250,1338,187,1913]" captionTargetId="figure-20@107.[244,1344,181,1919]" captionTargetPageId="107" captionText="FIGURE 54. AF. Dorsal and ventral surface: AB, Edessa (E.) accola sp. n.; CD, Edessa (E.) amapaensis sp. n.—yellow lines indicate where the length (L) and width (W) measurements of the humeral angles were taken; EF, Edessa (E.) amaurata sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146457" httpUri="https://zenodo.org/record/10146457/files/figure.png" pageId="9" pageNumber="10">Fig. 54 A</figureCitation>
). Posterolateral angles of connexivum with apices concolorous with the surrounding surface (
<figureCitation id="51D92A26FFDA9843FE1AD939774DF859" box="[494,606,1918,1945]" captionStart="FIGURE 54" captionStartId="107.[152,255,1943,1967]" captionTargetBox="[250,1338,187,1913]" captionTargetId="figure-20@107.[244,1344,181,1919]" captionTargetPageId="107" captionText="FIGURE 54. AF. Dorsal and ventral surface: AB, Edessa (E.) accola sp. n.; CD, Edessa (E.) amapaensis sp. n.—yellow lines indicate where the length (L) and width (W) measurements of the humeral angles were taken; EF, Edessa (E.) amaurata sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146457" httpUri="https://zenodo.org/record/10146457/files/figure.png" pageId="9" pageNumber="10">Fig. 54 A</figureCitation>
). Ventral surface, thorax with dark band of propleura covering 2/3 of the width of the sclerite (
<figureCitation id="51D92A26FFDA9843FE73D9E574E0F87D" box="[391,499,1954,1981]" captionStart="FIGURE 54" captionStartId="107.[152,255,1943,1967]" captionTargetBox="[250,1338,187,1913]" captionTargetId="figure-20@107.[244,1344,181,1919]" captionTargetPageId="107" captionText="FIGURE 54. AF. Dorsal and ventral surface: AB, Edessa (E.) accola sp. n.; CD, Edessa (E.) amapaensis sp. n.—yellow lines indicate where the length (L) and width (W) measurements of the humeral angles were taken; EF, Edessa (E.) amaurata sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146457" httpUri="https://zenodo.org/record/10146457/files/figure.png" pageId="9" pageNumber="10">Fig. 54 B</figureCitation>
). Evaporatorium concolorous with the surrounding surface; peritreme straight (
<figureCitation id="51D92A26FFDA9843FA9BD9E575DCF821" captionStart="FIGURE 54" captionStartId="107.[152,255,1943,1967]" captionTargetBox="[250,1338,187,1913]" captionTargetId="figure-20@107.[244,1344,181,1919]" captionTargetPageId="107" captionText="FIGURE 54. AF. Dorsal and ventral surface: AB, Edessa (E.) accola sp. n.; CD, Edessa (E.) amapaensis sp. n.—yellow lines indicate where the length (L) and width (W) measurements of the humeral angles were taken; EF, Edessa (E.) amaurata sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146457" httpUri="https://zenodo.org/record/10146457/files/figure.png" pageId="9" pageNumber="10">Fig. 54 B</figureCitation>
). Metasternal process (
<figureCitation id="51D92A26FFDA9843FE26D9817738F821" box="[466,555,1990,2016]" captionStart="FIGURE 1" captionStartId="10.[152,255,1943,1967]" captionTargetBox="[185,1408,189,1913]" captionTargetId="figure-22@10.[179,1408,181,1919]" captionTargetPageId="10" captionText="FIGURE 1. AF. Edessa (E.) accola sp. n.: AE, pygophore in dorsal, dorsolateral, ventral, posterior and posterolateral views, respectively; F, metasternal process. dr—dorsal rim; vr—ventral rim; proc—proctiger; pa—paramere; gp—superior processes of the genital cup. Scale= 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146345" httpUri="https://zenodo.org/record/10146345/files/figure.png" pageId="9" pageNumber="10">Fig. 1 F</figureCitation>
) with arms of anterior bifurcation straight and well expanded laterally at apex; anterior bifurcation broadly excavated receiving the fourth and a small part of the third rostral segment. Legs with tibiae and tarsi darker than the other articles (
<figureCitation id="51D92A26FFD89841FD6BDEFC761CFF14" box="[671,783,186,213]" captionStart="FIGURE 54" captionStartId="107.[152,255,1943,1967]" captionTargetBox="[250,1338,187,1913]" captionTargetId="figure-20@107.[244,1344,181,1919]" captionTargetPageId="107" captionText="FIGURE 54. AF. Dorsal and ventral surface: AB, Edessa (E.) accola sp. n.; CD, Edessa (E.) amapaensis sp. n.—yellow lines indicate where the length (L) and width (W) measurements of the humeral angles were taken; EF, Edessa (E.) amaurata sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146457" httpUri="https://zenodo.org/record/10146457/files/figure.png" pageId="11" pageNumber="12">Fig. 54 B</figureCitation>
). Abdomen with spine of segment III rounded (
<figureCitation id="51D92A26FFD89841FAC4DEFC709CFF14" box="[1328,1423,187,213]" captionStart="FIGURE 1" captionStartId="10.[152,255,1943,1967]" captionTargetBox="[185,1408,189,1913]" captionTargetId="figure-22@10.[179,1408,181,1919]" captionTargetPageId="10" captionText="FIGURE 1. AF. Edessa (E.) accola sp. n.: AE, pygophore in dorsal, dorsolateral, ventral, posterior and posterolateral views, respectively; F, metasternal process. dr—dorsal rim; vr—ventral rim; proc—proctiger; pa—paramere; gp—superior processes of the genital cup. Scale= 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146345" httpUri="https://zenodo.org/record/10146345/files/figure.png" pageId="11" pageNumber="12">Fig. 1 F</figureCitation>
). Intersegmental areas covered by wide smoky reddish band with smudged margins, reaching the lateral margin (
<figureCitation id="51D92A26FFD89841FF6BDF447407FEDC" box="[159,276,258,285]" captionStart="FIGURE 54" captionStartId="107.[152,255,1943,1967]" captionTargetBox="[250,1338,187,1913]" captionTargetId="figure-20@107.[244,1344,181,1919]" captionTargetPageId="107" captionText="FIGURE 54. AF. Dorsal and ventral surface: AB, Edessa (E.) accola sp. n.; CD, Edessa (E.) amapaensis sp. n.—yellow lines indicate where the length (L) and width (W) measurements of the humeral angles were taken; EF, Edessa (E.) amaurata sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146457" httpUri="https://zenodo.org/record/10146457/files/figure.png" pageId="11" pageNumber="12">Fig. 54 B</figureCitation>
). Pseudosutures smoky brown with irregular margins (
<figureCitation id="51D92A26FFD89841FC60DF44711BFEDC" box="[916,1032,258,285]" captionStart="FIGURE 54" captionStartId="107.[152,255,1943,1967]" captionTargetBox="[250,1338,187,1913]" captionTargetId="figure-20@107.[244,1344,181,1919]" captionTargetPageId="107" captionText="FIGURE 54. AF. Dorsal and ventral surface: AB, Edessa (E.) accola sp. n.; CD, Edessa (E.) amapaensis sp. n.—yellow lines indicate where the length (L) and width (W) measurements of the humeral angles were taken; EF, Edessa (E.) amaurata sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146457" httpUri="https://zenodo.org/record/10146457/files/figure.png" pageId="11" pageNumber="12">Fig. 54 B</figureCitation>
). Median longitudinal brown band almost complete and suffused (
<figureCitation id="51D92A26FFD89841FDF0DF60776AFE80" box="[516,633,294,321]" captionStart="FIGURE 54" captionStartId="107.[152,255,1943,1967]" captionTargetBox="[250,1338,187,1913]" captionTargetId="figure-20@107.[244,1344,181,1919]" captionTargetPageId="107" captionText="FIGURE 54. AF. Dorsal and ventral surface: AB, Edessa (E.) accola sp. n.; CD, Edessa (E.) amapaensis sp. n.—yellow lines indicate where the length (L) and width (W) measurements of the humeral angles were taken; EF, Edessa (E.) amaurata sp. n. Scale= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10146457" httpUri="https://zenodo.org/record/10146457/files/figure.png" pageId="11" pageNumber="12">Fig. 54 B</figureCitation>
). Trichobothria one in line with spiracle and the other laterad. Male genitalia, dorsal side of the pygophore with suffused brown area occupying more than 1/3 of the surface (
<figureCitation id="51D92A26FFD89841FA9BDF0C75F4FE48" captionStart="FIGURE 1" captionStartId="10.[152,255,1943,1967]" captionTargetBox="[185,1408,189,1913]" captionTargetId="figure-22@10.[179,1408,181,1919]" captionTargetPageId="10" captionText="FIGURE 1. AF. Edessa (E.) accola sp. n.: AE, pygophore in dorsal, dorsolateral, ventral, posterior and posterolateral views, respectively; F, metasternal process. dr—dorsal rim; vr—ventral rim; proc—proctiger; pa—paramere; gp—superior processes of the genital cup. Scale= 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146345" httpUri="https://zenodo.org/record/10146345/files/figure.png" pageId="11" pageNumber="12">Fig. 1 AB</figureCitation>
). Posterolateral angles of pygophore slightly developed (
<figureCitation id="51D92A26FFD89841FC82DF2876CFFE48" box="[886,988,367,393]" captionStart="FIGURE 1" captionStartId="10.[152,255,1943,1967]" captionTargetBox="[185,1408,189,1913]" captionTargetId="figure-22@10.[179,1408,181,1919]" captionTargetPageId="10" captionText="FIGURE 1. AF. Edessa (E.) accola sp. n.: AE, pygophore in dorsal, dorsolateral, ventral, posterior and posterolateral views, respectively; F, metasternal process. dr—dorsal rim; vr—ventral rim; proc—proctiger; pa—paramere; gp—superior processes of the genital cup. Scale= 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146345" httpUri="https://zenodo.org/record/10146345/files/figure.png" pageId="11" pageNumber="12">Fig. 1 A</figureCitation>
). Superior processes of the genital cup rectangular, narrow, grooved in posterior view, not continuing ventrally in a carina (
<figureCitation id="51D92A26FFD89841FBCEDFD471C9FE6C" box="[1082,1242,403,429]" captionStart="FIGURE 1" captionStartId="10.[152,255,1943,1967]" captionTargetBox="[185,1408,189,1913]" captionTargetId="figure-22@10.[179,1408,181,1919]" captionTargetPageId="10" captionText="FIGURE 1. AF. Edessa (E.) accola sp. n.: AE, pygophore in dorsal, dorsolateral, ventral, posterior and posterolateral views, respectively; F, metasternal process. dr—dorsal rim; vr—ventral rim; proc—proctiger; pa—paramere; gp—superior processes of the genital cup. Scale= 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146345" httpUri="https://zenodo.org/record/10146345/files/figure.png" pageId="11" pageNumber="12">Fig. 1 B, DE</figureCitation>
). Parameres (
<figureCitation id="51D92A26FFD89841FA9BDFD475F2FE10" captionStart="FIGURE 1" captionStartId="10.[152,255,1943,1967]" captionTargetBox="[185,1408,189,1913]" captionTargetId="figure-22@10.[179,1408,181,1919]" captionTargetPageId="10" captionText="FIGURE 1. AF. Edessa (E.) accola sp. n.: AE, pygophore in dorsal, dorsolateral, ventral, posterior and posterolateral views, respectively; F, metasternal process. dr—dorsal rim; vr—ventral rim; proc—proctiger; pa—paramere; gp—superior processes of the genital cup. Scale= 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146345" httpUri="https://zenodo.org/record/10146345/files/figure.png" pageId="11" pageNumber="12">Fig. 1 DE</figureCitation>
) concolorous with the surrounding surface; anterior lobe subtriangular and somewhat developed; dorsal lobe rounded, developed and curved; posterior lobe subtriangular and developed. Proctiger, posterior face subelliptical (
<figureCitation id="51D92A26FFD89841FF6BDFB8740CFDD8" box="[159,287,511,537]" captionStart="FIGURE 1" captionStartId="10.[152,255,1943,1967]" captionTargetBox="[185,1408,189,1913]" captionTargetId="figure-22@10.[179,1408,181,1919]" captionTargetPageId="10" captionText="FIGURE 1. AF. Edessa (E.) accola sp. n.: AE, pygophore in dorsal, dorsolateral, ventral, posterior and posterolateral views, respectively; F, metasternal process. dr—dorsal rim; vr—ventral rim; proc—proctiger; pa—paramere; gp—superior processes of the genital cup. Scale= 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146345" httpUri="https://zenodo.org/record/10146345/files/figure.png" pageId="11" pageNumber="12">Fig. 1 DE</figureCitation>
). Ventral rim setose, with medial tuft (
<figureCitation id="51D92A26FFD89841FD31DFB87630FDD8" box="[709,803,511,537]" captionStart="FIGURE 1" captionStartId="10.[152,255,1943,1967]" captionTargetBox="[185,1408,189,1913]" captionTargetId="figure-22@10.[179,1408,181,1919]" captionTargetPageId="10" captionText="FIGURE 1. AF. Edessa (E.) accola sp. n.: AE, pygophore in dorsal, dorsolateral, ventral, posterior and posterolateral views, respectively; F, metasternal process. dr—dorsal rim; vr—ventral rim; proc—proctiger; pa—paramere; gp—superior processes of the genital cup. Scale= 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146345" httpUri="https://zenodo.org/record/10146345/files/figure.png" pageId="11" pageNumber="12">Fig. 1 C</figureCitation>
); with expansions little developed, rounded, concolorous with the surrounding surface (
<figureCitation id="51D92A26FFD89841FE1CDC647754FDFC" box="[488,583,547,573]" captionStart="FIGURE 1" captionStartId="10.[152,255,1943,1967]" captionTargetBox="[185,1408,189,1913]" captionTargetId="figure-22@10.[179,1408,181,1919]" captionTargetPageId="10" captionText="FIGURE 1. AF. Edessa (E.) accola sp. n.: AE, pygophore in dorsal, dorsolateral, ventral, posterior and posterolateral views, respectively; F, metasternal process. dr—dorsal rim; vr—ventral rim; proc—proctiger; pa—paramere; gp—superior processes of the genital cup. Scale= 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146345" httpUri="https://zenodo.org/record/10146345/files/figure.png" pageId="11" pageNumber="12">Fig. 1 C</figureCitation>
). Female genitalia. Unknown.
</paragraph>
</subSubSection>
<caption id="9D9D662BFFD99840FF6CD9D074F1F836" ID-DOI="http://doi.org/10.5281/zenodo.10146345" ID-Zenodo-Dep="10146345" httpUri="https://zenodo.org/record/10146345/files/figure.png" pageId="10" pageNumber="11" startId="10.[152,255,1943,1967]" targetBox="[185,1408,189,1913]" targetPageId="10" targetType="figure">
<paragraph id="C95D36A3FFD99840FF6CD9D074F1F836" blockId="10.[151,1436,1943,2040]" pageId="10" pageNumber="11">
<emphasis id="FB96EAB1FFD99840FF6CD9D0740BF86E" bold="true" box="[152,280,1943,1967]" pageId="10" pageNumber="11">FIGURE 1.</emphasis>
AF.
<taxonomicName id="0EE24D20FFD99840FEA7D9DF7702F871" authority="Mendonça &amp; Silva &amp; Fernandes, 2023" authorityName="Mendonça &amp; Silva &amp; Fernandes" authorityYear="2023" box="[339,529,1944,1968]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="accola" status="sp. nov." subGenus="Edessa">
<emphasis id="FB96EAB1FFD99840FEA7D9DF7702F871" box="[339,529,1944,1968]" italics="true" pageId="10" pageNumber="11">Edessa (E.) accola</emphasis>
</taxonomicName>
<emphasis id="FB96EAB1FFD99840FDE3D9DF775CF871" bold="true" box="[535,591,1944,1968]" pageId="10" pageNumber="11">
<taxonomicNameLabel id="E0A557CAFFD99840FDE3D9DF775CF871" box="[535,591,1944,1968]" pageId="10" pageNumber="11" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
: AE, pygophore in dorsal, dorsolateral, ventral, posterior and posterolateral views, respectively; F, metasternal process. dr
<emphasis id="FB96EAB1FFD99840FDD0D9FB772EF815" bold="true" box="[548,573,1980,2004]" pageId="10" pageNumber="11"></emphasis>
dorsal rim; vr
<emphasis id="FB96EAB1FFD99840FD3DD9FB77F1F815" bold="true" box="[713,738,1980,2004]" pageId="10" pageNumber="11"></emphasis>
ventral rim; proc
<emphasis id="FB96EAB1FFD99840FC7AD9FB76B4F815" bold="true" box="[910,935,1980,2004]" pageId="10" pageNumber="11"></emphasis>
proctiger; pa
<emphasis id="FB96EAB1FFD99840FBDDD9FB7151F815" bold="true" box="[1065,1090,1980,2004]" pageId="10" pageNumber="11"></emphasis>
paramere; gp
<emphasis id="FB96EAB1FFD99840FB3CD9FB71F2F815" bold="true" box="[1224,1249,1980,2004]" pageId="10" pageNumber="11"></emphasis>
superior processes of the genital cup. Scale= 1 mm.
</paragraph>
</caption>
<subSubSection id="81F86528FFD89841FF33DC0171AAFCF8" pageId="11" pageNumber="12" type="discussion">
<paragraph id="C95D36A3FFD89841FF33DC0171AAFCF8" blockId="11.[151,1437,151,861]" pageId="11" pageNumber="12">
<emphasis id="FB96EAB1FFD89841FF33DC01745EFDA1" bold="true" box="[199,333,582,608]" pageId="11" pageNumber="12">Comments.</emphasis>
<taxonomicName id="0EE24D20FFD89841FEA6DC00770FFDA0" authority="Mendonça &amp; Silva &amp; Fernandes, 2023" authorityName="Mendonça &amp; Silva &amp; Fernandes" authorityYear="2023" box="[338,540,583,609]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="accola" status="sp. nov." subGenus="Edessa">
<emphasis id="FB96EAB1FFD89841FEA6DC00770FFDA0" box="[338,540,583,609]" italics="true" pageId="11" pageNumber="12">Edessa (E.) accola</emphasis>
</taxonomicName>
<emphasis id="FB96EAB1FFD89841FDD5DC00774EFDA0" bold="true" box="[545,605,583,609]" pageId="11" pageNumber="12">
<taxonomicNameLabel id="E0A557CAFFD89841FDD5DC00774EFDA0" box="[545,605,583,609]" pageId="11" pageNumber="12" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
resembles
<taxonomicName id="0EE24D20FFD89841FD21DC0076F2FDA0" authority="Stal, 1872" authorityName="Stal" authorityYear="1872" box="[725,993,582,609]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="fallenii" subGenus="Edessa">
<emphasis id="FB96EAB1FFD89841FD21DC00767CFDA0" box="[725,879,583,609]" italics="true" pageId="11" pageNumber="12">E. (E.) fallenii</emphasis>
<bibRefCitation id="AD734B52FFD89841FC87DC0176F2FDA0" author="Stal, C." box="[883,993,582,609]" pageId="11" pageNumber="12" pagination="1 - 159" refId="ref75136" refString="Stal, C. (1872) Enumeratio Hemipterorum. Bildrag till em f ˆ rteckning ˆ fver alla hittilis kanda Hemiptera, Jemte systematiska meddelanden. 2. Kongliga Svenska Vetenskaps - Akademiens Handlingar, Neue Folge, 10 (4), 1 - 159. https: // doi. org / 10.5962 / bhl. title. 12549" type="journal article" year="1872">Stål, 1872</bibRefCitation>
</taxonomicName>
(see
<bibRefCitation id="AD734B52FFD89841FBE3DC007018FDA0" author="Mendonca, M. T. S. &amp; Silva, V. J. &amp; Fernandes, J. A. M." box="[1047,1291,583,609]" pageId="11" pageNumber="12" pagination="1 - 63" refId="ref74748" refString="Mendonca, M. T. S., Silva, V. J. &amp; Fernandes, J. A. M. (2023) Diagnose of the nominal subgenus of Edessa and description of the E. sexdens group (Heteroptera, Pentatomidae, Edessinae). Zootaxa, 5240 (1), 1 - 63. https: // doi. org / 10.11646 / zootaxa. 5240.1.1" type="journal article" year="2023">
Mendonça
<emphasis id="FB96EAB1FFD89841FB64DC0071D6FDA0" box="[1168,1221,583,609]" italics="true" pageId="11" pageNumber="12">et al.</emphasis>
, 2023
</bibRefCitation>
,
<figureCitation id="51D92A26FFD89841FAE0DC007085FDA0" box="[1300,1430,583,609]" captionStart="FIGURE 6" captionStartId="20.[152,255,1943,1967]" captionTargetBox="[185,1402,191,1913]" captionTargetId="figure-22@20.[185,1402,181,1919]" captionTargetPageId="20" captionText="FIGURE 6. AG. Edessa (E.) brevihumeralis sp. n.:AE, pygophore in dorsal, dorsolateral, ventral, posterior and posterolateral views, respectively; F, genital plates; G, metasternal process. Scale= 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146358" httpUri="https://zenodo.org/record/10146358/files/figure.png" pageId="11" pageNumber="12">Figs. 6AG</figureCitation>
,
<figureCitation id="51D92A26FFD89841FF63DC2D75FBFD44" box="[151,232,618,645]" captionStart="FIGURE 25" captionStartId="55.[152,255,1943,1967]" captionTargetBox="[193,1394,187,1913]" captionTargetId="figure-20@55.[193,1394,181,1919]" captionTargetPageId="55" captionText="FIGURE 25. AG. Edessa (E.) immaculata sp. n.: AE, pygophore in dorsal, dorsolateral, ventral, posterior and posterolateral views, respectively; F, genital plates; G, metasternal process. Scale= 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146397" httpUri="https://zenodo.org/record/10146397/files/figure.png" pageId="11" pageNumber="12">25 EF</figureCitation>
) as it has the humeral angles light brown or concolorous with the pronotal surface; segments of the connexivum without spots and pygophore with the posterior face of the proctiger elliptical. On the other hand, both species can be easily separated:
<taxonomicName id="0EE24D20FFD89841FEACDCF474E7FD0C" authority="Mendonça &amp; Silva &amp; Fernandes, 2023" authorityName="Mendonça &amp; Silva &amp; Fernandes" authorityYear="2023" box="[344,500,691,717]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="accola" status="sp. n." subGenus="Edessa">
<emphasis id="FB96EAB1FFD89841FEACDCF474E7FD0C" box="[344,500,691,717]" italics="true" pageId="11" pageNumber="12">E. (E.) accola</emphasis>
</taxonomicName>
<emphasis id="FB96EAB1FFD89841FE09DCF4772FFD0C" bold="true" box="[509,572,691,717]" pageId="11" pageNumber="12">
<taxonomicNameLabel id="E0A557CAFFD89841FE09DCF4772FFD0C" box="[509,572,691,717]" pageId="11" pageNumber="12" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
has the connexivum with concavities on each segment (connexivum without concavities in
<taxonomicName id="0EE24D20FFD89841FEC1DC9074C1FD30" authorityName="Stal" authorityYear="1872" box="[309,466,727,753]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="fallenii" subGenus="Edessa">
<emphasis id="FB96EAB1FFD89841FEC1DC9074C1FD30" box="[309,466,727,753]" italics="true" pageId="11" pageNumber="12">E. (E.) fallenii</emphasis>
</taxonomicName>
); abdomen ventrally with intersegmental areas smoky reddish and pseudosutures smoky brown (intersegmental areas and pseudosutures black in
<taxonomicName id="0EE24D20FFD89841FCF5DCBC768EFCD4" authorityName="Stal" authorityYear="1872" box="[769,925,763,789]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="fallenii" subGenus="Edessa">
<emphasis id="FB96EAB1FFD89841FCF5DCBC768EFCD4" box="[769,925,763,789]" italics="true" pageId="11" pageNumber="12">E. (E.) fallenii</emphasis>
</taxonomicName>
) and pygophore with the superior processes of the genital cup without carina (superior processes of the genital cup carinated in
<taxonomicName id="0EE24D20FFD89841FBF8DD5871BAFCF8" authorityName="Stal" authorityYear="1872" box="[1036,1193,799,825]" class="Insecta" family="Pentatomidae" genus="Edessa" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="fallenii" subGenus="Edessa">
<emphasis id="FB96EAB1FFD89841FBF8DD5871BAFCF8" box="[1036,1193,799,825]" italics="true" pageId="11" pageNumber="12">E. (E.) fallenii</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="81F86528FFD89841FF33DD0577FDFC9C" box="[199,750,834,861]" pageId="11" pageNumber="12" type="distribution">
<paragraph id="C95D36A3FFD89841FF33DD0577FDFC9C" blockId="11.[151,1437,151,861]" box="[199,750,834,861]" pageId="11" pageNumber="12">
<emphasis id="FB96EAB1FFD89841FF33DD05744AFC9D" bold="true" box="[199,345,834,860]" pageId="11" pageNumber="12">Distribution</emphasis>
(
<figureCitation id="51D92A26FFD89841FE9DDD0474A9FC9C" box="[361,442,835,861]" captionStart="FIGURE 72" captionStartId="125.[152,255,1071,1095]" captionTargetBox="[195,1391,181,1040]" captionTargetId="figure-20@125.[195,1391,181,1050]" captionTargetPageId="125" captionText="FIGURE 72. Distribution map of the species." figureDoi="http://doi.org/10.5281/zenodo.10146495" httpUri="https://zenodo.org/record/10146495/files/figure.png" pageId="11" pageNumber="12">Fig. 72</figureCitation>
).
<collectingCountry id="B1F57633FFD89841FE24DD047729FC9C" box="[464,570,835,861]" name="Brazil" pageId="11" pageNumber="12">BRAZIL</collectingCountry>
:
<collectingRegion id="0B26F841FFD89841FDB0DD0477F9FC9C" box="[580,746,835,861]" country="Brazil" name="Rio de Janeiro" pageId="11" pageNumber="12">Rio de Janeiro</collectingRegion>
.
</paragraph>
</subSubSection>
</treatment>
</document>